253 lines
31 KiB
XML
253 lines
31 KiB
XML
<document id="5D0C4FA08C4610CDA0B5735B219B61EB" ID-DOI="10.5281/zenodo.5347271" ID-ISSN="2345-7600" ID-Zenodo-Dep="5347271" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630203671219" checkinUser="tatiana" docAuthor="Nguyen, Anh D." docDate="2012" docId="191D7C72CD26E812FC3F9239FF1CFA10" docLanguage="en" docName="60rbz289-311.pdf" docOrigin="Raffles Bulletin of Zoology 60 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Tylopus roseiparaterga Nguyen 2012, new species" docType="treatment" docVersion="3" lastPageNumber="298" masterDocId="E524040ACD22E81BFF859465FF88FFC5" masterDocTitle="Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species" masterLastPageNumber="311" masterPageNumber="289" pageNumber="293" updateTime="1699183855207" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="876BA712DE5B145EA585DF12467AEADC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2B55D9648ADECEE37DC8954E71D5311F">
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<mods:title id="39C756F75ACB53005EB0E12998859FD9">Tylopus Millipedes In Vietnam (Diplopoda: Polydesmida: Paradoxosomatidae: Sulciferini), With Descriptions Of Five New Species</mods:title>
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</mods:titleInfo>
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<mods:name id="A0310958249FB2DF5E832D9244FDD904" type="personal">
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<mods:roleTerm id="4A68C0309226DEF8AEC6810CDA434171">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="09BD2EB89CEC8F0EB3579C13DABA9B11">Nguyen, Anh D.</mods:namePart>
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<mods:typeOfResource id="2BF4E9082EAE3C2CE4885E86EA33F448">text</mods:typeOfResource>
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<mods:title id="A29660A1666EC104248F281F4EF14FB6">Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="86F21CEF39EC5CDFDEA730B817A81ED0">
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<mods:date id="0A80A24684A282B9506262BC771DD323">2012</mods:date>
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<mods:detail id="9C637F890C6D5136A28AFDD410E8515E" type="pubDate">
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<mods:number id="2D51F29152FE60B0068B135CC1C3AD80">2012-08-31</mods:number>
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</mods:detail>
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<mods:detail id="3332A3E6B088C10E4E1EAA7AFEF38C2E" type="volume">
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<mods:number id="1ECD2828EF0DC3E1A1E09E8DEBBF96D8">60</mods:number>
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</mods:detail>
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<mods:detail id="75882A90A9DB50686E64A094D7049A88" type="issue">
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<mods:number id="466B604BDBFEB59138A384512E674381">2</mods:number>
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</mods:detail>
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<mods:extent id="9EE2E88ED7B1AC338BE48A3F77BDCA11" unit="page">
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<mods:start id="F8FB899E581F81FC9C6EA7B08CA04F35">289</mods:start>
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<mods:end id="5A1B4F4A7DF433E48BA7E69E275319F2">311</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="FE5A6B6C4E91951C7377D70ACB62608A">journal article</mods:classification>
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<mods:identifier id="FA10DD82BCE17DB68002B3EF37ADCAE6" type="DOI">10.5281/zenodo.5347271</mods:identifier>
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<mods:identifier id="303C527B5A2D629F68254E2CDAB865DC" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="59D0E2348C62A68A863707E335805133" type="Zenodo-Dep">5347271</mods:identifier>
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</mods:mods>
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<treatment id="191D7C72CD26E812FC3F9239FF1CFA10" LSID="urn:lsid:plazi:treatment:191D7C72CD26E812FC3F9239FF1CFA10" httpUri="http://treatment.plazi.org/id/191D7C72CD26E812FC3F9239FF1CFA10" lastPageId="9" lastPageNumber="298" pageId="4" pageNumber="293">
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<subSubSection id="D9AE9EEFCD26E81FFC3F9239FAC4F9B0" box="[954,1356,1628,1653]" pageId="4" pageNumber="293" type="nomenclature">
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<paragraph id="910BCD64CD26E81FFC3F9239FAC4F9B0" blockId="4.[954,1356,1628,1685]" box="[954,1356,1628,1653]" pageId="4" pageNumber="293">
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<heading id="CA437A08CD26E81FFC3F9239FAC4F9B0" bold="true" box="[954,1356,1628,1653]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="293" reason="2">
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<emphasis id="A3C01176CD26E81FFC3F9239FAC4F9B0" bold="true" box="[954,1356,1628,1653]" pageId="4" pageNumber="293">
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<taxonomicName id="56B4B6E7CD26E81FFC3F9239FB30F9B0" ID-CoL="7CZPV" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[954,1208,1628,1653]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="roseiparaterga" status="sp. nov.">
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<emphasis id="A3C01176CD26E81FFC3F9239FB30F9B0" bold="true" box="[954,1208,1628,1653]" italics="true" pageId="4" pageNumber="293">Tylopus roseiparaterga</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="B8F3AC0DCD26E81FFB429238FAC4F9B0" box="[1223,1356,1629,1653]" pageId="4" pageNumber="293" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D9AE9EEFCD26E81FFBB79218FB5AF950" box="[1074,1234,1661,1685]" pageId="4" pageNumber="293" type="description">
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<paragraph id="910BCD64CD26E81FFBB79218FB5AF950" blockId="4.[954,1356,1628,1685]" box="[1074,1234,1661,1685]" pageId="4" pageNumber="293">
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(
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<figureCitation id="098FD1E1CD26E81FFBBF9218FB0EF950" box="[1082,1158,1661,1685]" captionStart="Fig" captionStartId="7.[113,149,2062,2084]" captionTargetBox="[122,1471,329,1918]" captionTargetId="figure-7@7.[90,1499,323,1976]" captionTargetPageId="7" captionText="Fig. 5. Tylopus roseiparaterga, new species, holotype: body rings 10–11, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); legs 9, (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347281" httpUri="https://zenodo.org/record/5347281/files/figure.png" pageId="4" pageNumber="293">Figs. 5</figureCitation>
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,
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<figureCitation id="098FD1E1CD26E81FFB169218FB2AF950" box="[1171,1186,1661,1685]" captionStart="Fig" captionStartId="8.[137,172,2062,2084]" captionTargetBox="[136,1474,240,2024]" captionTargetId="figure-8@8.[136,1475,240,2025]" captionTargetPageId="8" captionText="Fig. 6. Tylopus roseiparaterga, new species, holotype: left gonopod, lateral view (A); postfemoral region, lateral view (B) and submesal view (C)." figureDoi="http://doi.org/10.5281/zenodo.5347283" httpUri="https://zenodo.org/record/5347283/files/figure.png" pageId="4" pageNumber="293">6</figureCitation>
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,
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<figureCitation id="098FD1E1CD26E81FFB2A9218FB42F950" box="[1199,1226,1661,1685]" captionStart="Fig" captionStartId="21.[113,147,1802,1824]" captionTargetBox="[113,753,1083,1781]" captionTargetId="figure-736@21.[113,753,1083,1779]" captionTargetPageId="21" captionText="Fig. 18.Distribution map ofthe millipede genus Tylopus Jeekel, 1968 in Vietnam. 1: Sa Pa, Lao Cai Prov. (T. crassipes, T. procurvus, T. nodulipes, T. sigma, T. sapaensis, T. maculatus, T. magnicus); 2: Van Ban, Lao Cai Prov. (T. crassipes, T. nodulipes); 3: Xuan Son NP (T. strongylosomoides, T. golovatchi); 4: Tam Dao NP (T. tamdaoensis, T. roseiparaterga, T. strongylosomoides); 5: Cuc Phuong NP (T. granulatus, T. hilaroides, T. topali); 6: Pu Mat NP (T. nodulipes); 7: Huong Son, Ha Tinh prov. (T. nodulipes); 8: Bach Ma–Ba Na NP (T. hilaris, T. multilatus); 9: Ngoc Linh Mts., Kon Tum prov. (T. hilaris, T. phanluongi); 10: Bi Doup–Nui Ba NP (T. spinisterna)." figureDoi="http://doi.org/10.5281/zenodo.5347309" httpUri="https://zenodo.org/record/5347309/files/figure.png" pageId="4" pageNumber="293">18</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D9AE9EEFCD26E81FFCC692DEFC16F8A7" pageId="4" pageNumber="293" type="materials_examined">
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<paragraph id="910BCD64CD26E81FFCC692DEFA9AF8E2" blockId="4.[835,1474,1723,1890]" pageId="4" pageNumber="293">
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<emphasis id="A3C01176CD26E81FFCC692DEFB8FF914" bold="true" box="[835,1031,1723,1745]" italics="true" pageId="4" pageNumber="293">Material examined.</emphasis>
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—
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<materialsCitation id="21DCC739CD26E81FFBAB92D9FA9AF8E2" collectingDate="2010-10-15" collectingDateMax="2010-10-18" collectingDateMin="2010-10-15" collectorName="Nguyen Duc Anh" elevation="950" latitude="21.7" location="Male" longLatPrecision="1262" longitude="105.73333" pageId="4" pageNumber="293" specimenCount="1" stateProvince="Vinh Phuc" typeStatus="holotype">
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<typeStatus id="4E0F73C6CD26E81FFBAB92D9FB03F917" box="[1070,1163,1724,1746]" pageId="4" pageNumber="293" type="holotype">Holotype</typeStatus>
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:
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||
<location id="946B9BBFCD26E81FFB1392D9FB4FF917" LSID="urn:lsid:plazi:treatment:191D7C72CD26E812FC3F9239FF1CFA10:946B9BBFCD26E81FFB1392D9FB4FF917" box="[1174,1223,1724,1746]" latitude="21.7" longLatPrecision="1262" longitude="105.73333" name="Male" pageId="4" pageNumber="293" stateProvince="Vinh Phuc">Male</location>
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in the bottle
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||
<emphasis id="A3C01176CD26E81FFACB92D9FA33F914" bold="true" box="[1358,1467,1724,1745]" pageId="4" pageNumber="293">IEBR-97H</emphasis>
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, with a label “
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<collectingRegion id="53700386CD26E81FFC3A92BCFB0AF92A" box="[959,1154,1753,1775]" country="Vietnam" name="Vinh Phuc" pageId="4" pageNumber="293">Vinh Phuc Province</collectingRegion>
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, Tam Dao National Park (21°21'–
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<geoCoordinate id="F480ABA3CD26E81FFCC69293FC1BF8CE" box="[835,915,1782,1803]" degrees="21" direction="north" minutes="42" orientation="latitude" pageId="4" pageNumber="293" precision="925" value="21.7">21°42'N</geoCoordinate>
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||
, 105°23'–
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||
<geoCoordinate id="F480ABA3CD26E81FFC759290FBC2F8CE" box="[1008,1098,1781,1803]" degrees="105" direction="east" minutes="44" orientation="longitude" pageId="4" pageNumber="293" precision="925" value="105.73333">105°44'E</geoCoordinate>
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), forest,
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||
<elevation id="1A992A57CD26E81FFB1C9293FAC8F8CE" box="[1177,1344,1781,1803]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="4" pageNumber="293" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
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<quantity id="564C6081CD26E81FFB1C9293FA87F8CE" box="[1177,1295,1782,1803]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="4" pageNumber="293" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</quantity>
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a.s.l.
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||
</elevation>
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||
, coll.
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<collectorName id="3C41A8B2CD26E81FFAF29293FC12F8ED" pageId="4" pageNumber="293">Nguyen Duc Anh</collectorName>
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||
& Jun-ichi Kojima,
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||
<date id="E50AEBA4CD26E81FFBE09377FA8DF8E2" box="[1125,1285,1810,1832]" pageId="4" pageNumber="293" value="2010-10-15" valueMax="2010-10-18" valueMin="2010-10-15">
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||
<collectingDate id="F54E124CCD26E81FFBE09377FA8DF8E2" box="[1125,1285,1810,1832]" pageId="4" pageNumber="293" value="2010-10-15" valueMax="2010-10-18" valueMin="2010-10-15">15–18 Oct.2010</collectingDate>
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||
</date>
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||
”.
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||
</materialsCitation>
|
||
</paragraph>
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||
<paragraph id="910BCD64CD26E81FFCC6934AFC16F8A7" blockId="4.[835,1474,1723,1890]" pageId="4" pageNumber="293">
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||
<materialsCitation id="21DCC739CD26E81FFCC6934AFC12F8A7" collectingDate="2010-10-15" collectingDateMax="2010-10-18" collectingDateMin="2010-10-15" collectorName="Nguyen Duc Anh" elevation="950" latitude="21.7" location="Male" longLatPrecision="1262" longitude="105.73333" pageId="4" pageNumber="293" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="Vinh Phuc" typeStatus="paratype">
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||
<typeStatus id="4E0F73C6CD26E81FFCC6934AFC39F881" box="[835,945,1839,1860]" pageId="4" pageNumber="293" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="87B206EDCD26E81FFC44934AFB92F880" box="[961,1050,1839,1861]" pageId="4" pageNumber="293" type="male">6 males</specimenCount>
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,
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<specimenCount id="87B206EDCD26E81FFBAF934AFB03F880" box="[1066,1163,1839,1861]" pageId="4" pageNumber="293" type="female">1 female</specimenCount>
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(
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<emphasis id="A3C01176CD26E81FFB24934AFA9DF881" bold="true" box="[1185,1301,1839,1860]" pageId="4" pageNumber="293">IEBR-97P</emphasis>
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), same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="D9AE9EEFCD26E81FFCC693ECFB18F807" pageId="4" pageNumber="293" type="etymology">
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<paragraph id="910BCD64CD26E81FFCC693ECFB18F807" blockId="4.[835,1474,1929,1986]" pageId="4" pageNumber="293">
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<emphasis id="A3C01176CD26E81FFCC693ECFC36F867" bold="true" box="[835,958,1929,1954]" italics="true" pageId="4" pageNumber="293">Etymology.</emphasis>
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— “
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<taxonomicName id="56B4B6E7CD26E81FFC7093EEFB1BF867" authorityName="Nguyen" authorityYear="2012" box="[1013,1171,1931,1954]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="roseiparaterga">
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<emphasis id="A3C01176CD26E81FFC7093EEFB1BF867" box="[1013,1171,1931,1954]" italics="true" pageId="4" pageNumber="293">roseiparaterga</emphasis>
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</taxonomicName>
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”, a noun in apposition, and emphasizes the pink paraterga.
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</paragraph>
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</subSubSection>
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<subSubSection id="D9AE9EEFCD26E81DFCC6938FFE2CFF33" lastPageId="6" lastPageNumber="295" pageId="4" pageNumber="293" type="diagnosis">
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||
<paragraph id="910BCD64CD26E81FFCC6938FFA4AF787" blockId="4.[835,1474,2026,2114]" pageId="4" pageNumber="293">
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||
<emphasis id="A3C01176CD26E81FFCC6938FFC48F7C7" bold="true" box="[835,960,2026,2050]" italics="true" pageId="4" pageNumber="293">Diagnosis.</emphasis>
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— The species can be distinguished from its congeners by the pink paraterga and by gonopod conformation: lamina
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<emphasis id="A3C01176CD26E81FFBA89C4FFBBCF787" bold="true" box="[1069,1076,2090,2114]" pageId="4" pageNumber="293">l</emphasis>
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||
subtrapeziform; process
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<emphasis id="A3C01176CD26E81FFAB99C4FFAC3F787" bold="true" box="[1340,1355,2090,2114]" pageId="4" pageNumber="293">h</emphasis>
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||
short, thin,
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</paragraph>
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||
<caption id="C5CB9DECCD27E81EFFF49C6AFB0CF784" ID-DOI="http://doi.org/10.5281/zenodo.5347277" ID-Zenodo-Dep="5347277" httpUri="https://zenodo.org/record/5347277/files/figure.png" pageId="5" pageNumber="294" startId="5.[113,148,2063,2085]" targetBox="[109,1452,324,1968]" targetPageId="5">
|
||
<paragraph id="910BCD64CD27E81EFFF49C6AFB0CF784" blockId="5.[113,1452,2063,2114]" pageId="5" pageNumber="294">
|
||
Fig. 3.
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||
<taxonomicName id="56B4B6E7CD27E81EFF309C75FEF8F7E1" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[181,368,2064,2084]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="spinisterna" status="sp. nov.">
|
||
<emphasis id="A3C01176CD27E81EFF309C75FEF8F7E1" box="[181,368,2064,2084]" italics="true" pageId="5" pageNumber="294">Tylopus spinisterna</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="B8F3AC0DCD27E81EFEF99C75FE7AF7E0" box="[380,498,2063,2085]" pageId="5" pageNumber="294" rank="species">new species</taxonomicNameLabel>
|
||
, holotype: body ring 10, dorsal view (A); hypoproct, ventral view (B); sternite X, ventral view (C); sternite 5, posterioventral view (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
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||
<paragraph id="910BCD64CD24E81DFF0C94DBFE2CFF33" blockId="6.[137,775,190,246]" pageId="6" pageNumber="295">
|
||
finger-shaped; solenophore strongly coiled; lobe
|
||
<emphasis id="A3C01176CD24E81DFD1094DBFD23FF13" bold="true" box="[661,683,190,214]" pageId="6" pageNumber="295">m</emphasis>
|
||
present; lobe
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||
<emphasis id="A3C01176CD24E81DFF3B94BBFF45FF33" bold="true" box="[190,205,222,246]" pageId="6" pageNumber="295">n</emphasis>
|
||
and spine
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||
<emphasis id="A3C01176CD24E81DFEC194BBFED8FF33" bold="true" box="[324,336,222,246]" pageId="6" pageNumber="295">z</emphasis>
|
||
absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AE9EEFCD24E812FF0C957BFDAFFAB0" lastPageId="9" lastPageNumber="298" pageId="6" pageNumber="295" type="description">
|
||
<paragraph id="910BCD64CD24E81DFF0C957BFEFCFE53" blockId="6.[137,775,286,630]" pageId="6" pageNumber="295">
|
||
<emphasis id="A3C01176CD24E81DFF0C957BFE9DFEF3" bold="true" box="[137,277,286,310]" italics="true" pageId="6" pageNumber="295">Description.</emphasis>
|
||
— Size: Body length 34–[36] mm (male),
|
||
<quantity id="564C6081CD24E81DFF0C955BFF72FE93" box="[137,250,318,342]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.1" metricValueMin="3.0" pageId="6" pageNumber="295" unit="mm" value="30.5" valueMax="31.0" valueMin="30.0">30–31 mm</quantity>
|
||
(female). Width of prozona [2.6]–
|
||
<quantity id="564C6081CD24E81DFDE1955BFD3DFE93" box="[612,693,318,342]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="295" unit="mm" value="2.8">2.8 mm</quantity>
|
||
(male),
|
||
<quantity id="564C6081CD24E81DFF0C953BFE85FEB3" box="[137,269,350,374]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="6" pageNumber="295" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
(female) and metazona 3.4–[3.5] mm (male),
|
||
<quantity id="564C6081CD24E81DFF0C951BFE82FE53" box="[137,266,382,406]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="6" pageNumber="295" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.2–3.3 mm</quantity>
|
||
(female).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD24E81DFF0C95FBFF5CFDB3" blockId="6.[137,775,286,630]" pageId="6" pageNumber="295">Colouration: Prozona and metaterga with a median broad whitish-yellow stripe and two paramedian small brown regions. Paraterga pink (living specimens) or paler (preserved specimens). Pleura and distal podomeres brown. Sterna yellowish brown. Most antennomeres brown except for distal part of antennomere 6 and whole antennomere 7 blackish brown.</paragraph>
|
||
<paragraph id="910BCD64CD24E81DFCC694DBFBE7FE93" blockId="6.[835,1474,190,631]" pageId="6" pageNumber="295">
|
||
Head: Slightly broader than collum. Labrum densely setose. Epicranial suture thin, evident, dividing frons into two parts; each part with 2 setae along epicranial suture. Antenna claviform, long, reaching body ring 4 laterally. Antennomere 3>2=4=5>6>1&
|
||
<quantity id="564C6081CD24E81DFC77955BFB95FE93" box="[1010,1053,318,342]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="6" pageNumber="295" unit="in" value="7.0">7 in</quantity>
|
||
length.
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD24E81DFCC6953BFC74FE13" blockId="6.[835,1474,190,631]" pageId="6" pageNumber="295">Collum: Slightly narrower than body ring 2; surface shining, more or less smooth, with two rows of 3+3 and 1+1 setae. Axial line evident. Paratergum very small, with a setiferous incision laterally.</paragraph>
|
||
<paragraph id="910BCD64CD24E812FCC695BBFD0DFE50" blockId="6.[835,1474,190,631]" lastBlockId="9.[113,752,189,1397]" lastPageId="9" lastPageNumber="298" pageId="6" pageNumber="295">
|
||
Body ring 4<3<2=
|
||
<quantity id="564C6081CD24E81DFB9595BBFBB2FE33" box="[1040,1082,478,502]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="295" unit="in" value="5.0">5 in</quantity>
|
||
width, body parallel-sided on rings 5-16, thereafter gradually tapering towards telson. Surface of metaterga 2–4 more or less smooth whereas other metaterga considerably rugose. Transverse sulcus narrow, concave, present on metaterga 4-19, but fully developed (reaching base of paraterga) from metatergum 5. Anterior half of metaterga 2–19 with row of 2+2 setae (
|
||
<figureCitation id="098FD1E1CD2BE812FD9E94B8FDFBFF30" box="[539,627,221,245]" captionStart="Fig" captionStartId="7.[113,149,2062,2084]" captionTargetBox="[122,1471,329,1918]" captionTargetId="figure-7@7.[90,1499,323,1976]" captionTargetPageId="7" captionText="Fig. 5. Tylopus roseiparaterga, new species, holotype: body rings 10–11, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); legs 9, (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347281" httpUri="https://zenodo.org/record/5347281/files/figure.png" pageId="9" pageNumber="298">Fig. 5A</figureCitation>
|
||
). Axial line only evident in anterior half of metaterga. Waist between pro- and metazona broad, finely striate. Pleura with dense microgranules. Pleurosternal keels well developed on body rings 2–4, reduced as small caudal spine on body rings 5–9, and completely absent on subsequent body rings.
