259 lines
31 KiB
XML
259 lines
31 KiB
XML
<document id="0C46179B87C1F6060FADF6729B0A5EF6" ID-CLB-Dataset="29706" ID-DOI="10.11646/zootaxa.4444.3.10" ID-GBIF-Dataset="95cf5bec-8224-45bc-a584-7e1d3179b8b5" ID-ISSN="1175-5326" ID-Zenodo-Dep="1309636" ID-ZooBank="B79193AA-92FD-45F3-AA21-DAB4A312D50F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1531313877711" checkinUser="plazi" docAuthor="Gildenkov, Mikhail Yu." docDate="2018" docId="7C21A33FFFE3FF8CFF7B6C01FCD2F8B3" docLanguage="en" docName="zootaxa.4444.3.10.pdf" docOrigin="Zootaxa 4444 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carpelimus (Troginus) varius Gildenkov 2018, sp. n." docType="treatment" docUuid="EB583261-75D5-4486-961C-162AB62DDB12" docUuidSource="ZooBank" docVersion="7" lastPageNumber="349" masterDocId="8018DB47FFE3FF8EFFEC6B20FFEBFFE7" masterDocTitle="A new species of the subgenus Troginus Mulsant et Rey 1878 (Coleoptera: Staphylinidae: Oxytelinae: Carpelimus) from Borneo" masterLastPageNumber="350" masterPageNumber="347" pageNumber="347" updateTime="1698552932971" updateUser="ExternalLinkService">
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<mods:title id="10F4094F4FF0466E36D03847C1235050">A new species of the subgenus Troginus Mulsant et Rey 1878 (Coleoptera: Staphylinidae: Oxytelinae: Carpelimus) from Borneo</mods:title>
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<mods:name id="5419147D4D6F5ED41621F4B10931580C" type="personal">
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<mods:namePart id="3AD17C514A80FDDD848D72AF94F51E62">Gildenkov, Mikhail Yu.</mods:namePart>
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<mods:typeOfResource id="42EF9D9C28DAD51DE185934FE668B48F">text</mods:typeOfResource>
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<mods:title id="AE0EA4A0625F41D1601813346D779F65">Zootaxa</mods:title>
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<mods:part id="AE5D0A72626B15E123B63E26C1029FC6">
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<mods:date id="54D1DF6D7F7580DEB6B37662337E6F50">2018</mods:date>
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<mods:number id="CED4D9546F1617D47745B9CEBBBFC597">2018-07-10</mods:number>
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<mods:number id="0BC11B765F7F34E031B7E08858844924">4444</mods:number>
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<mods:detail id="9EEF3F63DA3B07B97C3929BC74E83EC0" type="issue">
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<mods:number id="24F20008B3D756401B82E257B96B70D9">3</mods:number>
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<mods:identifier id="06296DA90FD0D714D094345233309484" type="DOI">10.11646/zootaxa.4444.3.10</mods:identifier>
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<treatment id="7C21A33FFFE3FF8CFF7B6C01FCD2F8B3" ID-DOI="http://doi.org/10.5281/zenodo.5967789" ID-GBIF-Taxon="146045251" ID-Zenodo-Dep="5967789" LSID="urn:lsid:zoobank.org:act:EB583261-75D5-4486-961C-162AB62DDB12" httpUri="http://treatment.plazi.org/id/7C21A33FFFE3FF8CFF7B6C01FCD2F8B3" lastPageId="2" lastPageNumber="349" pageId="0" pageNumber="347">
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<subSubSection id="BC9241A2FFE3FF8EFF7B6C01FC89F8DC" box="[151,866,1825,1851]" pageId="0" pageNumber="347" type="nomenclature">
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<paragraph id="F4371229FFE3FF8EFF7B6C01FC89F8DC" blockId="0.[151,1436,1825,2011]" box="[151,866,1825,1851]" pageId="0" pageNumber="347">
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<heading id="AF7FA545FFE3FF8EFF7B6C01FC89F8DC" bold="true" box="[151,866,1825,1851]" fontSize="11" level="1" pageId="0" pageNumber="347" reason="1">
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<emphasis id="C6FCCE3BFFE3FF8EFF7B6C01FC89F8DC" bold="true" box="[151,866,1825,1851]" pageId="0" pageNumber="347">
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<taxonomicName id="338869AAFFE3FF8EFF7B6C01FD63F8DC" ID-CoL="RFCN" authority="Gildenkov" authorityName="Gildenkov" authorityYear="2018" box="[151,648,1825,1851]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="347" phylum="Arthropoda" rank="species" species="varius" status="sp. nov." subGenus="Troginus">
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<emphasis id="C6FCCE3BFFE3FF8EFF7B6C01FEC8F8DC" bold="true" box="[151,291,1825,1851]" italics="true" pageId="0" pageNumber="347">Carpelimus</emphasis>
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(
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<emphasis id="C6FCCE3BFFE3FF8EFED86C01FE49F8DC" bold="true" box="[308,418,1825,1851]" italics="true" pageId="0" pageNumber="347">Troginus</emphasis>
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)
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<emphasis id="C6FCCE3BFFE3FF8EFE5F6C01FE15F8DC" bold="true" box="[435,510,1825,1851]" italics="true" pageId="0" pageNumber="347">varius</emphasis>
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Gildenkov
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</taxonomicName>
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,
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<taxonomicNameLabel id="DDCF7340FFE3FF8EFD786C01FD3DF8DC" box="[660,726,1825,1851]" pageId="0" pageNumber="347" rank="species">sp. n.</taxonomicNameLabel>
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||
(
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<figureCitation id="6CB30EACFFE3FF8EFD0A6C01FCB2F8DC" box="[742,857,1825,1851]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="0" pageNumber="347">Figs. 1−5</figureCitation>
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)
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="BC9241A2FFE3FF8FFF7B6C64FCA4FEC8" lastPageId="1" lastPageNumber="348" pageId="0" pageNumber="347" type="materials_examined">
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||
<paragraph id="F4371229FFE3FF8EFF7B6C64FEC2F85C" blockId="0.[151,1436,1825,2011]" pageId="0" pageNumber="347">
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<emphasis id="C6FCCE3BFFE3FF8EFF7B6C64FE5EF8BC" bold="true" box="[151,437,1860,1883]" pageId="0" pageNumber="347">Type material examined:</emphasis>
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Holotype ♂: “BORNEO, Sabah, ca.
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<quantity id="3370BFCCFFE3FF8EFCA46C64FC6FF8BC" box="[840,900,1860,1883]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="347" unit="m" value="300.0">300m</quantity>
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SAFE Project
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<date id="803634E9FFE3FF8EFBC46C64FB77F8BC" box="[1064,1180,1860,1883]" pageId="0" pageNumber="347" value="2011-02" valueMax="2011-03-31" valueMin="2011-02-00">ii.–iii.2011</date>
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Logged forest/Palm oil N4°63´50´´–N4°77´16´´ E117°43´83´´–E117°70´31´´ Malaise/Pitfall Combination” “SAFE Project Ewers
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<emphasis id="C6FCCE3BFFE3FF8EFAE46C44FAD5F89C" box="[1288,1342,1892,1915]" italics="true" pageId="0" pageNumber="347">et al.</emphasis>
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<date id="803634E9FFE3FF8EFAA86C44FA7FF89C" box="[1348,1428,1892,1915]" pageId="0" pageNumber="347" value="23-11-02">110223</date>
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- 201b 6017” “
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<taxonomicName id="338869AAFFE3FF8EFEC76CA4FE4CF87C" box="[299,423,1924,1947]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="347" phylum="Arthropoda" rank="genus">
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<emphasis id="C6FCCE3BFFE3FF8EFEC76CA4FE4CF87C" box="[299,423,1924,1947]" italics="true" pageId="0" pageNumber="347">Carpelimus</emphasis>
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</taxonomicName>
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det. 2015 G. de Rougemont” “Holotypus
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<taxonomicName id="338869AAFFE3FF8EFC996CA4FB5EF87C" authorityName="Gildenkov" authorityYear="2018" box="[885,1205,1924,1947]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="347" phylum="Arthropoda" rank="species" species="varius" subGenus="Troginus">
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<emphasis id="C6FCCE3BFFE3FF8EFC996CA4FC1AF87C" box="[885,1009,1924,1947]" italics="true" pageId="0" pageNumber="347">Carpelimus</emphasis>
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(
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<emphasis id="C6FCCE3BFFE3FF8EFBEF6CA4FB8AF87C" box="[1027,1121,1924,1947]" italics="true" pageId="0" pageNumber="347">Troginus</emphasis>
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)
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<emphasis id="C6FCCE3BFFE3FF8EFB9E6CA4FB5EF87C" box="[1138,1205,1924,1947]" italics="true" pageId="0" pageNumber="347">varius</emphasis>
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</taxonomicName>
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| des. M. Gildenkov,
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<docYear id="ECEEFED2FFE3FF8EFF7B6C84FF25F85C" box="[151,206,1956,1979]" pageId="0" pageNumber="347">2018</docYear>
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” (cGR).
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</paragraph>
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<paragraph id="F4371229FFE3FF8EFF2B6CE4FA77F83C" blockId="0.[151,1436,1825,2011]" box="[199,1436,1988,2011]" pageId="0" pageNumber="347">
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<materialsCitation id="44E01874FFE3FF8EFF2B6CE4FA77F83C" ID-GBIF-Occurrence="1880439628" box="[199,1436,1988,2011]" collectingDate="2011-11" collectingDateMax="2011-12-31" collectingDateMin="2011-11" elevation="300" location="Primary forest" pageId="0" pageNumber="347" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
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<typeStatus id="2B33AC8BFFE3FF8EFF2B6CE4FED8F83C" box="[199,307,1988,2011]" pageId="0" pageNumber="347" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="E28ED9A0FFE3FF8EFEBF6CE4FE9EF83C" box="[339,373,1988,2011]" pageId="0" pageNumber="347" type="female">1♀</specimenCount>
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||
“BORNEO,
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||
<collectingRegion id="364CDCCBFFE3FF8EFDDC6CE4FD9FF83C" box="[560,628,1988,2011]" country="Malaysia" name="Sabah" pageId="0" pageNumber="347">Sabah</collectingRegion>
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, ca.
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<elevation id="7FA5F51AFFE3FF8EFD3E6CE4FCE4F83C" box="[722,783,1988,2011]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="347" unit="m" value="300.0">
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||
<quantity id="3370BFCCFFE3FF8EFD3E6CE4FCE4F83C" box="[722,783,1988,2011]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="347" unit="m" value="300.0">300m</quantity>
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</elevation>
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<collectingDate id="9072CD01FFE3FF8EFCC06CE4FC47F83C" box="[812,940,1988,2011]" pageId="0" pageNumber="347" value="2011-11" valueMax="2011-12-31" valueMin="2011-11-00">
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||
<date id="803634E9FFE3FF8EFCC06CE4FC47F83C" box="[812,940,1988,2011]" pageId="0" pageNumber="347" value="2011-11" valueMax="2011-12-31" valueMin="2011-11-00">xi.–xii.2011</date>
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</collectingDate>
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||
<location id="F15744F2FFE3FF8EFC266CE4FB91F83C" LSID="urn:lsid:plazi:treatment:7C21A33FFFE3FF8CFF7B6C01FCD2F8B3:F15744F2FFE3FF8EFC266CE4FB91F83C" box="[970,1146,1988,2011]" name="Primary forest" pageId="0" pageNumber="347" stateProvince="Sabah">Primary forest</location>
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N4°73´38´´–N4°75´36´´
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</materialsCitation>
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</paragraph>
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<paragraph id="F4371229FFE2FF8FFF7B6BB8FCA4FEC8" blockId="1.[151,1436,152,303]" pageId="1" pageNumber="348">
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E116°94´74´´–E116°96´87´´ Malaise/Pitfall Combination” “
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<collectionCode id="92998AECFFE2FF8FFCAC6BB8FC68FF48" box="[832,899,152,175]" pageId="1" pageNumber="348">SAFE</collectionCode>
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Project Ewers
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<emphasis id="C6FCCE3BFFE2FF8FFBA06BB8FB64FF48" box="[1100,1167,152,175]" italics="true" pageId="1" pageNumber="348">et al.</emphasis>
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<date id="803634E9FFE2FF8FFB4D6BB8FB1BFF48" box="[1185,1264,152,175]" pageId="1" pageNumber="348" value="08-11-11">111108</date>
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-410b 019509” “
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<typeStatus id="2B33AC8BFFE2FF8FFF4C6B98FEE7FF28" box="[160,268,184,207]" pageId="1" pageNumber="348" type="paratype">Paratypus</typeStatus>
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<taxonomicName id="338869AAFFE2FF8FFEF96B98FDBCFF28" authorityName="Gildenkov" authorityYear="2018" box="[277,599,184,207]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="varius" subGenus="Troginus">
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<emphasis id="C6FCCE3BFFE2FF8FFEF96B98FE79FF28" box="[277,402,184,207]" italics="true" pageId="1" pageNumber="348">Carpelimus</emphasis>
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(
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<emphasis id="C6FCCE3BFFE2FF8FFE486B98FDE9FF28" box="[420,514,184,207]" italics="true" pageId="1" pageNumber="348">Troginus</emphasis>
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||
)
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<emphasis id="C6FCCE3BFFE2FF8FFDF86B98FDBCFF28" box="[532,599,184,207]" italics="true" pageId="1" pageNumber="348">varius</emphasis>
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||
</taxonomicName>
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||
| des. M. Gildenkov, 2018” (cGR);
|
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<specimenCount id="E28ED9A0FFE2FF8FFC0B6B97FBE2FF28" box="[999,1033,183,207]" pageId="1" pageNumber="348" type="male">1♂</specimenCount>
|
||
“BORNEO,
|
||
<collectingRegion id="364CDCCBFFE2FF8FFB776B97FB34FF28" box="[1179,1247,183,207]" country="Malaysia" name="Sabah" pageId="1" pageNumber="348">Sabah</collectingRegion>
|
||
, ca.
|
||
<quantity id="3370BFCCFFE2FF8FFAF86B97FABAFF28" box="[1300,1361,183,207]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="348" unit="m" value="300.0">300m</quantity>
|
||
<collectionCode id="92998AECFFE2FF8FFAB66B97FA70FF28" box="[1370,1435,183,207]" pageId="1" pageNumber="348">SAFE</collectionCode>
|
||
Project
|
||
<date id="803634E9FFE2FF8FFF186BF8FE9FFF08" box="[244,372,216,239]" pageId="1" pageNumber="348" value="2011-11" valueMax="2011-12-31" valueMin="2011-11-00">xi.–xii.2011</date>
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||
Logged forest/Palm oil N4°63´50´´–N4°77´16´´
|
||
<geoCoordinate id="91BC74EEFFE2FF8FFC576BF8FBC9FF08" box="[955,1058,216,239]" direction="east" orientation="longitude" pageId="1" pageNumber="348" precision="925" value="117.71667">E117°43´</geoCoordinate>
|
||
83´´–E117°70´31´´ Malaise/Pitfall Combination” “
|
||
<collectionCode id="92998AECFFE2FF8FFED36BD8FE68FEE8" box="[319,387,248,271]" pageId="1" pageNumber="348">SAFE</collectionCode>
|
||
Project Ewers
|
||
<emphasis id="C6FCCE3BFFE2FF8FFDC46BD8FDB4FEE8" box="[552,607,248,271]" italics="true" pageId="1" pageNumber="348">et al.</emphasis>
|
||
<date id="803634E9FFE2FF8FFD8A6BD8FD5EFEE8" box="[614,693,248,271]" pageId="1" pageNumber="348" value="14-11-11">111114</date>
|
||
-244b 017057” “
|
||
<taxonomicName id="338869AAFFE2FF8FFC8B6BD8FC08FEE8" box="[871,995,248,271]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="348" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C6FCCE3BFFE2FF8FFC8B6BD8FC08FEE8" box="[871,995,248,271]" italics="true" pageId="1" pageNumber="348">Carpelimus</emphasis>
|
||
</taxonomicName>
|
||
det. 2015 G. de Rougemont” “
|
||
<typeStatus id="2B33AC8BFFE2FF8FFADC6BD8FA70FEE8" box="[1328,1435,248,271]" pageId="1" pageNumber="348" type="paratype">Paratypus</typeStatus>
|
||
<taxonomicName id="338869AAFFE2FF8FFF7B6A38FE38FEC8" authorityName="Gildenkov" authorityYear="2018" box="[151,467,280,303]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="varius" subGenus="Troginus">
|
||
<emphasis id="C6FCCE3BFFE2FF8FFF7B6A38FEFFFEC8" box="[151,276,280,303]" italics="true" pageId="1" pageNumber="348">Carpelimus</emphasis>
|
||
(
|
||
<emphasis id="C6FCCE3BFFE2FF8FFECF6A38FE6BFEC8" box="[291,384,280,303]" italics="true" pageId="1" pageNumber="348">Troginus</emphasis>
|
||
)
|
||
<emphasis id="C6FCCE3BFFE2FF8FFE7C6A38FE38FEC8" box="[400,467,280,303]" italics="true" pageId="1" pageNumber="348">varius</emphasis>
|
||
</taxonomicName>
|
||
| des. M. Gildenkov, 2018” (cMG).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A0F742A1FFE2FF8FFF7B6D5BFA8BF949" httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="1" pageNumber="348" startId="1.[151,258,1659,1681]" targetBox="[189,1386,418,1586]" targetPageId="1">
|
||
<paragraph id="F4371229FFE2FF8FFF7B6D5BFA8BF949" blockId="1.[151,1436,1659,1710]" pageId="1" pageNumber="348">
|
||
<emphasis id="C6FCCE3BFFE2FF8FFF7B6D5BFED9F976" bold="true" box="[151,306,1659,1681]" pageId="1" pageNumber="348">FIGURES 1–5.</emphasis>
|
||
<taxonomicName id="338869AAFFE2FF8FFEDB6D5CFD54F976" authority="Gildenkov" authorityName="Gildenkov" authorityYear="2018" box="[311,703,1660,1681]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="varius" status="sp. nov." subGenus="Troginus">
|
||
<emphasis id="C6FCCE3BFFE2FF8FFEDB6D5CFDB8F976" box="[311,595,1660,1681]" italics="true" pageId="1" pageNumber="348">Carpelimus (Troginus) varius</emphasis>
|
||
Gildenkov
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="DDCF7340FFE2FF8FFD246D5BFD15F976" box="[712,766,1659,1681]" pageId="1" pageNumber="348" rank="species">
|
||
<emphasis id="C6FCCE3BFFE2FF8FFD246D5BFD13F976" bold="true" box="[712,760,1659,1681]" pageId="1" pageNumber="348">sp. n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="BC9241A2FFE2FF8CFF2B6DF7FDEBF973" lastPageId="2" lastPageNumber="349" pageId="1" pageNumber="348" type="description">
|
||
<paragraph id="F4371229FFE2FF8FFF2B6DF7FCA7F8E9" blockId="1.[151,1436,1751,2030]" pageId="1" pageNumber="348">
|
||
<emphasis id="C6FCCE3BFFE2FF8FFF2B6DF7FEB9F909" bold="true" box="[199,338,1751,1774]" pageId="1" pageNumber="348">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="2B33AC8BFFE2FF8FFE4E6DF7FDEDF909" box="[418,518,1751,1774]" pageId="1" pageNumber="348" type="holotype">holotype</typeStatus>
|
||
): TL: 1.77; WH: 0.43; LH: 0.29; LA: 0.59; LE: 0.10; LT: 0.10; LP: 0.29; WPmax: 0.34; WPmin: 0.22; LEl: 0.37; WE: 0.43; WA: 0.36; LAed: 0.23.
|
||
</paragraph>
|
||
<paragraph id="F4371229FFE2FF8FFF2B6C37FB7FF8A9" blockId="1.[151,1436,1751,2030]" pageId="1" pageNumber="348">Coloration brown. Head, pronotum, most of elytra and abdomen brown; antennae, legs and apices of elytra yellowish brown; scutellum light brown. Body slightly shining, with short, light-coloured setae.</paragraph>
|
||
<paragraph id="F4371229FFE2FF8FFF2B6C77FC0EF809" blockId="1.[151,1436,1751,2030]" pageId="1" pageNumber="348">
|
||
Head large, widest between temples, wider than pronotum (
|
||
<figureCitation id="6CB30EACFFE2FF8FFCAE6C77FC90F889" box="[834,891,1879,1902]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="1" pageNumber="348">Fig 1</figureCitation>
|
||
). Eyes rather large, slightly convex. Temples welldeveloped, round, diameter of eye in dorsal view equal to length of temple. Surface of head densely shagreened. Antennae rather long, antennal segments 1–2 large, elongate; segment 3 elongate; segments 4–5 slightly elongate; segments 6–7 about as wide as long; segment 8 slightly transverse; segments 9–10 transverse; segment 11 elongate, conical. Last 3 segments more massive than others and form loose club (
|
||
<figureCitation id="6CB30EACFFE2FF8FFC706CF7FC3EF809" box="[924,981,2007,2030]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="1" pageNumber="348">Fig 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F4371229FFE1FF8CFF2B6BB8FE1AFF08" blockId="2.[151,1436,152,399]" pageId="2" pageNumber="349">
|
||
Pronotum widest in anterior 1/3, gradually widened from base (
|
||
<figureCitation id="6CB30EACFFE1FF8CFC996BB8FC44FF48" box="[885,943,152,175]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="2" pageNumber="349">Fig 1</figureCitation>
|
||
). Surface of pronotum densely shagreened as that on head. Base of pronotal disc with flat median depression, central part of pronotum with 2 small, symmetrical longitudinal depressions (
|
||
<figureCitation id="6CB30EACFFE1FF8CFE456BF8FE0AFF08" box="[425,481,216,239]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="2" pageNumber="349">Fig 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F4371229FFE1FF8CFF2B6BD8FAFAFEC8" blockId="2.[151,1436,152,399]" pageId="2" pageNumber="349">
|
||
Elytra rather short, only 1.3 times as long as pronotum. Surface of elytra with very delicate and dense, fine punctation, some punctures almost indistinct, microsculpture with grainy, shagreen-like appearance (
|
||
<figureCitation id="6CB30EACFFE1FF8CFB246A38FAEAFEC8" box="[1224,1281,280,303]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="2" pageNumber="349">Fig 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F4371229FFE1FF8CFF2B6A18FD4DFEA8" blockId="2.[151,1436,152,399]" box="[199,678,312,335]" pageId="2" pageNumber="349">Abdomen delicate and smoothly shagreened.</paragraph>
|
||
<paragraph id="F4371229FFE1FF8CFF2B6A78FF2BFE68" blockId="2.[151,1436,152,399]" pageId="2" pageNumber="349">
|
||
Aedeagus (
|
||
<figureCitation id="6CB30EACFFE1FF8CFED36A78FE4BFE88" box="[319,416,344,367]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="2" pageNumber="349">Figs 3, 4</figureCitation>
|
||
) with expanded apical portions of parameres and with poorly sclerotized structures of internal sac.
|
||
</paragraph>
|
||
<paragraph id="F4371229FFE1FF8CFF7B6E42FE47FA90" blockId="2.[151,428,1378,1400]" box="[151,428,1378,1400]" pageId="2" pageNumber="349">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFF7B6E42FED9FA9F" bold="true" box="[151,306,1378,1400]" pageId="2" pageNumber="349">
|
||
<figureCitation id="6CB30EACFFE1FF8CFF7B6E42FEC6FA9F" box="[151,301,1378,1400]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="2" pageNumber="349">FIGURES 1–5</figureCitation>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="C6FCCE3BFFE1FF8CFEDB6E42FE4DFA90" box="[311,422,1378,1399]" italics="true" pageId="2" pageNumber="349">(continued)</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F4371229FFE1FF8CFF2B6E9DFC9CF9D3" blockId="2.[151,1436,1469,1876]" pageId="2" pageNumber="349">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFF2B6E9DFEF1FA33" bold="true" box="[199,282,1469,1492]" pageId="2" pageNumber="349">Female</emphasis>
|
||
(
|
||
<typeStatus id="2B33AC8BFFE1FF8CFEC06E9DFE65FA33" box="[300,398,1469,1492]" pageId="2" pageNumber="349" type="paratype">paratype</typeStatus>
|
||
). Paler in coloration. Head, pronotum and most part of elytra light brown; abdomen brown; antennae, legs and apices of elytra yellowish brown. Female shows sexual dimorphism in body dimensions (
|
||
<figureCitation id="6CB30EACFFE1FF8CFAF16EFDFAB4FA13" box="[1309,1375,1501,1524]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="2" pageNumber="349">Fig. 2</figureCitation>
|
||
): TL: 1.75; WH: 0.30; LH: 0.23; LA: 0.57; LE: 0.07; LT: 0.07; LP: 0.29; WPmax: 0.31; WPmin: 0.19; LEl: 0.33; WE: 0.38; WA: 0.39; LSp: 0.09. Spermatheca of characteristic structure (
|
||
<figureCitation id="6CB30EACFFE1FF8CFCC36D3DFC82F9D3" box="[815,873,1565,1588]" captionStart="FIGURES 1–5" captionStartId="1.[151,258,1659,1681]" captionTargetBox="[189,1386,418,1586]" captionTargetId="figure@1.[158,1428,354,1636]" captionTargetPageId="1" captionText="FIGURES 1–5. Carpelimus (Troginus) varius Gildenkov, sp. n.: 1— habitus of holotype, dorsal view; 2— habitus of paratype, dorsal view; 3— aedeagus, ventral view; 4— spermatheca; 4— aedeagus, lateral view. Scale bar: figs. 1, 2: 1 mm; figs. 3–5: 0.25 mm." httpUri="https://zenodo.org/record/1309638/files/figure.png" pageId="2" pageNumber="349">Fig 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F4371229FFE1FF8CFF2B6D1DFDEBF973" blockId="2.[151,1436,1469,1876]" pageId="2" pageNumber="349">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFF2B6D1DFED2F9B3" bold="true" box="[199,313,1597,1620]" pageId="2" pageNumber="349">Variation.</emphasis>
|
||
The second male from the study material (
|
||
<typeStatus id="2B33AC8BFFE1FF8CFCEA6D1DFC8EF9B3" box="[774,869,1597,1620]" pageId="2" pageNumber="349" type="paratype">paratype</typeStatus>
|
||
) is smaller and does not differ significantly from the female: TL: 1.65; WH: 0.30; LH: 0.23; LA: 0.56; LE: 0.09; LT: 0.07; LP: 0.27; WPmax: 0.30; WPmin: 0.19; LEl: 0.36; WE: 0.41; WA: 0.34; LAed: 0.20.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BC9241A2FFE1FF8CFF2B6DBDFC78F8F3" pageId="2" pageNumber="349" type="diagnosis">
|
||
<paragraph id="F4371229FFE1FF8CFF2B6DBDFC78F8F3" blockId="2.[151,1436,1469,1876]" pageId="2" pageNumber="349">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFF2B6DBDFE29F953" bold="true" box="[199,450,1693,1716]" pageId="2" pageNumber="349">Differential diagnosis.</emphasis>
|
||
The new species differs markedly from all representatives of the subgenus
|
||
<taxonomicName id="338869AAFFE1FF8CFB1B6DBDFABFF953" authorityName="Mulsant et Rey" authorityYear="1878" box="[1271,1364,1693,1716]" class="Insecta" family="Staphylinidae" genus="Troginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFB1B6DBDFABFF953" box="[1271,1364,1693,1716]" italics="true" pageId="2" pageNumber="349">Troginus</emphasis>
|
||
</taxonomicName>
|
||
of the Oriental region by the shagreened surface of the head and pronotum, by the lack of distinct punctation on the elytra and by yellowish brown apices of the elytra. It can be easily distinguished from all representatives of
|
||
<taxonomicName id="338869AAFFE1FF8CFAF96DFDFA98F913" authorityName="Mulsant et Rey" authorityYear="1878" box="[1301,1395,1757,1780]" class="Insecta" family="Staphylinidae" genus="Troginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFAF96DFDFA98F913" box="[1301,1395,1757,1780]" italics="true" pageId="2" pageNumber="349">Troginus</emphasis>
|
||
</taxonomicName>
|
||
by characteristic shapes and location of internal structures of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BC9241A2FFE1FF8CFF2B6C3DFDC4F8D3" box="[199,559,1821,1844]" pageId="2" pageNumber="349" type="distribution">
|
||
<paragraph id="F4371229FFE1FF8CFF2B6C3DFDC4F8D3" blockId="2.[151,1436,1469,1876]" box="[199,559,1821,1844]" pageId="2" pageNumber="349">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFF2B6C3DFEB2F8D3" bold="true" box="[199,345,1821,1844]" pageId="2" pageNumber="349">Distribution.</emphasis>
|
||
Borneo (
|
||
<collectingCountry id="8C9F52B9FFE1FF8CFE506C3DFDCFF8D3" box="[444,548,1821,1844]" name="Malaysia" pageId="2" pageNumber="349">Malaysia</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BC9241A2FFE1FF8CFF2B6C1DFCD2F8B3" box="[199,825,1853,1876]" pageId="2" pageNumber="349" type="etymology">
|
||
<paragraph id="F4371229FFE1FF8CFF2B6C1DFCD2F8B3" blockId="2.[151,1436,1469,1876]" box="[199,825,1853,1876]" pageId="2" pageNumber="349">
|
||
<emphasis id="C6FCCE3BFFE1FF8CFF2B6C1DFEACF8B3" bold="true" box="[199,327,1853,1876]" pageId="2" pageNumber="349">Etymology.</emphasis>
|
||
From Latin “
|
||
<taxonomicName id="338869AAFFE1FF8CFE3A6C1DFDF0F8B3" authorityName="Gildenkov" authorityYear="2018" box="[470,539,1853,1876]" class="Insecta" family="Staphylinidae" genus="Carpelimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="varius">varius</taxonomicName>
|
||
” (motley, multi-coloured).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |