469 lines
63 KiB
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469 lines
63 KiB
XML
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<mods:title id="F0EDE55FFFE735E6DA38778D068FB21E">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<treatment id="03CE87D9FF98FFF1CC8B0749FE10FB2D" ID-DOI="http://doi.org/10.5281/zenodo.4428346" ID-GBIF-Taxon="176485737" ID-Zenodo-Dep="4428346" LSID="urn:lsid:zoobank.org:act:A199BE76-0028-4AD8-AE10-0C75FE3705CC" httpUri="http://treatment.plazi.org/id/03CE87D9FF98FFF1CC8B0749FE10FB2D" lastPageId="34" lastPageNumber="35" pageId="30" pageNumber="31">
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<taxonomicName id="4C674D4CFF98FFCDCC8B0749FD5DF8C3" authority="Abrams and Harvey" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,701,1845,1872]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="belalugosii" status="sp. nov.">
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<emphasis id="B913EADDFF98FFCDCC8B0749FE55F8DC" bold="true" box="[151,437,1845,1872]" italics="true" pageId="30" pageNumber="31">Draculoides belalugosii</emphasis>
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Abrams and Harvey
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,
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<paragraph id="8BD836CFFF98FFCDCC8B0725FE86F8F8" blockId="30.[151,1316,1845,1944]" box="[151,358,1881,1908]" pageId="30" pageNumber="31">
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(
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<figureCitation id="135C2A4AFF98FFCDCC830725FEEBF8FF" box="[159,267,1881,1907]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support <80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="30" pageNumber="31">Figs. 1–8</figureCitation>
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,
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<figureCitation id="135C2A4AFF98FFCDCD0B0725FEBDF8F8" box="[279,349,1881,1908]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStartId-0="31.[151,250,1460,1484]" captionStartId-1="32.[151,250,1481,1505]" captionStartId-2="33.[151,250,1852,1876]" captionTargetBox-0="[151,1436,381,1436]" captionTargetBox-1="[151,1437,489,1458]" captionTargetBox-2="[150,1430,181,1817]" captionTargetId-0="figure-99@31.[151,1436,381,1436]" captionTargetId-1="figure-182@32.[151,1437,489,1458]" captionTargetId-2="figure-17@33.[150,1437,181,1827]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionTargetPageId-2="33" captionText-0="FIGURE 13. Draculoides belalugosii sp. nov., holotype male (WAM T135952): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." captionText-1="FIGURE 14. Draculoides belalugosii sp. nov., paratype female (WAM T135954): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." captionText-2="FIGURE 15. Draculoides belalugosii sp. nov.: A–C, holotype male (WAM T135952):A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. D–G, paratype female (WAM T135954): D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral; G. Spermathecae, ventral. Scale bars: A–C: 0.1 mm, D–F: 0.1mm, G: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4416883" figureDoi-1="http://doi.org/10.5281/zenodo.4416885" figureDoi-2="http://doi.org/10.5281/zenodo.4416889" httpUri-0="https://zenodo.org/record/4416883/files/figure.png" httpUri-1="https://zenodo.org/record/4416885/files/figure.png" httpUri-2="https://zenodo.org/record/4416889/files/figure.png" pageId="30" pageNumber="31">13–15</figureCitation>
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)
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</paragraph>
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<paragraph id="8BD836CFFF98FFCDCC8B0701FAC4F814" blockId="30.[151,1316,1845,1944]" box="[151,1316,1917,1944]" pageId="30" pageNumber="31">
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Zoobank Code: http://zoobank.org/NomenclaturalActs/
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<uuid id="FFC10C1AFF98FFCDCF140701FAC4F814" box="[776,1316,1917,1944]" pageId="30" pageNumber="31">A199BE76-0028-4AD8-AE10-0C75FE3705CC</uuid>
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</paragraph>
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</subSubSection>
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<paragraph id="8BD836CFFF98FFCCCC8B07BEFB03FF75" blockId="30.[151,861,1986,2010]" lastBlockId="31.[151,1436,150,358]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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<taxonomicName id="4C674D4CFF98FFCDCC8B07BEFE9CF856" authority="SCH" authorityName="SCH" box="[151,380,1986,2010]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis id="B913EADDFF98FFCDCC8B07BEFEA1F856" box="[151,321,1986,2010]" italics="true" pageId="30" pageNumber="31">Paradraculoides</emphasis>
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<materialsCitation id="3B0F3C92FF98FFCDCD5407BEFE9CF856" ID-GBIF-Occurrence="3013449399" box="[328,380,1986,2010]" collectingDate="2014-10-27" collectionCode="WAM, COI" collectorName="J. Alexander & J. King" country="Australia" county="Material" latitude="-22.005886" location="Pannawonica" longLatPrecision="1" longitude="116.51459" municipality="Buckland Hills" pageId="30" pageNumber="31" specimenCode="BHDD004-20141027, T135952, ITS2" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">SCH</materialsCitation>
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</taxonomicName>
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077:
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<bibRefCitation id="EFF64B3EFF98FFCDCDB707BEFD8EF856" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[427,622,1986,2010]" pageId="30" pageNumber="31" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
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Abrams
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<emphasis id="B913EADDFF98FFCDCE1E07BEFDCEF856" box="[514,558,1986,2010]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
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. 2019
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</bibRefCitation>
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MPE 106532: 8, fig. 2.
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<emphasis id="B913EADDFF99FFCCCC8B00EAFE63FF3C" bold="true" box="[151,387,150,176]" pageId="31" pageNumber="32">
|
||
<collectingCounty id="62B94E43FF99FFCCCC8B00EAFF1FFF3C" box="[151,255,150,176]" pageId="31" pageNumber="32">Material</collectingCounty>
|
||
examined.
|
||
</emphasis>
|
||
<emphasis id="B913EADDFF99FFCCCD9000EBFDDAFF3D" box="[396,570,151,177]" italics="true" pageId="31" pageNumber="32">
|
||
<typeStatus id="54DC886DFF99FFCCCD9000EBFE13FF3D" box="[396,499,151,177]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
<collectingCountry id="F370765FFF99FFCCCE5D00EBFD04FF3D" box="[577,740,151,177]" name="Australia" pageId="31" pageNumber="32">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFF99FFCCCEEC00EBFC5AFF3D" box="[752,954,151,177]" italics="true" pageId="31" pageNumber="32">
|
||
<collectingRegion id="49A3F82DFF99FFCCCEEC00EBFC5AFF3D" box="[752,954,151,177]" country="Australia" name="Western Australia" pageId="31" pageNumber="32">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality id="6BBCACB5FF99FFCCCFD700EBFB99FF3D" box="[971,1145,151,177]" pageId="31" pageNumber="32">Buckland Hills</collectingMunicipality>
|
||
,
|
||
<quantity id="4C9F9B2AFF99FFCCC89A00EAFB05FF3D" box="[1158,1253,150,177]" metricMagnitude="4" metricUnit="m" metricValue="4.54" pageId="31" pageNumber="32" unit="km" value="45.4">45.4 km</quantity>
|
||
SSE. of
|
||
<location id="8EB86014FF99FFCCC95200EBFF0BFF59" LSID="urn:lsid:plazi:treatment:03CE87D9FF98FFF1CC8B0749FE10FB2D:8EB86014FF99FFCCC95200EBFF0BFF59" country="Australia" county="Material" latitude="-22.005886" longLatPrecision="1" longitude="116.51459" municipality="Buckland Hills" name="Pannawonica" pageId="31" pageNumber="32" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FF99FFCCCCEB00C7FE40FF59" box="[247,416,187,213]" degrees="22" direction="south" minutes="00" orientation="latitude" pageId="31" pageNumber="32" precision="1" seconds="21.19" value="-22.005886">22°00’21.19”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF99FFCCCDB100C6FD86FF59" box="[429,614,186,213]" degrees="116" direction="east" minutes="30" orientation="longitude" pageId="31" pageNumber="32" precision="1" seconds="52.52" value="116.51459">116°30’52.52”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFF99FFCCCE6E00C7FCD2FF59" box="[626,818,187,213]" pageId="31" pageNumber="32" value="2014-10-27">
|
||
<collectingDate id="EF9DE9E7FF99FFCCCE6E00C7FCD2FF59" box="[626,818,187,213]" pageId="31" pageNumber="32" value="2014-10-27">27 October 2014</collectingDate>
|
||
</date>
|
||
, troglofauna scrape,
|
||
<collectorName id="26925319FF99FFCCC80100C7FB4BFF59" box="[1053,1195,187,213]" pageId="31" pageNumber="32">J. Alexander</collectorName>
|
||
,
|
||
<collectorName id="26925319FF99FFCCC8AB00C7FAE9FF59" box="[1207,1289,187,213]" pageId="31" pageNumber="32">J. King</collectorName>
|
||
(Biota Environmental Sciences,
|
||
<specimenCode id="DBC19EB4FF99FFCCCD9C00A3FD99FF75" box="[384,633,223,249]" collectionCode="BHDD" pageId="31" pageNumber="32">BHDD004-20141027</specimenCode>
|
||
-SC01) (
|
||
<collectionCode id="ED76AE0AFF99FFCCCECD00A3FCF8FF75" box="[721,792,223,249]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="31" pageNumber="32" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FF99FFCCCF0200A2FC66FF75" box="[798,902,222,249]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="31" pageNumber="32">T135952</specimenCode>
|
||
) (DNA:
|
||
<collectionCode id="ED76AE0AFF99FFCCCFF500A3FBFCFF75" box="[1001,1052,223,249]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="31" pageNumber="32" type="Herbarium">COI</collectionCode>
|
||
, 18S, 28S,
|
||
<specimenCode id="DBC19EB4FF99FFCCC88600A3FB37FF75" box="[1178,1239,223,249]" collectionCode="ITS" pageId="31" pageNumber="32">ITS2</specimenCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF99FFCCCCDB017FFB63FEE9" blockId="31.[151,1436,150,358]" pageId="31" pageNumber="32">
|
||
<materialsCitation id="3B0F3C92FF99FFCCCCDB017FFD14FEE9" ID-GBIF-Occurrence="3013449398" collectingDate="2014-10-26" collectingDateMax="2014-12-16" collectingDateMin="2014-10-26" collectionCode="WAM" collectorName="J. Alexander & J. King" country="Australia" latitude="-22.14" location="Pannawonica" longLatPrecision="21" longitude="116.23499" municipality="Buckland Hills" pageId="31" pageNumber="32" specimenCode="BHRC120-20141216, T1-02, T135954" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<emphasis id="B913EADDFF99FFCCCCDB017FFED9FE90" box="[199,313,259,284]" italics="true" pageId="31" pageNumber="32">
|
||
<typeStatus id="54DC886DFF99FFCCCCDB017FFED9FE90" box="[199,313,259,284]" pageId="31" pageNumber="32" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="F370765FFF99FFCCCD58017FFE07FE91" box="[324,487,259,285]" name="Australia" pageId="31" pageNumber="32">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFF99FFCCCDED017FFD58FE91" box="[497,696,259,285]" italics="true" pageId="31" pageNumber="32">
|
||
<collectingRegion id="49A3F82DFF99FFCCCDED017FFD58FE91" box="[497,696,259,285]" country="Australia" name="Western Australia" pageId="31" pageNumber="32">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D61FD46FF99FFCCCEDA017FFD0FFE91" box="[710,751,259,285]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FF99FFCCCEE0017FFC48FE91" box="[764,936,259,285]" pageId="31" pageNumber="32">Buckland Hills</collectingMunicipality>
|
||
;
|
||
<quantity id="4C9F9B2AFF99FFCCCFAE017FFBEDFE91" box="[946,1037,259,285]" metricMagnitude="4" metricUnit="m" metricValue="4.76" pageId="31" pageNumber="32" unit="km" value="47.6">47.6 km</quantity>
|
||
SSE. of
|
||
<location id="8EB86014FF99FFCCC872017FFAE6FE91" LSID="urn:lsid:plazi:treatment:03CE87D9FF98FFF1CC8B0749FE10FB2D:8EB86014FF99FFCCC872017FFAE6FE91" box="[1134,1286,259,285]" country="Australia" latitude="-22.14" longLatPrecision="21" longitude="116.23499" municipality="Buckland Hills" name="Pannawonica" pageId="31" pageNumber="32" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FF99FFCCC90D017FFA77FE91" box="[1297,1431,259,285]" degrees="22" direction="south" minutes="08" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="24" value="-22.14">22°08’24”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF99FFCCCC8B015BFECCFECD" box="[151,300,295,321]" degrees="116" direction="east" minutes="14" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="06" value="116.23499">116°14’06”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFF99FFCCCD26015BFD44FECD" box="[314,676,295,321]" pageId="31" pageNumber="32" value="2014-10-26" valueMax="2014-12-16" valueMin="2014-10-26">
|
||
<collectingDate id="EF9DE9E7FF99FFCCCD26015BFD44FECD" box="[314,676,295,321]" pageId="31" pageNumber="32" value="2014-10-26" valueMax="2014-12-16" valueMin="2014-10-26">26 October–16 December 2014</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="52264ED8FF99FFCCCEAD015BFC89FECD" box="[689,873,295,321]" pageId="31" pageNumber="32">troglofauna trap</collectingMethod>
|
||
,
|
||
<collectorName id="26925319FF99FFCCCF6B015BFBE7FECD" box="[887,1031,295,321]" pageId="31" pageNumber="32">J. Alexander</collectorName>
|
||
,
|
||
<collectorName id="26925319FF99FFCCC808015BFB87FECD" box="[1044,1127,295,321]" pageId="31" pageNumber="32">J. King</collectorName>
|
||
(Biota Environmental Sciences,
|
||
<specimenCode id="DBC19EB4FF99FFCCCCFF0137FE3DFEEA" box="[227,477,331,358]" collectionCode="BHRC" pageId="31" pageNumber="32">BHRC120-20141216</specimenCode>
|
||
-
|
||
<specimenCode id="DBC19EB4FF99FFCCCDF80137FDC2FEEA" box="[484,546,331,358]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="31" pageNumber="32">T1-02</specimenCode>
|
||
) (
|
||
<collectionCode id="ED76AE0AFF99FFCCCE240137FD9FFEE9" box="[568,639,331,357]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="31" pageNumber="32" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FF99FFCCCE990136FD0CFEE9" box="[645,748,330,357]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="31" pageNumber="32">T135954</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="3B0F3C92FF99FFCCCEE70137FB9FFEE9" ID-GBIF-Occurrence="3013449384" box="[763,1151,331,357]" collectingDate="2014-10-26" collectingDateMax="2014-12-16" collectingDateMin="2014-10-26" collectionCode="COI" collectorName="J. Alexander & J. King" country="Australia" isTailEnumeration="true" latitude="-22.14" location="Pannawonica" longLatPrecision="21" longitude="116.23499" municipality="Buckland Hills" pageId="31" pageNumber="32" specimenCode="ITS2" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
(DNA: 12S,
|
||
<collectionCode id="ED76AE0AFF99FFCCCF950137FC5CFEE9" box="[905,956,331,357]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="31" pageNumber="32" type="Herbarium">COI</collectionCode>
|
||
, 18S, 28S,
|
||
<specimenCode id="DBC19EB4FF99FFCCC8260137FB97FEE9" box="[1082,1143,331,357]" collectionCode="ITS" pageId="31" pageNumber="32">ITS2</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF186647FF99FFCCCC8B05C8FBD2F99F" ID-DOI="http://doi.org/10.5281/zenodo.4416883" ID-Zenodo-Dep="4416883" httpUri="https://zenodo.org/record/4416883/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1460,1484]" targetBox="[151,1436,381,1436]" targetPageId="31">
|
||
<paragraph id="8BD836CFFF99FFCCCC8B05C8FBD2F99F" blockId="31.[151,1436,1460,1556]" pageId="31" pageNumber="32">
|
||
<emphasis id="B913EADDFF99FFCCCC8B05C8FEC1FA40" bold="true" box="[151,289,1460,1484]" pageId="31" pageNumber="32">FIGURE 13.</emphasis>
|
||
<taxonomicName id="4C674D4CFF99FFCCCD3405C8FDFEFA40" authority="Abrams & Huey & Hillyer & Didham & Harvey, 2020" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[296,542,1460,1484]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="belalugosii" status="sp. nov.">
|
||
<emphasis id="B913EADDFF99FFCCCD3405C8FDFEFA40" box="[296,542,1460,1484]" italics="true" pageId="31" pageNumber="32">Draculoides belalugosii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EADDFF99FFCCCE3905C8FD97FA40" bold="true" box="[549,631,1460,1484]" pageId="31" pageNumber="32">
|
||
<taxonomicNameLabel id="A22057A6FF99FFCCCE3905C8FD97FA40" box="[549,631,1460,1484]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (WAM T135952): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37D6544FF99FFCCCCDB061AFA86F82C" pageId="31" pageNumber="32" type="diagnosis">
|
||
<paragraph id="8BD836CFFF99FFCCCCDB061AFA86F82C" blockId="31.[151,1437,1638,2025]" pageId="31" pageNumber="32">
|
||
<emphasis id="B913EADDFF99FFCCCCDB061AFEA2F90C" bold="true" box="[199,322,1638,1664]" pageId="31" pageNumber="32">Diagnosis.</emphasis>
|
||
The shape of the male flagellum is distinctive as it is short, broad, and rounded. It most closely resembles
|
||
<taxonomicName id="4C674D4CFF99FFCCCCE406F6FE0BF928" box="[248,491,1674,1700]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="B913EADDFF99FFCCCCE406F6FE0BF928" box="[248,491,1674,1700]" italics="true" pageId="31" pageNumber="32">Draculoides confusus</emphasis>
|
||
</taxonomicName>
|
||
in lateral profile but differs in the distal position of dm
|
||
<quantity id="4C9F9B2AFF99FFCCC84D06F6FB9BF928" box="[1105,1147,1674,1700]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="31" pageNumber="32" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="4C674D4CFF99FFCCC89D06F7FAFFF928" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[1153,1311,1674,1700]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="belalugosii">
|
||
<emphasis id="B913EADDFF99FFCCC89D06F7FAFFF928" box="[1153,1311,1674,1700]" italics="true" pageId="31" pageNumber="32">D. belalugosii</emphasis>
|
||
</taxonomicName>
|
||
(sub-distal in
|
||
<taxonomicName id="4C674D4CFF99FFCCCCAF06D3FED7F944" box="[179,311,1710,1736]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="B913EADDFF99FFCCCCAF06D3FED7F944" box="[179,311,1710,1736]" italics="true" pageId="31" pageNumber="32">D. confusus</emphasis>
|
||
</taxonomicName>
|
||
) and the close proximity of dl
|
||
<quantity id="4C9F9B2AFF99FFCCCE9A06D2FD4FF944" box="[646,687,1710,1736]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="31" pageNumber="32" unit="in" value="3.0">3 in</quantity>
|
||
relation to vl1 (dl3 is distally placed and widely separated from vl
|
||
<quantity id="4C9F9B2AFF99FFCCC98E06D2FF4DF960" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="31" pageNumber="32" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C674D4CFF99FFCCCCA906AFFED9F960" box="[181,313,1746,1772]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="confusus">
|
||
<emphasis id="B913EADDFF99FFCCCCA906AFFED9F960" box="[181,313,1746,1772]" italics="true" pageId="31" pageNumber="32">D. confusus</emphasis>
|
||
</taxonomicName>
|
||
). Females differ from all other species of
|
||
<taxonomicName id="4C674D4CFF99FFCCCF0A06AEFC40F960" authorityName="Harvey" authorityYear="1992" box="[790,928,1746,1772]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFF99FFCCCF0A06AEFC40F960" box="[790,928,1746,1772]" italics="true" pageId="31" pageNumber="32">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
by the comparatively short, rounded, unconnected spermathecae of the female genitalia.
|
||
<taxonomicName id="4C674D4CFF99FFCCCE8E068AFC7FF89C" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[658,927,1782,1808]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="belalugosii">
|
||
<emphasis id="B913EADDFF99FFCCCE8E068AFC7FF89C" box="[658,927,1782,1808]" italics="true" pageId="31" pageNumber="32">Draculoides belalugosii</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFF99FFCCC90E068AFA7CF89C" authorityName="Harvey" authorityYear="1992" box="[1298,1436,1782,1808]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFF99FFCCC90E068AFA7CF89C" box="[1298,1436,1782,1808]" italics="true" pageId="31" pageNumber="32">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at COI and 12S by the 50bp mini-barcodes shown in
|
||
<figureCitation id="135C2A4AFF99FFCCC8510766FB58F8B8" box="[1101,1208,1818,1844]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="31" pageNumber="32">Figures 3</figureCitation>
|
||
and
|
||
<figureCitation id="135C2A4AFF99FFCCC8EF0766FAE3F8B9" box="[1267,1283,1818,1845]" captionStart="FIGURE 5" captionStartId="20.[151,250,1216,1240]" captionTargetBox="[158,1424,183,1189]" captionTargetId="figure-20@20.[151,1436,181,1192]" captionTargetPageId="20" captionText="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416863" httpUri="https://zenodo.org/record/4416863/files/figure.png" pageId="31" pageNumber="32">5</figureCitation>
|
||
.
|
||
<taxonomicName id="4C674D4CFF99FFCCC90E0766FEF3F8D4" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="belalugosii">
|
||
<emphasis id="B913EADDFF99FFCCC90E0766FEF3F8D4" italics="true" pageId="31" pageNumber="32">Draculoides belalugosii</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFF99FFCCCE9E0742FCECF8D4" authorityName="Harvey" authorityYear="1992" box="[642,780,1854,1880]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFF99FFCCCE9E0742FCECF8D4" box="[642,780,1854,1880]" italics="true" pageId="31" pageNumber="32">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at ITS2 (
|
||
<figureCitation id="135C2A4AFF99FFCCC8AB0742FB1AF8D5" box="[1207,1274,1854,1881]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="31" pageNumber="32">Fig. 7</figureCitation>
|
||
) except for
|
||
<taxonomicName id="4C674D4CFF99FFCCC99D0743FEC0F8F0" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFF99FFCCC99D0743FEC0F8F0" italics="true" pageId="31" pageNumber="32">D. anachoretus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCCD32071FFE41F8F0" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[302,417,1890,1916]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bythius">
|
||
<emphasis id="B913EADDFF99FFCCCD32071FFE41F8F0" box="[302,417,1890,1916]" italics="true" pageId="31" pageNumber="32">D. bythius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCCDB3071FFDC9F8F0" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[431,553,1891,1916]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="eremius">
|
||
<emphasis id="B913EADDFF99FFCCCDB3071FFDC9F8F0" box="[431,553,1891,1916]" italics="true" pageId="31" pageNumber="32">D. eremius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCCE2A071FFD36F8F0" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[566,726,1890,1916]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="gnophicola">
|
||
<emphasis id="B913EADDFF99FFCCCE2A071FFD36F8F0" box="[566,726,1890,1916]" italics="true" pageId="31" pageNumber="32">D. gnophicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCCEF8071FFCB8F8F0" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[740,856,1890,1916]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kryptus">
|
||
<emphasis id="B913EADDFF99FFCCCEF8071FFCB8F8F0" box="[740,856,1890,1916]" italics="true" pageId="31" pageNumber="32">D. kryptus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCCF7A071FFBD9F8F0" authorityName="Abrams and Harvey" authorityYear="2020" box="[870,1081,1890,1916]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mckechnieorum">
|
||
<emphasis id="B913EADDFF99FFCCCF7A071FFBD9F8F0" box="[870,1081,1890,1916]" italics="true" pageId="31" pageNumber="32">D. mckechnieorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCC85B071FFB0CF8F0" authorityName="Abrams and Harvey" authorityYear="2020" box="[1095,1260,1890,1916]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="warramboo">
|
||
<emphasis id="B913EADDFF99FFCCC85B071FFB0CF8F0" box="[1095,1260,1890,1916]" italics="true" pageId="31" pageNumber="32">D. warramboo</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCC8E6071FFA75F8F0" authorityName="Abrams and Harvey" authorityYear="2020" box="[1274,1429,1890,1916]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFF99FFCCC8E6071FFA75F8F0" box="[1274,1429,1890,1916]" italics="true" pageId="31" pageNumber="32">D. immortalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCCC8B07FBFE83F82C" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,355,1926,1952]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="christopherleei">
|
||
<emphasis id="B913EADDFF99FFCCCC8B07FBFE83F82C" box="[151,355,1926,1952]" italics="true" pageId="31" pageNumber="32">D. christopherleei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF99FFCCCD6D07FBFDEBF82C" authorityName="Abrams and Harvey" authorityYear="2020" box="[369,523,1926,1952]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="piscivultus">
|
||
<emphasis id="B913EADDFF99FFCCCD6D07FBFDEBF82C" box="[369,523,1926,1952]" italics="true" pageId="31" pageNumber="32">D. piscivultus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B913EADDFF99FFCCCE5D07FBFD26F82C" box="[577,710,1926,1952]" italics="true" pageId="31" pageNumber="32">
|
||
<taxonomicName id="4C674D4CFF99FFCCCE5D07FBFD23F82C" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[577,707,1926,1952]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="akashae">D. akashae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which are not distinguishable using the ITS2 mini-barcode.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FF99FFF1CCDB07D6FEECFC29" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32" type="description">
|
||
<paragraph id="8BD836CFFF99FFCCCCDB07D6FBCCF848" blockId="31.[151,1437,1638,2025]" box="[199,1068,1962,1989]" pageId="31" pageNumber="32">
|
||
<emphasis id="B913EADDFF99FFCCCCDB07D6FE5BF848" bold="true" box="[199,443,1962,1988]" pageId="31" pageNumber="32">Description (adults).</emphasis>
|
||
<emphasis id="B913EADDFF99FFCCCDDE07D6FDF1F848" box="[450,529,1962,1988]" italics="true" pageId="31" pageNumber="32">Colour</emphasis>
|
||
. Yellow-brown; propeltidium somewhat darker.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF99FFF3CCDB07B2FB20FF59" blockId="31.[151,1437,1638,2025]" lastBlockId="32.[151,1436,151,465]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
|
||
<emphasis id="B913EADDFF99FFCCCCDB07B2FE8FF864" box="[199,367,1998,2024]" italics="true" pageId="31" pageNumber="32">Cephalothorax</emphasis>
|
||
. Propeltidium with 2 +1 apical setae in a triangular formation on anterior process and 2+1+2 setae; eye spots absent. Mesopeltidia widely separated. Metapeltidium partially divided. Anterior sternum with 13 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CCBD00C7FF58FF59" box="[161,184,187,213]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
), 14 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CCE700C7FEEDFF59" box="[251,269,187,213]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) setae (including 2 sternapophysial setae); posterior sternum triangular with 6 setae.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA6FFF3CCDB00A3FC47FE5D" blockId="32.[151,1436,151,465]" pageId="32" pageNumber="33">
|
||
<emphasis id="B913EADDFFA6FFF3CCDB00A3FED5FF75" box="[199,309,223,249]" italics="true" pageId="32" pageNumber="33">Chelicera</emphasis>
|
||
. Fixed finger with 2 large teeth plus 5 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CEF500A3FCE1FF75" box="[745,769,223,249]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
)(
|
||
<specimenCount id="9D61FD46FFA6FFF3CF0800A3FCC4FF75" box="[788,804,223,249]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) smaller teeth between these; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 composed of 5 (
|
||
<specimenCount id="9D61FD46FFA6FFF3C8F3017FFAE6FE91" box="[1263,1286,259,285]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
), 4 (
|
||
<specimenCount id="9D61FD46FFA6FFF3C925017FFAABFE91" box="[1337,1355,259,285]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) setae; G3 composed of 5 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CD6D015BFE67FECD" box="[369,391,295,321]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
),4 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CDA9015BFE27FECD" box="[437,455,295,321]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) setae; internal face of chelicera with 4 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CF96015BFC40FECD" box="[906,928,295,321]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
) (
|
||
<specimenCount id="9D61FD46FFA6FFF3CFA6015BFC2CFECD" box="[954,972,295,321]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) short whip-like setae (G4); brush at base of fixed finger composed of 6 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CDE60137FDF1FEE9" box="[506,529,331,357]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
), 5 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CE5B0137FDB9FEE9" box="[583,601,331,357]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) setae (G5A), each densely pilose in distal half and G5B composed of 10 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CCBD0113FF58FE05" box="[161,184,367,393]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
), 9 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CCF00113FF1EFE05" box="[236,254,367,393]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) setae; G6 with one seta; G7 composed of 5 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CEE70113FCF3FE05" box="[763,787,367,393]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
)(
|
||
<specimenCount id="9D61FD46FFA6FFF3CF3B0113FCD7FE05" box="[807,823,367,393]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) setae. Movable finger serrula composed of 16 (
|
||
<specimenCount id="9D61FD46FFA6FFF3C9480113FA8BFE05" box="[1364,1387,367,393]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
), 17 (
|
||
<specimenCount id="9D61FD46FFA6FFF3CCBE01EFFF54FE21" box="[162,180,403,429]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) long lamellae, blunt guard tooth present subdistally; 1 large accessory tooth present at two-thirds from base of serrula (
|
||
<specimenCount id="9D61FD46FFA6FFF3CCE801CBFEEBFE5D" box="[244,267,439,465]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
), 1 large and 1 slightly smaller accessory tooth present (
|
||
<specimenCount id="9D61FD46FFA6FFF3CF9801CBFC77FE5D" box="[900,919,439,465]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF186647FFA6FFF3CC8B05B5FBD2F9A5" ID-DOI="http://doi.org/10.5281/zenodo.4416885" ID-Zenodo-Dep="4416885" httpUri="https://zenodo.org/record/4416885/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1481,1505]" targetBox="[151,1437,489,1458]" targetPageId="32">
|
||
<paragraph id="8BD836CFFFA6FFF3CC8B05B5FBD2F9A5" blockId="32.[151,1436,1481,1578]" pageId="32" pageNumber="33">
|
||
<emphasis id="B913EADDFFA6FFF3CC8B05B5FEC0FA6D" bold="true" box="[151,288,1481,1505]" pageId="32" pageNumber="33">FIGURE 14.</emphasis>
|
||
<taxonomicName id="4C674D4CFFA6FFF3CD3A05B5FDFBFA6D" authority="Abrams & Huey & Hillyer & Didham & Harvey, 2020" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[294,539,1481,1505]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="belalugosii" status="sp. nov.">
|
||
<emphasis id="B913EADDFFA6FFF3CD3A05B5FDFBFA6D" box="[294,539,1481,1505]" italics="true" pageId="32" pageNumber="33">Draculoides belalugosii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EADDFFA6FFF3CE3D05B5FD91FA6D" bold="true" box="[545,625,1481,1505]" pageId="32" pageNumber="33">
|
||
<taxonomicNameLabel id="A22057A6FFA6FFF3CE3D05B5FD91FA6D" box="[545,625,1481,1505]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype female (WAM T135954): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836CFFFA6FFF3CCDB0607FEABF952" blockId="32.[151,1436,1659,1758]" pageId="32" pageNumber="33">
|
||
<emphasis id="B913EADDFFA6FFF3CCDB0607FECCF919" box="[199,300,1659,1685]" italics="true" pageId="32" pageNumber="33">Pedipalp</emphasis>
|
||
. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 8 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.42 (
|
||
<specimenCount id="9D61FD46FFA6FFF3C8E506DCFAF0F936" box="[1273,1296,1696,1722]" pageId="32" pageNumber="33" type="male">♂</specimenCount>
|
||
), 0.55 (
|
||
<specimenCount id="9D61FD46FFA6FFF3C97406DCFA9AF936" box="[1384,1402,1696,1722]" pageId="32" pageNumber="33" type="female">♀</specimenCount>
|
||
) × length of tarsus.
|
||
</paragraph>
|
||
<caption id="DF186647FFA7FFF2CC8B0740FC1EF817" ID-DOI="http://doi.org/10.5281/zenodo.4416889" ID-Zenodo-Dep="4416889" httpUri="https://zenodo.org/record/4416889/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1852,1876]" targetBox="[150,1430,181,1817]" targetPageId="33">
|
||
<paragraph id="8BD836CFFFA7FFF2CC8B0740FC1EF817" blockId="33.[151,1437,1852,1948]" pageId="33" pageNumber="34">
|
||
<emphasis id="B913EADDFFA7FFF2CC8B0740FEFFF8D8" bold="true" box="[151,287,1852,1876]" pageId="33" pageNumber="34">FIGURE 15.</emphasis>
|
||
<taxonomicName id="4C674D4CFFA7FFF2CD390740FDF8F8D8" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[293,536,1852,1876]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="belalugosii">
|
||
<emphasis id="B913EADDFFA7FFF2CD390740FDF8F8D8" box="[293,536,1852,1876]" italics="true" pageId="33" pageNumber="34">Draculoides belalugosii</emphasis>
|
||
</taxonomicName>
|
||
s
|
||
<emphasis id="B913EADDFFA7FFF2CE3B0740FD8DF8D8" bold="true" box="[551,621,1852,1876]" pageId="33" pageNumber="34">p. nov.</emphasis>
|
||
: A–C, holotype male (WAM T135952): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. D–G, paratype female (WAM T135954): D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral; G. Spermathecae, ventral. Scale bars: A–C: 0.1 mm, D–F: 0.1mm, G: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB00EBFB0EFF3D" blockId="34.[151,1437,151,1185]" box="[199,1262,151,177]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB00EBFEE4FF3C" box="[199,260,151,176]" italics="true" pageId="34" pageNumber="35">Legs.</emphasis>
|
||
Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB00C7FF3AFE92" blockId="34.[151,1437,151,1185]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB00C7FED1FF59" box="[199,305,187,213]" italics="true" pageId="34" pageNumber="35">Abdomen</emphasis>
|
||
. Chaetotaxy of tergite I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II: 3 macrosetae + 6 microsetae (microsetae in column), tergites III–IX: 2: 2: 2: 2: 2: 2: 2; segment XII with small dorsal process (
|
||
<specimenCount id="9D61FD46FFA4FFF1C99B00A3FA7CFF75" box="[1415,1436,223,249]" pageId="34" pageNumber="35" type="male">♂</specimenCount>
|
||
only).
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB015BFCF7FE05" blockId="34.[151,1437,151,1185]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB015BFE69FECD" box="[199,393,295,321]" italics="true" pageId="34" pageNumber="35">Female genitalia</emphasis>
|
||
. Two pairs of spermathecae with short circular, equal-sized lobes, each pair not connected basally before connection with bursa (
|
||
<figureCitation id="135C2A4AFFA4FFF1CE5F0137FD4AFEE9" box="[579,682,330,357]" captionStart="FIGURE 15" captionStartId="33.[151,250,1852,1876]" captionTargetBox="[150,1430,181,1817]" captionTargetId="figure-17@33.[150,1437,181,1827]" captionTargetPageId="33" captionText="FIGURE 15. Draculoides belalugosii sp. nov.: A–C, holotype male (WAM T135952):A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. D–G, paratype female (WAM T135954): D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral; G. Spermathecae, ventral. Scale bars: A–C: 0.1 mm, D–F: 0.1mm, G: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4416889" httpUri="https://zenodo.org/record/4416889/files/figure.png" pageId="34" pageNumber="35">Fig. 15G</figureCitation>
|
||
), distally round and smooth; sparsely covered with small pores and numerous tiny folds; gonopod short, roughly rectangular.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB01EFFBB4FD41" blockId="34.[151,1437,151,1185]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB01EFFEDAFE21" box="[199,314,403,429]" italics="true" pageId="34" pageNumber="35">Flagellum</emphasis>
|
||
. Male: Dorsoventrally compressed (
|
||
<figureCitation id="135C2A4AFFA4FFF1CEC901EFFCCEFE21" box="[725,814,403,429]" captionStart="FIGURE 13" captionStartId="31.[151,250,1460,1484]" captionTargetBox="[151,1436,381,1436]" captionTargetId="figure-99@31.[151,1436,381,1436]" captionTargetPageId="31" captionText="FIGURE 13. Draculoides belalugosii sp. nov., holotype male (WAM T135952): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." figureDoi="http://doi.org/10.5281/zenodo.4416883" httpUri="https://zenodo.org/record/4416883/files/figure.png" pageId="34" pageNumber="35">Figs. 13</figureCitation>
|
||
D–F, 15A–C); 1.86 × longer than broad; seta dm1 situated dorso-medially; seta dm4 situated close to posterior margin; paired setae dl3 ‘staggered’ with left lower than right; dl3 on posterior margin, level with dm4; vm2 situated slightly posterior to vm1; vm5 situated slightly anterior to vl1, midway between vm3 and vl2; at least 4 pairs of microsetae between vl1 and dl3. Female: 4.25 × longer than broad; seta dm1 situated towards posterior end of flagellomere II, level with vm2; setae dl1 small, situated anterior to dm4, dm4 situated at four fifth length of flagellomere III; dl3 situated almost at posterior margin slightly more posterior than vl2, vm1 situated level with vm2, vm3 situated closer to vm1 than to vm5, vm5 closer to vm3 than vl2, vl1 situated posterior to vm3 and anterior to dl1; 1 pair of microsetae posterior to flagellomere III annulus, 1 pair of microsetae posterior to dl1, 1 pair of microsetae laterally between dl3 and vl2.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB02ABFE61FCB5" blockId="34.[151,1437,151,1185]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB02ABFE6EFD7D" box="[199,398,727,753]" italics="true" pageId="34" pageNumber="35">Dimensions (mm)</emphasis>
|
||
.
|
||
<typeStatus id="54DC886DFFA4FFF1CD8002ABFDE4FD7D" box="[412,516,727,753]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
male (WAM T135952): Body length 2.90. Propeltidium 0.96/0.50. Chelicera 0.6. Flagellum 0.38/0.20. Pedipalp: trochanter 0.35, femur 0.40, patella 0.44, tibia 0.52, tarsus 0.23, claw 0.10, total excluding claw 2.04.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB033FFEECFC29" blockId="34.[151,1437,151,1185]" pageId="34" pageNumber="35">
|
||
<typeStatus id="54DC886DFFA4FFF1CCDB033FFEC9FCD1" box="[199,297,835,861]" pageId="34" pageNumber="35" type="paratype">Paratype</typeStatus>
|
||
female (WAM T135954): Body length 3.65. Propeltidium 1.00/0.58. Chelicera 0.75. Flagellum 0.16/0.04. Pedipalp: trochanter 0.46, femur 0.67, patella 0.48, tibia 0.40, tarsus 0.21, claw 0.12, total excluding claw 2.35.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFA4FFF1CCDB03D3FC29FBD5" pageId="34" pageNumber="35" type="discussion">
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB03D3FC87FBB9" blockId="34.[151,1437,151,1185]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB03D3FEDBFC45" bold="true" box="[199,315,943,969]" pageId="34" pageNumber="35">Remarks.</emphasis>
|
||
<taxonomicName id="4C674D4CFFA4FFF1CD5F03D3FDB2FC45" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[323,594,943,969]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="belalugosii">
|
||
<emphasis id="B913EADDFFA4FFF1CD5F03D3FDB2FC45" box="[323,594,943,969]" italics="true" pageId="34" pageNumber="35">Draculoides belalugosii</emphasis>
|
||
</taxonomicName>
|
||
is known from several locations within an area known as Buckland Hills, situated in the eastern part of the Hamersley Range,
|
||
<collectingRegion id="49A3F82DFFA4FFF1CEF403AFFC50FC61" box="[744,944,979,1005]" country="Australia" name="Western Australia" pageId="34" pageNumber="35">Western Australia</collectingRegion>
|
||
(
|
||
<figureCitation id="135C2A4AFFA4FFF1CFA303AFFBF5FC61" box="[959,1045,979,1005]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="34" pageNumber="35">Fig. 1E</figureCitation>
|
||
).
|
||
<taxonomicName id="4C674D4CFFA4FFF1C83003AFFA88FC61" authorityName="Abrams and Harvey" authorityYear="2020" box="[1068,1384,979,1005]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="christopherleei">
|
||
<emphasis id="B913EADDFFA4FFF1C83003AFFA88FC61" box="[1068,1384,979,1005]" italics="true" pageId="34" pageNumber="35">Draculoides christopherleei</emphasis>
|
||
</taxonomicName>
|
||
also inhabits this area as well as two distinctive lineages (SCH100 and SCH101) which may represent new species but for which we do not have sufficient data to describe at this time.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB0442FC29FBD5" blockId="34.[151,1437,151,1185]" box="[199,969,1086,1113]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB0442FE81FBD5" bold="true" box="[199,353,1086,1113]" pageId="34" pageNumber="35">Other names</emphasis>
|
||
. WAM SCH077 (
|
||
<bibRefCitation id="EFF64B3EFFA4FFF1CE350443FCEDFBD5" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[553,781,1087,1113]" pageId="34" pageNumber="35" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
|
||
Abrams
|
||
<emphasis id="B913EADDFFA4FFF1CE970443FD5CFBD5" box="[651,700,1087,1113]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
|
||
., 2019
|
||
</bibRefCitation>
|
||
), Helix sp. SAL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FFA4FFF1CCDB041FFE10FB2D" pageId="34" pageNumber="35" type="etymology">
|
||
<paragraph id="8BD836CFFFA4FFF1CCDB041FFE10FB2D" blockId="34.[151,1437,151,1185]" pageId="34" pageNumber="35">
|
||
<emphasis id="B913EADDFFA4FFF1CCDB041FFEAEFBF1" bold="true" box="[199,334,1123,1149]" pageId="34" pageNumber="35">Etymology.</emphasis>
|
||
This species is named for actor Bela Lugosi (1882–1956) who played the role of Dracula in the 1931 movie of the same name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |