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60 KiB
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475 lines
60 KiB
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<mods:title id="499ED0AC55131067B8A5FD882717A6A9">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<treatment id="03CE87D9FFBAFFEDCC8B0362FD05FECE" ID-DOI="http://doi.org/10.5281/zenodo.4428369" ID-GBIF-Taxon="176485718" ID-Zenodo-Dep="4428369" LSID="urn:lsid:zoobank.org:act:FF4FEC13-E55A-4653-AC55-FE453AE90BAB" httpUri="http://treatment.plazi.org/id/03CE87D9FFBAFFEDCC8B0362FD05FECE" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61">
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<taxonomicName id="4C674D4CFFBAFFEFCC8B0362FD04FCB5" authority="Abrams and Harvey" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,740,798,825]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noctigrassator" status="sp. nov.">
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<emphasis id="B913EADDFFBAFFEFCC8B0362FE3EFCB5" bold="true" box="[151,478,798,825]" italics="true" pageId="60" pageNumber="61">Draculoides noctigrassator</emphasis>
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Abrams and Harvey
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,
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<paragraph id="8BD836CFFFBAFFEFCC8B033FFE86FCD1" blockId="60.[151,1325,798,897]" box="[151,358,834,861]" pageId="60" pageNumber="61">
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(
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<figureCitation id="135C2A4AFFBAFFEFCC83033FFEEBFCD1" box="[159,267,835,861]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support <80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="60" pageNumber="61">Figs. 1–8</figureCitation>
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,
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<figureCitation id="135C2A4AFFBAFFEFCD0B033EFEBFFCD1" box="[279,351,834,861]" captionStart-0="FIGURE 34" captionStart-1="FIGURE 35" captionStartId-0="61.[151,250,1546,1570]" captionStartId-1="62.[151,250,1137,1161]" captionTargetBox-0="[152,1435,381,1523]" captionTargetBox-1="[151,1437,345,1113]" captionTargetId-0="figure-107@61.[151,1436,381,1523]" captionTargetId-1="figure-86@62.[151,1437,345,1113]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="FIGURE 34. Draculoides noctigrassator sp. nov., holotype male (WAM T142871):A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." captionText-1="FIGURE 35. Draculoides noctigrassator sp. nov., holotype male (WAM T142871): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. Scale bar A–C: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4416933" figureDoi-1="http://doi.org/10.5281/zenodo.4416937" httpUri-0="https://zenodo.org/record/4416933/files/figure.png" httpUri-1="https://zenodo.org/record/4416937/files/figure.png" pageId="60" pageNumber="61">34–35</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD836CFFFBAFFEFCC8B031BFACDFC0D" blockId="60.[151,1325,798,897]" box="[151,1325,870,897]" pageId="60" pageNumber="61">
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Zoobank Code: http://zoobank.org/NomenclaturalActs/
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<uuid id="FFC10C1AFFBAFFEFCF15031AFACDFC0D" box="[777,1325,870,897]" pageId="60" pageNumber="61">FF4FEC13-E55A-4653-AC55-FE453AE90BAB</uuid>
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</paragraph>
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<paragraph id="8BD836CFFFBAFFEFCC8B03D7FCBDFC4F" blockId="60.[151,861,939,963]" box="[151,861,939,963]" pageId="60" pageNumber="61">
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<taxonomicName id="4C674D4CFFBAFFEFCC8B03D7FE9DFC4F" authority="SCH" authorityName="SCH" box="[151,381,939,963]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
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<emphasis id="B913EADDFFBAFFEFCC8B03D7FEA1FC4F" box="[151,321,939,963]" italics="true" pageId="60" pageNumber="61">Paradraculoides</emphasis>
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SCH
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</taxonomicName>
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102:
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<bibRefCitation id="EFF64B3EFFBAFFEFCDB703D7FD8EFC4F" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[427,622,939,963]" pageId="60" pageNumber="61" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
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Abrams
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<emphasis id="B913EADDFFBAFFEFCE1E03D0FDCEFC4F" box="[514,558,939,963]" italics="true" pageId="60" pageNumber="61">et al</emphasis>
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. 2019
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</bibRefCitation>
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MPE 106532: 8, fig. 2.
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</paragraph>
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</subSubSection>
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<emphasis id="B913EADDFFBAFFEFCC8B0396FE60FB88" bold="true" box="[151,384,1002,1028]" pageId="60" pageNumber="61">Material examined.</emphasis>
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<materialsCitation id="3B0F3C92FFBAFFEFCD9A0397FE3AFBC1" ID-GBIF-Occurrence="3013449377" collectingDate="2012-10-07" collectionCode="WAM" collectorName="J. W. Quartermaine & G. B. Pearson" country="Australia" elevation="430" latitude="-22.013721" location="Dragon" longLatPrecision="1" longitude="116.55586" pageId="60" pageNumber="61" specimenCode="T142871, ITS2" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B913EADDFFBAFFEFCD9A0397FDD1FB89" box="[390,561,1003,1029]" italics="true" pageId="60" pageNumber="61">
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<typeStatus id="54DC886DFFBAFFEFCD9A0397FE0DFB89" box="[390,493,1003,1029]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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male.
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:
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|
||
<collectingRegion id="49A3F82DFFBAFFEFCEFE0397FC49FB89" box="[738,937,1003,1029]" country="Australia" name="Western Australia" pageId="60" pageNumber="61">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D61FD46FFBAFFEFCFA40397FC03FB89" box="[952,995,1003,1029]" pageId="60" pageNumber="61" type="male">1 ♂</specimenCount>
|
||
,
|
||
<location id="8EB86014FFBAFFEFCFF30397FBA6FB89" LSID="urn:lsid:plazi:treatment:03CE87D9FFBAFFEDCC8B0362FD05FECE:8EB86014FFBAFFEFCFF30397FBA6FB89" box="[1007,1094,1003,1029]" country="Australia" latitude="-22.013721" longLatPrecision="1" longitude="116.55586" name="Dragon" pageId="60" pageNumber="61" stateProvince="Western Australia">Dragon</location>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFBAFFEFC86A0397FB5DFB89" box="[1142,1213,1003,1029]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="60" pageNumber="61" unit="km" value="44.0">44 km</quantity>
|
||
SSE. of Pannawon- ica,
|
||
<geoCoordinate id="EE535008FFBAFFEFCCD40473FE84FBA5" box="[200,356,1039,1065]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="49.4" value="-22.013721">22°00’49.4”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFCD6E0473FDFCFBA5" box="[370,540,1039,1065]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="21.1" value="116.55586">116°33’21.1”E</geoCoordinate>
|
||
,
|
||
<quantity id="4C9F9B2AFFBAFFEFCE360473FD95FBA5" box="[554,629,1039,1065]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="60" pageNumber="61" unit="m" value="430.0">
|
||
<elevation id="004AD1FCFFBAFFEFCE360473FD95FBA5" box="[554,629,1039,1065]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="60" pageNumber="61" unit="m" value="430.0">430 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFD9100FFFBAFFEFCE9F0473FCA6FBA5" box="[643,838,1039,1065]" pageId="60" pageNumber="61" value="2012-10-07">
|
||
<collectingDate id="EF9DE9E7FFBAFFEFCE9F0473FCA6FBA5" box="[643,838,1039,1065]" pageId="60" pageNumber="61" value="2012-10-07">07 October 2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925319FFBAFFEFCF480473FBCAFBA5" box="[852,1066,1039,1065]" pageId="60" pageNumber="61">J.W. Quartermaine</collectorName>
|
||
,
|
||
<collectorName id="26925319FFBAFFEFC8240473FB2FFBA5" box="[1080,1231,1039,1065]" pageId="60" pageNumber="61">G.B. Pearson</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFFBAFFEFC8C30473FAC6FBA5" box="[1247,1318,1039,1065]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="60" pageNumber="61" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFBAFFEFC9320473FA74FBA5" box="[1326,1428,1039,1065]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="60" pageNumber="61">T142871</specimenCode>
|
||
) (DNA:12S, 18S, 28S,
|
||
<specimenCode id="DBC19EB4FFBAFFEFCD8E044FFE2EFBC1" box="[402,462,1075,1101]" collectionCode="ITS" pageId="60" pageNumber="61">ITS2</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBAFFEFCCDB042BFC00FB51" blockId="60.[151,1437,1002,2037]" pageId="60" pageNumber="61">
|
||
<materialsCitation id="3B0F3C92FFBAFFEFCCDB042BFA63FB19" ID-GBIF-Occurrence="3013449446" collectionCode="WAM" country="Australia" latitude="-22.013556" location="Pannawonica" longLatPrecision="1" longitude="116.56456" municipality="Dragon" pageId="60" pageNumber="61" specimenCode="T129673, ITS2" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<emphasis id="B913EADDFFBAFFEFCCDB042BFED9FBFC" box="[199,313,1111,1136]" italics="true" pageId="60" pageNumber="61">
|
||
<typeStatus id="54DC886DFFBAFFEFCCDB042BFED9FBFC" box="[199,313,1111,1136]" pageId="60" pageNumber="61" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="F370765FFFBAFFEFCD56042BFE0DFBFD" box="[330,493,1111,1137]" name="Australia" pageId="60" pageNumber="61">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFFBAFFEFCDE1042BFD2AFBFD" box="[509,714,1111,1137]" italics="true" pageId="60" pageNumber="61">
|
||
<collectingRegion id="49A3F82DFFBAFFEFCDE1042BFD2AFBFD" box="[509,714,1111,1137]" country="Australia" name="Western Australia" pageId="60" pageNumber="61">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D61FD46FFBAFFEFCEC2042BFCF0FBFD" box="[734,784,1111,1137]" pageId="60" pageNumber="61" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FFBAFFEFCF3D042BFC98FBFD" box="[801,888,1111,1137]" pageId="60" pageNumber="61">Dragon</collectingMunicipality>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFBAFFEFCFA8042AFBE1FBFD" box="[948,1025,1110,1137]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="60" pageNumber="61" unit="km" value="45.0">45 km</quantity>
|
||
S. of
|
||
<location id="8EB86014FFBAFFEFC84F042BFB0BFBFD" LSID="urn:lsid:plazi:treatment:03CE87D9FFBAFFEDCC8B0362FD05FECE:8EB86014FFBAFFEFC84F042BFB0BFBFD" box="[1107,1259,1111,1137]" country="Australia" latitude="-22.013556" longLatPrecision="1" longitude="116.56456" municipality="Dragon" name="Pannawonica" pageId="60" pageNumber="61" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFC8E0042BFA78FBFD" box="[1276,1432,1111,1137]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="48.8" value="-22.013556">22°00’48.8”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFCC8B0406FEA2FB19" box="[151,322,1146,1173]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="52.4" value="116.56456">116°33’52.4”E</geoCoordinate>
|
||
, 06
|
||
<collectorName id="26925319FFBAFFEFCD6C0407FE2CFB19" box="[368,460,1147,1173]" pageId="60" pageNumber="61">October</collectorName>
|
||
, scrape, 75 metres,
|
||
<collectorName id="26925319FFBAFFEFCEBB0407FCDEFB19" box="[679,830,1147,1173]" pageId="60" pageNumber="61">G.B. Pearson</collectorName>
|
||
,
|
||
<collectorName id="26925319FFBAFFEFCF550407FBFAFB19" box="[841,1050,1147,1173]" pageId="60" pageNumber="61">J.W. Quartermaine</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFFBAFFEFC8340407FB8FFB19" box="[1064,1135,1147,1173]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="60" pageNumber="61" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFBAFFEFC8690407FB3BFB1A" box="[1141,1243,1147,1174]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="60" pageNumber="61">T129673</specimenCode>
|
||
) (DNA:
|
||
<specimenCode id="DBC19EB4FFBAFFEFC9220407FA9BFB19" box="[1342,1403,1147,1173]" collectionCode="ITS" pageId="60" pageNumber="61">ITS2</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FFBAFFEFC9920407FF24FB51" ID-GBIF-Occurrence="3013449392" collectingDate="2012-07-19" collectionCode="R" country="Australia" elevation="71" latitude="-22.0125" longLatPrecision="15" longitude="116.558754" pageId="60" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<specimenCount id="9D61FD46FFBAFFEFC9920407FF4EFB35" pageId="60" pageNumber="61" type="male">1 ♂</specimenCount>
|
||
, Dragon, ca.
|
||
<quantity id="4C9F9B2AFFBAFFEFCD5804E3FE6BFB35" box="[324,395,1183,1209]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="60" pageNumber="61" unit="km" value="44.0">44 km</quantity>
|
||
SSE. of Pannawonica,
|
||
<geoCoordinate id="EE535008FFBAFFEFCE8F04E2FCFAFB35" box="[659,794,1182,1209]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="60" pageNumber="61" precision="15" seconds="45" value="-22.0125">22°00’45”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFCF3A04E2FC30FB35" box="[806,976,1182,1209]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="31.5" value="116.558754">116°33’31.5”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFFBAFFEFCFC004E3FB91FB35" box="[988,1137,1183,1209]" pageId="60" pageNumber="61" value="2012-07-19">
|
||
<collectingDate id="EF9DE9E7FFBAFFEFCFC004E3FB91FB35" box="[988,1137,1183,1209]" pageId="60" pageNumber="61" value="2012-07-19">19 July 2012</collectingDate>
|
||
</date>
|
||
, troglofauna scrape, -
|
||
<quantity id="4C9F9B2AFFBAFFEFC97E04E3FA78FB35" box="[1378,1432,1183,1209]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="60" pageNumber="61" unit="m" value="71.0">
|
||
<elevation id="004AD1FCFFBAFFEFC97E04E3FA78FB35" box="[1378,1432,1183,1209]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="60" pageNumber="61" unit="m" value="71.0">71m</elevation>
|
||
</quantity>
|
||
, S.
|
||
<collectionCode id="ED76AE0AFFBAFFEFCCB004BFFF24FB51" box="[172,196,1219,1245]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="60" pageNumber="61">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0F3C92FFBAFFEFCCD204BFFC3BFB51" ID-GBIF-Occurrence="3013449371" box="[206,987,1219,1245]" collectionCode="WAM" collectorName="Bennett, S. G" country="Australia" pageId="60" pageNumber="61" specimenCode="T142893, ITS2" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<collectorName id="26925319FFBAFFEFCCD204BFFE81FB51" box="[206,353,1219,1245]" pageId="60" pageNumber="61">Bennett, S.G</collectorName>
|
||
. Pearson (
|
||
<collectionCode id="ED76AE0AFFBAFFEFCDCF04BFFDFAFB51" box="[467,538,1219,1245]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="60" pageNumber="61" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFBAFFEFCE3C04BFFD67FB51" box="[544,647,1219,1245]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="60" pageNumber="61">T142893</specimenCode>
|
||
) (DNA: 12S, 18S, 28S,
|
||
<specimenCode id="DBC19EB4FFBAFFEFCF8B04BFFC34FB51" box="[919,980,1219,1245]" collectionCode="ITS" pageId="60" pageNumber="61">ITS2</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBAFFEFCCDB049BFE21FAE1" blockId="60.[151,1437,1002,2037]" pageId="60" pageNumber="61">
|
||
<emphasis id="B913EADDFFBAFFEFCCDB049BFE9CFA8D" box="[199,380,1255,1281]" italics="true" pageId="60" pageNumber="61">Other material.</emphasis>
|
||
<materialsCitation id="3B0F3C92FFBAFFEFCD9B049BFD2FFAA9" ID-GBIF-Occurrence="3013449379" collectingDate="2012-07-19" collectionCode="R" country="Australia" latitude="-22.0125" location="Pannawonica" longLatPrecision="1" longitude="116.558754" municipality="Dragon" pageId="60" pageNumber="61" specimenCount="1" specimenCount-juvenile="1" stateProvince="Western Australia">
|
||
<collectingCountry id="F370765FFFBAFFEFCD9B049BFDCAFA8D" box="[391,554,1255,1281]" name="Australia" pageId="60" pageNumber="61">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFFBAFFEFCE27049BFCE9FA8D" box="[571,777,1255,1281]" italics="true" pageId="60" pageNumber="61">
|
||
<collectingRegion id="49A3F82DFFBAFFEFCE27049BFCE9FA8D" box="[571,777,1255,1281]" country="Australia" name="Western Australia" pageId="60" pageNumber="61">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D61FD46FFBAFFEFCF02049BFC76FA8D" box="[798,918,1255,1281]" pageId="60" pageNumber="61" type="juvenile">1 juvenile</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FFBAFFEFCFB4049BFC1FFA8D" box="[936,1023,1255,1281]" pageId="60" pageNumber="61">Dragon</collectingMunicipality>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFBAFFEFC821049BFB6AFA8D" box="[1085,1162,1255,1281]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="60" pageNumber="61" unit="km" value="44.0">44 km</quantity>
|
||
SSE. of
|
||
<location id="8EB86014FFBAFFEFC91C049BFA78FA8D" LSID="urn:lsid:plazi:treatment:03CE87D9FFBAFFEDCC8B0362FD05FECE:8EB86014FFBAFFEFC91C049BFA78FA8D" box="[1280,1432,1255,1281]" country="Australia" latitude="-22.0125" longLatPrecision="1" longitude="116.558754" municipality="Dragon" name="Pannawonica" pageId="60" pageNumber="61" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFCC8B0576FED4FAA9" box="[151,308,1290,1317]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="45.0" value="-22.0125">22°00’45.0”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFCD5E0576FE0CFAA9" box="[322,492,1290,1317]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="31.5" value="116.558754">116°33’31.5”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFFBAFFEFCDE70577FD75FAA9" box="[507,661,1291,1317]" pageId="60" pageNumber="61" value="2012-07-19">
|
||
<collectingDate id="EF9DE9E7FFBAFFEFCDE70577FD75FAA9" box="[507,661,1291,1317]" pageId="60" pageNumber="61" value="2012-07-19">19 July 2012</collectingDate>
|
||
</date>
|
||
, S.
|
||
<collectionCode id="ED76AE0AFFBAFFEFCEAB0577FD2FFAA9" box="[695,719,1291,1317]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="60" pageNumber="61">R</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0F3C92FFBAFFEFCEC10577FB50FAA9" ID-GBIF-Occurrence="3013449349" box="[733,1200,1290,1317]" collectionCode="WAM" collectorName="Bennett, G. B" country="Australia" pageId="60" pageNumber="61" specimenCode="T144530" specimenCount="1" stateProvince="Western Australia">
|
||
<collectorName id="26925319FFBAFFEFCEC10577FC96FAA9" box="[733,886,1291,1317]" pageId="60" pageNumber="61">Bennett, G.B</collectorName>
|
||
. Pearson (
|
||
<collectionCode id="ED76AE0AFFBAFFEFCFF20577FBD5FAA9" box="[1006,1077,1291,1317]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="60" pageNumber="61" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFBAFFEFC8220576FB48FAA9" box="[1086,1192,1290,1317]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="60" pageNumber="61">T144530</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FFBAFFEFC8A20577FE5DFAE1" ID-GBIF-Occurrence="3013449401" collectingDate="2012-10-07" collectionCode="WAM" collectorName="J. W. Quartermaine & G. B. Pearson" country="Australia" elevation="430" latitude="-22.013721" location="Pannawonica" longLatPrecision="1" longitude="116.55586" municipality="Dragon" pageId="60" pageNumber="61" specimenCode="T144531" specimenCount="1" specimenCount-juvenile="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D61FD46FFBAFFEFC8A20577FAD3FAA9" box="[1214,1331,1291,1317]" pageId="60" pageNumber="61" type="juvenile">1 juvenile</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FFBAFFEFC95D0577FA78FAA9" box="[1345,1432,1291,1317]" pageId="60" pageNumber="61">Dragon</collectingMunicipality>
|
||
, ca.
|
||
<quantity id="4C9F9B2AFFBAFFEFCCA30553FEE7FAC5" box="[191,263,1327,1353]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="60" pageNumber="61" unit="km" value="44.0">44 km</quantity>
|
||
SSE. of
|
||
<location id="8EB86014FFBAFFEFCD710553FDE5FAC5" LSID="urn:lsid:plazi:treatment:03CE87D9FFBAFFEDCC8B0362FD05FECE:8EB86014FFBAFFEFCD710553FDE5FAC5" box="[365,517,1327,1353]" country="Australia" latitude="-22.013721" longLatPrecision="1" longitude="116.55586" municipality="Dragon" name="Pannawonica" pageId="60" pageNumber="61" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFCE0D0553FD4DFAC5" box="[529,685,1327,1353]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="49.4" value="-22.013721">22°00’49.4”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FFBAFFEFCEA50553FC84FAC5" box="[697,868,1327,1353]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="21.1" value="116.55586">116°33’21.1”E</geoCoordinate>
|
||
,
|
||
<quantity id="4C9F9B2AFFBAFFEFCF6C0553FC5AFAC5" box="[880,954,1327,1353]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="60" pageNumber="61" unit="m" value="430.0">
|
||
<elevation id="004AD1FCFFBAFFEFCF6C0553FC5AFAC5" box="[880,954,1327,1353]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="60" pageNumber="61" unit="m" value="430.0">430 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFD9100FFFBAFFEFCFDB0553FB9AFAC5" box="[967,1146,1327,1353]" pageId="60" pageNumber="61" value="2012-10-07">
|
||
<collectingDate id="EF9DE9E7FFBAFFEFCFDB0553FB9AFAC5" box="[967,1146,1327,1353]" pageId="60" pageNumber="61" value="2012-10-07">7 October 2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925319FFBAFFEFC89A0553FABCFAC5" box="[1158,1372,1327,1353]" pageId="60" pageNumber="61">J.W. Quartermaine</collectorName>
|
||
,
|
||
<collectorName id="26925319FFBAFFEFC97B0553FF10FAE1" pageId="60" pageNumber="61">G.B. Pearson</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFFBAFFEFCCE2052FFEA5FAE1" box="[254,325,1363,1389]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="60" pageNumber="61" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FFBAFFEFCD57052EFE55FAE1" box="[331,437,1362,1389]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="60" pageNumber="61">T144531</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFBAFFEFCCDB050AFD62F901" pageId="60" pageNumber="61" type="diagnosis">
|
||
<paragraph id="8BD836CFFFBAFFEFCCDB050AFD62F901" blockId="60.[151,1437,1002,2037]" pageId="60" pageNumber="61">
|
||
<emphasis id="B913EADDFFBAFFEFCCDB050AFEA2FA1C" bold="true" box="[199,322,1398,1424]" pageId="60" pageNumber="61">Diagnosis.</emphasis>
|
||
The shape of the male flagellum of
|
||
<taxonomicName id="4C674D4CFFBAFFEFCEFC050BFBF5FA1C" box="[736,1045,1399,1425]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noctigrassator">
|
||
<emphasis id="B913EADDFFBAFFEFCEFC050BFBF5FA1C" box="[736,1045,1399,1425]" italics="true" pageId="60" pageNumber="61">Draculoides noctigrassator</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName id="4C674D4CFFBAFFEFC930050BFA75FA1D" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[1324,1429,1399,1425]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B913EADDFFBAFFEFC930050BFA75FA1D" box="[1324,1429,1399,1425]" italics="true" pageId="60" pageNumber="61">D. affinis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFBAFFEFCC8B05E7FEA3FA39" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,323,1435,1461]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFFBAFFEFCC8B05E7FEA3FA39" box="[151,323,1435,1461]" italics="true" pageId="60" pageNumber="61">D. anachoretus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFBAFFEFCD4E05E7FE25FA39" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[338,453,1435,1461]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="bythius">
|
||
<emphasis id="B913EADDFFBAFFEFCD4E05E7FE25FA39" box="[338,453,1435,1461]" italics="true" pageId="60" pageNumber="61">D. bythius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFBAFFEFCDC805E7FD8BFA39" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[468,619,1435,1461]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="cochranus">
|
||
<emphasis id="B913EADDFFBAFFEFCDC805E7FD8BFA39" box="[468,619,1435,1461]" italics="true" pageId="60" pageNumber="61">D. cochranus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B913EADDFFBAFFEFCE6505E7FC43FA39" box="[633,931,1435,1461]" italics="true" pageId="60" pageNumber="61">
|
||
<taxonomicName id="4C674D4CFFBAFFEFCE6505E7FD16FA38" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[633,758,1435,1460]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="eremius">D. eremius</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFBAFFEFCF1E05E7FC43FA39" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[770,931,1435,1461]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="gnophicola">D. gnophicola</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C674D4CFFBAFFEFCFC605E7FBAEFA39" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[986,1102,1435,1461]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="kryptus">
|
||
<emphasis id="B913EADDFFBAFFEFCFC605E7FBAEFA39" box="[986,1102,1435,1461]" italics="true" pageId="60" pageNumber="61">D. kryptus</emphasis>
|
||
</taxonomicName>
|
||
, especially in the presence of a broad base. It differs from
|
||
<taxonomicName id="4C674D4CFFBAFFEFCDC505C3FDA2FA55" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[473,578,1471,1497]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B913EADDFFBAFFEFCDC505C3FDA2FA55" box="[473,578,1471,1497]" italics="true" pageId="60" pageNumber="61">D. affinis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B913EADDFFBAFFEFCE4C05C3FB2CFA55" box="[592,1228,1471,1497]" italics="true" pageId="60" pageNumber="61">
|
||
<taxonomicName id="4C674D4CFFBAFFEFCE4C05C3FD1CFA55" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[592,764,1471,1497]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="anachoretus">D. anachoretus</taxonomicName>
|
||
<taxonomicName id="4C674D4CFFBAFFEFCF1805C3FC9AFA55" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[772,890,1471,1497]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="bythius">D. bythius</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFBAFFEFCF9A05C3FBFFFA55" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[902,1055,1471,1497]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="cochranus">D. cochranus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFFBAFFEFC83005C3FB2CFA55" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1068,1228,1471,1497]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="gnophicola">D. gnophicola</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C674D4CFFBAFFEFC91805C3FA98FA55" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1284,1400,1471,1497]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="kryptus">
|
||
<emphasis id="B913EADDFFBAFFEFC91805C3FA98FA55" box="[1284,1400,1471,1497]" italics="true" pageId="60" pageNumber="61">D. kryptus</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the dorsal margin of the male flagellum, which is quite straight in
|
||
<taxonomicName id="4C674D4CFFBAFFEFC80E059FFB37FA70" box="[1042,1239,1507,1532]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noctigrassator">
|
||
<emphasis id="B913EADDFFBAFFEFC80E059FFB37FA70" box="[1042,1239,1507,1532]" italics="true" pageId="60" pageNumber="61">D. noctigrassator</emphasis>
|
||
</taxonomicName>
|
||
and more curved and posteriorly tapering in the aforementioned species. It differs from
|
||
<taxonomicName id="4C674D4CFFBAFFEFCFB2067BFBC9F9AC" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[942,1065,1543,1568]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="eremius">
|
||
<emphasis id="B913EADDFFBAFFEFCFB2067BFBC9F9AC" box="[942,1065,1543,1568]" italics="true" pageId="60" pageNumber="61">D. eremius</emphasis>
|
||
</taxonomicName>
|
||
by the position of dm4, which is anterior to dl
|
||
<quantity id="4C9F9B2AFFBAFFEFCD340657FEB2F9C9" box="[296,338,1579,1605]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="60" pageNumber="61" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="4C674D4CFFBAFFEFCD440657FDFDF9C8" box="[344,541,1579,1604]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noctigrassator">
|
||
<emphasis id="B913EADDFFBAFFEFCD440657FDFDF9C8" box="[344,541,1579,1604]" italics="true" pageId="60" pageNumber="61">D. noctigrassator</emphasis>
|
||
</taxonomicName>
|
||
and level with dl
|
||
<quantity id="4C9F9B2AFFBAFFEFCEC30657FCE8F9C9" box="[735,776,1579,1605]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="60" pageNumber="61" unit="in" value="3.0">3 in</quantity>
|
||
<emphasis id="B913EADDFFBAFFEFCF130657FC70F9C8" box="[783,912,1579,1604]" italics="true" pageId="60" pageNumber="61">
|
||
<taxonomicName id="4C674D4CFFBAFFEFCF130657FC6BF9C8" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[783,907,1579,1604]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="eremius">D. eremius</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Females are unknown.
|
||
<taxonomicName id="4C674D4CFFBAFFEFC8840657FF31F9E4" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="noctigrassator">
|
||
<emphasis id="B913EADDFFBAFFEFC8840657FF31F9E4" italics="true" pageId="60" pageNumber="61">Draculoides noctigrassator</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFFBAFFEFCE580633FD2EF9E5" box="[580,718,1615,1641]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFFBAFFEFCE580633FD2EF9E5" box="[580,718,1615,1641]" italics="true" pageId="60" pageNumber="61">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at COI, 12S and ITS2 by the 50bp mini-barcodes shown in
|
||
<figureCitation id="135C2A4AFFBAFFEFCDB7060FFDF5F901" box="[427,533,1651,1677]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="60" pageNumber="61">Figures 3</figureCitation>
|
||
,
|
||
<figureCitation id="135C2A4AFFBAFFEFCE3E060EFDD2F901" box="[546,562,1650,1677]" captionStart="FIGURE 5" captionStartId="20.[151,250,1216,1240]" captionTargetBox="[158,1424,183,1189]" captionTargetId="figure-20@20.[151,1436,181,1192]" captionTargetPageId="20" captionText="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416863" httpUri="https://zenodo.org/record/4416863/files/figure.png" pageId="60" pageNumber="61">5</figureCitation>
|
||
, and 7.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFBAFFEDCCDB06EAFBF1FF3D" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61" type="description">
|
||
<paragraph id="8BD836CFFFBAFFEFCCDB06EAFB2EF93D" blockId="60.[151,1437,1002,2037]" box="[199,1230,1686,1713]" pageId="60" pageNumber="61">
|
||
<emphasis id="B913EADDFFBAFFEFCCDB06EAFE5BF93C" bold="true" box="[199,443,1686,1712]" pageId="60" pageNumber="61">Description (adults).</emphasis>
|
||
<emphasis id="B913EADDFFBAFFEFCDDE06EBFDF1F93D" box="[450,529,1687,1713]" italics="true" pageId="60" pageNumber="61">Colour</emphasis>
|
||
. Yellow-brown; propeltidium and pedipalps somewhat darker.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBAFFEFCCDB06C7FD84F891" blockId="60.[151,1437,1002,2037]" pageId="60" pageNumber="61">
|
||
<emphasis id="B913EADDFFBAFFEFCCDB06C7FE8FF959" box="[199,367,1723,1749]" italics="true" pageId="60" pageNumber="61">Cephalothorax</emphasis>
|
||
. Propeltidium with 2 apical setae on anterior process and 2 + 2 + 2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 12-13, setae (including 2 sternapophysial setae); posterior sternum triangular with 6 setae.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBAFFEFCCDB075BFED0F879" blockId="60.[151,1437,1002,2037]" pageId="60" pageNumber="61">
|
||
<emphasis id="B913EADDFFBAFFEFCCDB075BFED5F8CD" box="[199,309,1831,1857]" italics="true" pageId="60" pageNumber="61">Chelicera</emphasis>
|
||
. Fixed finger with 2 large teeth plus 6 smaller teeth between these; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 composed of 6 setae; G3 composed of 5 setae; internal face of chelicera with 4 short whip-like setae (G4); brush at base of fixed finger composed of 7 setae (G5A), each densely pilose in distal half and G5B composed of 9 setae; G6 with one seta; G7 composed of 4 setae. Movable finger serrula composed of 17 long lamellae, blunt guard tooth present subdistally; 1 large accessory tooth present.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBBFFEECCDB00EBFA7DFF3D" blockId="61.[151,1437,151,358]" box="[199,1437,151,177]" pageId="61" pageNumber="62">
|
||
<emphasis id="B913EADDFFBBFFEECCDB00EBFECCFF3D" box="[199,300,151,177]" italics="true" pageId="61" pageNumber="62">Pedipalp</emphasis>
|
||
. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBBFFEECC8B00C7FA7CFE91" blockId="61.[151,1437,151,358]" pageId="61" pageNumber="62">
|
||
8 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; 0.50 × length of tarsus.
|
||
<emphasis id="B913EADDFFBBFFEECCDB00A3FEE4FF74" box="[199,260,223,248]" italics="true" pageId="61" pageNumber="62">Legs.</emphasis>
|
||
Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
|
||
<emphasis id="B913EADDFFBBFFEECCDB017FFED1FE91" box="[199,305,259,285]" italics="true" pageId="61" pageNumber="62">Abdomen</emphasis>
|
||
. Chaetotaxy of tergite I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II: 3 macrosetae
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBBFFEECC8B015BFA7CFECD" blockId="61.[151,1437,151,358]" box="[151,1436,295,322]" pageId="61" pageNumber="62">+ 6 microsetae (microsetae in column), tergites III–IX: 2: 2: 2: 2: 2: 2; segment XII with very small dorsal process</paragraph>
|
||
<paragraph id="8BD836CFFFBBFFEECC8B0137FF1FFEEA" blockId="61.[151,1437,151,358]" box="[151,255,331,358]" pageId="61" pageNumber="62">
|
||
(
|
||
<specimenCount id="9D61FD46FFBBFFEECCBC0137FF55FEE9" box="[160,181,331,357]" pageId="61" pageNumber="62" type="male">♂</specimenCount>
|
||
only).
|
||
</paragraph>
|
||
<caption id="DF186647FFBBFFEECC8B0676FBD2F9E6" ID-DOI="http://doi.org/10.5281/zenodo.4416933" ID-Zenodo-Dep="4416933" httpUri="https://zenodo.org/record/4416933/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1546,1570]" targetBox="[152,1435,381,1523]" targetPageId="61">
|
||
<paragraph id="8BD836CFFFBBFFEECC8B0676FBD2F9E6" blockId="61.[151,1436,1546,1643]" pageId="61" pageNumber="62">
|
||
<emphasis id="B913EADDFFBBFFEECC8B0676FEFFF9AE" bold="true" box="[151,287,1546,1570]" pageId="61" pageNumber="62">FIGURE 34.</emphasis>
|
||
<taxonomicName id="4C674D4CFFBBFFEECD380677FDDBF9AE" authority="Abrams & Huey & Hillyer & Didham & Harvey, 2020" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[292,571,1547,1571]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="noctigrassator" status="sp. nov.">
|
||
<emphasis id="B913EADDFFBBFFEECD380677FDDBF9AE" box="[292,571,1547,1571]" italics="true" pageId="61" pageNumber="62">Draculoides noctigrassator</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EADDFFBBFFEECE5C0677FD6FF9AF" bold="true" box="[576,655,1547,1571]" pageId="61" pageNumber="62">
|
||
<taxonomicNameLabel id="A22057A6FFBBFFEECE5C0677FD6FF9AF" box="[576,655,1547,1571]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (WAM T142871): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836CFFFBBFFEECCDB06E5FAC2F893" blockId="61.[151,1437,1689,2040]" pageId="61" pageNumber="62">
|
||
<emphasis id="B913EADDFFBBFFEECCDB06E5FEDAF93F" box="[199,314,1689,1715]" italics="true" pageId="61" pageNumber="62">Flagellum</emphasis>
|
||
. Male: Dorsoventrally compressed (
|
||
<figureCitation id="135C2A4AFFBBFFEECEC506E5FCD5F93F" box="[729,821,1689,1715]" captionStart="FIGURE 34" captionStartId="61.[151,250,1546,1570]" captionTargetBox="[152,1435,381,1523]" captionTargetId="figure-107@61.[151,1436,381,1523]" captionTargetPageId="61" captionText="FIGURE 34. Draculoides noctigrassator sp. nov., holotype male (WAM T142871):A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." figureDoi="http://doi.org/10.5281/zenodo.4416933" httpUri="https://zenodo.org/record/4416933/files/figure.png" pageId="61" pageNumber="62">Figs. 34</figureCitation>
|
||
D–F, 35A–C); 1.9 × longer than broad; seta dm1 situated dorso-medially, closer to anterior margin; seta dm4 set back from posterior margin, anterior to dl3; dl1slightly posterior to vl1; dl3 close to posterior margin; vm2 situated slightly anterior to vm1; vm5 situated in line with vl1; left vm3 slightly anterior to right vm3; four pairs of microsetae between vl1 and dl3. Female: unknown.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBBFFEECCDB0756FADDF8EB" blockId="61.[151,1437,1689,2040]" pageId="61" pageNumber="62">
|
||
<emphasis id="B913EADDFFBBFFEECCDB0756FE6DF8CF" box="[199,397,1833,1859]" italics="true" pageId="61" pageNumber="62">Dimensions (mm)</emphasis>
|
||
.
|
||
<typeStatus id="54DC886DFFBBFFEECD860755FDE2F8CF" box="[410,514,1833,1859]" pageId="61" pageNumber="62" type="holotype">Holotype</typeStatus>
|
||
male (WAM T142871): Body length 3.46. Propeltidium 1.15/0.62. Chelicera 0.73. Flagellum 0.37/0.19. Pedipalp: trochanter 0.54, femur 0.54, patella 0.46, tibia 0.42, tarsus 0.23, claw 0.13.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFBBFFEECCDB070EFDCCF87B" blockId="61.[151,1437,1689,2040]" pageId="61" pageNumber="62">
|
||
<emphasis id="B913EADDFFBBFFEECCDB070EFECFF807" box="[199,303,1906,1931]" italics="true" pageId="61" pageNumber="62">Variation</emphasis>
|
||
. Body length (males) 3.06–4.04 (n = 3). Propeltidium setae: single seta following 2 on apical process and one extra pair of setae. Tergite II chaetotaxy: 2 setae instead of 3 as in
|
||
<typeStatus id="54DC886DFFBBFFEECFF207E9FBB4F823" box="[1006,1108,1941,1967]" pageId="61" pageNumber="62" type="holotype">holotype</typeStatus>
|
||
. Anterior sternum number of setae 12-13, including sternapophysial setae (n=3). Position of vm
|
||
<quantity id="4C9F9B2AFFBBFFEECF9007C5FC57F85F" box="[908,951,1977,2004]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="61" pageNumber="62" unit="in" value="5.0">5 in</quantity>
|
||
relation to vl1 varies, sometimes closer to vl1, sometimes vm5 is closer to vl2.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFB8FFEDCCDB00EAFBF1FF3D" blockId="62.[151,1437,150,322]" box="[199,1041,150,177]" pageId="62" pageNumber="63">
|
||
<emphasis id="B913EADDFFB8FFEDCCDB00EAFE88FF3D" bold="true" box="[199,360,150,177]" pageId="62" pageNumber="63">Other names.</emphasis>
|
||
WAM SCH102 (
|
||
<bibRefCitation id="EFF64B3EFFB8FFEDCE3500EBFCEDFF3D" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[553,781,151,177]" pageId="62" pageNumber="63" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
|
||
Abrams
|
||
<emphasis id="B913EADDFFB8FFEDCE9700EBFD5CFF3D" box="[651,700,151,177]" italics="true" pageId="62" pageNumber="63">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
), Bennelongia sp. B37.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFB8FFEDCCDB00C7FC5CFF75" pageId="62" pageNumber="63" type="discussion">
|
||
<paragraph id="8BD836CFFFB8FFEDCCDB00C7FC5CFF75" blockId="62.[151,1437,150,322]" pageId="62" pageNumber="63">
|
||
<emphasis id="B913EADDFFB8FFEDCCDB00C7FEDBFF59" bold="true" box="[199,315,187,213]" pageId="62" pageNumber="63">Remarks.</emphasis>
|
||
<taxonomicName id="4C674D4CFFB8FFEDCD5E00C7FD97FF58" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[322,631,187,213]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="noctigrassator">
|
||
<emphasis id="B913EADDFFB8FFEDCD5E00C7FD97FF58" box="[322,631,187,213]" italics="true" pageId="62" pageNumber="63">Draculoides noctigrassator</emphasis>
|
||
</taxonomicName>
|
||
is known from several locations within the ‘Dragon’ deposit within the Bungaroo South (Buckland project) area of
|
||
<collectingRegion id="49A3F82DFFB8FFEDCE9E00A3FCAAFF75" box="[642,842,223,249]" country="Australia" name="Western Australia" pageId="62" pageNumber="63">Western Australia</collectingRegion>
|
||
(
|
||
<figureCitation id="135C2A4AFFB8FFEDCF4500A3FC4EFF75" box="[857,942,223,249]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="62" pageNumber="63">Fig. 1E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFB8FFEDCCDB017FFD05FECE" pageId="62" pageNumber="63" type="etymology">
|
||
<paragraph id="8BD836CFFFB8FFEDCCDB017FFD05FECE" blockId="62.[151,1437,150,322]" pageId="62" pageNumber="63">
|
||
<emphasis id="B913EADDFFB8FFEDCCDB017FFEAEFE91" bold="true" box="[199,334,259,285]" pageId="62" pageNumber="63">Etymology.</emphasis>
|
||
The specific epithet is a verb referring to the occurrence of this species in permanently dark habitats (Latin,
|
||
<emphasis id="B913EADDFFB8FFEDCD0B015BFEBAFECC" box="[279,346,295,320]" italics="true" pageId="62" pageNumber="63">noctis</emphasis>
|
||
, night;
|
||
<emphasis id="B913EADDFFB8FFEDCDAD015BFDFCFECC" box="[433,540,295,320]" italics="true" pageId="62" pageNumber="63">grassatus</emphasis>
|
||
, go about, loiter).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |