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<document id="C462BAFA4D5032E517C88CBD9084CD69" ID="10.11646/zootaxa.4254.4.3SLASH10855" ID-CLB-Dataset="33201" ID-DOI="10.11646/zootaxa.4254.4.3" ID-GBIF-Dataset="d25360d8-7a6b-4441-8ce3-4fd2b89121f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="556045" ID-ZooBank="29A83502-70C0-4A8F-82D5-CD692FC3219E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1492587321329" checkinUser="plazi" docAuthor="Hara, Marcos R., Bragagnolo, Cibele &amp; Pinto-Da-Rocha, Ricardo" docDate="2017" docId="F179007AFF9FFFCA6CC5CF64FE7CF832" docLanguage="en" docName="zootaxa.4254.4.3SLASH10855.pdf" docOrigin="Zootaxa 4254 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thaumatocranaus Roewer 1932" docType="treatment" docVersion="7" lastPageNumber="459" masterDocId="0D407802FF9DFFC86C52CE0CFFD4FF97" masterDocTitle="Revision of Thaumatocranaus Roewer, 1932, with description of two new species (Opiliones: Laniatores: Gonyleptoidea)" masterLastPageNumber="472" masterPageNumber="457" pageNumber="459" updateTime="1698438960372" updateUser="ExternalLinkService">
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<mods:title id="D8BAFE29998E6BFF458269D21C62D0D8">Revision of Thaumatocranaus Roewer, 1932, with description of two new species (Opiliones: Laniatores: Gonyleptoidea)</mods:title>
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<subSubSection id="31CAE2E7FF9FFFCA6CC5CF64FC50FE15" box="[151,900,360,386]" pageId="2" pageNumber="459" type="nomenclature">
<paragraph id="796FB16CFF9FFFCA6CC5CF64FC50FE15" blockId="2.[151,900,360,386]" box="[151,900,360,386]" pageId="2" pageNumber="459">
<heading id="22270600FF9FFFCA6CC5CF64FC50FE15" bold="true" box="[151,900,360,386]" fontSize="11" level="1" pageId="2" pageNumber="459" reason="1">
<emphasis id="4BA46D7EFF9FFFCA6CC5CF64FC50FE15" bold="true" box="[151,900,360,386]" pageId="2" pageNumber="459">
<taxonomicName id="BED0CAEFFF9FFFCA6CC5CF64FDF0FE15" authority="Roewer, 1932" authorityName="Roewer" authorityYear="1932" box="[151,548,360,386]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="4BA46D7EFF9FFFCA6CC5CF64FEA2FE15" bold="true" box="[151,374,360,386]" italics="true" pageId="2" pageNumber="459">Thaumatocranaus</emphasis>
<bibRefCitation id="1D41CC9DFF9FFFCA6D2FCF64FDF0FE15" author="Roewer" box="[381,548,360,386]" pageId="2" pageNumber="459" refString="Roewer, C. F. (1932) Weitere Weberknechte VII (7. Erganzung der Weberknechte der Erde, 1923). Archiv fur Naturgeschichte, 1 (2), 275 - 350." type="journal article" year="1932">Roewer, 1932</bibRefCitation>
</taxonomicName>
new subfamilial assignment
</emphasis>
</heading>
</paragraph>
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<subSubSection id="31CAE2E7FF9FFFCA6CC5CFA2FDDBFE74" pageId="2" pageNumber="459" type="reference_group">
<paragraph id="796FB16CFF9FFFCA6CC5CFA2FDDBFE74" blockId="2.[151,1436,430,483]" pageId="2" pageNumber="459">
<treatmentCitationGroup id="59C09642FF9FFFCA6CC5CFA2FDDBFE74" pageId="2" pageNumber="459">
<taxonomicName id="BED0CAEFFF9FFFCA6CC5CFA2FDCEFE53" authority="Roewer, 1932: 277" authorityName="Roewer" authorityPageNumber="277" authorityYear="1932" box="[151,538,430,452]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="4BA46D7EFF9FFFCA6CC5CFA2FE9BFE53" box="[151,335,430,452]" italics="true" pageId="2" pageNumber="459">Thaumatocranaus</emphasis>
<treatmentCitation id="F871977DFF9FFFCA6D05CFA3FDCEFE53" author="Roewer" box="[343,538,430,452]" page="277" pageId="2" pageNumber="459" year="1932">
<bibRefCitation id="1D41CC9DFF9FFFCA6D05CFA3FE3DFE53" author="Roewer" box="[343,489,430,452]" pageId="2" pageNumber="459" refString="Roewer, C. F. (1932) Weitere Weberknechte VII (7. Erganzung der Weberknechte der Erde, 1923). Archiv fur Naturgeschichte, 1 (2), 275 - 350." type="journal article" year="1932">Roewer, 1932</bibRefCitation>
: 277
</treatmentCitation>
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(key), 307 (desc);
<treatmentCitation id="F871977DFF9FFFCA6EB2CFA2FC1CFE53" author="Mello-Leitao" box="[736,968,430,452]" page="97" pageId="2" pageNumber="459" year="1935">
<bibRefCitation id="1D41CC9DFF9FFFCA6EB2CFA2FC70FE53" author="Mello-Leitao" box="[736,932,430,452]" pageId="2" pageNumber="459" refString="Mello-Leitao, C. F. (1935) Algumas notas sobre Laniatores. Archivos do Museu Nacional, 36, 89 - 116." type="journal article" year="1935">Mello-Leitão 1935</bibRefCitation>
: 97
</treatmentCitation>
(sys);
<treatmentCitation id="F871977DFF9FFFCA6843CFA2FAFEFE53" author="Soares" box="[1041,1322,430,452]" page="586" pageId="2" pageNumber="459" year="1948">
<bibRefCitation id="1D41CC9DFF9FFFCA6843CFA2FB2DFE53" author="Soares" box="[1041,1273,430,452]" pageId="2" pageNumber="459" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1948) Monografia dos generos de opilioes neotropicos I. Arquivos de Zoologia do Estado de Sao Paulo, 5 (9), 553 - 636." type="journal article" year="1948">Soares &amp; Soares 1948</bibRefCitation>
: 586
</treatmentCitation>
(key), 619 (cat, diag);
<treatmentCitation id="F871977DFF9FFFCA6D69CFC2FE07FE74" author="Kury" box="[315,467,461,483]" page="99" pageId="2" pageNumber="459" year="2003">
<bibRefCitation id="1D41CC9DFF9FFFCA6D69CFC2FE64FE74" author="Kury" box="[315,432,461,483]" pageId="2" pageNumber="459" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, Special Volume, Monografico n o 1, 5 - 337." type="journal article" year="2003">Kury, 2003</bibRefCitation>
: 99
</treatmentCitation>
(cat).
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</paragraph>
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<paragraph id="796FB16CFF9FFFCA6CC5CC03FC68FDBF" blockId="2.[151,1437,527,1957]" box="[151,956,527,552]" pageId="2" pageNumber="459">
<emphasis id="4BA46D7EFF9FFFCA6CC5CC03FEE6FDBF" bold="true" box="[151,306,527,552]" pageId="2" pageNumber="459">Type species.</emphasis>
<taxonomicName id="BED0CAEFFF9FFFCA6D6BCC03FCC3FDBF" authority="Roewer, 1932" authorityName="Roewer" authorityYear="1932" box="[313,791,527,552]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF9FFFCA6D6BCC03FDA5FDBF" box="[313,625,527,552]" italics="true" pageId="2" pageNumber="459">Thaumatocranaus mirabilis</emphasis>
<bibRefCitation id="1D41CC9DFF9FFFCA6E2ACC1CFCC3FDBF" author="Roewer" box="[632,791,528,552]" pageId="2" pageNumber="459" refString="Roewer, C. F. (1932) Weitere Weberknechte VII (7. Erganzung der Weberknechte der Erde, 1923). Archiv fur Naturgeschichte, 1 (2), 275 - 350." type="journal article" year="1932">Roewer, 1932</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="796FB16CFF9FFFCA6C95CC38FBD0FDDA" blockId="2.[151,1437,527,1957]" box="[199,1028,564,589]" pageId="2" pageNumber="459">
<emphasis id="4BA46D7EFF9FFFCA6C95CC38FE09FDDA" bold="true" box="[199,477,564,589]" pageId="2" pageNumber="459">Other species included.</emphasis>
<taxonomicName id="BED0CAEFFF9FFFCA6DB6CC39FDAEFDDB" box="[484,634,564,588]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="magnificus" status="sp. nov.">
<emphasis id="4BA46D7EFF9FFFCA6DB6CC39FDAEFDDB" box="[484,634,564,588]" italics="true" pageId="2" pageNumber="459">T. magnificus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5097D005FF9FFFCA6ED3CC38FD0FFDDA" box="[641,731,564,589]" pageId="2" pageNumber="459" rank="species">
<emphasis id="4BA46D7EFF9FFFCA6ED3CC38FD0FFDDA" bold="true" box="[641,731,564,589]" pageId="2" pageNumber="459">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="BED0CAEFFF9FFFCA6F43CC39FC70FDDA" box="[785,932,564,589]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="splendidus" status="sp. nov.">
<emphasis id="4BA46D7EFF9FFFCA6F43CC39FC70FDDA" box="[785,932,564,589]" italics="true" pageId="2" pageNumber="459">T. splendidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5097D005FF9FFFCA6FF9CC38FBD0FDDA" box="[939,1028,564,589]" pageId="2" pageNumber="459" rank="species">
<emphasis id="4BA46D7EFF9FFFCA6FF9CC38FBD0FDDA" bold="true" box="[939,1028,564,589]" pageId="2" pageNumber="459">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="31CAE2E7FF9FFFCA6C95CC5BFB5AFCDF" pageId="2" pageNumber="459" type="diagnosis">
<paragraph id="796FB16CFF9FFFCA6C95CC5BFB5AFCDF" blockId="2.[151,1437,527,1957]" pageId="2" pageNumber="459">
<emphasis id="4BA46D7EFF9FFFCA6C95CC5BFE95FDE7" bold="true" box="[199,321,599,624]" pageId="2" pageNumber="459">Diagnosis.</emphasis>
Ampycinae (see diagnosis in Villarreal M. &amp; Kury, 2012) with a pair of enlarged tubercles on each corner of anterior margin of DS, low ocularium with a pair of enlarged tubercles, lateral margins of DS with external row of tubercles increasing in size posteriorly, becoming slightly large and blunt or large, conical and slightly acuminated. Scutal grooves I and IV conspicuous; scutal grooves IIIII inconspicuous in the middle and shallow on the sides, weakly delimitating three scutal areas, scutal areas III with a paramedian pair of slightly enlarged pointed tubercles, scutal area III with a pair of spiniform apophyses. Only free tergite III armed with a median robust apophysis. Coxa IV with prolateral apical and retrolateral apical apophyses.
</paragraph>
</subSubSection>
<subSubSection id="31CAE2E7FF9FFFCA6C95CD58FE7CF832" pageId="2" pageNumber="459" type="description">
<paragraph id="796FB16CFF9FFFCA6C95CD58FECFFB6F" blockId="2.[151,1437,527,1957]" pageId="2" pageNumber="459">
<emphasis id="4BA46D7EFF9FFFCA6C95CD58FEA0FCFA" bold="true" box="[199,372,852,877]" pageId="2" pageNumber="459">Redescription.</emphasis>
<emphasis id="4BA46D7EFF9FFFCA6D2ECD58FE6BFCFA" box="[380,447,852,877]" italics="true" pageId="2" pageNumber="459">Male:</emphasis>
Dorsum (
<figureCitation id="E1EBADE9FF9FFFCA6E63CD58FDA1FCFB" box="[561,629,852,877]" captionStart="FIGURE 1" captionStartId="4.[151,250,1857,1879]" captionTargetBox="[207,1331,223,1803]" captionTargetId="figure@4.[199,1365,193,1834]" captionTargetPageId="4" captionText="FIGURE 1. Thaumatocranaus mirabilis Roewer. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femur tarsus prolateral view. Abbreviations: A 1 3, scutal areas I III; Fh, frontal hump; Ft 1 3, free tergites I III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: A C: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm." httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="2" pageNumber="459">Figs 1</figureCitation>
A, CE, 2E, 3AE, 5AE): Anterior margin of carapace with a weakly marked frontal hump, a row of tubercles on each side and 1 pair of enlarged tubercles on each corner (internal largest). Ocularium low, with tubercles sparsely distributed and 1 paramedian pair of enlarged tubercles. Dorsal scutum with well-marked lateral longitudinal grooves and scutal grooves I and IV; scutal grooves IIIII inconspicuous in the middle and shallow on the sides, weakly delimitating three scutal areas. Scutal areas III with a paramedian pair of slightly enlarged pointed tubercles; scutal area III with a paramedian pair of spiniform apophyses. Body shape
<typeStatus id="A66B0FCEFF9FFFCA6DF7CA21FE01FBD2" box="[421,469,1069,1093]" pageId="2" pageNumber="459">type</typeStatus>
gamma. Lateral margin of dorsal scutum with an external row of tubercles increasing in size posteriorly, becoming slightly large and blunt (
<taxonomicName id="BED0CAEFFF9FFFCA6EA2CA5DFCA4FBFF" box="[752,880,1103,1128]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF9FFFCA6EA2CA5DFCA4FBFF" box="[752,880,1103,1128]" italics="true" pageId="2" pageNumber="459">T. mirabilis</emphasis>
</taxonomicName>
) or large, conical and slightly acuminated, and an internal irregular row of tubercles. Only one ozopore on each side. Posterior margin of dorsal scutum and free tergites IIII each with a row of tubercles; free tergites III unarmed; free tergite III with a robust bifid (
<taxonomicName id="BED0CAEFFF9FFFCA69D5CA95FF28FB42" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF9FFFCA69D5CA95FF28FB42" italics="true" pageId="2" pageNumber="459">T. mirabilis</emphasis>
</taxonomicName>
) or unbranched median apophysis and few sparsely distributed tubercles. Anal operculum unarmed, tuberculate.
</paragraph>
<paragraph id="796FB16CFF9FFFCA6C95CB08FA44FA8A" blockId="2.[151,1437,527,1957]" box="[199,1424,1284,1309]" pageId="2" pageNumber="459">Chelicerae: Segment I with well-marked bulla; both fingers toothed (triangle- to laminar-shaped projections).</paragraph>
<paragraph id="796FB16CFF9FFFCA6C95CB2BFBB5FA1F" blockId="2.[151,1437,527,1957]" pageId="2" pageNumber="459">
Pedipalps (
<figureCitation id="E1EBADE9FF9FFFCA6D1ACB2BFE44FAD7" box="[328,400,1319,1344]" captionStart="FIGURE 1" captionStartId="4.[151,250,1857,1879]" captionTargetBox="[207,1331,223,1803]" captionTargetId="figure@4.[199,1365,193,1834]" captionTargetPageId="4" captionText="FIGURE 1. Thaumatocranaus mirabilis Roewer. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femur tarsus prolateral view. Abbreviations: A 1 3, scutal areas I III; Fh, frontal hump; Ft 1 3, free tergites I III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: A C: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm." httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="2" pageNumber="459">Figs 1</figureCitation>
F, G, 3F, G, 5F, G): Trochanter inflated, ventral face with a pair of enlarged setiferous tubercles. Femur unarmed, with sparse tubercles and 1 ventrobasal slightly enlarged tubercle, without prolateral subapical seta. Tibia and tarsus dorso-lateral face with few tubercles, variable setation.
</paragraph>
<paragraph id="796FB16CFF9FFFCA6C95CB99FA98F93F" blockId="2.[151,1437,527,1957]" pageId="2" pageNumber="459">
Legs (
<figureCitation id="E1EBADE9FF9FFFCA6D42CB98FE8EFA3B" box="[272,346,1428,1453]" captionStart="FIGURE 2" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[214,1391,186,1893]" captionTargetId="figure@5.[176,1411,167,1906]" captionTargetPageId="5" captionText="FIGURE 2. Thaumatocranaus mirabilis Roewer. Male (holotype): A, right leg I, apex of metatarsus tarsus I, prolateral view; B, right femur III, prolateral view; C, right trochanter IV, dorsal view; D, apex of right femur IV, dorsal view; E, anal operculum, posterior view; F, right trochanter tarsus IV, dorsal view; G, ditto, ventral view. Abbreviations: Pro, prolateral; Ret, retrolateral. A, B at same scale; F, G at same scale. Scale bars: A, B, F, G, 1 mm; C E, 0.5 mm." httpUri="https://zenodo.org/record/556047/files/figure.png" pageId="2" pageNumber="459">Figs 2</figureCitation>
AD, F, G, 4, 6): Coxa I with 1 prolateral, 1 retrolateral apophyses; coxa II ditto, retrolateral apophysis apex fused to prolateral apophysis of coxa III; coxa III with 1 prolateral, 1 retrolateral apophyses; coxa IV tuberculate, with 1 prolateral and 1 retrolateral apical, conical apophyses. Coxa IV sides slightly visible in dorsal view, only apical region most visible (in dorsal view). Trochanters IIV tuberculate; trochanter IV unarmed prolaterally (except
<taxonomicName id="BED0CAEFFF9FFFCA6D2EC829FE2BF9AA" box="[380,511,1572,1597]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="459" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF9FFFCA6D2EC829FE2BF9AA" box="[380,511,1572,1597]" italics="true" pageId="2" pageNumber="459">T. mirabilis</emphasis>
</taxonomicName>
). Femoratibiae IIV with tubercles roughly organized in 6 longitudinal rows (a pro- and a retrodorsal rows, a pro- and a retrolateral rows and a pro- and a retroventral rows), tubercles on femora tibiae III smaller than those on IIIIV. Femora IIII roughly straight, femur IV straight to slightly curved inwards (in dorsal view). Patellaemetatarsi IIII minute tuberculate. Metatarsi IIV unarmed. Tarsal claws smooth.
</paragraph>
<paragraph id="796FB16CFF9FFFCA6C95C8B8FE7DF8AF" blockId="2.[151,1437,527,1957]" pageId="2" pageNumber="459">
Penis (
<figureCitation id="E1EBADE9FF9FFFCA6D46C8B8FE8EF95B" box="[276,346,1716,1741]" captionStart="FIGURE 7" captionStartId="12.[151,250,1877,1899]" captionTargetBox="[260,1351,245,1808]" captionTargetId="figure@12.[218,1385,200,1840]" captionTargetPageId="12" captionText="FIGURE 7. Distal part of Thaumatocranaus spp. penises: A, T. mirabilis, dorsal view, B, ditto, right lateral view, C, ditto, ventral view; D, T. magnificus sp. nov., dorsal view, E, ditto, left lateral view; F, T. splendidus sp. nov., dorsal view, G, ditto, left lateral view. Abbreviations: Gl, glans; MS A E, ventral plate macrosetae A E; St, glans stylus; Vpl, ventral plate. A C at same scale. D, E at same scale. F, G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/556052/files/figure.png" pageId="2" pageNumber="459">Fig. 7</figureCitation>
): Glans with sub cylindrical stylus, without dorsal or ventral process. Ventral plate conspicuous, subhexagonal shaped. Distal margin of ventral plate with a median narrow cleft, and a dorso mesal projection on its mesal apical corner (
<figureCitation id="E1EBADE9FF9FFFCA6DD4C8F0FE1FF883" box="[390,459,1788,1813]" captionStart="FIGURE 7" captionStartId="12.[151,250,1877,1899]" captionTargetBox="[260,1351,245,1808]" captionTargetId="figure@12.[218,1385,200,1840]" captionTargetPageId="12" captionText="FIGURE 7. Distal part of Thaumatocranaus spp. penises: A, T. mirabilis, dorsal view, B, ditto, right lateral view, C, ditto, ventral view; D, T. magnificus sp. nov., dorsal view, E, ditto, left lateral view; F, T. splendidus sp. nov., dorsal view, G, ditto, left lateral view. Abbreviations: Gl, glans; MS A E, ventral plate macrosetae A E; St, glans stylus; Vpl, ventral plate. A C at same scale. D, E at same scale. F, G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/556052/files/figure.png" pageId="2" pageNumber="459">Fig. 7</figureCitation>
C, D, F), with MS C, dorsal subdistalmedian MS D, ventral subdistalmedian small MS E, MS A and MS B.
</paragraph>
<paragraph id="796FB16CFF9FFFCA6C95C948FE4AF8CA" blockId="2.[151,1437,527,1957]" box="[199,414,1860,1885]" pageId="2" pageNumber="459">Female: Unknown.</paragraph>
<paragraph id="796FB16CFF9FFFCA6C95C96BFE7CF832" blockId="2.[151,1437,527,1957]" pageId="2" pageNumber="459">
<materialsCitation id="C9B8BB31FF9FFFCA6C95C96BFE7CF832" ID-GBIF-Occurrence="1457229156" county="Southeastern Colombia" location="Geographical" municipality="Central Ecuador" pageId="2" pageNumber="459" specimenCount="1" typeStatus="holotype">
<emphasis id="4BA46D7EFF9FFFCA6C95C96BFDD0F817" bold="true" box="[199,516,1895,1920]" pageId="2" pageNumber="459">
<location id="7C0FE7B7FF9FFFCA6C95C96BFEBDF817" LSID="urn:lsid:plazi:treatment:F179007AFF9FFFCA6CC5CF64FE7CF832:7C0FE7B7FF9FFFCA6C95C96BFEBDF817" box="[199,361,1895,1920]" county="Southeastern Colombia" municipality="Central Ecuador" name="Geographical" pageId="2" pageNumber="459">Geographical</location>
distribution.
</emphasis>
<collectingMunicipality id="990B2B16FF9FFFCA6E59C96BFD15F817" box="[523,705,1895,1920]" pageId="2" pageNumber="459">Central Ecuador</collectingMunicipality>
and
<collectingCounty id="900EC9E0FF9FFFCA6EAAC96BFC29F817" box="[760,1021,1895,1920]" pageId="2" pageNumber="459">Southeastern Colombia</collectingCounty>
(known only from the
<typeStatus id="A66B0FCEFF9FFFCA6952C965FAE4F816" box="[1280,1328,1897,1921]" pageId="2" pageNumber="459">type</typeStatus>
localities of the included species).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>