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<document id="8DEAE1D1C18B8FA583881C8A4BE4911F" ID-CLB-Dataset="9009" ID-DOI="10.11646/zootaxa.4864.1.1" ID-GBIF-Dataset="710afcac-f353-49bf-a397-3c71ba1508be" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416849" ID-ZooBank="A5F51A7F-83DA-4C77-A85C-0FCF8A400CF2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609802491702" checkinUser="plazi" docAuthor="Abrams, Kym M., Huey, Joel A., Hillyer, Mia J., Didham, Raphael K. &amp; Harvey, Mark S." docDate="2020" docId="03CE87D9FFA0FFFBCC8B05EAFC56FC29" docLanguage="en" docName="zootaxa.4864.1.1.pdf" docOrigin="Zootaxa 4864 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Draculoides christopherleei Abrams and Harvey 2020, n. sp." docType="treatment" docUuid="5AADC536-FD7C-4792-B6B6-CB6BED0D9176" docUuidSource="ZooBank" docVersion="8" lastPageNumber="41" masterDocId="FFF7FFA1FF86FFD3CC1C007CFFE0FF8C" masterDocTitle="A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara" masterLastPageNumber="75" masterPageNumber="1" pageNumber="39" updateTime="1698852268950" updateUser="ExternalLinkService">
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<mods:title id="9F8296D8D74CE9C8DD59545D6B11B4EF">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<emphasis id="B913EADDFFA0FFF5CC8B05EAFE06FA3D" bold="true" box="[151,486,1430,1457]" italics="true" pageId="38" pageNumber="39">Draculoides christopherleei</emphasis>
Abrams and Harvey
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(
<figureCitation id="135C2A4AFFA0FFF5CC8305C7FEEBFA59" box="[159,267,1467,1493]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support &lt;80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="38" pageNumber="39">Figs. 18</figureCitation>
,
<figureCitation id="135C2A4AFFA0FFF5CD0B05C7FEBEFA59" box="[279,350,1467,1493]" captionStart-0="FIGURE 18" captionStart-1="FIGURE 19" captionStartId-0="39.[151,250,1687,1711]" captionStartId-1="40.[151,250,1677,1701]" captionTargetBox-0="[151,1436,705,1664]" captionTargetBox-1="[156,1424,957,1647]" captionTargetId-0="figure-248@39.[151,1436,705,1664]" captionTargetId-1="figure-341@40.[156,1432,957,1654]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionText-0="FIGURE 18. Draculoides christopherleei sp. nov., holotype male (WAM T129676): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 1 mm and also applies to E and F." captionText-1="FIGURE 19. Draculoides christopherleei sp. nov., holotype male (WAM T129676): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. Scale bar: AC: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4416895" figureDoi-1="http://doi.org/10.5281/zenodo.4416897" httpUri-0="https://zenodo.org/record/4416895/files/figure.png" httpUri-1="https://zenodo.org/record/4416897/files/figure.png" pageId="38" pageNumber="39">1819</figureCitation>
)
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Zoobank Code: http://zoobank.org/NomenclaturalActs/
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<taxonomicName id="4C674D4CFFA0FFF5CC8B065FFE9BF9B7" authority="SCH" authorityName="SCH" box="[151,379,1571,1595]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFA0FFF5CC8B065FFEA1F9B7" box="[151,321,1571,1595]" italics="true" pageId="38" pageNumber="39">Paradraculoides</emphasis>
SCH
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080:
<bibRefCitation id="EFF64B3EFFA0FFF5CDB7065FFD8EF9B7" author="Abrams, K. M. &amp; Huey, J. A. &amp; Hillyer, M. J. &amp; Humphreys, W. F. &amp; Didham, R. K. &amp; Harvey, M. S." box="[427,622,1571,1595]" pageId="38" pageNumber="39" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. &amp; Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
Abrams
<emphasis id="B913EADDFFA0FFF5CE1E0658FDCEF9B7" box="[514,558,1571,1595]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
. 2019
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MPE 106532: 8, fig. 2.
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<emphasis id="B913EADDFFA0FFF5CC8B0615FE63F90F" bold="true" box="[151,387,1641,1667]" pageId="38" pageNumber="39">Material examined.</emphasis>
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male.
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,
<geoCoordinate id="EE535008FFA0FFF5CD5C06F1FE0BF924" box="[320,491,1677,1704]" degrees="116" direction="east" minutes="30" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="35.7" value="116.50992">116°3035.7”E</geoCoordinate>
,
<date id="FFD9100FFFA0FFF5CDEB06F1FD4BF92B" box="[503,683,1677,1703]" pageId="38" pageNumber="39" value="2012-10-09">
<collectingDate id="EF9DE9E7FFA0FFF5CDEB06F1FD4BF92B" box="[503,683,1677,1703]" pageId="38" pageNumber="39" value="2012-10-09">9 October 2012</collectingDate>
</date>
, scrape,
<quantity id="4C9F9B2AFFA0FFF5CF1206F1FCABF924" box="[782,843,1677,1704]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="38" pageNumber="39" unit="m" value="37.0">
<elevation id="004AD1FCFFA0FFF5CF1206F1FCABF924" box="[782,843,1677,1704]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="38" pageNumber="39" unit="m" value="37.0">37 m</elevation>
</quantity>
,
<collectorName id="26925319FFA0FFF5CF4406F1FC11F924" box="[856,1009,1677,1704]" pageId="38" pageNumber="39">G.B. Pearson</collectorName>
,
<collectorName id="26925319FFA0FFF5CFE106F2FB30F92B" box="[1021,1232,1677,1704]" pageId="38" pageNumber="39">J.W. Quartermaine</collectorName>
(
<collectionCode id="ED76AE0AFFA0FFF5C8C306F1FAC6F92B" box="[1247,1318,1677,1703]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="38" pageNumber="39" type="Museum">WAM</collectionCode>
<specimenCode id="DBC19EB4FFA0FFF5C93206F1FA75F924" box="[1326,1429,1677,1704]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="38" pageNumber="39">T129676</specimenCode>
) (DNA:
<collectionCode id="ED76AE0AFFA0FFF5CCF006CDFEFCF947" box="[236,284,1713,1739]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="38" pageNumber="39" type="Herbarium">COI</collectionCode>
from
<bibRefCitation id="EFF64B3EFFA0FFF5CD7D06CEFDD4F947" author="Harms, D. &amp; Curran, M. K. &amp; Klesser, R. &amp; Finston, T. L. &amp; Halse, S. A." box="[353,564,1713,1740]" pageId="38" pageNumber="39" pagination="355 - 367" refId="ref46587" refString="Harms, D., Curran, M. K., Klesser, R., Finston, T. L. &amp; Halse, S. A. (2018) Speciation patterns in complex subterranean environments: a case study using short-tailed whipscorpions (Schizomida: Hubbardiidae). Biological Journal of the Linnean Society, 125 (2), 355 - 367. https: // doi. org / 10.1093 / biolinnean / bly 102" type="journal article" year="2018">
Harms
<emphasis id="B913EADDFFA0FFF5CDAE06CEFE03F947" box="[434,483,1713,1739]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
., 2018
</bibRefCitation>
:
<accessionNumber id="9434AB2CFFA0FFF5CE2306CDFD25F940" box="[575,709,1713,1740]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH620848" pageId="38" pageNumber="39">
<collectionCode id="ED76AE0AFFA0FFF5CE2306CDFD8EF940" box="[575,622,1713,1740]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="38" pageNumber="39">MH</collectionCode>
620848
</accessionNumber>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD836CFFFA0FFF4CCDB06A9FCA7FF59" blockId="38.[151,1437,1641,2028]" lastBlockId="39.[151,1437,150,681]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="B913EADDFFA0FFF5CCDB06A9FE9AF963" box="[199,378,1749,1775]" italics="true" pageId="38" pageNumber="39">Other material.</emphasis>
<materialsCitation id="3B0F3C92FFA0FFF5CD9E06A9FE75F8D0" ID-GBIF-Occurrence="3013449397" accessionNumber="MH620857" collectingDate="2012-07-20" collectingDateMax="2012-09-11" collectingDateMin="2012-07-20" collectionCode="WAM, COI, MH" country="Australia" latitude="-21.991806" location="Pannawonica" longLatPrecision="1" longitude="116.50753" pageId="38" pageNumber="39" specimenCode="T142837" specimenCount="1" stateProvince="Western Australia">
<collectingCountry id="F370765FFFA0FFF5CD9E06A9FDC5F963" box="[386,549,1749,1775]" name="Australia" pageId="38" pageNumber="39">AUSTRALIA</collectingCountry>
:
<emphasis id="B913EADDFFA0FFF5CE2806AAFD1FF963" box="[564,767,1749,1775]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="49A3F82DFFA0FFF5CE2806AAFD1FF963" box="[564,767,1749,1775]" country="Australia" name="Western Australia" pageId="38" pageNumber="39">Western Australia</collectingRegion>
</emphasis>
: sex indeterminate, partial specimen, cephalothorax only, “West Pit”, ca.
<quantity id="4C9F9B2AFFA0FFF5CD540685FE71F89F" box="[328,401,1785,1811]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="38" pageNumber="39" unit="km" value="40.0">40 km</quantity>
SSE. of
<location id="8EB86014FFA0FFF5CDE70685FD73F89F" LSID="urn:lsid:plazi:treatment:03CE87D9FFA0FFFBCC8B05EAFC56FC29:8EB86014FFA0FFF5CDE70685FD73F89F" box="[507,659,1785,1811]" country="Australia" latitude="-21.991806" longLatPrecision="1" longitude="116.50753" name="Pannawonica" pageId="38" pageNumber="39" stateProvince="Western Australia">Pannawonica</location>
,
<geoCoordinate id="EE535008FFA0FFF5CEBC0685FCDCF898" box="[672,828,1785,1812]" degrees="21" direction="south" minutes="59" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="30.5" value="-21.991806">21°5930.5”S</geoCoordinate>
,
<geoCoordinate id="EE535008FFA0FFF5CF560685FC14F89F" box="[842,1012,1785,1811]" degrees="116" direction="east" minutes="30" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="27.1" value="116.50753">116°3027.1”E</geoCoordinate>
,
<date id="FFD9100FFFA0FFF5C81E0685FAA3F89F" box="[1026,1347,1785,1811]" pageId="38" pageNumber="39" value="2012-07-20" valueMax="2012-09-11" valueMin="2012-07-20">
<collectingDate id="EF9DE9E7FFA0FFF5C81E0685FAA3F89F" box="[1026,1347,1785,1811]" pageId="38" pageNumber="39" value="2012-07-20" valueMax="2012-09-11" valueMin="2012-07-20">20 July11 September 2012</collectingDate>
</date>
,
<collectingMethod id="52264ED8FFA0FFF5C94C0685FEEBF8B4" pageId="38" pageNumber="39">troglofauna trap</collectingMethod>
,
<collectorName id="26925319FFA0FFF5CD040762FE0DF8BB" box="[280,493,1821,1848]" pageId="38" pageNumber="39">J.W. Quartermaine</collectorName>
,
<collectorName id="26925319FFA0FFF5CDE50761FD6FF8B4" box="[505,655,1821,1848]" pageId="38" pageNumber="39">G.B. Pearson</collectorName>
(
<collectorName id="26925319FFA0FFF5CE820761FCD0F8BB" box="[670,816,1821,1847]" pageId="38" pageNumber="39">Bennelongia</collectorName>
, BH045) (
<collectionCode id="ED76AE0AFFA0FFF5CFB90761FC0CF8BB" box="[933,1004,1821,1847]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="38" pageNumber="39" type="Museum">WAM</collectionCode>
<specimenCode id="DBC19EB4FFA0FFF5CFEF0761FBBAF8BB" box="[1011,1114,1821,1847]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="38" pageNumber="39">T142837</specimenCode>
) (DNA:
<collectionCode id="ED76AE0AFFA0FFF5C8A30761FB0FF8BB" box="[1215,1263,1821,1847]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="38" pageNumber="39" type="Herbarium">COI</collectionCode>
from
<bibRefCitation id="EFF64B3EFFA0FFF5C92A0762FF1DF8D7" author="Harms, D. &amp; Curran, M. K. &amp; Klesser, R. &amp; Finston, T. L. &amp; Halse, S. A." pageId="38" pageNumber="39" pagination="355 - 367" refId="ref46587" refString="Harms, D., Curran, M. K., Klesser, R., Finston, T. L. &amp; Halse, S. A. (2018) Speciation patterns in complex subterranean environments: a case study using short-tailed whipscorpions (Schizomida: Hubbardiidae). Biological Journal of the Linnean Society, 125 (2), 355 - 367. https: // doi. org / 10.1093 / biolinnean / bly 102" type="journal article" year="2018">
Harms
<emphasis id="B913EADDFFA0FFF5C9940762FF4DF8D7" italics="true" pageId="38" pageNumber="39">et al</emphasis>
., 2018
</bibRefCitation>
:
<accessionNumber id="9434AB2CFFA0FFF5CD1A073DFE6CF8D0" box="[262,396,1857,1884]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH620857" pageId="38" pageNumber="39">
<collectionCode id="ED76AE0AFFA0FFF5CD1A073DFED5F8D0" box="[262,309,1857,1884]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="38" pageNumber="39">MH</collectionCode>
620857
</accessionNumber>
)
</materialsCitation>
;
<materialsCitation id="3B0F3C92FFA0FFF5CD82073DFE4DF82F" ID-GBIF-Occurrence="3013449347" collectingDate="2012-06-22" collectionCode="R, WAM, COI" country="Australia" latitude="-22.013334" location="Pannawonica" longLatPrecision="1" longitude="116.53125" pageId="38" pageNumber="39" specimenCode="T129677, ITS2" specimenCount="1" stateProvince="Western Australia">
1 sex indeterminate, partial specimen, cephalothorax only, “East Pit”, ca.
<quantity id="4C9F9B2AFFA0FFF5C8D6073DFAEFF8D7" box="[1226,1295,1857,1884]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="38" pageNumber="39" unit="km" value="45.0">45 km</quantity>
S. of
<location id="8EB86014FFA0FFF5C952073EFF0BF8F3" LSID="urn:lsid:plazi:treatment:03CE87D9FFA0FFFBCC8B05EAFC56FC29:8EB86014FFA0FFF5C952073EFF0BF8F3" country="Australia" latitude="-22.013334" longLatPrecision="1" longitude="116.53125" name="Pannawonica" pageId="38" pageNumber="39" stateProvince="Western Australia">Pannawonica</location>
,
<geoCoordinate id="EE535008FFA0FFF5CCE90719FE72F8F3" box="[245,402,1893,1919]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="48.0" value="-22.013334">22°0048.0”S</geoCoordinate>
,
<geoCoordinate id="EE535008FFA0FFF5CD800719FDA7F80C" box="[412,583,1893,1920]" degrees="116" direction="east" minutes="31" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="52.5" value="116.53125">116°3152.5”E</geoCoordinate>
,
<date id="FFD9100FFFA0FFF5CE4E0719FD09F8F3" box="[594,745,1893,1920]" pageId="38" pageNumber="39" value="2012-06-22">
<collectingDate id="EF9DE9E7FFA0FFF5CE4E0719FD09F8F3" box="[594,745,1893,1920]" pageId="38" pageNumber="39" value="2012-06-22">22 June 2012</collectingDate>
</date>
,
<collectorName id="26925319FFA0FFF5CEEF0719FCA2F8F3" box="[755,834,1893,1919]" pageId="38" pageNumber="39">Scrape</collectorName>
, metres,
<collectorName id="26925319FFA0FFF5CFB90719FBDCF80C" box="[933,1084,1893,1920]" pageId="38" pageNumber="39">G.B. Pearson</collectorName>
, S.
<collectionCode id="ED76AE0AFFA0FFF5C8470719FB93F8F3" box="[1115,1139,1893,1919]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="38" pageNumber="39">R</collectionCode>
.
<collectorName id="26925319FFA0FFF5C861071AFB34F80C" box="[1149,1236,1894,1920]" pageId="38" pageNumber="39">Bennett</collectorName>
(
<collectionCode id="ED76AE0AFFA0FFF5C8FD0719FAC8F8F3" box="[1249,1320,1893,1919]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="38" pageNumber="39" type="Museum">WAM</collectionCode>
<specimenCode id="DBC19EB4FFA0FFF5C9320719FA74F80C" box="[1326,1428,1893,1920]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="38" pageNumber="39">T129677</specimenCode>
) (DNA:
<collectionCode id="ED76AE0AFFA0FFF5CCF207F5FEC1F82F" box="[238,289,1929,1955]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="38" pageNumber="39" type="Herbarium">COI</collectionCode>
, 18S,
<specimenCode id="DBC19EB4FFA0FFF5CD7507F5FE45F82F" box="[361,421,1929,1955]" collectionCode="ITS" pageId="38" pageNumber="39">ITS2</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B0F3C92FFA0FFF5CDA607F5FE8CF860" ID-GBIF-Occurrence="3013449376" accessionNumber="MH620858" collectingDate="2012-10-07" collectionCode="WAM, COI, MH" collectorName="J. W. Quartermaine &amp; G. B. Pearson" country="Australia" elevation="16" latitude="-22.009445" location="Pannawonica" longLatPrecision="21" longitude="116.51945" pageId="38" pageNumber="39" specimenCode="T142838" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount id="9D61FD46FFA0FFF5CDA607F5FE08F828" box="[442,488,1929,1956]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
, “East Pit”, ca.
<quantity id="4C9F9B2AFFA0FFF5CE8307F5FD07F82F" box="[671,743,1929,1955]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="38" pageNumber="39" unit="km" value="42.0">42 km</quantity>
SSE. of
<location id="8EB86014FFA0FFF5CF4C07F5FC08F82F" LSID="urn:lsid:plazi:treatment:03CE87D9FFA0FFFBCC8B05EAFC56FC29:8EB86014FFA0FFF5CF4C07F5FC08F82F" box="[848,1000,1929,1955]" country="Australia" latitude="-22.009445" longLatPrecision="21" longitude="116.51945" name="Pannawonica" pageId="38" pageNumber="39" stateProvince="Western Australia">Pannawonica</location>
,
<geoCoordinate id="EE535008FFA0FFF5CFE907F5FB9BF82F" box="[1013,1147,1929,1955]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="34" value="-22.009445">22°0034”S</geoCoordinate>
,
<geoCoordinate id="EE535008FFA0FFF5C89507F5FAFEF82F" box="[1161,1310,1929,1955]" degrees="116" direction="east" minutes="31" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="10" value="116.51945">116°3110”E</geoCoordinate>
,
<date id="FFD9100FFFA0FFF5C93707F6FF32F84B" pageId="38" pageNumber="39" value="2012-10-07">
<collectingDate id="EF9DE9E7FFA0FFF5C93707F6FF32F84B" pageId="38" pageNumber="39" value="2012-10-07">7 October 2012</collectingDate>
</date>
, troglofauna scrape, -
<quantity id="4C9F9B2AFFA0FFF5CDDE07D1FE17F84B" box="[450,503,1965,1991]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="m" value="16.0">
<elevation id="004AD1FCFFA0FFF5CDDE07D1FE17F84B" box="[450,503,1965,1991]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="m" value="16.0">16m</elevation>
</quantity>
,
<collectorName id="26925319FFA0FFF5CE1E07D2FD36F84B" box="[514,726,1965,1992]" pageId="38" pageNumber="39">J.W. Quartermaine</collectorName>
,
<collectorName id="26925319FFA0FFF5CEFD07D1FC95F844" box="[737,885,1965,1992]" pageId="38" pageNumber="39">G.B. Pearson</collectorName>
(
<collectionCode id="ED76AE0AFFA0FFF5CF9F07D1FC2AF84B" box="[899,970,1965,1991]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="38" pageNumber="39" type="Museum">WAM</collectionCode>
<specimenCode id="DBC19EB4FFA0FFF5CFD307D1FBD6F84B" box="[975,1078,1965,1991]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="38" pageNumber="39">T142838</specimenCode>
) (DNA:
<collectionCode id="ED76AE0AFFA0FFF5C88507D1FB29F84B" box="[1177,1225,1965,1991]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="38" pageNumber="39" type="Herbarium">COI</collectionCode>
from
<bibRefCitation id="EFF64B3EFFA0FFF5C91007D2FF33F867" author="Harms, D. &amp; Curran, M. K. &amp; Klesser, R. &amp; Finston, T. L. &amp; Halse, S. A." pageId="38" pageNumber="39" pagination="355 - 367" refId="ref46587" refString="Harms, D., Curran, M. K., Klesser, R., Finston, T. L. &amp; Halse, S. A. (2018) Speciation patterns in complex subterranean environments: a case study using short-tailed whipscorpions (Schizomida: Hubbardiidae). Biological Journal of the Linnean Society, 125 (2), 355 - 367. https: // doi. org / 10.1093 / biolinnean / bly 102" type="journal article" year="2018">
Harms
<emphasis id="B913EADDFFA0FFF5C94207D2FA6EF84B" box="[1374,1422,1965,1991]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
., 2018
</bibRefCitation>
:
<accessionNumber id="9434AB2CFFA0FFF5CCC107ADFE83F860" box="[221,355,2001,2028]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH620858" pageId="38" pageNumber="39">
<collectionCode id="ED76AE0AFFA0FFF5CCC107ADFEECF860" box="[221,268,2001,2028]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="38" pageNumber="39">MH</collectionCode>
620858
</accessionNumber>
)
</materialsCitation>
;
<materialsCitation id="3B0F3C92FFA0FFF4CD6A07ADFEE6FF59" ID-GBIF-Occurrence="3013449307" collectingDate="2012-06-22" collectionCode="R, WAM" collectorName="Bennett, G. B &amp; Pearson" country="Australia" elevation="8" lastPageId="39" lastPageNumber="40" latitude="-22.012362" location="East Pit" longLatPrecision="1" longitude="116.523285" pageId="38" pageNumber="39" specimenCode="T142839" specimenCount="1" stateProvince="Western Australia">
1 sex indeterminate, partial specimen, cephalothorax only, “
<location id="8EB86014FFA0FFF5C81207ADFB8BF867" LSID="urn:lsid:plazi:treatment:03CE87D9FFA0FFFBCC8B05EAFC56FC29:8EB86014FFA0FFF5C81207ADFB8BF867" box="[1038,1131,2001,2027]" country="Australia" latitude="-22.012362" longLatPrecision="1" longitude="116.523285" name="East Pit" pageId="38" pageNumber="39" stateProvince="Western Australia">East Pit</location>
”, ca.
<quantity id="4C9F9B2AFFA0FFF5C8BB07ADFB0DF867" box="[1191,1261,2001,2027]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="38" pageNumber="39" unit="km" value="42.0">42 km</quantity>
SSE. of Panna- wonica,
<geoCoordinate id="EE535008FFA1FFF4CCEB00EAFE73FF3D" box="[247,403,150,177]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="39" pageNumber="40" precision="1" seconds="44.5" value="-22.012362">22°0044.5”S</geoCoordinate>
,
<geoCoordinate id="EE535008FFA1FFF4CD8300EBFDA9FF3D" box="[415,585,151,177]" degrees="116" direction="east" minutes="31" orientation="longitude" pageId="39" pageNumber="40" precision="1" seconds="23.8" value="116.523285">116°3123.8”E</geoCoordinate>
,
<date id="FFD9100FFFA1FFF4CE4900EBFD0FFF3D" box="[597,751,151,177]" pageId="39" pageNumber="40" value="2012-06-22">
<collectingDate id="EF9DE9E7FFA1FFF4CE4900EBFD0FFF3D" box="[597,751,151,177]" pageId="39" pageNumber="40" value="2012-06-22">22 June 2012</collectingDate>
</date>
, troglofauna scrape, -
<quantity id="4C9F9B2AFFA1FFF4CFFD00EBFBE8FF3D" box="[993,1032,151,177]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="40" unit="m" value="8.0">
<elevation id="004AD1FCFFA1FFF4CFFD00EBFBE8FF3D" box="[993,1032,151,177]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="40" unit="m" value="8.0">8m</elevation>
</quantity>
, S.
<collectionCode id="ED76AE0AFFA1FFF4C83500EBFBA1FF3D" box="[1065,1089,151,177]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="39" pageNumber="40">R</collectionCode>
.
<collectorName id="26925319FFA1FFF4C85000EBFB03FF3D" box="[1100,1251,151,177]" pageId="39" pageNumber="40">Bennett, G.B</collectorName>
.
<collectorName id="26925319FFA1FFF4C8F200EBFAA7FF3D" box="[1262,1351,151,177]" pageId="39" pageNumber="40">Pearson</collectorName>
(
<collectionCode id="ED76AE0AFFA1FFF4C94A00EBFA7DFF3D" box="[1366,1437,151,177]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="39" pageNumber="40" type="Museum">WAM</collectionCode>
<specimenCode id="DBC19EB4FFA1FFF4CC8B00C7FF1DFF59" box="[151,253,187,213]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="40">T142839</specimenCode>
)
</materialsCitation>
<materialsCitation id="3B0F3C92FFA1FFF4CD1100C7FCA3FF59" ID-GBIF-Occurrence="3013449453" accessionNumber="MH620859" box="[269,835,186,213]" collectingDate="2012-06-22" collectionCode="COI, MH" collectorName="Bennett, G. B &amp; Pearson" country="Australia" elevation="8" isTailEnumeration="true" latitude="-22.012362" location="East Pit" longLatPrecision="1" longitude="116.523285" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Western Australia">
(DNA:
<collectionCode id="ED76AE0AFFA1FFF4CD7D00C7FE71FF59" box="[353,401,187,213]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="39" pageNumber="40" type="Herbarium">COI</collectionCode>
from
<bibRefCitation id="EFF64B3EFFA1FFF4CDCA00C7FD4AFF59" author="Harms, D. &amp; Curran, M. K. &amp; Klesser, R. &amp; Finston, T. L. &amp; Halse, S. A." box="[470,682,187,213]" pageId="39" pageNumber="40" pagination="355 - 367" refId="ref46587" refString="Harms, D., Curran, M. K., Klesser, R., Finston, T. L. &amp; Halse, S. A. (2018) Speciation patterns in complex subterranean environments: a case study using short-tailed whipscorpions (Schizomida: Hubbardiidae). Biological Journal of the Linnean Society, 125 (2), 355 - 367. https: // doi. org / 10.1093 / biolinnean / bly 102" type="journal article" year="2018">
Harms
<emphasis id="B913EADDFFA1FFF4CE3400C7FDB9FF59" box="[552,601,187,213]" italics="true" pageId="39" pageNumber="40">et al</emphasis>
., 2018
</bibRefCitation>
:
<accessionNumber id="9434AB2CFFA1FFF4CEA800C6FCDBFF59" box="[692,827,186,213]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH620859" pageId="39" pageNumber="40">
<collectionCode id="ED76AE0AFFA1FFF4CEA800C6FD03FF59" box="[692,739,186,213]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="39" pageNumber="40">MH</collectionCode>
620859
</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFA1FFF4CCDB00A2FC7AFD95" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BD836CFFFA1FFF4CCDB00A2FC7AFD95" blockId="39.[151,1437,150,681]" pageId="39" pageNumber="40">
<emphasis id="B913EADDFFA1FFF4CCDB00A2FEA2FF74" bold="true" box="[199,322,222,248]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
Females are unknown. The shape of the male flagellum of
<taxonomicName id="4C674D4CFFA1FFF4CFD300A3FAE9FF75" authorityName="Abrams and Harvey" authorityYear="2020" box="[975,1289,223,249]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFA1FFF4CFD300A3FAE9FF75" box="[975,1289,223,249]" italics="true" pageId="39" pageNumber="40">Draculoides christopherleei</emphasis>
</taxonomicName>
is distinctive due to its small lateral lobes, very broad stalk and the absence of dl1.
<taxonomicName id="4C674D4CFFA1FFF4CF93017FFBB5FE90" authorityName="Harvey" authorityYear="1992" box="[911,1109,259,285]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vinei">
<emphasis id="B913EADDFFA1FFF4CF93017FFBB5FE90" box="[911,1109,259,285]" italics="true" pageId="39" pageNumber="40">Draculoides vinei</emphasis>
</taxonomicName>
is the only other
<taxonomicName id="4C674D4CFFA1FFF4C90E017FFA7CFE91" box="[1298,1436,259,285]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFA1FFF4C90E017FFA7CFE91" box="[1298,1436,259,285]" italics="true" pageId="39" pageNumber="40">Draculoides</emphasis>
</taxonomicName>
species lacking dl1 but clearly differs from
<taxonomicName id="4C674D4CFFA1FFF4CE98015BFC20FECD" authorityName="Abrams and Harvey" authorityYear="2020" box="[644,960,295,321]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFA1FFF4CE98015BFC20FECD" box="[644,960,295,321]" italics="true" pageId="39" pageNumber="40">Draculoides christopherleei</emphasis>
</taxonomicName>
in the shape of the male flagellum, which is dorsolaterally compressed in
<taxonomicName id="4C674D4CFFA1FFF4CE1C0137FD29FEE8" authorityName="Harvey" authorityYear="1992" box="[512,713,331,357]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vinei">
<emphasis id="B913EADDFFA1FFF4CE1C0137FD29FEE8" box="[512,713,331,357]" italics="true" pageId="39" pageNumber="40">Draculoides vinei</emphasis>
</taxonomicName>
and dorsoventrally compressed in
<taxonomicName id="4C674D4CFFA1FFF4C8450137FA76FEE9" authorityName="Abrams and Harvey" authorityYear="2020" box="[1113,1430,331,357]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFA1FFF4C8450137FA76FEE9" box="[1113,1430,331,357]" italics="true" pageId="39" pageNumber="40">Draculoides christopherleei</emphasis>
</taxonomicName>
.
<taxonomicName id="4C674D4CFFA1FFF4CC8B0113FE33FE05" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,467,367,393]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFA1FFF4CC8B0113FE33FE05" box="[151,467,367,393]" italics="true" pageId="39" pageNumber="40">Draculoides christopherleei</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFA1FFF4CF540113FC32FE05" box="[840,978,367,393]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFA1FFF4CF540113FC32FE05" box="[840,978,367,393]" italics="true" pageId="39" pageNumber="40">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at COI and ITS2 by the 50bp mini-barcodes shown in
<figureCitation id="135C2A4AFFA1FFF4CE6501EFFD02FE21" box="[633,738,403,429]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="39" pageNumber="40">Figures 3</figureCitation>
and
<figureCitation id="135C2A4AFFA1FFF4CF0601EEFCCAFE21" box="[794,810,402,429]" captionStart="FIGURE 5" captionStartId="20.[151,250,1216,1240]" captionTargetBox="[158,1424,183,1189]" captionTargetId="figure-20@20.[151,1436,181,1192]" captionTargetPageId="20" captionText="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416863" httpUri="https://zenodo.org/record/4416863/files/figure.png" pageId="39" pageNumber="40">5</figureCitation>
.
<taxonomicName id="4C674D4CFFA1FFF4CF2A01EFFB92FE21" authorityName="Abrams and Harvey" authorityYear="2020" box="[822,1138,403,429]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFA1FFF4CF2A01EFFB92FE21" box="[822,1138,403,429]" italics="true" pageId="39" pageNumber="40">Draculoides christopherleei</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFA1FFF4CCC401CBFE82FE5D" box="[216,354,439,465]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFA1FFF4CCC401CBFE82FE5D" box="[216,354,439,465]" italics="true" pageId="39" pageNumber="40">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at ITS2 (
<figureCitation id="135C2A4AFFA1FFF4CF0E01CBFCB5FE5D" box="[786,853,439,465]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="39" pageNumber="40">Fig. 7</figureCitation>
) except for
<taxonomicName id="4C674D4CFFA1FFF4CFC201CBFB6AFE5D" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[990,1162,439,465]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="B913EADDFFA1FFF4CFC201CBFB6AFE5D" box="[990,1162,439,465]" italics="true" pageId="39" pageNumber="40">D. anachoretus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4C88501CBFAECFE5D" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1177,1292,439,465]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFA1FFF4C88501CBFAECFE5D" box="[1177,1292,439,465]" italics="true" pageId="39" pageNumber="40">D. bythius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4C90701CBFA76FE5C" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[1307,1430,439,464]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFA1FFF4C90701CBFA76FE5C" box="[1307,1430,439,464]" italics="true" pageId="39" pageNumber="40">D. eremius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4CC8B01A7FED7FE79" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,311,475,501]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFA1FFF4CC8B01A7FED7FE79" box="[151,311,475,501]" italics="true" pageId="39" pageNumber="40">D. gnophicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4CD5901A7FE59FE79" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[325,441,475,501]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFA1FFF4CD5901A7FE59FE79" box="[325,441,475,501]" italics="true" pageId="39" pageNumber="40">D. kryptus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4CDDB01A7FD7BFE79" authorityName="Abrams and Harvey" authorityYear="2020" box="[455,667,475,501]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFA1FFF4CDDB01A7FD7BFE79" box="[455,667,475,501]" italics="true" pageId="39" pageNumber="40">D. mckechnieorum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4CEB501A7FCAEFE79" authorityName="Abrams and Harvey" authorityYear="2020" box="[681,846,475,501]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFA1FFF4CEB501A7FCAEFE79" box="[681,846,475,501]" italics="true" pageId="39" pageNumber="40">D. warramboo</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4CF4701A7FC16FE79" authorityName="Abrams and Harvey" authorityYear="2020" box="[859,1014,475,501]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="immortalis">
<emphasis id="B913EADDFFA1FFF4CF4701A7FC16FE79" box="[859,1014,475,501]" italics="true" pageId="39" pageNumber="40">D. immortalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4C81901A7FB43FE79" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[1029,1187,475,501]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="belalugosii">
<emphasis id="B913EADDFFA1FFF4C81901A7FB43FE79" box="[1029,1187,475,501]" italics="true" pageId="39" pageNumber="40">D. belalugosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFA1FFF4C8AD01A7FAABFE79" authorityName="Abrams and Harvey" authorityYear="2020" box="[1201,1355,475,501]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="piscivultus">
<emphasis id="B913EADDFFA1FFF4C8AD01A7FAABFE79" box="[1201,1355,475,501]" italics="true" pageId="39" pageNumber="40">D. piscivultus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFA1FFF4C99D01A7FF13FD95" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="akashae">
<emphasis id="B913EADDFFA1FFF4C99D01A7FF13FD95" italics="true" pageId="39" pageNumber="40">D. akashae</emphasis>
</taxonomicName>
, which are not distinguishable using the ITS2 mini-barcode.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFA1FFFBCCDB025EFE61FD41" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="8BD836CFFFA1FFF4CCDB025EFA9CFDB1" blockId="39.[151,1437,150,681]" box="[199,1404,546,573]" pageId="39" pageNumber="40">
<emphasis id="B913EADDFFA1FFF4CCDB025EFE5BFDB0" bold="true" box="[199,443,546,572]" pageId="39" pageNumber="40">Description (adults).</emphasis>
<emphasis id="B913EADDFFA1FFF4CDDE025FFDF1FDB1" box="[450,529,547,573]" italics="true" pageId="39" pageNumber="40">Colour</emphasis>
. Pale yellow-brown; propeltidium, pedipalps and flagellum somewhat darker.
</paragraph>
<paragraph id="8BD836CFFFA1FFF4CCDB023BFC60FD25" blockId="39.[151,1437,150,681]" pageId="39" pageNumber="40">
<emphasis id="B913EADDFFA1FFF4CCDB023BFE8FFDED" box="[199,367,583,609]" italics="true" pageId="39" pageNumber="40">Cephalothorax</emphasis>
. Propeltidium with 2+1 apical setae in a triangular formation on anterior process and 2+2+2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 12 (
<specimenCount id="9D61FD46FFA1FFF4C8DB0217FB3DFD09" box="[1223,1245,619,645]" pageId="39" pageNumber="40" type="male"></specimenCount>
) setae (including 2 sternapophysial setae); posterior sternum triangular with 6 setae.
</paragraph>
<caption id="DF186647FFA1FFF4CC8B06EBFB85F97B" ID-DOI="http://doi.org/10.5281/zenodo.4416895" ID-Zenodo-Dep="4416895" httpUri="https://zenodo.org/record/4416895/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1687,1711]" targetBox="[151,1436,705,1664]" targetPageId="39">
<paragraph id="8BD836CFFFA1FFF4CC8B06EBFB85F97B" blockId="39.[151,1437,1687,1784]" pageId="39" pageNumber="40">
<emphasis id="B913EADDFFA1FFF4CC8B06EBFEC2F93C" bold="true" box="[151,290,1687,1712]" pageId="39" pageNumber="40">FIGURE 18.</emphasis>
<taxonomicName id="4C674D4CFFA1FFF4CD3006E4FDABF93C" authority="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey, 2020" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[300,587,1688,1712]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="christopherleei" status="sp. nov.">
<emphasis id="B913EADDFFA1FFF4CD3006E4FDABF93C" box="[300,587,1688,1712]" italics="true" pageId="39" pageNumber="40">Draculoides christopherleei</emphasis>
</taxonomicName>
<emphasis id="B913EADDFFA1FFF4CE4806E4FD48F93C" bold="true" box="[596,680,1688,1712]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="A22057A6FFA1FFF4CE4806E4FD48F93C" box="[596,680,1688,1712]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (WAM T129676): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 1 mm and also applies to E and F.
</paragraph>
</caption>
<paragraph id="8BD836CFFFA1FFF4CCDB075AFE88F878" blockId="39.[151,1437,1830,2036]" pageId="39" pageNumber="40">
<emphasis id="B913EADDFFA1FFF4CCDB075AFED5F8CC" box="[199,309,1830,1856]" italics="true" pageId="39" pageNumber="40">Chelicera</emphasis>
. Fixed finger with 2 large teeth plus 4 smaller teeth between these; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 composed of 5 setae; G3 composed of 6 setae; internal face of chelicera with 5 short whip-like setae (G4); brush at base of fixed finger composed of 8 setae (G5A), each densely pilose in distal half and G5B composed of 10 setae; G6 with one seta; G7 composed of 3 setae. movable finger serrula composed of ~16 long lamellae, blunt guard tooth present subdistally; 1 large accessory tooth present.
</paragraph>
<paragraph id="8BD836CFFFAEFFFBCCDB00EBFA6FFF59" blockId="40.[151,1437,151,933]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB00EBFECCFF3D" box="[199,300,151,177]" italics="true" pageId="40" pageNumber="41">Pedipalp</emphasis>
. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 7 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.27 × length of tarsus.
</paragraph>
<paragraph id="8BD836CFFFAEFFFBCCDB00A3FB0EFF75" blockId="40.[151,1437,151,933]" box="[199,1262,223,249]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB00A3FEE4FF74" box="[199,260,223,248]" italics="true" pageId="40" pageNumber="41">Legs.</emphasis>
Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
</paragraph>
<paragraph id="8BD836CFFFAEFFFBCCDB017FFF3AFEEA" blockId="40.[151,1437,151,933]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB017FFED1FE91" box="[199,305,259,285]" italics="true" pageId="40" pageNumber="41">Abdomen</emphasis>
. Chaetotaxy of tergites I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II: 3 macrosetae + 6 microsetae (microsetae in column), tergites IIIIX: 2: 2: 2: 2: 2: 2: 3; segment XII with small dorsal process (
<specimenCount id="9D61FD46FFAEFFFBC99B015BFA7CFECD" box="[1415,1436,295,321]" pageId="40" pageNumber="41" type="male"></specimenCount>
only).
</paragraph>
<paragraph id="8BD836CFFFAEFFFBCCDB0113FE5EFDED" blockId="40.[151,1437,151,933]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB0113FEDAFE05" box="[199,314,367,393]" italics="true" pageId="40" pageNumber="41">Flagellum</emphasis>
. Moderately dorsoventrally compressed (
<figureCitation id="135C2A4AFFAEFFFBCF120113FC86FE05" box="[782,870,367,393]" captionStart="FIGURE 18" captionStartId="39.[151,250,1687,1711]" captionTargetBox="[151,1436,705,1664]" captionTargetId="figure-248@39.[151,1436,705,1664]" captionTargetPageId="39" captionText="FIGURE 18. Draculoides christopherleei sp. nov., holotype male (WAM T129676): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 1 mm and also applies to E and F." figureDoi="http://doi.org/10.5281/zenodo.4416895" httpUri="https://zenodo.org/record/4416895/files/figure.png" pageId="40" pageNumber="41">Figs. 18</figureCitation>
DF, 19AC), with small lateral lobes, very broad stalk; in lateral view, dorsal margin very slightly concave with very slight dorsal concavity above vl1 terminating in small distal triangular tip; ventral margin evenly convex with shallow concavity between vm3 and vl2; 2.13 × longer than broad; seta dm1 situated dorso-medially, slightly closer to anterior margin; seta dm4 anterior to dl3; dl1 absent; dl3 on posterior margin; vm2 situated slightly anterior to vm1; paired vm3 staggered, with right seta below left; vm5 situated slightly posterior to dl1, closer to vm3 than vl2; 6 pairs of microsetae near anterior end, and three pairs between vl1 and vl2.
</paragraph>
<paragraph id="8BD836CFFFAEFFFBCCDB0217FE61FD41" blockId="40.[151,1437,151,933]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB0217FE6DFD09" box="[199,397,619,645]" italics="true" pageId="40" pageNumber="41">Dimensions (mm)</emphasis>
.
<typeStatus id="54DC886DFFAEFFFBCD860217FDE2FD09" box="[410,514,619,645]" pageId="40" pageNumber="41" type="holotype">Holotype</typeStatus>
male (WAM T129676): Body length 3.13. Propeltidium 0.88/0.58. Chelicera 0.48. Flagellum 0.33/0.15. Pedipalp: trochanter 0.46, femur 0.44, patella 0.50, tibia 0.42, tarsus 0.25, claw 0.07, total excluding claw 2.14.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFAEFFFBCCDB02ABFC0BFCD1" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BD836CFFFAEFFFBCCDB02ABFC4DFCB5" blockId="40.[151,1437,151,933]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB02ABFEDBFD7D" bold="true" box="[199,315,727,753]" pageId="40" pageNumber="41">Remarks.</emphasis>
<taxonomicName id="4C674D4CFFAEFFFBCD5D02ABFD9BFD7D" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[321,635,727,753]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFAEFFFBCD5D02ABFD9BFD7D" box="[321,635,727,753]" italics="true" pageId="40" pageNumber="41">Draculoides christopherleei</emphasis>
</taxonomicName>
is known from several locations within the Buckland Hills area, situated in the eastern Hamersley Range,
<collectingRegion id="49A3F82DFFAEFFFBCE190287FD2CFC99" box="[517,716,763,789]" country="Australia" name="Western Australia" pageId="40" pageNumber="41">Western Australia</collectingRegion>
(
<figureCitation id="135C2A4AFFAEFFFBCEC60287FCCEFC99" box="[730,814,763,789]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="40" pageNumber="41">Fig. 1E</figureCitation>
). The partial specimens listed above are associated with this species by locality and, in some cases, by sequence data (
<figureCitation id="135C2A4AFFAEFFFBCF550363FC7FFCB5" box="[841,927,799,825]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="40" pageNumber="41">Fig. 1E</figureCitation>
).
</paragraph>
<paragraph id="8BD836CFFFAEFFFBCCDB033EFC0BFCD1" blockId="40.[151,1437,151,933]" box="[199,1003,834,861]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB033EFE81FCD1" bold="true" box="[199,353,834,861]" pageId="40" pageNumber="41">Other names</emphasis>
. WAM SCH080 (
<bibRefCitation id="EFF64B3EFFAEFFFBCE35033FFCEDFCD1" author="Abrams, K. M. &amp; Huey, J. A. &amp; Hillyer, M. J. &amp; Humphreys, W. F. &amp; Didham, R. K. &amp; Harvey, M. S." box="[553,781,835,861]" pageId="40" pageNumber="41" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. &amp; Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
Abrams
<emphasis id="B913EADDFFAEFFFBCE97033FFD5CFCD1" box="[651,700,835,861]" italics="true" pageId="40" pageNumber="41">et al</emphasis>
., 2019
</bibRefCitation>
), Bennelongia B40.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFAEFFFBCCDB031BFC56FC29" pageId="40" pageNumber="41" type="etymology">
<paragraph id="8BD836CFFFAEFFFBCCDB031BFC56FC29" blockId="40.[151,1437,151,933]" pageId="40" pageNumber="41">
<emphasis id="B913EADDFFAEFFFBCCDB031BFEAEFC0D" bold="true" box="[199,334,871,897]" pageId="40" pageNumber="41">Etymology.</emphasis>
This species is named for actor Christopher Lee (19222015) who played the role of Dracula in the 1958 movie “Horror of Dracula” and the 1970 movie “Count Dracula.”
</paragraph>
</subSubSection>
</treatment>
</document>