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<mods:title id="D58919214DAA294D6EDEE0F33FF12DF3">Description of a new long-tailed skink (Scincidae: Trachylepis) from Angola and the Democratic Republic of the Congo</mods:title>
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<emphasis id="20BBCD532513320F84CC5BDBFE47FF4F" bold="true" box="[151,503,221,247]" italics="true" pageId="8" pageNumber="59">Trachylepis raymondlaurenti</emphasis>
Marques, Ceríaco, Bandeira, Pauwels &amp; Bauer
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(
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,
<figureCitation id="8AF40DC42513320F84A15BF9FEB9FEA0" box="[250,265,255,280]" captionStart="FIGURE 6" captionStartId="10.[151,250,1947,1969]" captionTargetBox="[173,1415,346,1928]" captionTargetId="figure@10.[165,1425,337,1937]" captionTargetPageId="10" captionText="FIGURE 6. Holotype of Trachylepis raymondlaurenti sp. nov. (CAS 258401) from Cangandala National Park, Angola in life. Photo by Luis M. P. Ceríaco." httpUri="https://zenodo.org/record/2599242/files/figure.png" pageId="8" pageNumber="59">6</figureCitation>
)
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<taxonomicName id="D5CF6AC22513320F84CC5A4EFD99FEE6" authority="(de Witte 1953: 107)" authorityName="de Witte" authorityPageNumber="107" authorityYear="1953" baseAuthorityName="de Witte" baseAuthorityPageNumber="107" baseAuthorityYear="1953" box="[151,553,328,350]" class="Reptilia" family="Scincidae" genus="Mabuya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532513320F84CC5A4EFEE1FEE6" box="[151,337,328,350]" italics="true" pageId="8" pageNumber="59">Mabuya megalura</emphasis>
(de
<treatmentCitation id="936E37502513320F85255A4EFD92FEE6" author="de Witte, G. - F." box="[382,546,328,350]" page="107" pageId="8" pageNumber="59" year="1953">
<bibRefCitation id="765E6CB02513320F85255A4EFE41FEE6" author="de Witte, G. - F." box="[382,497,328,350]" pageId="8" pageNumber="59" refId="ref7347" refString="de Witte, G. - F. (1953) Exploration du Parc National de l'Upemba. Fol. 6. Reptiles. Institut des Parcs Nationaux du Congo Belge, Brussels, 322 pp." type="book" year="1953">Witte 1953</bibRefCitation>
: 107
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)
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<emphasis id="20BBCD532513320F84CC5A60FEE1FEC4" box="[151,337,358,380]" italics="true" pageId="8" pageNumber="59">Mabuya megalura</emphasis>
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subsp. (
<treatmentCitation id="936E37502513320F85FF5A60FDE5FEC4" author="Laurent, R." box="[420,597,358,380]" page="74" pageId="8" pageNumber="59" year="1964">
<bibRefCitation id="765E6CB02513320F85FF5A60FDE5FEC4" author="Laurent, R." box="[420,597,358,380]" pageId="8" pageNumber="59" pagination="1 - 165" refId="ref8224" refString="Laurent, R. (1964) Reptiles et amphibiens de l'Angola (troisieme contribution). Publicacoes Culturais. Companhia de Diamantes de Angola, 67, 1 - 165." type="journal article" year="1964">Laurent 1964: 74</bibRefCitation>
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)
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<taxonomicName id="D5CF6AC22513320F84CC5A83FEC4FE23" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[151,372,389,411]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532513320F84CC5A83FEC4FE23" box="[151,372,389,411]" italics="true" pageId="8" pageNumber="59">Trachylepis megalura</emphasis>
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(
<bibRefCitation id="765E6CB02513320F85DA5A83FCBDFE23" author="Broadley, D. G. &amp; Cotterill, F. P. D." box="[385,781,389,411]" pageId="8" pageNumber="59" pagination="35 - 61" refId="ref6377" refString="Broadley, D. G. &amp; Cotterill, F. P. D. (2004) The reptiles of southeast Katanga, an overlooked ' hot spot'. African Journal of Herpetology, 53 (1), 35 - 61. https: // doi. org / 10.1080 / 21564574.2004.9635497" type="journal article" year="2004">
Broadley &amp; Cotterill 2004: 42 [
<emphasis id="20BBCD532513320F86985A80FCB7FE23" box="[707,775,390,411]" italics="true" pageId="8" pageNumber="59">partim</emphasis>
]
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<treatmentCitationGroup id="32DF366F2513320F84CC5AA2FB9BFE02" box="[151,1067,420,442]" pageId="8" pageNumber="59">
<taxonomicName id="D5CF6AC22513320F84CC5AA2FE25FE02" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[151,405,420,442]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532513320F84CC5AA2FEBDFE02" box="[151,269,420,442]" italics="true" pageId="8" pageNumber="59">Trachylepis</emphasis>
cf.
<emphasis id="20BBCD532513320F856F5AA2FE25FE02" box="[308,405,420,442]" italics="true" pageId="8" pageNumber="59">megalura</emphasis>
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(
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<bibRefCitation id="765E6CB02513320F85F95AA2FDC5FE02" author="Ceriaco, L. M. P. &amp; Marques, M. P. &amp; Bandeira, S. A." box="[418,629,420,442]" pageId="8" pageNumber="59" refId="ref6649" refString="Ceriaco, L. M. P., Marques, M. P. &amp; Bandeira, S. A. (2016 b) Anfibios e Repteis do Parque Nacional da Cangandala. Instituto Nacional da Biodiversidade e Areas de Conservacao, Angola &amp; Museu Nacional de Historia Nacional e da Ciencia da Universidade de Lisboa, Portugal, 97 pp." type="book" year="2016">
Ceríaco
<emphasis id="20BBCD532513320F85A35AA3FD9CFE02" box="[504,556,420,442]" italics="true" pageId="8" pageNumber="59">et al.</emphasis>
2016b
</bibRefCitation>
: 71
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; 2018b: 423;
<treatmentCitation id="936E37502513320F877E5AA3FB93FE02" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[805,1059,420,442]" page="264" pageId="8" pageNumber="59" year="2018">
<bibRefCitation id="765E6CB02513320F877E5AA3FC42FE02" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[805,1010,420,442]" pageId="8" pageNumber="59" pagination="1 - 501" refId="ref8373" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola-Atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (2), 1 - 501." type="journal article" year="2018">
<collectingRegion id="D00BDFA32513320F877E5AA3FCCEFE02" box="[805,894,421,442]" country="Italy" name="Marche" pageId="8" pageNumber="59">Marques</collectingRegion>
<emphasis id="20BBCD532513320F87DF5AA3FC07FE02" box="[900,951,420,442]" italics="true" pageId="8" pageNumber="59">et al.</emphasis>
2018
</bibRefCitation>
: 264
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)
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</paragraph>
</subSubSection>
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<materialsCitation id="A2A71B1C2513320F84CC5AE0FAE9FDFF" ID-GBIF-Occurrence="2013824459" collectingDate="2015-09-17" collectionCode="CAS" country="Angola" county="Jens Vindum" elevation="1101" latitude="9.84606" location="Giant Sable Sanctuary of Cangandala National Park" longLatPrecision="5" longitude="16.7223" municipality="Edward Stanley" pageId="8" pageNumber="59" specimenCode="CAS 258401" specimenCount="1" stateProvince="Malanje" typeStatus="holotype">
<emphasis id="20BBCD532513320F84CC5AE0FEBBFE47" bold="true" box="[151,267,486,511]" pageId="8" pageNumber="59">
<typeStatus id="CD74AFE32513320F84CC5AE0FEB6FE47" box="[151,262,486,511]" pageId="8" pageNumber="59" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="4269B93A2513320F85485AE1FE16FE47" box="[275,422,486,511]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="8" pageNumber="59" type="Museum">CAS 258401</specimenCode>
(
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,
<figureCitation id="8AF40DC42513320F864E5AE0FD95FE47" box="[533,549,486,511]" captionStart="FIGURE 6" captionStartId="10.[151,250,1947,1969]" captionTargetBox="[173,1415,346,1928]" captionTargetId="figure@10.[165,1425,337,1937]" captionTargetPageId="10" captionText="FIGURE 6. Holotype of Trachylepis raymondlaurenti sp. nov. (CAS 258401) from Cangandala National Park, Angola in life. Photo by Luis M. P. Ceríaco." httpUri="https://zenodo.org/record/2599242/files/figure.png" pageId="8" pageNumber="59">6</figureCitation>
), adult female collected in the
<location id="1710479A2513320F87D25AE0FF7BFD9B" LSID="urn:lsid:plazi:treatment:9A66A0572513320C84CC5BDBFB4FFD84:1710479A2513320F87D25AE0FF7BFD9B" country="Angola" county="Jens Vindum" latitude="9.84606" longLatPrecision="5" longitude="16.7223" municipality="Edward Stanley" name="Giant Sable Sanctuary of Cangandala National Park" pageId="8" pageNumber="59" stateProvince="Malanje">Giant Sable Sanctuary of Cangandala National Park</location>
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<geoCoordinate id="77FB77862513320F8534590CFE5DFD9A" box="[367,493,522,547]" direction="east" orientation="longitude" pageId="8" pageNumber="59" precision="5" value="16.7223">16.7223° E</geoCoordinate>
, WGS-84;
<elevation id="99E2F6722513320F8631590CFC6DFD9B" box="[618,989,522,547]" metricMagnitude="3" metricUnit="m" metricValue="1.101" pageId="8" pageNumber="59" unit="m" value="1101.0">
elevation
<quantity id="D537BCA42513320F868C590DFC9BFD9A" box="[727,811,523,547]" metricMagnitude="3" metricUnit="m" metricValue="1.101" pageId="8" pageNumber="59" unit="m" value="1101.0">1101 m</quantity>
above sea level
</elevation>
) (
<figureCitation id="8AF40DC42513320F87AF590CFB88FD9B" box="[1012,1080,522,547]" captionStart="FIGURE 4" captionStartId="6.[151,250,1979,2001]" captionTargetBox="[218,1369,1114,1956]" captionTargetId="figure@6.[210,1379,1104,1966]" captionTargetPageId="6" captionText="FIGURE 4. Distribution map of Trachylepis raymondlaurenti sp. nov. (blue squares) and T. megalura (yellow dots) in sub- Saharan Africa. For distributional data see Appendices I and II." httpUri="https://zenodo.org/record/2599236/files/figure.png" pageId="8" pageNumber="59">Fig. 4</figureCitation>
),
<collectingRegion id="D00BDFA32513320F8016590CFAA5FD9B" box="[1101,1301,522,547]" country="Angola" name="Malanje" pageId="8" pageNumber="59">Malanje Province</collectingRegion>
,
<collectingCountry id="6AD851D12513320F817B590CFAC5FD9B" box="[1312,1397,522,547]" name="Angola" pageId="8" pageNumber="59">Angola</collectingCountry>
, by
<location id="1710479A2513320F84CC5928FED0FDFE" LSID="urn:lsid:plazi:treatment:9A66A0572513320C84CC5BDBFB4FFD84:1710479A2513320F84CC5928FED0FDFE" box="[151,352,558,583]" country="Angola" county="Jens Vindum" latitude="9.84606" longLatPrecision="5" longitude="16.7223" municipality="Edward Stanley" name="Mariana Marques" pageId="8" pageNumber="59" stateProvince="Malanje">Mariana Marques</location>
,
<location id="1710479A2513320F85305928FE4EFDFF" LSID="urn:lsid:plazi:treatment:9A66A0572513320C84CC5BDBFB4FFD84:1710479A2513320F85305928FE4EFDFF" box="[363,510,558,583]" country="Angola" county="Jens Vindum" latitude="9.84606" longLatPrecision="5" longitude="16.7223" municipality="Edward Stanley" name="Luis Ceriaco" pageId="8" pageNumber="59" stateProvince="Malanje">Luis Ceríaco</location>
,
<location id="1710479A2513320F86525929FD79FDFF" LSID="urn:lsid:plazi:treatment:9A66A0572513320C84CC5BDBFB4FFD84:1710479A2513320F86525929FD79FDFF" box="[521,713,558,583]" country="Angola" county="Jens Vindum" latitude="9.84606" longLatPrecision="5" longitude="16.7223" municipality="Edward Stanley" name="Suzana Bandeira" pageId="8" pageNumber="59" stateProvince="Malanje">Suzana Bandeira</location>
,
<collectingMunicipality id="F2148B3B2513320F868F5928FC36FDFF" box="[724,902,558,583]" pageId="8" pageNumber="59">Edward Stanley</collectingMunicipality>
and
<collectingCounty id="FB1169CD2513320F87E75929FBFDFDFF" box="[956,1101,558,583]" pageId="8" pageNumber="59">Jens Vindum</collectingCounty>
on
<date id="667137812513320F802C5929FAE5FDFF" box="[1143,1365,558,583]" pageId="8" pageNumber="59" value="2015-09-17">
<collectingDate id="7635CE692513320F802C5929FAE5FDFF" box="[1143,1365,558,583]" pageId="8" pageNumber="59" value="2015-09-17">17 September 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="127011412513320F849C5954FD4CFC33" blockId="8.[151,1437,486,907]" pageId="8" pageNumber="59">
<materialsCitation id="A2A71B1C2513320F849C5954FC9AFD37" ID-GBIF-Occurrence="2013824457" collectingDate="1954-06" collectionCode="MD" country="Angola" elevation="1101" latitude="10.01667" location="Alto Cuilo" longLatPrecision="547" longitude="19.55" pageId="8" pageNumber="59" specimenCode="MD 5309" specimenCount="1" stateProvince="Lunda Sul" typeStatus="paratype">
<emphasis id="20BBCD532513320F849C5954FEF6FDD3" bold="true" box="[199,326,594,619]" pageId="8" pageNumber="59">
<typeStatus id="CD74AFE32513320F849C5954FEF1FDD3" box="[199,321,594,619]" pageId="8" pageNumber="59" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="4269B93A2513320F85165955FE0DFDD3" box="[333,445,594,619]" collectionCode="MD" country="Germany" name="Museum Donaueschingen" pageId="8" pageNumber="59">MD 5309</specimenCode>
, adult female collected in
<location id="1710479A2513320F86BC5954FCEEFDD3" LSID="urn:lsid:plazi:treatment:9A66A0572513320C84CC5BDBFB4FFD84:1710479A2513320F86BC5954FCEEFDD3" box="[743,862,594,619]" country="Angola" latitude="10.01667" longLatPrecision="547" longitude="19.55" name="Alto Cuilo" pageId="8" pageNumber="59" stateProvince="Lunda Sul">Alto Cuílo</location>
(-
<geoCoordinate id="77FB77862513320F872E5954FBB9FDD2" box="[885,1033,594,619]" direction="north" orientation="latitude" pageId="8" pageNumber="59" precision="1" value="10.01667">10.01667° N</geoCoordinate>
,
<geoCoordinate id="77FB77862513320F804E5954FBC7FDD2" box="[1045,1143,594,619]" direction="east" orientation="longitude" pageId="8" pageNumber="59" precision="555" value="19.55">19.55° E</geoCoordinate>
, WGS-84;
<elevation id="99E2F6722513320F80AC5954FF00FD37" metricMagnitude="3" metricUnit="m" metricValue="1.101" pageId="8" pageNumber="59" unit="m" value="1101.0">
elevation
<quantity id="D537BCA42513320F813E5955FF00FD37" metricMagnitude="3" metricUnit="m" metricValue="1.101" pageId="8" pageNumber="59" unit="m" value="1101.0">1101 m</quantity>
</elevation>
),
<collectingRegion id="D00BDFA32513320F849D5970FE1CFD37" box="[198,428,630,655]" country="Angola" name="Lunda Sul" pageId="8" pageNumber="59">Lunda Sul Province</collectingRegion>
,
<collectingCountry id="6AD851D12513320F85E35970FDBDFD37" box="[440,525,630,655]" name="Angola" pageId="8" pageNumber="59">Angola</collectingCountry>
, by a local in
<date id="667137812513320F86E85971FC9AFD37" box="[691,810,630,655]" pageId="8" pageNumber="59" value="1954-06">
<collectingDate id="7635CE692513320F86E85971FC9AFD37" box="[691,810,630,655]" pageId="8" pageNumber="59" value="1954-06">June 1954</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="A2A71B1C2513320F87635971FED2FD0B" ID-GBIF-Occurrence="2013824460" collectingDate="1954-06" collectionCode="MCZ" country="Angola" elevation="1101" latitude="10.01667" location="Alto Cuilo" longLatPrecision="547" longitude="19.55" pageId="8" pageNumber="59" specimenCount="1" stateProvince="Lunda Sul" typeStatus="paratype">
<collectionCode id="74DE89842513320F87635971FCC4FD37" box="[824,884,631,655]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="59" type="Museum">MCZ</collectionCode>
R-67627, adult female with the same collection data as the former
</materialsCitation>
;
<materialsCitation id="A2A71B1C2513320F8536599DFA28FD6F" ID-GBIF-Occurrence="2013824463" collectingDate="1947-03-16" collectionCode="RBINS" collectorName="Mission G. &amp; F. de Witte" country="Democratic Republic of the Congo" elevation="1810" latitude="8.933333" location="Lusinga" longLatPrecision="5488" longitude="27.2" municipality="Upemba National Park" pageId="8" pageNumber="59" specimenCode="RBINS 2691, RBINS 6998" specimenCount="1" typeStatus="paratype">
<specimenCode id="4269B93A2513320F8536599DFE4EFD0B" box="[365,510,666,691]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 2691</specimenCode>
(formerly
<specimenCode id="4269B93A2513320F862D599DFCBBFD0B" box="[630,779,666,691]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 6998</specimenCode>
), adult female collected in
<location id="1710479A2513320F806C599CFB21FD0B" LSID="urn:lsid:plazi:treatment:9A66A0572513320C84CC5BDBFB4FFD84:1710479A2513320F806C599CFB21FD0B" box="[1079,1169,666,691]" country="Democratic Republic of the Congo" latitude="8.933333" longLatPrecision="5488" longitude="27.2" municipality="Upemba National Park" name="Lusinga" pageId="8" pageNumber="59">Lusinga</location>
(-
<geoCoordinate id="77FB77862513320F80FD599DFA89FD0A" box="[1190,1337,667,691]" direction="north" orientation="latitude" pageId="8" pageNumber="59" precision="1" value="8.933333">8.933333° N</geoCoordinate>
,
<geoCoordinate id="77FB77862513320F811F599DFA27FD0A" box="[1348,1431,667,691]" direction="east" orientation="longitude" pageId="8" pageNumber="59" precision="5555" value="27.2">27.2° E</geoCoordinate>
, WGS-84;
<elevation id="99E2F6722513320F855159B8FE62FD6F" box="[266,466,702,727]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="8" pageNumber="59" unit="m" value="1810.0">
elevation
<quantity id="D537BCA42513320F852259B9FE62FD6F" box="[377,466,703,727]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="8" pageNumber="59" unit="m" value="1810.0">1810 m</quantity>
</elevation>
),
<collectingMunicipality id="F2148B3B2513320F85BC59B8FD5CFD6F" box="[487,748,702,727]" pageId="8" pageNumber="59">Upemba National Park</collectingMunicipality>
,
<collectingCountry id="6AD851D12513320F86A259B9FC85FD6F" box="[761,821,703,727]" name="Democratic Republic of the Congo" pageId="8" pageNumber="59">DRC</collectingCountry>
, during the
<collectorName id="BF3A74972513320F87E759B8FB88FD6F" box="[956,1080,702,727]" pageId="8" pageNumber="59">Mission G.</collectorName>
-
<collectorName id="BF3A74972513320F801B59B9FB0BFD6F" box="[1088,1211,702,727]" pageId="8" pageNumber="59">F. de Witte</collectorName>
on
<date id="667137812513320F80BC59B8FA28FD6F" box="[1255,1432,702,727]" pageId="8" pageNumber="59" value="1947-03-16">
<collectingDate id="7635CE692513320F80BC59B8FA28FD6F" box="[1255,1432,702,727]" pageId="8" pageNumber="59" value="1947-03-16">16 March 1947</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="A2A71B1C2513320F84CC59E5FF63FCA7" ID-GBIF-Occurrence="2013824456" collectingDate="1947-04-09" collectingDateMax="1947-04-17" collectingDateMin="1947-04-09" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="8" pageNumber="59" specimenCode="RBINS 2692, RBINS 6999" specimenCount="1" typeStatus="paratype">
<specimenCode id="4269B93A2513320F84CC59E5FE99FD43" box="[151,297,738,763]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 2692</specimenCode>
(formerly
<specimenCode id="4269B93A2513320F85FE59E5FD8CFD43" box="[421,572,738,763]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 6999</specimenCode>
), adult male with locality data as for the former but collected on
<date id="667137812513320F817A59E5FF63FCA7" pageId="8" pageNumber="59" value="1947-04-09" valueMax="1947-04-17" valueMin="1947-04-09">
<collectingDate id="7635CE692513320F817A59E5FF63FCA7" pageId="8" pageNumber="59" value="1947-04-09" valueMax="1947-04-17" valueMin="1947-04-09">917 April 1947</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="A2A71B1C2513320F84855801FEC1FCFB" ID-GBIF-Occurrence="2013824462" collectingDate="1947-06-21" collectingDateMax="1947-07-21" collectingDateMin="1947-06-21" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="8" pageNumber="59" specimenCode="RBINS 2693, RBINS 7000" specimenCount="1" typeStatus="paratype">
<specimenCode id="4269B93A2513320F84855801FEC1FCA7" box="[222,369,774,799]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 2693</specimenCode>
(formerly
<specimenCode id="4269B93A2513320F85B75801FD32FCA7" box="[492,642,775,799]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 7000</specimenCode>
), adult male with locality data as for RBINS 2691 but collected on
<date id="667137812513320F81DB5801FEC1FCFB" pageId="8" pageNumber="59" value="1947-06-21" valueMax="1947-07-21" valueMin="1947-06-21">
<collectingDate id="7635CE692513320F81DB5801FEC1FCFB" pageId="8" pageNumber="59" value="1947-06-21" valueMax="1947-07-21" valueMin="1947-06-21">21 June21 July 1947</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="A2A71B1C2513320F8524582DFDF5FCDF" ID-GBIF-Occurrence="2013824458" collectingDate="1947-12" collectingDateMax="1947-07-21" collectingDateMin="1947-06-21" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="8" pageNumber="59" specimenCode="RBINS 2694, RBINS 7001" specimenCount="1" typeStatus="paratype">
<specimenCode id="4269B93A2513320F8524582DFDA3FCFB" box="[383,531,810,835]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 2694</specimenCode>
(formerly
<specimenCode id="4269B93A2513320F86CA582DFC99FCFB" box="[657,809,811,835]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 7001</specimenCode>
), adult male with the same locality data as for
<collectionCode id="74DE89842513320F8111582DFA2DFCFB" box="[1354,1437,811,835]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS</collectionCode>
2691 but collected in
<date id="667137812513320F85D45848FDF5FCDF" box="[399,581,846,871]" pageId="8" pageNumber="59" value="1947-12">
<collectingDate id="7635CE692513320F85D45848FDF5FCDF" box="[399,581,846,871]" pageId="8" pageNumber="59" value="1947-12">December 1947</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="A2A71B1C2513320F86095849FD48FC33" ID-GBIF-Occurrence="2013824461" collectingDate="1948-01" collectingDateMax="1947-07-21" collectingDateMin="1947-06-21" collectionCode="RBINS" country="Democratic Republic of the Congo" pageId="8" pageNumber="59" specimenCode="RBINS 2695, RBINS 7002" specimenCount="1" typeStatus="paratype">
<specimenCode id="4269B93A2513320F86095849FD55FCDF" box="[594,741,846,871]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 2695</specimenCode>
(formerly
<specimenCode id="4269B93A2513320F873A5849FC48FCDF" box="[865,1016,847,871]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS 7002</specimenCode>
), adult female with the same locality data as for
<collectionCode id="74DE89842513320F85495875FED5FC33" box="[274,357,883,907]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="8" pageNumber="59" type="Museum">RBINS</collectionCode>
2691 but collected in
<date id="667137812513320F86045875FD48FC33" box="[607,760,883,907]" pageId="8" pageNumber="59" value="1948-01">
<collectingDate id="7635CE692513320F86045875FD48FC33" box="[607,760,883,907]" pageId="8" pageNumber="59" value="1948-01">January 1948</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="46B041C92513320F84CC5CCDFAF7F859" box="[151,1351,1995,2017]" httpUri="https://zenodo.org/record/2599240/files/figure.png" pageId="8" pageNumber="59" startId="8.[151,250,1995,2017]" targetBox="[156,1426,978,1952]" targetPageId="8">
<paragraph id="127011412513320F84CC5CCDFAF7F859" blockId="8.[151,1351,1995,2017]" box="[151,1351,1995,2017]" pageId="8" pageNumber="59">
<emphasis id="20BBCD532513320F84CC5CCDFEA2F859" bold="true" box="[151,274,1995,2017]" pageId="8" pageNumber="59">FIGURE 5.</emphasis>
Preserved holotype of
<taxonomicName id="D5CF6AC22513320F85A65CCDFC92F859" authority="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[509,802,1995,2017]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="59" phylum="Chordata" rank="species" species="raymondlaurenti" status="sp. nov.">
<emphasis id="20BBCD532513320F85A65CCDFC92F859" box="[509,802,1995,2017]" italics="true" pageId="8" pageNumber="59">Trachylepis raymondlaurenti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3B8870282513320F87725CCDFCCAF859" box="[809,890,1995,2017]" pageId="8" pageNumber="59" rank="species">
<emphasis id="20BBCD532513320F87725CCDFCC4F859" bold="true" box="[809,884,1995,2017]" pageId="8" pageNumber="59">sp. nov</emphasis>
.
</taxonomicNameLabel>
(CAS 258401). Photo by Luis M. P. Ceríaco.
</paragraph>
</caption>
<subSubSection id="5AD542CA2512320E849C5B91FC2AFE3F" pageId="9" pageNumber="60" type="diagnosis">
<paragraph id="127011412512320E849C5B91FC2AFE3F" blockId="9.[151,1437,151,2035]" pageId="9" pageNumber="60">
<emphasis id="20BBCD532512320E849C5B91FEF1FF08" bold="true" box="[199,321,151,176]" pageId="9" pageNumber="60">Diagnosis.</emphasis>
<taxonomicName id="D5CF6AC22512320E85165B91FD26FF08" authority="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[333,662,151,176]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti" status="sp. nov.">
<emphasis id="20BBCD532512320E85165B91FD26FF08" box="[333,662,151,176]" italics="true" pageId="9" pageNumber="60">Trachylepis raymondlaurenti</emphasis>
</taxonomicName>
<emphasis id="20BBCD532512320E86F95B91FCB0FF08" bold="true" box="[674,768,151,176]" pageId="9" pageNumber="60">
<taxonomicNameLabel id="3B8870282512320E86F95B91FCB0FF08" box="[674,768,151,176]" pageId="9" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is readily distinguished from all other congeners by the following combination of characters: (1) slender body of medium size, SVL up to
<quantity id="D537BCA42512320E80665BBAFB15FF6B" box="[1085,1189,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="7.93" pageId="9" pageNumber="60" unit="mm" value="79.3">79.3 mm</quantity>
; (2) long-tailed, up to
<quantity id="D537BCA42512320E84CC5BD9FEBDFF4F" box="[151,269,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="2.158" pageId="9" pageNumber="60" unit="mm" value="215.8">215.8 mm</quantity>
, greater than twice body length; (3) well-defined neck constriction; (4) color homogeneously greyishbrown dorsally, with some dark speckles on sides of tail; ventral surface with some scattered dark speckles near the flanks and under the tail; supralabials and infralabials dotted with white; no distinct longitudinal or transverse dorsal bands; (5) MSR 2428, SAD 4853, SAV 5058; (6) KDS 03; (7) scales on palms of hands and soles of feet smooth; (8) supranasals separated; and (9) prefrontals separated.
</paragraph>
</subSubSection>
<subSubSection id="5AD542CA2512320E849C5A94FC5BFB58" pageId="9" pageNumber="60" type="materials_examined">
<paragraph id="127011412512320E849C5A94FC5BFB58" blockId="9.[151,1437,151,2035]" pageId="9" pageNumber="60">
<emphasis id="20BBCD532512320E849C5A94FE7CFE13" bold="true" box="[199,460,402,427]" pageId="9" pageNumber="60">
<typeStatus id="CD74AFE32512320E849C5A94FE84FE13" box="[199,308,402,427]" pageId="9" pageNumber="60" type="holotype">Holotype</typeStatus>
description.
</emphasis>
Individual in good condition. Gravid female, with 10 fetuses in advanced stage of development. Habitus and relative sizes of head, body and tail comparable to
<taxonomicName id="D5CF6AC22512320E80155AB3FAFAFE76" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[1102,1354,437,462]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532512320E80155AB3FAFAFE76" box="[1102,1354,437,462]" italics="true" pageId="9" pageNumber="60">Trachylepis megalura</emphasis>
</taxonomicName>
. Body cylindrical and slender with a well-defined neck and thin limbs; tail very long, its length greater than twice the SVL, smoothly tapering. Fore- and hind limbs overlap when depressed against the body. SVL
<quantity id="D537BCA42512320E809A5AF8FA97FDAD" box="[1217,1319,510,534]" metricMagnitude="-2" metricUnit="m" metricValue="7.93" pageId="9" pageNumber="60" unit="mm" value="79.3">79.3 mm</quantity>
, TL
<quantity id="D537BCA42512320E81065AFBFF76FD81" metricMagnitude="-1" metricUnit="m" metricValue="1.523" pageId="9" pageNumber="60" unit="mm" value="152.3">152.3 mm</quantity>
. HL
<quantity id="D537BCA42512320E84A65927FED4FD81" box="[253,356,545,569]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="9" pageNumber="60" unit="mm" value="13.1">13.1 mm</quantity>
, with relatively long snout (HL/HW 65.1%). Additional measurements are presented in
<tableCitation id="5F4D24FA2512320E81195927FA28FD81" box="[1346,1432,545,570]" captionStart="TABLE 2" captionStartId="7.[147,233,558,580]" captionText="TABLE 2. Mensural (in mm) and meristic data for the holotype and paratypes of Trachylepis raymondlaurenti sp. nov., the holotype of Eumeces massaianus Fischer, 1884," pageId="9" pageNumber="60">Table 2</tableCitation>
. Ear opening small, without anterior lobes. Rostral visible from above. Small nostrils set posteriorly so that postnasal effectively borders nostril. Supranasals without contact. Frontonasal wider than long, in contact with loreal scale. Prefrontals pentagonal, separated from one another, each in contact with the following head shields: frontonasal, loreal, first and second supraocular, second supraciliary and frontal. Two loreals. Frontal longer than the distance between anterior tip of frontal and tip of snout. Frontal in contact with three supraoculars on each side. Two frontoparietals, in contact with each other and the frontal, third and fourth supraoculars, parietal and interparietal. Frontoparietal plus interparietal length greater than frontal length. Interparietal twice as long as wide, with a visible parietal foramen. Parietals of same length as frontoparietals. Parietals not in contact with one another. Five supraciliaries, second largest. Seven supralabials, fifth being subocular. Six infralabials. Postmental bordering seven scales. Transparent scale present in lower eyelid, as is usual for
<taxonomicName id="D5CF6AC22512320E87975880FBFFFC27" authorityName="Fitzinger" authorityYear="1843" box="[972,1103,902,927]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="20BBCD532512320E87975880FBFFFC27" box="[972,1103,902,927]" italics="true" pageId="9" pageNumber="60">Trachylepis</emphasis>
</taxonomicName>
. Tympanum visible, at same level as mouth. Dorsal scales each with three smooth keels, sometimes very indistinct. Ventral scales smooth. MSR 28, SAD 52, SAV 58. Limbs with five digits; scales on soles of hands and feet smooth. Relative length of fingers IV&gt;III&gt;II&gt;V&gt;I, relative length of toes IV&gt;III&gt;V&gt;II&gt;I. Finger-IV lamellae 13, Toe-IV lamellae 17. Color in life is homogenous light greenish-brown on the entire dorsum; head, neck, legs and tail uniformly light grey-brown. No visible spots, markings or dorsolateral stripes. Gular region white, remaining ventral areas yellowish, except the brown palmar regions of hand and feet. In preservative, background color of flanks and upper side of head, neck, dorsum, legs and tail homogenously brown, with some white speckles running from the labials through the anterior half of the tail. Above the labials the head is uniformly brown. Eyelids dark-brown. Ventral surfaces uniformly whitish with some scattered dark speckles near the flanks and under the tail.
</paragraph>
</subSubSection>
<subSubSection id="5AD542CA2512320E849C5FEDFCA0F84B" pageId="9" pageNumber="60" type="description">
<paragraph id="127011412512320E849C5FEDFDA2FA2B" blockId="9.[151,1437,151,2035]" pageId="9" pageNumber="60">
<emphasis id="20BBCD532512320E849C5FEDFE8CFABC" bold="true" box="[199,316,1259,1284]" pageId="9" pageNumber="60">Variation.</emphasis>
Variation in measurements and scalation of the
<typeStatus id="CD74AFE32512320E873A5FEAFC7DFABC" box="[865,973,1260,1284]" pageId="9" pageNumber="60" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="D5CF6AC22512320E87AF5FEDFA89FABC" authority="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[1012,1337,1259,1284]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti" status="sp. nov.">
<emphasis id="20BBCD532512320E87AF5FEDFA89FABC" box="[1012,1337,1259,1284]" italics="true" pageId="9" pageNumber="60">Trachylepis raymondlaurenti</emphasis>
</taxonomicName>
<emphasis id="20BBCD532512320E811A5FEDFA2CFABC" bold="true" box="[1345,1436,1259,1284]" pageId="9" pageNumber="60">
<taxonomicNameLabel id="3B8870282512320E811A5FEDFA2CFABC" box="[1345,1436,1259,1284]" pageId="9" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are reported in
<tableCitation id="5F4D24FA2512320E85165E09FE17FA90" box="[333,423,1295,1320]" captionStart="TABLE 2" captionStartId="7.[147,233,558,580]" captionText="TABLE 2. Mensural (in mm) and meristic data for the holotype and paratypes of Trachylepis raymondlaurenti sp. nov., the holotype of Eumeces massaianus Fischer, 1884," pageId="9" pageNumber="60">Table 2</tableCitation>
. Head scalation similar to
<typeStatus id="CD74AFE32512320E86BE5E09FCF7FA90" box="[741,839,1295,1320]" pageId="9" pageNumber="60" type="holotype">holotype</typeStatus>
in all
<typeStatus id="CD74AFE32512320E87C25E16FBB4FA90" box="[921,1028,1296,1320]" pageId="9" pageNumber="60" type="paratype">paratypes</typeStatus>
except for prefrontals, which vary considerably; in RBINS 2 691 and RBINS 2695 the prefrontals are separated by an additional median scale, and in RBINS 2693 they are separated. Coloration of
<typeStatus id="CD74AFE32512320E86EB5E5EFCABFAC8" box="[688,795,1368,1392]" pageId="9" pageNumber="60" type="paratype">paratypes</typeStatus>
agrees entirely with that of
<typeStatus id="CD74AFE32512320E80045E50FB75FAD7" box="[1119,1221,1366,1391]" pageId="9" pageNumber="60" type="holotype">holotype</typeStatus>
. Hatchlings have a reddish tail and hind legs (
<figureCitation id="8AF40DC42512320E859B5E7CFDB4FA2B" box="[448,516,1402,1427]" captionStart="FIGURE 7" captionStartId="11.[151,250,1750,1772]" captionTargetBox="[187,1395,679,1708]" captionTargetId="figure@11.[151,1436,628,1729]" captionTargetPageId="11" captionText="FIGURE 7. General view of five out of the ten fetuses found in the reproductive tract of the holotype of Trachylepis raymondlaurenti sp. nov. Photos by Luis M. P. Ceríaco." httpUri="https://zenodo.org/record/2599244/files/figure.png" pageId="9" pageNumber="60">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="127011412512320E849C5E98FCA0F84B" blockId="9.[151,1437,151,2035]" pageId="9" pageNumber="60">
<emphasis id="20BBCD532512320E849C5E98FD8EFA0F" bold="true" box="[199,574,1438,1463]" pageId="9" pageNumber="60">Comparison with other species</emphasis>
.
<taxonomicName id="D5CF6AC22512320E86155E98FC26FA0F" authority="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[590,918,1438,1463]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti" status="sp. nov.">
<emphasis id="20BBCD532512320E86155E98FC26FA0F" box="[590,918,1438,1463]" italics="true" pageId="9" pageNumber="60">Trachylepis raymondlaurenti</emphasis>
</taxonomicName>
<emphasis id="20BBCD532512320E87C45E98FC4CFA0F" bold="true" box="[927,1020,1438,1463]" pageId="9" pageNumber="60">
<taxonomicNameLabel id="3B8870282512320E87C45E98FC4CFA0F" box="[927,1020,1438,1463]" pageId="9" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all congeners, with the exception of
<taxonomicName id="D5CF6AC22512320E85725EC5FE01FA63" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[297,433,1474,1499]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532512320E85725EC5FE01FA63" box="[297,433,1474,1499]" italics="true" pageId="9" pageNumber="60">T. megalura</emphasis>
</taxonomicName>
and
<taxonomicName id="D5CF6AC22512320E85BC5EC5FDCCFA63" baseAuthorityName="Sternfeld" baseAuthorityYear="1911" box="[487,636,1474,1499]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="20BBCD532512320E85BC5EC5FDCCFA63" box="[487,636,1474,1499]" italics="true" pageId="9" pageNumber="60">T. boulengeri</emphasis>
</taxonomicName>
, in being a slender skink with a well-marked neck, a very long tail (TL generally more than twice the SVL), and a low number of midbody scale rows (2428) in comparison with the majority of
<taxonomicName id="D5CF6AC22512320E857B5D0FFE13F99A" authorityName="Fitzinger" authorityYear="1843" box="[288,419,1545,1570]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="20BBCD532512320E857B5D0FFE13F99A" box="[288,419,1545,1570]" italics="true" pageId="9" pageNumber="60">Trachylepis</emphasis>
</taxonomicName>
species (
<bibRefCitation id="765E6CB02512320E86555D0FFD0CF99A" author="Broadley, D. G." box="[526,700,1545,1570]" pageId="9" pageNumber="60" pagination="87 - 110" refId="ref6329" refString="Broadley, D. G. (2000) A review of the genus Mabuya in southeastern Africa (Sauria: Scincidae). African Journal of Herpetology, 49 (2), 87 - 110. https: // doi. org / 10.1080 / 21564574.2000.9635437" type="journal article" year="2000">Broadley 2000</bibRefCitation>
).
<taxonomicName id="D5CF6AC22512320E868A5D0FFBA8F99A" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[721,1048,1545,1570]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti">
<emphasis id="20BBCD532512320E868A5D0FFBA8F99A" box="[721,1048,1545,1570]" italics="true" pageId="9" pageNumber="60">Trachylepis raymondlaurenti</emphasis>
</taxonomicName>
is readily distinguishable from
<taxonomicName id="D5CF6AC22512320E81DC5D0CFEB3F9FE" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532512320E81DC5D0CFEB3F9FE" italics="true" pageId="9" pageNumber="60">T. megalura</emphasis>
</taxonomicName>
by its coloration and scalation.
<taxonomicName id="D5CF6AC22512320E86335D2BFCEEF9FE" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[616,862,1581,1606]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532512320E86335D2BFCEEF9FE" box="[616,862,1581,1606]" italics="true" pageId="9" pageNumber="60">Trachylepis megalura</emphasis>
</taxonomicName>
is characterized by a striped pattern with a number of fine dorsal longitudinal black or white stripes (grey-brown, light orange-brown or golden or red-brown) including a very characteristic prominent white stripe on the flanks, bordered above by a black stripe running from the upper lip along the entire body onto the tail, while the new species is characterized by the lack of the lateral stripe on the flanks, being homogeneously greyish-brown dorsally with scattered dark speckles. In terms of scalation, the supranasals and prefrontals are never in contact in
<taxonomicName id="D5CF6AC22512320E87375DE7FBF3F940" authority="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[876,1091,1759,1784]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti" status="sp. nov.">
<emphasis id="20BBCD532512320E87375DE7FBF3F940" box="[876,1091,1759,1784]" italics="true" pageId="9" pageNumber="60">T. raymondlaurenti</emphasis>
</taxonomicName>
<emphasis id="20BBCD532512320E80115DD9FB14F940" bold="true" box="[1098,1188,1759,1784]" pageId="9" pageNumber="60">
<taxonomicNameLabel id="3B8870282512320E80115DD9FB14F940" box="[1098,1188,1759,1784]" pageId="9" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, while in
<taxonomicName id="D5CF6AC22512320E814E5DE7FA2CF940" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[1301,1436,1759,1784]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532512320E814E5DE7FA2CF940" box="[1301,1436,1759,1784]" italics="true" pageId="9" pageNumber="60">T. megalura</emphasis>
</taxonomicName>
they are both in contact.
<taxonomicName id="D5CF6AC22512320E85F45C05FD03F8A4" baseAuthorityName="Sternfeld" baseAuthorityYear="1911" box="[431,691,1795,1820]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="20BBCD532512320E85F45C05FD03F8A4" box="[431,691,1795,1820]" italics="true" pageId="9" pageNumber="60">Trachylepis boulengeri</emphasis>
</taxonomicName>
shares more morphological similarities with the new species than with
<taxonomicName id="D5CF6AC22512320E84895C2EFEE9F8F8" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[210,345,1831,1856]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532512320E84895C2EFEE9F8F8" box="[210,345,1831,1856]" italics="true" pageId="9" pageNumber="60">T. megalura</emphasis>
</taxonomicName>
. Both
<taxonomicName id="D5CF6AC22512320E85F05C21FD00F8F8" baseAuthorityName="Sternfeld" baseAuthorityYear="1911" box="[427,688,1831,1856]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="20BBCD532512320E85F05C21FD00F8F8" box="[427,688,1831,1856]" italics="true" pageId="9" pageNumber="60">Trachylepis boulengeri</emphasis>
</taxonomicName>
and
<taxonomicName id="D5CF6AC22512320E86B05C2EFC74F8F8" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[747,964,1831,1856]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti">
<emphasis id="20BBCD532512320E86B05C2EFC74F8F8" box="[747,964,1831,1856]" italics="true" pageId="9" pageNumber="60">T. raymondlaurenti</emphasis>
</taxonomicName>
have a uniform grey-brown background, sometimes with a few scattered dark flecks, but
<taxonomicName id="D5CF6AC22512320E86F45C4AFCF4F8DC" baseAuthorityName="Sternfeld" baseAuthorityYear="1911" box="[687,836,1867,1892]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="20BBCD532512320E86F45C4AFCF4F8DC" box="[687,836,1867,1892]" italics="true" pageId="9" pageNumber="60">T. boulengeri</emphasis>
</taxonomicName>
also exhibits a black streak from below the eye to the ear opening, which is not visible in
<taxonomicName id="D5CF6AC22512320E866E5C76FCBEF83F" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[565,782,1902,1927]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti">
<emphasis id="20BBCD532512320E866E5C76FCBEF83F" box="[565,782,1902,1927]" italics="true" pageId="9" pageNumber="60">T. raymondlaurenti</emphasis>
</taxonomicName>
. In both taxa the supranasals and prefrontals are without contact, although
<taxonomicName id="D5CF6AC22512320E85045C92FE44F813" baseAuthorityName="Sternfeld" baseAuthorityYear="1911" box="[351,500,1938,1963]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="20BBCD532512320E85045C92FE44F813" box="[351,500,1938,1963]" italics="true" pageId="9" pageNumber="60">T. boulengeri</emphasis>
</taxonomicName>
frequently has only one pretemporal (
<bibRefCitation id="765E6CB02512320E87F95C94FBFEF813" author="Broadley, D. G." box="[930,1102,1938,1963]" pageId="9" pageNumber="60" pagination="87 - 110" refId="ref6329" refString="Broadley, D. G. (2000) A review of the genus Mabuya in southeastern Africa (Sauria: Scincidae). African Journal of Herpetology, 49 (2), 87 - 110. https: // doi. org / 10.1080 / 21564574.2000.9635437" type="journal article" year="2000">Broadley 2000</bibRefCitation>
), whereas
<taxonomicName id="D5CF6AC22512320E809E5C92FA2CF813" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[1221,1436,1938,1963]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti">
<emphasis id="20BBCD532512320E809E5C92FA2CF813" box="[1221,1436,1938,1963]" italics="true" pageId="9" pageNumber="60">T. raymondlaurenti</emphasis>
</taxonomicName>
has two.
<taxonomicName id="D5CF6AC22512320E84A75CB0FE4FF877" baseAuthorityName="Sternfeld" baseAuthorityYear="1911" box="[252,511,1974,1999]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="boulengeri">
<emphasis id="20BBCD532512320E84A75CB0FE4FF877" box="[252,511,1974,1999]" italics="true" pageId="9" pageNumber="60">Trachylepis boulengeri</emphasis>
</taxonomicName>
has five to 11 keels per dorsal scale, whereas
<taxonomicName id="D5CF6AC22512320E805D5CBEFB6DF877" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[1030,1245,1974,1999]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="raymondlaurenti">
<emphasis id="20BBCD532512320E805D5CBEFB6DF877" box="[1030,1245,1974,1999]" italics="true" pageId="9" pageNumber="60">T. raymondlaurenti</emphasis>
</taxonomicName>
and
<taxonomicName id="D5CF6AC22512320E814E5CBEFA2DF877" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[1301,1437,1974,1999]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="60" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532512320E814E5CBEFA2DF877" box="[1301,1437,1974,1999]" italics="true" pageId="9" pageNumber="60">T. megalura</emphasis>
</taxonomicName>
have only three weak keels, sometimes nearly indistinct.
</paragraph>
</subSubSection>
<subSubSection id="5AD542CA2511320C849C5B91FE5EFE14" lastPageId="11" lastPageNumber="62" pageId="10" pageNumber="61" type="distribution">
<paragraph id="127011412511320D849C5B91FF44FEA5" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="61">
<emphasis id="20BBCD532511320D849C5B91FED0FF08" bold="true" box="[199,352,151,176]" pageId="10" pageNumber="61">Distribution.</emphasis>
<taxonomicName id="D5CF6AC22511320D85315B91FD01FF08" authority="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[362,689,151,176]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="61" phylum="Chordata" rank="species" species="raymondlaurenti" status="sp. nov.">
<emphasis id="20BBCD532511320D85315B91FD01FF08" box="[362,689,151,176]" italics="true" pageId="10" pageNumber="61">Trachylepis raymondlaurenti</emphasis>
</taxonomicName>
<emphasis id="20BBCD532511320D86E05B91FCA7FF08" bold="true" box="[699,791,151,176]" pageId="10" pageNumber="61">
<taxonomicNameLabel id="3B8870282511320D86E05B91FCA7FF08" box="[699,791,151,176]" pageId="10" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
appears to be restricted to the southeastern
<collectingCountry id="6AD851D12511320D814C5B9EFAE1FF08" box="[1303,1361,152,176]" name="Democratic Republic of the Congo" pageId="10" pageNumber="61">DRC</collectingCountry>
in the
<collectingRegion id="D00BDFA32511320D84CC5BBBFF44FF6D" box="[151,244,189,213]" country="Democratic Republic of the Congo" name="Katanga" pageId="10" pageNumber="61">Katanga</collectingRegion>
Region, and to northern
<collectingCountry id="6AD851D12511320D86735BBAFDCDFF6D" box="[552,637,188,213]" name="Angola" pageId="10" pageNumber="61">Angola</collectingCountry>
, with records for
<collectingRegion id="D00BDFA32511320D87035BBAFC04FF6D" box="[856,948,188,213]" country="Angola" name="Malanje" pageId="10" pageNumber="61">Malanje</collectingRegion>
and
<collectingRegion id="D00BDFA32511320D87AD5BBAFB37FF6D" box="[1014,1159,188,213]" country="Angola" name="Lunda Norte" pageId="10" pageNumber="61">Lunda-Norte</collectingRegion>
provinces. The species potentially occurs in the neighboring provinces of
<collectingRegion id="D00BDFA32511320D868A5BD9FD4BFF40" box="[721,763,223,248]" country="Angola" name="Bie" pageId="10" pageNumber="61">Bié</collectingRegion>
,
<collectingRegion id="D00BDFA32511320D875C5BD9FCCDFF40" box="[775,893,223,248]" country="Angola" name="Lunda Sul" pageId="10" pageNumber="61">Lunda-Sul</collectingRegion>
and
<collectingRegion id="D00BDFA32511320D87EE5BD9FBBCFF40" box="[949,1036,223,248]" country="Angola" name="Moxico" pageId="10" pageNumber="61">Moxico</collectingRegion>
in
<collectingCountry id="6AD851D12511320D806A5BD9FB36FF40" box="[1073,1158,223,248]" name="Angola" pageId="10" pageNumber="61">Angola</collectingCountry>
, and in northern parts of
<collectingCountry id="6AD851D12511320D84CC5A02FF40FEA5" box="[151,240,260,285]" name="Zambia" pageId="10" pageNumber="61">Zambia</collectingCountry>
.
</paragraph>
<caption id="46B041C92511320D84CC5C9DFE0CF868" httpUri="https://zenodo.org/record/2599242/files/figure.png" pageId="10" pageNumber="61" startId="10.[151,250,1947,1969]" targetBox="[173,1415,346,1928]" targetPageId="10">
<paragraph id="127011412511320D84CC5C9DFE0CF868" blockId="10.[151,1437,1947,2000]" pageId="10" pageNumber="61">
<emphasis id="20BBCD532511320D84CC5C9DFEA3F809" bold="true" box="[151,275,1947,1969]" pageId="10" pageNumber="61">FIGURE 6.</emphasis>
Holotype of
<taxonomicName id="D5CF6AC22511320D85C15C9DFD70F809" authority="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer, 2019" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[410,704,1947,1969]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="61" phylum="Chordata" rank="species" species="raymondlaurenti" status="sp. nov.">
<emphasis id="20BBCD532511320D85C15C9DFD70F809" box="[410,704,1947,1969]" italics="true" pageId="10" pageNumber="61">Trachylepis raymondlaurenti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3B8870282511320D869D5C9DFCA7F809" box="[710,791,1947,1969]" pageId="10" pageNumber="61" rank="species">
<emphasis id="20BBCD532511320D869D5C9DFCA1F809" bold="true" box="[710,785,1947,1969]" pageId="10" pageNumber="61">sp. nov</emphasis>
.
</taxonomicNameLabel>
(CAS 258401) from Cangandala National Park, Angola in life. Photo by Luis M. P. Ceríaco.
</paragraph>
</caption>
<paragraph id="127011412510320C849C5B91FE5EFE14" blockId="11.[151,1437,151,573]" pageId="11" pageNumber="62">
<emphasis id="20BBCD532510320C849C5B91FD35FF08" bold="true" box="[199,645,151,176]" pageId="11" pageNumber="62">Habitat and Natural History notes.</emphasis>
The
<typeStatus id="CD74AFE32510320C86965B91FC9FFF08" box="[717,815,151,176]" pageId="11" pageNumber="62" type="holotype">holotype</typeStatus>
was found while it was basking under a shrub of
<taxonomicName id="D5CF6AC22510320C84CC5BBAFD3BFF6D" authority="Oliv." authorityName="Oliv." box="[151,651,188,213]" class="Magnoliopsida" family="Fabaceae" genus="Cryptosepalum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="62" phylum="Tracheophyta" rank="species" species="maraviense">
<emphasis id="20BBCD532510320C84CC5BBAFE7AFF6C" box="[151,458,188,213]" italics="true" pageId="11" pageNumber="62">Cryptosepalum maraviense</emphasis>
Oliv. (Fabaceae)
</taxonomicName>
in a Miombo woodland habitat (
<figureCitation id="8AF40DC42510320C87A05BBAFB8FFF6C" box="[1019,1087,188,213]" captionStart="FIGURE 8" captionStartId="12.[151,250,1942,1964]" captionTargetBox="[159,1432,200,1917]" captionTargetId="figure@12.[151,1437,193,1921]" captionTargetPageId="12" captionText="FIGURE 8. A: General view of the Miombo woodland habitat in the Giant Sable Sanctuary area in the Cangandala National Park where the holotype was collected. B: Cryptosepalum maraviense Oliv. shrub under which the holotype was found basking. Photos by Luis M. P. Ceríaco." httpUri="https://zenodo.org/record/2599246/files/figure.png" pageId="11" pageNumber="62">Fig. 8</figureCitation>
). Dissection revealed ten welldeveloped fetuses in the
<typeStatus id="CD74AFE32510320C85E85BD9FDA5FF40" box="[435,533,223,248]" pageId="11" pageNumber="62" type="holotype">holotype</typeStatus>
(
<figureCitation id="8AF40DC42510320C867D5BD9FDDCFF40" box="[550,620,223,248]" captionStart="FIGURE 7" captionStartId="11.[151,250,1750,1772]" captionTargetBox="[187,1395,679,1708]" captionTargetId="figure@11.[151,1436,628,1729]" captionTargetPageId="11" captionText="FIGURE 7. General view of five out of the ten fetuses found in the reproductive tract of the holotype of Trachylepis raymondlaurenti sp. nov. Photos by Luis M. P. Ceríaco." httpUri="https://zenodo.org/record/2599244/files/figure.png" pageId="11" pageNumber="62">Fig. 7</figureCitation>
), indicating that the young of this diurnal, viviparous species are likely born in late September or October. The fetuses, and presumably neonates, have bright reddish-orange tails that are proportionally shorter than in the adults. The relative length changes in tail length during ontogeny (and presumably also does so in
<taxonomicName id="D5CF6AC22510320C85895A4BFDE8FEDD" authorityName="Marques &amp; Ceríaco &amp; Bandeira &amp; Pauwels &amp; Bauer" authorityYear="2019" box="[466,600,332,357]" class="Reptilia" family="Scincidae" genus="Trachylepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="megalura">
<emphasis id="20BBCD532510320C85895A4BFDE8FEDD" box="[466,600,332,357]" italics="true" pageId="11" pageNumber="62">T. megalura</emphasis>
</taxonomicName>
), explain why the
<typeStatus id="CD74AFE32510320C876B5A4AFC22FEDD" box="[816,914,332,357]" pageId="11" pageNumber="62" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D5CF6AC22510320C87E15A4BFB18FEDC" authorityName="Fischer" authorityYear="1884" box="[954,1192,333,356]" class="Reptilia" family="Scincidae" genus="Eumeces" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="massaianus">
<emphasis id="20BBCD532510320C87E15A4BFB18FEDC" box="[954,1192,333,356]" italics="true" pageId="11" pageNumber="62">Eumeces massaianus</emphasis>
</taxonomicName>
, a probable subadult, has a proportionally shorter tail than the types of
<emphasis id="20BBCD532510320C86B45A77FC52FE30" box="[751,994,367,392]" italics="true" pageId="11" pageNumber="62">
<taxonomicName id="D5CF6AC22510320C86B45A77FC6EFE30" authorityName="Peters" authorityYear="1878" box="[751,990,367,392]" class="Reptilia" family="Scincidae" genus="Euprepes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="62" phylum="Chordata" rank="species" species="megalurus">Euprepes megalurus</taxonomicName>
,
</emphasis>
a fact which contributed to its early recognition as a distinct taxon.
</paragraph>
</subSubSection>
<subSubSection id="5AD542CA2510320C849C5AB1FB4FFD84" pageId="11" pageNumber="62" type="etymology">
<paragraph id="127011412510320C849C5AB1FB4FFD84" blockId="11.[151,1437,151,573]" pageId="11" pageNumber="62">
<emphasis id="20BBCD532510320C849C5AB1FEFEFE68" bold="true" box="[199,334,439,464]" pageId="11" pageNumber="62">Etymology.</emphasis>
The species is named after Raymond F. Laurent (
<date id="667137812510320C879E5AB1FBF8FE68" box="[965,1096,439,464]" pageId="11" pageNumber="62" value="1917" valueMax="2005">19172005</date>
), Belgian herpetologist who specialized in African amphibians and reptiles and contributed significantly to current knowledge of the Angolan and Congolese herpetofaunas. We propose the English name &quot;Laurents Long Tailed Skink&quot;, the Portuguese name &quot;Lagartixa de Cauda Longa de Laurent&quot;, and the French name &quot;Scinque à longue queue de Laurent&quot;.
</paragraph>
</subSubSection>
</treatment>
</document>