treatments-xml/data/03/D8/87/03D887BE8168360A2A55FE9F44600BDC.xml

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<mods:title id="19CDF460CDEFDD0BB49329D681072765">Revision of the New World Ceratoculicoides Wirth &amp; Ratanaworabhan (Diptera, Ceratopogonidae, Ceratopogonini)</mods:title>
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<mods:namePart id="63CFFAD00B362DACF8F27C9CCB495986">Fasbender, Andrew</mods:namePart>
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<treatment id="03D887BE8168360A2A55FE9F44600BDC" ID-DOI="http://doi.org/10.5281/zenodo.8083824" ID-GBIF-Taxon="210477699" ID-Zenodo-Dep="8083824" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC" httpUri="http://treatment.plazi.org/id/03D887BE8168360A2A55FE9F44600BDC" lastPageId="32" lastPageNumber="190" pageId="29" pageNumber="188">
<subSubSection id="C36B6523816836372A55FE9F451F0854" box="[536,1052,264,292]" pageId="29" pageNumber="188" type="nomenclature">
<paragraph id="8BCE36A8816836372A55FE9F451F0854" blockId="29.[536,1052,264,328]" box="[536,1052,264,292]" pageId="29" pageNumber="188">
<heading id="D08681C4816836372A55FE9F451F0854" box="[536,1052,264,292]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="188" reason="2">
<taxonomicName id="4C714D2B816836372A55FE9F451F0854" ID-CoL="SMSD" authority="(Wirth, 1951)" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[536,1052,264,292]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoidesvirginianus" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA816836372A55FE9F42770853" bold="true" box="[536,884,264,291]" italics="true" pageId="29" pageNumber="188">Ceratoculicoidesvirginianus</emphasis>
(
<bibRefCitation id="EFE04B59816836372BCEFE9E45100854" author="Wirth W. W." box="[899,1043,265,292]" pageId="29" pageNumber="188" pagination="313 - 326" refId="ref23778" refString="Wirth W. W. 1951. New species and records of Virginia Heleidae. Proceedings of the Entomological Society of Washington 53: 313 - 326." type="journal article" year="1951">Wirth, 1951</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B6523816836372AD4FEBA42980838" box="[665,923,301,328]" pageId="29" pageNumber="188" type="description">
<paragraph id="8BCE36A8816836372AD4FEBA42980838" blockId="29.[536,1052,264,328]" box="[665,923,301,328]" pageId="29" pageNumber="188">
<figureCitation id="134A2A2D816836372AD4FEBA43F10837" box="[665,754,301,327]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[195,1398,668,1908]" captionTargetId="figure-118@9.[195,1398,668,1908]" captionTargetPageId="9" captionText="Fig. 4. Male heads I. a. Ceratoculicoides confusus sp. nov. b. C. pacificus sp. nov. c. C. confusus (Colombia). d. C.virginianus (Wirth, 1951). Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083785" httpUri="https://zenodo.org/record/8083785/files/figure.png" pageId="29" pageNumber="188">Figs 4d</figureCitation>
, 
<figureCitation id="134A2A2D816836372AB3FEBA421D0837" box="[766,798,301,327]" captionStart="Fig" captionStartId="11.[189,232,1963,1989]" captionTargetBox="[222,1365,830,1930]" captionTargetId="figure-172@11.[222,1365,830,1930]" captionTargetPageId="11" captionText="Fig. 6. Female heads I. a. Ceratoculicoidesconfusus sp. nov. b. Ceratoculicoides sp. F2 (Colombia). c. C. pacificus sp. nov. d. C.virginianus (Wirth, 1951). Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083791" httpUri="https://zenodo.org/record/8083791/files/figure.png" pageId="29" pageNumber="188">6d</figureCitation>
, 
<figureCitation id="134A2A2D816836372B67FEB942670838" box="[810,868,302,328]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="29" pageNumber="188">9a–c</figureCitation>
, 
<figureCitation id="134A2A2D816836372B3DFEB942980838" box="[880,923,302,328]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="29" pageNumber="188">12c</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B65238168363728F0FEE740F50BBD" pageId="29" pageNumber="188" type="reference_group">
<paragraph id="8BCE36A88168363728F0FEE7451F08FB" blockId="29.[189,1385,368,431]" box="[189,1052,368,395]" pageId="29" pageNumber="188">
<taxonomicName id="4C714D2B8168363728F0FEE743CA08FA" ID-CoL="88BYB" authority="Wirth 1951: 318" authorityName="Wirth" authorityPageNumber="318" authorityYear="1951" box="[189,713,368,395]" class="Insecta" family="Ceratopogonidae" genus="Helea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="virginiana" subGenus="Isohelea">
<emphasis id="B905EABA8168363728F0FEE7400008FA" box="[189,259,368,394]" italics="true" pageId="29" pageNumber="188">Helea</emphasis>
(
<emphasis id="B905EABA816836372959FEE7407408FA" box="[276,375,368,394]" italics="true" pageId="29" pageNumber="188">Isohelea</emphasis>
) 
<emphasis id="B905EABA8168363729C4FEE6430108FA" box="[393,514,369,394]" italics="true" pageId="29" pageNumber="188">virginiana</emphasis>
<bibRefCitation id="EFE04B59816836372A44FEE743CA08FA" author="Wirth W. W." box="[521,713,368,395]" pageId="29" pageNumber="188" pagination="313 - 326" refId="ref23778" refString="Wirth W. W. 1951. New species and records of Virginia Heleidae. Proceedings of the Entomological Society of Washington 53: 313 - 326." type="journal article" year="1951">Wirth 1951: 318</bibRefCitation>
</taxonomicName>
(original description), fig. 5.
</paragraph>
<paragraph id="8BCE36A88168363728F0FE02446A08DF" blockId="29.[189,1385,368,431]" box="[189,1385,405,431]" pageId="29" pageNumber="188">
<taxonomicName id="4C714D2B8168363728F0FE02429D08DF" ID-CoL="SMS7" authority="With &amp; Ratanaworabhan, 1971: 172" authorityName="Wirth &amp; Ratanaworabhan" authorityPageNumber="172" authorityYear="1971" box="[189,926,405,431]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA8168363728F0FE0240EC08DF" box="[189,495,405,431]" italics="true" pageId="29" pageNumber="188">Ceratoculicoides blantoni</emphasis>
With &amp; Ratanaworabhan, 1971: 172
</taxonomicName>
(original description), fig. 2. 
<emphasis id="B905EABA816836372CB5FE02446A08DF" bold="true" box="[1272,1385,405,431]" pageId="29" pageNumber="188">Syn. nov.</emphasis>
</paragraph>
<paragraph id="8BCE36A88168363728F0FE4F45860882" blockId="29.[189,1399,471,718]" box="[189,1157,471,498]" pageId="29" pageNumber="188">
<taxonomicName id="4C714D2B8168363728F0FE4F43720881" box="[189,625,472,498]" class="Insecta" family="Ceratopogonidae" genus="Ceratopogon" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="virginianus" subGenus="Nilohelea">
<emphasis id="B905EABA8168363728F0FE4F405B0882" box="[189,344,472,498]" italics="true" pageId="29" pageNumber="188">Ceratopogon</emphasis>
(
<emphasis id="B905EABA816836372924FE4F40D80882" box="[361,475,472,498]" italics="true" pageId="29" pageNumber="188">Nilohelea</emphasis>
) 
<emphasis id="B905EABA8168363729A1FE4F43720881" box="[492,625,472,497]" italics="true" pageId="29" pageNumber="188">virginianus</emphasis>
</taxonomicName>
<emphasis id="B905EABA816836372A35FE4F43840881" box="[632,647,472,497]" italics="true" pageId="29" pageNumber="188">–</emphasis>
<bibRefCitation id="EFE04B59816836372AC3FE4F424C0882" author="Wirth W. W." box="[654,847,471,498]" pageId="29" pageNumber="188" pagination="121 - 142" refId="ref23830" refString="Wirth W. W. 1965. Family Ceratopogonidae. In: Stone A., Sabrosky C. W., Wirth W. W., Foote R. H. &amp; Coulson J. R. (eds) A Catalog of the Diptera of America North of Mexico: 121 - 142. U. S. Government Printing Office, Washington D. C." type="book chapter" year="1965">Wirth 1965: 133</bibRefCitation>
(combination, in catalog).
</paragraph>
<paragraph id="8BCE36A88168363728F0FE6B44720B2F" blockId="29.[189,1399,471,718]" pageId="29" pageNumber="188">
<taxonomicName id="4C714D2B8168363728F0FE6B43120B66" ID-CoL="SMSD" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[189,529,508,534]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA8168363728F0FE6B43120B66" box="[189,529,508,534]" italics="true" pageId="29" pageNumber="188">Ceratoculicoidesvirginianus</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFE04B59816836372A63FE6B42DA0B66" author="Wirth W. W. &amp; Ratanaworabhan N. C." box="[558,985,508,534]" pageId="29" pageNumber="188" pagination="170 - 177" refId="ref23938" refString="Wirth W. W. &amp; Ratanaworabhan N. C. 1971. Ceratoculicoides, a new genus related to Ceratopogon Meigen (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 73: 170 - 177." type="journal article" year="1971">Wirth &amp; Ratanaworabhan 1971: 172</bibRefCitation>
(redescription), fig. 3. — 
<bibRefCitation id="EFE04B59816836372D41FE6A40630B4B" author="Knoz J." pageId="29" pageNumber="188" pagination="388 - 392" refId="ref23379" refString="Knoz J. 1987. Description of a new species of Ceratoculicoides (Diptera, Ceratopogonidae) from Czechoslovakia and a review of the genus. Acta Entomologica Bohemoslovaca 84: 388 - 392." type="journal article" year="1987">Knoz 1987: 391</bibRefCitation>
(key). — 
<bibRefCitation id="EFE04B59816836372994FDB6420D0B4C" author="Borkent A. &amp; Wirth W. W." box="[473,782,545,572]" pageId="29" pageNumber="188" pagination="1 - 257" refId="ref22357" refString="Borkent A. &amp; Wirth W. W. 1997. World species of biting midges (Diptera: Ceratopogonidae). Bulletin of the American Museum of Natural History 233: 1 - 257." type="journal article" year="1997">Borkent &amp; Wirth 1997: 95</bibRefCitation>
(in catalog). — 
<bibRefCitation id="EFE04B59816836372B9DFDB644230B4B" author="Huerta H. &amp; Borkent A." box="[976,1312,545,572]" pageId="29" pageNumber="188" pagination="111 - 119" refId="ref23309" refString="Huerta H. &amp; Borkent A. 2005. A new species and first record of Ceratoculicoides Wirth &amp; Ratanaworabhan from the Neotropical region and new species and records of Brachypogon Kieffer from Mexico (Diptera: Ceratopogonidae). Folia Entomologica Mexicana 44: 111 - 119." type="journal article" year="2005">Huerta &amp; Borkent 2005: 114</bibRefCitation>
(catalog). — 
<bibRefCitation id="EFE04B598168363729CBFDD243A70B2F" author="Borkent A. &amp; Grogan W. L." box="[390,676,581,607]" pageId="29" pageNumber="188" pagination="1 - 48" refId="ref22308" refString="Borkent A. &amp; Grogan W. L. 2009. Catalog of the New World biting midges north of Mexico (Diptera: Ceratopogonidae). Zootaxa 2273 (1): 1 - 48. https: // doi. org / 10.11646 / zootaxa. 2273.1.1" type="journal article" year="2009">Borkent &amp; Grogan 2009</bibRefCitation>
(in catalog). — 
<bibRefCitation id="EFE04B59816836372B28FDD245DF0B10" author="Borkent A. &amp; Dominiak P." box="[869,1244,581,608]" pageId="29" pageNumber="188" pagination="1 - 377" refId="ref22263" refString="Borkent A. &amp; Dominiak P. 2020. Catalog of the biting midges of the world (Diptera: Ceratopogonidae). Zootaxa 4787 (1): 1 - 377. https: // doi. org / 10.11646 / zootaxa. 4787.1.1" type="journal article" year="2020">Borkent &amp; Dominiak 2020: 157</bibRefCitation>
(in catalog).
</paragraph>
<paragraph id="8BCE36A88168363728F0FDFD40F50BBD" blockId="29.[189,1399,471,718]" pageId="29" pageNumber="188">
<taxonomicName id="4C714D2B8168363728F0FDFD40EE0BF4" ID-CoL="SMS7" authority="With &amp; Ratanaworabhan, 1971: 172" authorityName="Wirth &amp; Ratanaworabhan" authorityPageNumber="172" authorityYear="1971" box="[189,493,618,644]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA8168363728F0FDFD40EE0BF4" box="[189,493,618,644]" italics="true" pageId="29" pageNumber="188">Ceratoculicoidesblantoni</emphasis>
</taxonomicName>
– 
<bibRefCitation id="EFE04B59816836372A40FDFD43C40BF4" author="Knoz J." box="[525,711,618,644]" pageId="29" pageNumber="188" pagination="388 - 392" refId="ref23379" refString="Knoz J. 1987. Description of a new species of Ceratoculicoides (Diptera, Ceratopogonidae) from Czechoslovakia and a review of the genus. Acta Entomologica Bohemoslovaca 84: 388 - 392." type="journal article" year="1987">Knoz 1987: 391</bibRefCitation>
(key). — 
<bibRefCitation id="EFE04B59816836372B71FDFD45680BF5" author="Borkent A. &amp; Wirth W. W." box="[828,1131,618,645]" pageId="29" pageNumber="188" pagination="1 - 257" refId="ref22357" refString="Borkent A. &amp; Wirth W. W. 1997. World species of biting midges (Diptera: Ceratopogonidae). Bulletin of the American Museum of Natural History 233: 1 - 257." type="journal article" year="1997">Borkent &amp; Wirth 1997: 95</bibRefCitation>
(in catalog). — 
<bibRefCitation id="EFE04B59816836372D6AFDFD40EB0BD9" author="Huerta H. &amp; Borkent A." pageId="29" pageNumber="188" pagination="111 - 119" refId="ref23309" refString="Huerta H. &amp; Borkent A. 2005. A new species and first record of Ceratoculicoides Wirth &amp; Ratanaworabhan from the Neotropical region and new species and records of Brachypogon Kieffer from Mexico (Diptera: Ceratopogonidae). Folia Entomologica Mexicana 44: 111 - 119." type="journal article" year="2005">Huerta &amp; Borkent 2005: 114</bibRefCitation>
(catalog). — 
<bibRefCitation id="EFE04B59816836372ADDFD1942B00BD9" author="Borkent A. &amp; Grogan W. L." box="[656,947,654,681]" pageId="29" pageNumber="188" pagination="1 - 48" refId="ref22308" refString="Borkent A. &amp; Grogan W. L. 2009. Catalog of the New World biting midges north of Mexico (Diptera: Ceratopogonidae). Zootaxa 2273 (1): 1 - 48. https: // doi. org / 10.11646 / zootaxa. 2273.1.1" type="journal article" year="2009">Borkent &amp; Grogan 2009</bibRefCitation>
(in catalog). — 
<bibRefCitation id="EFE04B59816836372C37FD1940630BBE" author="Borkent A. &amp; Dominiak P." pageId="29" pageNumber="188" pagination="1 - 377" refId="ref22263" refString="Borkent A. &amp; Dominiak P. 2020. Catalog of the biting midges of the world (Diptera: Ceratopogonidae). Zootaxa 4787 (1): 1 - 377. https: // doi. org / 10.11646 / zootaxa. 4787.1.1" type="journal article" year="2020">Borkent &amp; Dominiak 2020: 157</bibRefCitation>
(in catalog).
</paragraph>
</subSubSection>
<subSubSection id="C36B65238168363728F0FD6143760D2E" pageId="29" pageNumber="188" type="diagnosis">
<paragraph id="8BCE36A88168363728F0FD6140350A60" blockId="29.[189,310,758,784]" box="[189,310,758,784]" pageId="29" pageNumber="188">
<heading id="D08681C48168363728F0FD6140350A60" bold="true" box="[189,310,758,784]" fontSize="11" level="3" pageId="29" pageNumber="188" reason="3">
<emphasis id="B905EABA8168363728F0FD6140350A60" bold="true" box="[189,310,758,784]" pageId="29" pageNumber="188">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A88168363728A7FCB140290A30" blockId="29.[189,1398,806,942]" box="[234,298,806,832]" pageId="29" pageNumber="188">
<emphasis id="B905EABA8168363728A7FCB140290A30" bold="true" box="[234,298,806,832]" pageId="29" pageNumber="188">Male</emphasis>
</paragraph>
<paragraph id="8BCE36A88168363728F0FCDD441B0ADD" blockId="29.[189,1398,806,942]" pageId="29" pageNumber="188">
<taxonomicName id="4C714D2B8168363728F0FCDD43190A14" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[189,538,842,868]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA8168363728F0FCDD43190A14" box="[189,538,842,868]" italics="true" pageId="29" pageNumber="188">Ceratoculicoides virginianus</emphasis>
</taxonomicName>
can be separated from congeners by the following combination of characters: femora and tibiae yellow or brown; apices of paramere broadly rounded, not tapering, minute apical point displaced laterally; aedeagus subquadrate, slightly constricted apically (
<figureCitation id="134A2A2D816836372CD5FC0444080ADD" box="[1176,1291,915,942]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="29" pageNumber="188">Fig. 9b–c</figureCitation>
).
</paragraph>
<paragraph id="8BCE36A88168363728A7FC4140460A80" blockId="29.[189,1399,982,1118]" box="[234,325,982,1008]" pageId="29" pageNumber="188">
<emphasis id="B905EABA8168363728A7FC4140460A80" bold="true" box="[234,325,982,1008]" pageId="29" pageNumber="188">Female</emphasis>
</paragraph>
<paragraph id="8BCE36A88168363728F0FC6C43760D2E" blockId="29.[189,1399,982,1118]" pageId="29" pageNumber="188">
Only species of 
<taxonomicName id="4C714D2B816836372933FC6C43450D65" authorityName=", Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[382,582,1019,1045]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA816836372933FC6C43450D65" box="[382,582,1019,1045]" italics="true" pageId="29" pageNumber="188">Ceratoculicoides</emphasis>
</taxonomicName>
with the following combination of characters: FR 1.57–1.75; femora and tibiae yellow or brown, wing length ~ 
<quantity id="4C899B4D816836372AF0FB8842530D4A" box="[701,848,1055,1082]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="29" pageNumber="188" unit="mm" value="1.1" valueMax="1.3" valueMin="0.9">0.9–1.3 mm</quantity>
; 2 major spermathecae, largest 52–69; medial margin of 9 
<superScript id="7C049BE0816836372908FBD440510D21" attach="left" box="[325,338,1091,1105]" fontSize="6" pageId="29" pageNumber="188">th</superScript>
sternite deeply concave.
</paragraph>
</subSubSection>
<subSubSection id="C36B65238168363728F0FB1145B00E98" pageId="29" pageNumber="188" type="materials_examined">
<paragraph id="8BCE36A88168363728F0FB1140A90DD0" blockId="29.[189,426,1158,1184]" box="[189,426,1158,1184]" pageId="29" pageNumber="188">
<heading id="D08681C48168363728F0FB1140A90DD0" bold="true" box="[189,426,1158,1184]" fontSize="11" level="3" pageId="29" pageNumber="188" reason="3">
<emphasis id="B905EABA8168363728F0FB1140A90DD0" bold="true" box="[189,426,1158,1184]" pageId="29" pageNumber="188">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B193CF58168363728A7FB2045B00D85" ID-GBIF-Occurrence="4135858320" collectingDate="1950-07-11" collectionCode="USNM" collectorName="W. W. Wirth" country="United States of America" county="Augusta Co." location="Mount Solon" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Virginia" typeStatus="holotype">
<paragraph id="8BCE36A88168363728A7FB20405F0DA1" blockId="29.[189,1203,1207,1270]" box="[234,348,1207,1233]" pageId="29" pageNumber="188">
<emphasis id="B905EABA8168363728A7FB20405F0DA1" bold="true" box="[234,348,1207,1233]" pageId="29" pageNumber="188">
<typeStatus id="54CA880A8168363728A7FB20405F0DA1" box="[234,348,1207,1233]" pageId="29" pageNumber="188" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCE36A88168363728F0FB4C45B00D85" blockId="29.[189,1203,1207,1270]" box="[189,1203,1243,1270]" pageId="29" pageNumber="188">
<collectingCountry id="F36676388168363728F0FB4C41F50D85" box="[189,246,1243,1269]" name="United States of America" pageId="29" pageNumber="188">USA</collectingCountry>
• 
<specimenCount id="9D77FD21816836372942FB4B402B0D86" box="[271,296,1244,1270]" pageId="29" pageNumber="188" type="male">♂</specimenCount>
; 
<collectingRegion id="49B5F84A816836372979FB4C40940D85" box="[308,407,1243,1269]" country="United States of America" name="Virginia" pageId="29" pageNumber="188">Virginia</collectingRegion>
, 
<collectingCounty id="62AF4E248168363729EDFB4C43360D85" box="[416,565,1243,1269]" pageId="29" pageNumber="188">Augusta Co.</collectingCounty>
, 
<location id="8EAE6073816836372A0FFB4B43DC0D85" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372A0FFB4B43DC0D85" box="[578,735,1243,1270]" country="United States of America" county="Augusta Co." name="Mount Solon" pageId="29" pageNumber="188" stateProvince="Virginia">Mount Solon</location>
; 
<date id="FFCF1068816836372AA6FB4C427D0D86" box="[747,894,1243,1270]" pageId="29" pageNumber="188" value="1950-07-11">
<collectingDate id="EF8BE980816836372AA6FB4C427D0D86" box="[747,894,1243,1270]" pageId="29" pageNumber="188" value="1950-07-11">11 Jul. 1950</collectingDate>
</date>
; 
<collectorName id="2684537E816836372BC4FB4B45170D85" box="[905,1044,1243,1270]" pageId="29" pageNumber="188">W.W. Wirth</collectorName>
leg.; 
<collectionCode id="ED60AE6D816836372C1BFB4C45AC0D85" box="[1110,1199,1243,1269]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8BCE36A88168363728A7FA8940A50C48" blockId="29.[189,1398,1310,1446]" box="[234,422,1310,1336]" pageId="29" pageNumber="188">
<emphasis id="B905EABA8168363728A7FA8940A50C48" bold="true" box="[234,422,1310,1336]" pageId="29" pageNumber="188">Other material</emphasis>
</paragraph>
<paragraph id="8BCE36A88168363728F0FAD443AD0CD6" blockId="29.[189,1398,1310,1446]" pageId="29" pageNumber="188">
<materialsCitation id="3B193CF58168363728F0FAD440000CF1" ID-GBIF-Occurrence="4135858302" collectingDate="1984-07-09" collectionCode="CNCI" collectorName="A. Borkent" country="Canada" county="Elgin Co." location="Springwater Conservation Area" pageId="29" pageNumber="188" specimenCount="1" specimenCount-female="1" stateProvince="Ontario">
<collectingCountry id="F36676388168363728F0FAD4403A0C2D" box="[189,313,1347,1373]" name="Canada" pageId="29" pageNumber="188">CANADA</collectingCountry>
• 
<specimenCount id="9D77FD21816836372914FAD4408A0C2D" box="[345,393,1347,1373]" pageId="29" pageNumber="188" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="49B5F84A8168363729D6FAD540FB0C2C" box="[411,504,1346,1372]" country="Canada" name="Ontario" pageId="29" pageNumber="188">Ontario</collectingRegion>
, 
<collectingCounty id="62AF4E24816836372A4AFAD5437E0C2D" box="[519,637,1346,1373]" pageId="29" pageNumber="188">Elgin Co.</collectingCounty>
, 
<location id="8EAE6073816836372AC0FAD545090C2D" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372AC0FAD545090C2D" box="[653,1034,1346,1373]" country="Canada" county="Elgin Co." name="Springwater Conservation Area" pageId="29" pageNumber="188" stateProvince="Ontario">Springwater Conservation Area</location>
; 
<date id="FFCF1068816836372C54FAD445A70C2D" box="[1049,1188,1346,1373]" pageId="29" pageNumber="188" value="1984-07-09">
<collectingDate id="EF8BE980816836372C54FAD445A70C2D" box="[1049,1188,1346,1373]" pageId="29" pageNumber="188" value="1984-07-09">9 Jul. 1984</collectingDate>
</date>
; 
<collectorName id="2684537E816836372CFFFAD4443A0C2C" box="[1202,1337,1346,1373]" pageId="29" pageNumber="188">A. Borkent</collectorName>
leg.; 
<collectionCode id="ED60AE6D8168363728F0FAF040000CF1" box="[189,259,1383,1409]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="29" pageNumber="188">CNCI</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF581683637296EFAF043AD0CD6" ID-GBIF-Occurrence="4135858346" collectingDate="1985-06-27" collectionCode="CNCI" collectorName="L. Forster" location="Gatineau Park" municipality="Outaouais Region" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Quebec">
<specimenCount id="9D77FD2181683637296EFAF040560CF2" box="[291,341,1383,1410]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<collectingRegion id="49B5F84A816836372928FAF040C10CF1" box="[357,450,1383,1409]" country="Canada" name="Quebec" pageId="29" pageNumber="188">Quebec</collectingRegion>
, 
<collectingMunicipality id="6BAAACD281683637299DFAF043A90CF1" box="[464,682,1383,1409]" pageId="29" pageNumber="188">Outaouais Region</collectingMunicipality>
, 
<location id="8EAE6073816836372AF4FAF042650CF1" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372AF4FAF042650CF1" box="[697,870,1383,1409]" municipality="Outaouais Region" name="Gatineau Park" pageId="29" pageNumber="188" stateProvince="Quebec">Gatineau Park</location>
, 
<location id="8EAE6073816836372B38FAF045010CF1" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372B38FAF045010CF1" box="[885,1026,1383,1409]" municipality="Outaouais Region" name="Black Lake" pageId="29" pageNumber="188" stateProvince="Quebec">Black Lake</location>
; 
<date id="FFCF1068816836372C5CFAF045B10CF2" box="[1041,1202,1383,1410]" pageId="29" pageNumber="188" value="1985-06-27">
<collectingDate id="EF8BE980816836372C5CFAF045B10CF2" box="[1041,1202,1383,1410]" pageId="29" pageNumber="188" value="1985-06-27">27 Jun. 1985</collectingDate>
</date>
; 
<collectorName id="2684537E816836372C8CFAFF443B0CF2" box="[1217,1336,1384,1410]" pageId="29" pageNumber="188">L. Forster</collectorName>
leg.; originally identified as 
<taxonomicName id="4C714D2B816836372981FA1B43530CD6" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[460,592,1420,1446]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372981FA1B43530CD6" box="[460,592,1420,1446]" italics="true" pageId="29" pageNumber="188">C. blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372A2DFA1B43A90CD6" box="[608,682,1420,1446]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="29" pageNumber="188">CNCI</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCE36A88168363728F0FA5845B00E98" blockId="29.[189,1399,1486,2025]" pageId="29" pageNumber="188">
<materialsCitation id="3B193CF58168363728F0FA5845B00C99" ID-GBIF-Occurrence="4135858321" box="[189,1203,1486,1513]" collectingDate="1967-04-20" collectionCode="USNM" collectorName="W. W. Wirth" country="United States of America" county="Alachua Co." location="Gainesville" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Florida">
<collectingCountry id="F36676388168363728F0FA5841F50C99" box="[189,246,1487,1513]" name="United States of America" pageId="29" pageNumber="188">USA</collectingCountry>
• – 
<emphasis id="B905EABA81683637296CFA59407D0C98" bold="true" box="[289,382,1486,1512]" pageId="29" pageNumber="188">
<collectingRegion id="49B5F84A81683637296CFA59407D0C98" box="[289,382,1486,1512]" country="United States of America" name="Florida" pageId="29" pageNumber="188">Florida</collectingRegion>
</emphasis>
<specimenCount id="9D77FD218168363729C8FA5840B00C99" box="[389,435,1487,1513]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<collectingCounty id="62AF4E248168363729F0FA5943520C99" box="[445,593,1486,1513]" pageId="29" pageNumber="188">Alachua Co.</collectingCounty>
, 
<location id="8EAE6073816836372A11FA5943E60C98" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372A11FA5943E60C98" box="[604,741,1486,1512]" country="United States of America" county="Alachua Co." name="Gainesville" pageId="29" pageNumber="188" stateProvince="Florida">Gainesville</location>
; 
<date id="FFCF1068816836372ABDFA5842890C98" box="[752,906,1486,1513]" pageId="29" pageNumber="188" value="1967-04-20">
<collectingDate id="EF8BE980816836372ABDFA5842890C98" box="[752,906,1486,1513]" pageId="29" pageNumber="188" value="1967-04-20">20 Apr. 1967</collectingDate>
</date>
; 
<collectorName id="2684537E816836372BD9FA58451D0C98" box="[916,1054,1486,1513]" pageId="29" pageNumber="188">W.W. Wirth</collectorName>
leg.; 
<collectionCode id="ED60AE6D816836372C13FA5845B00C99" box="[1118,1203,1487,1513]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372C87FA5840060F42" ID-GBIF-Occurrence="4135858378" collectingDate="1983-01-21" collectingMethod="light trap with CO 2" collectionCode="FSCA" collectorName="A. Wilkening" county="Levy Co." location="Yankeetown" pageId="29" pageNumber="188" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D77FD21816836372C87FA5845F50C99" box="[1226,1270,1487,1513]" pageId="29" pageNumber="188" type="female">1 ♀</specimenCount>
; 
<collectingCounty id="62AF4E24816836372D4EFA5844720C99" box="[1283,1393,1487,1513]" pageId="29" pageNumber="188">Levy Co.</collectingCounty>
, 
<location id="8EAE60738168363728F0FA6440510F7D" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE60738168363728F0FA6440510F7D" box="[189,338,1523,1549]" county="Levy Co." name="Yankeetown" pageId="29" pageNumber="188">Yankeetown</location>
; 
<date id="FFCF1068816836372912FA6440FA0F7D" box="[351,505,1523,1550]" pageId="29" pageNumber="188" value="1983-01-21">
<collectingDate id="EF8BE980816836372912FA6440FA0F7D" box="[351,505,1523,1550]" pageId="29" pageNumber="188" value="1983-01-21">21 Jan. 1983</collectingDate>
</date>
; 
<collectorName id="2684537E816836372A49FA6443A00F7D" box="[516,675,1523,1549]" pageId="29" pageNumber="188">A. Wilkening</collectorName>
leg.; 
<collectingMethod id="52304EBF816836372AABFA6442CB0F64" box="[742,968,1523,1556]" pageId="29" pageNumber="188">
light trap with CO 
<subScript id="17F534ED816836372BF2F99142CB0F64" attach="left" box="[959,968,1542,1556]" fontSize="6" pageId="29" pageNumber="188">2</subScript>
</collectingMethod>
; originally identified as 
<taxonomicName id="4C714D2B816836372CA4FA64446D0F7D" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[1257,1390,1523,1549]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372CA4FA64446D0F7D" box="[1257,1390,1523,1549]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D8168363728F0F98F40060F42" box="[189,261,1560,1586]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="29" pageNumber="188" type="Museum">FSCA</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372952F98F44740F42" ID-GBIF-Occurrence="4135858375" box="[287,1399,1559,1586]" collectingDate="1971-05-15" collectionCode="USNM" collectorName="G. B. Fairchild &amp; B. L. Trap" county="Liberty Co." location="Torreya State Park" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816836372952F98F404C0F42" box="[287,335,1560,1586]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<collectingCounty id="62AF4E24816836372910F98040E60F42" box="[349,485,1559,1586]" pageId="29" pageNumber="188">Liberty Co.</collectingCounty>
<location id="8EAE60738168363729A0F98F43CE0F41" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE60738168363729A0F98F43CE0F41" box="[493,717,1559,1586]" county="Liberty Co." name="Torreya State Park" pageId="29" pageNumber="188">Torreya State Park</location>
; 
<date id="FFCF1068816836372A94F98042790F42" box="[729,890,1559,1586]" pageId="29" pageNumber="188" value="1971-05-15">
<collectingDate id="EF8BE980816836372A94F98042790F42" box="[729,890,1559,1586]" pageId="29" pageNumber="188" value="1971-05-15">15 May 1971</collectingDate>
</date>
; 
<collectorName id="2684537E816836372BCAF98F45320F41" box="[903,1073,1559,1586]" pageId="29" pageNumber="188">G.B. Fairchild</collectorName>
and 
<collectorName id="2684537E816836372C21F98F45DD0F42" box="[1132,1246,1560,1586]" pageId="29" pageNumber="188">B.L. Trap</collectorName>
leg.; 
<collectionCode id="ED60AE6D816836372D6FF98F44740F42" box="[1314,1399,1560,1586]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF581683637289CF9AB451A0F26" ID-GBIF-Occurrence="4135858324" box="[209,1049,1596,1623]" collectingDate="1979-05-13" collectingDateMax="1979-05-14" collectingDateMin="1979-05-13" collectionCode="CNCI" collectorName="J. A. Downes" county="Wakulla Co." location="Sopchoppy" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD2181683637289CF9AB40010F27" box="[209,258,1596,1623]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<collectingCounty id="62AF4E2481683637295DF9AB40A50F26" box="[272,422,1596,1622]" pageId="29" pageNumber="188">Wakulla Co.</collectingCounty>
, 
<location id="8EAE60738168363729F8F9AB433E0F26" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE60738168363729F8F9AB433E0F26" box="[437,573,1596,1622]" county="Wakulla Co." name="Sopchoppy" pageId="29" pageNumber="188">Sopchoppy</location>
; 
<date id="FFCF1068816836372A07F9AB42180F26" box="[586,795,1596,1623]" pageId="29" pageNumber="188" value="1979-05-13" valueMax="1979-05-14" valueMin="1979-05-13">
<collectingDate id="EF8BE980816836372A07F9AB42180F26" box="[586,795,1596,1623]" pageId="29" pageNumber="188" value="1979-05-13" valueMax="1979-05-14" valueMin="1979-05-13">13–14 May 1979</collectingDate>
</date>
; 
<collectorName id="2684537E816836372B64F9AB42C50F27" box="[809,966,1596,1623]" pageId="29" pageNumber="188">J.A. Downes</collectorName>
; 
<collectionCode id="ED60AE6D816836372B9EF9AB451A0F26" box="[979,1049,1596,1622]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="29" pageNumber="188">CNCI</collectionCode>
</materialsCitation>
– 
<materialsCitation id="3B193CF5816836372C76F9AB40C50FEF" ID-GBIF-Occurrence="4135858351" collectingDate="1976-04-28" collectionCode="FSCA" collectorName="W. L. Grogan" county="Prince Georges Co." location="Patuxent Wildlife Research Center" pageId="29" pageNumber="188" specimenCount="2" specimenCount-male="2" stateProvince="Maryland">
<emphasis id="B905EABA816836372C76F9AB45BB0F26" bold="true" box="[1083,1208,1596,1622]" pageId="29" pageNumber="188">
<collectingRegion id="49B5F84A816836372C76F9AB45BB0F26" box="[1083,1208,1596,1622]" country="United States of America" name="Maryland" pageId="29" pageNumber="188">Maryland</collectingRegion>
</emphasis>
• 
<specimenCount id="9D77FD21816836372C9BF9AB441E0F27" box="[1238,1309,1596,1623]" pageId="29" pageNumber="188" type="male">2 ♂♂</specimenCount>
; 
<collectingCounty id="62AF4E24816836372D66F9AB40510F0B" pageId="29" pageNumber="188">Prince Georges Co.</collectingCounty>
, 
<location id="8EAE6073816836372912F9F643F80F0B" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372912F9F643F80F0B" box="[351,763,1633,1659]" county="Prince Georges Co." name="Patuxent Wildlife Research Center" pageId="29" pageNumber="188" stateProvince="Maryland">Patuxent Wildlife Research Center</location>
; 
<date id="FFCF1068816836372B4AF9F642A60F0B" box="[775,933,1633,1659]" pageId="29" pageNumber="188" value="1976-04-28">
<collectingDate id="EF8BE980816836372B4AF9F642A60F0B" box="[775,933,1633,1659]" pageId="29" pageNumber="188" value="1976-04-28">28 Apr. 1976</collectingDate>
</date>
; 
<collectorName id="2684537E816836372BFCF9F645490F0B" box="[945,1098,1633,1659]" pageId="29" pageNumber="188">W.L. Grogan</collectorName>
leg.; originally identified as 
<taxonomicName id="4C714D2B816836372893F911406E0FEF" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[222,365,1669,1696]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372893F911406E0FEF" box="[222,365,1669,1696]" italics="true" pageId="29" pageNumber="188">C. blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372933F91240C50FEF" box="[382,454,1669,1695]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="29" pageNumber="188" type="Museum">FSCA</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF58168363729AFF91245B60FEF" ID-GBIF-Occurrence="4135858327" box="[482,1205,1669,1696]" collectingDate="1976-05-06" collectionCode="FSCA" pageId="29" pageNumber="188" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D77FD218168363729AFF91243130FD0" box="[482,528,1669,1696]" pageId="29" pageNumber="188" type="female">1 ♀</specimenCount>
; 
<date id="FFCF1068816836372A6DF91243B60FEF" box="[544,693,1669,1696]" pageId="29" pageNumber="188" value="1976-05-06">
<collectingDate id="EF8BE980816836372A6DF91243B60FEF" box="[544,693,1669,1696]" pageId="29" pageNumber="188" value="1976-05-06">6 May 1976</collectingDate>
</date>
; originally identified as 
<taxonomicName id="4C714D2B816836372B9AF911455F0FEF" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[983,1116,1669,1696]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372B9AF911455F0FEF" box="[983,1116,1669,1696]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372C20F91245B60FEF" box="[1133,1205,1669,1695]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="29" pageNumber="188" type="Museum">FSCA</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372C9CF91244010FB4" ID-GBIF-Occurrence="4135858328" collectingDate="1978-05-10" collectionCode="USNM" collectorName="W. W. Wirth" location="Patuxent Wildlife Refuge" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816836372C9CF91244010FD0" box="[1233,1282,1669,1696]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<location id="8EAE6073816836372D5DF911407C0FB4" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372D5DF911407C0FB4" name="Patuxent Wildlife Refuge" pageId="29" pageNumber="188">Patuxent Wildlife Refuge</location>
; 
<date id="FFCF10688168363729C7F93D432F0FB4" box="[394,556,1706,1732]" pageId="29" pageNumber="188" value="1978-05-10">
<collectingDate id="EF8BE9808168363729C7F93D432F0FB4" box="[394,556,1706,1732]" pageId="29" pageNumber="188" value="1978-05-10">10 May 1978</collectingDate>
</date>
; 
<collectorName id="2684537E816836372A75F93D43C00FB4" box="[568,707,1706,1732]" pageId="29" pageNumber="188">W.W. Wirth</collectorName>
leg.; originally identified as 
<taxonomicName id="4C714D2B816836372C55F93D459E0FB4" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[1048,1181,1706,1732]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372C55F93D459E0FB4" box="[1048,1181,1706,1732]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372CE0F93D44010FB4" box="[1197,1282,1706,1732]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372D51F93D424C0F99" ID-GBIF-Occurrence="4135858343" collectingDate="1979-05-12" collectionCode="USNM" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816836372D51F93D444F0FB4" box="[1308,1356,1706,1732]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<date id="FFCF1068816836372D14F93D40380F99" pageId="29" pageNumber="188" value="1979-05-12">
<collectingDate id="EF8BE980816836372D14F93D40380F99" pageId="29" pageNumber="188" value="1979-05-12">12 May 1979</collectingDate>
</date>
; originally identified as 
<taxonomicName id="4C714D2B816836372A2CF95843EB0F98" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[609,744,1742,1769]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372A2CF95843EB0F98" box="[609,744,1742,1769]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372AB7F958424C0F99" box="[762,847,1743,1769]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372B23F95843C40E7D" ID-GBIF-Occurrence="4135858372" collectingDate="1979-05-15" collectingDateMax="1979-05-21" collectingDateMin="1979-05-15" collectingMethod="Malaise trap" collectionCode="FSCA" collectorName="W. L. Grogan" county="Wicomico Co." location="Salisbury" municipality="Salisbury" pageId="29" pageNumber="188" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D77FD21816836372B23F958429E0F99" box="[878,925,1743,1769]" pageId="29" pageNumber="188" type="female">1 ♀</specimenCount>
; 
<collectingCounty id="62AF4E24816836372BE3F959455E0F99" box="[942,1117,1742,1769]" pageId="29" pageNumber="188">Wicomico Co.</collectingCounty>
, 
<collectingMunicipality id="6BAAACD2816836372C21F95945DC0F98" box="[1132,1247,1742,1768]" pageId="29" pageNumber="188">Salisbury</collectingMunicipality>
; 
<date id="FFCF1068816836372CA0F95941F80E7D" pageId="29" pageNumber="188" value="1979-05-15" valueMax="1979-05-21" valueMin="1979-05-15">
<collectingDate id="EF8BE980816836372CA0F95941F80E7D" pageId="29" pageNumber="188" value="1979-05-15" valueMax="1979-05-21" valueMin="1979-05-15">15–21 May 1979</collectingDate>
</date>
; 
<collectorName id="2684537E81683637294BF964409D0E7D" box="[262,414,1779,1805]" pageId="29" pageNumber="188">W.L. Grogan</collectorName>
leg.; 
<collectingMethod id="52304EBF816836372992F96443770E7D" box="[479,628,1779,1805]" pageId="29" pageNumber="188">Malaise trap</collectingMethod>
; 
<collectionCode id="ED60AE6D816836372A32F96443C40E7D" box="[639,711,1779,1805]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="29" pageNumber="188" type="Museum">FSCA</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372A92F96443B60E42" ID-GBIF-Occurrence="4135858363" collectingDate="1968-05-19" collectionCode="USNM" collectorName="W. H. Anderson" county="Worcester Co." location="Snow Hill" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D77FD21816836372A92F96442090E7D" box="[735,778,1779,1805]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
(
<typeStatus id="54CA880A816836372B54F96442810E7D" box="[793,898,1779,1805]" pageId="29" pageNumber="188" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C714D2B816836372BEAF96445280E7D" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[935,1067,1779,1805]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372BEAF96445280E7D" box="[935,1067,1779,1805]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
); 
<collectingCounty id="62AF4E24816836372C0EF96445E80E7D" box="[1091,1259,1779,1805]" pageId="29" pageNumber="188">Worcester Co.</collectingCounty>
, 
<location id="8EAE6073816836372CBAF96444710E7D" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372CBAF96444710E7D" box="[1271,1394,1779,1805]" county="Worcester Co." name="Snow Hill" pageId="29" pageNumber="188">Snow Hill</location>
; 
<date id="FFCF10688168363728F0F88F405E0E41" box="[189,349,1815,1842]" pageId="29" pageNumber="188" value="1968-05-19">
<collectingDate id="EF8BE9808168363728F0F88F405E0E41" box="[189,349,1815,1842]" pageId="29" pageNumber="188" value="1968-05-19">19 May 1968</collectingDate>
</date>
; 
<collectorName id="2684537E816836372924F88F431E0E41" box="[361,541,1815,1842]" pageId="29" pageNumber="188">W.H. Anderson</collectorName>
leg.; 
<collectionCode id="ED60AE6D816836372A2DF88F43B60E42" box="[608,693,1816,1842]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
– 
<materialsCitation id="3B193CF5816836372A9FF88F421C0E26" ID-GBIF-Occurrence="4135858374" collectingDate="1986-06-18" collectionCode="USNM" collectorName="W. W. Wirth" county="Jackson Co." location="Cashiers" municipality="Dunaley" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
<emphasis id="B905EABA816836372A9FF88F42910E41" bold="true" box="[722,914,1815,1842]" pageId="29" pageNumber="188">
<collectingRegion id="49B5F84A816836372A9FF88F42910E41" box="[722,914,1815,1842]" country="United States of America" name="North Carolina" pageId="29" pageNumber="188">North Carolina</collectingRegion>
</emphasis>
• 
<specimenCount id="9D77FD21816836372BE1F88F42D80E42" box="[940,987,1816,1842]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<collectingCounty id="62AF4E24816836372BA5F880457B0E42" box="[1000,1144,1815,1842]" pageId="29" pageNumber="188">Jackson Co.</collectingCounty>
, 
<collectingMunicipality id="6BAAACD2816836372CC9F88045E40E41" box="[1156,1255,1815,1841]" pageId="29" pageNumber="188">Dunaley</collectingMunicipality>
bog, 
<locationDeviation id="9A7C2F8A816836372D67F88F41D60E26" bearing="180.0" distance="11265.408" pageId="29" pageNumber="188">
<quantity id="4C899B4D816836372D67F88F445C0E41" box="[1322,1375,1815,1842]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="29" pageNumber="188" unit="mi" value="7.0">7 mi</quantity>
S of
</locationDeviation>
<location id="8EAE6073816836372893F8AB40440E26" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372893F8AB40440E26" box="[222,327,1852,1878]" county="Jackson Co." municipality="Dunaley" name="Cashiers" pageId="29" pageNumber="188" stateProvince="North Carolina">Cashiers</location>
; 
<date id="FFCF106881683637291EF8AB40ED0E26" box="[339,494,1852,1879]" pageId="29" pageNumber="188" value="1986-06-18">
<collectingDate id="EF8BE98081683637291EF8AB40ED0E26" box="[339,494,1852,1879]" pageId="29" pageNumber="188" value="1986-06-18">18 Jun. 1986</collectingDate>
</date>
; 
<collectorName id="2684537E8168363729B6F8AA43850E26" box="[507,646,1852,1879]" pageId="29" pageNumber="188">W.W. Wirth</collectorName>
leg.; 
<collectionCode id="ED60AE6D816836372A87F8AB421C0E26" box="[714,799,1852,1878]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372B74F8AB43950E0B" ID-GBIF-Occurrence="4135858319" collectingDate="1986-06-21" collectionCode="USNM" collectorName="W. W. Wirth" county="Macon Co." location="Wightman Cottage" municipality="Highlands" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816836372B74F8AB426A0E27" box="[825,873,1852,1879]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<collectingCounty id="62AF4E24816836372B3BF8AA42FF0E26" box="[886,1020,1852,1879]" pageId="29" pageNumber="188">Macon Co.</collectingCounty>
, 
<collectingMunicipality id="6BAAACD2816836372C44F8AB45860E26" box="[1033,1157,1852,1878]" pageId="29" pageNumber="188">Highlands</collectingMunicipality>
, 
<location id="8EAE6073816836372CDDF8AB44710E26" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372CDDF8AB44710E26" box="[1168,1394,1852,1878]" county="Macon Co." municipality="Highlands" name="Wightman Cottage" pageId="29" pageNumber="188">Wightman Cottage</location>
; 
<date id="FFCF10688168363728F0F8F6405E0E0A" box="[189,349,1888,1915]" pageId="29" pageNumber="188" value="1986-06-21">
<collectingDate id="EF8BE9808168363728F0F8F6405E0E0A" box="[189,349,1888,1915]" pageId="29" pageNumber="188" value="1986-06-21">21 Jun. 1986</collectingDate>
</date>
; 
<collectorName id="2684537E816836372926F8F640FA0E0B" box="[363,505,1889,1915]" pageId="29" pageNumber="188">W.W. Wirth</collectorName>
leg.; 
<collectionCode id="ED60AE6D816836372A0CF8F643950E0B" box="[577,662,1889,1915]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372AF9F8F6445E0E0B" ID-GBIF-Occurrence="4135858359" box="[692,1373,1888,1915]" collectingDate="1957-06-03" collectionCode="CNCI" collectorName="J. R. Vockeroth" location="Highlands" municipality="Highlands" pageId="29" pageNumber="188" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D77FD21816836372AF9F8F643E70E0B" box="[692,740,1889,1915]" pageId="29" pageNumber="188" type="female">1 ♀</specimenCount>
; 
<collectingMunicipality id="6BAAACD2816836372AB8F8F642720E0B" box="[757,881,1889,1915]" pageId="29" pageNumber="188">Highlands</collectingMunicipality>
; 
<date id="FFCF1068816836372BCDF8F645120E0B" box="[896,1041,1888,1915]" pageId="29" pageNumber="188" value="1957-06-03">
<collectingDate id="EF8BE980816836372BCDF8F645120E0B" box="[896,1041,1888,1915]" pageId="29" pageNumber="188" value="1957-06-03">3 Jun. 1957</collectingDate>
</date>
; 
<collectorName id="2684537E816836372C6DF8F645D30E0B" box="[1056,1232,1889,1915]" pageId="29" pageNumber="188">J.R. Vockeroth</collectorName>
leg.; 
<collectionCode id="ED60AE6D816836372D5AF8F6445E0E0B" box="[1303,1373,1889,1915]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="29" pageNumber="188">CNCI</collectionCode>
</materialsCitation>
– 
<materialsCitation id="3B193CF58168363728F0F81240A70EB4" ID-GBIF-Occurrence="4135858371" collectingDate="1959-05-04" collectionCode="USNM" collectorName="W. W. Wirth" location="Falls Church" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<emphasis id="B905EABA8168363728F0F81240270EEF" bold="true" box="[189,292,1925,1951]" pageId="29" pageNumber="188">
<collectingRegion id="49B5F84A8168363728F0F81240270EEF" box="[189,292,1925,1951]" country="United States of America" name="Virginia" pageId="29" pageNumber="188">Virginia</collectingRegion>
</emphasis>
• 
<specimenCount id="9D77FD21816836372973F812406D0ED0" box="[318,366,1925,1952]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<location id="8EAE6073816836372936F81243110EEF" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE6073816836372936F81243110EEF" box="[379,530,1925,1951]" name="Falls Church" pageId="29" pageNumber="188" stateProvince="Virginia">Falls Church</location>
[independent city]; 
<date id="FFCF1068816836372B4CF81242900ED0" box="[769,915,1925,1952]" pageId="29" pageNumber="188" value="1959-05-04">
<collectingDate id="EF8BE980816836372B4CF81242900ED0" box="[769,915,1925,1952]" pageId="29" pageNumber="188" value="1959-05-04">4 May 1959</collectingDate>
</date>
; 
<collectorName id="2684537E816836372BD2F81145290EEF" box="[927,1066,1925,1952]" pageId="29" pageNumber="188">W.W. Wirth</collectorName>
leg.; originally identified as 
<taxonomicName id="4C714D2B8168363728F0F83D40420EB4" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[189,321,1962,1988]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA8168363728F0F83D40420EB4" box="[189,321,1962,1988]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372902F83D40A70EB4" box="[335,420,1962,1988]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF58168363729F1F83D45840EB4" ID-GBIF-Occurrence="4135858356" box="[444,1159,1962,1988]" collectingDate="1960-05-07" collectionCode="USNM" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD218168363729F1F83D40E90EB4" box="[444,490,1962,1988]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<date id="FFCF10688168363729BBF83D43850EB4" box="[502,646,1962,1988]" pageId="29" pageNumber="188" value="1960-05-07">
<collectingDate id="EF8BE9808168363729BBF83D43850EB4" box="[502,646,1962,1988]" pageId="29" pageNumber="188" value="1960-05-07">7 May 1960</collectingDate>
</date>
; originally identified as 
<taxonomicName id="4C714D2B816836372BEDF83D45200EB4" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[928,1059,1962,1988]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372BEDF83D45200EB4" box="[928,1059,1962,1988]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372C7FF83D45840EB4" box="[1074,1159,1962,1988]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B193CF5816836372CD2F83D45AC0E98" ID-GBIF-Occurrence="4135858301" collectingDate="1962-05" collectionCode="USNM" collectorName="W. W. Wirth" location="Holmes Run" municipality="Falls Church" pageId="29" pageNumber="188" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D77FD21816836372CD2F83D45CE0EB4" box="[1183,1229,1962,1988]" pageId="29" pageNumber="188" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BAAACD2816836372C94F83D44710EB4" box="[1241,1394,1962,1988]" pageId="29" pageNumber="188">Falls Church</collectingMunicipality>
, 
<location id="8EAE60738168363728F0F85940560E99" LSID="urn:lsid:plazi:treatment:03D887BE8168360A2A55FE9F44600BDC:8EAE60738168363728F0F85940560E99" box="[189,341,1998,2025]" municipality="Falls Church" name="Holmes Run" pageId="29" pageNumber="188">Holmes Run</location>
; 
<date id="FFCF106881683637292DF85840D80E98" box="[352,475,1998,2025]" pageId="29" pageNumber="188" value="1962-05">
<collectingDate id="EF8BE98081683637292DF85840D80E98" box="[352,475,1998,2025]" pageId="29" pageNumber="188" value="1962-05">May 1962</collectingDate>
</date>
; 
<collectorName id="2684537E8168363729AAF85843720E98" box="[487,625,1998,2025]" pageId="29" pageNumber="188">W.W. Wirth</collectorName>
leg.; originally identified as 
<taxonomicName id="4C714D2B816836372B8EF85845440E98" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[963,1095,1998,2025]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="188" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816836372B8EF85845440E98" box="[963,1095,1998,2025]" italics="true" pageId="29" pageNumber="188">C.blantoni</emphasis>
</taxonomicName>
; 
<collectionCode id="ED60AE6D816836372C1BF85945AC0E98" box="[1110,1199,1998,2024]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="188" type="Museum">USNM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0E6620816B363428F0F88943190E99" ID-DOI="http://doi.org/10.5281/zenodo.8083812" ID-Zenodo-Dep="8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="30" pageNumber="189" startId="30.[189,232,1822,1848]" targetBox="[225,1362,265,1749]" targetPageId="30">
<paragraph id="8BCE36A8816B363428F0F88943190E99" blockId="30.[189,1399,1822,2025]" pageId="30" pageNumber="189">
<emphasis id="B905EABA816B363428F0F88940190E48" bold="true" box="[189,282,1822,1848]" pageId="30" pageNumber="189">Fig. 12.</emphasis>
Female sternite 9 II. 
<emphasis id="B905EABA816B36342A5CF88943230E48" bold="true" box="[529,544,1822,1848]" pageId="30" pageNumber="189">a</emphasis>
. 
<taxonomicName id="4C714D2B816B36342A63F88942660E48" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[558,869,1822,1848]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="189" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov.">
<emphasis id="B905EABA816B36342A63F88942660E48" box="[558,869,1822,1848]" italics="true" pageId="30" pageNumber="189">Ceratoculicoides pacificus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C1816B36342B21F88842C50E49" box="[876,966,1823,1849]" pageId="30" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (CNCI), Carmanah Valley, British Columbia, Canada. 
<emphasis id="B905EABA816B363429BBF8D643050E2B" bold="true" box="[502,518,1857,1883]" pageId="30" pageNumber="189">b</emphasis>
. 
<taxonomicName id="4C714D2B816B36342A58F8D5439E0E2B" authorityName="Fasbender" authorityYear="2023" box="[533,669,1857,1884]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="189" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B905EABA816B36342A58F8D5439E0E2B" box="[533,669,1857,1884]" italics="true" pageId="30" pageNumber="189">C. pacificus</emphasis>
</taxonomicName>
paratype (CNCI), East Sooke Peak, British Columbia, Canada. 
<emphasis id="B905EABA816B363428F0F8F241C90E0F" bold="true" box="[189,202,1893,1919]" pageId="30" pageNumber="189">c</emphasis>
. 
<taxonomicName id="4C714D2B816B36342897F8F2432E0E0F" authority="(Wirth, 1951)" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[218,557,1892,1919]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="189" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA816B36342897F8F240810E0E" box="[218,386,1893,1919]" italics="true" pageId="30" pageNumber="189">C.virginianus</emphasis>
(
<bibRefCitation id="EFE04B59816B363429DCF8F343270E0F" author="Wirth W. W." box="[401,548,1892,1919]" pageId="30" pageNumber="189" pagination="313 - 326" refId="ref23778" refString="Wirth W. W. 1951. New species and records of Virginia Heleidae. Proceedings of the Entomological Society of Washington 53: 313 - 326." type="journal article" year="1951">Wirth, 1951</bibRefCitation>
)
</taxonomicName>
, Springwater Conservation Area, Ontario, Canada. 
<emphasis id="B905EABA816B36342CC3F8F2459D0E0F" bold="true" box="[1166,1182,1893,1919]" pageId="30" pageNumber="189">d</emphasis>
. 
<taxonomicName id="4C714D2B816B36342CE3F8F341F90ED2" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="189" phylum="Arthropoda" rank="species" species="undefined-f">
<emphasis id="B905EABA816B36342CE3F8F344750E0E" box="[1198,1398,1892,1918]" italics="true" pageId="30" pageNumber="189">Ceratoculicoides</emphasis>
sp. F
</taxonomicName>
1, Tuparro National Park, Colombia. 
<emphasis id="B905EABA816B36342AEAF81F43B70ED2" bold="true" box="[679,692,1928,1954]" pageId="30" pageNumber="189">e</emphasis>
. 
<taxonomicName id="4C714D2B816B36342A8EF81F42CE0ED2" box="[707,973,1928,1954]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="189" phylum="Arthropoda" rank="species" species="undefined-f">
<emphasis id="B905EABA816B36342A8EF81F42880ED2" box="[707,907,1928,1954]" italics="true" pageId="30" pageNumber="189">Ceratoculicoides</emphasis>
sp. F
</taxonomicName>
2, Tuparro National Park, Colombia. 
<emphasis id="B905EABA816B363428F0F83C41C40EB5" bold="true" box="[189,199,1963,1989]" pageId="30" pageNumber="189">f</emphasis>
. 
<taxonomicName id="4C714D2B816B3634289AF83C40E50EB5" box="[215,486,1963,1989]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="189" phylum="Arthropoda" rank="species" species="undefined-f">
<emphasis id="B905EABA816B3634289AF83C409C0EB5" box="[215,415,1963,1989]" italics="true" pageId="30" pageNumber="189">Ceratoculicoides</emphasis>
sp. F
</taxonomicName>
3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm.
</paragraph>
</caption>
<subSubSection id="C36B6523816A363528F0FE9F40B30DC0" pageId="31" pageNumber="190" type="description">
<paragraph id="8BCE36A8816A363528F0FE9F404D0852" blockId="31.[189,334,264,290]" box="[189,334,264,290]" pageId="31" pageNumber="190">
<heading id="D08681C4816A363528F0FE9F404D0852" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="31" pageNumber="190" reason="3">
<emphasis id="B905EABA816A363528F0FE9F404D0852" bold="true" box="[189,334,264,290]" pageId="31" pageNumber="190">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A8816A363528A7FEA040290821" blockId="31.[188,1399,311,480]" box="[234,298,311,337]" pageId="31" pageNumber="190">
<emphasis id="B905EABA816A363528A7FEA040290821" bold="true" box="[234,298,311,337]" pageId="31" pageNumber="190">Male</emphasis>
</paragraph>
<paragraph id="8BCE36A8816A363528F0FECC45E008AF" blockId="31.[188,1399,311,480]" pageId="31" pageNumber="190">
<smallCapsWord id="8D28A074816A363528F0FECC40670803" baselines="366,366" box="[189,356,347,373]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="31" pageNumber="190">MEASUREMENTS</smallCapsWord>
(n = 3). Head width282–295(289); flagellomeres50–54(52), 25–30(28), 22–31(26), 25–28(26), 21–27(25), 22–32(26), 24–28(26), 23–36(27), 29–33(32), 26–37(28), 81–114(103), 63–77(69), 51–69(59); AR 0.74–0.8(0.75); FR 1.63; wing length (n = 10) 0.81–1.21(1.04); wing width (n = 10) 0.25–0.4(0.34); costal ratio (n = 1) 0.49; GCR 1.87–2.17(2.08); GSR 0.97–1.14(1.08); aedeagus ratio 0.66–0.91(0.78).
</paragraph>
<paragraph id="8BCE36A8816A363528F0FD9F43D40B35" blockId="31.[189,1399,519,582]" pageId="31" pageNumber="190">
<smallCapsWord id="8D28A074816A363528F0FD9F401A0B50" baselines="539,539" box="[189,281,520,546]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="31" pageNumber="190">THORAX</smallCapsWord>
. Dorsocentral punctations prominent and interspersed in posterior half of dorsocentral setae. Legs with femora and tibiae yellow or brown.
</paragraph>
<paragraph id="8BCE36A8816A363528F0FDFA45E10B81" blockId="31.[188,1399,621,754]" pageId="31" pageNumber="190">
<smallCapsWord id="8D28A074816A363528F0FDFA40320BF5" baselines="640,640" box="[189,305,621,647]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="31" pageNumber="190">GENITALIA</smallCapsWord>
(
<figureCitation id="134A2A2D816A36352904FDFA40BC0BF7" box="[329,447,621,647]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="31" pageNumber="190">Fig. 9a–c</figureCitation>
). Distal portion of parameres not tapered, apex broadly rounded, with small subacute point displaced laterally. Aedeagus lateral margins convex, rounded, posterior margin straight, notched medially without hyaline incision, posterolateral point triangular, shifted medially nearly to lateral margin of medial notch, curving smoothly into dorsally directed, acute apical point.
</paragraph>
<paragraph id="8BCE36A8816A363528A7FC8E40460A43" blockId="31.[189,1399,793,998]" box="[234,325,793,819]" pageId="31" pageNumber="190">
<emphasis id="B905EABA816A363528A7FC8E40460A43" bold="true" box="[234,325,793,819]" pageId="31" pageNumber="190">Female</emphasis>
</paragraph>
<paragraph id="8BCE36A8816A363528F0FCAA453D0A95" blockId="31.[189,1399,793,998]" pageId="31" pageNumber="190">
<smallCapsWord id="8D28A074816A363528F0FCAA406D0A25" baselines="848,848" box="[189,366,829,855]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="31" pageNumber="190">MEASUREMENTS</smallCapsWord>
(n = 5). Head width (n = 3) 267–306(274); flagellomeres (n = 3) 27–30(27), 17–25(18), 18–25(22), 23–28(26), 24–32(28), 24–32(27), 23–32(31), 24–37(31), 42–52(47), 44–50(47), 48–54(52, 50–54(54), 55–70(66); AR 1.15–1.33(1.29); FR 1.57–1.75(1.69); wing length (n = 6) 0.92–1.29(1.19); wing width (n = 6) 0.34–0.49(0.45); costal ratio 0.56; spermathecal length 52–69(64); spermathecal width 37–56(47); spermathecal neck 12–23(19); spermatheca/neck ratio 0.3.
</paragraph>
<paragraph id="8BCE36A8816A363528F0FB9A42410D57" blockId="31.[189,834,1037,1063]" box="[189,834,1037,1063]" pageId="31" pageNumber="190">
<smallCapsWord id="8D28A074816A363528F0FB9A401A0D55" baselines="1056,1056" box="[189,281,1037,1063]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="31" pageNumber="190">THORAX</smallCapsWord>
. Legs with femora and tibiae yellow or brown.
</paragraph>
<paragraph id="8BCE36A8816A363528F0FBD840B30DC0" blockId="31.[189,1399,1102,1200]" pageId="31" pageNumber="190">
<smallCapsWord id="8D28A074816A363528F0FBD840320D17" baselines="1122,1122" box="[189,305,1103,1129]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="31" pageNumber="190">GENITALIA</smallCapsWord>
(
<figureCitation id="134A2A2D816A3635290EFBD840A60D19" box="[323,421,1103,1129]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="31" pageNumber="190">Fig. 12c</figureCitation>
). 9 
<superScript id="7C049BE0816A36352986FBD940DB0D2C" attach="left" box="[459,472,1102,1116]" fontSize="6" pageId="31" pageNumber="190">th</superScript>
sternite anterior branch truncate or obtusely rounded, apices widely separated; base of posterior branch directed posteriorly, distal half curving medially, tip rounded or pointed. 2 major spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="C36B6523816A363528F0FB4042240C51" box="[189,807,1239,1313]" pageId="31" pageNumber="190" type="distribution">
<paragraph id="8BCE36A8816A363528F0FB4040550D81" blockId="31.[189,342,1239,1265]" box="[189,342,1239,1265]" pageId="31" pageNumber="190">
<heading id="D08681C4816A363528F0FB4040550D81" bold="true" box="[189,342,1239,1265]" fontSize="11" level="3" pageId="31" pageNumber="190" reason="3">
<emphasis id="B905EABA816A363528F0FB4040550D81" bold="true" box="[189,342,1239,1265]" pageId="31" pageNumber="190">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A8816A363528F0FA9042240C51" blockId="31.[189,807,1286,1313]" box="[189,807,1286,1313]" pageId="31" pageNumber="190">
Eastern North America (
<collectingCountry id="F3667638816A36352996FA9143360C50" box="[475,565,1286,1312]" name="Canada" pageId="31" pageNumber="190">Canada</collectingCountry>
and 
<collectingCountry id="F3667638816A36352A23FA9043A90C51" box="[622,682,1287,1313]" name="United States of America" pageId="31" pageNumber="190">USA</collectingCountry>
) (
<figureCitation id="134A2A2D816A36352A8EFA9142150C51" box="[707,790,1286,1313]" captionStart="Fig" captionStartId="38.[189,232,1822,1848]" captionTargetBox="[283,1385,608,1735]" captionTargetId="figure-200@38.[203,1385,608,1735]" captionTargetPageId="38" captionText="Fig. 13. Distribution. a. Western North America. Solid star =Ceratoculicoides aliciae Huerta &amp; Borkent, 2005; solid circles = C.borkenti sp. nov.; open triangles = C.grogani sp. nov.; open star = C. longipennis (Wirth, 1952); open square = C. pacificus sp. nov.; solid triangle = C.propinquus sp. nov.; solid cross C. sp. M1. b. Eastern North America. Solid circles = C. confusus sp. nov.; open squares = C.virginianus (Wirth, 1951). c. Central and northern South America. Solid circle = C.confusus; open squares = C. sp. F1; open triangles = C. sp. F2; solid triangle = C. sp. F3." figureDoi="http://doi.org/10.5281/zenodo.8083814" httpUri="https://zenodo.org/record/8083814/files/figure.png" pageId="31" pageNumber="190">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B6523816A360A28F0FADF44600BDC" lastPageId="32" lastPageNumber="191" pageId="31" pageNumber="190" type="discussion">
<paragraph id="8BCE36A8816A363528F0FADF402C0C12" blockId="31.[189,303,1352,1378]" box="[189,303,1352,1378]" pageId="31" pageNumber="190">
<heading id="D08681C4816A363528F0FADF402C0C12" bold="true" box="[189,303,1352,1378]" fontSize="11" level="3" pageId="31" pageNumber="190" reason="3">
<emphasis id="B905EABA816A363528F0FADF402C0C12" bold="true" box="[189,303,1352,1378]" pageId="31" pageNumber="190">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A8816A363528F0FAE045B70EF3" blockId="31.[188,1401,1399,1923]" pageId="31" pageNumber="190">
<bibRefCitation id="EFE04B59816A363528F0FAE0433C0CE1" author="Wirth W. W. &amp; Ratanaworabhan N. C." box="[189,575,1399,1425]" pageId="31" pageNumber="190" pagination="170 - 177" refId="ref23938" refString="Wirth W. W. &amp; Ratanaworabhan N. C. 1971. Ceratoculicoides, a new genus related to Ceratopogon Meigen (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 73: 170 - 177." type="journal article" year="1971">Wirth &amp; Ratanaworabhan (1971)</bibRefCitation>
described 
<taxonomicName id="4C714D2B816A36352A8CFAEF42460CE1" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[705,837,1399,1426]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816A36352A8CFAEF42460CE1" box="[705,837,1399,1426]" italics="true" pageId="31" pageNumber="190">C. blantoni</emphasis>
</taxonomicName>
, distinguishing it from 
<taxonomicName id="4C714D2B816A36352C12FAEF44000CE1" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[1119,1283,1400,1426]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA816A36352C12FAEF44000CE1" box="[1119,1283,1400,1426]" italics="true" pageId="31" pageNumber="190">C.virginianus</emphasis>
</taxonomicName>
based on leg coloration (brown in 
<taxonomicName id="4C714D2B816A363529AFFA0C43660CC5" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[482,613,1435,1461]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816A363529AFFA0C43660CC5" box="[482,613,1435,1461]" italics="true" pageId="31" pageNumber="190">C.blantoni</emphasis>
</taxonomicName>
, yellow in 
<taxonomicName id="4C714D2B816A36352AA0FA0C42920CC4" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[749,913,1435,1461]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA816A36352AA0FA0C42920CC4" box="[749,913,1435,1461]" italics="true" pageId="31" pageNumber="190">C.virginianus</emphasis>
</taxonomicName>
), orientation of the apex of the paramere (posterolateral in 
<taxonomicName id="4C714D2B816A363529CFFA2843020CA8" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[386,513,1470,1497]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816A363529CFFA2843020CA8" box="[386,513,1470,1497]" italics="true" pageId="31" pageNumber="190">C.blantoni</emphasis>
</taxonomicName>
, ventroposterior in 
<taxonomicName id="4C714D2B816A36352A90FA28427D0CA8" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[733,894,1471,1497]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA816A36352A90FA28427D0CA8" box="[733,894,1471,1497]" italics="true" pageId="31" pageNumber="190">C.virginianus</emphasis>
</taxonomicName>
), and the aedeagal proportions (broader than long in 
<taxonomicName id="4C714D2B816A36352940FA75408F0C8C" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[269,396,1506,1532]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816A36352940FA75408F0C8C" box="[269,396,1506,1532]" italics="true" pageId="31" pageNumber="190">C.blantoni</emphasis>
</taxonomicName>
, as long as broad in 
<taxonomicName id="4C714D2B816A36352A2AFA75420A0C8C" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[615,777,1506,1532]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA816A36352A2AFA75420A0C8C" box="[615,777,1506,1532]" italics="true" pageId="31" pageNumber="190">C.virginianus</emphasis>
</taxonomicName>
). My examination of additional material of both species found that these characters are not diagnostic. Leg color appears to be a spectrum; while some specimens have completely yellow and others completely brown legs, there are intermediates with solid brown femora and yellow tibiae or femora proximally brown and yellow distally. The legs of the 
<typeStatus id="54CA880A816A36352C26F9DB45CC0F16" box="[1131,1231,1612,1638]" pageId="31" pageNumber="190" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C714D2B816A36352CBBF9DA44740F16" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[1270,1399,1612,1639]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816A36352CBBF9DA44740F16" box="[1270,1399,1612,1639]" italics="true" pageId="31" pageNumber="190">C.blantoni</emphasis>
</taxonomicName>
are now yellow with brown bands on the femora, probably a result of the slide mounting process. The male genitalia characters used by Wirth &amp; Ratanaworabhan appear to be artifacts of slide mounting. Specifically, the direction of the apex of the parameres is directly correlated to the compression of the genitalia by the coverslip during mounting: when there is little pressure the parameres point ventroposteriorly, but when compressed the apices are displaced laterally. The proportions of the aedeagus are similarly affected by the mounting of the specimen, the degree of tilt of the aedeagus in the dorsal plane changes its apparent proportions on the slide mount due to foreshortening. As no characters consistently separate 
<taxonomicName id="4C714D2B816A36352CBBF8D144740E2F" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[1270,1399,1861,1888]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816A36352CBBF8D144740E2F" box="[1270,1399,1861,1888]" italics="true" pageId="31" pageNumber="190">C.blantoni</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C714D2B816A363528B6F8FE409F0EF3" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[251,412,1897,1923]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA816A363528B6F8FE409F0EF3" box="[251,412,1897,1923]" italics="true" pageId="31" pageNumber="190">C.virginianus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C714D2B816A363529E7F8FE43290EF3" authorityName="Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[426,554,1897,1923]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="190" phylum="Arthropoda" rank="species" species="blantoni">
<emphasis id="B905EABA816A363529E7F8FE43290EF3" box="[426,554,1897,1923]" italics="true" pageId="31" pageNumber="190">C.blantoni</emphasis>
</taxonomicName>
is here considered a junior synonym of the latter species.
</paragraph>
<paragraph id="8BCE36A8816A360A28F0F83C44600BDC" blockId="31.[189,1399,1963,2025]" lastBlockId="32.[187,1400,265,684]" lastPageId="32" lastPageNumber="191" pageId="31" pageNumber="190">
Currently, females of this species can be recognized by being the only known species in eastern North America with spermathecae &lt;70 μm long. There are several morphospecies from the western Nearctic and Neotropics with similarly sized spermathecae. 
<taxonomicName id="4C714D2B8155360A2B7AFE9E45920853" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[823,1169,265,291]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="191" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA8155360A2B7AFE9E45920853" box="[823,1169,265,291]" italics="true" pageId="32" pageNumber="191">Ceratoculicoides virginianus</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A2D8155360A2CEAFE9E440D0853" box="[1191,1294,265,291]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="32" pageNumber="191">Fig. 12c</figureCitation>
) can be separated from 
<taxonomicName id="4C714D2B8155360A2939FEBA40F10836" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[372,498,301,327]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="191" phylum="Arthropoda" rank="species" species="grogani" status="sp. nov.">
<emphasis id="B905EABA8155360A2939FEBA40F10836" box="[372,498,301,327]" italics="true" pageId="32" pageNumber="191">C.grogani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C18155360A29B4FEBA43570837" box="[505,596,301,327]" pageId="32" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
by the deeply concave and rounded medial margin of the 9 
<superScript id="7C049BE08155360A2D46FEBB441B084A" attach="left" box="[1291,1304,300,314]" fontSize="6" pageId="32" pageNumber="191">th</superScript>
sternite (vs weakly concave and sinuous, 
<figureCitation id="134A2A2D8155360A2A07FEC743AE081A" box="[586,685,336,362]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta &amp; Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="32" pageNumber="191">Fig. 11e</figureCitation>
), the base of the posterior branch being posteriorly from the base, curving medially in its apical half (vs directed posteromedially in an even curve in 
<taxonomicName id="4C714D2B8155360A2CA4FEE3446C08FE" authorityName="Fasbender" authorityYear="2023" box="[1257,1391,372,398]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="191" phylum="Arthropoda" rank="species" species="borkenti">
<emphasis id="B905EABA8155360A2CA4FEE3446C08FE" box="[1257,1391,372,398]" italics="true" pageId="32" pageNumber="191">C.borkenti</emphasis>
</taxonomicName>
, 
<figureCitation id="134A2A2D8155360A28F0FE0F402008C2" box="[189,291,408,434]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta &amp; Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="32" pageNumber="191">Fig. 11b</figureCitation>
), the posterior branch tapering only in its apical half (vs 
<taxonomicName id="4C714D2B8155360A2B96FE0F456108C2" authority="Fasbender, 2023" authorityName="Fasbender" authorityYear="2023" box="[987,1122,408,434]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="191" phylum="Arthropoda" rank="species" species="borkenti" status="sp. nov.">
<emphasis id="B905EABA8155360A2B96FE0F456108C2" box="[987,1122,408,434]" italics="true" pageId="32" pageNumber="191">C.borkenti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23657C18155360A2C21FE0F45CA08C2" box="[1132,1225,408,434]" pageId="32" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
and 
<emphasis id="B905EABA8155360A2D4AFE0F442108C2" box="[1287,1314,408,434]" italics="true" pageId="32" pageNumber="191">C.</emphasis>
sp. F3 where it tapers to an acute spine in the basal half, 
<figureCitation id="134A2A2D8155360A2B4BFE2C426E08A5" box="[774,877,443,469]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[238,1398,265,1781]" captionTargetId="figure-88@24.[238,1398,265,1781]" captionTargetPageId="24" captionText="Fig. 11. Female sternite 9 I. a. Ceratoculicoides aliciae Huerta &amp; Borkent, 2005, paratype, Chamela Biological Station, Jalisco, Mexico. b. C. borkenti sp. nov., paratype, Nogales, Arizona, USA. c. C. confusus Kouchibouguac National Park, New Brunswick, Canada. d. C.confusus, paratype, Great Smoky Mountains National Park, Tennessee, USA. e. C.grogani sp. nov., paratype, Corvallis, Oregon, USA. f. C.moravicus Bornova, Izmir, Turkey. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083808" httpUri="https://zenodo.org/record/8083808/files/figure.png" pageId="32" pageNumber="191">Figs 11b</figureCitation>
, 
<figureCitation id="134A2A2D8155360A2B37FE2C42A608A5" box="[890,933,443,469]" captionStart="Fig" captionStartId="30.[189,232,1822,1848]" captionTargetBox="[225,1362,265,1749]" captionTargetId="figure-86@30.[225,1362,265,1749]" captionTargetPageId="30" captionText="Fig. 12. Female sternite 9 II. a. Ceratoculicoides pacificus sp. nov., paratype (CNCI), Carmanah Valley, British Columbia, Canada.b. C. pacificus paratype (CNCI), East Sooke Peak, British Columbia, Canada. c. C.virginianus (Wirth, 1951), Springwater Conservation Area, Ontario, Canada. d. Ceratoculicoides sp. F1, Tuparro National Park, Colombia. e. Ceratoculicoides sp. F2, Tuparro National Park, Colombia. f. Ceratoculicoides sp. F3, Rincon, Costa Rica. Linework has been added to make sternite 9 easier to discern. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083812" httpUri="https://zenodo.org/record/8083812/files/figure.png" pageId="32" pageNumber="191">12f</figureCitation>
). Accordingly, 
<taxonomicName id="4C714D2B8155360A2C1AFE2B45FD08A5" baseAuthorityName="Wirth" baseAuthorityYear="1951" box="[1111,1278,444,470]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="191" phylum="Arthropoda" rank="species" species="virginianus">
<emphasis id="B905EABA8155360A2C1AFE2B45FD08A5" box="[1111,1278,444,470]" italics="true" pageId="32" pageNumber="191">C.virginianus</emphasis>
</taxonomicName>
cannot be distinguished from 
<emphasis id="B905EABA8155360A29D6FE4840B50889" box="[411,438,479,505]" italics="true" pageId="32" pageNumber="191">C.</emphasis>
sp. F1 based on morphology from our current knowledge. There is some variability in the shape of the apices of the anterior and posterior branches of the 9 
<superScript id="7C049BE08155360A2C5FFD95451C0B60" attach="left" box="[1042,1055,514,528]" fontSize="6" pageId="32" pageNumber="191">th</superScript>
sternite. The anterior branch can be obtusely rounded or somewhat flattened and truncate, while the tip of the posterior branch may be either pointed or acutely rounded. Males can be separated from all other 
<taxonomicName id="4C714D2B8155360A2C4DFDDD45CB0B14" authorityName=", Wirth &amp; Ratanaworabhan" authorityYear="1971" box="[1024,1224,586,612]" class="Insecta" family="Ceratopogonidae" genus="Ceratoculicoides" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B905EABA8155360A2C4DFDDD45CB0B14" box="[1024,1224,586,612]" italics="true" pageId="32" pageNumber="191">Ceratoculicoides</emphasis>
</taxonomicName>
by the broadly rounded apex of the paramere with subacute triangular point (
<figureCitation id="134A2A2D8155360A2BC1FDF942E00BF8" box="[908,995,622,648]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="32" pageNumber="191">Fig. 9b</figureCitation>
), the aedeagus with convex lateral margins, no hyaline medial incision, and the dorsolateral and apical points directed dorsally (
<figureCitation id="134A2A2D8155360A2D4DFD0644550BDC" box="[1280,1366,657,684]" captionStart="Fig" captionStartId="16.[189,232,1787,1813]" captionTargetBox="[224,1364,267,1733]" captionTargetId="figure-118@16.[224,1364,267,1733]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia II. Ceratoculicoides virginianus (Wirth, 1951): a. Genitalia, aedeagus and parameres removed.b. Parameres.c. Aedeagus.C.borkenti sp. nov:d. Genitalia, aedeagus and parameres removed. e. Parameres.f. Aedeagus.C.grogani sp. nov.: g. Genitalia, aedeagus and parameres removed. h. Parameres. i. Aedeagus. Abbreviations: ALP = apicolateral point of epandrium; AP = apical point of aedeagus; BA = basal arch of aedeagus; CRC = cercus; GC = gonocoxite; GCA = gonocoxal apodeme; GNS = gonostylus; HI = hyaline incision of aedeagus; HYP = hypandrium; PA = paramere apex; PB = paramere base; PLP = posterolateral point of aedeagus. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8083800" httpUri="https://zenodo.org/record/8083800/files/figure.png" pageId="32" pageNumber="191">Fig. 9c</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>