332 lines
62 KiB
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332 lines
62 KiB
XML
<document id="CAFF4ACAC2E89CB0256238CC76DB1168" ID-DOI="10.5281/zenodo.5303571" ID-ISSN="0374-1036" ID-Zenodo-Dep="5303571" ID-ZooBank="0360453D-E09E-4CC4-9BB0-981AEC7BED66S" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630104360660" checkinUser="marcus" docAuthor="Tsai, Jing-Fu & Rédei, Dávid" docDate="2015" docId="03B63F659F581C4DFEEBFD19FBC6FC07" docLanguage="en" docName="ActaEntMusNatPra.2015.55.2.625-664.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Acanthosoma Curtis 1824" docType="key" docVersion="4" lastPageNumber="629" masterDocId="FF8F471D9F5A1C49FFADFFB2FFA0FFF4" masterDocTitle="The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)" masterLastPageNumber="664" masterPageNumber="625" pageNumber="627" updateTime="1699153447944" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6883F317495351D8AB55F2D9641637D2">The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)</mods:title>
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<mods:name id="4B3FB99A023C98B0B524F5C0996AD7C8" type="personal">
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<mods:namePart id="28821EDA9CE5960BCB38B080F4C8D209">Tsai, Jing-Fu</mods:namePart>
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<mods:affiliation id="9DB2CDC71BDA4431524890EA03DD6201">Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589 Japan; e-mail: jingfu. tsai @ gmail. com</mods:affiliation>
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<mods:roleTerm id="D487148E4CC43C256A367EF3A3181654">Author</mods:roleTerm>
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<mods:namePart id="07E725F77B38D89916D0AF2FF264A3E4">Rédei, Dávid</mods:namePart>
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<mods:affiliation id="6648EEDB1FEB2D22629E65F54CC38CFE">Institute of Entomology, College of Life Sciences, Nankai University, Weijin Road 94, 300071 Tianjin, China; e-mail: david. redei @ gmail. com & Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross u. 13, Hungary</mods:affiliation>
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<mods:title id="C59C0905740CF5AB73AE9D0B8387CFFC">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="710FFB567B7252A69136FDAFC0F4D63E">
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<mods:date id="0FF86CE67B42A785DD0CD782FF4E0E1C">2015</mods:date>
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<mods:number id="B18FE15B72C3348EF2982A51F52D2924">2015-12-31</mods:number>
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<mods:identifier id="E29717FDA647636F3D7D47B717A5E950" type="DOI">10.5281/zenodo.5303571</mods:identifier>
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<subSubSection id="C305DDF89F581C4BFEEBFD19FC17FD36" box="[326,951,682,706]" pageId="2" pageNumber="627" type="nomenclature">
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<paragraph id="8BA08E739F581C4BFEEBFD19FC17FD36" blockId="2.[326,951,682,706]" box="[326,951,682,706]" pageId="2" pageNumber="627">
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<heading id="D0E8391F9F581C4BFEEBFD19FC17FD36" bold="true" box="[326,951,682,706]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="627" reason="3">
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<emphasis id="B96B52619F581C4BFEEBFD19FC17FD36" bold="true" box="[326,951,682,706]" pageId="2" pageNumber="627">
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Key to the species of
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<taxonomicName id="4C1FF5F09F581C4BFD99FD18FD6BFD36" ID-CoL="7NMN9" authorityName="Curtis" authorityYear="1824" box="[564,715,682,706]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="627" phylum="Arthropoda" rank="genus">
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<emphasis id="B96B52619F581C4BFD99FD18FD6BFD36" bold="true" box="[564,715,682,706]" italics="true" pageId="2" pageNumber="627">Acanthosoma</emphasis>
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</taxonomicName>
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occurring in
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<collectingCountry id="F308CEE39F581C4BFCC9FD19FC17FD36" box="[868,951,683,706]" name="Taiwan" pageId="2" pageNumber="627">Taiwan</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C305DDF89F581C4DFF3AFD68FBC6FC07" lastPageId="4" lastPageNumber="629" pageId="2" pageNumber="627" type="key">
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<keyStep id="30EB96D69F581C4BFF3AFD68FBC6FCE5" pageId="2" pageNumber="627">
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<paragraph id="8BA08E739F581C4BFF3AFD68FBC6FD05" blockId="2.[151,1127,730,1553]" box="[151,1126,730,753]" pageId="2" pageNumber="627">
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<keyLead id="30EE2D469F581C4BFF3AFD68FBC6FD05" box="[151,1126,730,753]" pageId="2" pageNumber="627">1 Humeri rounded, slightly surpassing costal margins of fore wing laterally. ................ 2</keyLead>
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</paragraph>
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<paragraph id="8BA08E739F581C4BFF3AFD48FBC6FCE5" blockId="2.[151,1127,730,1553]" box="[151,1126,762,785]" pageId="2" pageNumber="627">
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<keyLead id="30EE2D469F581C4BFF3AFD48FBC6FCE5" box="[151,1126,762,785]" pageId="2" pageNumber="627">
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– Humeri produced into distinct processes of various length (
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<figureCitation id="132492F69F581C4BFCEEFD48FC61FCE5" box="[835,961,762,785]" captionStart-0="Figs 56–59" captionStart-1="Figs 60–63" captionStart-2="Figs 64–67" captionStart-3="Figs 68–71" captionStart-4="Figs 72–75" captionStart-5="Figs 76–79" captionStart-6="Figs 80–83" captionStart-7="Figs 84–87" captionStart-8="Figs 88–91" captionStartId-0="22.[151,188,1539,1558]" captionStartId-1="23.[113,150,1540,1559]" captionStartId-2="24.[151,188,1541,1560]" captionStartId-3="25.[113,150,1545,1564]" captionStartId-4="26.[151,188,1547,1566]" captionStartId-5="27.[113,150,1553,1572]" captionStartId-6="28.[151,188,1553,1572]" captionStartId-7="29.[113,150,1546,1565]" captionStartId-8="30.[151,188,1546,1565]" captionTargetBox-0="[191,1126,235,1500]" captionTargetBox-1="[144,1044,236,1495]" captionTargetBox-2="[182,1088,227,1493]" captionTargetBox-3="[164,1023,231,1494]" captionTargetBox-4="[194,1071,246,1496]" captionTargetBox-5="[155,1044,235,1489]" captionTargetBox-6="[184,1077,234,1502]" captionTargetBox-7="[141,1052,239,1495]" captionTargetBox-8="[186,1062,229,1509]" captionTargetId-0="figure-10@22.[151,1126,220,1520]" captionTargetId-1="figure-12@23.[113,1088,219,1519]" captionTargetId-2="figure-10@24.[151,1126,212,1512]" captionTargetId-3="figure-12@25.[113,1088,220,1520]" captionTargetId-4="figure-10@26.[151,1126,217,1516]" captionTargetId-5="figure-12@27.[113,1088,220,1520]" captionTargetId-6="figure-10@28.[151,1126,221,1521]" captionTargetId-7="figure-12@29.[113,1088,219,1519]" captionTargetId-8="figure-10@30.[151,1126,219,1519]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="25" captionTargetPageId-4="26" captionTargetPageId-5="27" captionTargetPageId-6="28" captionTargetPageId-7="29" captionTargetPageId-8="30" captionText-0="Figs 56–59. Acanthosoma atayal sp. nov. 56–57 – male; 58–59 – female. Scales in mm." captionText-1="Figs 60–63. Acanthosoma haemorrhoidale formosanum subsp. nov. 60–61 – male (holotype); 62–63 – female (paratype, right antennal segment IV reconstructed digitally). Scales in mm." captionText-2="Figs 64–67. Acanthosoma asahinai Ishihara, 1943. 64–65 – male; 66–67 – female. Scales in mm." captionText-3="Figs 68–71. Acanthosoma fallax sp. nov. 68–69 – male (holotype, right antennal segments III–IV reconstructed digitally); 70–71 – female. Scales in mm." captionText-4="Figs 72–75. Acanthosoma pugnax sp. nov. 72–73 – male (holotype); 74–75 – female (paratype). Scales in mm." captionText-5="Figs 76–79. Acanthosoma axicia sp. nov. 76–77 – male; 78–79 – female. Scales in mm." captionText-6="Figs 80–83. Acanthosoma laevicorne Dallas, 1851. 80–81 – male; 82–83 – female. Scales in mm." captionText-7="Figs 84–87. Acanthosoma spp. 84–85 – A. sichuanense (Liu, 1980); 86 – A. expansum Horváth, 1905, male (Japan: Sapporo); 87 – same, female (Japan: Sapporo). Scales in mm." captionText-8="Figs 88–91. Acanthosoma expansum Horváth, 1905. 88–89 – male (Taiwan: Nantou Co., Sunkang); 90–91 – female (Taiwan: Nantou Co., Tsuifeng–Sungkang). Scales in mm." figureDoi-0="http://doi.org/10.5281/zenodo.5303583" figureDoi-1="http://doi.org/10.5281/zenodo.5303585" figureDoi-2="http://doi.org/10.5281/zenodo.5303587" figureDoi-3="http://doi.org/10.5281/zenodo.5303589" figureDoi-4="http://doi.org/10.5281/zenodo.5303591" figureDoi-5="http://doi.org/10.5281/zenodo.5303593" figureDoi-6="http://doi.org/10.5281/zenodo.5303595" figureDoi-7="http://doi.org/10.5281/zenodo.5303597" figureDoi-8="http://doi.org/10.5281/zenodo.5303599" httpUri-0="https://zenodo.org/record/5303583/files/figure.png" httpUri-1="https://zenodo.org/record/5303585/files/figure.png" httpUri-2="https://zenodo.org/record/5303587/files/figure.png" httpUri-3="https://zenodo.org/record/5303589/files/figure.png" httpUri-4="https://zenodo.org/record/5303591/files/figure.png" httpUri-5="https://zenodo.org/record/5303593/files/figure.png" httpUri-6="https://zenodo.org/record/5303595/files/figure.png" httpUri-7="https://zenodo.org/record/5303597/files/figure.png" httpUri-8="https://zenodo.org/record/5303599/files/figure.png" pageId="2" pageNumber="627">Figs 56–91</figureCitation>
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). ................... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30EB96D69F581C4BFF3AFCA8FBC6FB05" pageId="2" pageNumber="627">
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<paragraph id="8BA08E739F581C4BFF3AFCA8FBC6FBE5" blockId="2.[151,1127,730,1553]" pageId="2" pageNumber="627">
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<keyLead id="30EE2D469F581C4BFF3AFCA8FBC6FBE5" pageId="2" pageNumber="627">
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2 Body large (
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<quantity id="4CE723969F581C4BFEFEFCA8FE55FCC5" box="[339,501,794,817]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.95" metricValueMin="1.75" pageId="2" pageNumber="627" unit="mm" value="18.5" valueMax="19.5" valueMin="17.5">17.5–19.5 mm</quantity>
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). Posterolateral angle of pronotum provided with a small, triangular process directed posteriad and extending over base of corium. Black markings of connexival plates enclosing intersegmental sutures broad (as wide as or wider than tibia). Genital capsule of male large, robust, with short, broad, apically rounded lateral projections not surpassing apex of membrane, ventrally densely pilose, provided with 1+1 pigmented denticles subapically and 1+1 denticles proximally. Posterior margin of laterotergites VIII of female broadly rounded, ventrites VI and VII each with a pair of large Pendergrast’s organs. ............................
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<taxonomicName id="4C1FF5F09F581C4BFCA5FC4BFBC6FBE5" ID-CoL="7Q9JP" authority="Jakovlev, 1880" authorityName="Jakovlev" authorityYear="1880" box="[776,1126,1017,1041]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="627" phylum="Arthropoda" rank="species" species="crassicaudum">
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<emphasis id="B96B52619F581C4BFCA5FC4BFC60FBE5" bold="true" box="[776,960,1017,1041]" italics="true" pageId="2" pageNumber="627">A. crassicaudum</emphasis>
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<bibRefCitation id="EF8EF3829F581C4BFC6BFC48FBC6FBE5" author="JAKOVLEV V. E." box="[966,1126,1018,1041]" pageId="2" pageNumber="627" pagination="385 - 398" refId="ref19702" refString="JAKOVLEV V. E. 1880: Contributions to the fauna of bugs of Russia and the neighbouring countries. IV. Bulletin de la Societe Imperiale des Naturalistes de Moscou 55 (2): 385 - 398 (in Russian and German)." type="journal article" year="1880">Jakovlev, 1880</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BA08E739F581C4BFF3AFBA8FBC6FB05" blockId="2.[151,1127,730,1553]" pageId="2" pageNumber="627">
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<keyLead id="30EE2D469F581C4BFF3AFBA8FBC6FB05" pageId="2" pageNumber="627">
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– Body of medium size (
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<quantity id="4CE723969F581C4BFE12FBA8FDFFFBC5" box="[447,607,1050,1073]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.65" metricValueMin="1.45" pageId="2" pageNumber="627" unit="mm" value="15.5" valueMax="16.5" valueMin="14.5">14.5–16.5 mm</quantity>
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). Posterolateral angle of pronotum without small triangular process. Black markings of connexival plates enclosing intersegmental sutures narrow (narrower than tibia). Genital capsule of male provided with a pair of long, slender, posteriorly strongly diverging posterolateral projections, apical halves of projections surpass apex of membrane, ventral surface lacking pigmented denticles. Posterior margin of laterotergite VIII of female truncate, ventrites VI and VII each with a pair of small Pendergrast’s organs. ..................................
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<taxonomicName id="4C1FF5F09F581C4BFCEFFB6BFBC6FB05" ID-CoL="7Q9K8" authority="Jakovlev, 1880" authorityName="Jakovlev" authorityYear="1880" box="[834,1126,1241,1265]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="627" phylum="Arthropoda" rank="species" species="forficula">
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<emphasis id="B96B52619F581C4BFCEFFB6BFC1FFB05" bold="true" box="[834,959,1241,1265]" italics="true" pageId="2" pageNumber="627">A. forficula</emphasis>
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<bibRefCitation id="EF8EF3829F581C4BFC6BFB68FBC6FB05" author="JAKOVLEV V. E." box="[966,1126,1242,1265]" pageId="2" pageNumber="627" pagination="385 - 398" refId="ref19702" refString="JAKOVLEV V. E. 1880: Contributions to the fauna of bugs of Russia and the neighbouring countries. IV. Bulletin de la Societe Imperiale des Naturalistes de Moscou 55 (2): 385 - 398 (in Russian and German)." type="journal article" year="1880">Jakovlev, 1880</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30EB96D69F581C4AFF3AFB48FBE0FEE7" lastPageId="3" lastPageNumber="628" pageId="2" pageNumber="627">
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<paragraph id="8BA08E739F581C4BFF3AFB48FBC6F9E5" blockId="2.[151,1127,730,1553]" pageId="2" pageNumber="627">
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<keyLead id="30EE2D469F581C4BFF3AFB48FBC6F9E5" pageId="2" pageNumber="627">
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3 Posterolateral angles of pregenital abdominal ventrites not produced (
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<figureCitation id="132492F69F581C4BFC63FB48FB81FAE5" box="[974,1057,1274,1297]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="2" pageNumber="627">Fig. 16</figureCitation>
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). Posterolateral angle of pronotum horizontal, subtriangularly produced and broadly obtuse apically (population in
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<collectingCountry id="F308CEE39F581C4BFE68FA88FDA2FAA5" box="[453,514,1338,1361]" name="Japan" pageId="2" pageNumber="627">Japan</collectingCountry>
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and the Asian mainland) or elevated, finger-shaped, with rounded apex (population in Taiwan), in both cases with black punctures nearly to their tip (
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<figureCitation id="132492F69F581C4BFF57FAC8FEDAFA65" box="[250,378,1402,1425]" captionStart-0="Figs 84–87" captionStart-1="Figs 88–91" captionStartId-0="29.[113,150,1546,1565]" captionStartId-1="30.[151,188,1546,1565]" captionTargetBox-0="[141,1052,239,1495]" captionTargetBox-1="[186,1062,229,1509]" captionTargetId-0="figure-12@29.[113,1088,219,1519]" captionTargetId-1="figure-10@30.[151,1126,219,1519]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="Figs 84–87. Acanthosoma spp. 84–85 – A. sichuanense (Liu, 1980); 86 – A. expansum Horváth, 1905, male (Japan: Sapporo); 87 – same, female (Japan: Sapporo). Scales in mm." captionText-1="Figs 88–91. Acanthosoma expansum Horváth, 1905. 88–89 – male (Taiwan: Nantou Co., Sunkang); 90–91 – female (Taiwan: Nantou Co., Tsuifeng–Sungkang). Scales in mm." figureDoi-0="http://doi.org/10.5281/zenodo.5303597" figureDoi-1="http://doi.org/10.5281/zenodo.5303599" httpUri-0="https://zenodo.org/record/5303597/files/figure.png" httpUri-1="https://zenodo.org/record/5303599/files/figure.png" pageId="2" pageNumber="627">Figs 86–91</figureCitation>
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). Genital capsule of male without lateral projection, subquadrate in posterior view, ventral rim with 1+1 pigmented denticles laterally (
|
||
<figureCitation id="132492F69F581C4BFC0EFA28FB81FA45" box="[931,1057,1434,1457]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="2" pageNumber="627">Figs 16–19</figureCitation>
|
||
). Ventrite VII of female with 1+1 small, black, rounded spots laterad of median incision of posterior margin, Pendergrast’s organs on ventrites VI and VII narrowly separated (
|
||
<figureCitation id="132492F69F581C4BFB96FA68FF4AF9E5" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="2" pageNumber="627">Fig. 55</figureCitation>
|
||
). ...................................................................................
|
||
<taxonomicName id="4C1FF5F09F581C4BFC98FA4BFBC6F9E5" ID-CoL="7Q9K2" authority="Horvath, 1905" authorityName="Horvath" authorityYear="1905" box="[821,1126,1529,1553]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="627" phylum="Arthropoda" rank="species" species="expansum">
|
||
<emphasis id="B96B52619F581C4BFC98FA4BFC66F9E5" bold="true" box="[821,966,1529,1553]" italics="true" pageId="2" pageNumber="627">A. expansum</emphasis>
|
||
<bibRefCitation id="EF8EF3829F581C4BFC61FA48FBC6F9E5" author="HORVATH G." box="[972,1126,1530,1553]" pageId="2" pageNumber="627" pagination="413 - 423" refId="ref19504" refString="HORVATH G. 1905: Hemipteres nouveaux de Japon. Annales Musei Nationalis Hungarici 3: 413 - 423." type="journal article" year="1905">Horváth, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F591C4AFFDCFF6EFBE0FEE7" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFF6EFBE0FEE7" pageId="3" pageNumber="628">
|
||
– Posterolateral angles of abdominal ventrites II–VI produced into small, sharp denticles (as in
|
||
<figureCitation id="132492F69F591C4AFF4AFF4EFEC0FEE7" box="[231,352,252,275]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="3" pageNumber="628">Figs 1, 6, 8</figureCitation>
|
||
,
|
||
<figureCitation id="132492F69F591C4AFEC1FF4EFE29FEE7" box="[364,393,252,275]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="3" pageNumber="628">15</figureCitation>
|
||
,
|
||
<figureCitation id="132492F69F591C4AFE39FF4EFD8CFEE7" box="[404,556,252,275]" captionStart-0="Figs 45–51" captionStart-1="Figs 52–55" captionStartId-0="17.[113,150,1468,1487]" captionStartId-1="18.[151,188,1522,1541]" captionTargetBox-0="[115,1086,506,1453]" captionTargetBox-1="[151,1124,840,1491]" captionTargetId-0="figure-12@17.[113,1088,506,1462]" captionTargetId-1="figure-10@18.[151,1126,839,1496]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." captionText-1="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi-0="http://doi.org/10.5281/zenodo.5303579" figureDoi-1="http://doi.org/10.5281/zenodo.5303581" httpUri-0="https://zenodo.org/record/5303579/files/figure.png" httpUri-1="https://zenodo.org/record/5303581/files/figure.png" pageId="3" pageNumber="628">45–46, 48–55</figureCitation>
|
||
). .......................................................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F591C4AFFDCFEAEFBE0FE87" pageId="3" pageNumber="628">
|
||
<paragraph id="8BA08E739F591C4AFFDCFEAEFBE0FEC7" blockId="3.[113,1089,220,1587]" box="[113,1088,284,307]" pageId="3" pageNumber="628">
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||
<keyLead id="30EE2D469F591C4AFFDCFEAEFBE0FEC7" box="[113,1088,284,307]" pageId="3" pageNumber="628">4 Connexivum uniformly pale. ....................................................................................... 5</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F591C4AFFDCFE8EFBE0FE87" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFE8EFBE0FE87" pageId="3" pageNumber="628">– Connexival plates with conspicuous black markings anteriorly and posteriorly, enclosing intersegmental sutures. ......................................................................................... 10</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F591C4AFFDCFECEFBE0FE07" pageId="3" pageNumber="628">
|
||
<paragraph id="8BA08E739F591C4AFFDCFECEFBE0FE47" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFECEFBE0FE47" pageId="3" pageNumber="628">5 Head unpunctured or at most with very few punctures which are pale, never contrasting black. ............................................................................................................................ 6</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F591C4AFFDCFE0EFBE0FE07" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFE0EFBE0FE07" pageId="3" pageNumber="628">– Head with irregularly distributed black punctures on mandibular plates and between eyes. ............................................................................................................................. 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F591C4AFFDCFE4EFBE1FC67" pageId="3" pageNumber="628">
|
||
<paragraph id="8BA08E739F591C4AFFDCFE4EFBE0FD27" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
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<keyLead id="30EE2D469F591C4AFFDCFE4EFBE0FD27" pageId="3" pageNumber="628">
|
||
6 Body very large (
|
||
<quantity id="4CE723969F591C4AFECAFE4EFDA7FDE7" box="[359,519,508,531]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" metricValueMax="2.07" metricValueMin="1.65" pageId="3" pageNumber="628" unit="mm" value="18.6" valueMax="20.7" valueMin="16.5">16.5–20.7 mm</quantity>
|
||
) (
|
||
<figureCitation id="132492F69F591C4AFDB2FE4EFD3CFDE7" box="[543,668,508,531]" captionStart="Figs 80–83" captionStartId="28.[151,188,1553,1572]" captionTargetBox="[184,1077,234,1502]" captionTargetId="figure-10@28.[151,1126,221,1521]" captionTargetPageId="28" captionText="Figs 80–83. Acanthosoma laevicorne Dallas, 1851. 80–81 – male; 82–83 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303595" httpUri="https://zenodo.org/record/5303595/files/figure.png" pageId="3" pageNumber="628">Figs 80–83</figureCitation>
|
||
,
|
||
<figureCitation id="132492F69F591C4AFD04FE4EFD50FDE7" box="[681,752,508,531]" captionStart="Figs 92–99" captionStartId="31.[113,150,1517,1536]" captionTargetBox="[123,1076,233,1475]" captionTargetId="figure-12@31.[113,1088,216,1489]" captionTargetPageId="31" captionText="Figs 92–99. Type specimens of Acanthosoma spp. and their labels.92–93 – A. laevicorne Dallas, 1851, lectotype; 94 – same, labels; 95–96 – A. expansum Horváth, 1905, lectotype; 97 – same, labels; 98 – A. expansum, paralectotype; 99 – same, labels. Scales in mm. © BMNH (92–94), HNHM (95–99)." figureDoi="http://doi.org/10.5281/zenodo.5303601" httpUri="https://zenodo.org/record/5303601/files/figure.png" pageId="3" pageNumber="628">92–93</figureCitation>
|
||
). Genital capsule of male with a pair of robust, apically rather acute posterolateral projections, with 2+2 denticles on dorsal rim due to extension of dorsal wall over ventral infolding, and 1+1 of protuberances on ventral rim laterally (
|
||
<figureCitation id="132492F69F591C4AFE57FDEEFDEAFD87" box="[506,586,604,627]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="3" pageNumber="628">Fig. 15</figureCitation>
|
||
). Paramere as in
|
||
<figureCitation id="132492F69F591C4AFCAAFDEEFC22FD87" box="[775,898,604,627]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="3" pageNumber="628">Figs 40–41</figureCitation>
|
||
. Posterior margin of laterotergites VIII of female truncate, reaching or slightly surpassing posterolateral angles of ventrite VII; Pendergrast’s organs of ventrite VII larger (diameter more than
|
||
<quantity id="4CE723969F591C4AFF0BFD0EFF59FD27" box="[166,249,700,723]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="628" unit="mm" value="0.5">0.5 mm</quantity>
|
||
) (
|
||
<figureCitation id="132492F69F591C4AFEA2FD0EFEFFFD27" box="[271,351,700,723]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="3" pageNumber="628">Fig. 54</figureCitation>
|
||
). ...............................................................
|
||
<taxonomicName id="4C1FF5F09F591C4AFC8CFD09FBE0FD27" ID-CoL="7Q9KQ" authority="Dallas, 1851" authorityName="Dallas" authorityYear="1851" box="[801,1088,699,723]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="628" phylum="Arthropoda" rank="species" species="laevicorne">
|
||
<emphasis id="B96B52619F591C4AFC8CFD09FC13FD27" bold="true" box="[801,947,699,723]" italics="true" pageId="3" pageNumber="628">A. laevicorne</emphasis>
|
||
<bibRefCitation id="EF8EF3829F591C4AFC14FD0EFBE0FD27" author="DALLAS W. S." box="[953,1088,700,723]" pageId="3" pageNumber="628" refId="ref18627" refString="DALLAS W. S. 1851: List of the specimens of hemipterous insects in the collection of the British Museum. Part 1. Trustees of the British Museum, London, 368 pp, XI plates." type="book" year="1851">Dallas, 1851</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F591C4AFFDCFD6EFBE1FC67" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFD6EFBE1FC67" pageId="3" pageNumber="628">
|
||
– Body relatively small (
|
||
<quantity id="4CE723969F591C4AFE0FFD6EFDE2FD07" box="[418,578,732,755]" metricMagnitude="-2" metricUnit="m" metricValue="1.515" metricValueMax="1.65" metricValueMin="1.38" pageId="3" pageNumber="628" unit="mm" value="15.15" valueMax="16.5" valueMin="13.8">13.8–16.5 mm</quantity>
|
||
) (
|
||
<figureCitation id="132492F69F591C4AFDF7FD6EFD7AFD07" box="[602,730,732,755]" captionStart="Figs 84–87" captionStartId="29.[113,150,1546,1565]" captionTargetBox="[141,1052,239,1495]" captionTargetId="figure-12@29.[113,1088,219,1519]" captionTargetPageId="29" captionText="Figs 84–87. Acanthosoma spp. 84–85 – A. sichuanense (Liu, 1980); 86 – A. expansum Horváth, 1905, male (Japan: Sapporo); 87 – same, female (Japan: Sapporo). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303597" httpUri="https://zenodo.org/record/5303597/files/figure.png" pageId="3" pageNumber="628">Figs 84–85</figureCitation>
|
||
). Genital capsule of male with a pair of short posterolateral projections, dorsal rim broadly U-shaped, apex of paramere bifurcated. Posterior margin of laterotergites VIII of female broadly rounded, far surpassing posterolateral angles of ventrite VII, posterior margin of ventrite VII with a deep, U-shaped incision margined with black; Pendergrast’s organ of ventrite VI small (diameter less than
|
||
<quantity id="4CE723969F591C4AFEDBFCCEFE6CFC67" box="[374,460,892,915]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="628" unit="mm" value="0.5">0.5 mm</quantity>
|
||
). .............................................
|
||
<taxonomicName id="4C1FF5F09F591C4AFCB4FCC9FBE1FC67" ID-CoL="7Q9LB" authority="(Liu, 1980)" baseAuthorityName="Liu" baseAuthorityYear="1980" box="[793,1089,891,915]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="628" phylum="Arthropoda" rank="species" species="sichuanense">
|
||
<emphasis id="B96B52619F591C4AFCB4FCC9FC60FC67" bold="true" box="[793,960,891,915]" italics="true" pageId="3" pageNumber="628">A. sichuanense</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8EF3829F591C4AFC63FCCEFB99FC67" author="LIU S. L." box="[974,1081,892,915]" pageId="3" pageNumber="628" pagination="233 - 242" refId="ref20554" refString="LIU S. L. 1980: Six new species of Acanthosomatidae from China (Hemiptera: Heteroptera). Zoological Research 1: 233 - 242 (in Chinese, English summary)." type="journal article" year="1980">Liu, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F591C4AFFDCFC2EFBE0FBE7" pageId="3" pageNumber="628">
|
||
<paragraph id="8BA08E739F591C4AFFDCFC2EFBE0FC27" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFC2EFBE0FC27" pageId="3" pageNumber="628">
|
||
7 Humeral process long, directed distinctly anterolaterad, apically sharp (
|
||
<figureCitation id="132492F69F591C4AFC13FC2EFB9DFC47" box="[958,1085,924,947]" captionStart="Figs 60–63" captionStartId="23.[113,150,1540,1559]" captionTargetBox="[144,1044,236,1495]" captionTargetId="figure-12@23.[113,1088,219,1519]" captionTargetPageId="23" captionText="Figs 60–63. Acanthosoma haemorrhoidale formosanum subsp. nov. 60–61 – male (holotype); 62–63 – female (paratype, right antennal segment IV reconstructed digitally). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303585" httpUri="https://zenodo.org/record/5303585/files/figure.png" pageId="3" pageNumber="628">Figs 60–63</figureCitation>
|
||
,
|
||
<figureCitation id="132492F69F591C4AFF0BFC0EFF4EFC27" box="[166,238,956,979]" captionStart="Figs 72–75" captionStartId="26.[151,188,1547,1566]" captionTargetBox="[194,1071,246,1496]" captionTargetId="figure-10@26.[151,1126,217,1516]" captionTargetPageId="26" captionText="Figs 72–75. Acanthosoma pugnax sp. nov. 72–73 – male (holotype); 74–75 – female (paratype). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303591" httpUri="https://zenodo.org/record/5303591/files/figure.png" pageId="3" pageNumber="628">72–75</figureCitation>
|
||
). ......................................................................................................................... 8
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F591C4AFFDCFC6EFBE0FBE7" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFC6EFBE0FBE7" pageId="3" pageNumber="628">
|
||
– Humeral process short, directed laterad, apically sharp (
|
||
<figureCitation id="132492F69F591C4AFCA0FC6EFC2AFC07" box="[781,906,988,1011]" captionStart="Figs 68–71" captionStartId="25.[113,150,1545,1564]" captionTargetBox="[164,1023,231,1494]" captionTargetId="figure-12@25.[113,1088,220,1520]" captionTargetPageId="25" captionText="Figs 68–71. Acanthosoma fallax sp. nov. 68–69 – male (holotype, right antennal segments III–IV reconstructed digitally); 70–71 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303589" httpUri="https://zenodo.org/record/5303589/files/figure.png" pageId="3" pageNumber="628">Figs 68–71</figureCitation>
|
||
) or obtuse (
|
||
<figureCitation id="132492F69F591C4AFBBCFC6EFF4DFBE7" captionStart="Figs 56–59" captionStartId="22.[151,188,1539,1558]" captionTargetBox="[191,1126,235,1500]" captionTargetId="figure-10@22.[151,1126,220,1520]" captionTargetPageId="22" captionText="Figs 56–59. Acanthosoma atayal sp. nov. 56–57 – male; 58–59 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303583" httpUri="https://zenodo.org/record/5303583/files/figure.png" pageId="3" pageNumber="628">Figs 56–59</figureCitation>
|
||
). ......................................................................................................................... 9
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F591C4AFFDCFBAEFBE0FB07" pageId="3" pageNumber="628">
|
||
<paragraph id="8BA08E739F591C4AFFDCFBAEFBE0FB87" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFBAEFBE0FB87" pageId="3" pageNumber="628">
|
||
8 Humeral process bright red (
|
||
<figureCitation id="132492F69F591C4AFE74FBAEFDF6FBC7" box="[473,598,1052,1075]" captionStart="Figs 60–63" captionStartId="23.[113,150,1540,1559]" captionTargetBox="[144,1044,236,1495]" captionTargetId="figure-12@23.[113,1088,219,1519]" captionTargetPageId="23" captionText="Figs 60–63. Acanthosoma haemorrhoidale formosanum subsp. nov. 60–61 – male (holotype); 62–63 – female (paratype, right antennal segment IV reconstructed digitally). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303585" httpUri="https://zenodo.org/record/5303585/files/figure.png" pageId="3" pageNumber="628">Figs 60–63</figureCitation>
|
||
). Head punctured, interocular area with a pair of longitudinal rows of black punctures, maxillary plate without tubercle. ................... ..................................................................
|
||
<emphasis id="B96B52619F591C4AFDC5FBE9FBE0FB87" bold="true" box="[616,1088,1115,1139]" pageId="3" pageNumber="628">
|
||
<taxonomicName id="4C1FF5F09F591C4AFDC5FBE9FC1CFB87" authority="Tsai & Rédei, 2015" authorityName="Tsai & Rédei" authorityYear="2015" box="[616,956,1115,1139]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="628" phylum="Arthropoda" rank="subSpecies" species="haemorroidale" status="subsp. nov." subSpecies="formosanum">
|
||
<emphasis id="B96B52619F591C4AFDC5FBE9FC1CFB87" bold="true" box="[616,956,1115,1139]" italics="true" pageId="3" pageNumber="628">A. haemorroidale formosanum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A258EF1A9F591C4AFC6EFBEEFBE0FB87" box="[963,1088,1116,1139]" pageId="3" pageNumber="628" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F591C4AFFDCFBCEFBE0FB07" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFBCEFBE0FB07" pageId="3" pageNumber="628">
|
||
– Humeral process black or brownish (
|
||
<figureCitation id="132492F69F591C4AFD9FFBCEFD10FB67" box="[562,688,1148,1171]" captionStart="Figs 72–75" captionStartId="26.[151,188,1547,1566]" captionTargetBox="[194,1071,246,1496]" captionTargetId="figure-10@26.[151,1126,217,1516]" captionTargetPageId="26" captionText="Figs 72–75. Acanthosoma pugnax sp. nov. 72–73 – male (holotype); 74–75 – female (paratype). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303591" httpUri="https://zenodo.org/record/5303591/files/figure.png" pageId="3" pageNumber="628">Figs 72–75</figureCitation>
|
||
). Head with very few and faint punctures, interocular area without longitudinal rows of black punctures, maxillary plate with a conspicuous, obtuse tubercle anteriad and mesad of antennal insertion. ............. ..........................................................................................................
|
||
<emphasis id="B96B52619F591C4AFCDFFB69FBE0FB07" bold="true" box="[882,1088,1243,1267]" pageId="3" pageNumber="628">
|
||
<taxonomicName id="4C1FF5F09F591C4AFCDFFB69FC44FB07" authority="Tsai & Rédei, 2015" authorityName="Tsai & Rédei" authorityYear="2015" box="[882,996,1243,1267]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="628" phylum="Arthropoda" rank="species" species="pugnax" status="sp. nov.">
|
||
<emphasis id="B96B52619F591C4AFCDFFB69FC44FB07" bold="true" box="[882,996,1243,1267]" italics="true" pageId="3" pageNumber="628">A. pugnax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A258EF1A9F591C4AFC46FB6EFBE0FB07" box="[1003,1088,1244,1267]" pageId="3" pageNumber="628" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F591C4DFFDCFB4EFBC6FEC7" lastPageId="4" lastPageNumber="629" pageId="3" pageNumber="628">
|
||
<paragraph id="8BA08E739F591C4AFFDCFB4EFBE0FA47" blockId="3.[113,1089,220,1587]" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4AFFDCFB4EFBE0FA47" pageId="3" pageNumber="628">
|
||
9 Maxillary plate without tubercle. Humeral process short and broad, subtriangular, apically obtuse, reddish brown to red with more or less distinct dark suffusion (
|
||
<figureCitation id="132492F69F591C4AFC4CFAAEFF64FAA7" captionStart="Figs 56–59" captionStartId="22.[151,188,1539,1558]" captionTargetBox="[191,1126,235,1500]" captionTargetId="figure-10@22.[151,1126,220,1520]" captionTargetPageId="22" captionText="Figs 56–59. Acanthosoma atayal sp. nov. 56–57 – male; 58–59 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303583" httpUri="https://zenodo.org/record/5303583/files/figure.png" pageId="3" pageNumber="628">Figs 56– 59</figureCitation>
|
||
). Genital capsule of male with a pair of short and broad posterolateral projections, dorsal rim broadly U-shaped, greatly covered by membrane (
|
||
<figureCitation id="132492F69F591C4AFC9EFAEEFC33FA87" box="[819,915,1372,1395]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="3" pageNumber="628">Figs 1–3</figureCitation>
|
||
). Posterior margin of laterotergites VIII of female strongly protruding; Pendergrast’s organ on ventrite VII of female small (
|
||
<figureCitation id="132492F69F591C4AFE2BFA2EFE76FA47" box="[390,470,1436,1459]" captionStart="Figs 45–51" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="3" pageNumber="628">Fig. 45</figureCitation>
|
||
). ...........................................................
|
||
<emphasis id="B96B52619F591C4AFC2FFA29FBE0FA47" bold="true" box="[898,1088,1435,1459]" pageId="3" pageNumber="628">
|
||
<taxonomicName id="4C1FF5F09F591C4AFC2FFA29FC45FA47" authority="Tsai & Rédei, 2015" authorityName="Tsai & Rédei" authorityYear="2015" box="[898,997,1435,1459]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="628" phylum="Arthropoda" rank="species" species="atayal" status="sp. nov.">
|
||
<emphasis id="B96B52619F591C4AFC2FFA29FC45FA47" bold="true" box="[898,997,1435,1459]" italics="true" pageId="3" pageNumber="628">A. atayal</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A258EF1A9F591C4AFC46FA2EFBE0FA47" box="[1003,1088,1436,1459]" pageId="3" pageNumber="628" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F591C4DFFDCFA0EFBC6FEC7" blockId="3.[113,1089,220,1587]" lastBlockId="4.[151,1127,220,1011]" lastPageId="4" lastPageNumber="629" pageId="3" pageNumber="628">
|
||
<keyLead id="30EE2D469F591C4DFFDCFA0EFBC6FEC7" lastPageId="4" lastPageNumber="629" pageId="3" pageNumber="628">
|
||
– Maxillary plate with a conspicuous, obtuse tubercle anteriad and mesad of antennal insertion. Humeral process spiniform, directed laterad, yellow to pale orange (
|
||
<figureCitation id="132492F69F591C4AFC4EFA6EFF64F9E7" captionStart="Figs 68–71" captionStartId="25.[113,150,1545,1564]" captionTargetBox="[164,1023,231,1494]" captionTargetId="figure-12@25.[113,1088,220,1520]" captionTargetPageId="25" captionText="Figs 68–71. Acanthosoma fallax sp. nov. 68–69 – male (holotype, right antennal segments III–IV reconstructed digitally); 70–71 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303589" httpUri="https://zenodo.org/record/5303589/files/figure.png" pageId="3" pageNumber="628">Figs 68– 71</figureCitation>
|
||
). Genital capsule with a pair of elongate, gracile, arched posterolateral projections far surpassing apex of membrane, dorsal rim narrowly U-shaped, concave at midline (
|
||
<figureCitation id="132492F69F5E1C4DFF79FF6EFE95FF07" box="[212,309,220,243]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="4" pageNumber="629">Figs 8–9</figureCitation>
|
||
). Posterior margin of tergite VIII of female truncate, not surpassing posterolateral angles of abdominal segment VII; Pendergrast’s organs of ventrites VI and VII of female elliptical, of equal size (
|
||
<figureCitation id="132492F69F5E1C4DFD95FEAEFD28FEC7" box="[568,648,284,307]" captionStart="Figs 45–51" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="4" pageNumber="629">Fig. 51</figureCitation>
|
||
). .......................................
|
||
<emphasis id="B96B52619F5E1C4DFC06FEA9FBC6FEC7" bold="true" box="[939,1126,283,307]" pageId="4" pageNumber="629">
|
||
<taxonomicName id="4C1FF5F09F5E1C4DFC06FEA9FBAAFEC7" authority="Tsai & Rédei, 2015" authorityName="Tsai & Rédei" authorityYear="2015" box="[939,1034,283,307]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="fallax" status="sp. nov.">
|
||
<emphasis id="B96B52619F5E1C4DFC06FEA9FBAAFEC7" bold="true" box="[939,1034,283,307]" italics="true" pageId="4" pageNumber="629">A. fallax</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A258EF1A9F5E1C4DFBBCFEAEFBC6FEC7" box="[1041,1126,284,307]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F5E1C4DFF3AFE8EFBC6FDE7" pageId="4" pageNumber="629">
|
||
<paragraph id="8BA08E739F5E1C4DFF3AFE8EFBC6FE27" blockId="4.[151,1127,220,1011]" pageId="4" pageNumber="629">
|
||
<keyLead id="30EE2D469F5E1C4DFF3AFE8EFBC6FE27" pageId="4" pageNumber="629">
|
||
10 Head with irregularly distributed black punctures on mandibular plates and between eyes. Hind tibia of male dilated subbasally (
|
||
<figureCitation id="132492F69F5E1C4DFD35FEEEFD47FE87" box="[664,743,348,371]" captionStart="Figs 45–51" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="4" pageNumber="629">Fig. 47</figureCitation>
|
||
). Genital capsule of male with a pair of long, subparallel posterolateral projections (
|
||
<figureCitation id="132492F69F5E1C4DFD66FECEFC90FE67" box="[715,816,380,403]" captionStart="Figs 1–10" captionStartId="10.[151,188,1450,1469]" captionTargetBox="[154,1125,221,1407]" captionTargetId="figure-10@10.[151,1126,221,1420]" captionTargetPageId="10" captionText="Figs 1–10.Acanthosoma spp., males. 1 – A.atayal sp.nov., posterior part of abdomen, ventral view; 2 – same, genital capsule and exposed part of segment VIII, ventral view; 3 – same, genital capsule, lateral view; 4 – A. haemorrhoidale formosanum subsp. nov., holotype, genital capsule, posterior view; 5 – A.haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM), genital capsule, posterior view; 6 – A. asahinai Ishihara, 1943, posterior part of abdomen, ventral view; 7 – same, genital capsule, ventral view; 8 – A. fallax sp. nov., posterior part of abdomen, ventral view; 9 – same, genital capsule, ventral view; 10 – A. pugnax sp. nov., genital capsule and exposed part of segment VIII, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303573" httpUri="https://zenodo.org/record/5303573/files/figure.png" pageId="4" pageNumber="629">Figs 6–7</figureCitation>
|
||
). Posterior margin of tergite VIII of female prominent, far surpassing posterolateral angles of segment VII (
|
||
<figureCitation id="132492F69F5E1C4DFB96FE2EFEB3FE27" captionStart="Figs 45–51" captionStartId="17.[113,150,1468,1487]" captionTargetBox="[115,1086,506,1453]" captionTargetId="figure-12@17.[113,1088,506,1462]" captionTargetPageId="17" captionText="Figs 45–51. Acanthosoma spp. 45 – A. atayal sp. nov., female terminalia, posteroventral view; 46 – A. haemorrhoidale formosanum subsp. nov., paratype, female terminalia, posteroventral view; 47 – A. asahinai Ishihara, 1943, hind tibia of male; 48 – same, female terminalia, dorsal view; 49 – same, posteroventral view; 50 – A. fallax sp. nov., female terminalia, dorsal view; 51 – same, posteroventral view. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303579" httpUri="https://zenodo.org/record/5303579/files/figure.png" pageId="4" pageNumber="629">Fig. 48–49</figureCitation>
|
||
). ................................................................................
|
||
<taxonomicName id="4C1FF5F09F5E1C4DFCE4FE09FBC6FE27" ID-CoL="7NYCZ" authority="Ishihara, 1943" authorityName="Ishihara" authorityYear="1943" box="[841,1126,443,467]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="asahinai">
|
||
<emphasis id="B96B52619F5E1C4DFCE4FE09FC67FE27" bold="true" box="[841,967,443,467]" italics="true" pageId="4" pageNumber="629">A. asahinai</emphasis>
|
||
<bibRefCitation id="EF8EF3829F5E1C4DFC60FE0EFBC6FE27" author="ISHIHARA T." box="[973,1126,444,467]" pageId="4" pageNumber="629" pagination="495 - 497" refId="ref19649" refString="ISHIHARA T. 1943: Zwei Acanthosoma-arten der Insel Formosa (Hemiptera: Pentatomidae). Transactions of the Natural History Society of Taiwan 33: 495 - 497." type="journal article" year="1943">Ishihara, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F5E1C4DFF3AFE6EFBC6FDE7" blockId="4.[151,1127,220,1011]" pageId="4" pageNumber="629">
|
||
<keyLead id="30EE2D469F5E1C4DFF3AFE6EFBC6FDE7" pageId="4" pageNumber="629">– Head unpunctured or at most with very few punctures which are pale, never contrasting black. Hind tibia of male not dilated. ......................................................................... 11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30EB96D69F5E1C4DFF3AFDAEFBC6FC07" pageId="4" pageNumber="629">
|
||
<paragraph id="8BA08E739F5E1C4DFF3AFDAEFBC6FCE7" blockId="4.[151,1127,220,1011]" pageId="4" pageNumber="629">
|
||
<keyLead id="30EE2D469F5E1C4DFF3AFDAEFBC6FCE7" pageId="4" pageNumber="629">
|
||
11 Humeral process relatively long, directed laterad (population in Taiwan) or more or less anterolaterad (population in
|
||
<collectingCountry id="F308CEE39F5E1C4DFD82FD8EFDCCFDA7" box="[559,620,572,595]" name="Japan" pageId="4" pageNumber="629">Japan</collectingCountry>
|
||
and the Asian mainland), frequently red. Male genital capsule with a robust, apically obtusely rounded posterolateral projections not surpassing apex of membrane, widely separated by a broad median incision, ventral rim broadly excavate, of rather rectangular outline, densely pilose, with a pair of small, pigmented denticles. Posterior margin of ventrite VII of female with a pentagonal median incision; Pendergrast’s organs of ventrites VI and VII small (diameter less than
|
||
<quantity id="4CE723969F5E1C4DFBE8FD6EFF59FCE7" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="629" unit="mm" value="0.5">0.5 mm</quantity>
|
||
), rather broadly separated. .........................................
|
||
<taxonomicName id="4C1FF5F09F5E1C4DFC97FD49FBC6FCE7" ID-CoL="7Q9K4" authority="(Walker, 1868)" baseAuthorityName="Walker" baseAuthorityYear="1868" box="[826,1126,763,787]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="firmatum">
|
||
<emphasis id="B96B52619F5E1C4DFC97FD49FC1FFCE7" bold="true" box="[826,959,763,787]" italics="true" pageId="4" pageNumber="629">A. firmatum</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8EF3829F5E1C4DFC60FD4EFBFFFCE7" author="WALKER F." box="[973,1119,764,787]" pageId="4" pageNumber="629" pagination="419 - 599" refId="ref21358" refString="WALKER F. 1868: Catalogue of the specimens of Hemiptera Heteroptera in the collection of the British Museum. Part III. Trustees of the British Museum, London, pp. 419 - 599." type="book chapter" year="1868">Walker, 1868</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F5E1C4DFF3AFCAEFBC6FC07" blockId="4.[151,1127,220,1011]" pageId="4" pageNumber="629">
|
||
<keyLead id="30EE2D469F5E1C4DFF3AFCAEFBC6FC07" pageId="4" pageNumber="629">
|
||
– Humeral process short, invariably directed laterad, yellow to pale orange, never red (
|
||
<figureCitation id="132492F69F5E1C4DFF79FC8EFEF3FCA7" box="[212,339,828,851]" captionStart="Figs 76–79" captionStartId="27.[113,150,1553,1572]" captionTargetBox="[155,1044,235,1489]" captionTargetId="figure-12@27.[113,1088,220,1520]" captionTargetPageId="27" captionText="Figs 76–79. Acanthosoma axicia sp. nov. 76–77 – male; 78–79 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303593" httpUri="https://zenodo.org/record/5303593/files/figure.png" pageId="4" pageNumber="629">Figs 76–79</figureCitation>
|
||
). Male genital capsule with a pair of scissors-like posterolateral projections surpassing apex of membrane by their apical halves, dorsal rim nearly V-shaped, ventral surface densely covered by long setae (
|
||
<figureCitation id="132492F69F5E1C4DFD1AFCCEFC91FC67" box="[695,817,892,915]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="4" pageNumber="629">Figs 11–14</figureCitation>
|
||
). Posterior margin of ventrite VII of female with a shield-shaped median incision; Pendergrast’s organs of ventrites VI and VII large (diameter more than
|
||
<quantity id="4CE723969F5E1C4DFDCFFC0EFD17FC27" box="[610,695,956,979]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="629" unit="mm" value="0.5">0.5 mm</quantity>
|
||
), narrowly separated (
|
||
<figureCitation id="132492F69F5E1C4DFC09FC0EFC53FC27" box="[932,1011,956,979]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="4" pageNumber="629">Fig. 52</figureCitation>
|
||
). .............. ............................................................................................................
|
||
<emphasis id="B96B52619F5E1C4DFC05FC69FBC6FC07" bold="true" box="[936,1126,987,1011]" pageId="4" pageNumber="629">
|
||
<taxonomicName id="4C1FF5F09F5E1C4DFC05FC69FBAAFC07" authority="Tsai & Rédei, 2015" authorityName="Tsai & Rédei" authorityYear="2015" box="[936,1034,987,1011]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="629" phylum="Arthropoda" rank="species" species="axicia" status="sp. nov.">
|
||
<emphasis id="B96B52619F5E1C4DFC05FC69FBAAFC07" bold="true" box="[936,1034,987,1011]" italics="true" pageId="4" pageNumber="629">A. axicia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A258EF1A9F5E1C4DFBBCFC6EFBC6FC07" box="[1041,1126,988,1011]" pageId="4" pageNumber="629" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |