752 lines
102 KiB
XML
752 lines
102 KiB
XML
<document id="32DCB53BC22AD1D9CA18BA5578DBCE53" ID-DOI="10.5281/zenodo.179171" ID-GBIF-Dataset="2f58270b-64da-4865-9b8c-e8af94dc3275" ID-ISSN="1175-5326" ID-Zenodo-Dep="179171" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459941252152" checkinUser="plazi" docAuthor="Kłonowska-Olejnik, Małgorzata, Prokopov, Grigorii A. & Godunko, Roman J." docDate="2007" docId="03A3C7011E69FF8A54C2FCB8BB6956EE" docLanguage="en" docName="zt01620p062.pdf" docOrigin="Zootaxa 1620" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ecdyonurus solus Godunko, Klonowska-Olejnik and Prokopov, sp. nov." docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="FF9ABF791E68FF835455FFAFBE385634" masterDocTitle="Description of Ecdyonurus solus sp. nov., a new species of the Ecdyonurus venosus species-group (Ephemeroptera: Heptageniidae) from the Crimean Peninsula, Ukraine" masterLastPageNumber="62" masterPageNumber="53" pageNumber="54" updateTime="1698230474167" updateUser="plazi">
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<mods:title id="01395CEE49C33C407A15509B375EDE9A">Description of Ecdyonurus solus sp. nov., a new species of the Ecdyonurus venosus species-group (Ephemeroptera: Heptageniidae) from the Crimean Peninsula, Ukraine</mods:title>
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<mods:namePart id="7271CC9C2FCEB9DA3F895B83F7BE1CCC">Kłonowska-Olejnik, Małgorzata</mods:namePart>
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<mods:namePart id="6A413F1A2939B5660DF12585C5D53F00">Prokopov, Grigorii A.</mods:namePart>
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<mods:namePart id="8865F5015D01CCD471F64D3CB70F1B79">Godunko, Roman J.</mods:namePart>
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<mods:title id="FA7F7324C1862F2D532884CCA89A6DA3">Zootaxa</mods:title>
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<mods:part id="6800021B0462557CA171646935DE8EB8">
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<mods:date id="6C94E39A7F566448E01F852416D37B3A">2007</mods:date>
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<mods:number id="2A38FF9B892F9CF929B68E178CAC3A95">1620</mods:number>
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<mods:identifier id="C751BB49AD2228005FC427C4779C5367" type="DOI">10.5281/zenodo.179171</mods:identifier>
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<treatment id="03A3C7011E69FF8A54C2FCB8BB6956EE" ID-DOI="http://doi.org/10.5281/zenodo.6240749" ID-GBIF-Taxon="119375952" ID-Zenodo-Dep="6240749" LSID="urn:lsid:plazi:treatment:03A3C7011E69FF8A54C2FCB8BB6956EE" httpUri="http://treatment.plazi.org/id/03A3C7011E69FF8A54C2FCB8BB6956EE" lastPageId="9" lastPageNumber="61" pageId="1" pageNumber="54">
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<subSubSection id="C310259C1E69FF8254C2FCB8BF275560" pageId="1" pageNumber="54" type="nomenclature">
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<paragraph id="8BB576171E69FF8254C2FCB8BA285505" blockId="1.[151,1040,791,852]" box="[151,1040,791,817]" pageId="1" pageNumber="54">
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<heading id="D0FDC17B1E69FF8254C2FCB8BA285505" bold="true" box="[151,1040,791,817]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
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<emphasis id="B97EAA051E69FF8254C2FCB8BA285505" bold="true" box="[151,1040,791,817]" pageId="1" pageNumber="54">
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<taxonomicName id="4C0A0D941E69FF8254C2FCB8BD9E5505" ID-CoL="6DST3" authority="Godunko, Klonowska-Olejnik and Prokopov" authorityName="Godunko, Klonowska-Olejnik and Prokopov" box="[151,934,791,817]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
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<emphasis id="B97EAA051E69FF8254C2FCB8BF575505" bold="true" box="[151,367,791,817]" italics="true" pageId="1" pageNumber="54">Ecdyonurus solus</emphasis>
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Godunko, Kłonowska-Olejnik and Prokopov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24D177E1E69FF8257E7FCB8BA285505" box="[946,1040,791,817]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB576171E69FF8254C2FC95BF275560" blockId="1.[151,1040,791,852]" box="[151,287,826,852]" pageId="1" pageNumber="54">
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(
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<figureCitation id="13316A921E69FF8254CAFC95BF2E5560" box="[159,278,826,852]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURES 4 – 6" captionStart-2="FIGURES 7 – 14" captionStart-3="FIGURES 15 – 19" captionStart-4="FIGURES 20 – 25" captionStartId-0="2.[151,269,1136,1160]" captionStartId-1="3.[151,269,1306,1330]" captionStartId-2="4.[151,269,1614,1638]" captionStartId-3="5.[151,269,1773,1797]" captionStartId-4="7.[151,269,1850,1874]" captionTargetBox-0="[280,1325,199,1102]" captionTargetBox-1="[297,1248,309,1278]" captionTargetBox-2="[248,1325,199,1575]" captionTargetBox-3="[227,1281,309,1772]" captionTargetBox-4="[200,1397,314,1828]" captionTargetId-0="figure@2.[236,1351,132,1150]" captionTargetId-1="figure@3.[237,1347,296,1300]" captionTargetId-2="figure@4.[224,1359,172,1608]" captionTargetId-3="figure@5.[227,1362,308,1775]" captionTargetId-4="figure@7.[200,1398,314,1829]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="7" captionText-0="FIGURES 1 – 3. Ecdyonurus solus sp. nov., male imago, paratype: 1, head, dorsal view; 2, head, lateral view; 3, abdominal segments 4 - 7, lateral view." captionText-1="FIGURES 4 – 6. Ecdyonurus solus sp. nov. Figs 4 – 5, male imago, holotype: 4, male genitalia, ventral view; 5, penis lobe, dorsal view. Fig 6, female imago, paratype: tip of abdomen, ventral view." captionText-2="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." captionText-3="FIGURES 15 – 19. Ecdyonurus solus sp. nov., larva, paratype: 15, abdomen, dorsal view; 16, two kinds of posterior margin of terga; 17, gill 1; 18, gill 4; 19, gill 7." captionText-4="FIGURES 20 – 25. Ecdyonurus solus sp. nov., egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S- 4700 scanning electron microscope)." httpUri-0="https://zenodo.org/record/179172/files/figure.png" httpUri-1="https://zenodo.org/record/179173/files/figure.png" httpUri-2="https://zenodo.org/record/179174/files/figure.png" httpUri-3="https://zenodo.org/record/179175/files/figure.png" httpUri-4="https://zenodo.org/record/179176/files/figure.png" pageId="1" pageNumber="54">Figs 1–25</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C310259C1E69FF8254C2FC25BA8155F5" pageId="1" pageNumber="54" type="reference_group">
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<paragraph id="8BB576171E69FF8254C2FC25BA8155F5" blockId="1.[151,1209,906,961]" pageId="1" pageNumber="54">
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<treatmentCitationGroup id="AB1A51391E69FF8254C2FC25BA8155F5" pageId="1" pageNumber="54">
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<taxonomicName id="4C0A0D941E69FF8254C2FC25BCC05595" ID-CoL="6DT4T" authority="Kiseleva and Ezernitskii 1985: 112" authorityName="Kiseleva and Ezernitskii" authorityPageNumber="112" authorityYear="1985" box="[151,760,906,929]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="venosus">
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<emphasis id="B97EAA051E69FF8254C2FC25BF495595" box="[151,369,906,929]" italics="true" pageId="1" pageNumber="54">Ecdyonurus venosus</emphasis>
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:
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<treatmentCitation id="0AAB50061E69FF82552AFC25BCC05595" author="Kiseleva" box="[383,760,906,929]" page="112" pageId="1" pageNumber="54" year="1985">
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<bibRefCitation id="EF9B0BE61E69FF82552AFC25BCFC5595" author="Kiseleva" box="[383,708,906,929]" pageId="1" pageNumber="62" refString="Kiseleva, G. A. & Ezernitskii, E. V. (1985) Distribution of the water entomofauna in the upper part of the Salgir riverbasin. Ekologicheskie i prirodookhrannye aspekty izucheniya Gornogo Kryma. Izd-vo SGU, 110 - 119. (in Russian)" type="book chapter" year="1985">Kiseleva and Ezernitskii 1985</bibRefCitation>
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: 112
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0AAB50061E69FF825756FC25BDED5595" author="Kiseleva" box="[771,981,906,929]" page="163" pageId="1" pageNumber="54" year="1993">
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<bibRefCitation id="EF9B0BE61E69FF825756FC25BD985595" author="Kiseleva" box="[771,928,906,929]" pageId="1" pageNumber="62" refString="Kiseleva, G. A. (1993) Aquatic insects in the water ecosystems of the small river of pre-mountain zone of the Crimea. Uspekhi entomologii v SSSR: Ekologiya i faunistika. Nebol'shie otryady nasekomykh. Izd-vo RAN, 163. (in Russian)" type="book chapter" year="1993">Kiseleva 1993</bibRefCitation>
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||
: 163
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||
</treatmentCitation>
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||
;
|
||
<treatmentCitation id="0AAB50061E69FF8257B5FC25BA905595" author="Godunko" box="[992,1192,906,929]" page="81" pageId="1" pageNumber="54" year="2001">
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<bibRefCitation id="EF9B0BE61E69FF8257B5FC25BABB5595" author="Godunko" box="[992,1155,906,929]" pageId="1" pageNumber="62" refString="Godunko, R. J. (2001) Biodiversity of mayflies (Insecta: Ephemeroptera) of the Crimean Peninsula. In: Gaino, E. (Ed.), 2001 International Joint Meeting. X International Conference on Ephemeroptera and XIV International Symposium on Plecoptera, University of Perugia, Perugia (Italy), pp. 80 - 81." type="journal article" year="2001">Godunko 2001</bibRefCitation>
|
||
: 81
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||
</treatmentCitation>
|
||
<taxonomicName id="4C0A0D941E69FF8254C2FC05BD3755F5" authority="Kiseleva and Ezernitsikii 1985: 112" authorityName="Kiseleva and Ezernitsikii" authorityPageNumber="112" authorityYear="1985" box="[151,783,938,961]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="species" species="fluminum">
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<emphasis id="B97EAA051E69FF8254C2FC05BFB955F5" box="[151,385,938,961]" italics="true" pageId="1" pageNumber="54">Ecdyonurus fluminum</emphasis>
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||
:
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<treatmentCitation id="0AAB50061E69FF8255DAFC05BD3755F5" author="Kiseleva" box="[399,783,938,961]" page="112" pageId="1" pageNumber="54" year="1985">
|
||
<bibRefCitation id="EF9B0BE61E69FF8255DAFC05BCE355F5" author="Kiseleva" box="[399,731,938,961]" pageId="1" pageNumber="62" refString="Kiseleva, G. A. & Ezernitskii, E. V. (1985) Distribution of the water entomofauna in the upper part of the Salgir riverbasin. Ekologicheskie i prirodookhrannye aspekty izucheniya Gornogo Kryma. Izd-vo SGU, 110 - 119. (in Russian)" type="book chapter" year="1985">Kiseleva and Ezernitsikii 1985</bibRefCitation>
|
||
: 112
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||
</treatmentCitation>
|
||
</taxonomicName>
|
||
;
|
||
<treatmentCitation id="0AAB50061E69FF82574FFC05BDD455F5" author="Kiseleva" box="[794,1004,938,961]" page="116" pageId="1" pageNumber="54" year="1992">
|
||
<bibRefCitation id="EF9B0BE61E69FF82574FFC05BD8F55F5" author="Kiseleva" box="[794,951,938,961]" pageId="1" pageNumber="62" refString="Kiseleva, G. A. (1992) Benthic fauna of the small rivers of the mountain and pre-mountain zone of the Crimea. Ratsional'noe ispol'zovanie i okhrana ekosistem Kryma. UMK VO, 116. (in Russian)" type="book chapter" year="1992">Kiseleva 1992</bibRefCitation>
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||
: 116
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||
</treatmentCitation>
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||
; Kiseleva 1997: 39
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||
</treatmentCitationGroup>
|
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</paragraph>
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</subSubSection>
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<subSubSection id="C310259C1E69FF8554C2FC5DBDE25216" lastPageId="6" lastPageNumber="59" pageId="1" pageNumber="54" type="description">
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<paragraph id="8BB576171E69FF8254C2FC5DBA5352B0" blockId="1.[151,1437,1010,2036]" pageId="1" pageNumber="54">
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<emphasis id="B97EAA051E69FF8254C2FC5DBF175238" bold="true" box="[151,303,1010,1036]" pageId="1" pageNumber="54">Description.</emphasis>
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<emphasis id="B97EAA051E69FF825563FC5DBFFA523F" box="[310,450,1010,1035]" italics="true" pageId="1" pageNumber="54">Male imago</emphasis>
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(all adults were reared from larvae). Size: body 10.0–
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||
<quantity id="4CF2DBF21E69FF82506FFC5DBA9D5238" box="[1082,1189,1010,1036]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="1" pageNumber="54" unit="mm" value="10.4">10.4 mm</quantity>
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; fore wing 11.0–
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||
<quantity id="4CF2DBF21E69FF82513FFC5DBEF05200" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="54" unit="mm" value="11.2">11.2 mm</quantity>
|
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; hind wings
|
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<quantity id="4CF2DBF21E69FF825509FBB5BFD45200" box="[348,492,1050,1076]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.3" metricValueMin="3.9" pageId="1" pageNumber="54" unit="mm" value="4.1" valueMax="4.3" valueMin="3.9">3.9–4.3 mm</quantity>
|
||
; fore legs
|
||
<quantity id="4CF2DBF21E69FF825630FBB5BD375200" box="[613,783,1050,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.09" metricValueMin="1.02" pageId="1" pageNumber="54" unit="mm" value="10.55" valueMax="10.9" valueMin="10.2">10.2–10.9 mm</quantity>
|
||
(femur
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||
<quantity id="4CF2DBF21E69FF825738FBB5BDC55200" box="[877,1021,1050,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="1" pageNumber="54" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.6–2.7 mm</quantity>
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; tibia
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||
<quantity id="4CF2DBF21E69FF825011FBB5BAEC5200" box="[1092,1236,1050,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="1" pageNumber="54" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
|
||
; tarsal segments: T1 = 0.8–1.0 mm; T2 =
|
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<quantity id="4CF2DBF21E69FF8255E2FBEDBC705268" box="[439,584,1090,1116]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="54" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
|
||
; T3 =
|
||
<quantity id="4CF2DBF21E69FF8256C2FBEDBD105268" box="[663,808,1090,1116]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="54" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
|
||
; T4 =
|
||
<quantity id="4CF2DBF21E69FF825723FBEDBDEC5268" box="[886,980,1090,1116]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="54" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; T5 =
|
||
<quantity id="4CF2DBF21E69FF825077FBEDBAB85268" box="[1058,1152,1090,1116]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="54" unit="mm" value="0.5">0.5 mm</quantity>
|
||
; gradation of tarsal segments: 2>=3>4>1>5); cerci
|
||
<quantity id="4CF2DBF21E69FF8255B4FBC5BCB752B0" box="[481,655,1130,1156]" metricMagnitude="-2" metricUnit="m" metricValue="3.005" metricValueMax="3.05" metricValueMin="2.96" pageId="1" pageNumber="54" unit="mm" value="30.05" valueMax="30.5" valueMin="29.6">29.6–30.5 mm</quantity>
|
||
. General color of body yellow to brown.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E69FF825492FB3DBD695310" blockId="1.[151,1437,1010,2036]" pageId="1" pageNumber="54">
|
||
Head yellowish with brownish and brownish-gray smudges on short facial keel. Antennae brown. Ocelli black basally with subapical yellowish-brown ring and whitish or white-gray apically. Eyes are separated by a distinct gap comparable in size with the central ocellus, gray-blackish, with light gray ring around of margin. Lateral portion of eyes with two light gray rings (
|
||
<figureCitation id="13316A921E69FF82568CFAA5BD7C5310" box="[729,836,1290,1316]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,269,1136,1160]" captionTargetBox="[280,1325,199,1102]" captionTargetId="figure@2.[236,1351,132,1150]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Ecdyonurus solus sp. nov., male imago, paratype: 1, head, dorsal view; 2, head, lateral view; 3, abdominal segments 4 - 7, lateral view." httpUri="https://zenodo.org/record/179172/files/figure.png" pageId="1" pageNumber="54">Figs 1, 2</figureCitation>
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||
).
|
||
</paragraph>
|
||
<paragraph id="8BB576171E69FF825492FA9DBF385008" blockId="1.[151,1437,1010,2036]" pageId="1" pageNumber="54">Thorax yellow to brown, dorsally with dark brown maculation on metanotum and hind part of mesonotum. Pronotum light yellowish-brown. Lateral part of thorax generally whitish to yellowish-brown. Ventral part of thorax distinctly brown to brownish-black. Wing hyaline, transparent, unicolorous. Pterostigmatic area clearly yellow, costal area yellowish-white. Venation brown to blackish, lighter basally. Fore legs distinctly darker than middle and hind legs: femora dark brown; tibiae and tarsi brown, with slightly dark last tarsal segment. Middle and hind legs with yellow femora and tibiae; tarsi yellowish-brown basally with brownish last segment.</paragraph>
|
||
<paragraph id="8BB576171E69FF825492F9E5BB545118" blockId="1.[151,1437,1010,2036]" pageId="1" pageNumber="54">
|
||
Abdominal terga yellow to yellowish-brown. Tergum 1 brown, with blackish-brown strips near hind margin of segment; tergum 2 with central longitudinal light band and two brownish paramedian bands; terga 3–10 yellowish, with two longitudinal narrow paramedia bands (terga 7–10 slightly darker that others ones). Laterally terga 2–8 with distinct blackish-brown stripes connected dorsally in posterior part of segments (
|
||
<figureCitation id="13316A921E69FF825116F96DBBB450E8" box="[1347,1420,1730,1756]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,269,1136,1160]" captionTargetBox="[280,1325,199,1102]" captionTargetId="figure@2.[236,1351,132,1150]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Ecdyonurus solus sp. nov., male imago, paratype: 1, head, dorsal view; 2, head, lateral view; 3, abdominal segments 4 - 7, lateral view." httpUri="https://zenodo.org/record/179172/files/figure.png" pageId="1" pageNumber="54">Fig. 3</figureCitation>
|
||
). Sterna 1 and 9 brown, sterna 2–8 uniformly yellowish with two unclear brownish spots. Violet or blackish nerve ganglia well visible on the sterna 2–8. Cerci brownish, lighter apically. Joints of segments blackish.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E69FF825492F895BC5451C0" blockId="1.[151,1437,1010,2036]" pageId="1" pageNumber="54">
|
||
Styliger plate light brown, with two small and rounded apically protuberance near forceps base (
|
||
<figureCitation id="13316A921E69FF825111F895BBB65160" box="[1348,1422,1850,1876]" captionStart="FIGURES 4 – 6" captionStartId="3.[151,269,1306,1330]" captionTargetBox="[297,1248,309,1278]" captionTargetId="figure@3.[237,1347,296,1300]" captionTargetPageId="3" captionText="FIGURES 4 – 6. Ecdyonurus solus sp. nov. Figs 4 – 5, male imago, holotype: 4, male genitalia, ventral view; 5, penis lobe, dorsal view. Fig 6, female imago, paratype: tip of abdomen, ventral view." httpUri="https://zenodo.org/record/179173/files/figure.png" pageId="1" pageNumber="54">Fig. 4</figureCitation>
|
||
). Forceps brown. Penis lobes light brown, slightly stretched laterally and convergent toward outer margins, with rounded distal part. Penis light brown. Basal sclerite large, partially cover the basal part of lateral sclerite. Lateral sclerite narrow and curved, distinctly convergent distally. Apical sclerite relatively wide, not convergent apically. Titilators wide, brown (
|
||
<figureCitation id="13316A921E69FF825640F875BC6451C0" box="[533,604,2010,2036]" captionStart="FIGURES 4 – 6" captionStartId="3.[151,269,1306,1330]" captionTargetBox="[297,1248,309,1278]" captionTargetId="figure@3.[237,1347,296,1300]" captionTargetPageId="3" captionText="FIGURES 4 – 6. Ecdyonurus solus sp. nov. Figs 4 – 5, male imago, holotype: 4, male genitalia, ventral view; 5, penis lobe, dorsal view. Fig 6, female imago, paratype: tip of abdomen, ventral view." httpUri="https://zenodo.org/record/179173/files/figure.png" pageId="1" pageNumber="54">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF75269F1E6AFF8154C2FBDFBFE4529E" httpUri="https://zenodo.org/record/179172/files/figure.png" pageId="2" pageNumber="55" targetBox="[280,1325,199,1102]" targetPageId="2">
|
||
<paragraph id="8BB576171E6AFF8154C2FBDFBFE4529E" blockId="2.[151,1436,1136,1194]" pageId="2" pageNumber="55">
|
||
<emphasis id="B97EAA051E6AFF8154C2FBDFBF7A52BC" bold="true" box="[151,322,1136,1160]" pageId="2" pageNumber="55">FIGURES 1–3.</emphasis>
|
||
<taxonomicName id="4C0A0D941E6AFF81551CFBDFBC3E52B3" box="[329,518,1136,1159]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E6AFF81551CFBDFBC3E52B3" box="[329,518,1136,1159]" italics="true" pageId="2" pageNumber="55">Ecdyonurus solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E6AFF815659FBDFBC5952B3" box="[524,609,1136,1159]" pageId="2" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male imago, paratype: 1, head, dorsal view; 2, head, lateral view; 3, abdominal segments 4-7, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB576171E6AFF815492FB76BCDC5376" blockId="2.[151,1437,1240,2026]" pageId="2" pageNumber="55">
|
||
<emphasis id="B97EAA051E6AFF815492FB76BF4F52C6" box="[199,375,1241,1266]" italics="true" pageId="2" pageNumber="55">Female imago.</emphasis>
|
||
Size: body
|
||
<quantity id="4CF2DBF21E6AFF8155AAFB77BC5252C6" box="[511,618,1240,1266]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="2" pageNumber="55" unit="mm" value="12.2">12.2 mm</quantity>
|
||
; fore wing 12.0 mm; hind wings 4.0 mm; cerci
|
||
<quantity id="4CF2DBF21E6AFF8150C7FB77BAC652C6" box="[1170,1278,1240,1266]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="2" pageNumber="55" unit="mm" value="19.5">19.5 mm</quantity>
|
||
. Head yellow with blackish smudges on facial keel. Eyes and apical part of ocelli grayish-black. Central part of ocelli light yellow; apical part whitish. Antennae light brown.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6AFF815492FAFFBC9F5006" blockId="2.[151,1437,1240,2026]" pageId="2" pageNumber="55">Thorax yellow to dark brown. Pronotum yellowish-gray with two brownish spots centrally. Mesonotum yellowish-brown with brown to dark brown distal part. Metanotum dark brown. Lateral part of thorax yellowish-brown. Sterna of thorax brown to dark brown. Wings transparent, hyaline slightly yellowish color. Pterostigmatic area yellowish, opaque. Venation yellow to dark brown. Fore legs with light brown femora, and brown to dark brown tibiae and tarsi. Middle and hind legs with unicolorous yellow femora and yellowish-brown tibiae. Tarsi brown to dark-brown.</paragraph>
|
||
<paragraph id="8BB576171E6AFF815492F9EFBCBB50CE" blockId="2.[151,1437,1240,2026]" pageId="2" pageNumber="55">
|
||
Abdominal terga with central longitudinal bands and two short reddish strokes on the segments. Two distinct triangular reddish or brownish spots are present on central part of tergum 1; lateral part of terga 2–8 with blackish stripes. Abdominal sterna yellow to reddish-brown, with uniformly pattern on the segments 1–7 (generally present two central light short strokes). Sterna 2-8 with visible violet or blackish nerve ganglia. Subgenital and subanal plates as in
|
||
<figureCitation id="13316A921E6AFF815662F94FBC4750CE" box="[567,639,1760,1786]" captionStart="FIGURES 4 – 6" captionStartId="3.[151,269,1306,1330]" captionTargetBox="[297,1248,309,1278]" captionTargetId="figure@3.[237,1347,296,1300]" captionTargetPageId="3" captionText="FIGURES 4 – 6. Ecdyonurus solus sp. nov. Figs 4 – 5, male imago, holotype: 4, male genitalia, ventral view; 5, penis lobe, dorsal view. Fig 6, female imago, paratype: tip of abdomen, ventral view." httpUri="https://zenodo.org/record/179173/files/figure.png" pageId="2" pageNumber="55">Fig. 6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6AFF815492F8A7BD94517E" blockId="2.[151,1437,1240,2026]" pageId="2" pageNumber="55">
|
||
<emphasis id="B97EAA051E6AFF815492F8A7BFBF5115" box="[199,391,1800,1825]" italics="true" pageId="2" pageNumber="55">Male subimago.</emphasis>
|
||
Size: body
|
||
<quantity id="4CF2DBF21E6AFF81564CF8A7BC415116" box="[537,633,1800,1826]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="55" unit="mm" value="9.5">9.5 mm</quantity>
|
||
; fore wing 12.0 mm; hind wings 4.0 mm; cerci
|
||
<quantity id="4CF2DBF21E6AFF815096F8A7BB095116" box="[1219,1329,1800,1826]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="2" pageNumber="55" unit="mm" value="19.5">19.5 mm</quantity>
|
||
. General color of body yellow and yellowish-gray to brown and dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6AFF815492F8F7BD3051AE" blockId="2.[151,1437,1240,2026]" pageId="2" pageNumber="55">Head grayish brown, antennae brownish. Eyes grayish-black with one hard visible grayish-white ring laterally. Ocelli blackish basally, with grayish-white tip.</paragraph>
|
||
<paragraph id="8BB576171E6AFF805492F807BD825736" blockId="2.[151,1437,1240,2026]" lastBlockId="3.[151,1436,152,258]" lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55">Pronotum brownish-gray. Mesonotum and metanotum yellowish-white with distinct dark brown bands. Wings unicolorous yellowish, relatively transparent. Venation whitish-gray to brownish. Transversal veins darker than longitudinal with grayish-brown smudges around veins. Fore legs dark: femora brown slightly darker distally; tibiae dark brown; tarsi unicolorous brown. Middle and hind legs: femora yellow with elongate brownish spot apically; tibiae yellowish-brown; tarsi yellowish.</paragraph>
|
||
<caption id="DF75269F1E6BFF8054C2FAB5BD9D5360" httpUri="https://zenodo.org/record/179173/files/figure.png" pageId="3" pageNumber="56" targetBox="[297,1248,309,1278]" targetPageId="3">
|
||
<paragraph id="8BB576171E6BFF8054C2FAB5BD9D5360" blockId="3.[151,1436,1306,1364]" pageId="3" pageNumber="56">
|
||
<emphasis id="B97EAA051E6BFF8054C2FAB5BF7A5306" bold="true" box="[151,322,1306,1330]" pageId="3" pageNumber="56">FIGURES 4–6.</emphasis>
|
||
<taxonomicName id="4C0A0D941E6BFF80551DFAB4BC3D5306" box="[328,517,1307,1330]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E6BFF80551DFAB4BC3D5306" box="[328,517,1307,1330]" italics="true" pageId="3" pageNumber="56">Ecdyonurus solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E6BFF80565EFAB4BC655306" box="[523,605,1307,1330]" pageId="3" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Figs 4–5, male imago, holotype: 4, male genitalia, ventral view; 5, penis lobe, dorsal view. Fig 6, female imago, paratype: tip of abdomen, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB576171E6BFF805492FA2CBDEA5021" blockId="3.[151,1437,1411,2037]" pageId="3" pageNumber="56">Abdominal terga yellow to yellowish-brown. Sterna 1–8 slightly darker than terga with two light short strokes and two light spots centrally. Sternum 8 with violet nerve ganglia. Lateral part of terga 1–8 with distinct blackish-brown stripes connected distally in posterior part of segments. Styliger plate yellowish-brown, forceps slightly darker. Cerci light brown. Joints of segments blackish.</paragraph>
|
||
<paragraph id="8BB576171E6BFF805492F98CBB195051" blockId="3.[151,1437,1411,2037]" pageId="3" pageNumber="56">
|
||
<emphasis id="B97EAA051E6BFF805492F98CBF9B5008" box="[199,419,1571,1596]" italics="true" pageId="3" pageNumber="56">Female subimago.</emphasis>
|
||
Size: body
|
||
<quantity id="4CF2DBF21E6BFF805660F98CBCDE5009" box="[565,742,1571,1597]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.38" metricValueMin="1.27" pageId="3" pageNumber="56" unit="mm" value="13.25" valueMax="13.8" valueMin="12.7">12.7–13.8 mm</quantity>
|
||
; fore wing 13.0–
|
||
<quantity id="4CF2DBF21E6BFF8057E0F98CBA1B5009" box="[949,1059,1571,1597]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="3" pageNumber="56" unit="mm" value="14.2">14.2 mm</quantity>
|
||
; hind wings 4.2–5.0 mm; cerci
|
||
<quantity id="4CF2DBF21E6BFF8054C2F9E4BF7D5051" box="[151,325,1611,1637]" metricMagnitude="-2" metricUnit="m" metricValue="1.295" metricValueMax="1.3699999999999999" metricValueMin="1.22" pageId="3" pageNumber="56" unit="mm" value="12.95" valueMax="13.7" valueMin="12.2">12.2–13.7 mm</quantity>
|
||
. General color of body very similar to those of female imago except wings and legs.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6BFF805492F9DCBB2650E9" blockId="3.[151,1437,1411,2037]" pageId="3" pageNumber="56">Thorax yellow to dark brown. Wings distinctly yellowish to light brown, slightly opaque, especially in pterostigmatic area. Veins dark brown to black. Brownish smudges around veins are presented. Fore legs unicolorous dark brown with black tarsi. Middle and hind legs light brown. Tarsi dark brown to black.</paragraph>
|
||
<paragraph id="8BB576171E6BFF805492F944BF1D5191" blockId="3.[151,1437,1411,2037]" pageId="3" pageNumber="56">Abdominal terga with distinct drawing: tergum 1 with diffuse central brownish spots; terga 2–3 with central diffuse brownish and two yellow spots; terga 4–5 with longitudinal brownish spot; terga 6–7 with two pair of small brownish spots near anterior part of segment; terga 8–10 uniformly yellowish with central brownish smudges. Lateral side of terga 1–8 with distinct dark brown strips. Sternum 8 with violet nerve ganglia. Cerci dark brown.</paragraph>
|
||
<paragraph id="8BB576171E6BFF805492F81CBAA751C1" blockId="3.[151,1437,1411,2037]" pageId="3" pageNumber="56">
|
||
<emphasis id="B97EAA051E6BFF805492F81CBF5351F8" box="[199,363,1971,1996]" italics="true" pageId="3" pageNumber="56">Mature larva.</emphasis>
|
||
Body length
|
||
<quantity id="4CF2DBF21E6BFF805658F81CBC8451F9" box="[525,700,1971,1997]" metricMagnitude="-2" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.35" metricValueMin="1.1199999999999999" pageId="3" pageNumber="56" unit="mm" value="12.35" valueMax="13.5" valueMin="11.2">11.2–13.5 mm</quantity>
|
||
; caudal filaments 7.4–10.0 mm. General color yellow to light brown. Abdominal segments of mature larvae with well visible pattern similar to adults.
|
||
</paragraph>
|
||
<caption id="DF75269F1E6CFF8754C2F9E1BBA550BC" httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="4" pageNumber="57" targetBox="[248,1325,199,1575]" targetPageId="4">
|
||
<paragraph id="8BB576171E6CFF8754C2F9E1BBA550BC" blockId="4.[151,1437,1614,1672]" pageId="4" pageNumber="57">
|
||
<emphasis id="B97EAA051E6CFF8754C2F9E1BF695052" bold="true" box="[151,337,1614,1638]" pageId="4" pageNumber="57">FIGURES 7–14.</emphasis>
|
||
<taxonomicName id="4C0A0D941E6CFF87550CF9E0BC2F5052" box="[345,535,1615,1638]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E6CFF87550CF9E0BC2F5052" box="[345,535,1615,1638]" italics="true" pageId="4" pageNumber="57">Ecdyonurus solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E6CFF87564AF9E0BC4D5052" box="[543,629,1615,1638]" pageId="4" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB576171E6CFF865492F913BBA4571E" blockId="4.[151,1437,1724,2030]" lastBlockId="5.[151,1436,152,298]" lastPageId="5" lastPageNumber="58" pageId="4" pageNumber="57">
|
||
Head yellow, oval, relatively large, the part at the eyes level not widest. Antennae yellow to light brown. Mouthparts: Labrum not large, slightly stretched laterally, dorsally with one row of 7–9 median strong bristles (
|
||
<figureCitation id="13316A921E6CFF8754CAF8A3BEDF5112" box="[159,231,1804,1830]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1614,1638]" captionTargetBox="[248,1325,199,1575]" captionTargetId="figure@4.[224,1359,172,1608]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="4" pageNumber="57">Fig. 7</figureCitation>
|
||
), typical for the
|
||
<taxonomicName id="4C0A0D941E6CFF8755FBF8A3BC155111" box="[430,557,1804,1829]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="venosus">
|
||
<emphasis id="B97EAA051E6CFF8755FBF8A3BC155111" box="[430,557,1804,1829]" italics="true" pageId="4" pageNumber="57">E. venosus</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<bibRefCitation id="EF9B0BE61E6CFF8756BDF8A3BA185112" author="Belfiore" box="[744,1056,1804,1830]" pageId="4" pageNumber="62" refString="Belfiore, C. & Buffagni, A. (1994) Revision of the Italian species of the Ecdyonurus helveticus - group; taxonomy of the nymphs (Ephemeroptera: Heptageniidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 67, 143 - 149." type="journal article" year="1994">Belfiore & Buffagni 1994</bibRefCitation>
|
||
). Hypopharynx without specific features, generally with long hair on the outer margin and distal part, typical for the
|
||
<taxonomicName id="4C0A0D941E6CFF875021F89BBACA5179" box="[1140,1266,1844,1869]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="venosus">
|
||
<emphasis id="B97EAA051E6CFF875021F89BBACA5179" box="[1140,1266,1844,1869]" italics="true" pageId="4" pageNumber="57">E. venosus</emphasis>
|
||
</taxonomicName>
|
||
species-group (
|
||
<figureCitation id="13316A921E6CFF8754CAF8F3BED15142" box="[159,233,1884,1910]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1614,1638]" captionTargetBox="[248,1325,199,1575]" captionTargetId="figure@4.[224,1359,172,1608]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="4" pageNumber="57">Fig. 8</figureCitation>
|
||
). Mandibles (n = 20) with 6–9 prosthecal bristles. Maxillae (n = 20) (characters listed by
|
||
<bibRefCitation id="EF9B0BE61E6CFF875161F8F3BECB51AA" author="Haybach" pageId="4" pageNumber="62" refString="Haybach, A. (1999) Contribution to larval taxonomy of Ecdyonurus venosus - group in Germany. Lauterbornia, 37, 113 - 150. (in German, Engl. sum.)" type="journal article" year="1999">Haybach (1999))</bibRefCitation>
|
||
: number of comb-shaped bristles (N_CBS) = 17–22; number of teeth on 5th comb-shaped bristle (N_TCB5) = 14–17; number of hairs on dorsal upper side (N_DOR) = 14–18; outer margin of maxillae without hairs (N_OUT = 0); number of hairs on ventral basal part of maxillae (N_VEN) = 21–24; number of hairs at the base of maxillary palps (N_PLBas) = 16–20; outer base of the first segment of maxillary palps without hairs (N_PLH = 0); number of setae on the outer margin of the first segment of maxillary palps (N_PLS)>50; number of setae on the inner side of the first segment of maxillary palps (N_PLP)>40. Labial palps with numerous hairs of the group on the dorsal side of its first segment (N_LPH = 28–36) arranged on 2–3 rows.
|
||
</paragraph>
|
||
<caption id="DF75269F1E6DFF8654C2F942BC595113" httpUri="https://zenodo.org/record/179175/files/figure.png" pageId="5" pageNumber="58" targetBox="[227,1281,309,1772]" targetPageId="5">
|
||
<paragraph id="8BB576171E6DFF8654C2F942BC595113" blockId="5.[151,1436,1773,1831]" pageId="5" pageNumber="58">
|
||
<emphasis id="B97EAA051E6DFF8654C2F942BF655131" bold="true" box="[151,349,1773,1797]" pageId="5" pageNumber="58">FIGURES 15–19.</emphasis>
|
||
<taxonomicName id="4C0A0D941E6DFF865531F941BC185131" box="[356,544,1774,1797]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E6DFF865531F941BC185131" box="[356,544,1774,1797]" italics="true" pageId="5" pageNumber="58">Ecdyonurus solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E6DFF865673F942BC435130" box="[550,635,1773,1796]" pageId="5" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, larva, paratype: 15, abdomen, dorsal view; 16, two kinds of posterior margin of terga; 17, gill 1; 18, gill 4; 19, gill 7.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB576171E6DFF855492F8F7BC3E5766" blockId="5.[151,1436,1880,2026]" lastBlockId="6.[151,1437,152,2016]" lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58">
|
||
Pronotum yellow with some light central smudges. Lateral projection of pronotum short, distinctly asymmetrical, with apical part curved towards the body (
|
||
<figureCitation id="13316A921E6DFF8656AFF82FBD6B51AE" box="[762,851,1920,1946]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1614,1638]" captionTargetBox="[248,1325,199,1575]" captionTargetId="figure@4.[224,1359,172,1608]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="5" pageNumber="58">Fig. 10</figureCitation>
|
||
). The width/length ratio of semipronotum to caudal projection (see
|
||
<bibRefCitation id="EF9B0BE61E6DFF86552CF807BCCB51F6" author="Bauernfeind" box="[377,755,1960,1986]" pageId="5" pageNumber="62" refString="Bauernfeind, E. & Humpesch, U. H. (2001) Eintagsfliegen Zentraluropas (Insecta: Ephemeroptera): Bestimmung und Okologie. Verlag des Naturhistorischen Museums Wien, Wien, 239 pp." type="book" year="2001">Bauernfeind & Humpesch 2001</bibRefCitation>
|
||
) is 3.8–4.2. Mesonotum yellow to yellowish-brown, with whitish spots laterally. Legs yellow to light brown. Femora relatively slender and long. The length/width ratio of metafemora on average 2.67. Dorsal surface of femora mainly with distinct pattern (four yellowish lateral spots on a dark field) (
|
||
<figureCitation id="13316A921E6EFF8555C8FF6FBFCB56EE" box="[413,499,192,218]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1614,1638]" captionTargetBox="[248,1325,199,1575]" captionTargetId="figure@4.[224,1359,172,1608]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="6" pageNumber="59">Fig. 11</figureCitation>
|
||
). Femoral scales of various
|
||
<typeStatus id="54B1C8B51E6EFF855769FF6FBD4256EE" box="[828,890,192,218]" pageId="6" pageNumber="59">types</typeStatus>
|
||
(
|
||
<figureCitation id="13316A921E6EFF8557DFFF6FBDD956EE" box="[906,993,192,218]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1614,1638]" captionTargetBox="[248,1325,199,1575]" captionTargetId="figure@4.[224,1359,172,1608]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="6" pageNumber="59">Fig. 12</figureCitation>
|
||
). Outer margin of femora with row of long slender bristles and small pointed spines; subapical spines is pointed or bluntly pointed (
|
||
<figureCitation id="13316A921E6EFF8550B1FF47BB045736" box="[1252,1340,232,258]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1614,1638]" captionTargetBox="[248,1325,199,1575]" captionTargetId="figure@4.[224,1359,172,1608]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="6" pageNumber="59">Fig. 13</figureCitation>
|
||
). Tibiae yellow. Tarsi yellow to light brown basally with brown or dark brown apical ring. Tarsal claw brown to dark brown with 2–3 teeth (
|
||
<figureCitation id="13316A921E6EFF8555F4FE97BFC05766" box="[417,504,312,338]" captionStart="FIGURES 7 – 14" captionStartId="4.[151,269,1614,1638]" captionTargetBox="[248,1325,199,1575]" captionTargetId="figure@4.[224,1359,172,1608]" captionTargetPageId="4" captionText="FIGURES 7 – 14. Ecdyonurus solus sp. nov., larva, paratype: 7, two kinds of lateral part of labrum; 8, hypopharynx; 9, glossa and paraglossa; 10, pronotum; 11, hind leg; 12, shape of femoral scales; 13, outer margin of femur; 14, tarsal claw." httpUri="https://zenodo.org/record/179174/files/figure.png" pageId="6" pageNumber="59">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6EFF855492FECFBD995476" blockId="6.[151,1437,152,2016]" pageId="6" pageNumber="59">
|
||
Abdominal terga yellow to light brown. Tergum 1 uniformly light; terga 2–7 and 10 (sometime 8) with two light central longitudinal spots (sometimes, with two pair of light spots); Tergum 9 with wide central light spot (
|
||
<figureCitation id="13316A921E6EFF85548CFE1FBF0857FE" box="[217,304,432,458]" captionStart="FIGURES 15 – 19" captionStartId="5.[151,269,1773,1797]" captionTargetBox="[227,1281,309,1772]" captionTargetId="figure@5.[227,1362,308,1775]" captionTargetPageId="5" captionText="FIGURES 15 – 19. Ecdyonurus solus sp. nov., larva, paratype: 15, abdomen, dorsal view; 16, two kinds of posterior margin of terga; 17, gill 1; 18, gill 4; 19, gill 7." httpUri="https://zenodo.org/record/179175/files/figure.png" pageId="6" pageNumber="59">Fig. 15</figureCitation>
|
||
). Posterior margins of terga with large pointed marginal teeth (
|
||
<figureCitation id="13316A921E6EFF85504EFE1FBA4C57FE" box="[1051,1140,432,458]" captionStart="FIGURES 15 – 19" captionStartId="5.[151,269,1773,1797]" captionTargetBox="[227,1281,309,1772]" captionTargetId="figure@5.[227,1362,308,1775]" captionTargetPageId="5" captionText="FIGURES 15 – 19. Ecdyonurus solus sp. nov., larva, paratype: 15, abdomen, dorsal view; 16, two kinds of posterior margin of terga; 17, gill 1; 18, gill 4; 19, gill 7." httpUri="https://zenodo.org/record/179175/files/figure.png" pageId="6" pageNumber="59">Fig. 16</figureCitation>
|
||
). Sterna unicolorous yellowish-brown with well visible violet nerve ganglia on the segments 2–8. Gills yellowish to light brown. Gill 1 relatively long with broadly rounded apical part (
|
||
<figureCitation id="13316A921E6EFF8556B9FDAFBD7A542E" box="[748,834,512,538]" captionStart="FIGURES 15 – 19" captionStartId="5.[151,269,1773,1797]" captionTargetBox="[227,1281,309,1772]" captionTargetId="figure@5.[227,1362,308,1775]" captionTargetPageId="5" captionText="FIGURES 15 – 19. Ecdyonurus solus sp. nov., larva, paratype: 15, abdomen, dorsal view; 16, two kinds of posterior margin of terga; 17, gill 1; 18, gill 4; 19, gill 7." httpUri="https://zenodo.org/record/179175/files/figure.png" pageId="6" pageNumber="59">Fig. 17</figureCitation>
|
||
). Gill 4 distinctly wide and asymmetrical (
|
||
<figureCitation id="13316A921E6EFF855163FDAFBBB5542E" box="[1334,1421,512,538]" captionStart="FIGURES 15 – 19" captionStartId="5.[151,269,1773,1797]" captionTargetBox="[227,1281,309,1772]" captionTargetId="figure@5.[227,1362,308,1775]" captionTargetPageId="5" captionText="FIGURES 15 – 19. Ecdyonurus solus sp. nov., larva, paratype: 15, abdomen, dorsal view; 16, two kinds of posterior margin of terga; 17, gill 1; 18, gill 4; 19, gill 7." httpUri="https://zenodo.org/record/179175/files/figure.png" pageId="6" pageNumber="59">Fig. 18</figureCitation>
|
||
). Gill 7 as in
|
||
<figureCitation id="13316A921E6EFF855575FD87BF4D5476" box="[288,373,552,578]" captionStart="FIGURES 15 – 19" captionStartId="5.[151,269,1773,1797]" captionTargetBox="[227,1281,309,1772]" captionTargetId="figure@5.[227,1362,308,1775]" captionTargetPageId="5" captionText="FIGURES 15 – 19. Ecdyonurus solus sp. nov., larva, paratype: 15, abdomen, dorsal view; 16, two kinds of posterior margin of terga; 17, gill 1; 18, gill 4; 19, gill 7." httpUri="https://zenodo.org/record/179175/files/figure.png" pageId="6" pageNumber="59">Fig. 19</figureCitation>
|
||
, without tuft. Cerci yellow to yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6EFF855492FDFFBDE25216" blockId="6.[151,1437,152,2016]" pageId="6" pageNumber="59">
|
||
<emphasis id="B97EAA051E6EFF855492FDFFBEC6545D" box="[199,254,592,617]" italics="true" pageId="6" pageNumber="59">Egg.</emphasis>
|
||
Size: length: 164–171 µm; width 108–116 µm. Egg oval. The eggs are characterized by attachment structures, tubercles, and very small granules (
|
||
<figureCitation id="13316A921E6EFF8556E6FDD7BD3354A6" box="[691,779,632,658]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,269,1850,1874]" captionTargetBox="[200,1397,314,1828]" captionTargetId="figure@7.[200,1398,314,1829]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Ecdyonurus solus sp. nov., egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S- 4700 scanning electron microscope)." httpUri="https://zenodo.org/record/179176/files/figure.png" pageId="6" pageNumber="59">Fig. 20</figureCitation>
|
||
). The attachment structures are represented by knob-terminated coiled threads (KCTs) (
|
||
<bibRefCitation id="EF9B0BE61E6EFF855642FD0FBD0A548E" author="Koss" box="[535,818,672,698]" pageId="6" pageNumber="62" refString="Koss, R. W. & Edmunds, G. F. Jr. (1974) Ephemeroptera eggs and their contribution to phylogenetic studies of the order. Zoological Journal of the Linnaean Society, 55, 267 - 349, pl. 1 - 24." type="journal article" year="1974">Koss & Edmunds 1974</bibRefCitation>
|
||
) and are of two different kinds. The first are larger (diameter 3.9–5.3 µm) and concentrated (aggregated) at one egg pole, covering a densely chorionic surface (spaced at a distance of 2.6–3.9 µm) (
|
||
<figureCitation id="13316A921E6EFF855602FD5FBC97553E" box="[599,687,752,778]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,269,1850,1874]" captionTargetBox="[200,1397,314,1828]" captionTargetId="figure@7.[200,1398,314,1829]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Ecdyonurus solus sp. nov., egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S- 4700 scanning electron microscope)." httpUri="https://zenodo.org/record/179176/files/figure.png" pageId="6" pageNumber="59">Fig. 21</figureCitation>
|
||
). This structure is similar to that of
|
||
<taxonomicName id="4C0A0D941E6EFF85500CFD5FBAEF553D" box="[1113,1239,752,777]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="venosus">
|
||
<emphasis id="B97EAA051E6EFF85500CFD5FBAEF553D" box="[1113,1239,752,777]" italics="true" pageId="6" pageNumber="59">E. venosus</emphasis>
|
||
</taxonomicName>
|
||
KCTs (
|
||
<bibRefCitation id="EF9B0BE61E6EFF855166FD5FBF175506" author="Gaino" pageId="6" pageNumber="62" refString="Gaino, E. & Rebora, M. (2003) Adhesiveness of the eggs of Ecdyonurus venosus to siliceous and calcareous substrate. In: Gaino, E. (Ed.), Research update on Ephemeroptera & Plecoptera, University of Perugia, Perugia (Italy), pp. 437 - 443." type="book chapter" year="2003">Gaino & Rebora 2003</bibRefCitation>
|
||
) (
|
||
<figureCitation id="13316A921E6EFF855512FCB7BFA75506" box="[327,415,792,818]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,269,1850,1874]" captionTargetBox="[200,1397,314,1828]" captionTargetId="figure@7.[200,1398,314,1829]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Ecdyonurus solus sp. nov., egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S- 4700 scanning electron microscope)." httpUri="https://zenodo.org/record/179176/files/figure.png" pageId="6" pageNumber="59">Fig. 22</figureCitation>
|
||
). The second attachment structure is slightly smaller (diameter 2.7–3.3 µm) and is regularly distributed over the chorion surface (spaced at a distance of 6.7–12.0 µm) (
|
||
<figureCitation id="13316A921E6EFF855003FCEFBA96556E" box="[1110,1198,832,858]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,269,1850,1874]" captionTargetBox="[200,1397,314,1828]" captionTargetId="figure@7.[200,1398,314,1829]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Ecdyonurus solus sp. nov., egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S- 4700 scanning electron microscope)." httpUri="https://zenodo.org/record/179176/files/figure.png" pageId="6" pageNumber="59">Fig. 23</figureCitation>
|
||
). The chorionic surface has many small rounded tubercles (0.7–0.9 µm), spaced at a distance of 0.4–1.3 µm. The chorionic surface and tubercles are covered by very small granules (granular ground matrix) 0.09–0.17 µm in diameter (
|
||
<figureCitation id="13316A921E6EFF8554CAFC17BECF55E6" box="[159,247,952,978]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,269,1850,1874]" captionTargetBox="[200,1397,314,1828]" captionTargetId="figure@7.[200,1398,314,1829]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Ecdyonurus solus sp. nov., egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S- 4700 scanning electron microscope)." httpUri="https://zenodo.org/record/179176/files/figure.png" pageId="6" pageNumber="59">Fig. 24</figureCitation>
|
||
). Five to eight micropyles are visible in the subequatorial area. The sperm guide ovoidal is 11.1–11.8 µm in length and 7.9–9.0 µm in width (micropylar opening situated at the side). The micropylar rim is narrow with tubercles similar to those found on the chorionic surface (
|
||
<figureCitation id="13316A921E6EFF855720FBA7BDF55216" box="[885,973,1032,1058]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,269,1850,1874]" captionTargetBox="[200,1397,314,1828]" captionTargetId="figure@7.[200,1398,314,1829]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Ecdyonurus solus sp. nov., egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S- 4700 scanning electron microscope)." httpUri="https://zenodo.org/record/179176/files/figure.png" pageId="6" pageNumber="59">Fig. 25</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C310259C1E6EFF8B5493FB80BF645036" lastPageId="8" lastPageNumber="61" pageId="6" pageNumber="59" type="etymology">
|
||
<paragraph id="8BB576171E6EFF855493FB80BD285246" blockId="6.[151,1437,152,2016]" pageId="6" pageNumber="59">
|
||
<emphasis id="B97EAA051E6EFF855493FB80BF6B527D" bold="true" box="[198,339,1071,1097]" pageId="6" pageNumber="59">Etymology.</emphasis>
|
||
The species name originates from the Latin noun
|
||
<emphasis id="B97EAA051E6EFF8557F1FB9FBDDF527D" box="[932,999,1072,1097]" italics="true" pageId="6" pageNumber="59">
|
||
<taxonomicName id="4C0A0D941E6EFF8557F1FB9FBDDB527D" box="[932,995,1072,1097]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="species" species="solus">solus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
i.e. single, solitary. It is the only one species of the genus
|
||
<taxonomicName id="4C0A0D941E6EFF8555DCFBF7BC2D5245" box="[393,533,1112,1137]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="59" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97EAA051E6EFF8555DCFBF7BC2D5245" box="[393,533,1112,1137]" italics="true" pageId="6" pageNumber="59">Ecdyonurus</emphasis>
|
||
</taxonomicName>
|
||
living in the Crimea.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6EFF855493FBD0BF3751A4" blockId="6.[151,1437,152,2016]" pageId="6" pageNumber="59">
|
||
<emphasis id="B97EAA051E6EFF855493FBD0BFEF52AD" bold="true" box="[198,471,1151,1177]" pageId="6" pageNumber="59">Specimens examined.</emphasis>
|
||
<typeStatus id="54B1C8B51E6EFF8555B6FB2FBCBA52AE" box="[483,642,1152,1178]" pageId="6" pageNumber="59" type="holotype">HOLOTYPE</typeStatus>
|
||
: male imago,
|
||
<collectingCountry id="F31D36871E6EFF855763FB2FBD8552AE" box="[822,957,1152,1178]" name="Ukraine" pageId="6" pageNumber="59">UKRAINE</collectingCountry>
|
||
, Autonomous Republic of the Crimea, Chorna River, Chornorichens’kyi gap, 8.06.
|
||
<quantity id="4CF2DBF21E6EFF8556F6FB05BD7852F0" box="[675,832,1194,1220]" metricMagnitude="6" metricUnit="m" metricValue="2.0033" pageId="6" pageNumber="59" unit="m" value="2003300.0">
|
||
2003,
|
||
<specimenCount id="9D0CBD9E1E6EFF8556A5FB05BD7852F0" box="[752,832,1194,1220]" pageId="6" pageNumber="59" type="generic">300 m</specimenCount>
|
||
</quantity>
|
||
. a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF8557B0FB05BA4652F0" box="[997,1150,1192,1220]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="33.771946">33o46’19’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF855090FB05BB5A52F0" box="[1221,1378,1192,1220]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.50639">44o30’23’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov.
|
||
<typeStatus id="54B1C8B51E6EFF85550EFB7DBC3A52D8" box="[347,514,1234,1260]" pageId="6" pageNumber="59" type="paratype">PARATYPES</typeStatus>
|
||
:
|
||
<specimenCount id="9D0CBD9E1E6EFF855644FB7DBC4B52D8" box="[529,627,1234,1260]" pageId="6" pageNumber="59" type="larva">3 larvae</specimenCount>
|
||
(two on slides), ibid, Stilia River, 7.07.
|
||
<quantity id="4CF2DBF21E6EFF85500AFB7DBAC552D8" box="[1119,1277,1234,1260]" metricMagnitude="6" metricUnit="m" metricValue="1.98915" pageId="6" pageNumber="59" unit="m" value="1989150.0">
|
||
1989,
|
||
<specimenCount id="9D0CBD9E1E6EFF8550F9FB7DBAC552D8" box="[1196,1277,1234,1260]" pageId="6" pageNumber="59" type="generic">150 m</specimenCount>
|
||
</quantity>
|
||
. a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF8554C2FB51BF08532C" box="[151,304,1275,1304]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="34.047222">34o02’50’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF855523FB51BC2A532C" box="[374,530,1275,1304]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.63639">44o38’11’’ N</geoCoordinate>
|
||
, leg. G.A. Kiseleva;
|
||
<specimenCount id="9D0CBD9E1E6EFF855758FB51BD48532C" box="[781,880,1278,1304]" pageId="6" pageNumber="59" type="larva">2 larvae</specimenCount>
|
||
, ibid, Upper Chorna River, Chornorichens’kyi gap,
|
||
<date id="FFB450D71E6EFF855486FA86BF625377" box="[211,346,1321,1347]" pageId="6" pageNumber="59" value="1998-06-15">
|
||
15.06.
|
||
<quantity id="4CF2DBF21E6EFF855548FA86BF625377" box="[285,346,1321,1347]" metricMagnitude="6" metricUnit="m" metricValue="1.9982999999999997" pageId="6" pageNumber="59" unit="m" value="1998300.0">1998</quantity>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0CBD9E1E6EFF85553FFA86BFF85377" box="[362,448,1321,1347]" pageId="6" pageNumber="59" type="generic">300 m.</specimenCount>
|
||
a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF855631FA86BCC65377" box="[612,766,1317,1347]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="33.771946">33o46’19’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF85571DFA86BDDF5377" box="[840,999,1317,1347]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.50639">44o30’23’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov;
|
||
<specimenCount id="9D0CBD9E1E6EFF8550A6FA86BB605377" box="[1267,1368,1321,1347]" pageId="6" pageNumber="59" type="larva">5 larvae</specimenCount>
|
||
, ibid, Upper Chorna River, Chornorichens’kyi gap, 3.01.
|
||
<quantity id="4CF2DBF21E6EFF8556BEFAFCBDBC5359" box="[747,900,1363,1389]" metricMagnitude="6" metricUnit="m" metricValue="2.0012499999999998" pageId="6" pageNumber="59" unit="m" value="2001250.0">
|
||
2001,
|
||
<specimenCount id="9D0CBD9E1E6EFF855763FAFCBDBC5359" box="[822,900,1363,1389]" pageId="6" pageNumber="59" type="generic">250 m</specimenCount>
|
||
</quantity>
|
||
. a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF855076FAFCBA815359" box="[1059,1209,1360,1389]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="33.771946">33o46’19’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF8550A9FAFCBBAF5359" box="[1276,1431,1360,1389]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.50639">44o30’23’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov;
|
||
<specimenCount id="9D0CBD9E1E6EFF8555DCFAD4BFD253A1" box="[393,490,1403,1429]" pageId="6" pageNumber="59" type="larva">6 larvae</specimenCount>
|
||
, ibid, Upper Chorna River near Chornorichens’kyi gap, 4.01.
|
||
<quantity id="4CF2DBF21E6EFF8550EFFAD4BB6A53A1" box="[1210,1362,1403,1429]" metricMagnitude="6" metricUnit="m" metricValue="2.0012999999999996" pageId="6" pageNumber="59" unit="m" value="2001300.0">
|
||
2001,
|
||
<specimenCount id="9D0CBD9E1E6EFF855151FAD4BB6A53A1" box="[1284,1362,1403,1429]" pageId="6" pageNumber="59" type="generic">300 m</specimenCount>
|
||
</quantity>
|
||
. a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF8554BCFA09BFBD53F4" box="[233,389,1443,1472]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="33.771946">33o46’19’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF855587FA09BC4953F4" box="[466,625,1443,1472]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.50639">44o30’23’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov;
|
||
<specimenCount id="9D0CBD9E1E6EFF8557D6FA09BDD253F4" box="[899,1002,1446,1472]" pageId="6" pageNumber="59" type="larva">9 larvae</specimenCount>
|
||
, ibid, Al’ma River, downstream of “Asport” Boundary,
|
||
<date id="FFB450D71E6EFF8555DCFA7EBC2853DF" box="[393,528,1489,1515]" pageId="6" pageNumber="59" value="2001-06-29">
|
||
29.06.
|
||
<quantity id="4CF2DBF21E6EFF855581FA7EBC2853DF" box="[468,528,1489,1515]" metricMagnitude="6" metricUnit="m" metricValue="2.00145" pageId="6" pageNumber="59" unit="m" value="2001450.0">2001</quantity>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0CBD9E1E6EFF855648FA7EBC5753DF" box="[541,623,1489,1515]" pageId="6" pageNumber="59" type="generic">450 m.</specimenCount>
|
||
a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF85575FFA7EBD9953DF" box="[778,929,1486,1515]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="34.239166">34o14’21’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF8557B0FA7EBA4753DF" box="[997,1151,1486,1515]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.720554">44o43’14’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov;
|
||
<specimenCount id="9D0CBD9E1E6EFF85512AFA7EBEDA5022" pageId="6" pageNumber="59" type="larva">29 larvae</specimenCount>
|
||
, ibid, Al’ma River downstream of “Tar’er” Boundary,
|
||
<date id="FFB450D71E6EFF855723FA53BDC45022" box="[886,1020,1532,1558]" pageId="6" pageNumber="59" value="2001-06-30">
|
||
30.06.
|
||
<quantity id="4CF2DBF21E6EFF855795FA53BDC45022" box="[960,1020,1532,1558]" metricMagnitude="6" metricUnit="m" metricValue="2.0014" pageId="6" pageNumber="59" unit="m" value="2001400.0">2001</quantity>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0CBD9E1E6EFF85505FFA53BA665022" box="[1034,1118,1532,1558]" pageId="6" pageNumber="59" type="generic">400 m.</specimenCount>
|
||
a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF8550ABFA53BBAE5022" box="[1278,1430,1529,1558]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="34.17861">34o10’43’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF855484F988BF565075" box="[209,366,1572,1601]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.719723">44o43’11’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov;
|
||
<specimenCount id="9D0CBD9E1E6EFF85562EF988BCE75075" box="[635,735,1575,1601]" pageId="6" pageNumber="59" type="larva">4 larvae</specimenCount>
|
||
, ibid, Al’ma River downstream of “Sosnovyi” Boundary, 1.07.2001, Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF85553BF9FDBC305058" box="[366,520,1615,1644]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="34.134445">34o08’04’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF855604F9FDBCD65058" box="[593,750,1615,1644]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.759167">44o45’33’’ N</geoCoordinate>
|
||
,
|
||
<specimenCount id="9D0CBD9E1E6EFF8556ABF9FDBD6B5058" box="[766,851,1618,1644]" pageId="6" pageNumber="59" type="generic">
|
||
<quantity id="4CF2DBF21E6EFF8556ABF9FDBD765058" box="[766,846,1618,1644]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="59" unit="m" value="350.0">350 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
a.s.l., leg. G. A. Prokopov; 1 larva, ibid, Al’ma River near Partyzans’ke Reservoir, 1.07.
|
||
<quantity id="4CF2DBF21E6EFF855621F9D2BD3650A3" box="[628,782,1661,1687]" metricMagnitude="6" metricUnit="m" metricValue="2.0012999999999996" pageId="6" pageNumber="59" unit="m" value="2001300.0">
|
||
2001,
|
||
<specimenCount id="9D0CBD9E1E6EFF855695F9D2BD3650A3" box="[704,782,1661,1687]" pageId="6" pageNumber="59" type="generic">300 m</specimenCount>
|
||
</quantity>
|
||
. a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF8557FAF9D2BA7F50A3" box="[943,1095,1658,1687]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="34.095276">34o05’43’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF8550DEF9D2BB1E50A3" box="[1163,1318,1658,1687]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.791668">44o47’30’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov; 1 larva, ibid, Upper Chorna River, Chornorichens’kyi gap,
|
||
<date id="FFB450D71E6EFF85505FF90ABAA9508B" box="[1034,1169,1701,1727]" pageId="6" pageNumber="59" value="2001-08-16">
|
||
16.08.
|
||
<quantity id="4CF2DBF21E6EFF855006F90ABAA9508B" box="[1107,1169,1701,1727]" metricMagnitude="6" metricUnit="m" metricValue="2.0012999999999996" pageId="6" pageNumber="59" unit="m" value="2001300.0">2001</quantity>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0CBD9E1E6EFF8550F1F90ABAC5508B" box="[1188,1277,1701,1727]" pageId="6" pageNumber="59" type="generic">300 m.</specimenCount>
|
||
a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF8554C2F97FBF1750DE" box="[151,303,1741,1770]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="33.771946">33o46’19’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF855524F97FBC3450DE" box="[369,524,1741,1770]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.50639">44o30’23’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov;
|
||
<specimenCount id="9D0CBD9E1E6EFF855759F97FBD5450DE" box="[780,876,1744,1770]" pageId="6" pageNumber="59" type="larva">6 larvae</specimenCount>
|
||
, ibid, Chorna River near Chornorichyns’ke village,
|
||
<date id="FFB450D71E6EFF855482F954BF655121" box="[215,349,1787,1813]" pageId="6" pageNumber="59" value="2001-08-17">
|
||
17.08.
|
||
<quantity id="4CF2DBF21E6EFF85554AF954BF655121" box="[287,349,1787,1813]" metricMagnitude="6" metricUnit="m" metricValue="2.0012" pageId="6" pageNumber="59" unit="m" value="2001200.0">2001</quantity>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D0CBD9E1E6EFF85553FF954BF845121" box="[362,444,1787,1813]" pageId="6" pageNumber="59" type="generic">200 m.</specimenCount>
|
||
a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF855602F954BCD65121" box="[599,750,1784,1813]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="33.689724">33o41’23’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF855764F954BDF45121" box="[817,972,1784,1813]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.536667">44o32’12’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov;
|
||
<specimenCount id="9D0CBD9E1E6EFF85509EF954BB235121" box="[1227,1307,1787,1813]" pageId="6" pageNumber="59" type="male">1 male</specimenCount>
|
||
subimago,
|
||
<specimenCount id="9D0CBD9E1E6EFF8554C2F88CBF395109" box="[151,257,1827,1853]" pageId="6" pageNumber="59" type="female">1 female</specimenCount>
|
||
imago,
|
||
<specimenCount id="9D0CBD9E1E6EFF855532F88CBFE95109" box="[359,465,1827,1853]" pageId="6" pageNumber="59" type="female">3 female</specimenCount>
|
||
subimagines,
|
||
<specimenCount id="9D0CBD9E1E6EFF8556D5F88CBCCC5109" box="[640,756,1827,1853]" pageId="6" pageNumber="59" type="larva">25 larvae</specimenCount>
|
||
,
|
||
<collectingCountry id="F31D36871E6EFF855756F88CBD5D5109" box="[771,869,1827,1853]" name="Ukraine" pageId="6" pageNumber="59">Ukraine</collectingCountry>
|
||
, Autonomous Republic of the Crimea, Chorna River, Chornorichens’kyi gap, 8.06.
|
||
<quantity id="4CF2DBF21E6EFF85561FF8E1BCD3515C" box="[586,747,1870,1896]" metricMagnitude="6" metricUnit="m" metricValue="2.0033" pageId="6" pageNumber="59" unit="m" value="2003300.0">
|
||
2003,
|
||
<specimenCount id="9D0CBD9E1E6EFF8556CCF8E1BCD3515C" box="[665,747,1870,1896]" pageId="6" pageNumber="59" type="generic">300 m</specimenCount>
|
||
</quantity>
|
||
. a.s.l., Long.
|
||
<geoCoordinate id="EE3E10D01E6EFF8557C2F8E1BA0B515C" box="[919,1075,1867,1896]" direction="east" orientation="longitude" pageId="6" pageNumber="59" precision="15" value="33.771946">33o46’19’’ E</geoCoordinate>
|
||
, Lat.
|
||
<geoCoordinate id="EE3E10D01E6EFF85502AF8E1BB25515C" box="[1151,1309,1867,1896]" direction="north" orientation="latitude" pageId="6" pageNumber="59" precision="15" value="44.50639">44o30’23’’ N</geoCoordinate>
|
||
, leg. G.A. Prokopov.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E6EFF845493F831BC375736" blockId="6.[151,1437,152,2016]" lastBlockId="7.[151,1436,152,258]" lastPageId="7" lastPageNumber="60" pageId="6" pageNumber="59">
|
||
All specimens were preserved in 75% alcohol. All adults were reared from larvae. The
|
||
<typeStatus id="54B1C8B51E6EFF855098F831BB0C518C" box="[1229,1332,1950,1976]" pageId="6" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
and part of the
|
||
<typeStatus id="54B1C8B51E6EFF8554B0F869BF6D51D4" box="[229,341,1990,2016]" pageId="6" pageNumber="59" type="paratype">paratypes</typeStatus>
|
||
are housed in the collection of the State Museum of Natural History, National Academy of Sciences of
|
||
<collectingCountry id="F31D36871E6FFF845570FF37BFBB5686" box="[293,387,152,178]" name="Ukraine" pageId="7" pageNumber="60">Ukraine</collectingCountry>
|
||
(Lviv,
|
||
<collectingCountry id="F31D36871E6FFF84558CFF37BC055686" box="[473,573,152,178]" name="Ukraine" pageId="7" pageNumber="60">Ukraine</collectingCountry>
|
||
). The other
|
||
<typeStatus id="54B1C8B51E6FFF84569CFF37BD015686" box="[713,825,152,178]" pageId="7" pageNumber="60" type="paratype">paratypes</typeStatus>
|
||
are stored in the Biology Centre of the Academy of Science of the
|
||
<collectingCountry id="F31D36871E6FFF845512FF6FBC3D56EE" box="[327,517,192,218]" name="Czech Republic" pageId="7" pageNumber="60">Czech Republic</collectingCountry>
|
||
, Institute of Entomology (Č eské Budĕjovice,
|
||
<collectingCountry id="F31D36871E6FFF845077FF6FBAD856EE" box="[1058,1248,192,218]" name="Czech Republic" pageId="7" pageNumber="60">Czech Republic</collectingCountry>
|
||
) and in the first and second author’s collections.
|
||
</paragraph>
|
||
<caption id="DF75269F1E6FFF8454C2F895BF1A51DD" httpUri="https://zenodo.org/record/179176/files/figure.png" pageId="7" pageNumber="60" targetBox="[200,1397,314,1828]" targetPageId="7">
|
||
<paragraph id="8BB576171E6FFF8454C2F895BF1A51DD" blockId="7.[151,1436,1850,2025]" pageId="7" pageNumber="60">
|
||
<emphasis id="B97EAA051E6FFF8454C2F895BF665166" bold="true" box="[151,350,1850,1874]" pageId="7" pageNumber="60">FIGURES 20–25.</emphasis>
|
||
<taxonomicName id="4C0A0D941E6FFF845531F894BC1A5166" box="[356,546,1851,1874]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="60" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E6FFF845531F894BC1A5166" box="[356,546,1851,1874]" italics="true" pageId="7" pageNumber="60">Ecdyonurus solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E6FFF84567CF894BC465166" box="[553,638,1851,1874]" pageId="7" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, egg: 20, general outline of the egg, scale bar = 100 µm; 21, egg pole with gather of large KCTs attachment structures, scale bar = 50 µm; 22, detail of large KCTs, scale bar = 10 µm; 23, chorionic surface with KCTs attachment structures, tubercles and small granules, scale bar = 20 µm; 24, tubercles and small granules at high magnification, scale bar = 3 µm; 25, micropyle, scale bar = 10 µm (HITACHI S-4700 scanning electron microscope).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB576171E60FF8B5493FF38BB5D54D6" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">
|
||
<emphasis id="B97EAA051E60FF8B5493FF38BF795685" bold="true" box="[198,321,151,177]" pageId="8" pageNumber="61">Affinities.</emphasis>
|
||
<taxonomicName id="4C0A0D941E60FF8B551CFF37BF9E5685" box="[329,422,152,177]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B551CFF37BF9E5685" box="[329,422,152,177]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B55FBFF37BC315686" box="[430,521,152,178]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the
|
||
<taxonomicName id="4C0A0D941E60FF8B56E8FF37BD035685" box="[701,827,152,177]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="venosus">
|
||
<emphasis id="B97EAA051E60FF8B56E8FF37BD035685" box="[701,827,152,177]" italics="true" pageId="8" pageNumber="61">E. venosus</emphasis>
|
||
</taxonomicName>
|
||
species-group by the presence of following characteristics:
|
||
<emphasis id="B97EAA051E60FF8B5556FF6FBF9556ED" box="[259,429,192,217]" italics="true" pageId="8" pageNumber="61">in male imago</emphasis>
|
||
: (1) apical sclerite of penis subparalel to the body axis, straight, relatively short and blunt; (2) abdominal terga with mainly visible darker pattern;
|
||
<emphasis id="B97EAA051E60FF8B5724FF47BDF65735" box="[881,974,232,257]" italics="true" pageId="8" pageNumber="61">in larva</emphasis>
|
||
: (3) superlinguae of hypopharynx with long fine setae on the anterior margin (up to the lateral tips); (4) labrum with one row of setae ventrally; (5) lateral prolongation of sterna are markedly visible. The new species can be easily distinguished from all other representatives of this species-group by the following features:
|
||
<emphasis id="B97EAA051E60FF8B57C5FECFBA04574D" box="[912,1084,352,377]" italics="true" pageId="8" pageNumber="61">in male imago</emphasis>
|
||
: (1) eyes gray-blackish, with two light gray rings laterally, divided by a small gap; (2) wing hyaline, transparent and unicolorous; (3) venation brown to blackish, lighter basally, without maculation around veins; (4) color of abdominal terga and sterna; (5) styliger plate with two small rounded apically protuberance; (6) penis lobes slightly stretched laterally and convergent toward outer margins, with rounded distal part; (7) peculiarities of the penis structure;
|
||
<emphasis id="B97EAA051E60FF8B51D0FDAFBEED5475" italics="true" pageId="8" pageNumber="61">in larva</emphasis>
|
||
: (8) number of some groups of setae, bristles and hairs on maxillae and labial palps (9) lateral projection of pronotum short, distinctly asymmetrical, with curved towards the body apical part; (10) femora relatively slender and long; (11) outer margin of femora with row of long slender bristles and small pointed spines; (12) coloration of abdominal terga and sterna; (13) coloration of tarsi; (14) denticulation of posterior margins of terga; (15) shape of scales of dorsal surface of femora and (16) shape of gills (especially gills 1, 4 and 7).
|
||
</paragraph>
|
||
<paragraph id="8BB576171E60FF8B5492FD5FBA1255B6" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">
|
||
The penis structure discriminates the new species from the all other European species of the
|
||
<taxonomicName id="4C0A0D941E60FF8B5148FD5FBBA4553D" box="[1309,1436,752,777]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="venosus">
|
||
<emphasis id="B97EAA051E60FF8B5148FD5FBBA4553D" box="[1309,1436,752,777]" italics="true" pageId="8" pageNumber="61">E. venosus</emphasis>
|
||
</taxonomicName>
|
||
species-group (except for
|
||
<taxonomicName id="4C0A0D941E60FF8B5593FCB7BC425505" box="[454,634,792,817]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="ornatipennis">
|
||
<emphasis id="B97EAA051E60FF8B5593FCB7BC425505" box="[454,634,792,817]" italics="true" pageId="8" pageNumber="61">E. ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
, see below). Some aspects of the penis shape of
|
||
<taxonomicName id="4C0A0D941E60FF8B50FBFCB7BB325505" box="[1198,1290,792,817]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B50FBFCB7BB325505" box="[1198,1290,792,817]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B5144FCB7BB555506" box="[1297,1389,792,818]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, are similar to
|
||
<taxonomicName id="4C0A0D941E60FF8B555AFCEFBC37556E" authority="Grandi, 1953" authorityName="Grandi" authorityYear="1953" box="[271,527,832,858]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="ruffii">
|
||
<emphasis id="B97EAA051E60FF8B555AFCEFBF53556D" box="[271,363,832,857]" italics="true" pageId="8" pageNumber="61">E. ruffii</emphasis>
|
||
Grandi, 1953
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9B0BE61E60FF8B564BFCEFBA73556E" author="Bauernfeind" box="[542,1099,832,858]" pageId="8" pageNumber="62" refString="Bauernfeind, E. & Humpesch, U. H. (2001) Eintagsfliegen Zentraluropas (Insecta: Ephemeroptera): Bestimmung und Okologie. Verlag des Naturhistorischen Museums Wien, Wien, 239 pp." type="book" year="2001">Bauernfeind & Humpesch 2001: 139, Fig. 474</bibRefCitation>
|
||
), but the new species can be distinguished by its body coloration and especially by the pattern on the terga.
|
||
</paragraph>
|
||
<paragraph id="8BB576171E60FF8B5492FC3FBB1552DE" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">
|
||
The male imago of
|
||
<taxonomicName id="4C0A0D941E60FF8B55E7FC3FBC28559D" box="[434,528,912,937]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B55E7FC3FBC28559D" box="[434,528,912,937]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B564CFC3FBC4D559E" box="[537,629,912,938]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be easily distinguished from the two Caucasian species, viz.
|
||
<taxonomicName id="4C0A0D941E60FF8B51D6FC3FBF2255E5" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="B97EAA051E60FF8B51D6FC3FBF2255E5" italics="true" pageId="8" pageNumber="61">E. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A0D941E60FF8B5506FC17BC3055E5" box="[339,520,952,977]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="ornatipennis">
|
||
<emphasis id="B97EAA051E60FF8B5506FC17BC3055E5" box="[339,520,952,977]" italics="true" pageId="8" pageNumber="61">E. ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
. By laterally stretched penis lobes the new species is similar in appearance to
|
||
<emphasis id="B97EAA051E60FF8B54C2FC4FBF6E55CD" box="[151,342,992,1017]" italics="true" pageId="8" pageNumber="61">
|
||
<taxonomicName id="4C0A0D941E60FF8B54C2FC4FBF6B55CD" box="[151,339,992,1017]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="ornatipennis">E. ornatipennis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The combination of following features markedly differs
|
||
<taxonomicName id="4C0A0D941E60FF8B505EFC4FBAFA55CD" box="[1035,1218,992,1017]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="ornatipennis">
|
||
<emphasis id="B97EAA051E60FF8B505EFC4FBAFA55CD" box="[1035,1218,992,1017]" italics="true" pageId="8" pageNumber="61">E. ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C0A0D941E60FF8B5145FC4FBB4855CD" box="[1296,1392,992,1017]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B5145FC4FBB4855CD" box="[1296,1392,992,1017]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B512EFC4FBEF55216" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
: (1) unicolorous eyes without any rings, not close to each other; (2) penis lobes relatively square-shape distally; (3) abdominal sterna with brown longitudinal central band (
|
||
<bibRefCitation id="EF9B0BE61E60FF8B579AFB9FBF2C5246" author="Tshernova" pageId="8" pageNumber="62" refString="Tshernova, O. (1938) To the knowledge of mayflies (Ephemeroptera) of Eastern Transcaucasus. Trudy Azerbaidzhanskoi Akademii Nauk, 8 (42), 55 - 64. (in Russian and German)" type="journal article" year="1938">
|
||
Tshernova 1938, 57–58,
|
||
<figureCitation id="13316A921E60FF8B50A9FB9FBB7D527E" box="[1276,1349,1072,1098]" captionStart="FIGURES 4 – 6" captionStartId="3.[151,269,1306,1330]" captionTargetBox="[297,1248,309,1278]" captionTargetId="figure@3.[237,1347,296,1300]" captionTargetPageId="3" captionText="FIGURES 4 – 6. Ecdyonurus solus sp. nov. Figs 4 – 5, male imago, holotype: 4, male genitalia, ventral view; 5, penis lobe, dorsal view. Fig 6, female imago, paratype: tip of abdomen, ventral view." httpUri="https://zenodo.org/record/179173/files/figure.png" pageId="8" pageNumber="61">Fig. 4</figureCitation>
|
||
, Kluge pers. com.
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C0A0D941E60FF8B5572FBF7BFF25245" box="[295,458,1112,1137]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="B97EAA051E60FF8B5572FBF7BFF25245" box="[295,458,1112,1137]" italics="true" pageId="8" pageNumber="61">E. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0A0D941E60FF8B5636FBF7BC875245" box="[611,703,1112,1137]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B5636FBF7BC875245" box="[611,703,1112,1137]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B5690FBF7BD275246" box="[709,799,1112,1138]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by (1) the distinctly darker color of body; (2) the presence one light ring on eyes; (3) the presence of brown longitudinal central band on abdominal sterna; (4) the shape of penis, with large and not stretched laterally lobes; (5) the distinctly convergent towards the tip apical sclerites; (6) the shape of lateral sclerites (
|
||
<bibRefCitation id="EF9B0BE61E60FF8B56D1FB7FBD0F52DE" author="Braasch" box="[644,823,1232,1258]" pageId="8" pageNumber="62" refString="Braasch, D. (1980 b) Neue Ecdyonurus - Arten (Heptageniidae, Ephemeroptera) aus dem Kaukasus und Transkaukasien (Sowjetunion) (3). Entomologische Nachrichten, 7, 103 - 110." type="journal article" year="1980" yearSuffix="b">Braasch 1980b</bibRefCitation>
|
||
, 103–104,
|
||
<figureCitation id="13316A921E60FF8B57EDFB7FBA2F52DE" box="[952,1047,1232,1258]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,269,1136,1160]" captionTargetBox="[280,1325,199,1102]" captionTargetId="figure@2.[236,1351,132,1150]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Ecdyonurus solus sp. nov., male imago, paratype: 1, head, dorsal view; 2, head, lateral view; 3, abdominal segments 4 - 7, lateral view." httpUri="https://zenodo.org/record/179172/files/figure.png" pageId="8" pageNumber="61">Fig. 1, 1</figureCitation>
|
||
a, 2, Kluge pers. com.).
|
||
</paragraph>
|
||
<paragraph id="8BB576171E60FF8B5492FB57BF645036" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="61">
|
||
The larvae of
|
||
<taxonomicName id="4C0A0D941E60FF8B553AFB57BFF55325" box="[367,461,1272,1297]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B553AFB57BFF55325" box="[367,461,1272,1297]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B5583FB57BC0D5326" box="[470,565,1272,1298]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, partly resembles to
|
||
<taxonomicName id="4C0A0D941E60FF8B5778FB57BAFF5326" authority="Burmeister, 1839" authorityName="Burmeister" authorityYear="1839" box="[813,1223,1272,1298]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="aurantiacus">
|
||
<emphasis id="B97EAA051E60FF8B5778FB57BDE35325" box="[813,987,1272,1297]" italics="true" pageId="8" pageNumber="61">E. aurantiacus</emphasis>
|
||
(Burmeister, 1839)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E60FF8B5081FB57BF95530E" authority="Haybach and Thomas, 2002" authorityName="Haybach and Thomas" authorityYear="2002" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="belfiorei">
|
||
<emphasis id="B97EAA051E60FF8B5081FB57BB605325" box="[1236,1368,1272,1297]" italics="true" pageId="8" pageNumber="61">E. belfiorei</emphasis>
|
||
<bibRefCitation id="EF9B0BE61E60FF8B5134FB57BF95530E" author="Haybach" pageId="8" pageNumber="62" refString="Haybach, A. & Thomas, A. G. B. (2002) Ecdyonurus belfiorei nov. sp. from Italy, with a note on E. aurantiacus androsianus Braasch, 1983 (Ephemeroptera: Heptageniidae). Ephemera, 2 (2), 79 - 91." type="journal article" year="2002">Haybach and Thomas, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A0D941E60FF8B55B1FA8FBCC1530E" authority="Curtis, 1834" authorityName="Curtis" authorityYear="1834" box="[484,761,1312,1338]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="dispar">
|
||
<emphasis id="B97EAA051E60FF8B55B1FA8FBC77530D" box="[484,591,1312,1337]" italics="true" pageId="8" pageNumber="61">E. dispar</emphasis>
|
||
(Curtis, 1834)
|
||
</taxonomicName>
|
||
by the structure of the pronotal projection and gills (especially gill 1). However, larval characteristics 8, 11, 14 and 16 easily differs new species from
|
||
<taxonomicName id="4C0A0D941E60FF8B50B3FAE7BBAB5355" box="[1254,1427,1352,1377]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="aurantiacus">
|
||
<emphasis id="B97EAA051E60FF8B50B3FAE7BBAB5355" box="[1254,1427,1352,1377]" italics="true" pageId="8" pageNumber="61">E. aurantiacus</emphasis>
|
||
</taxonomicName>
|
||
; characters 8, 10, 12, 13 and 14 from
|
||
<taxonomicName id="4C0A0D941E60FF8B5603FADFBCFC53BD" box="[598,708,1392,1417]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="dispar">
|
||
<emphasis id="B97EAA051E60FF8B5603FADFBCFC53BD" box="[598,708,1392,1417]" italics="true" pageId="8" pageNumber="61">E. dispar</emphasis>
|
||
</taxonomicName>
|
||
; characters 8, 12 and 16 from
|
||
<taxonomicName id="4C0A0D941E60FF8B506CFADFBA8653BD" box="[1081,1214,1392,1417]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="belfiorei">
|
||
<emphasis id="B97EAA051E60FF8B506CFADFBA8653BD" box="[1081,1214,1392,1417]" italics="true" pageId="8" pageNumber="61">E. belfiorei</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF9B0BE61E60FF8B5155FADFBF055386" author="Haybach" pageId="8" pageNumber="62" refString="Haybach, A. & Thomas, A. G. B. (2002) Ecdyonurus belfiorei nov. sp. from Italy, with a note on E. aurantiacus androsianus Braasch, 1983 (Ephemeroptera: Heptageniidae). Ephemera, 2 (2), 79 - 91." type="journal article" year="2002">Haybach and Thomas 2002</bibRefCitation>
|
||
). The larvae of
|
||
<taxonomicName id="4C0A0D941E60FF8B55A0FA37BC6A5385" box="[501,594,1432,1457]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B55A0FA37BC6A5385" box="[501,594,1432,1457]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B560FFA37BC8D5386" box="[602,693,1432,1458]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be separated from larvae described as
|
||
<taxonomicName id="4C0A0D941E60FF8B50E5FA37BB5D5385" box="[1200,1381,1432,1457]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="ornatipennis">
|
||
<emphasis id="B97EAA051E60FF8B50E5FA37BB5D5385" box="[1200,1381,1432,1457]" italics="true" pageId="8" pageNumber="61">E. ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF9B0BE61E60FF8B54C2FA6FBF7353EE" author="Braasch" box="[151,331,1472,1498]" pageId="8" pageNumber="62" refString="Braasch, D. (1980 a) Neue Ecdyonurus - Arten (Heptageniidae, Ephemeroptera) aus dem Kaukasus und Transkaukasien (Sowjetunion) II. Entomologische Nachrichten, 2, 23 - 28." type="journal article" year="1980" yearSuffix="a">Braasch 1980a</bibRefCitation>
|
||
), by the shape of the pronotum expansion, femoral scales, gills and by the coloration of the abdominal terga.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C310259C1E60FF8A5493F9A0BB6956EE" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61" type="distribution">
|
||
<paragraph id="8BB576171E60FF8A5493F9A0BB6956EE" blockId="8.[151,1437,151,2020]" lastBlockId="9.[151,1436,152,218]" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61">
|
||
<emphasis id="B97EAA051E60FF8B5493F9A0BC38501D" bold="true" box="[198,512,1551,1577]" pageId="8" pageNumber="61">Distribution and biology.</emphasis>
|
||
The larvae of
|
||
<taxonomicName id="4C0A0D941E60FF8B56E5F9BFBD36501D" box="[688,782,1552,1577]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B56E5F9BFBD36501D" box="[688,782,1552,1577]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B5743F9BFBD4A501E" box="[790,882,1552,1578]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are found in metarithral and hyporithral rivers on the northern slopes of the Crimean mountains at altitudes of
|
||
<quantity id="4CF2DBF21E60FF8B57F0F997BA085066" box="[933,1072,1592,1618]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.5" metricValueMin="1.5" pageId="8" pageNumber="61" unit="m" value="300.0" valueMax="450.0" valueMin="150.0">
|
||
150–
|
||
<specimenCount id="9D0CBD9E1E60FF8B578AF997BA085066" box="[991,1072,1592,1618]" pageId="8" pageNumber="61" type="generic">450 m</specimenCount>
|
||
</quantity>
|
||
. a.s.l. In summer, the average water temperature is 12 0C and the water is of hydrogen carbonate
|
||
<typeStatus id="54B1C8B51E60FF8B57EBF9CDBDCC5048" box="[958,1012,1634,1660]" pageId="8" pageNumber="61">type</typeStatus>
|
||
. The larvae inhabit the waters with current velocities of about
|
||
<quantity id="4CF2DBF21E60FF8B5585F925BC385090" box="[464,512,1674,1700]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="61" unit="m" value="1.0">1 m</quantity>
|
||
/s. Width of river bed width is around
|
||
<quantity id="4CF2DBF21E60FF8B57EEF925BA305090" box="[955,1032,1674,1700]" metricMagnitude="0" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="8" pageNumber="61" unit="m" value="4.5" valueMax="6.0" valueMin="3.0">3–6 m</quantity>
|
||
, depth
|
||
<quantity id="4CF2DBF21E60FF8B5009F925BAED5090" box="[1116,1237,1674,1700]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="8" pageNumber="61" unit="m" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 m</quantity>
|
||
, with cobble and gravel bottoms. The banks of beds are partly shaded by
|
||
<taxonomicName id="4C0A0D941E60FF8B5766F91CBA6850F8" authority="Lipsky" authorityName="Lipsky" box="[819,1104,1714,1740]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="61" phylum="Tracheophyta" rank="species" species="orientalis">
|
||
<emphasis id="B97EAA051E60FF8B5766F91CBDCD50F8" box="[819,1013,1715,1740]" italics="true" pageId="8" pageNumber="61">Fagus orientalis</emphasis>
|
||
Lipsky
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E60FF8B5008F91CBBA450F8" authority="Linnaeus" authorityName="Linnaeus" box="[1117,1436,1714,1740]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="61" phylum="Tracheophyta" rank="species" species="betulus">
|
||
<emphasis id="B97EAA051E60FF8B5008F91CBB1050F8" box="[1117,1320,1715,1740]" italics="true" pageId="8" pageNumber="61">Carpinus betulus</emphasis>
|
||
Linnaeus
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A0D941E60FF8B549FF974BC3850C0" authority="Willd." authorityName="Willd." box="[202,512,1754,1780]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="61" phylum="Tracheophyta" rank="species" species="pubescens">
|
||
<emphasis id="B97EAA051E60FF8B549FF974BF9550C0" box="[202,429,1755,1780]" italics="true" pageId="8" pageNumber="61">Quercus pubescens</emphasis>
|
||
Willd.
|
||
</taxonomicName>
|
||
<taxonomicName id="4C0A0D941E60FF8B565DF974BC5E50C0" box="[520,614,1755,1780]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B565DF974BC5E50C0" box="[520,614,1755,1780]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B563AF975BCF350C0" box="[623,715,1754,1780]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a univoltine winter cycle (Uw), with one generation per year (
|
||
<bibRefCitation id="EF9B0BE61E60FF8B5482F8ADBF455128" author="Clifford" box="[215,381,1794,1820]" pageId="8" pageNumber="62" refString="Clifford, H. F. (1982) Life cycles of mayflies (Ephemeroptera), with special reference to voltinism. Quaestiones Entomologicae, 18, 15 - 90." type="journal article" year="1982">Clifford 1982</bibRefCitation>
|
||
). The population overwinters in the nymphal stage and larval growth is very slow. The flying period extends from May to July, with most of the population emerging by the end of June or the beginning of July. Other insects groups found in these rivers with
|
||
<taxonomicName id="4C0A0D941E60FF8B572EF8FCBDE25158" box="[891,986,1875,1900]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="solus" status="sp. nov.">
|
||
<emphasis id="B97EAA051E60FF8B572EF8FCBDE25158" box="[891,986,1875,1900]" italics="true" pageId="8" pageNumber="61">E. solus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D177E1E60FF8B57B1F8FDBA7A5158" box="[996,1090,1874,1900]" pageId="8" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
include: stoneflies
|
||
<taxonomicName id="4C0A0D941E60FF8B517AF8FCBF9451A0" authority="Retzius, 1783" authorityName="Retzius" authorityYear="1783" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="cinerea">
|
||
<emphasis id="B97EAA051E60FF8B517AF8FCBED651A0" italics="true" pageId="8" pageNumber="61">Nemoura cinerea</emphasis>
|
||
(Retzius, 1783)
|
||
</taxonomicName>
|
||
; dragonflies
|
||
<taxonomicName id="4C0A0D941E60FF8B5614F8D4BD8851A0" authority="Selys, 1853" authorityName="Selys" authorityYear="1853" box="[577,944,1914,1940]" class="Insecta" family="Calopterygidae" genus="Calopteryx" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="taurica">
|
||
<emphasis id="B97EAA051E60FF8B5614F8D4BD2751A0" box="[577,799,1915,1940]" italics="true" pageId="8" pageNumber="61">Calopteryx taurica</emphasis>
|
||
Selys, 1853
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E60FF8B57EEF8D4BBA051A0" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[955,1432,1914,1940]" class="Insecta" family="Gomphidae" genus="Gomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="vulgatissima">
|
||
<emphasis id="B97EAA051E60FF8B57EEF8D4BAFE51A0" box="[955,1222,1915,1940]" italics="true" pageId="8" pageNumber="61">Gomphus vulgatissima</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E60FF8B54C2F80CBC945188" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,684,1954,1980]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="forcipata">
|
||
<emphasis id="B97EAA051E60FF8B54C2F80CBFE95188" box="[151,465,1955,1980]" italics="true" pageId="8" pageNumber="61">Onychogomphus forcipata</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
; mayflies
|
||
<taxonomicName id="4C0A0D941E60FF8B5779F80CBB295188" authority="Demoulin, 1973" authorityName="Demoulin" authorityYear="1973" box="[812,1297,1954,1980]" class="Insecta" family="Heptageniidae" genus="Heptagenia" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="samochai">
|
||
<emphasis id="B97EAA051E60FF8B5779F80CBA085188" box="[812,1072,1955,1980]" italics="true" pageId="8" pageNumber="61">Heptagenia samochai</emphasis>
|
||
(Demoulin, 1973)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A0D941E60FF8B5106F80CBCFF51D0" authority="Godunko and Prokopov, 2003" authorityName="Godunko and Prokopov" authorityYear="2003" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="rhodani" subSpecies="tauricus">
|
||
<emphasis id="B97EAA051E60FF8B5106F80CBF6351D0" italics="true" pageId="8" pageNumber="61">Baetis rhodani tauricus</emphasis>
|
||
Godunko and Prokopov, 2003
|
||
</taxonomicName>
|
||
, caddisflies
|
||
<taxonomicName id="4C0A0D941E60FF8B570CF864BAC751D0" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[857,1279,1994,2020]" class="Insecta" family="Limnephilidae" genus="Halesus" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="tessellatus">
|
||
<emphasis id="B97EAA051E60FF8B570CF864BA0151D0" box="[857,1081,1995,2020]" italics="true" pageId="8" pageNumber="61">Halesus tessellatus</emphasis>
|
||
(Rambur, 1842)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E60FF8A515EF864BFB25686" authority="Curtis, 1834" authorityName="Curtis" authorityYear="1834" class="Insecta" family="Limnephilidae" genus="Limnephilus" kingdom="Animalia" lastPageId="9" lastPageNumber="62" order="Trichoptera" pageId="8" pageNumber="61" phylum="Arthropoda" rank="species" species="lunatus">
|
||
<emphasis id="B97EAA051E60FF8A515EF864BED55685" italics="true" lastPageId="9" lastPageNumber="62" pageId="8" pageNumber="61">Limnephilus lunatus</emphasis>
|
||
Curtis, 1834
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E61FF8A55C2FF37BD785686" authority="Martynov, 1909" authorityName="Martynov" authorityYear="1909" box="[407,832,152,178]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="species" species="acuta">
|
||
<emphasis id="B97EAA051E61FF8A55C2FF37BC425685" box="[407,634,152,177]" italics="true" pageId="9" pageNumber="62">Hydropsyche acuta</emphasis>
|
||
Martynov, 1909
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E61FF8A571BFF37BB205686" authority="Enderlein, 1924" authorityName="Enderlein" authorityYear="1924" box="[846,1304,152,178]" class="Insecta" family="Simuliidae" genus="Wilhelmia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="species" species="balcanica">
|
||
<emphasis id="B97EAA051E61FF8A571BFF37BA795685" box="[846,1089,152,177]" italics="true" pageId="9" pageNumber="62">Wilhelmia balcanica</emphasis>
|
||
(Enderlein, 1924)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A0D941E61FF8A517DFF37BF4856EE" authority="Puri, 1925" authorityName="Puri" authorityYear="1925" class="Insecta" family="Simuliidae" genus="Wilhelmia" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="species" species="paraequinum">
|
||
<emphasis id="B97EAA051E61FF8A517DFF37BED756ED" italics="true" pageId="9" pageNumber="62">W. paraequinum</emphasis>
|
||
Puri, 1925
|
||
</taxonomicName>
|
||
and blackflies
|
||
<emphasis id="B97EAA051E61FF8A5677FF6FBD1156ED" box="[546,809,192,217]" italics="true" pageId="9" pageNumber="62">Simuluim acutiphallus</emphasis>
|
||
(Rubtsov, 1956),
|
||
<emphasis id="B97EAA051E61FF8A57A9FF6FBABE56ED" box="[1020,1158,192,217]" italics="true" pageId="9" pageNumber="62">S. ponticum</emphasis>
|
||
(Rubtsov, 1956).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |