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<document id="56A170E85C3A09CE68EFC5EA8B70C936" ID-CLB-Dataset="29262" ID-DOI="10.11646/zootaxa.4496.1.22" ID-GBIF-Dataset="ae180d87-c93d-4839-bb4d-5d504167601b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446803" ID-ZooBank="8DC29B3B-EF7C-4002-B7B0-EE67D84AE40F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747275108" checkinUser="plazi" docAuthor="Jiang, Ji-Bao, Dong, Yan, Yuan, Zhu &amp; Qiu, Jiang-Ping" docDate="2018" docId="03FCEC28DB6CFFCFFF1FFD8666B97A15" docLanguage="en" docName="zootaxa.4496.1.22.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas marsupiformis Jiang &amp; Dong &amp; Yuan &amp; Qiu 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="275" masterDocId="FFC59450DB69FFC8FF88FFEA67097828" masterDocTitle="Three new earthworm species of the tokioensis - group in the genus Amynthas (Oligochaeta: Megascolecidae) from Guangxi Province, China" masterLastPageNumber="278" masterPageNumber="269" pageNumber="274" updateTime="1698566712894" updateUser="ExternalLinkService">
<mods:mods id="D2A047CE0BAF3E4BE82251E87B1811AB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="324FB4ADC76E703C44733F55A97AB559">
<mods:title id="7A70F3A21E0B3505CF0978C8BDF8D180">Three new earthworm species of the tokioensis - group in the genus Amynthas (Oligochaeta: Megascolecidae) from Guangxi Province, China</mods:title>
</mods:titleInfo>
<mods:name id="1CD866B9C591B6E70CFFF00E8BB5B40D" type="personal">
<mods:role id="206B712197350628829E0C6C7EFDE55E">
<mods:roleTerm id="42E3F19D3438CECEB1AD4BE336650E12">Author</mods:roleTerm>
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<mods:namePart id="AED2574ACD78C7E8E1F662AF9ED22AF7">Jiang, Ji-Bao</mods:namePart>
</mods:name>
<mods:name id="0391CE1B7C5D1D9162328A90F2B2F15F" type="personal">
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<mods:namePart id="992CDF62E1D927D26EF55E2914F530B3">Dong, Yan</mods:namePart>
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<mods:name id="7319B35C5968AFC4656F9790821535AE" type="personal">
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<mods:namePart id="9FEBF4945D0B60E8A66E9B889F3D9AEA">Yuan, Zhu</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="E0E88C0B171BC082E10F7A2F96CA9DDB">Qiu, Jiang-Ping</mods:namePart>
</mods:name>
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<mods:title id="420CFBDA37FCC55BD11A01144A516B21">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4E35D7A16D4C9F71725C333862D99015">
<mods:date id="4930E42EDA7EBDA0C242F5654313C8C4">2018</mods:date>
<mods:detail id="322B4FB6542BCE2D1E68B9AB3CA93F27" type="pubDate">
<mods:number id="1F00A858164410C320ADC044CE7CC404">2018-10-05</mods:number>
</mods:detail>
<mods:detail id="E494FE80FA3D7B3036D10FAEEB06DC0D" type="volume">
<mods:number id="26B647CB18A777B1315547F65A116FE9">4496</mods:number>
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<mods:number id="13D40A88FAE6D8D5FD0F9B6203451CCE">1</mods:number>
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<mods:classification id="64A68F7549EC5BA20EF3FD2C4E4AD11A">journal article</mods:classification>
<mods:identifier id="0C446456C14D3C0C1CF43D2FF9F54245" type="CLB-Dataset">29262</mods:identifier>
<mods:identifier id="BD4C64007B5D5E46E370CD497372FD78" type="DOI">10.11646/zootaxa.4496.1.22</mods:identifier>
<mods:identifier id="D5485F124BC0FE42387481EC111E3C76" type="GBIF-Dataset">ae180d87-c93d-4839-bb4d-5d504167601b</mods:identifier>
<mods:identifier id="8254D09C902ED3695FC9F3F98A2EE928" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6E5E73E50D56E1468ADC3C7A8E72F0BA" type="Zenodo-Dep">1446803</mods:identifier>
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</mods:mods>
<treatment id="03FCEC28DB6CFFCFFF1FFD8666B97A15" ID-DOI="http://doi.org/10.5281/zenodo.5957937" ID-GBIF-Taxon="148639856" ID-Zenodo-Dep="5957937" LSID="urn:lsid:plazi:treatment:03FCEC28DB6CFFCFFF1FFD8666B97A15" httpUri="http://treatment.plazi.org/id/03FCEC28DB6CFFCFFF1FFD8666B97A15" lastPageId="7" lastPageNumber="275" pageId="5" pageNumber="274">
<subSubSection id="C34F0EB5DB6CFFCDFF1FFD86660E7A8F" pageId="5" pageNumber="274" type="nomenclature">
<paragraph id="8BEA5D3EDB6CFFCDFF1FFD8665E37AAE" blockId="5.[151,746,620,679]" box="[151,746,620,646]" pageId="5" pageNumber="274">
<heading id="D0A2EA52DB6CFFCDFF1FFD8665E37AAE" bold="true" box="[151,746,620,646]" fontSize="11" level="1" pageId="5" pageNumber="274" reason="1">
<emphasis id="B921812CDB6CFFCDFF1FFD8665E37AAE" bold="true" box="[151,746,620,646]" pageId="5" pageNumber="274">
<taxonomicName id="4C5526BDDB6CFFCDFF1FFD8665897AAE" authority="Jiang &amp; Yuan" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[151,640,620,646]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="274" phylum="Annelida" rank="species" species="marsupiformis" status="sp. nov.">
<emphasis id="B921812CDB6CFFCDFF1FFD8666C07AAE" bold="true" box="[151,457,620,646]" italics="true" pageId="5" pageNumber="274">Amynthas marsupiformis</emphasis>
Jiang &amp; Yuan
</taxonomicName>
,
<taxonomicNameLabel id="A2123C57DB6CFFCDFD05FD8665E37AAE" box="[653,746,620,646]" pageId="5" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA5D3EDB6CFFCDFF1FFD64660E7A8F" blockId="5.[151,746,620,679]" box="[151,263,654,679]" pageId="5" pageNumber="274">
(
<figureCitation id="136E41BBDB6CFFCDFF16FD6467F77A8F" box="[158,254,654,679]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[163,1424,721,1840]" captionTargetId="figure@6.[151,1436,700,1840]" captionTargetPageId="6" captionText="FIGURE 3. Amynthas marsupiformis Jiang &amp; Yuan, sp. nov. A. Region of spermathecal pores, ventral view. B. Region of male pores, ventral view. C. Spermatheca. D. Intestinal caeca (ventral margin is on the left)." httpUri="https://zenodo.org/record/1446809/files/figure.png" pageId="5" pageNumber="274">Figure 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F0EB5DB6CFFCDFF1FFD3D66077BEF" pageId="5" pageNumber="274" type="materials_examined">
<paragraph id="8BEA5D3EDB6CFFCDFF1FFD3D655B7B1F" blockId="5.[151,1437,726,1255]" pageId="5" pageNumber="274">
<materialsCitation id="3B3D5763DB6CFFCDFF1FFD3D655B7B1F" ID-GBIF-Occurrence="1914265367" collectingDate="15 th May" collectorName="Qiu &amp; Jiang &amp; Zhang &amp; Y. Dong" country="China" county="Clitellate" elevation="907" latitude="23.98077" location="Dayao Mountain" longLatPrecision="1" longitude="110.11883" municipality="Dayao Mountain Nature Reserve" pageId="5" pageNumber="274" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B921812CDB6CFFCDFF1FFD3D66027AC7" bold="true" box="[151,267,727,751]" pageId="5" pageNumber="274">
<typeStatus id="54EEE39CDB6CFFCDFF1FFD3D660F7AC7" box="[151,262,727,751]" pageId="5" pageNumber="274" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="628B25B2DB6CFFCDFE99FD3C66717AC7" box="[273,376,726,751]" pageId="5" pageNumber="274">Clitellate</collectingCounty>
specimen (C-GX201314-04A):
<collectingCountry id="F3421DAEDB6CFFCDFD69FD3C642F7AC7" box="[737,806,726,751]" name="China" pageId="5" pageNumber="274">China</collectingCountry>
,
<collectingRegion id="499193DCDB6CFFCDFCB9FD3C649B7AC7" box="[817,914,726,751]" country="China" name="Guangxi" pageId="5" pageNumber="274">Guangxi</collectingRegion>
Province,
<collectingMunicipality id="6B8EC744DB6CFFCDFB81FD3D62717AC6" box="[1033,1400,726,751]" pageId="5" pageNumber="274">
<location id="8E8A0BE5DB6CFFCDFB81FD3D63CC7AC7" LSID="urn:lsid:plazi:treatment:03FCEC28DB6CFFCFFF1FFD8666B97A15:8E8A0BE5DB6CFFCDFB81FD3D63CC7AC7" box="[1033,1221,726,751]" country="China" county="Clitellate" latitude="23.98077" longLatPrecision="1" longitude="110.11883" municipality="Dayao Mountain Nature Reserve" name="Dayao Mountain" pageId="5" pageNumber="274" stateProvince="Guangxi">
Dayao
<location id="8E8A0BE5DB6CFFCDFBD0FD3C63CC7AC7" LSID="urn:lsid:plazi:treatment:03FCEC28DB6CFFCFFF1FFD8666B97A15:8E8A0BE5DB6CFFCDFBD0FD3C63CC7AC7" box="[1112,1221,726,751]" country="China" county="Clitellate" latitude="23.98077" longLatPrecision="1" longitude="110.11883" municipality="Dayao Mountain Nature Reserve" name="Mountain Nature Reserve" pageId="5" pageNumber="274" stateProvince="Guangxi">Mountain</location>
</location>
Nature Reserve
</collectingMunicipality>
(
<geoCoordinate id="EE613BF9DB6CFFCDFA0FFD3D66077B3B" direction="north" orientation="latitude" pageId="5" pageNumber="274" precision="1" value="23.98077">N 23.98077°</geoCoordinate>
,
<geoCoordinate id="EE613BF9DB6CFFCDFE91FD1166BF7B3B" box="[281,438,763,787]" direction="east" orientation="longitude" pageId="5" pageNumber="274" precision="1" value="110.11883">E 110.11883°</geoCoordinate>
),
<elevation id="0078BA0DDB6CFFCDFE40FD1165427B3B" box="[456,587,762,787]" metricMagnitude="2" metricUnit="m" metricValue="9.07" pageId="5" pageNumber="274" unit="m" value="907.0">
<quantity id="4CADF0DBDB6CFFCDFE40FD1165067B3A" box="[456,527,763,787]" metricMagnitude="2" metricUnit="m" metricValue="9.07" pageId="5" pageNumber="274" unit="m" value="907.0">907 m</quantity>
a.s.l.
</elevation>
, yellow sandy topsoil under bush besides road,
<collectingDate id="EFAF8216DB6CFFCDFBE9FD1063C77B3B" box="[1121,1230,762,787]" pageId="5" pageNumber="274">
<date id="FFEB7BFEDB6CFFCDFBE9FD1063C77B3B" box="[1121,1230,762,787]" pageId="5" pageNumber="274">15th May</date>
</collectingDate>
, 2013, J. P.
<collectorName id="26A038E8DB6CFFCDFADAFD1062767B3B" box="[1362,1407,762,787]" pageId="5" pageNumber="274">Qiu</collectorName>
, J. B.
<collectorName id="26A038E8DB6CFFCDFF3FFCF467FC7B1F" box="[183,245,798,823]" pageId="5" pageNumber="274">Jiang</collectorName>
, L. L.
<collectorName id="26A038E8DB6CFFCDFEB7FCF4668E7B1F" box="[319,391,798,823]" pageId="5" pageNumber="274">Zhang</collectorName>
and
<collectorName id="26A038E8DB6CFFCDFE35FCF565107B1F" box="[445,537,799,823]" pageId="5" pageNumber="274">Y. Dong</collectorName>
coll.
</materialsCitation>
</paragraph>
<paragraph id="8BEA5D3EDB6CFFCDFF4FFCA966077BEF" blockId="5.[151,1437,726,1255]" pageId="5" pageNumber="274">
<materialsCitation id="3B3D5763DB6CFFCDFF4FFCA966077BEF" ID-GBIF-Occurrence="1914265364" collectingDate="16 th May" collectorName="Qiu &amp; Jiang &amp; Zhang &amp; Y. Dong" country="China" county="Dayao Mountain Nature Reserve" elevation="1196" latitude="24.115622" location="Mountain Nature Reserve" longLatPrecision="1" longitude="110.21279" municipality="Dayao Mountain" pageId="5" pageNumber="274" specimenCount="2" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="B921812CDB6CFFCDFF4FFCA9664F7B73" bold="true" box="[199,326,835,859]" pageId="5" pageNumber="274">
<typeStatus id="54EEE39CDB6CFFCDFF4FFCA966487B73" box="[199,321,835,859]" pageId="5" pageNumber="274" type="paratype">Paratypes</typeStatus>
.
</emphasis>
3 clitellate specimens (C-GX201314-04B), same data as for
<typeStatus id="54EEE39CDB6CFFCDFB88FCA8636F7B73" box="[1024,1126,834,859]" pageId="5" pageNumber="274" type="holotype">holotype</typeStatus>
. 2 clitellate (C-GX201318- 02):
<collectingCountry id="F3421DAEDB6CFFCDFF47FC8C661C7B57" box="[207,277,870,895]" name="China" pageId="5" pageNumber="274">China</collectingCountry>
,
<collectingRegion id="499193DCDB6CFFCDFEABFC8C668D7B57" box="[291,388,870,895]" country="China" name="Guangxi" pageId="5" pageNumber="274">Guangxi</collectingRegion>
Province,
<collectingCounty id="628B25B2DB6CFFCDFD8CFC8D64897B56" box="[516,896,870,895]" pageId="5" pageNumber="274">
<collectingMunicipality id="6B8EC744DB6CFFCDFD8CFC8D65CC7B57" box="[516,709,870,895]" pageId="5" pageNumber="274">
Dayao
<location id="8E8A0BE5DB6CFFCDFDD0FC8C65CC7B57" LSID="urn:lsid:plazi:treatment:03FCEC28DB6CFFCFFF1FFD8666B97A15:8E8A0BE5DB6CFFCDFDD0FC8C65CC7B57" box="[600,709,870,895]" country="China" county="Dayao Mountain Nature Reserve" latitude="24.115622" longLatPrecision="1" longitude="110.21279" municipality="Dayao Mountain" name="Mountain Nature Reserve" pageId="5" pageNumber="274" stateProvince="Guangxi">Mountain</location>
</collectingMunicipality>
Nature Reserve
</collectingCounty>
(
<geoCoordinate id="EE613BF9DB6CFFCDFC1BFC8D633E7B57" box="[915,1079,870,895]" direction="north" orientation="latitude" pageId="5" pageNumber="274" precision="1" value="24.115622">N 24.115622°</geoCoordinate>
,
<geoCoordinate id="EE613BF9DB6CFFCDFBCFFC8D63E17B57" box="[1095,1256,871,895]" direction="east" orientation="longitude" pageId="5" pageNumber="274" precision="1" value="110.21279">E 110.21279°</geoCoordinate>
),
<elevation id="0078BA0DDB6CFFCDFB77FC8C629E7B57" box="[1279,1431,870,895]" metricMagnitude="3" metricUnit="m" metricValue="1.196" pageId="5" pageNumber="274" unit="m" value="1196.0">
<quantity id="4CADF0DBDB6CFFCDFB77FC8C625E7B56" box="[1279,1367,870,895]" metricMagnitude="3" metricUnit="m" metricValue="1.196" pageId="5" pageNumber="274" unit="m" value="1196.0">1196 m</quantity>
a.s.l.
</elevation>
, black sandy topsoil besides road in
<location id="8E8A0BE5DB6CFFCDFDA4FC6064087B8B" LSID="urn:lsid:plazi:treatment:03FCEC28DB6CFFCFFF1FFD8666B97A15:8E8A0BE5DB6CFFCDFDA4FC6064087B8B" box="[556,769,906,931]" country="China" county="Dayao Mountain Nature Reserve" latitude="24.115622" longLatPrecision="1" longitude="110.21279" municipality="Dayao Mountain" name="Monkey Mountain" pageId="5" pageNumber="274" stateProvince="Guangxi">Monkey Mountain</location>
,
<collectingDate id="EFAF8216DB6CFFCDFC85FC6064757B8B" box="[781,892,906,931]" pageId="5" pageNumber="274">
<date id="FFEB7BFEDB6CFFCDFC85FC6064757B8B" box="[781,892,906,931]" pageId="5" pageNumber="274">16th May</date>
</collectingDate>
, 2013, J. P.
<collectorName id="26A038E8DB6CFFCDFB8DFC60633B7B8B" box="[1029,1074,906,931]" pageId="5" pageNumber="274">Qiu</collectorName>
, J. B.
<collectorName id="26A038E8DB6CFFCDFBF2FC6063B17B8B" box="[1146,1208,906,931]" pageId="5" pageNumber="274">Jiang</collectorName>
, L. L.
<collectorName id="26A038E8DB6CFFCDFA8CFC6062427B8B" box="[1284,1355,906,931]" pageId="5" pageNumber="274">Zhang</collectorName>
and
<collectorName id="26A038E8DB6CFFCDFA0DFC6167DC7BEF" pageId="5" pageNumber="274">Y. Dong</collectorName>
coll.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F0EB5DB6CFFCDFF4FFC3966127C27" pageId="5" pageNumber="274" type="etymology">
<paragraph id="8BEA5D3EDB6CFFCDFF4FFC3966127C27" blockId="5.[151,1437,726,1255]" pageId="5" pageNumber="274">
<emphasis id="B921812CDB6CFFCDFF4FFC3966477BC3" bold="true" box="[199,334,979,1003]" pageId="5" pageNumber="274">Etymology.</emphasis>
The species is named after the characteristics of the spermathecal diverticulum, its diverticulum is long baggy.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EB5DB6CFFCEFF4FFBF0639979A0" lastPageId="6" lastPageNumber="275" pageId="5" pageNumber="274" type="diagnosis">
<paragraph id="8BEA5D3EDB6CFFCDFF4FFBF066557CCF" blockId="5.[151,1437,726,1255]" pageId="5" pageNumber="274">
<emphasis id="B921812CDB6CFFCDFF4FFBF066487C1B" bold="true" box="[199,321,1050,1075]" pageId="5" pageNumber="274">Diagnosis.</emphasis>
Dimensions
<quantity id="4CADF0DBDB6CFFCDFE5CFBF065457C1A" box="[468,588,1050,1075]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" metricValueMax="7.0" metricValueMin="5.8" pageId="5" pageNumber="274" unit="mm" value="64.0" valueMax="70.0" valueMin="58.0">58–70 mm</quantity>
by 2.4–3.0 mm at clitellum, segments 60–73. Setae numbering 24–28 at III, 34–37 at V, 44–47 at VIII, 35–40 at XX, 36–40 at XXV; 13–16 between spermathecal pores. Spermathecal pores in 6/7–7/8, 0.33 body circumference ventrally apart. Male pores absent. 12 tiny presetal or postsetal genital papillae in parallel rows present on the ventrolateral sides in XVII–XIX. Ampulla ovoid; duct as long as 2.5 ampulla. Diverticulum slightly shorter than main pouch (ampulla and duct), terminal 2/5 dilated into a long marsupial seminal chamber.
</paragraph>
<caption id="DF2A0DB6DB6CFFCDFF1FFAFC67FA7D65" pageId="5" pageNumber="274">
<paragraph id="8BEA5D3EDB6CFFCDFF1FFAFC67FA7D65" blockId="5.[151,1436,1301,1357]" pageId="5" pageNumber="274">
<emphasis id="B921812CDB6CFFCDFF1FFAFC66077D05" bold="true" box="[151,270,1302,1325]" pageId="5" pageNumber="274">TABLE 2.</emphasis>
Comparison of
<taxonomicName id="4C5526BDDB6CFFCDFE42FAFC65817D05" authority="Jiang &amp; Dong &amp; Yuan &amp; Qiu, 2018" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[458,648,1301,1325]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="274" phylum="Annelida" rank="species" species="marsupiformis" status="sp. nov.">
<emphasis id="B921812CDB6CFFCDFE42FAFC65817D05" box="[458,648,1301,1325]" italics="true" pageId="5" pageNumber="274">A. marsupiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C57DB6CFFCDFD1CFAFC65E67D05" box="[660,751,1302,1325]" pageId="5" pageNumber="274" rank="species">
<emphasis id="B921812CDB6CFFCDFD1CFAFC65E67D05" bold="true" box="[660,751,1302,1325]" pageId="5" pageNumber="274">sp. nov.</emphasis>
</taxonomicNameLabel>
and two species with the same arrangement of post-clitellar papillae.
</paragraph>
</caption>
<paragraph id="8BEA5D3EDB6CFFCDFF17FA83626F7FE8" pageId="5" pageNumber="274">
<table id="F955AF9EDB6C0037FF17FA8362857FF7" box="[159,1420,1385,2015]" gridcols="4" gridrows="12" pageId="5" pageNumber="274">
<tr id="35655F7CDB6C0037FF17FA8362857D57" box="[159,1420,1385,1407]" gridrow="0" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17FA8366737D57" box="[159,378,1385,1407]" gridcol="0" gridrow="0" pageId="5" pageNumber="274">Characteristics</th>
<th id="76B43600DB6C0037FE02FA8365FF7D57" box="[394,758,1385,1407]" gridcol="1" gridrow="0" pageId="5" pageNumber="274">
<taxonomicName id="4C5526BDDB6CFFCDFE02FA8065337D57" authority="Jiang &amp; Dong &amp; Yuan &amp; Qiu, 2018" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[394,570,1385,1407]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="274" phylum="Annelida" rank="species" species="marsupiformis" status="sp. nov.">
<emphasis id="B921812CDB6CFFCDFE02FA8065337D57" box="[394,570,1385,1407]" italics="true" pageId="5" pageNumber="274">A. marsupiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C57DB6CFFCDFDC8FA8365997D56" box="[576,656,1385,1406]" pageId="5" pageNumber="274" rank="species">
<emphasis id="B921812CDB6CFFCDFDC8FA8365997D56" bold="true" box="[576,656,1385,1406]" pageId="5" pageNumber="274">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<th id="76B43600DB6C0037FC85FA8363517D57" box="[781,1112,1385,1407]" gridcol="2" gridrow="0" pageId="5" pageNumber="274">
<taxonomicName id="4C5526BDDB6CFFCDFC85FA8063167D57" authority="(Chen, 1946)" authorityName="Chen" authorityYear="1946" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[781,1055,1385,1407]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="274" phylum="Annelida" rank="species" species="moniliatus">
<emphasis id="B921812CDB6CFFCDFC85FA80649D7D57" box="[781,916,1385,1407]" italics="true" pageId="5" pageNumber="274">A. moniliatus</emphasis>
(Chen, 1946)
</taxonomicName>
</th>
<th id="76B43600DB6C0037FBE7FA8362857D57" box="[1135,1420,1385,1407]" gridcol="3" gridrow="0" pageId="5" pageNumber="274">
<taxonomicName id="4C5526BDDB6CFFCDFBE7FA80625D7D57" authority="(Chen, 1946)" authorityName="Chen" authorityYear="1946" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[1135,1364,1385,1407]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="274" phylum="Annelida" rank="species" species="axillis">
<emphasis id="B921812CDB6CFFCDFBE7FA8063C07D57" box="[1135,1225,1385,1407]" italics="true" pageId="5" pageNumber="274">A. axillis</emphasis>
(Chen, 1946)
</taxonomicName>
</th>
</tr>
<tr id="35655F7CDB6C0037FF17FA7862857D80" box="[159,1420,1426,1448]" gridrow="1" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17FA7866737D80" box="[159,378,1426,1448]" gridcol="0" gridrow="1" pageId="5" pageNumber="274">First dorsal pore</th>
<td id="76B43600DB6C0037FE02FA7865FF7D80" box="[394,758,1426,1448]" gridcol="1" gridrow="1" pageId="5" pageNumber="274">12/13</td>
<td id="76B43600DB6C0037FC85FA7863517D80" box="[781,1112,1426,1448]" gridcol="2" gridrow="1" pageId="5" pageNumber="274">12/13</td>
<td id="76B43600DB6C0037FBE7FA7862857D80" box="[1135,1420,1426,1448]" gridcol="3" gridrow="1" pageId="5" pageNumber="274">11/12</td>
</tr>
<tr id="35655F7CDB6C0037FF17FA5662857DFA" box="[159,1420,1468,1490]" gridrow="2" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17FA5666737DFA" box="[159,378,1468,1490]" gridcol="0" gridrow="2" pageId="5" pageNumber="274">Setal formula</th>
<td id="76B43600DB6C0037FE02FA5665FF7DFA" box="[394,758,1468,1490]" gridcol="1" gridrow="2" pageId="5" pageNumber="274">AA=1.8–2.0AB, ZZ=2.0–3.0ZY</td>
<td id="76B43600DB6C0037FC85FA5663517DFA" box="[781,1112,1468,1490]" gridcol="2" gridrow="2" pageId="5" pageNumber="274">AA=1.3AB, ZZ=1.5–2.0ZY</td>
<td id="76B43600DB6C0037FBE7FA5662857DFA" box="[1135,1420,1468,1490]" gridcol="3" gridrow="2" pageId="5" pageNumber="274">AA=1.2AB, ZZ=1.2–1.5ZY</td>
</tr>
<tr id="35655F7CDB6C0037FF17FA0F62857DD3" box="[159,1420,1509,1531]" gridrow="3" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17FA0F66737DD3" box="[159,378,1509,1531]" gridcol="0" gridrow="3" pageId="5" pageNumber="274">Spermathecal pores</th>
<td id="76B43600DB6C0037FE02FA0F65FF7DD3" box="[394,758,1509,1531]" gridcol="1" gridrow="3" pageId="5" pageNumber="274">2 pairs, in 6/7–7/8</td>
<td id="76B43600DB6C0037FC85FA0F63517DD3" box="[781,1112,1509,1531]" gridcol="2" gridrow="3" pageId="5" pageNumber="274">3 pairs, in 6/7–8/9</td>
<td id="76B43600DB6C0037FBE7FA0F62857DD3" box="[1135,1420,1509,1531]" gridcol="3" gridrow="3" pageId="5" pageNumber="274">2 pairs, in 6/7–7/8</td>
</tr>
<tr id="35655F7CDB6C0037FF17F9E462857E0C" box="[159,1420,1550,1572]" gridrow="4" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F9E466737E0C" box="[159,378,1550,1572]" gridcol="0" gridrow="4" pageId="5" pageNumber="274">Papillae pre-clitellum</th>
<td id="76B43600DB6C0037FE02F9E465FF7E0C" box="[394,758,1550,1572]" gridcol="1" gridrow="4" pageId="5" pageNumber="274">none</td>
<td id="76B43600DB6C0037FC85F9E463517E0C" box="[781,1112,1550,1572]" gridcol="2" gridrow="4" pageId="5" pageNumber="274">many small papillae</td>
<td id="76B43600DB6C0037FBE7F9E462857E0C" box="[1135,1420,1550,1572]" gridcol="3" gridrow="4" pageId="5" pageNumber="274">none</td>
</tr>
<tr id="35655F7CDB6C0037FF17F9D262857E44" box="[159,1420,1592,1644]" gridrow="5" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F9D266737E44" box="[159,378,1592,1644]" gridcol="0" gridrow="5" pageId="5" pageNumber="274">Male pores</th>
<td id="76B43600DB6C0037FE02F9D265FF7E44" box="[394,758,1592,1644]" gridcol="1" gridrow="5" pageId="5" pageNumber="274">absent</td>
<td id="76B43600DB6C0037FC85F9D263517E44" box="[781,1112,1592,1644]" gridcol="2" gridrow="5" pageId="5" pageNumber="274">as a wart in large depressed (or 2 coalesced) papilla</td>
<td id="76B43600DB6C0037FBE7F9D262857E44" box="[1135,1420,1592,1644]" gridcol="3" gridrow="5" pageId="5" pageNumber="274">on raised conical porophore</td>
</tr>
<tr id="35655F7CDB6C0037FF17F96A62857EBE" box="[159,1420,1664,1686]" gridrow="6" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F96A66737EBE" box="[159,378,1664,1686]" gridcol="0" gridrow="6" pageId="5" pageNumber="274">Testis sacs</th>
<td id="76B43600DB6C0037FE02F96A65FF7EBE" box="[394,758,1664,1686]" gridcol="1" gridrow="6" pageId="5" pageNumber="274">absent</td>
<td id="76B43600DB6C0037FC85F96A63517EBE" box="[781,1112,1664,1686]" gridcol="2" gridrow="6" pageId="5" pageNumber="274">developed</td>
<td id="76B43600DB6C0037FBE7F96A62857EBE" box="[1135,1420,1664,1686]" gridcol="3" gridrow="6" pageId="5" pageNumber="274">developed</td>
</tr>
<tr id="35655F7CDB6C0037FF17F94362857E97" box="[159,1420,1705,1727]" gridrow="7" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F94366737E97" box="[159,378,1705,1727]" gridcol="0" gridrow="7" pageId="5" pageNumber="274">Seminal vesicles</th>
<td id="76B43600DB6C0037FE02F94365FF7E97" box="[394,758,1705,1727]" gridcol="1" gridrow="7" pageId="5" pageNumber="274">small</td>
<td id="76B43600DB6C0037FC85F94363517E97" box="[781,1112,1705,1727]" gridcol="2" gridrow="7" pageId="5" pageNumber="274">well-developed</td>
<td id="76B43600DB6C0037FBE7F94362857E97" box="[1135,1420,1705,1727]" gridcol="3" gridrow="7" pageId="5" pageNumber="274">developed</td>
</tr>
<tr id="35655F7CDB6C0037FF17F93862857EC0" box="[159,1420,1746,1768]" gridrow="8" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F93866737EC0" box="[159,378,1746,1768]" gridcol="0" gridrow="8" pageId="5" pageNumber="274">Prostatic duct</th>
<td id="76B43600DB6C0037FE02F93865FF7EC0" box="[394,758,1746,1768]" gridcol="1" gridrow="8" pageId="5" pageNumber="274">absent</td>
<td id="76B43600DB6C0037FC85F93863517EC0" box="[781,1112,1746,1768]" gridcol="2" gridrow="8" pageId="5" pageNumber="274">S-shaped</td>
<td id="76B43600DB6C0037FBE7F93862857EC0" box="[1135,1420,1746,1768]" gridcol="3" gridrow="8" pageId="5" pageNumber="274">absent</td>
</tr>
<tr id="35655F7CDB6C0037FF17F91662857F18" box="[159,1420,1788,1840]" gridrow="9" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F91666737F18" box="[159,378,1788,1840]" gridcol="0" gridrow="9" pageId="5" pageNumber="274">Accessory glands</th>
<td id="76B43600DB6C0037FE02F91665FF7F18" box="[394,758,1788,1840]" gridcol="1" gridrow="9" pageId="5" pageNumber="274">none</td>
<td id="76B43600DB6C0037FC85F91663517F18" box="[781,1112,1788,1840]" gridcol="2" gridrow="9" pageId="5" pageNumber="274">globular, stalked, gland ovoid</td>
<td id="76B43600DB6C0037FBE7F91662857F18" box="[1135,1420,1788,1840]" gridcol="3" gridrow="9" pageId="5" pageNumber="274">very small, with spherical gland portion and long duct</td>
</tr>
<tr id="35655F7CDB6C0037FF17F8AE62857F50" box="[159,1420,1860,1912]" gridrow="10" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F8AE66737F50" box="[159,378,1860,1912]" gridcol="0" gridrow="10" pageId="5" pageNumber="274">Spermathecae</th>
<td id="76B43600DB6C0037FE02F8AE65FF7F50" box="[394,758,1860,1912]" gridcol="1" gridrow="10" pageId="5" pageNumber="274">1.4–1.6 mm; ampulla ovoid; duct as long as 2.5 ampulla</td>
<td id="76B43600DB6C0037FC85F8AE63517F50" box="[781,1112,1860,1912]" gridcol="2" gridrow="10" pageId="5" pageNumber="274">absent or in VII–IX, 1.0 mm; ampulla elongate, its ductlonger</td>
<td id="76B43600DB6C0037FBE7F8AE62857F50" box="[1135,1420,1860,1912]" gridcol="3" gridrow="10" pageId="5" pageNumber="274">ampulla date-shaped, with long duct</td>
</tr>
<tr id="35655F7CDB6C0037FF17F86662857FF7" box="[159,1420,1932,2015]" gridrow="11" pageId="5" pageNumber="274">
<th id="76B43600DB6C0037FF17F86666737FF7" box="[159,378,1932,2015]" gridcol="0" gridrow="11" pageId="5" pageNumber="274">Diverticulum</th>
<td id="76B43600DB6C0037FE02F86665FF7FF7" box="[394,758,1932,2015]" gridcol="1" gridrow="11" pageId="5" pageNumber="274">slightly shorter than main pouch, terminal 2/5 dilated into a long marsupial chamber</td>
<td id="76B43600DB6C0037FC85F86663517FF7" box="[781,1112,1932,2015]" gridcol="2" gridrow="11" pageId="5" pageNumber="274">twice as long, zigzagly constricted, moniliform, transparent</td>
<td id="76B43600DB6C0037FBE7F86662857FF7" box="[1135,1420,1932,2015]" gridcol="3" gridrow="11" pageId="5" pageNumber="274">only by a bud-like structure at ental 1/5 of main duct</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEA5D3EDB6FFFCEFF4FFF72639979A0" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="275">
<emphasis id="B921812CDB6FFFCEFF4FFF7266B57898" bold="true" box="[199,444,152,176]" pageId="6" pageNumber="275">External characters.</emphasis>
Pinky pigment generally. Dimensions
<quantity id="4CADF0DBDB6FFFCEFCFAFF7D64E57887" box="[882,1004,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" metricValueMax="7.0" metricValueMin="5.8" pageId="6" pageNumber="275" unit="mm" value="64.0" valueMax="70.0" valueMin="58.0">58–70 mm</quantity>
by 2.4–3.0 mm at clitellum, segments 60–73. Prostomium 1/2 epilobous. First dorsal pore in 12/13. Setae numbering 24–28 at III, 34–37 at V, 44–47 at VIII, 35–40 at XX, 36–40 at XXV; 13–16 between spermathecal pores; setal formula: AA=1.8–2.0AB, ZZ=2.0– 3.0ZY. Clitellum annular in XIV–XVI, light brown, swollen, setae invisible externally. Spermathecal pores two pairs in 6/7–7/8, ventral, eye-like, 0.33 circumference ventrally apart from each other (
<figureCitation id="136E41BBDB6FFFCEFBD3FECD63BC7968" box="[1115,1205,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[163,1424,721,1840]" captionTargetId="figure@6.[151,1436,700,1840]" captionTargetPageId="6" captionText="FIGURE 3. Amynthas marsupiformis Jiang &amp; Yuan, sp. nov. A. Region of spermathecal pores, ventral view. B. Region of male pores, ventral view. C. Spermatheca. D. Intestinal caeca (ventral margin is on the left)." httpUri="https://zenodo.org/record/1446809/files/figure.png" pageId="6" pageNumber="275">Fig. 3A</figureCitation>
). No genital papillae observed. Male pores absent. 12 tiny presetal or postsetal genital papillae in parallel rows present on the ventrolateral sides of XVII–XIX (
<figureCitation id="136E41BBDB6FFFCEFD9BFE85656379A0" box="[531,618,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[163,1424,721,1840]" captionTargetId="figure@6.[151,1436,700,1840]" captionTargetPageId="6" captionText="FIGURE 3. Amynthas marsupiformis Jiang &amp; Yuan, sp. nov. A. Region of spermathecal pores, ventral view. B. Region of male pores, ventral view. C. Spermatheca. D. Intestinal caeca (ventral margin is on the left)." httpUri="https://zenodo.org/record/1446809/files/figure.png" pageId="6" pageNumber="275">Fig. 3B</figureCitation>
). Female pore single in XIV, ovoid, milky white.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EB5DB6FFFCEFF4FFE7E632A7AAD" pageId="6" pageNumber="275" type="description">
<paragraph id="8BEA5D3EDB6FFFCEFF4FFE7E632A7AAD" blockId="6.[151,1437,151,645]" pageId="6" pageNumber="275">
<emphasis id="B921812CDB6FFFCEFF4FFE7E66B37984" bold="true" box="[199,442,404,428]" pageId="6" pageNumber="275">Internal characters.</emphasis>
Septa 5/6–7/8 thick and muscular,
<date id="FFEB7BFEDB6FFFCEFCECFE7E64F97984" box="[868,1008,404,428]" pageId="6" pageNumber="275" value="2014-10-11" valueMax="2014-10-13" valueMin="2014-10-11">10/11–13/14</date>
slightly thickened,
<date id="FFEB7BFEDB6FFFCEFB55FE7E62487984" box="[1245,1345,404,428]" pageId="6" pageNumber="275" value="2010-08-09" valueMax="2010-08-09" valueMin="2010-08-09">8/9–9/10</date>
absent. Gizzard ball-shaped, in IX–X. Intestine enlarged distinctly from XV. Typhlosole medium from XXVII. Intestinal caeca paired in XXVII, simple, smooth, extending anteriorly to XXIII (
<figureCitation id="136E41BBDB6FFFCEFC5EFE36633A79DC" box="[982,1075,476,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[163,1424,721,1840]" captionTargetId="figure@6.[151,1436,700,1840]" captionTargetPageId="6" captionText="FIGURE 3. Amynthas marsupiformis Jiang &amp; Yuan, sp. nov. A. Region of spermathecal pores, ventral view. B. Region of male pores, ventral view. C. Spermatheca. D. Intestinal caeca (ventral margin is on the left)." httpUri="https://zenodo.org/record/1446809/files/figure.png" pageId="6" pageNumber="275">Fig. 3D</figureCitation>
). Esophageal hearts in X–XIII. Ovaries in XIII. Spermathecae two pairs in VII and VIII, small, about
<quantity id="4CADF0DBDB6FFFCEFC4CFE1563447A3F" box="[964,1101,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="6" pageNumber="275" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
long, ampulla ovoid; duct as long as 2.5 ampulla. Diverticulum slightly shorter than main pouch (ampulla and duct), terminal 2/5 dilated into a long marsupial seminal chamber (
<figureCitation id="136E41BBDB6FFFCEFD9FFDAD65797A48" box="[535,624,583,608]" captionStart="FIGURE 3" captionStartId="6.[151,250,1861,1883]" captionTargetBox="[163,1424,721,1840]" captionTargetId="figure@6.[151,1436,700,1840]" captionTargetPageId="6" captionText="FIGURE 3. Amynthas marsupiformis Jiang &amp; Yuan, sp. nov. A. Region of spermathecal pores, ventral view. B. Region of male pores, ventral view. C. Spermatheca. D. Intestinal caeca (ventral margin is on the left)." httpUri="https://zenodo.org/record/1446809/files/figure.png" pageId="6" pageNumber="275">Fig. 3C</figureCitation>
). No accessory glands observed. Seminal vesicles two pairs, in XI–XII, very small. No testis sacs, prostate glands, accessory glands and sperm observed.
</paragraph>
</subSubSection>
<caption id="DF2A0DB6DB6FFFCEFF1FF8AF630A7F52" httpUri="https://zenodo.org/record/1446809/files/figure.png" pageId="6" pageNumber="275" startId="6.[151,250,1861,1883]" targetBox="[163,1424,721,1840]" targetPageId="6">
<paragraph id="8BEA5D3EDB6FFFCEFF1FF8AF630A7F52" blockId="6.[151,1436,1861,1914]" pageId="6" pageNumber="275">
<emphasis id="B921812CDB6FFFCEFF1FF8AF661B7F73" bold="true" box="[151,274,1861,1883]" pageId="6" pageNumber="275">FIGURE 3.</emphasis>
<taxonomicName id="4C5526BDDB6FFFCEFE90F8AF65AF7F73" authority="Jiang &amp; Yuan" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[280,678,1861,1883]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="275" phylum="Annelida" rank="species" species="marsupiformis" status="sp. nov.">
<emphasis id="B921812CDB6FFFCEFE90F8AF651C7F73" box="[280,533,1861,1883]" italics="true" pageId="6" pageNumber="275">Amynthas marsupiformis</emphasis>
Jiang &amp; Yuan
</taxonomicName>
,
<emphasis id="B921812CDB6FFFCEFD38F8AC64177F73" bold="true" box="[688,798,1862,1883]" pageId="6" pageNumber="275">
<taxonomicNameLabel id="A2123C57DB6FFFCEFD38F8AC64097F73" box="[688,768,1862,1883]" pageId="6" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Region of spermathecal pores, ventral view.
<emphasis id="B921812CDB6FFFCEFB6CF8AC63F27F73" bold="true" box="[1252,1275,1862,1883]" pageId="6" pageNumber="275">B.</emphasis>
Region of male pores, ventral view.
<emphasis id="B921812CDB6FFFCEFEEBF88E66727F52" bold="true" box="[355,379,1892,1914]" pageId="6" pageNumber="275">C.</emphasis>
Spermatheca.
<emphasis id="B921812CDB6FFFCEFD99F88F65237F52" bold="true" box="[529,554,1893,1914]" pageId="6" pageNumber="275">D.</emphasis>
Intestinal caeca (ventral margin is on the left).
</paragraph>
</caption>
<subSubSection id="C34F0EB5DB6FFFCFFF4FF84D66B97A15" lastPageId="7" lastPageNumber="276" pageId="6" pageNumber="275" type="discussion">
<paragraph id="8BEA5D3EDB6FFFCFFF4FF84D66B97A15" blockId="6.[151,1436,1959,2020]" lastBlockId="7.[151,1437,151,573]" lastPageId="7" lastPageNumber="276" pageId="6" pageNumber="275">
<emphasis id="B921812CDB6FFFCEFF4FF84D66327F97" bold="true" box="[199,315,1959,1983]" pageId="6" pageNumber="275">Remarks.</emphasis>
Absence of sperm and main reproductive organs recorded suggest parthenogenesis (
<bibRefCitation id="EFC420CFDB6FFFCEFB65F84D62787F97" box="[1261,1393,1959,1983]" pageId="6" pageNumber="275" refId="ref4688" refString="Gates, G. E. (1972) Burmese earthworms: an tntroduction to the systematics and biology of Megadrile Oligochaetes with special reference to Southeast Asia. Transactions of the American Philosophical Society, 62, 1 - 326. https: // doi. org / 10.2307 / 1006214" type="journal article">Gates 1972</bibRefCitation>
).
<taxonomicName id="4C5526BDDB6FFFCEFA0CF84D66327FCB" authority="Jiang &amp; Dong &amp; Yuan &amp; Qiu, 2018" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="6" pageNumber="275" phylum="Annelida" rank="species" species="marsupiformis" status="sp. nov.">
<emphasis id="B921812CDB6FFFCEFA0CF84D66327FCB" italics="true" pageId="6" pageNumber="275">A. marsupiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2123C57DB6FFFCEFECCF82166A97FCB" box="[324,416,1995,2019]" pageId="6" pageNumber="275" rank="species">
<emphasis id="B921812CDB6FFFCEFECCF82166A97FCB" bold="true" box="[324,416,1995,2019]" pageId="6" pageNumber="275">sp. nov.</emphasis>
</taxonomicNameLabel>
which has no male pores, testis sacs and prostate glands is certainly parthenogenetic. In terms of external characters,
<taxonomicName id="4C5526BDDB6EFFCFFE68FF7265AD7898" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[480,676,152,176]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="marsupiformis">
<emphasis id="B921812CDB6EFFCFFE68FF7265AD7898" box="[480,676,152,176]" italics="true" pageId="7" pageNumber="276">A. marsupiformis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C5526BDDB6EFFCFFCFEFF7263717898" authority="(Chen, 1946)" authorityName="Chen" authorityYear="1946" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[886,1144,151,176]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="axillis">
<emphasis id="B921812CDB6EFFCFFCFEFF7264D27898" box="[886,987,152,176]" italics="true" pageId="7" pageNumber="276">A. axillis</emphasis>
(
<bibRefCitation id="EFC420CFDB6EFFCFFC63FF7D63797898" box="[1003,1136,151,176]" pageId="7" pageNumber="276" refId="ref4463" refString="Chen, Y. (1946) On the terrestrial Oligochaeta from Szechuan?. Journal of the West China Border Research Society, 16, 83 - 141." type="journal article">Chen, 1946</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C5526BDDB6EFFCFFB39FF7267D178FD" authority="(Chen, 1946)" authorityName="Chen" authorityYear="1946" baseAuthorityName="Chen" baseAuthorityYear="1946" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="moniliatus">
<emphasis id="B921812CDB6EFFCFFB39FF7262417898" box="[1201,1352,152,176]" italics="true" pageId="7" pageNumber="276">A. moniliatus</emphasis>
(
<bibRefCitation id="EFC420CFDB6EFFCFFAD0FF7D67D978FD" pageId="7" pageNumber="276" refId="ref4463" refString="Chen, Y. (1946) On the terrestrial Oligochaeta from Szechuan?. Journal of the West China Border Research Society, 16, 83 - 141." type="journal article">Chen, 1946</bibRefCitation>
)
</taxonomicName>
in the characteristics of the post-clitellum papillae. All of them have many small papillae in ventrolateral lines on segments XVII–XIX. Besides, they have similar small body size, reddish or purple pigments, sparse setae and no prostate glands. The main differences of
<taxonomicName id="4C5526BDDB6EFFCFFD55FEEE64AC7934" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[733,933,260,284]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="marsupiformis">
<emphasis id="B921812CDB6EFFCFFD55FEEE64AC7934" box="[733,933,260,284]" italics="true" pageId="7" pageNumber="276">A. marsupiformis</emphasis>
</taxonomicName>
from these two species are small seminal vesicles, absence of male pores, testis sacs and accessory glands. In contrast,
<taxonomicName id="4C5526BDDB6EFFCFFC7DFEC263507968" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[1013,1113,296,320]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="axillis">
<emphasis id="B921812CDB6EFFCFFC7DFEC263507968" box="[1013,1113,296,320]" italics="true" pageId="7" pageNumber="276">A. axillis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5526BDDB6EFFCFFB06FEC2622A7968" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[1166,1315,296,320]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="moniliatus">
<emphasis id="B921812CDB6EFFCFFB06FEC2622A7968" box="[1166,1315,296,320]" italics="true" pageId="7" pageNumber="276">A. moniliatus</emphasis>
</taxonomicName>
have male pores and stalked accessory glands, and their testis sacs and seminal vesicles are developed or well developed.
<tableCitation id="C6D76885DB6EFFCFFF1FFE8567E779A0" box="[151,238,367,392]" captionStart="TABLE 2" captionStartId="5.[151,239,1302,1325]" captionText="TABLE 2. Comparison of A. marsupiformis sp. nov. and two species with the same arrangement of post-clitellar papillae." pageId="7" pageNumber="276">Table 2</tableCitation>
lists the differences among them in detail. In addition,
<taxonomicName id="4C5526BDDB6EFFCFFC08FE9A634F79A0" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[896,1094,368,392]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="marsupiformis">
<emphasis id="B921812CDB6EFFCFFC08FE9A634F79A0" box="[896,1094,368,392]" italics="true" pageId="7" pageNumber="276">A. marsupiformis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5526BDDB6EFFCFFB6EFE9A624779A0" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[1254,1358,368,392]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="axillis">
<emphasis id="B921812CDB6EFFCFFB6EFE9A624779A0" box="[1254,1358,368,392]" italics="true" pageId="7" pageNumber="276">A. axillis</emphasis>
</taxonomicName>
in the features of the first dorsal pore, setal formula and diverticulum. The diverticulum of
<taxonomicName id="4C5526BDDB6EFFCFFBEEFE7E63C47984" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[1126,1229,404,428]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="axillis">
<emphasis id="B921812CDB6EFFCFFBEEFE7E63C47984" box="[1126,1229,404,428]" italics="true" pageId="7" pageNumber="276">A. axillis</emphasis>
</taxonomicName>
is quite short and bud-like. Besides,
<taxonomicName id="4C5526BDDB6EFFCFFEFBFE52653379F8" authorityName="Jiang &amp; Dong &amp; Yuan &amp; Qiu" authorityYear="2018" box="[371,570,440,464]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="marsupiformis">
<emphasis id="B921812CDB6EFFCFFEFBFE52653379F8" box="[371,570,440,464]" italics="true" pageId="7" pageNumber="276">A. marsupiformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5526BDDB6EFFCFFDF4FE52641E79F8" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[636,791,440,464]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="moniliatus">
<emphasis id="B921812CDB6EFFCFFDF4FE52641E79F8" box="[636,791,440,464]" italics="true" pageId="7" pageNumber="276">A. moniliatus</emphasis>
</taxonomicName>
have different setal formula, spermathecal pores, preclitellum papillae, prostatic duct and spermathecae.
<taxonomicName id="4C5526BDDB6EFFCFFD6EFE36647479DC" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[742,893,476,500]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="276" phylum="Annelida" rank="species" species="moniliatus">
<emphasis id="B921812CDB6EFFCFFD6EFE36647479DC" box="[742,893,476,500]" italics="true" pageId="7" pageNumber="276">A. moniliatus</emphasis>
</taxonomicName>
, belonging to the
<emphasis id="B921812CDB6EFFCFFBC6FE3663AE79DC" box="[1102,1191,476,500]" italics="true" pageId="7" pageNumber="276">sieboldi</emphasis>
-group, has 3 pairs of spermathecal pores with many small papillae, S-shaped prostatic duct, stout spermathecal duct and moniliformshaped seminal chamber.
</paragraph>
</subSubSection>
</treatment>
</document>
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