190 lines
21 KiB
XML
190 lines
21 KiB
XML
<document id="5210407BDE90D5570AC7D2E3EF1D5005" ID-CLB-Dataset="55533" ID-DOI="10.5281/zenodo.5174222" ID-GBIF-Dataset="b0a0b4e7-7c0d-4b51-9a59-156975677df7" ID-ISSN="1942-1354" ID-Zenodo-Dep="5174222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628335594538" checkinUser="felipe" docAuthor="Dellacasa, Marco, Dellacasa, Giovanni & Gordon, Robert D." docDate="2012" docId="03E28783FF91EE72FF16F900F4792F88" docLanguage="en" docName="InsectaMundi.0230.1-41.pdf" docOrigin="Insecta Mundi 2012 (230)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Gonaphodiellus Schmidt 1913" docType="treatment" docVersion="7" lastPageNumber="3" masterDocId="FFDBFFFBFF92EE76FFD6FFE8F1332B10" masterDocTitle="Systematic revision of Gonaphodiellus taxa, with description of two new genera and fourteen new species (Coleoptera: Scarabaeidae: Aphodiinae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="2" updateTime="1699069380811" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="0148EBC856E4711540E710F0587DF67C">
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<mods:title id="DEF0035A49ABBAF45F83E1A84F721691">Systematic revision of Gonaphodiellus taxa, with description of two new genera and fourteen new species (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<mods:name id="FB38ACFDA45953E82C6968CEFE731A26" type="personal">
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<mods:namePart id="D264B44BFDB0B227A5252ADE4F7A51DF">Dellacasa, Marco</mods:namePart>
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<mods:affiliation id="C55828A3C6209D9165B46D6369A1AC58">Museo di Storia Naturale e del Territorio Università di Pisa Via Roma, 79 I- 56011 Calci (Pisa), Italy</mods:affiliation>
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<mods:nameIdentifier id="191DB67E8562E43919D3FFB70BA21E6B" type="email">dellacasa@museo.unipi.it</mods:nameIdentifier>
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<mods:namePart id="82AC162689CE648D646F3E7DC720ED07">Dellacasa, Giovanni</mods:namePart>
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<mods:affiliation id="58A1E4676F98F98CCE33C5827E5AE284">Via Talamone 31 / 19 I- 16127 Genova, Italy</mods:affiliation>
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<mods:nameIdentifier id="024E51F40A832B825AA8A51DB0C465AC" type="email">dellacasag@alice.it</mods:nameIdentifier>
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<mods:namePart id="0F17002350ED2142C2D40CB6F48E09F2">Gordon, Robert D.</mods:namePart>
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<mods:affiliation id="5CF76FA32C2C2F4810AAC93A0A20AFC8">Northern Plains Entomology P. O. Box 65 Willow City, ND 58384 USA</mods:affiliation>
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<mods:nameIdentifier id="8554A7F2E1E45CBDA77E1C589A81B934" type="email">rdgordon@utma.com</mods:nameIdentifier>
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<mods:title id="2866E142CD32639B27F4CFC03DBA8F83">Insecta Mundi</mods:title>
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<mods:date id="498DA1E73B6DCEFE2150ABB8A0B4B1D6">2012</mods:date>
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<mods:number id="3F6E6DD591E36C6B4D5DE0973F5B2472">2012-04-06</mods:number>
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<mods:number id="0E682365E80538170EBA7DCE7C5CBBE9">2012</mods:number>
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<treatment id="03E28783FF91EE72FF16F900F4792F88" ID-DOI="http://doi.org/10.5281/zenodo.6547451" ID-GBIF-Taxon="195308681" ID-Zenodo-Dep="6547451" LSID="urn:lsid:plazi:treatment:03E28783FF91EE72FF16F900F4792F88" httpUri="http://treatment.plazi.org/id/03E28783FF91EE72FF16F900F4792F88" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<subSubSection id="C351651EFF91EE75FF16F900F3F92DEB" box="[192,714,1768,1787]" pageId="3" pageNumber="2" type="nomenclature">
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<heading id="D0BC81F9FF91EE75FF16F900F3F92DEB" bold="true" box="[192,714,1768,1787]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<emphasis id="B93FEA87FF91EE75FF16F900F3F92DEB" bold="true" box="[192,714,1768,1787]" pageId="3" pageNumber="2">
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Genus
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<taxonomicName id="4C4B4D16FF91EE75FEF6F900F3F92DEB" authority="Schmidt, 1913" authorityName="Schmidt" authorityYear="1913" box="[288,714,1768,1787]" class="Insecta" family="Aphodiidae" genus="Gonaphodiellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA87FF91EE75FEF6F900F3332DEB" bold="true" box="[288,512,1768,1787]" italics="true" pageId="3" pageNumber="2">Gonaphodiellus</emphasis>
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<bibRefCitation id="EFDA4B64FF91EE75FDD1F900F3F92DEB" author="Schmidt, A." box="[519,714,1768,1787]" pageId="3" pageNumber="2" pagination="117 - 178" refId="ref23232" refString="Schmidt, A. 1913. Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv fur Naturgeschichte, Abt. A, 79: 117 - 178." type="journal article" year="1913">Schmidt, 1913</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF43695FF91EE75FF16F8CFF3DB2C28" blockId="3.[192,1096,1831,1906]" box="[192,744,1831,1848]" pageId="3" pageNumber="2">
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<treatmentCitationGroup id="AB5B11BBFF91EE75FF16F8CFF3DB2C28" box="[192,744,1831,1848]" pageId="3" pageNumber="2">
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<treatmentCitation id="0AEA1084FF91EE75FF16F8CFF3D72C28" author="Schmidt, A." box="[192,740,1831,1848]" page="133" pageId="3" pageNumber="2" year="1913">
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<taxonomicName id="4C4B4D16FF91EE75FF16F8CFF3D72C28" authority="Schmidt, 1913: 133" authorityName="Schmidt" authorityPageNumber="133" authorityYear="1913" box="[192,740,1831,1848]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Gonaphodiellus">
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<emphasis id="B93FEA87FF91EE75FF16F8CFF01D2C28" box="[192,302,1831,1848]" italics="true" pageId="3" pageNumber="2">Aphodius</emphasis>
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(
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<emphasis id="B93FEA87FF91EE75FEEBF8CFF0C72C28" box="[317,500,1831,1848]" italics="true" pageId="3" pageNumber="2">Gonaphodiellus</emphasis>
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)
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<bibRefCitation id="EFDA4B64FF91EE75FDD2F8CFF3D72C28" author="Schmidt, A." box="[516,740,1831,1848]" pageId="3" pageNumber="2" pagination="117 - 178" refId="ref23232" refString="Schmidt, A. 1913. Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv fur Naturgeschichte, Abt. A, 79: 117 - 178." type="journal article" year="1913">Schmidt, 1913: 133</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BF43695FF91EE75FF16F8ACF3632C45" blockId="3.[192,1096,1831,1906]" box="[192,592,1860,1877]" pageId="3" pageNumber="2">
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<treatmentCitationGroup id="AB5B11BBFF91EE75FF16F8ACF3632C45" box="[192,592,1860,1877]" pageId="3" pageNumber="2">
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<taxonomicName id="4C4B4D16FF91EE75FF16F8ACF0442C45" authorityName="Schmidt" authorityYear="1913" box="[192,375,1860,1877]" class="Insecta" family="Aphodiidae" genus="Gonaphodiellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA87FF91EE75FF16F8ACF0442C45" box="[192,375,1860,1877]" italics="true" pageId="3" pageNumber="2">Gonaphodiellus</emphasis>
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</taxonomicName>
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;
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<treatmentCitation id="0AEA1084FF91EE75FE53F8ACF37F2C45" author="Paulian, R." box="[389,588,1860,1877]" page="73" pageId="3" pageNumber="2" year="1942">
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<bibRefCitation id="EFDA4B64FF91EE75FE53F8ACF37F2C45" author="Paulian, R." box="[389,588,1860,1877]" pageId="3" pageNumber="2" pagination="1 - 143" refId="ref23139" refString="Paulian, R. 1942. Aphodiinae (Coleoptera Lamellicornia) Fam. Scarabaeidae. Exploration du Parc National Albert, Mission G. F. de Witte (19331935). Bruxelles, 35: 1 - 143." type="journal article" year="1942">Paulian 1942: 73</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF43695FF91EE75FF16F889F57B2C62" blockId="3.[192,1096,1831,1906]" box="[192,1096,1889,1906]" pageId="3" pageNumber="2">
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<treatmentCitationGroup id="AB5B11BBFF91EE75FF16F889F57B2C62" box="[192,1096,1889,1906]" pageId="3" pageNumber="2">
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<taxonomicName id="4C4B4D16FF91EE75FF16F889F0CC2C62" authority="" authorityName="Schmidt" authorityYear="1913" box="[192,511,1889,1906]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Gonaphodiellus">
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<emphasis id="B93FEA87FF91EE75FF16F889F01D2C62" box="[192,302,1889,1906]" italics="true" pageId="3" pageNumber="2">Aphodius</emphasis>
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(
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<emphasis id="B93FEA87FF91EE75FEEBF889F0C72C62" box="[317,500,1889,1906]" italics="true" pageId="3" pageNumber="2">Gonaphodiellus</emphasis>
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)
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</taxonomicName>
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;
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<treatmentCitation id="0AEA1084FF91EE75FDDDF889F3D52C62" author="Balthasar, V." box="[523,742,1889,1906]" page="58" pageId="3" pageNumber="2" year="1960">
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<bibRefCitation id="EFDA4B64FF91EE75FDDDF889F3D52C62" author="Balthasar, V." box="[523,742,1889,1906]" pageId="3" pageNumber="2" pagination="55 - 63" refId="ref22717" refString="Balthasar, V. 1960. Neue Aphodius-Arten aus der orientalischen, aethiopischen und neotropischen Region. Mitteilungen der Munchener Entomologischen Gesellschaft 50: 55 - 63." type="journal article" year="1960">Balthasar 1960: 58</bibRefCitation>
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</treatmentCitation>
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(as synonym of
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<taxonomicName id="4C4B4D16FF91EE75FC48F889F50B2C62" authorityName="Reitter" authorityYear="1892" box="[926,1080,1889,1906]" class="Insecta" family="Aphodiidae" genus="Gonaphodius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA87FF91EE75FC48F889F50B2C62" box="[926,1080,1889,1906]" italics="true" pageId="3" pageNumber="2">Gonaphodius</emphasis>
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</taxonomicName>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF43695FF96EE72FF16FF2FF3CB2BC8" blockId="4.[192,956,199,247]" box="[192,760,199,216]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="AB5B11BBFF96EE72FF16FF2FF3CB2BC8" box="[192,760,199,216]" pageId="4" pageNumber="3">
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<taxonomicName id="4C4B4D16FF96EE72FF16FF2FF0CC2BC8" authority="" authorityName="Schmidt" authorityYear="1913" box="[192,511,199,216]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Gonaphodiellus">
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<emphasis id="B93FEA87FF96EE72FF16FF2FF01D2BC8" box="[192,302,199,216]" italics="true" pageId="4" pageNumber="3">Aphodius</emphasis>
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(
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<emphasis id="B93FEA87FF96EE72FEEBFF2FF0C72BC8" box="[317,500,199,216]" italics="true" pageId="4" pageNumber="3">Gonaphodiellus</emphasis>
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)
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</taxonomicName>
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;
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<treatmentCitation id="0AEA1084FF96EE72FDDDFF2FF3C72BC8" author="Dellacasa, M." box="[523,756,199,216]" page="239" pageId="4" pageNumber="3" year="1988">
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<bibRefCitation id="EFDA4B64FF96EE72FDDDFF2FF3C72BC8" author="Dellacasa, M." box="[523,756,199,216]" pageId="4" pageNumber="3" pagination="1 - 455" refId="ref22906" refString="Dellacasa, M. 1988. Contribution to a world-wide Catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae. (Part I). Memorie della Societa Entomologica Italiana [1987] 66: 1 - 455." type="journal article" year="1988">Dellacasa 1988: 239</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF43695FF96EE72FF16FF0EF28F2BE7" blockId="4.[192,956,199,247]" box="[192,956,230,247]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="AB5B11BBFF96EE72FF16FF0EF28F2BE7" box="[192,956,230,247]" pageId="4" pageNumber="3">
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<taxonomicName id="4C4B4D16FF96EE72FF16FF0EF0442BE7" authorityName="Schmidt" authorityYear="1913" box="[192,375,230,247]" class="Insecta" family="Aphodiidae" genus="Gonaphodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B93FEA87FF96EE72FF16FF0EF0442BE7" box="[192,375,230,247]" italics="true" pageId="4" pageNumber="3">Gonaphodiellus</emphasis>
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</taxonomicName>
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;
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<treatmentCitation id="0AEA1084FF96EE72FE50FF0EF3872BE7" author="Dellacasa, M. & al." box="[390,692,230,247]" page="154" pageId="4" pageNumber="3" year="2001">
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<bibRefCitation id="EFDA4B64FF96EE72FE50FF0EF3872BE7" author="Dellacasa, G. & P. Bordat & M. Dellacasa" box="[390,692,230,247]" firstAuthor="Dellacasa" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="4" pageNumber="3" pagination="1 - 482" part="79" refId="ref22866" refString="Dellacasa, G., P. Bordat, and M. Dellacasa. 2001. A revisional essay of world genus-group taxa of Aphodiinae. Memorie della Societa Entomologica Italiana [2000] 79: 1 - 482." title="A revisional essay of world genus-group taxa of Aphodiinae" type="journal article" year="2001">Dellacasa et al. 2001: 154</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AEA1084FF96EE72FD17FF0EF28B2BE7" author="Skelley, P. E. & Dellacasa M. & Dellacasa G. & R. D. Gordon" box="[705,952,230,247]" httpUri="http://treatment.plazi.org/id/03CF900C4B43FFD203F6FB5DFB07FF41" page="4" pageId="4" pageNumber="3" year="2007">
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<bibRefCitation id="EFDA4B64FF96EE72FD17FF0EF28B2BE7" author="Skelley, P. E. & Dellacasa M. & Dellacasa G. & R. D. Gordon" box="[705,952,230,247]" pageId="4" pageNumber="3" pagination="1 - 14" refId="ref23381" refString="Skelley, P. E., Dellacasa M., Dellacasa G. and R. D. Gordon. 2007. Checklist of the Aphodiini of Mexico, Central and South America. Insecta Mundi 0014: 1 - 14." type="book chapter" year="2007">Skelley et al. 2007: 4</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C351651EFF96EE72FF16FECDF46B2A28" box="[192,1368,293,312]" pageId="4" pageNumber="3" type="type_taxon">
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<paragraph id="8BF43695FF96EE72FF16FECDF46B2A28" blockId="4.[192,1368,293,312]" box="[192,1368,293,312]" pageId="4" pageNumber="3">
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<emphasis id="B93FEA87FF96EE72FF16FECDF0482A28" bold="true" box="[192,379,293,312]" pageId="4" pageNumber="3">
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<typeStatus id="54F08837FF96EE72FF16FECDF0352A28" box="[192,262,293,312]" pageId="4" pageNumber="3">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C4B4D16FF96EE72FE54FECDF23E2A28" authority="Harold, 1880" authorityName="Harold" authorityYear="1880" box="[386,781,293,312]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="pacatus">
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<emphasis id="B93FEA87FF96EE72FE54FECDF3502A28" box="[386,611,293,312]" italics="true" pageId="4" pageNumber="3">Aphodius pacatus</emphasis>
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<bibRefCitation id="EFDA4B64FF96EE72FDBCFECDF23E2A28" author="Harold, E." box="[618,781,293,312]" pageId="4" pageNumber="3" pagination="13 - 43" refId="ref23038" refString="Harold, E. 1880. Verzeichniss der von E. Steinheil in New-Granada gesammelten coprophagen Lamellicornien. Stettiner Entomologische Zeitschift 41: 13 - 43." type="journal article" year="1880">Harold, 1880</bibRefCitation>
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</taxonomicName>
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(subsequent designation by
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<bibRefCitation id="EFDA4B64FF96EE72FBA5FECDF4792A28" author="Paulian, R." box="[1139,1354,293,312]" pageId="4" pageNumber="3" pagination="1 - 143" refId="ref23139" refString="Paulian, R. 1942. Aphodiinae (Coleoptera Lamellicornia) Fam. Scarabaeidae. Exploration du Parc National Albert, Mission G. F. de Witte (19331935). Bruxelles, 35: 1 - 143." type="journal article" year="1942">Paulian 1942: 73</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C351651EFF96EE72FF16FE8DF2FF28E8" pageId="4" pageNumber="3" type="diagnosis">
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<paragraph id="8BF43695FF96EE72FF16FE8DF2FF28E8" blockId="4.[192,1441,357,1016]" pageId="4" pageNumber="3">
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<emphasis id="B93FEA87FF96EE72FF16FE8DF0662A68" bold="true" box="[192,341,357,376]" pageId="4" pageNumber="3">Diagnosis.</emphasis>
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Rather small to medium size species (length 3.5-6.0 mm), feebly or moderately convex, short or elongate oval, feebly shiny, glabrous or almost glabrous. More or less dark brownish, with or without elytral paler spots. Head with epistome punctured; clypeus subtruncate anteriorly or faintly sinuate at middle, round at sides, quite finely bordered, edge bristled; genae obtusely round, very shortly ciliate, protruding from the eyes or not; eyes rather large; frontal suture finely impressed, not tuberculate; first segment of labial palp as long as the second. Pronotum transverse, punctured; hind angles truncate and more or less distinctly inwardly sinuate; basal margin not bordered. Scutellum small, narrow, regularly triangular. Elytra more or less strongly denticulate at shoulder, rarely lacking denticle; striae fine and punctured, first one more superficial than the others and sometimes faint toward apex; interstriae slightly convex, finely punctured, rarely shortly sparsely pubescent on preapical declivity. Fore tibiae distally tridentate and proximally serrulate on outer margin; upper side rather densely and more or less coarsely punctured. Middle and hind tibiae with feeble transverse carinae on outer face, apically fimbriate with spinules elongate and irregularly unequal. Hind tibiae upper spur shorter than first tarsal segment. Pygidium with straight pubescence. Sexual dimorphism shown in males mainly by shape of fore tibiae and respective spur; by sparser punctation of head and pronotum; and pronotum with hind angles more strongly inwardly sinuate. Aedeagus with parameres short, at least as long as half of tegmen, or somewhat more elongate, always with clump of more or less dense hairs toward apex. Epipharynx with anterior margin distinctly sinuate at middle; epitorma drop-shaped; corypha with two strong apical spiculae; pedia densely pubescent, intermixed few stout and large chaetae; chaetopariae slender and elongate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C351651EFF96EE72FF16FBCDF25B2F28" box="[192,872,1061,1080]" pageId="4" pageNumber="3" type="distribution">
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<paragraph id="8BF43695FF96EE72FF16FBCDF25B2F28" blockId="4.[192,872,1061,1080]" box="[192,872,1061,1080]" pageId="4" pageNumber="3">
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<emphasis id="B93FEA87FF96EE72FF16FBCDF0492F28" bold="true" box="[192,378,1061,1080]" pageId="4" pageNumber="3">Distribution.</emphasis>
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Mesoamerican and Neotropical region.
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</paragraph>
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</subSubSection>
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<subSubSection id="C351651EFF96EE72FF16FB8DF4792F88" pageId="4" pageNumber="3" type="discussion">
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<paragraph id="8BF43695FF96EE72FF16FB8DF4792F88" blockId="4.[192,1439,1125,1176]" pageId="4" pageNumber="3">
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<emphasis id="B93FEA87FF96EE72FF16FB8DF07B2F68" bold="true" box="[192,328,1125,1144]" pageId="4" pageNumber="3">Remarks.</emphasis>
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The correct identification of unassociated females of several
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<taxonomicName id="4C4B4D16FF96EE72FB80FB8DF4B02F68" box="[1110,1411,1125,1144]" class="Insecta" family="Aphodiidae" genus="Gonaphodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B93FEA87FF96EE72FB80FB8DF4132F68" box="[1110,1312,1125,1144]" italics="true" pageId="4" pageNumber="3">Gonaphodiellus</emphasis>
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species
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</taxonomicName>
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is more or less impossible due to their weakly differentiated external morphological characters.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |