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<document id="62EC301DC434B3DBA0C42BFF5AB9DB0B" ID-DOI="10.5281/zenodo.189873" ID-GBIF-Dataset="c823cfc2-2f08-4ea9-8dbe-5fedb87c8123" ID-ISSN="1175-5326" ID-Zenodo-Dep="189873" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461105455337" checkinUser="plazi" docAuthor="Heinicke, Matthew P., Duellman, William E., Trueb, Linda, Means, Bruce, D, Ross &amp; Hedges, Blair" docDate="2009" docId="03D6631035561754FF3DFAA7FA14F81F" docLanguage="en" docName="zt02211p035.pdf" docOrigin="Zootaxa 2211" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ceuthomantis Heinicke, Duellman, Trueb, Means, D &amp; Hedges, 2009, new genus" docType="treatment" docVersion="8" lastPageNumber="6" masterDocId="FFEF1B6835531751FFAAFF9BFF9CFFA9" masterDocTitle="A new frog family (Anura: Terrarana) from South America and an expanded direct-developing clade revealed by molecular phylogeny" masterLastPageNumber="35" masterPageNumber="1" pageNumber="6" updateTime="1698593055785" updateUser="plazi">
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<mods:title id="2C4754F4BD2360668ACDFBA4FF62E7C6">A new frog family (Anura: Terrarana) from South America and an expanded direct-developing clade revealed by molecular phylogeny</mods:title>
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<mods:namePart id="3E547A1C7EC717DC152B1E4E5393AFF4">Heinicke, Matthew P.</mods:namePart>
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<mods:namePart id="CB63FA5000C050C220DBEFDC89A18D73">Trueb, Linda</mods:namePart>
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<mods:namePart id="B946B0010A5169F87ABA0A7AED997E75">Means, Bruce</mods:namePart>
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<mods:namePart id="D3A4A2C49393BD6F309A096E8638F1AE">D, Ross</mods:namePart>
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<mods:namePart id="F7A89F2271B6B9711C63798411C42791">Hedges, Blair</mods:namePart>
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<mods:date id="35856326253DC366628EFE52C6E69A69">2009</mods:date>
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<treatment id="03D6631035561754FF3DFAA7FA14F81F" ID-DOI="http://doi.org/10.5281/zenodo.5630193" ID-GBIF-Taxon="119622993" ID-Zenodo-Dep="5630193" LSID="urn:lsid:plazi:treatment:03D6631035561754FF3DFAA7FA14F81F" httpUri="http://treatment.plazi.org/id/03D6631035561754FF3DFAA7FA14F81F" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="C365818D35561754FF3DFAA7FE55FAFC" box="[151,457,1339,1366]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BC0D20635561754FF3DFAA7FE55FAFC" blockId="5.[151,457,1339,1366]" box="[151,457,1339,1366]" pageId="5" pageNumber="6">
<heading id="D088656A35561754FF3DFAA7FE55FAFC" bold="true" box="[151,457,1339,1366]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B90B0E1435561754FF3DFAA7FE55FAFC" bold="true" box="[151,457,1339,1366]" pageId="5" pageNumber="6">
<taxonomicName id="4C7FA98535561754FF3DFAA7FEDCFAFF" box="[151,320,1340,1366]" class="Amphibia" family="Ceuthomantidae" genus="Ceuthomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B90B0E1435561754FF3DFAA7FEDCFAFF" bold="true" box="[151,320,1340,1366]" italics="true" pageId="5" pageNumber="6">Ceuthomantis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238B36F35561754FEEDFAA0FE55FAFC" box="[327,457,1339,1365]" pageId="5" pageNumber="6" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BC0D20635561754FF3DFA1FFAA5FA35" blockId="5.[151,1337,1412,1436]" box="[151,1337,1412,1436]" pageId="5" pageNumber="6">
<emphasis id="B90B0E1435561754FF3DFA1FFEB7FA35" bold="true" box="[151,299,1412,1436]" pageId="5" pageNumber="6">
<typeStatus id="54C46CA435561754FF3DFA1FFF53FA35" box="[151,207,1412,1436]" pageId="5" pageNumber="6">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C7FA98535561754FE9BFA1EFBF4FA35" authority="Heinicke, Duellman, Trueb, Means, MacCulloch" authorityName="Heinicke, Duellman, Trueb, Means, MacCulloch" box="[305,1128,1413,1436]" class="Amphibia" family="Ceuthomantidae" genus="Ceuthomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="smaragdinus">
<emphasis id="B90B0E1435561754FE9BFA1EFDCAFA35" box="[305,598,1413,1436]" italics="true" pageId="5" pageNumber="6">Ceuthomantis smaragdinus</emphasis>
Heinicke, Duellman, Trueb, Means, MacCulloch
</taxonomicName>
, and Hedges, 2009.
</paragraph>
</subSubSection>
<subSubSection id="C365818D35561754FF3DFA50FACBF929" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BC0D20635561754FF3DFA50FC0DF99A" blockId="5.[151,1437,1483,1974]" pageId="5" pageNumber="6">
<emphasis id="B90B0E1435561754FF3DFA50FE8BFA4C" bold="true" box="[151,279,1483,1509]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Same as for family. Members of the genus
<taxonomicName id="4C7FA98535561754FCBEFA57FC2BFA4C" box="[788,951,1484,1509]" class="Amphibia" family="Ceuthomantidae" genus="Ceuthomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B90B0E1435561754FCBEFA57FC2BFA4C" box="[788,951,1484,1509]" italics="true" pageId="5" pageNumber="6">Ceuthomantis</emphasis>
</taxonomicName>
are unique compared to the strabomantid genera
<taxonomicName id="4C7FA98535561754FF5AFA68FE5EF9A5" box="[240,450,1523,1548]" class="Amphibia" family="Strabomantidae" genus="Dischidodactylus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B90B0E1435561754FF5AFA68FE5EF9A5" box="[240,450,1523,1548]" italics="true" pageId="5" pageNumber="6">Dischidodactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7FA98535561754FE54FA68FD0EF9A5" box="[510,658,1523,1548]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B90B0E1435561754FE54FA68FD0EF9A5" box="[510,658,1523,1548]" italics="true" pageId="5" pageNumber="6">Pristimantis</emphasis>
</taxonomicName>
in the
<collectingCountry id="F368929635561754FD42FA69FCA3F9A5" box="[744,831,1522,1548]" name="Guyana" pageId="5" pageNumber="6">Guiana</collectingCountry>
Highlands by having notched digital discs on the fingers and toes and by lacking dentigerous processes of vomers.
</paragraph>
<paragraph id="8BC0D20635561754FF6CF9A4FACBF929" blockId="5.[151,1437,1483,1974]" pageId="5" pageNumber="6">
<emphasis id="B90B0E1435561754FF6CF9A4FEAFF9F0" bold="true" box="[198,307,1599,1625]" pageId="5" pageNumber="6">Content.</emphasis>
Tentatively three species,
<taxonomicName id="4C7FA98535561754FDC2F9DBFBD1F9F3" authority="Barrio Amoros &amp; Molina" authorityName="Barrio Amoros &amp; Molina" box="[616,1101,1600,1626]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="aracamuni">
<emphasis id="B90B0E1435561754FDC2F9DBFC95F9F0" box="[616,777,1600,1625]" italics="true" pageId="5" pageNumber="6">C. aracamuni</emphasis>
(Barrio Amorós &amp; Molina)
</taxonomicName>
and
<taxonomicName id="4C7FA98535561754FB2FF9DBFEB2F929" authority="Myers &amp; Donnelly" authorityName="Myers &amp; Donnelly" class="Amphibia" family="Ceuthomantidae" genus="Ceuthomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="caveribardus">
<emphasis id="B90B0E1435561754FB2FF9DBFADCF9F0" box="[1157,1344,1600,1625]" italics="true" pageId="5" pageNumber="6">C. caveribardus</emphasis>
(Myers &amp; Donnelly)
</taxonomicName>
, new combinations, plus
<taxonomicName id="4C7FA98535561754FDFDF9FCFC8CF929" box="[599,784,1639,1664]" class="Amphibia" family="Ceuthomantidae" genus="Ceuthomantis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="smaragdinus" status="sp. nov.">
<emphasis id="B90B0E1435561754FDFDF9FCFC8CF929" box="[599,784,1639,1664]" italics="true" pageId="5" pageNumber="6">C. smaragdinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238B36F35561754FCB2F9FDFCCBF929" box="[792,855,1638,1664]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
described below, are assigned to the genus.
</paragraph>
</subSubSection>
<subSubSection id="C365818D35561754FF6FF916FD87F8EB" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BC0D20635561754FF6FF916FD87F8EB" blockId="5.[151,1437,1483,1974]" pageId="5" pageNumber="6">
<emphasis id="B90B0E1435561754FF6FF916FEFAF90E" bold="true" box="[197,358,1677,1703]" pageId="5" pageNumber="6">Distribution.</emphasis>
The genus is known only from elevations of
<quantity id="4C877FE335561754FC2BF916FB89F90E" box="[897,1045,1677,1703]" metricMagnitude="3" metricUnit="m" metricValue="1.0165" metricValueMax="1.54" metricValueMin="0.493" pageId="5" pageNumber="6" unit="m" value="1016.5" valueMax="1540.0" valueMin="493.0">4931540 m</quantity>
in the southern and eastern parts of the
<collectingCountry id="F368929635561754FF40F92FFED9F967" box="[234,325,1716,1742]" name="Guyana" pageId="5" pageNumber="6">Guiana</collectingCountry>
Highlands. These include Mt. Ayanganna and the Wokomung Massif in
<collectingCountry id="F368929635561754FB49F92FFAD4F967" box="[1251,1352,1716,1742]" name="Guyana" pageId="5" pageNumber="6">Guyana</collectingCountry>
, Cerro
<taxonomicName id="4C7FA98535561754FF3DF941FE87F95D" box="[151,283,1754,1780]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="16" phylum="Chordata" rank="species" species="aracamuni">Aracamuni</taxonomicName>
and Sierra Tapirapecó in the Cerro Neblina Massif on the Venezuela-Brazil border, and possibly Sarisariñama Tepui in southern
<collectingCountry id="F368929635561754FD8BF89AFD03F8B2" box="[545,671,1793,1819]" name="Venezuela" pageId="5" pageNumber="6">Venezuela</collectingCountry>
(see Remarks). The species are known from the slopes of the mountains and the tops of tepuis.
</paragraph>
</subSubSection>
<subSubSection id="C365818D35561754FF6FF8D5FA14F81F" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BC0D20635561754FF6FF8D5FA14F81F" blockId="5.[151,1437,1483,1974]" pageId="5" pageNumber="6">
<emphasis id="B90B0E1435561754FF6FF8D5FEC8F8C1" bold="true" box="[197,340,1870,1896]" pageId="5" pageNumber="6">Etymology.</emphasis>
The generic name is masculine and derived from the Greek noun
<emphasis id="B90B0E1435561754FBDAF8D4FB5BF8C1" box="[1136,1223,1871,1896]" italics="true" pageId="5" pageNumber="6">mantis,</emphasis>
meaning treefrog and the Greek adjective
<emphasis id="B90B0E1435561754FE6FF8EEFDB7F827" box="[453,555,1909,1934]" italics="true" pageId="5" pageNumber="6">keuthos,</emphasis>
meaning hidden and alludes to its hidden existence in the tepuis of the
<collectingCountry id="F368929635561754FF3DF807FF70F81F" box="[151,236,1948,1974]" name="Guyana" pageId="5" pageNumber="6">Guiana</collectingCountry>
Shield, which became known as the
<emphasis id="B90B0E1435561754FD34F807FC83F81C" box="[670,799,1948,1973]" italics="true" pageId="5" pageNumber="6">Lost World</emphasis>
through the writings of Arthur Conan
<bibRefCitation id="EFEEAFF735561754FB49F807FA18F81F" author="Doyle" box="[1251,1412,1948,1974]" pageId="5" pageNumber="27" refString="Doyle, A. C. (1912) The Lost World. Hodder &amp; Stoughten, London, 192 pp." type="book" year="1912">Doyle (1912)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>