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<document ID-DOI="10.1371/journal.pone.0149556" ID-GBIF-Dataset="f02345d5-4425-434a-8574-883833ae0b6f" ID-PMC="PMC4784953" ID-PubMed="26958842" ID-Zenodo-Dep="1161582" checkinTime="1517115034139" checkinUser="plazi" docAuthor="Benny K. K. Chan, H. - N. Chen, P. R. Dando, A. J. Southward &amp; E. C. Southward" docDate="2016" docId="03D887BE630C8402C7794EF60BB8FA77" docLanguage="en" docName="journal.pone.0149556.PDF" docOrigin="PLoS ONE 11 (3)" docStyle="DocumentStyle{}" docTitle="Chthamalus alani Chan, Chen, Dando, Southward &amp; Southward, 2016, nom. nov." docType="treatment" docVersion="11" lastPageNumber="13" masterDocId="FFE1FFC6630B840EC56C4C1A0E10FFDD" masterDocTitle="Biodiversity and Biogeography of Chthamalid Barnacles from the North-Eastern Pacific (Crustacea Cirripedia)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="8" updateTime="1668134660285" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Biodiversity and Biogeography of Chthamalid Barnacles from the North-Eastern Pacific (Crustacea Cirripedia)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Benny K. K. Chan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>H. - N. Chen</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>P. R. Dando</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>A. J. Southward</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>E. C. Southward</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PLoS ONE</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-03-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>11</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>51</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1371/journal.pone.0149556</mods:identifier>
<mods:identifier type="GBIF-Dataset">f02345d5-4425-434a-8574-883833ae0b6f</mods:identifier>
<mods:identifier type="PMC">PMC4784953</mods:identifier>
<mods:identifier type="PubMed">26958842</mods:identifier>
<mods:identifier type="Zenodo-Dep">1161582</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4426650" ID-GBIF-Taxon="140512817" ID-Zenodo-Dep="4426650" LSID="urn:lsid:plazi:treatment:03D887BE630C8402C7794EF60BB8FA77" httpUri="http://treatment.plazi.org/id/03D887BE630C8402C7794EF60BB8FA77" lastPageId="12" lastPageNumber="13" pageId="7" pageNumber="8">
<subSubSection pageId="7" pageNumber="8" type="nomenclature">
<paragraph blockId="7.[533,917,748,778]" box="[533,917,748,778]" pageId="7" pageNumber="8">
<heading box="[533,917,748,778]" fontSize="12" level="2" pageId="7" pageNumber="8" reason="1">
<taxonomicName authorityName="Chan" authorityYear="2016" box="[533,772,748,778]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alani" status="nom. nov.">
<emphasis box="[533,772,748,778]" italics="true" pageId="7" pageNumber="8">Chthamalus alani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[779,917,748,778]" pageId="7" pageNumber="8">nom. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph blockId="7.[533,1536,798,1897]" box="[533,1280,798,822]" pageId="7" pageNumber="8">urn:lsid:zoobank.org:act:4C460612-D362-49BA-9A20-F2895C4CE1F5</paragraph>
</subSubSection>
<subSubSection box="[565,1238,833,857]" pageId="7" pageNumber="8" type="etymology">
<paragraph blockId="7.[533,1536,798,1897]" box="[565,1238,833,857]" pageId="7" pageNumber="8">the authority for this species name change rests with BKK Chan</paragraph>
</subSubSection>
<subSubSection box="[565,817,867,892]" pageId="7" pageNumber="8" type="description">
<paragraph blockId="7.[533,1536,798,1897]" box="[565,817,867,892]" pageId="7" pageNumber="8">
<figureCitation box="[565,725,867,892]" captionStart="Fig2" captionStartId="8.[96,128,1511,1531]" captionTargetBox="[99,1277,211,1487]" captionTargetId="figure@8.[96,1280,208,1490]" captionTargetPageId="8" captionText="Fig2. LiveChthamalusspeciesphotographedunderwater.A,mixedpopulationsofCanisopoma(CAN),withwhite-edgedtergoscutalflaps,Balanus inexpectatatus(BI)withwhitetergoscutalflapsandChthamalusalaninom.nov.(CA)withorangetergoscutalflaps(fromLaPaz,BajaCalifornia).B,C.alani showingorangetergoscutalflaps(fromLaPaz).C,C.hedgcockishowingorangetergoscutalflaps(fromPointLobos,TodosSantos,BajaCalifornia).D, Chthamalusnewmanisp.nov.showingbrowntergoscutalflaps(fromFarfanPoint,Panama).E,F,mixedpopulationofC.panamensis(CP)showingorange tergoscutalflapsandC.newmani(CN)showingbrowntergoscutalflaps(E,BalboaStatue.PanamaCityandF,NaosIsland). doi:10.1371/journal.pone.0149556.g002" httpUri="https://zenodo.org/record/1161639/files/figure.png" pageId="7" pageNumber="8">Figs 2A and 2B</figureCitation>
and
<figureCitation box="[777,817,867,892]" captionStart-0="Fig3" captionStart-1="Fig4" captionStart-2="Fig5" captionStartId-0="9.[96,128,1270,1290]" captionStartId-1="10.[96,128,1733,1753]" captionStartId-2="11.[96,128,1733,1753]" captionTargetBox-0="[98,1502,210,1247]" captionTargetBox-1="[100,1122,212,1709]" captionTargetBox-2="[100,1138,211,1710]" captionTargetId-0="figure@9.[96,1504,208,1249]" captionTargetId-1="figure@10.[96,1126,208,1712]" captionTargetId-2="figure@11.[96,1144,208,1712]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Fig3. Chthamalusalaninom.nov.A,B,lightstereomicroscopyshowinginnersideofpairedscutumandtergum,LaPazspecimen.C,scanningElectron Micrograph(SEM)showingtheinnersideofscutumandtergum,Mazatlanspecimen.D,pairedarticulatedscutumandtergum,aLaPazspecimen,showing theobliquearticulationoftheplates(a). doi:10.1371/journal.pone.0149556.g003" captionText-1="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" captionText-2="Fig5. Chthamalusalaninom.nov.Matzalan,Mexico.A,maxilla.B,maxillule.C,magnifiedviewshowingthenotchregionofmaxillule.D,mandible.E, fourthtooth,pectenandinferiorangleofmandible.F,mandibularpalp.G,labrum.H,cuttingmarginoflabrumshowingthefineteeth.Scalebarsin μm. doi:10.1371/journal.pone.0149556.g005" httpUri-0="https://zenodo.org/record/1161641/files/figure.png" httpUri-1="https://zenodo.org/record/1161645/files/figure.png" httpUri-2="https://zenodo.org/record/1161606/files/figure.png" pageId="7" pageNumber="8">35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="reference_group">
<paragraph blockId="7.[533,1536,798,1897]" box="[565,1315,902,926]" pageId="7" pageNumber="8">
<taxonomicName authority="Pitombo &amp; Burton, 2007: 9" authorityName="Pitombo &amp; Burton" authorityPageNumber="9" authorityYear="2007" box="[565,1160,902,926]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[565,868,902,926]" italics="true" pageId="7" pageNumber="8">Chthamalus southwardorum</emphasis>
Pitombo &amp; Burton, 2007: 9
</taxonomicName>
, figs 914 [
<bibRefCitation author="Pitombo FB &amp; Burton R." box="[1280,1306,902,926]" journalOrPublisher="Zootaxa" pageId="7" pageNumber="8" pagination="1 - 30" part="1547" refString="12. Pitombo FB, Burton R. Systematics and biogeography of Tropical Eastern Pacific Chthamalus with descriptions of two new species (Cirripedia, Thoracica). Zootaxa. 2007; 1547: 1 - 30." title="Systematics and biogeography of Tropical Eastern Pacific Chthamalus with descriptions of two new species (Cirripedia, Thoracica)" type="journal article" year="2007">12</bibRefCitation>
]
</paragraph>
<paragraph blockId="7.[533,1536,798,1897]" box="[565,1254,936,961]" pageId="7" pageNumber="8">
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[565,868,937,961]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[565,868,937,961]" italics="true" pageId="7" pageNumber="8">Chthamalus southwardorum</emphasis>
</taxonomicName>
A.—Wares
<emphasis box="[997,1040,937,961]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
., 2009:
<tableCitation box="[1121,1199,937,961]" captionStart="Table1" captionStartId="4.[96,151,207,227]" captionTargetBox="[96,1536,250,1198]" captionTargetId="graphics@4.[96,1536,716,775]" captionText="Table1. Stationswherespecimenswerecollectedforcomparativestudiesofthespecies." httpUri="http://table.plazi.org/id/DF0E6620630F840AC50C4CD50DC6FF3E" pageId="7" pageNumber="8" tableUuid="DF0E6620630F840AC50C4CD50DC6FF3E">Table 1</tableCitation>
. [
<bibRefCitation author="Wares JP &amp; Pankey MS &amp; Pitombo F &amp; Daglio LG &amp; Achituv Y. A" box="[1220,1246,937,961]" journalOrPublisher="PLoS One" pageId="7" pageNumber="8" pagination="5567" part="4" publicationUrl="http://dx.doi.org/ 10.1371/journal.pone.0005567" refString="14. Wares JP, Pankey MS, Pitombo F, Daglio LG, Achituv Y. A &quot; Shallow Phylogeny &quot; of Shallow Barnacles (Chthamalus). PLoS One. 2009; 4: e 5567. doi: 10.1371 / journal. pone. 0005567 PMID: 19440543" title="Shallow Phylogeny &quot; of Shallow Barnacles (Chthamalus)" type="journal article" year="2009">14</bibRefCitation>
]
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="etymology">
<paragraph blockId="7.[533,1536,798,1897]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[565,691,971,995]" pageId="7" pageNumber="8">Etymology.</emphasis>
Pitombo &amp; Burton [
<bibRefCitation author="Pitombo FB &amp; Burton R." box="[925,951,972,996]" journalOrPublisher="Zootaxa" pageId="7" pageNumber="8" pagination="1 - 30" part="1547" refString="12. Pitombo FB, Burton R. Systematics and biogeography of Tropical Eastern Pacific Chthamalus with descriptions of two new species (Cirripedia, Thoracica). Zootaxa. 2007; 1547: 1 - 30." title="Systematics and biogeography of Tropical Eastern Pacific Chthamalus with descriptions of two new species (Cirripedia, Thoracica)" type="journal article" year="2007">12</bibRefCitation>
] stated explicitly that
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[1184,1486,971,995]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[1184,1486,971,995]" italics="true" pageId="7" pageNumber="8">Chthamalus southwardorum</emphasis>
</taxonomicName>
is named “in honor of Dr A.J. Southward, for his extensive contribution to the knowledge of chthamalids and his careful collecting during the Tropical eastern Pacific Expedition”. Unfortunately, the suffix
<emphasis box="[735,801,1076,1099]" italics="true" pageId="7" pageNumber="8">-orum</emphasis>
indicates “for men or for man (men) and woman (women) together” (ICZN, 4th edition, 1999 Article 31.1.2). Since there is clear evidence of a
<emphasis box="[1300,1441,1110,1134]" italics="true" pageId="7" pageNumber="8">lapsus calami</emphasis>
in the original publication (ICZN ART 32.5.1) the name
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[1063,1260,1145,1169]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[1063,1080,1145,1168]" italics="true" pageId="7" pageNumber="8">C</emphasis>
.
<emphasis box="[1092,1260,1145,1169]" italics="true" pageId="7" pageNumber="8">southwardorum</emphasis>
</taxonomicName>
is not admissible under ICZN Art. 21.13 and has to be treated as an incorrect original spelling that should be corrected to
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[561,679,1214,1238]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardi">
<emphasis box="[561,679,1214,1238]" italics="true" pageId="7" pageNumber="8">southwardi</emphasis>
</taxonomicName>
, taking the authorship and date of Pitombo &amp; Burton [
<bibRefCitation author="Pitombo FB &amp; Burton R." box="[1260,1286,1214,1238]" journalOrPublisher="Zootaxa" pageId="7" pageNumber="8" pagination="1 - 30" part="1547" refString="12. Pitombo FB, Burton R. Systematics and biogeography of Tropical Eastern Pacific Chthamalus with descriptions of two new species (Cirripedia, Thoracica). Zootaxa. 2007; 1547: 1 - 30." title="Systematics and biogeography of Tropical Eastern Pacific Chthamalus with descriptions of two new species (Cirripedia, Thoracica)" type="journal article" year="2007">12</bibRefCitation>
]. However,
<taxonomicName authority="Pitombo &amp; Burton, 2007" authorityName="Pitombo &amp; Burton" authorityYear="2007" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardi">
<emphasis italics="true" pageId="7" pageNumber="8">Chthamalus southwardi</emphasis>
Pitombo &amp; Burton, 2007
</taxonomicName>
is a junior primary homonym of
<taxonomicName authority="Poltaruka, 2000" authorityName="Poltaruka" authorityYear="2000" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardi">
<emphasis italics="true" pageId="7" pageNumber="8">Chthamalus southwardi</emphasis>
Poltaruka, 2000
</taxonomicName>
and therefore an invalid name (ICZN Art 57.2) and so a new replacement name is required for
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[890,1086,1318,1342]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[890,907,1319,1342]" italics="true" pageId="7" pageNumber="8">C</emphasis>
.
<emphasis box="[918,1086,1318,1342]" italics="true" pageId="7" pageNumber="8">southwardorum</emphasis>
</taxonomicName>
. In the present study, I (BKKC) rename
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[533,730,1353,1377]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[533,550,1353,1376]" italics="true" pageId="7" pageNumber="8">C</emphasis>
.
<emphasis box="[562,730,1353,1377]" italics="true" pageId="7" pageNumber="8">southwardorum</emphasis>
</taxonomicName>
as
<emphasis box="[763,952,1353,1377]" italics="true" pageId="7" pageNumber="8">
Chthamlaus
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[898,952,1353,1377]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alani">alani</taxonomicName>
</emphasis>
, again in honour of Prof. Alan Southward, but using his first name, for his contributions to barnacle distribution, ecology and taxonomy.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="materials_examined">
<paragraph blockId="7.[533,1536,798,1897]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[565,802,1422,1446]" pageId="7" pageNumber="8">Specimens examined.</emphasis>
AJS-2, 10 specimens, high intertidal shores, La Paz Bay, 24° 13' N, 110° 18.6' W, 3 Nov 1978; AJS-3, 7 specimens, intertidal shores, Point Lobos, Todos Santos, Baja California Sur, Mexico, 23° 24.8' N, 110° 57' W, 4 Nov 1978; TEPE78-36, 10 specimens, Ocean Front, Mazatlan, Sinaloa, Mexico, exposed rocky shores with large boulders, 23° 11.7' N, 106° 25.5' W, 19 April 1978.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="13" pageId="7" pageNumber="8" type="diagnosis">
<paragraph blockId="7.[533,1536,798,1897]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[565,682,1595,1619]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
<taxonomicName authorityName="Ranzani" authorityYear="1817" box="[703,831,1595,1619]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis box="[703,831,1595,1619]" italics="true" pageId="7" pageNumber="8">Chthamalus</emphasis>
</taxonomicName>
with cirrus I having bidenticulate setae with or without basal guards on the distalmost segment of the posterior ramus and cirrus II having bidenticulate setae with basal guards on the two distalmost segments of the posterior ramus and the distalmost segment of the anterior ramus. Tergoscutal flaps orange-brown in colour when alive. COI sequence (Gene Bank Number KU356723).
</paragraph>
<paragraph blockId="7.[533,1536,798,1897]" lastBlockId="9.[533,1536,1416,1891]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<emphasis bold="true" box="[565,704,1768,1792]" pageId="7" pageNumber="8">Description.</emphasis>
Shell depressed, sutures between plates visible (
<figureCitation box="[1222,1372,1769,1793]" captionStart="Fig2" captionStartId="8.[96,128,1511,1531]" captionTargetBox="[99,1277,211,1487]" captionTargetId="figure@8.[96,1280,208,1490]" captionTargetPageId="8" captionText="Fig2. LiveChthamalusspeciesphotographedunderwater.A,mixedpopulationsofCanisopoma(CAN),withwhite-edgedtergoscutalflaps,Balanus inexpectatatus(BI)withwhitetergoscutalflapsandChthamalusalaninom.nov.(CA)withorangetergoscutalflaps(fromLaPaz,BajaCalifornia).B,C.alani showingorangetergoscutalflaps(fromLaPaz).C,C.hedgcockishowingorangetergoscutalflaps(fromPointLobos,TodosSantos,BajaCalifornia).D, Chthamalusnewmanisp.nov.showingbrowntergoscutalflaps(fromFarfanPoint,Panama).E,F,mixedpopulationofC.panamensis(CP)showingorange tergoscutalflapsandC.newmani(CN)showingbrowntergoscutalflaps(E,BalboaStatue.PanamaCityandF,NaosIsland). doi:10.1371/journal.pone.0149556.g002" httpUri="https://zenodo.org/record/1161639/files/figure.png" pageId="7" pageNumber="8">Fig 2A and 2B</figureCitation>
). External surface white and smooth, sheath white, with ribs at the basal region, not extending to the apex (
<figureCitation box="[542,693,1838,1862]" captionStart="Fig2" captionStartId="8.[96,128,1511,1531]" captionTargetBox="[99,1277,211,1487]" captionTargetId="figure@8.[96,1280,208,1490]" captionTargetPageId="8" captionText="Fig2. LiveChthamalusspeciesphotographedunderwater.A,mixedpopulationsofCanisopoma(CAN),withwhite-edgedtergoscutalflaps,Balanus inexpectatatus(BI)withwhitetergoscutalflapsandChthamalusalaninom.nov.(CA)withorangetergoscutalflaps(fromLaPaz,BajaCalifornia).B,C.alani showingorangetergoscutalflaps(fromLaPaz).C,C.hedgcockishowingorangetergoscutalflaps(fromPointLobos,TodosSantos,BajaCalifornia).D, Chthamalusnewmanisp.nov.showingbrowntergoscutalflaps(fromFarfanPoint,Panama).E,F,mixedpopulationofC.panamensis(CP)showingorange tergoscutalflapsandC.newmani(CN)showingbrowntergoscutalflaps(E,BalboaStatue.PanamaCityandF,NaosIsland). doi:10.1371/journal.pone.0149556.g002" httpUri="https://zenodo.org/record/1161639/files/figure.png" pageId="7" pageNumber="8">Fig 2A and 2B</figureCitation>
). Internal colour of shell white. Orifice oval, scutum and tergum articulated with a marked angle (
<figureCitation box="[760,922,1873,1897]" captionStart="Fig2" captionStartId="8.[96,128,1511,1531]" captionTargetBox="[99,1277,211,1487]" captionTargetId="figure@8.[96,1280,208,1490]" captionTargetPageId="8" captionText="Fig2. LiveChthamalusspeciesphotographedunderwater.A,mixedpopulationsofCanisopoma(CAN),withwhite-edgedtergoscutalflaps,Balanus inexpectatatus(BI)withwhitetergoscutalflapsandChthamalusalaninom.nov.(CA)withorangetergoscutalflaps(fromLaPaz,BajaCalifornia).B,C.alani showingorangetergoscutalflaps(fromLaPaz).C,C.hedgcockishowingorangetergoscutalflaps(fromPointLobos,TodosSantos,BajaCalifornia).D, Chthamalusnewmanisp.nov.showingbrowntergoscutalflaps(fromFarfanPoint,Panama).E,F,mixedpopulationofC.panamensis(CP)showingorange tergoscutalflapsandC.newmani(CN)showingbrowntergoscutalflaps(E,BalboaStatue.PanamaCityandF,NaosIsland). doi:10.1371/journal.pone.0149556.g002" httpUri="https://zenodo.org/record/1161639/files/figure.png" pageId="7" pageNumber="8">Figs 2A and 2B</figureCitation>
and
<figureCitation box="[974,1006,1873,1897]" captionStart="Fig3" captionStartId="9.[96,128,1270,1290]" captionTargetBox="[98,1502,210,1247]" captionTargetId="figure@9.[96,1504,208,1249]" captionTargetPageId="9" captionText="Fig3. Chthamalusalaninom.nov.A,B,lightstereomicroscopyshowinginnersideofpairedscutumandtergum,LaPazspecimen.C,scanningElectron Micrograph(SEM)showingtheinnersideofscutumandtergum,Mazatlanspecimen.D,pairedarticulatedscutumandtergum,aLaPazspecimen,showing theobliquearticulationoftheplates(a). doi:10.1371/journal.pone.0149556.g003" httpUri="https://zenodo.org/record/1161641/files/figure.png" pageId="7" pageNumber="8">3D</figureCitation>
). Tergoscutal flaps orange-brown when alive (
<figureCitation captionStart="Fig2" captionStartId="8.[96,128,1511,1531]" captionTargetBox="[99,1277,211,1487]" captionTargetId="figure@8.[96,1280,208,1490]" captionTargetPageId="8" captionText="Fig2. LiveChthamalusspeciesphotographedunderwater.A,mixedpopulationsofCanisopoma(CAN),withwhite-edgedtergoscutalflaps,Balanus inexpectatatus(BI)withwhitetergoscutalflapsandChthamalusalaninom.nov.(CA)withorangetergoscutalflaps(fromLaPaz,BajaCalifornia).B,C.alani showingorangetergoscutalflaps(fromLaPaz).C,C.hedgcockishowingorangetergoscutalflaps(fromPointLobos,TodosSantos,BajaCalifornia).D, Chthamalusnewmanisp.nov.showingbrowntergoscutalflaps(fromFarfanPoint,Panama).E,F,mixedpopulationofC.panamensis(CP)showingorange tergoscutalflapsandC.newmani(CN)showingbrowntergoscutalflaps(E,BalboaStatue.PanamaCityandF,NaosIsland). doi:10.1371/journal.pone.0149556.g002" httpUri="https://zenodo.org/record/1161639/files/figure.png" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">Fig 2A and 2B</figureCitation>
), colour fades out when preserved in ethanol or formalin. Scutum triangular, basal margin about twice as long as tergal margin (
<figureCitation box="[1008,1060,1760,1784]" captionStart="Fig3" captionStartId="9.[96,128,1270,1290]" captionTargetBox="[98,1502,210,1247]" captionTargetId="figure@9.[96,1504,208,1249]" captionTargetPageId="9" captionText="Fig3. Chthamalusalaninom.nov.A,B,lightstereomicroscopyshowinginnersideofpairedscutumandtergum,LaPazspecimen.C,scanningElectron Micrograph(SEM)showingtheinnersideofscutumandtergum,Mazatlanspecimen.D,pairedarticulatedscutumandtergum,aLaPazspecimen,showing theobliquearticulationoftheplates(a). doi:10.1371/journal.pone.0149556.g003" httpUri="https://zenodo.org/record/1161641/files/figure.png" pageId="8" pageNumber="9">Fig 3</figureCitation>
). Tergal margin with a wide and elevated articular ridge, ridge extending beyond the tergal margin, articular furrow wide. Occludent margin straight, without teeth. Adductor muscle pit oval and deep, not extending to the basal margin. Lateral depressor muscle scar deep and smooth (
<figureCitation box="[1134,1186,1864,1888]" captionStart="Fig3" captionStartId="9.[96,128,1270,1290]" captionTargetBox="[98,1502,210,1247]" captionTargetId="figure@9.[96,1504,208,1249]" captionTargetPageId="9" captionText="Fig3. Chthamalusalaninom.nov.A,B,lightstereomicroscopyshowinginnersideofpairedscutumandtergum,LaPazspecimen.C,scanningElectron Micrograph(SEM)showingtheinnersideofscutumandtergum,Mazatlanspecimen.D,pairedarticulatedscutumandtergum,aLaPazspecimen,showing theobliquearticulationoftheplates(a). doi:10.1371/journal.pone.0149556.g003" httpUri="https://zenodo.org/record/1161641/files/figure.png" pageId="8" pageNumber="9">Fig 3</figureCitation>
). Tergum trapezoid, external surface smooth near the apex, basal region with growth lines or striations, spur wide and not obvious, basal margin with 4 rostral depressor muscle crests (
<figureCitation box="[1179,1231,1451,1475]" captionStart="Fig3" captionStartId="9.[96,128,1270,1290]" captionTargetBox="[98,1502,210,1247]" captionTargetId="figure@9.[96,1504,208,1249]" captionTargetPageId="9" captionText="Fig3. Chthamalusalaninom.nov.A,B,lightstereomicroscopyshowinginnersideofpairedscutumandtergum,LaPazspecimen.C,scanningElectron Micrograph(SEM)showingtheinnersideofscutumandtergum,Mazatlanspecimen.D,pairedarticulatedscutumandtergum,aLaPazspecimen,showing theobliquearticulationoftheplates(a). doi:10.1371/journal.pone.0149556.g003" httpUri="https://zenodo.org/record/1161641/files/figure.png" pageId="9" pageNumber="10">Fig 3</figureCitation>
).
</paragraph>
<caption httpUri="https://zenodo.org/record/1161639/files/figure.png" pageId="8" pageNumber="9" targetBox="[99,1277,211,1487]" targetPageId="8">
<paragraph blockId="8.[96,1524,1511,1673]" pageId="8" pageNumber="9">
<emphasis bold="true" box="[96,678,1511,1531]" pageId="8" pageNumber="9">
Fig 2. Live
<taxonomicName authorityName="Ranzani" authorityYear="1817" box="[203,325,1511,1531]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[203,325,1511,1531]" italics="true" pageId="8" pageNumber="9">Chthamalus</emphasis>
</taxonomicName>
species photographed underwater.
</emphasis>
A, mixed populations of
<emphasis box="[902,1025,1511,1531]" italics="true" pageId="8" pageNumber="9">C anisopoma</emphasis>
(CAN), with white-edged tergoscutal flaps,
<emphasis italics="true" pageId="8" pageNumber="9">
<taxonomicName box="[1423,1498,1511,1531]" class="Maxillopoda" family="Balanidae" genus="Balanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Balanus</taxonomicName>
inexpectatatus
</emphasis>
(BI) with white tergoscutal flaps and
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[568,727,1536,1556]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alani" status="nom. nov.">
<emphasis box="[568,727,1536,1556]" italics="true" pageId="8" pageNumber="9">Chthamalus alani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[732,825,1536,1556]" pageId="8" pageNumber="9">nom. nov.</taxonomicNameLabel>
(CA) with orange tergoscutal flaps (from La Paz, Baja California). B,
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[1451,1520,1536,1556]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="alani">
<emphasis box="[1451,1466,1536,1556]" italics="true" pageId="8" pageNumber="9">C</emphasis>
.
<emphasis box="[1477,1520,1536,1556]" italics="true" pageId="8" pageNumber="9">alani</emphasis>
</taxonomicName>
showing orange tergoscutal flaps (from La Paz). C,
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[567,689,1561,1581]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hedgcocki">
<emphasis box="[567,582,1561,1581]" italics="true" pageId="8" pageNumber="9">C</emphasis>
.
<emphasis box="[592,689,1561,1581]" italics="true" pageId="8" pageNumber="9">hedgcocki</emphasis>
</taxonomicName>
showing orange tergoscutal flaps (from Point Lobos, Todos Santos, Baja California). D,
<taxonomicName authority="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward, 2016" authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[96,296,1587,1607]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="newmani" status="sp. nov.">
<emphasis box="[96,296,1587,1607]" italics="true" pageId="8" pageNumber="9">Chthamalus newmani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[301,375,1587,1607]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
showing brown tergoscutal flaps (from Farfan Point, Panama). E, F, mixed population of
<emphasis box="[1186,1201,1587,1607]" italics="true" pageId="8" pageNumber="9">C</emphasis>
.
<emphasis box="[1212,1324,1587,1607]" italics="true" pageId="8" pageNumber="9">panamensis</emphasis>
(CP) showing orange tergoscutal flaps and
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[293,403,1612,1632]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="newmani">
<emphasis box="[293,308,1612,1632]" italics="true" pageId="8" pageNumber="9">C</emphasis>
.
<emphasis box="[319,403,1612,1632]" italics="true" pageId="8" pageNumber="9">newmani</emphasis>
</taxonomicName>
(CN) showing brown tergoscutal flaps (E,
<taxonomicName authority="Statue." authorityName="Statue." box="[789,928,1612,1632]" class="Insecta" family="Tettigoniidae" genus="Balboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Balboa Statue.</taxonomicName>
Panama City and F,
<taxonomicName box="[1118,1168,1612,1632]" class="Anthozoa" family="Arachnophyllidae" genus="Naos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rugosa" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">Naos</taxonomicName>
Island). doi:10.1371/journal.pone.0149556.g002
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/1161641/files/figure.png" pageId="9" pageNumber="10" targetBox="[98,1502,210,1247]" targetPageId="9">
<paragraph blockId="9.[96,1527,1270,1382]" pageId="9" pageNumber="10">
<emphasis bold="true" box="[96,435,1270,1290]" pageId="9" pageNumber="10">
Fig 3.
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[157,331,1270,1290]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="alani" status="nom. nov.">
<emphasis bold="true" box="[157,331,1270,1290]" italics="true" pageId="9" pageNumber="10">Chthamalus alani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[336,435,1270,1290]" pageId="9" pageNumber="10">nom. nov.</taxonomicNameLabel>
</emphasis>
A, B, light stereomicroscopy showing inner side of paired scutum and tergum, La Paz specimen. C, scanning Electron Micrograph (SEM) showing the inner side of scutum and tergum, Mazatlan specimen. D, paired articulated scutum and tergum, a La Paz specimen, showing the oblique articulation of the plates (a). doi:10.1371/journal.pone.0149556.g003
</paragraph>
</caption>
<paragraph blockId="9.[533,1536,1416,1891]" lastBlockId="12.[533,1535,872,1450]" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
Segment counts on cirri I-III were based on five specimens collected from Mazatlan, Mexico (TEPE-78-36). Segment counts on cirri IV-VI were based on a single specimen from Mazatlan, Mexico (
<tableCitation box="[627,705,1555,1579]" captionStart="Table4" captionStartId="12.[96,151,207,227]" captionTargetBox="[96,1536,250,661]" captionTargetId="graphics@12.[96,1536,380,505]" captionText="Table4. Segmentcounts(n=5individuals)ofChthamalusintheTropicalEasternPacificwaters." httpUri="http://table.plazi.org/id/DF0E662063078402C50C4CD50A0EFF3E" pageId="9" pageNumber="10" tableUuid="DF0E662063078402C50C4CD50A0EFF3E">Table 4</tableCitation>
). Cirrus I: posterior ramus 5 to 6-segmented (
<figureCitation box="[1191,1261,1555,1579]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="9" pageNumber="10">Fig 4A</figureCitation>
), the distalmost segment bears bidenticulate setae with or without basal guards (
<figureCitation box="[1111,1179,1590,1614]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="9" pageNumber="10">Fig 4B</figureCitation>
), serrulate type setae common on all segments of the rami, anterior ramus 6 to 8, serrulate type, setae common on all segments (
<figureCitation box="[542,613,1659,1683]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="9" pageNumber="10">Fig 4A</figureCitation>
,
<tableCitation box="[624,702,1659,1683]" captionStart="Table4" captionStartId="12.[96,151,207,227]" captionTargetBox="[96,1536,250,661]" captionTargetId="graphics@12.[96,1536,380,505]" captionText="Table4. Segmentcounts(n=5individuals)ofChthamalusintheTropicalEasternPacificwaters." httpUri="http://table.plazi.org/id/DF0E662063078402C50C4CD50A0EFF3E" pageId="9" pageNumber="10" tableUuid="DF0E662063078402C50C4CD50A0EFF3E">Table 4</tableCitation>
). Cirrus II: posterior ramus 4 to 7-segmented, the two distalmost segments bear bidenticulate setae with basal guards, anterior ramus 5- or 6-segmented, one or two of the distalmost segments have bidenticulate setae with basal guards, serrulate type setae common on both rami (
<figureCitation box="[687,840,1763,1787]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="9" pageNumber="10">Fig 4C and 4D</figureCitation>
,
<tableCitation box="[852,930,1763,1787]" captionStart="Table4" captionStartId="12.[96,151,207,227]" captionTargetBox="[96,1536,250,661]" captionTargetId="graphics@12.[96,1536,380,505]" captionText="Table4. Segmentcounts(n=5individuals)ofChthamalusintheTropicalEasternPacificwaters." httpUri="http://table.plazi.org/id/DF0E662063078402C50C4CD50A0EFF3E" pageId="9" pageNumber="10" tableUuid="DF0E662063078402C50C4CD50A0EFF3E">Table 4</tableCitation>
). Cirrus III: anterior and posterior rami similar in length (anterior and posterior rami length ratio = 0.9 ± 0.06), posterior ramus 11 to 16-segmented, anterior ramus 11 to 16-segmented (
<figureCitation box="[919,986,1832,1856]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="9" pageNumber="10">Fig 4E</figureCitation>
,
<tableCitation box="[998,1076,1832,1856]" captionStart="Table4" captionStartId="12.[96,151,207,227]" captionTargetBox="[96,1536,250,661]" captionTargetId="graphics@12.[96,1536,380,505]" captionText="Table4. Segmentcounts(n=5individuals)ofChthamalusintheTropicalEasternPacificwaters." httpUri="http://table.plazi.org/id/DF0E662063078402C50C4CD50A0EFF3E" pageId="9" pageNumber="10" tableUuid="DF0E662063078402C50C4CD50A0EFF3E">Table 4</tableCitation>
). Cirrus IV: both anterior and posterior rami 16-segmented (
<figureCitation box="[753,818,1867,1891]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="9" pageNumber="10">Fig 4F</figureCitation>
). Cirrus V: posterior ramus 18-segmented, anterior ramus 19-segmented (
<figureCitation box="[697,768,872,896]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="12" pageNumber="13">Fig 4G</figureCitation>
). Cirrus VI: posterior ramus 18-segmented, anterior ramus 19-segmented (
<figureCitation box="[629,701,906,930]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="12" pageNumber="13">Fig 4H</figureCitation>
). Intermediate segments of both rami on cirri III-VI bear two pairs of long serrulate setae and three pairs of shorter simple setae (
<figureCitation box="[1073,1187,941,965]" captionStart="Fig4" captionStartId="10.[96,128,1733,1753]" captionTargetBox="[100,1122,212,1709]" captionTargetId="figure@10.[96,1126,208,1712]" captionTargetPageId="10" captionText="Fig4. Chthamalusalaninom.nov.Matzalan,Mexico.A,B,cirrusI,showingthepresenceofbidenticulatesetae(bi)withorwithoutbasalguards(bg), enlargedinB.C,D,cirrusII,showingthepresenceofbidenticulatesetae(bi)withbasalguards(bg)onthefirstfewsegments(indicatedbydoubleend arrows)inD.E,cirrusIII.F,cirrusIV.G,cirrusV.H,cirrusVI.IntermediatesegmentsofanteriorramiofcirriIII-VIareshownasinsertsinE-H.Scalebars in μm. doi:10.1371/journal.pone.0149556.g004" httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="12" pageNumber="13">Fig 4E4H</figureCitation>
).
</paragraph>
<caption httpUri="https://zenodo.org/record/1161645/files/figure.png" pageId="10" pageNumber="11" targetBox="[100,1122,212,1709]" targetPageId="10">
<paragraph blockId="10.[96,1491,1733,1870]" pageId="10" pageNumber="11">
<emphasis bold="true" box="[96,616,1733,1753]" pageId="10" pageNumber="11">
Fig 4.
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[157,331,1733,1753]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alani" status="nom. nov.">
<emphasis bold="true" box="[157,331,1733,1753]" italics="true" pageId="10" pageNumber="11">Chthamalus alani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[336,435,1733,1753]" pageId="10" pageNumber="11">nom. nov.</taxonomicNameLabel>
Matzalan, Mexico.
</emphasis>
A, B, cirrus I, showing the presence of bidenticulate setae (bi) with or without basal guards (bg), enlarged in B. C, D, cirrus II, showing the presence of bidenticulate setae (bi) with basal guards (bg) on the first few segments (indicated by double end arrows) in D. E, cirrus III. F, cirrus IV. G, cirrus V. H, cirrus VI. Intermediate segments of anterior rami of cirri III-VI are shown as inserts in E-H. Scale bars in μm. doi:10.1371/journal.pone.0149556.g004
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/1161606/files/figure.png" pageId="11" pageNumber="12" targetBox="[100,1138,211,1710]" targetPageId="11">
<paragraph blockId="11.[96,1491,1733,1819]" pageId="11" pageNumber="12">
<emphasis bold="true" box="[96,435,1733,1753]" pageId="11" pageNumber="12">
Fig 5.
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[157,331,1733,1753]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="alani" status="nom. nov.">
<emphasis bold="true" box="[157,331,1733,1753]" italics="true" pageId="11" pageNumber="12">Chthamalus alani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[336,435,1733,1753]" pageId="11" pageNumber="12">nom. nov.</taxonomicNameLabel>
</emphasis>
Matzalan, Mexico. A, maxilla. B, maxillule. C, magnified view showing the notch region of maxillule. D, mandible. E, fourth tooth, pecten and inferior angle of mandible. F, mandibular palp. G, labrum. H, cutting margin of labrum showing the fine teeth. Scale bars in μm. doi:10.1371/journal.pone.0149556.g005
</paragraph>
</caption>
<caption ID-Table-UUID="DF0E662063078402C50C4CD50A0EFF3E" box="[96,1054,207,227]" httpUri="http://table.plazi.org/id/DF0E662063078402C50C4CD50A0EFF3E" pageId="12" pageNumber="13" targetBox="[96,1536,250,661]" targetIsTable="true" targetPageId="12">
<paragraph blockId="12.[96,1054,207,227]" box="[96,1054,207,227]" pageId="12" pageNumber="13">
Table 4. Segment counts (n = 5 individuals) of
<taxonomicName authorityName="Ranzani" authorityYear="1817" box="[553,675,207,227]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[553,675,207,227]" italics="true" pageId="12" pageNumber="13">Chthamalus</emphasis>
</taxonomicName>
in the Tropical Eastern Pacific waters.
</paragraph>
</caption>
<paragraph pageId="12" pageNumber="13">
<table box="[96,1536,250,661]" gridcols="6" gridrows="9" pageId="12" pageNumber="13">
<tr box="[96,1536,250,270]" gridrow="0" pageId="12" pageNumber="13" rowspan-0="1">
<th box="[858,1001,250,270]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">
<taxonomicName box="[858,1001,250,270]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="hedgecocki">
<emphasis bold="true" box="[858,1001,250,270]" italics="true" pageId="12" pageNumber="13">
C.
<emphasis bold="true" box="[885,1001,250,270]" italics="true" pageId="12" pageNumber="13">hedgecocki</emphasis>
</emphasis>
</taxonomicName>
</th>
<th box="[1015,1152,250,270]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">
<emphasis bold="true" box="[1015,1152,250,270]" pageId="12" pageNumber="13">
<emphasis bold="true" box="[1015,1031,250,270]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis bold="true" box="[1042,1152,250,270]" italics="true" pageId="12" pageNumber="13">anisopoma</emphasis>
</emphasis>
</th>
<th box="[1166,1241,250,270]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[1166,1241,250,270]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alani">
<emphasis bold="true" box="[1166,1241,250,270]" pageId="12" pageNumber="13">
<emphasis bold="true" box="[1166,1182,250,270]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis bold="true" box="[1193,1241,250,270]" italics="true" pageId="12" pageNumber="13">alani</emphasis>
</emphasis>
</taxonomicName>
</th>
<th box="[1254,1403,250,270]" gridcol="4" gridrow="0" pageId="12" pageNumber="13">
<emphasis bold="true" box="[1254,1403,250,270]" pageId="12" pageNumber="13">
<emphasis bold="true" box="[1254,1270,250,270]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis bold="true" box="[1282,1403,250,270]" italics="true" pageId="12" pageNumber="13">panamensis</emphasis>
</emphasis>
</th>
<th box="[1417,1536,250,270]" gridcol="5" gridrow="0" pageId="12" pageNumber="13">
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[1417,1536,250,270]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="newmani">
<emphasis bold="true" box="[1417,1536,250,270]" pageId="12" pageNumber="13">
<emphasis bold="true" box="[1417,1433,250,270]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis bold="true" box="[1444,1536,250,270]" italics="true" pageId="12" pageNumber="13">newmani</emphasis>
</emphasis>
</taxonomicName>
</th>
</tr>
<tr box="[96,1536,290,310]" gridrow="1" pageId="12" pageNumber="13">
<th box="[96,835,290,310]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">Cirrus I: No. of segments in posterior ramus</th>
<td box="[858,1001,290,310]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">57</td>
<td box="[1015,1152,290,310]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">67</td>
<td box="[1166,1241,290,310]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">56</td>
<td box="[1254,1403,290,310]" gridcol="4" gridrow="1" pageId="12" pageNumber="13">57</td>
<td box="[1417,1536,290,310]" gridcol="5" gridrow="1" pageId="12" pageNumber="13">67</td>
</tr>
<tr box="[96,1536,324,344]" gridrow="2" pageId="12" pageNumber="13">
<th box="[96,835,324,344]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">Cirrus I: No. of segmentsin anterior ramus</th>
<td box="[858,1001,324,344]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">79</td>
<td box="[1015,1152,324,344]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">78</td>
<td box="[1166,1241,324,344]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">68</td>
<td box="[1254,1403,324,344]" gridcol="4" gridrow="2" pageId="12" pageNumber="13">68</td>
<td box="[1417,1536,324,344]" gridcol="5" gridrow="2" pageId="12" pageNumber="13">89</td>
</tr>
<tr box="[96,1536,357,377]" gridrow="3" pageId="12" pageNumber="13">
<th box="[96,835,357,377]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">Cirrus I: No. of segments in posterior ramus with bidenticulate setae</th>
<td box="[858,1001,357,377]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">0 ± 0</td>
<td box="[1015,1152,357,377]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">1</td>
<td box="[1166,1241,357,377]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">1</td>
<td box="[1254,1403,357,377]" gridcol="4" gridrow="3" pageId="12" pageNumber="13">0</td>
<td box="[1417,1536,357,377]" gridcol="5" gridrow="3" pageId="12" pageNumber="13">1</td>
</tr>
<tr box="[96,1536,390,502]" gridrow="4" pageId="12" pageNumber="13">
<th box="[96,835,390,502]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">Cirrus II: No. of segments in posterior ramus Cirrus II: No. of segments in anterior ramus Cirrus II: No. of segments in posterior ramus with bidenticulate setae with basal guards</th>
<td box="[858,1001,390,502]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">56 56 12</td>
<td box="[1015,1152,390,502]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">5 67 4</td>
<td box="[1166,1241,390,502]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">47 56 2</td>
<td box="[1254,1403,390,502]" gridcol="4" gridrow="4" pageId="12" pageNumber="13">57 68 1</td>
<td box="[1417,1536,390,502]" gridcol="5" gridrow="4" pageId="12" pageNumber="13">57 78 3</td>
</tr>
<tr box="[96,1536,516,561]" gridrow="5" pageId="12" pageNumber="13">
<th box="[96,835,516,561]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">Cirrus II: No. of segments in anterior ramus with bidenticulate setae with basal guards</th>
<td box="[858,1001,516,561]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">1</td>
<td box="[1015,1152,516,561]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">2</td>
<td box="[1166,1241,516,561]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">12</td>
<td box="[1254,1403,516,561]" gridcol="4" gridrow="5" pageId="12" pageNumber="13">1</td>
<td box="[1417,1536,516,561]" gridcol="5" gridrow="5" pageId="12" pageNumber="13">2</td>
</tr>
<tr box="[96,1536,574,594]" gridrow="6" pageId="12" pageNumber="13">
<th box="[96,835,574,594]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">Cirrus III: No. of segments in posterior ramus</th>
<td box="[858,1001,574,594]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">1417</td>
<td box="[1015,1152,574,594]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">1314</td>
<td box="[1166,1241,574,594]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">1116</td>
<td box="[1254,1403,574,594]" gridcol="4" gridrow="6" pageId="12" pageNumber="13">1216</td>
<td box="[1417,1536,574,594]" gridcol="5" gridrow="6" pageId="12" pageNumber="13">1719</td>
</tr>
<tr box="[96,1536,608,628]" gridrow="7" pageId="12" pageNumber="13">
<th box="[96,835,608,628]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">Cirrus III: No. of segments in anterior ramus</th>
<td box="[858,1001,608,628]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">1319</td>
<td box="[1015,1152,608,628]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">1319</td>
<td box="[1166,1241,608,628]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">1116</td>
<td box="[1254,1403,608,628]" gridcol="4" gridrow="7" pageId="12" pageNumber="13">1622</td>
<td box="[1417,1536,608,628]" gridcol="5" gridrow="7" pageId="12" pageNumber="13">1823</td>
</tr>
<tr box="[96,1536,641,661]" gridrow="8" pageId="12" pageNumber="13">
<th box="[96,835,641,661]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">Cirris III, anterior/posterior length ratio</th>
<td box="[858,1001,641,661]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">0.91.0</td>
<td box="[1015,1152,641,661]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">0.81.2</td>
<td box="[1166,1241,641,661]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">0.80.9</td>
<td box="[1254,1403,641,661]" gridcol="4" gridrow="8" pageId="12" pageNumber="13">1.41.5</td>
<td box="[1417,1536,641,661]" gridcol="5" gridrow="8" pageId="12" pageNumber="13">0.91.2</td>
</tr>
</table>
</paragraph>
<paragraph blockId="12.[96,1530,692,744]" lastBlockId="12.[96,393,772,792]" pageId="12" pageNumber="13">
<tableNote pageId="12" pageNumber="13" targetBox="[96,1536,250,661]" targetPageId="12">
Note: cirrus I of
<emphasis box="[245,260,692,712]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis box="[272,375,692,712]" italics="true" pageId="12" pageNumber="13">anisopoma</emphasis>
has bidenticulate setae without basal guards, whilst cirrus I of
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[959,1029,692,712]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="alani" status="nom. nov.">
<emphasis box="[959,974,692,712]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis box="[986,1029,692,712]" italics="true" pageId="12" pageNumber="13">alani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1035,1129,692,712]" pageId="12" pageNumber="13">nom. nov.</taxonomicNameLabel>
and
<taxonomicName authority="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward, 2016" authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[1175,1286,692,712]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="newmani" status="sp. nov.">
<emphasis box="[1175,1190,692,712]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis box="[1202,1286,692,712]" italics="true" pageId="12" pageNumber="13">newmani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1292,1367,692,712]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
has bidenticulate setae with or without basal guards. doi:10.1371/journal.pone.0149556.t004
</tableNote>
</paragraph>
<paragraph blockId="12.[533,1535,872,1450]" pageId="12" pageNumber="13">
Maxilla bilobed, serrulate setae on apex of both lobes and on superior margin (
<figureCitation box="[1384,1453,975,1000]" captionStart="Fig5" captionStartId="11.[96,128,1733,1753]" captionTargetBox="[100,1138,211,1710]" captionTargetId="figure@11.[96,1144,208,1712]" captionTargetPageId="11" captionText="Fig5. Chthamalusalaninom.nov.Matzalan,Mexico.A,maxilla.B,maxillule.C,magnifiedviewshowingthenotchregionofmaxillule.D,mandible.E, fourthtooth,pectenandinferiorangleofmandible.F,mandibularpalp.G,labrum.H,cuttingmarginoflabrumshowingthefineteeth.Scalebarsin μm. doi:10.1371/journal.pone.0149556.g005" httpUri="https://zenodo.org/record/1161606/files/figure.png" pageId="12" pageNumber="13">Fig 5A</figureCitation>
). Maxillule notched, two large setae above notch, notch small, 8 setae on cutting margin below notch, inferior angle protuberant with bundles of simple setae (
<figureCitation box="[1120,1268,1045,1069]" captionStart="Fig5" captionStartId="11.[96,128,1733,1753]" captionTargetBox="[100,1138,211,1710]" captionTargetId="figure@11.[96,1144,208,1712]" captionTargetPageId="11" captionText="Fig5. Chthamalusalaninom.nov.Matzalan,Mexico.A,maxilla.B,maxillule.C,magnifiedviewshowingthenotchregionofmaxillule.D,mandible.E, fourthtooth,pectenandinferiorangleofmandible.F,mandibularpalp.G,labrum.H,cuttingmarginoflabrumshowingthefineteeth.Scalebarsin μm. doi:10.1371/journal.pone.0149556.g005" httpUri="https://zenodo.org/record/1161606/files/figure.png" pageId="12" pageNumber="13">Fig 5B and 5C</figureCitation>
). Mandible has 4 major teeth, fourth tooth bidenticulate, pecten with 16 small teeth, inferior angle tipped with two pointed teeth (
<figureCitation box="[687,836,1114,1138]" captionStart="Fig5" captionStartId="11.[96,128,1733,1753]" captionTargetBox="[100,1138,211,1710]" captionTargetId="figure@11.[96,1144,208,1712]" captionTargetPageId="11" captionText="Fig5. Chthamalusalaninom.nov.Matzalan,Mexico.A,maxilla.B,maxillule.C,magnifiedviewshowingthenotchregionofmaxillule.D,mandible.E, fourthtooth,pectenandinferiorangleofmandible.F,mandibularpalp.G,labrum.H,cuttingmarginoflabrumshowingthefineteeth.Scalebarsin μm. doi:10.1371/journal.pone.0149556.g005" httpUri="https://zenodo.org/record/1161606/files/figure.png" pageId="12" pageNumber="13">Fig 5D and 5E</figureCitation>
). Mandibular palp rectangular, serrulate setae on all margins (
<figureCitation captionStart="Fig5" captionStartId="11.[96,128,1733,1753]" captionTargetBox="[100,1138,211,1710]" captionTargetId="figure@11.[96,1144,208,1712]" captionTargetPageId="11" captionText="Fig5. Chthamalusalaninom.nov.Matzalan,Mexico.A,maxilla.B,maxillule.C,magnifiedviewshowingthenotchregionofmaxillule.D,mandible.E, fourthtooth,pectenandinferiorangleofmandible.F,mandibularpalp.G,labrum.H,cuttingmarginoflabrumshowingthefineteeth.Scalebarsin μm. doi:10.1371/journal.pone.0149556.g005" httpUri="https://zenodo.org/record/1161606/files/figure.png" pageId="12" pageNumber="13">Fig 5F</figureCitation>
). Labrum concave, with 67 small teeth on both sides of the cutting edge (
<figureCitation box="[1335,1489,1149,1173]" captionStart="Fig5" captionStartId="11.[96,128,1733,1753]" captionTargetBox="[100,1138,211,1710]" captionTargetId="figure@11.[96,1144,208,1712]" captionTargetPageId="11" captionText="Fig5. Chthamalusalaninom.nov.Matzalan,Mexico.A,maxilla.B,maxillule.C,magnifiedviewshowingthenotchregionofmaxillule.D,mandible.E, fourthtooth,pectenandinferiorangleofmandible.F,mandibularpalp.G,labrum.H,cuttingmarginoflabrumshowingthefineteeth.Scalebarsin μm. doi:10.1371/journal.pone.0149556.g005" httpUri="https://zenodo.org/record/1161606/files/figure.png" pageId="12" pageNumber="13">Fig 5G and 5H</figureCitation>
).
</paragraph>
<paragraph blockId="12.[533,1535,872,1450]" box="[565,902,1184,1208]" pageId="12" pageNumber="13">Penis without basi-dorsal point.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="distribution">
<paragraph blockId="12.[533,1535,872,1450]" pageId="12" pageNumber="13">
<emphasis bold="true" box="[565,712,1218,1242]" pageId="12" pageNumber="13">Distribution.</emphasis>
Northern limit probably at about 29° N inside the Gulf of California and 24° 40' N on the Pacific coast; southern limit unknown, but probably north of 15° N. The holotype and paratypes of
<taxonomicName authority="Pitombo &amp; Burton, 2007" authorityName="Pitombo &amp; Burton" authorityYear="2007" box="[714,1177,1288,1312]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[714,731,1288,1311]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis box="[743,911,1288,1312]" italics="true" pageId="12" pageNumber="13">southwardorum</emphasis>
Pitombo &amp; Burton, 2007
</taxonomicName>
were collected in the San Cristovan river estuary, San Blas, Nayarit, Mexico (21° 30' 54'' N, 105° 15' 53'' W), attached to an oyster shell, low midlittoral zone. Pitombo and Burton stated that the distribution of
<taxonomicName authorityName="Pitombo &amp; Burton" authorityYear="2007" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="southwardorum">
<emphasis box="[1392,1409,1357,1380]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis box="[533,701,1392,1416]" italics="true" pageId="12" pageNumber="13">southwardorum</emphasis>
</taxonomicName>
was from Bahía Magdalena, Baja California Sur, Mexico to Puerto Chicama, Peru, but they did not distinguish it from the new species
<taxonomicName authorityName="Benny K. K. Chan &amp; H. -N. Chen &amp; P. R. Dando &amp; A. J. Southward &amp; E. C. Southward" authorityYear="2016" box="[1140,1267,1427,1450]" class="Maxillopoda" family="Chthamalidae" genus="Chthamalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sessilia" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="newmani">
<emphasis box="[1140,1157,1427,1450]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis box="[1169,1267,1427,1450]" italics="true" pageId="12" pageNumber="13">newmani</emphasis>
</taxonomicName>
described below.
</paragraph>
</subSubSection>
</treatment>
</document>