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<document id="8EF2CBAD578B15AB11F18FBB8B999423" ID-DOI="10.5281/zenodo.192017" ID-GBIF-Dataset="5b26ee34-55ff-4da4-a200-444541506cd0" ID-ISSN="1175-5326" ID-Zenodo-Dep="192017" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461140117641" checkinUser="plazi" docAuthor="Imura, Jimpei &amp; Ishikawa, Tadashi" docDate="2009" docId="03CE87E8AB20676CFF6E80B412C7FD96" docLanguage="en" docName="zt02315p030.pdf" docOrigin="Zootaxa 2315" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Schiodtella formosana Takano &amp; Yanagihara 1939" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FFF7FF90AB266765FFF98370112EFFA2" masterDocTitle="A review of Schiodtella formosana, with some remarks on the genus Schiodtella and description of a Japanese new species (Hemiptera: Heteroptera: Cydnidae)" masterLastPageNumber="30" masterPageNumber="19" pageNumber="25" updateTime="1698598840229" updateUser="plazi">
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<mods:title id="E94E4AD09536F540F58D273707BE82D2">A review of Schiodtella formosana, with some remarks on the genus Schiodtella and description of a Japanese new species (Hemiptera: Heteroptera: Cydnidae)</mods:title>
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<mods:namePart id="50C80FE842A9E4468DB2D8C2AB31821A">Imura, Jimpei</mods:namePart>
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<mods:namePart id="4622C7E44CAEAF63E33388AC6439295C">Ishikawa, Tadashi</mods:namePart>
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<mods:date id="0AC2AEB79AB8AA810CB2A6AD92389FBE">2009</mods:date>
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<treatment id="03CE87E8AB20676CFF6E80B412C7FD96" ID-DOI="http://doi.org/10.5281/zenodo.6226907" ID-GBIF-Taxon="119627849" ID-Zenodo-Dep="6226907" LSID="urn:lsid:plazi:treatment:03CE87E8AB20676CFF6E80B412C7FD96" httpUri="http://treatment.plazi.org/id/03CE87E8AB20676CFF6E80B412C7FD96" lastPageId="9" lastPageNumber="28" pageId="6" pageNumber="25">
<subSubSection id="C37D6575AB206763FF6E80B410A2FBA2" pageId="6" pageNumber="25" type="nomenclature">
<paragraph id="8BD836FEAB206763FF6E80B4120EFC7F" blockId="6.[151,800,963,1024]" box="[151,800,963,990]" pageId="6" pageNumber="25">
<heading id="D0908192AB206763FF6E80B4120EFC7F" bold="true" box="[151,800,963,990]" fontSize="11" level="1" pageId="6" pageNumber="25" reason="1">
<taxonomicName id="4C674D7DAB206763FF6E80B4120EFC7F" authority="Takano &amp; Yanagihara, 1939" authorityName="Takano &amp; Yanagihara" authorityYear="1939" box="[151,800,963,990]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="25" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB206763FF6E80B4120EFC7F" bold="true" box="[151,800,963,990]" pageId="6" pageNumber="25">
<emphasis id="B913EAECAB206763FF6E80B41089FC7C" bold="true" box="[151,423,964,990]" italics="true" pageId="6" pageNumber="25">Schiodtella formosana</emphasis>
(
<bibRefCitation id="EFF64B0FAB206763FE4F80B31236FC7F" author="Takano" box="[438,792,963,989]" pageId="6" pageNumber="25" refString="Takano, S. &amp; Yanagihara, M. (1939) Researches on injurious and beneficial animals of sugarcane. Taiwan Sugar Experimental Station, Extra Reports, 2, 311 pp., 18 pls." type="book" year="1939">Takano &amp; Yanagihara, 1939</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD836FEAB206763FF6E809610A2FBA2" blockId="6.[151,800,963,1024]" box="[151,396,998,1024]" pageId="6" pageNumber="25">
(
<figureCitation id="135C2A7BAB206763FF6680961003FBA2" box="[159,301,998,1024]" captionStart-0="FIGURE 17 21" captionStart-1="FIGURE 22 30" captionStartId-0="7.[151,258,1851,1875]" captionStartId-1="8.[151,255,1497,1521]" captionTargetBox-0="[161,1434,836,1789]" captionTargetBox-1="[168,1420,201,1466]" captionTargetId-0="figure@7.[151,1436,815,1825]" captionTargetId-1="figure@8.[151,1436,194,1474]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." captionText-1="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri-0="https://zenodo.org/record/192021/files/figure.png" httpUri-1="https://zenodo.org/record/192022/files/figure.png" pageId="6" pageNumber="25">Figs. 1730</figureCitation>
,
<figureCitation id="135C2A7BAB206763FEC3809610ABFBA2" box="[314,389,998,1024]" captionStart="FIGURE 31 39" captionStartId="9.[151,256,1612,1636]" captionTargetBox="[151,1434,622,1586]" captionTargetId="figure@9.[151,1436,622,1587]" captionTargetPageId="9" captionText="FIGURE 31 39. Leg structures of Schiodtella spp. 31 33, Sc. japonica sp. nov., a form with shortened wings, apical part of pro- (31), meso- (32) and meta- (33) tarsi; 34 36, Sc. japonica sp. nov., macropterous form, apical part of pro- (34), meso- (35) and mata- (36) tarsi; 37 39, Sc. formosana, apical part of pro- (36), meso- (37) and meta- (38) tibiae. Abbreviations: Cl, claw; Pv, pulvillus; Ts 3, tarsal segment III. Scales: 20 μm for 31 33, 50 μm for 34 36, 100 μm for 37 39." httpUri="https://zenodo.org/record/192023/files/figure.png" pageId="6" pageNumber="25">3739</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D6575AB206763FF6E87441597FBE9" box="[151,1209,1076,1099]" pageId="6" pageNumber="25" type="reference_group">
<paragraph id="8BD836FEAB206763FF6E87441597FBE9" blockId="6.[151,1209,1076,1099]" box="[151,1209,1076,1099]" pageId="6" pageNumber="25">
<treatmentCitationGroup id="AB7711D0AB206763FF6E87441597FBE9" box="[151,1209,1076,1099]" pageId="6" pageNumber="25">
<taxonomicName id="4C674D7DAB206763FF6E87441201FBE9" authority="Takano and Yanagihara, 1939: 135" authorityName="Takano and Yanagihara" authorityPageNumber="135" authorityYear="1939" box="[151,815,1076,1099]" class="Insecta" family="Cydnidae" genus="Neostibaropus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="25" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B913EAECAB206763FF6E8744109AFBE9" box="[151,436,1076,1099]" italics="true" pageId="6" pageNumber="25">Neostibaropus formosanus</emphasis>
<treatmentCitation id="0AC610EFAB206763FE4287441201FBE9" author="Takano" box="[443,815,1076,1099]" page="135" pageId="6" pageNumber="25" year="1939">
<bibRefCitation id="EFF64B0FAB206763FE42874413D5FBE9" author="Takano" box="[443,763,1076,1099]" pageId="6" pageNumber="25" refString="Takano, S. &amp; Yanagihara, M. (1939) Researches on injurious and beneficial animals of sugarcane. Taiwan Sugar Experimental Station, Extra Reports, 2, 311 pp., 18 pls." type="book" year="1939">Takano and Yanagihara, 1939</bibRefCitation>
: 135
</treatmentCitation>
</taxonomicName>
, pl. 4, fig. 19.
<typeStatus id="54DC885CAB206763FC33874412D1FBE9" box="[970,1023,1076,1099]" pageId="6" pageNumber="25">Type</typeStatus>
locality:
<collectingCountry id="F370766EAB206763FB9D87441598FBE9" box="[1124,1206,1076,1099]" name="Taiwan" pageId="6" pageNumber="25">Taiwan</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37D6575AB206763FF6E870A102BFA8D" pageId="6" pageNumber="25" type="diagnosis">
<paragraph id="8BD836FEAB206763FF6E870A102BFA8D" blockId="6.[151,1437,1146,2023]" pageId="6" pageNumber="25">
<emphasis id="B913EAECAB206763FF6E870A1032FB36" bold="true" box="[151,284,1146,1172]" italics="true" pageId="6" pageNumber="25">Diagnosis.</emphasis>
Head strongly wrinkled in dorsum (
<figureCitation id="135C2A7BAB206763FD20870A1240FB36" box="[729,878,1146,1172]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Figs. 18, 19</figureCitation>
); eyes very small (
<figureCitation id="135C2A7BAB206763FBAE870A15C2FB36" box="[1111,1260,1146,1172]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Figs. 18, 19</figureCitation>
); ocular index 13.516. Scutellum short and rounded, almost as long as wide (
<figureCitation id="135C2A7BAB206763FC8487D112E3FB19" box="[893,973,1185,1211]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Fig.17</figureCitation>
). Peritreme lacking apical process (
<figureCitation id="135C2A7BAB206763FA9587D11197FB40" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Fig. 20</figureCitation>
). Tarsi absent from all legs (
<figureCitation id="135C2A7BAB206763FDF687B8138EFB40" box="[527,672,1224,1250]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="6" pageNumber="25">Figs. 2225</figureCitation>
,
<figureCitation id="135C2A7BAB206763FD5487B813D2FB40" box="[685,764,1224,1250]" captionStart="FIGURE 31 39" captionStartId="9.[151,256,1612,1636]" captionTargetBox="[151,1434,622,1586]" captionTargetId="figure@9.[151,1436,622,1587]" captionTargetPageId="9" captionText="FIGURE 31 39. Leg structures of Schiodtella spp. 31 33, Sc. japonica sp. nov., a form with shortened wings, apical part of pro- (31), meso- (32) and meta- (33) tarsi; 34 36, Sc. japonica sp. nov., macropterous form, apical part of pro- (34), meso- (35) and mata- (36) tarsi; 37 39, Sc. formosana, apical part of pro- (36), meso- (37) and meta- (38) tibiae. Abbreviations: Cl, claw; Pv, pulvillus; Ts 3, tarsal segment III. Scales: 20 μm for 31 33, 50 μm for 34 36, 100 μm for 37 39." httpUri="https://zenodo.org/record/192023/files/figure.png" pageId="6" pageNumber="25">3739</figureCitation>
). Median arm of basal plate of penis shorter than lateral arms (
<figureCitation id="135C2A7BAB206763FF19879E1019FAAA" box="[224,311,1262,1288]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="6" pageNumber="25">Fig. 26</figureCitation>
); conjunctival appendage strongly protruding from phallotreme and recurved downwards in rest (
<figureCitation id="135C2A7BAB206763FF66866511D9FA8D" box="[159,247,1301,1327]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="6" pageNumber="25">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D6575AB206762FF3C864C13A7FDF9" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25" type="description">
<paragraph id="8BD836FEAB206763FF3C864C126DFA68" blockId="6.[151,1437,1146,2023]" pageId="6" pageNumber="25">
<emphasis id="B913EAECAB206763FF3C864C109FFAF7" bold="true" box="[197,433,1339,1366]" pageId="6" pageNumber="25">
<emphasis id="B913EAECAB206763FF3C864C1073FAF4" bold="true" box="[197,349,1340,1366]" italics="true" pageId="6" pageNumber="25">Description.</emphasis>
Male:
</emphasis>
― total length 4.254.65; total width 2.83.0; body 2.352.5 high; head 0.951.0 long, 0.920.95 wide; length of antennal segments I 0.2, II 0.30.32, III 0.20.22, IV 0.350.38; length of rostral segments I 0.4, II 0.45, III 0.30.32, IV 0.32; pronotum 1.441.52 long, 2.362.7 wide; scutellum 1.851.95 long, 1.81.95 wide; hemelytron 2.82.86 long.
</paragraph>
<paragraph id="8BD836FEAB206763FF3C86A6152DF99C" blockId="6.[151,1437,1146,2023]" pageId="6" pageNumber="25">Body yellowish brown to brown; dorsum of head and anterior part of pronotum blackish. Eyes reddish; ocelli orange to reddish. Antennae and rostrum yellowish brown. Hemelytral membranes hyaline. Legs yellowish brown; apical half of protibia and corbicle of metatibia blackish.</paragraph>
<paragraph id="8BD836FEAB206762FF3C853A1292FD96" blockId="6.[151,1437,1146,2023]" lastBlockId="7.[151,1437,152,758]" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25">
Total length about 1.5 times as long as total width, 1.8 times as long as body height. Head (
<figureCitation id="135C2A7BAB206763FAFD853A14BDF9C6" box="[1284,1427,1610,1636]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Figs. 18, 19</figureCitation>
) slightly longer than wide, strongly wrinkled in dorsum; clypeus (
<figureCitation id="135C2A7BAB206763FC5685011526F929" box="[943,1032,1649,1675]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Fig. 18</figureCitation>
) slightly shorter than paraclypei; paraclypeus (
<figureCitation id="135C2A7BAB206763FECE85E810E6F910" box="[311,456,1688,1714]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Figs. 18, 19</figureCitation>
) fringed with 710 peg-bearing tubercles, and with several setae anteroventrally; primary setae IIIV (
<figureCitation id="135C2A7BAB206763FE6285CE1301F97A" box="[411,559,1726,1752]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Figs. 18, 19</figureCitation>
) peg-like, thinner than secondary ones. Eyes (
<figureCitation id="135C2A7BAB206763FB9485CE142FF97A" box="[1133,1281,1726,1752]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Figs. 18, 19</figureCitation>
) very small, weakly prominent; ocular index 13.516; ocellar index 6.5; interocellar index 3.33.5. Antenna covered with thin, sparse setae in segments I and II, and dense setae in III and IV; approximate proportion of segments I to IV 1.0: 1.5: 1.0: 1.8. Rostrum surpassing procoxae; approximate proportion of segments I to IV 1.3: 1.5: 1.0: 1.0. Pronotum (
<figureCitation id="135C2A7BAB206763FEA884291086F8D1" box="[337,424,1881,1907]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Fig. 17</figureCitation>
) distinctly wrinkled in anteromedial part and posterior half, mixed with punctures in posteromedial part, and with 1419 setae on lateral margin and 1 seta behind callus (
<figureCitation id="135C2A7BAB206763FB6984F015C6F838" box="[1168,1256,1920,1946]" captionStart="FIGURE 10 16" captionStartId="4.[151,256,1832,1856]" captionTargetBox="[181,1418,605,1790]" captionTargetId="figure@4.[171,1425,583,1810]" captionTargetPageId="4" captionText="FIGURE 10 16. Schiodtella japonica sp. nov. 10 11, Penis, dorsal (10) and lateral (11) views; 12 14, left paramere, ventral (12), dorsal (13) and inner (14) views; 15, spermatheca; 16, female terminalia (gonocoxa I and setae of left side removed), posterior view. Abbreviations: Ca, conjunctival appendage; Dd, dilation of spermathecal duct; Dpd, distal part of spermathecal duct; Gc 1 &amp; 2, gonocoxae I &amp; II; Hp, hypophysis; IP, intermediate part; La, lateral arm of basal plate; Ma, median arm of basal plate; Mt 8 &amp; 9, mediotergites VIII &amp; IX; Phtr, phallotreme; Ppd, proximal part of spermathecal duct; Pt 8 &amp; 9, paratergites VIII &amp; IX; R, receptacle; Th, theca. Scales: 0.2 mm for 10 15, 0.5 mm for 16." httpUri="https://zenodo.org/record/192019/files/figure.png" pageId="6" pageNumber="25">Fig. 16</figureCitation>
), on each side. Scutellum (
<figureCitation id="135C2A7BAB206763FEE484D6105FF862" box="[285,369,1958,1984]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Fig. 17</figureCitation>
) short, rounded apically, almost as long as wide, entirely wrinkled. Thoracic pleura smooth except evaporatoria; peritreme without apical process (
<figureCitation id="135C2A7BAB206763FCDA84BD1253F845" box="[803,893,1997,2023]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Fig. 20</figureCitation>
). Hemelytron (
<figureCitation id="135C2A7BAB206763FBD684BD15ABF845" box="[1071,1157,1997,2023]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="6" pageNumber="25">Fig. 17</figureCitation>
) short, not reaching tip of abdomen; corium (
<figureCitation id="135C2A7BAB216762FE6E83E810C2FF10" box="[407,492,152,178]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="7" pageNumber="26">Fig. 17</figureCitation>
) indistinctly covered with fine punctures and wrinkles, and with 6 or 7 setae on costal margin; clavus (
<figureCitation id="135C2A7BAB216762FE6683CE10DAFF7A" box="[415,500,190,216]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="7" pageNumber="26">Fig. 17</figureCitation>
) with 1 seta basally. Protibia (
<figureCitation id="135C2A7BAB216762FCAF83CE1282FF7A" box="[854,940,190,216]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Fig. 22</figureCitation>
) covered with setae between proximal 2/5 and distal 1/5 of anterior side, and with 5 spines on tibial edge. Setae of mesotibia gradually becoming longer and stouter toward tibial apex in anterior and dorsal sides (
<figureCitation id="135C2A7BAB216762FCBB827C12B7FE84" box="[834,921,268,294]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Fig. 23</figureCitation>
).
<collectingCountry id="F370766EAB216762FC57827C12C7FE84" box="[942,1001,268,294]" name="India" pageId="7" pageNumber="26">Hind</collectingCountry>
leg (
<figureCitation id="135C2A7BAB216762FBDA827C1580FE84" box="[1059,1198,268,294]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Figs. 24, 25</figureCitation>
) considerably stout; setae on tibia gradually becoming longer and stouter toward corbicle; corbicle bearing stout spines, with 1 rough, longitudinal line of spines. Tarsi absent from all legs. Abdominal sternites III to VII moderately covered with setae of various length and thickness, especially in lateral part. Genital capsule with medial process on apicoventral margin. Penis (
<figureCitation id="135C2A7BAB216762FD9482D613D0FE62" box="[621,766,422,448]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Figs. 26, 27</figureCitation>
) with strongly sclerotized and pigmented theca; median arm of basal plate shorter than lateral arms (
<figureCitation id="135C2A7BAB216762FD3882BD120EFE45" box="[705,800,461,487]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Fig. 26</figureCitation>
); conjunctival appendage strongly protruding from phallotreme and recurved downwards in rest (
<figureCitation id="135C2A7BAB216762FD1382841263FDAC" box="[746,845,500,526]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Fig. 27</figureCitation>
). Paramere (
<figureCitation id="135C2A7BAB216762FC09828415A5FDAC" box="[1008,1163,500,526]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Figs. 2830</figureCitation>
) somewhat elongate, moderately covered with setae; hypophysis sharpened (
<figureCitation id="135C2A7BAB216762FCE6816A1281FD96" box="[799,943,538,564]" captionStart="FIGURE 22 30" captionStartId="8.[151,255,1497,1521]" captionTargetBox="[168,1420,201,1466]" captionTargetId="figure@8.[151,1436,194,1474]" captionTargetPageId="8" captionText="FIGURE 22 30. Schiodtella formosana. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 24 25, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 26 27, penis, dorsal (26) and lateral (27) views; 28 30, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 22 23, 1.0 mm for 24 25, 0.2 mm for 26 30." httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="7" pageNumber="26">Figs. 2830</figureCitation>
).
</paragraph>
<paragraph id="8BD836FEAB216762FF3C813013A7FDF9" blockId="7.[151,1437,152,758]" box="[197,649,576,603]" pageId="7" pageNumber="26">
<emphasis id="B913EAECAB216762FF3C81301004FDF8" bold="true" box="[197,298,576,602]" pageId="7" pageNumber="26">Female:</emphasis>
— not available in this study.
</paragraph>
</subSubSection>
<subSubSection id="C37D6575AB216762FF3C8117102EFD0A" pageId="7" pageNumber="26" type="materials_examined">
<paragraph id="8BD836FEAB216762FF3C8117102EFD0A" blockId="7.[151,1437,152,758]" pageId="7" pageNumber="26">
<emphasis id="B913EAECAB216762FF3C81171090FD23" bold="true" box="[197,446,615,641]" italics="true" pageId="7" pageNumber="26">Material examined.</emphasis>
23, Chiwei, Chishan, Kaohsiung Hsien,
<collectingCountry id="F370766EAB216762FC3A8118150EFD20" box="[963,1056,616,642]" name="Taiwan" pageId="7" pageNumber="26">Taiwan</collectingCountry>
,
<date id="FFD9103EAB216762FBD48118158FFD20" box="[1069,1185,616,642]" pageId="7" pageNumber="26" value="1934-02-04">4.II.1934</date>
, M. Yanagihara leg. (
<collectionCode id="ED76AE3BAB216762FF6681FE11DAFD0A" box="[159,244,654,680]" pageId="7" pageNumber="26">ELKU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D6575AB216762FF3C81C5129FFD54" pageId="7" pageNumber="26" type="distribution">
<paragraph id="8BD836FEAB216762FF3C81C5129FFD54" blockId="7.[151,1437,152,758]" pageId="7" pageNumber="26">
<emphasis id="B913EAECAB216762FF3C81C5104BFD6D" bold="true" box="[197,357,693,719]" italics="true" pageId="7" pageNumber="26">Distribution.</emphasis>
<collectingCountry id="F370766EAB216762FE9481C510E8FD6D" box="[365,454,693,719]" name="Taiwan" pageId="7" pageNumber="26">Taiwan</collectingCountry>
is the definite locality of this species at present, because previous distributional records are most probably erroneous for mainland
<collectingCountry id="F370766EAB216762FD1181AC1200FD54" box="[744,814,732,758]" name="China" pageId="7" pageNumber="26">China</collectingCountry>
and
<collectingCountry id="F370766EAB216762FC9E81AC1280FD54" box="[871,942,732,758]" name="Japan" pageId="7" pageNumber="26">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF186676AB216762FF6E844B13BDF835" httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="7" pageNumber="26" targetBox="[161,1434,836,1789]" targetPageId="7">
<paragraph id="8BD836FEAB216762FF6E844B13BDF835" blockId="7.[151,1437,1851,1943]" pageId="7" pageNumber="26">
<emphasis id="B913EAECAB216762FF6E844B107BF8F1" bold="true" box="[151,341,1851,1875]" pageId="7" pageNumber="26">FIGURE 1721.</emphasis>
<taxonomicName id="4C674D7DAB216762FEA5844B1379F8F0" box="[348,599,1851,1874]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB216762FEA5844B1379F8F0" box="[348,599,1851,1874]" italics="true" pageId="7" pageNumber="26">Schiodtella formosana</emphasis>
</taxonomicName>
. 17, dorsal aspect; 1819, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna IIIVII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 1820.
</paragraph>
</caption>
<caption id="DF186676AB2E676DFF6E86A91071F9FB" httpUri="https://zenodo.org/record/192022/files/figure.png" pageId="8" pageNumber="27" targetBox="[168,1420,201,1466]" targetPageId="8">
<paragraph id="8BD836FEAB2E676DFF6E86A91071F9FB" blockId="8.[151,1436,1497,1625]" pageId="8" pageNumber="27">
<emphasis id="B913EAECAB2E676DFF6E86A91061FA53" bold="true" box="[151,335,1497,1521]" pageId="8" pageNumber="27">FIGURE 2230.</emphasis>
<taxonomicName id="4C674D7DAB2E676DFEAF86AA1366FA53" box="[342,584,1498,1521]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2E676DFEAF86AA1366FA53" box="[342,584,1498,1521]" italics="true" pageId="8" pageNumber="27">Schiodtella formosana</emphasis>
</taxonomicName>
. 22, left protibia, posteroventral view; 23, left mesotibia, anterior view; 2425, left hind leg, dorsal view of tibia (24) and anterior view of femur and tibia (25); 2627, penis, dorsal (26) and lateral (27) views; 2830, left paramere, ventral (28), dorsal (29) and inner (30) views. Scales: 0.5 mm for 2223, 1.0 mm for 2425, 0.2 mm for 2630.
</paragraph>
</caption>
<subSubSection id="C37D6575AB2E676CFF3C85F712C7FD96" lastPageId="9" lastPageNumber="28" pageId="8" pageNumber="27" type="discussion">
<paragraph id="8BD836FEAB2E676CFF3C85F71389FE62" blockId="8.[151,1437,1671,2006]" lastBlockId="9.[151,1437,152,564]" lastPageId="9" lastPageNumber="28" pageId="8" pageNumber="27">
<emphasis id="B913EAECAB2E676DFF3C85F71016F903" bold="true" box="[197,312,1671,1697]" italics="true" pageId="8" pageNumber="27">Remarks.</emphasis>
The most remarkable characteristic of
<taxonomicName id="4C674D7DAB2E676DFCF885F71286F902" box="[769,936,1671,1696]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2E676DFCF885F71286F902" box="[769,936,1671,1696]" italics="true" pageId="8" pageNumber="27">Sc. formosana</emphasis>
</taxonomicName>
is the absence of tarsi from all legs, which easily distinguishes this species from the other members of
<taxonomicName id="4C674D7DAB2E676DFC8C85DE12D0F965" box="[885,1022,1710,1735]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB2E676DFC8C85DE12D0F965" box="[885,1022,1710,1735]" italics="true" pageId="8" pageNumber="27">Schiodtella</emphasis>
</taxonomicName>
. Regarding the peculiarity of this characteristic for its taxonomic value,
<bibRefCitation id="EFF64B0FAB2E676DFD9685A412C6F94C" author="Takano" box="[623,1000,1748,1774]" pageId="8" pageNumber="27" refString="Takano, S. &amp; Yanagihara, M. (1939) Researches on injurious and beneficial animals of sugarcane. Taiwan Sugar Experimental Station, Extra Reports, 2, 311 pp., 18 pls." type="book" year="1939">Takano and Yanagihara (1939)</bibRefCitation>
appear to have established the new genus
<emphasis id="B913EAECAB2E676DFF1B858B10A2F8B6" box="[226,396,1787,1812]" italics="true" pageId="8" pageNumber="27">Neostibaropus</emphasis>
to accommodate the simultaneously described species
<emphasis id="B913EAECAB2E676DFBE8858B150AF8B6" box="[1041,1060,1787,1812]" italics="true" pageId="8" pageNumber="27">N</emphasis>
.
<taxonomicName id="4C674D7DAB2E676DFBCA858B1592F8B6" box="[1075,1212,1787,1812]" class="Insecta" family="Cydnidae" genus="Neostibaropus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B913EAECAB2E676DFBCA858B1592F8B6" box="[1075,1212,1787,1812]" italics="true" pageId="8" pageNumber="27">formosanus</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C674D7DAB2E676DFB1D858B14A4F8B6" box="[1252,1418,1787,1813]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2E676DFB1D858B142EF8B6" box="[1252,1280,1787,1812]" italics="true" pageId="8" pageNumber="27">Sc</emphasis>
.
<emphasis id="B913EAECAB2E676DFAF4858B14A4F8B6" box="[1293,1418,1787,1812]" italics="true" pageId="8" pageNumber="27">formosana</emphasis>
</taxonomicName>
). Actually, the lack of tarsi is considered a notable feature worthy of establishing a new genus for not only this species but also any other species of Scaptocorini. However,
<taxonomicName id="4C674D7DAB2E676DFC4684391554F8C0" box="[959,1146,1865,1891]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2E676DFC46843912F3F8C0" box="[959,989,1865,1890]" italics="true" pageId="8" pageNumber="27">Sc</emphasis>
.
<emphasis id="B913EAECAB2E676DFC0984391554F8C0" box="[1008,1146,1865,1890]" italics="true" pageId="8" pageNumber="27">formosana</emphasis>
</taxonomicName>
has several important characteristics which are common to other congeners of
<taxonomicName id="4C674D7DAB2E676DFCD384001280F82B" box="[810,942,1904,1929]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB2E676DFCD384001280F82B" box="[810,942,1904,1929]" italics="true" pageId="8" pageNumber="27">Schiodtella</emphasis>
</taxonomicName>
, such as the four-segmented antennae, the peg-bearing tubercles fringed along the margin of the paraclypeus, and the abdomen with a 1+1 trichobothrial formula on the sternites III to VII, as documented by Lis (1999) and
<bibRefCitation id="EFF64B0FAB2E676DFC3684CC1456F874" author="Lis" box="[975,1400,1980,2006]" pageId="8" pageNumber="27" refString="Lis, J. A. &amp; Hohol-Kilinkiewicz, A. (2001) Abdominal trichobothrial pattern and its taxonomic and phylogenetic significance in Cephalocteinae (Hemiptera: Heteroptera: Cydnidae). Annales zoologici (Warszawa), 51 (3), 293 - 297." type="journal article" year="2001">Lis and Hohol-Kilinkiewicz (2001)</bibRefCitation>
. In addition to these characteristics, the penis structure may be sufficiently distinct for this species to be included in
<taxonomicName id="4C674D7DAB2F676CFF4183CF1068FF7A" box="[184,326,191,216]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB2F676CFF4183CF1068FF7A" box="[184,326,191,216]" italics="true" pageId="9" pageNumber="28">Schiodtella</emphasis>
</taxonomicName>
. Every species of this genus possesses the conjunctival appendage on the venter of the endosoma of the penis. The appendage serially varies from externally invisible as in
<taxonomicName id="4C674D7DAB2F676CFB86839514B5FF5D" authority="(Lis 1991b)" baseAuthorityName="Lis" baseAuthorityYear="1991" box="[1151,1435,229,255]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="secunda">
<emphasis id="B913EAECAB2F676CFB86839515B5FF5C" box="[1151,1179,229,254]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFB5383951424FF5C" box="[1194,1290,229,254]" italics="true" pageId="9" pageNumber="28">secunda</emphasis>
(
<bibRefCitation id="EFF64B0FAB2F676CFAE3839514BDFF5D" author="Lis" box="[1306,1427,229,255]" pageId="9" pageNumber="28" refString="Lis, J. A. (1991 b) Schiodtella secunda n. sp., a new species of burrowing bugs from Palaearctic China (Heteroptera: Cydnidae: Scaptocorinae). Genus (Wroclaw), 2 (1), 27 - 31." type="journal article" year="1991" yearSuffix="b">Lis 1991b</bibRefCitation>
)
</taxonomicName>
to large and long as in
<taxonomicName id="4C674D7DAB2F676CFE59827C131FFE87" box="[416,561,268,294]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B913EAECAB2F676CFE59827C1095FE87" box="[416,443,268,293]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFE33827C131FFE87" box="[458,561,268,293]" italics="true" pageId="9" pageNumber="28">japonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2205797AB2F676CFDCE827C13BFFE84" box="[567,657,268,294]" pageId="9" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
through medium-sized as in
<taxonomicName id="4C674D7DAB2F676CFC1C827C1444FE84" authority="Montandon 1897" authorityName="Montandon" authorityYear="1897" box="[997,1386,268,294]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="laevicollis">
<emphasis id="B913EAECAB2F676CFC1C827C152EFE87" box="[997,1024,268,293]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFBF6827C15A6FE87" box="[1039,1160,268,293]" italics="true" pageId="9" pageNumber="28">laevicollis</emphasis>
(
<bibRefCitation id="EFF64B0FAB2F676CFB6F827C144CFE84" author="Montandon" box="[1174,1378,268,294]" pageId="9" pageNumber="28" refString="Montandon, A. L. (1897) Nouvelles especes d'Hemipteres-Heteropteres d'Algerie et de Tunisie. Revue d'Entomologie 16, 97 - 104." type="journal article" year="1897">Montandon 1897</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C674D7DAB2F676CFF6E824210D0FEEE" authority="Breddin 1900" authorityName="Breddin" authorityYear="1900" box="[151,510,306,332]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="subglabra">
<emphasis id="B913EAECAB2F676CFF6E8242119DFEE9" box="[151,179,306,331]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFF3A82421012FEE9" box="[195,316,306,331]" italics="true" pageId="9" pageNumber="28">subglabra</emphasis>
(
<bibRefCitation id="EFF64B0FAB2F676CFEB2824210DBFEEE" author="Breddin" box="[331,501,306,332]" pageId="9" pageNumber="28" refString="Breddin, G. (1900) Hemiptera Sumatrana collecta a dom. Henrico Dohrn, enumerate et descripta. Pars I. Stettiner Entomologische Zeitung, 61, 275 - 336." type="journal article" year="1900">Breddin 1900</bibRefCitation>
)
</taxonomicName>
. The appendage of
<taxonomicName id="4C674D7DAB2F676CFD11824212BDFEE9" box="[744,915,306,332]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2F676CFD118242122AFEE9" box="[744,772,306,331]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFCED824212BDFEE9" box="[788,915,306,331]" italics="true" pageId="9" pageNumber="28">formosana</emphasis>
</taxonomicName>
is large and long, being most similar to
<taxonomicName id="4C674D7DAB2F676CFA808242102FFED0" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B913EAECAB2F676CFA80824214BBFEE9" box="[1401,1429,306,331]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFF6E8229102FFED0" box="[151,257,345,370]" italics="true" pageId="9" pageNumber="28">japonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2205797AB2F676CFEF082291049FED1" box="[265,359,345,371]" pageId="9" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
among the congeners of
<taxonomicName id="4C674D7DAB2F676CFD638229120CFED0" box="[666,802,345,370]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB2F676CFD638229120CFED0" box="[666,802,345,370]" italics="true" pageId="9" pageNumber="28">Schiodtella</emphasis>
</taxonomicName>
. The range of variation in the appendage, then, also allows
<taxonomicName id="4C674D7DAB2F676CFF1282F010BDFE3B" box="[235,403,384,410]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2F676CFF1282F01029FE3B" box="[235,263,384,409]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFEEF82F010BDFE3B" box="[278,403,384,409]" italics="true" pageId="9" pageNumber="28">formosana</emphasis>
</taxonomicName>
to remain in
<taxonomicName id="4C674D7DAB2F676CFDCB82F01398FE3B" box="[562,694,384,409]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB2F676CFDCB82F01398FE3B" box="[562,694,384,409]" italics="true" pageId="9" pageNumber="28">Schiodtella</emphasis>
</taxonomicName>
. Perhaps
<taxonomicName id="4C674D7DAB2F676CFCD382F012FFFE3B" box="[810,977,384,410]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2F676CFCD382F01268FE3B" box="[810,838,384,409]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFCAD82F012FFFE3B" box="[852,977,384,409]" italics="true" pageId="9" pageNumber="28">formosana</emphasis>
</taxonomicName>
is more closely related to
<taxonomicName id="4C674D7DAB2F676CFAF282F014B5FE3B" box="[1291,1435,384,410]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="japonica" status="sp. nov.">
<emphasis id="B913EAECAB2F676CFAF282F01408FE3B" box="[1291,1318,384,409]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFACC82F014B5FE3B" box="[1333,1435,384,409]" italics="true" pageId="9" pageNumber="28">japonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2205797AB2F676CFF6E82D611DFFE62" box="[151,241,422,448]" pageId="9" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
than the other species of
<taxonomicName id="4C674D7DAB2F676CFDE582D7138EFE62" box="[540,672,423,448]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB2F676CFDE582D7138EFE62" box="[540,672,423,448]" italics="true" pageId="9" pageNumber="28">Schiodtella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD836FEAB2F676CFF3C82BD12C7FD96" blockId="9.[151,1437,152,564]" pageId="9" pageNumber="28">
The absence of tarsi from all legs is also observed in the Afrotropical
<emphasis id="B913EAECAB2F676CFBE182BD15D9FE44" box="[1048,1271,461,486]" italics="true" pageId="9" pageNumber="28">Afroropus bocagei</emphasis>
(de
<bibRefCitation id="EFF64B0FAB2F676CFAD782BD11FDFDAC" author="Carvalho" pageId="9" pageNumber="28" refString="De Carvalho, E. L. (1969) Um novo Cydnideo Escaptocorineo Africano (Heteroptera, Cydnidae). Boletim de Sociedade Portuguesa de Ciencias Naturais. 2 e serie, 12, 223 - 226." type="journal article" year="1969">Carvalho 1969</bibRefCitation>
) and the Neotropical
<emphasis id="B913EAECAB2F676CFE28828413E7FDAF" box="[465,713,500,525]" italics="true" pageId="9" pageNumber="28">Atarsocoris giselleae</emphasis>
(=
<emphasis id="B913EAECAB2F676CFD13828412BDFDAF" box="[746,915,500,525]" italics="true" pageId="9" pageNumber="28">A. macroptera</emphasis>
). However,
<taxonomicName id="4C674D7DAB2F676CFBDD828415E5FDAF" box="[1060,1227,500,526]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B913EAECAB2F676CFBDD8284156EFDAF" box="[1060,1088,500,525]" italics="true" pageId="9" pageNumber="28">Sc</emphasis>
.
<emphasis id="B913EAECAB2F676CFBB7828415E5FDAF" box="[1102,1227,500,525]" italics="true" pageId="9" pageNumber="28">formosana</emphasis>
</taxonomicName>
is separable from these two species by the paraclypeus fringed with peg-bearing tubercles.
</paragraph>
</subSubSection>
</treatment>
</document>