|
||
</paragraph>
|
||
<caption id="C5CB9DECCD24E81DFF0C9C6BFF27F784" ID-DOI="http://doi.org/10.5281/zenodo.5347279" ID-Zenodo-Dep="5347279" httpUri="https://zenodo.org/record/5347279/files/figure.png" pageId="6" pageNumber="295" startId="6.[137,172,2062,2084]" targetBox="[138,1474,724,2025]" targetPageId="6">
|
||
<paragraph id="910BCD64CD24E81DFF0C9C6BFF27F784" blockId="6.[137,1474,2062,2113]" pageId="6" pageNumber="295">
|
||
Fig. 4.
|
||
<taxonomicName id="56B4B6E7CD24E81DFF4B9C6AFE02F7E6" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[206,394,2063,2083]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="spinisterna" status="sp. nov.">
|
||
<emphasis id="A3C01176CD24E81DFF4B9C6AFE02F7E6" box="[206,394,2063,2083]" italics="true" pageId="6" pageNumber="295">Tylopus spinisterna</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B8F3AC0DCD24E81DFE129C6BFD85F7E1" box="[407,525,2062,2084]" pageId="6" pageNumber="295" rank="species">new species</taxonomicNameLabel>
|
||
, holotype: left gonopod, lateral view (A) and submesal view (B); postfemoral region, lateral view (C).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C5CB9DECCD25E81CFFF49C6BFB1AF784" ID-DOI="http://doi.org/10.5281/zenodo.5347281" ID-Zenodo-Dep="5347281" httpUri="https://zenodo.org/record/5347281/files/figure.png" pageId="7" pageNumber="296" startId="7.[113,149,2062,2084]" targetBox="[122,1471,329,1918]" targetPageId="7">
|
||
<paragraph id="910BCD64CD25E81CFFF49C6BFB1AF784" blockId="7.[113,1452,2062,2113]" pageId="7" pageNumber="296">
|
||
Fig. 5.
|
||
<taxonomicName id="56B4B6E7CD25E81CFF3B9C6AFE2EF7E6" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[190,422,2063,2083]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="296" phylum="Arthropoda" rank="species" species="roseiparaterga" status="sp. nov.">
|
||
<emphasis id="A3C01176CD25E81CFF3B9C6AFE2EF7E6" box="[190,422,2063,2083]" italics="true" pageId="7" pageNumber="296">Tylopus roseiparaterga</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B8F3AC0DCD25E81CFE339C6AFDBAF7E1" box="[438,562,2062,2084]" pageId="7" pageNumber="296" rank="species">new species</taxonomicNameLabel>
|
||
, holotype: body rings 10–11, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); legs 9, (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C5CB9DECCD2AE813FF0C9C6BFF6DF784" ID-DOI="http://doi.org/10.5281/zenodo.5347283" ID-Zenodo-Dep="5347283" httpUri="https://zenodo.org/record/5347283/files/figure.png" pageId="8" pageNumber="297" startId="8.[137,172,2062,2084]" targetBox="[136,1474,240,2024]" targetPageId="8">
|
||
<paragraph id="910BCD64CD2AE813FF0C9C6BFF6DF784" blockId="8.[137,1473,2062,2113]" pageId="8" pageNumber="297">
|
||
Fig. 6.
|
||
<taxonomicName id="56B4B6E7CD2AE813FF559C6AFE38F7E6" authority="Nguyen, 2012" authorityName="Nguyen" authorityYear="2012" box="[208,432,2063,2083]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="297" phylum="Arthropoda" rank="species" species="roseiparaterga" status="sp. nov.">
|
||
<emphasis id="A3C01176CD2AE813FF559C6AFE38F7E6" box="[208,432,2063,2083]" italics="true" pageId="8" pageNumber="297">Tylopus roseiparaterga</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B8F3AC0DCD2AE813FE3B9C6AFDBEF7E1" box="[446,566,2062,2084]" pageId="8" pageNumber="297" rank="species">new species</taxonomicNameLabel>
|
||
, holotype: left gonopod, lateral view (A); postfemoral region, lateral view (B) and submesal view (C).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="910BCD64CD2BE812FFF495F8FE4BFD50" blockId="9.[113,752,189,1397]" pageId="9" pageNumber="298">Paraterga: Well developed, slightly lower than metatergal surface. Caudal corner somewhat rounded on body rings 2–13, but gradually protruding, acute on body rings 14–19. Paraterga not surpassing posterior contour of metaterga 2–13, but surpassing posterior contour of metaterga 14–19. Calluses very small, somewhat reduced on poreless paraterga, but more developed on pore-bearing paraterga, with two traces of setiferous incisions laterally.</paragraph>
|
||
<paragraph id="910BCD64CD2BE812FFF496F8FF53FCD0" blockId="9.[113,752,189,1397]" pageId="9" pageNumber="298">
|
||
Telson: Epiproct long, truncated with two small terminal tubercles. Tip concave, with four spinnerets. Hypoproct subtrapeziform, with two small distolateral setiferous knobs (
|
||
<figureCitation id="098FD1E1CD2BE812FFFC9698FF46FCD0" box="[121,206,765,789]" captionStart="Fig" captionStartId="7.[113,149,2062,2084]" captionTargetBox="[122,1471,329,1918]" captionTargetId="figure-7@7.[90,1499,323,1976]" captionTargetPageId="7" captionText="Fig. 5. Tylopus roseiparaterga, new species, holotype: body rings 10–11, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); legs 9, (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347281" httpUri="https://zenodo.org/record/5347281/files/figure.png" pageId="9" pageNumber="298">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD2BE812FFF49778FEBFFC50" blockId="9.[113,752,189,1397]" pageId="9" pageNumber="298">
|
||
Sterna: Sparsely setose, without any modifications; longitudinal and transverse sutures obvious. Male sternum 5 with a highly elevated, setiferous, bifid lamina between coxae 4 (
|
||
<figureCitation id="098FD1E1CD2BE812FF519718FEA2FC50" box="[212,298,893,917]" captionStart="Fig" captionStartId="7.[113,149,2062,2084]" captionTargetBox="[122,1471,329,1918]" captionTargetId="figure-7@7.[90,1499,323,1976]" captionTargetPageId="7" captionText="Fig. 5. Tylopus roseiparaterga, new species, holotype: body rings 10–11, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); legs 9, (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347281" httpUri="https://zenodo.org/record/5347281/files/figure.png" pageId="9" pageNumber="298">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD2BE812FFF497F8FE92FB90" blockId="9.[113,752,189,1397]" pageId="9" pageNumber="298">
|
||
Legs: Thin and slender, about 1.3–1.4 times as long as midbody height. Prefemora not swollen. Ventral tubercles present on all legs. Tarsal brushes present on leg pairs 1–15, thinner on leg pairs 16–21 and absent on subsequent leg pairs. Ventral part of prefemora and femora with long setae (
|
||
<figureCitation id="098FD1E1CD2BE812FF339058FE85FB90" box="[182,269,1085,1109]" captionStart="Fig" captionStartId="7.[113,149,2062,2084]" captionTargetBox="[122,1471,329,1918]" captionTargetId="figure-7@7.[90,1499,323,1976]" captionTargetPageId="7" captionText="Fig. 5. Tylopus roseiparaterga, new species, holotype: body rings 10–11, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); legs 9, (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347281" httpUri="https://zenodo.org/record/5347281/files/figure.png" pageId="9" pageNumber="298">Fig. 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="910BCD64CD2BE812FFF49038FDAFFAB0" blockId="9.[113,752,189,1397]" pageId="9" pageNumber="298">
|
||
Gonopod (
|
||
<figureCitation id="098FD1E1CD2BE812FF6D9038FED5FBB0" box="[232,349,1117,1141]" captionStart="Fig" captionStartId="7.[113,149,2062,2084]" captionTargetBox="[122,1471,329,1918]" captionTargetId="figure-7@7.[90,1499,323,1976]" captionTargetPageId="7" captionText="Fig. 5. Tylopus roseiparaterga, new species, holotype: body rings 10–11, dorsal view (A); hypoproct, ventral view (B); sternite 5, posterioventral view (C); legs 9, (D); left gonopod, mesal view (E) and lateral view (F). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5347281" httpUri="https://zenodo.org/record/5347281/files/figure.png" pageId="9" pageNumber="298">Figs. 5E,F</figureCitation>
|
||
,
|
||
<figureCitation id="098FD1E1CD2BE812FEEE9038FE39FBB0" box="[363,433,1117,1141]" captionStart="Fig" captionStartId="8.[137,172,2062,2084]" captionTargetBox="[136,1474,240,2024]" captionTargetId="figure-8@8.[136,1475,240,2025]" captionTargetPageId="8" captionText="Fig. 6. Tylopus roseiparaterga, new species, holotype: left gonopod, lateral view (A); postfemoral region, lateral view (B) and submesal view (C)." figureDoi="http://doi.org/10.5281/zenodo.5347283" httpUri="https://zenodo.org/record/5347283/files/figure.png" pageId="9" pageNumber="298">6A–C</figureCitation>
|
||
): Coxite cylindrical, slightly shorter than femorite; distoventral part sparsely setose. Prefemorite densely setose, set off from femorite by an oblique sulcus laterally. Femorite slender, slightly twisted, and demarcated from postfemoral region by a distinct transverse sulcus laterally. Lamina
|
||
<emphasis id="A3C01176CD2BE812FE059098FE0FFAD0" bold="true" box="[384,391,1277,1301]" pageId="9" pageNumber="298">l</emphasis>
|
||
present, subtrapeziform; spine
|
||
<emphasis id="A3C01176CD2BE812FD619098FD78FAD0" bold="true" box="[740,752,1277,1301]" pageId="9" pageNumber="298">z</emphasis>
|
||
absent. Process
|
||
<emphasis id="A3C01176CD2BE812FEA29178FEBEFAF0" bold="true" box="[295,310,1309,1333]" pageId="9" pageNumber="298">h</emphasis>
|
||
thin, short and finger-shaped. Lobe
|
||
<emphasis id="A3C01176CD2BE812FD5F9178FD78FAF0" bold="true" box="[730,752,1309,1333]" pageId="9" pageNumber="298">m</emphasis>
|
||
moderate whereas lobe
|
||
<emphasis id="A3C01176CD2BE812FEFC9158FE00FA90" bold="true" box="[377,392,1341,1365]" pageId="9" pageNumber="298">n</emphasis>
|
||
absent. Solenophore completely sheathing solenomere and twisted twice.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AE9EEFCD2BE812FFF491F9FF1CFA10" pageId="9" pageNumber="298" type="distribution">
|
||
<paragraph id="910BCD64CD2BE812FFF491F9FF1CFA10" blockId="9.[113,752,1436,1493]" pageId="9" pageNumber="298">
|
||
<emphasis id="A3C01176CD2BE812FFF491F9FF76FA70" bold="true" box="[113,254,1436,1461]" italics="true" pageId="9" pageNumber="298">Distribution.</emphasis>
|
||
— Known only from the
|
||
<typeStatus id="4E0F73C6CD2BE812FDAC91F8FDDFFA70" box="[553,599,1437,1461]" pageId="9" pageNumber="298">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="098FD1E1CD2BE812FD4191F8FF04FA10" captionStart="Fig" captionStartId="21.[113,147,1802,1824]" captionTargetBox="[113,753,1083,1781]" captionTargetId="figure-736@21.[113,753,1083,1779]" captionTargetPageId="21" captionText="Fig. 18.Distribution map ofthe millipede genus Tylopus Jeekel, 1968 in Vietnam. 1: Sa Pa, Lao Cai Prov. (T. crassipes, T. procurvus, T. nodulipes, T. sigma, T. sapaensis, T. maculatus, T. magnicus); 2: Van Ban, Lao Cai Prov. (T. crassipes, T. nodulipes); 3: Xuan Son NP (T. strongylosomoides, T. golovatchi); 4: Tam Dao NP (T. tamdaoensis, T. roseiparaterga, T. strongylosomoides); 5: Cuc Phuong NP (T. granulatus, T. hilaroides, T. topali); 6: Pu Mat NP (T. nodulipes); 7: Huong Son, Ha Tinh prov. (T. nodulipes); 8: Bach Ma–Ba Na NP (T. hilaris, T. multilatus); 9: Ngoc Linh Mts., Kon Tum prov. (T. hilaris, T. phanluongi); 10: Bi Doup–Nui Ba NP (T. spinisterna)." figureDoi="http://doi.org/10.5281/zenodo.5347309" httpUri="https://zenodo.org/record/5347309/files/figure.png" pageId="9" pageNumber="298">Fig. 18</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |