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<document id="731CB8E01227CE78D30F17826B2BF4C7" ID-DOI="10.5281/zenodo.5375302" ID-ISSN="1638-9395" ID-Zenodo-Dep="5375302" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630277866243" checkinUser="marcus" docAuthor="Pacaud, J. - M." docDate="2003" docId="03CE87D4FFCBFFC54545FB5BA061ABBC" docLanguage="en" docName="Geodiversitas.2003.25.3.451-462.pdf" docOrigin="Geodiversitas 25 (3)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Pseudocypraea dolini Pacaud 2003, n. sp." docType="treatment" docVersion="3" lastPageNumber="456" masterDocId="FFF7FFACFFCEFFC3444BFFB0A359AD39" masterDocTitle="First fossil records of the Recent Ovulid genus Pseudocypraea Schilder, 1927 (Mollusca, Gastropoda) with description of a new species" masterLastPageNumber="462" masterPageNumber="451" pageNumber="456" updateTime="1699211013072" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="5D58369A788F70CE2E2F2CB4CC4F42AC">First fossil records of the Recent Ovulid genus Pseudocypraea Schilder, 1927 (Mollusca, Gastropoda) with description of a new species</mods:title>
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<mods:namePart id="6DF410FA171F3169FD7EA796BBA7090C">Pacaud, J. - M.</mods:namePart>
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<mods:date id="BE7F756E4228B0D08CD27B9DD6949674">2003</mods:date>
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<mods:identifier id="D897B5605091915A3D24A4A961042DB1" type="ISSN">1638-9395</mods:identifier>
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<treatment id="03CE87D4FFCBFFC54545FB5BA061ABBC" LSID="urn:lsid:plazi:treatment:03CE87D4FFCBFFC54545FB5BA061ABBC" httpUri="http://treatment.plazi.org/id/03CE87D4FFCBFFC54545FB5BA061ABBC" lastPageId="6" lastPageNumber="456" pageId="5" pageNumber="456">
<subSubSection id="C37D6549FFCBFFC64545FB5BA15FA81C" pageId="5" pageNumber="456" type="nomenclature">
<paragraph id="8BD836C2FFCBFFC64545FB5BA15FA81C" blockId="5.[270,565,1258,1317]" pageId="5" pageNumber="456">
<heading id="D09081AEFFCBFFC64545FB5BA15FA81C" centered="true" fontSize="11" level="2" pageId="5" pageNumber="456" reason="2">
<taxonomicName id="4C674D41FFCBFFC64545FB5BA2ABA83C" authority="Pacaud, 2003" authorityName="Pacaud" authorityYear="2003" box="[270,498,1259,1285]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="dolini" status="sp. nov.">
<emphasis id="B913EAD0FFCBFFC64545FB5BA2ABA83C" bold="true" box="[270,498,1259,1285]" italics="true" pageId="5" pageNumber="456">Pseudocypraea dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057ABFFCBFFC645B1FB5AA16CA83D" box="[506,565,1258,1284]" pageId="5" pageNumber="456" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="135C2A47FFCBFFC6450FFABBA2DDA81C" box="[324,388,1291,1317]" captionStart="FIG" captionStartId="6.[151,162,1442,1459]" captionTargetBox="[237,1174,174,1398]" captionTargetId="figure-69@6.[216,1179,144,1413]" captionTargetPageId="6" captionText="FIG. 3. — A, B, Pseudocypraea eratoformis (Hoernes &amp; Auinger, 1880) n. comb., Langhian (middle Miocene) of Lapugiu de Sus (Rumania), original pictures; C-E, Pseudocypraea dolini n. sp.; C, under ultraviolet light showing the trace of a colour pattern; D, E, ventral face and detail of the siphonal canal region. Scale bars: A, B, 3 mm; C-E, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5375308" httpUri="https://zenodo.org/record/5375308/files/figure.png" pageId="5" pageNumber="456">Figs 3</figureCitation>
C-E; 4A-C)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D6549FFCBFFC644DCFAF3A1F4A865" box="[151,685,1347,1372]" pageId="5" pageNumber="456" type="materials_examined">
<paragraph id="8BD836C2FFCBFFC644DCFAF3A1F4A865" blockId="5.[151,685,1347,1668]" box="[151,685,1347,1372]" pageId="5" pageNumber="456">
<materialsCitation id="3B0F3C9FFFCBFFC644DCFAF3A1F4A865" accessionNumber="R63008" box="[151,685,1347,1372]" collectionCode="MNHN-DHT" pageId="5" pageNumber="456" specimenCount="1" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus id="54DC8860FFCBFFC64519FAF4A2F5A865" box="[338,428,1348,1372]" pageId="5" pageNumber="456" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED76AE07FFCBFFC645F3FAF4A113A865" box="[440,586,1348,1372]" pageId="5" pageNumber="456">MNHN-DHT</collectionCode>
R63008).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D6549FFCBFFC644DCFADEA111A880" pageId="5" pageNumber="456" type="etymology">
<paragraph id="8BD836C2FFCBFFC644DCFADEA111A880" blockId="5.[151,685,1347,1668]" pageId="5" pageNumber="456">ETYMOLOGY. — Named after Luc Dolin, in recognition of his help and his numerous contributions to Tertiary Cypraeid and Ovulid paleontology.</paragraph>
</subSubSection>
<subSubSection id="C37D6549FFCBFFC644DCFA7BA2E3AB79" pageId="5" pageNumber="456" type="materials_examined">
<paragraph id="8BD836C2FFCBFFC644DCFA7BA26DA8C4" blockId="5.[151,685,1347,1668]" pageId="5" pageNumber="456">
<typeStatus id="54DC8860FFCBFFC644DCFA7BA39EA8D9" box="[151,199,1483,1506]" pageId="5" pageNumber="456">TYPE</typeStatus>
LOCALITY. — Le Quoniam near Haravilliers (Val dOise,
<collectingCountry id="F3707652FFCBFFC644A8FA55A273A8C4" box="[227,298,1509,1533]" name="France" pageId="5" pageNumber="456">France</collectingCountry>
).
</paragraph>
<paragraph id="8BD836C2FFCBFFC644DCF9BFA2E3AB79" blockId="5.[151,685,1347,1668]" pageId="5" pageNumber="456">STRATIGRAPHIC HORIZON. — Early Bartonian (Marinesian, middle Eocene).</paragraph>
</subSubSection>
<subSubSection id="C37D6549FFCBFFC644DCF9E2A056A9BC" pageId="5" pageNumber="456" type="description">
<paragraph id="8BD836C2FFCBFFC644DCF9E2A3B5ABBB" blockId="5.[151,685,1347,1668]" pageId="5" pageNumber="456">DIMENSIONS. — Height: 10.9 mm, maximal diameter: 6.6 mm.</paragraph>
<paragraph id="8BD836C2FFCBFFC64695FF1EA0BDADCD" blockId="5.[769,1247,169,245]" pageId="5" pageNumber="456">
IAGNOSIS. — A large fusiform
<taxonomicName id="4C674D41FFCBFFC64058FF1AA7C3ADFB" authorityName="Schilder" authorityYear="1927" box="[1043,1178,170,194]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="genus">
<emphasis id="B913EAD0FFCBFFC64058FF1AA7C3ADFB" box="[1043,1178,170,194]" italics="true" pageId="5" pageNumber="456">Pseudocypraea</emphasis>
</taxonomicName>
, ornamented with strong spiral cords, the outerlip labial being short and strong.
</paragraph>
<paragraph id="8BD836C2FFCBFFC64681FE9AA002AC7B" blockId="5.[714,1247,303,1157]" box="[714,859,298,324]" pageId="5" pageNumber="456">DESCRIPTION</paragraph>
<paragraph id="8BD836C2FFCBFFC64681FEFBA056A9BC" blockId="5.[714,1247,303,1157]" pageId="5" pageNumber="456">The shell is of a small size, fusiform, with an involuted protoconch. The dorsal area as well as the ventral area are deeply incised by a spiral sculpture (which the wear of the shell has not completely erased). The ventral sole is evenly convex, the neck, as pinched, forms an obtuse bottom. The aperture, quite broad, with subparallel sides, is evenly curved. The siphonal canal, cone-shaped, is deep and well defined. The terminal fold is lamellar, trimmed, long and straight, in an adaxial position. The auriform and concave fossula, which is reduced, slightly trimmed, and the convex columellar area, suggest the juvenile decoration. The angulation is slightly settled. The inner lip bears 18 strong, short teeth, which become thinner and more elongated adapically. The adapical edge, obscurely denticulated is thin, elongated and straight. The excurrent channel is deep, well defined and forms an adapical bridge, which is canaliculate and subaxially opening. The outer lip, crescent-shaped, slightly flattened by the callous development of the margin, bears 21 big short labral teeth, hardly more elongated in their abapical quarter. The outer lip shows spots of colour along the periphery.</paragraph>
</subSubSection>
<subSubSection id="C37D6549FFCBFFC54681FB1AA061ABBC" lastPageId="6" lastPageNumber="457" pageId="5" pageNumber="456" type="discussion">
<paragraph id="8BD836C2FFCBFFC64681FB1AA015A9FB" blockId="5.[714,1247,1194,1669]" box="[714,844,1194,1220]" pageId="5" pageNumber="456">DISCUSSION</paragraph>
<paragraph id="8BD836C2FFCBFFC54681FB7BA061ABBC" blockId="5.[714,1247,1194,1669]" lastBlockId="6.[714,1247,1578,1669]" lastPageId="6" lastPageNumber="457" pageId="5" pageNumber="456">
A l t h o u g h t h i s d e s c r i p t i o n i s b a s e d o n a n unique specimen,
<taxonomicName id="4C674D41FFCBFFC647EEFB5BA7CBA83C" authority="Pacaud, 2003" authorityName="Pacaud" authorityYear="2003" box="[933,1170,1259,1285]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="dolini" status="sp. nov.">
<emphasis id="B913EAD0FFCBFFC647EEFB5BA7CBA83C" box="[933,1170,1259,1285]" italics="true" pageId="5" pageNumber="456">Pseudocypraea dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057ABFFCBFFC640D6FB5AA786A83D" box="[1181,1247,1258,1284]" pageId="5" pageNumber="456" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C674D41FFCBFFC64720FABAA74CA81A" authorityName="Pacaud" authorityYear="2003" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[875,1045,1289,1316]" class="Gastropoda" family="Cypraeidae" genus="Cypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="eratoformis" status="comb. nov.">
<emphasis id="B913EAD0FFCBFFC64720FABAA023A81A" box="[875,890,1290,1315]" italics="true" pageId="5" pageNumber="456">P</emphasis>
.
<emphasis id="B913EAD0FFCBFFC647C4FAB9A74CA81A" box="[911,1045,1289,1315]" italics="true" pageId="5" pageNumber="456">eratoformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057ABFFCBFFC6406BFABAA7CDA81C" box="[1056,1172,1290,1317]" pageId="5" pageNumber="456" rank="species">n. comb.</taxonomicNameLabel>
, from
<taxonomicName id="4C674D41FFCBFFC64681FA9AA006A87C" authorityName="Sowerby" authorityYear="1832" box="[714,863,1322,1349]" class="Gastropoda" family="Cypraeidae" genus="Cypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="adamsonii">
<emphasis id="B913EAD0FFCBFFC64681FA9AA180A87A" box="[714,729,1322,1347]" italics="true" pageId="5" pageNumber="456">P</emphasis>
.
<emphasis id="B913EAD0FFCBFFC646A0FA9BA006A87C" box="[747,863,1323,1349]" italics="true" pageId="5" pageNumber="456">adamsonii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D41FFCBFFC647D4FA9AA77CA87A" authorityName="Petuch" authorityYear="1979" box="[927,1061,1322,1348]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="exquisita">
<emphasis id="B913EAD0FFCBFFC647D4FA9AA0F7A87A" box="[927,942,1322,1347]" italics="true" pageId="5" pageNumber="456">P</emphasis>
.
<emphasis id="B913EAD0FFCBFFC6478BFA9AA77CA87A" box="[960,1061,1322,1347]" italics="true" pageId="5" pageNumber="456">exquisita</emphasis>
</taxonomicName>
(
<figureCitation id="135C2A47FFCBFFC64073FA9BA7DAA87C" box="[1080,1155,1323,1349]" captionStart="FIG" captionStartId="7.[151,162,1545,1562]" captionTargetBox="[196,1201,253,1514]" captionTargetId="figure-18@7.[196,1201,222,1515]" captionTargetPageId="7" captionText="FIG. 4. — A-C, Pseudocypraea dolini n. sp., Bartonian (Marinesian, middle Eocene) of Le Quoniam (Val dOise), holotype (MNHN- DHT R63008) (leg. J.-M. Pacaud); D-F, Pseudocypraea eratoformis (Hoernes &amp; Auinger, 1880) n. comb., Langhian (middle Miocene) of Lapugiu de Sus (Rumania), holotype (NHMW 1999Z0077/0027). Scale bar: A-C, 5 mm; D-F, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5375310" httpUri="https://zenodo.org/record/5375310/files/figure.png" pageId="5" pageNumber="456">Figs 4</figureCitation>
D-F; 5) by numerous specific characters: tapering and non subglobose morphology, reduced fossula, adapical edge very thin, excurrent channel not curled and embayed, and indeed less numerous denticulation of the outer lip (21 shorter labral teeth in
<taxonomicName id="4C674D41FFCBFFC64765FA5AA0C9AB3C" authority="Pacaud, 2003" authorityName="Pacaud" authorityYear="2003" box="[814,912,1514,1541]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="dolini" status="sp. nov.">
<emphasis id="B913EAD0FFCBFFC64765FA5AA064AB3A" box="[814,829,1514,1539]" italics="true" pageId="5" pageNumber="456">P</emphasis>
.
<emphasis id="B913EAD0FFCBFFC64704FA5BA0C9AB3C" box="[847,912,1515,1541]" italics="true" pageId="5" pageNumber="456">dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057ABFFCBFFC647D1FA5AA084AB3D" box="[922,989,1514,1540]" pageId="5" pageNumber="456" rank="species">n. sp.</taxonomicNameLabel>
, against an average of
<quantity id="4C9F9B27FFCBFFC64681F9BBA051AB1C" box="[714,776,1547,1573]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="5" pageNumber="456" unit="in" value="26.0">26 in</quantity>
<taxonomicName id="4C674D41FFCBFFC64758F9BAA0FCAB1C" authorityName="Sowerby" authorityYear="1832" box="[787,933,1546,1573]" class="Gastropoda" family="Cypraeidae" genus="Cypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="adamsonii">
<emphasis id="B913EAD0FFCBFFC64758F9BAA07BAB1A" box="[787,802,1546,1571]" italics="true" pageId="5" pageNumber="456">P</emphasis>
.
<emphasis id="B913EAD0FFCBFFC6477FF9BBA0FCAB1C" box="[820,933,1547,1573]" italics="true" pageId="5" pageNumber="456">adamsonii</emphasis>
</taxonomicName>
,
<quantity id="4C9F9B27FFCBFFC647F3F9BBA0AEAB1C" box="[952,1015,1547,1573]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="456" unit="in" value="20.0">20 in</quantity>
<taxonomicName id="4C674D41FFCBFFC6404AF9BAA7DAAB1A" authorityName="Petuch" authorityYear="1979" box="[1025,1155,1546,1572]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="exquisita">
<emphasis id="B913EAD0FFCBFFC6404AF9BAA749AB1A" box="[1025,1040,1546,1571]" italics="true" pageId="5" pageNumber="456">P</emphasis>
.
<emphasis id="B913EAD0FFCBFFC6406AF9BAA7DAAB1A" box="[1057,1155,1546,1571]" italics="true" pageId="5" pageNumber="456">exquisita</emphasis>
</taxonomicName>
and
<quantity id="4C9F9B27FFCBFFC64089F9BBA1B8AB7C" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="5" pageNumber="456" unit="in" value="24.0">24 in</quantity>
<taxonomicName id="4C674D41FFCBFFC646A2F99AA0D8AB7A" authorityName="Pacaud" authorityYear="2003" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[745,897,1577,1604]" class="Gastropoda" family="Cypraeidae" genus="Cypraea" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="456" phylum="Mollusca" rank="species" species="eratoformis" status="comb. nov.">
<emphasis id="B913EAD0FFCBFFC646A2F99AA1A1AB7A" box="[745,760,1578,1603]" italics="true" pageId="5" pageNumber="456">P</emphasis>
.
<emphasis id="B913EAD0FFCBFFC64743F999A0D8AB7A" box="[776,897,1577,1603]" italics="true" pageId="5" pageNumber="456">eratoformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057ABFFCBFFC647C1F99AA0ADAB7C" box="[906,1012,1578,1605]" pageId="5" pageNumber="456" rank="species">n. comb.</taxonomicNameLabel>
). By the whole of its features, and in particular its subglobose general curve, excurrent channel curled and jagged,
<taxonomicName id="4C674D41FFC8FFC544BCF99AA2C5AB7A" authorityName="Pacaud" authorityYear="2003" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[247,412,1577,1604]" class="Gastropoda" family="Cypraeidae" genus="Cypraea" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="457" phylum="Mollusca" rank="species" species="eratoformis" status="comb. nov.">
<emphasis id="B913EAD0FFC8FFC544BCF99AA25FAB7A" box="[247,262,1578,1603]" italics="true" pageId="6" pageNumber="457">P</emphasis>
.
<emphasis id="B913EAD0FFC8FFC54551F999A2C5AB7A" box="[282,412,1577,1603]" italics="true" pageId="6" pageNumber="457">eratoformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057ABFFC8FFC545E3F99AA14DAB7C" box="[424,532,1578,1605]" pageId="6" pageNumber="457" rank="species">n. comb.</taxonomicNameLabel>
seems to be closer to the recent typical species. Under ultraviolet light (
<figureCitation id="135C2A47FFC8FFC5453EF9DBA280ABBC" box="[373,473,1643,1669]" captionStart="FIG" captionStartId="6.[151,162,1442,1459]" captionTargetBox="[237,1174,174,1398]" captionTargetId="figure-69@6.[216,1179,144,1413]" captionTargetPageId="6" captionText="FIG. 3. — A, B, Pseudocypraea eratoformis (Hoernes &amp; Auinger, 1880) n. comb., Langhian (middle Miocene) of Lapugiu de Sus (Rumania), original pictures; C-E, Pseudocypraea dolini n. sp.; C, under ultraviolet light showing the trace of a colour pattern; D, E, ventral face and detail of the siphonal canal region. Scale bars: A, B, 3 mm; C-E, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5375308" httpUri="https://zenodo.org/record/5375308/files/figure.png" pageId="6" pageNumber="457">Fig. 3C</figureCitation>
)
<taxonomicName id="4C674D41FFC8FFC545A5F9DAA103ABBC" authority="Pacaud, 2003" authorityName="Pacaud" authorityYear="2003" box="[494,602,1642,1669]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="457" phylum="Mollusca" rank="species" species="dolini" status="sp. nov.">
<emphasis id="B913EAD0FFC8FFC545A5F9DAA2A4ABBA" box="[494,509,1642,1667]" italics="true" pageId="6" pageNumber="457">P</emphasis>
.
<emphasis id="B913EAD0FFC8FFC54659F9DBA103ABBC" box="[530,602,1643,1669]" italics="true" pageId="6" pageNumber="457">dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057ABFFC8FFC5462DF9DAA1F5ABBD" box="[614,684,1642,1668]" pageId="6" pageNumber="457" rank="species">n. sp.</taxonomicNameLabel>
shows spots along the periphery of the outer lip that are typical of the genus
<taxonomicName id="4C674D41FFC8FFC5400FF9FBA786AB5C" authorityName="Schilder" authorityYear="1927" box="[1092,1247,1611,1637]" class="Gastropoda" family="Ovulidae" genus="Pseudocypraea" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="457" phylum="Mollusca" rank="genus">
<emphasis id="B913EAD0FFC8FFC5400FF9FBA786AB5C" box="[1092,1247,1611,1637]" italics="true" pageId="6" pageNumber="457">Pseudocypraea</emphasis>
</taxonomicName>
(
<figureCitation id="135C2A47FFC8FFC54699F9DBA072ABBC" box="[722,811,1643,1669]" captionStart="FIG" captionStartId="8.[151,162,1407,1424]" captionTargetBox="[184,1219,171,1367]" captionTargetId="figure-86@8.[184,1219,122,1410]" captionTargetPageId="8" captionText="FIG. 5. — A-C, Pseudocypraea adamsonii (Sowerby, 1832) of Huahine Island (Polynésie) (MNHN-DSE) (leg. L. Dolin); D-F, Pseudocypraea exquisita Petuch, 1979 of Bohol Island (Philippines) (MNHN-DSE) (leg. L. Dolin). Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5375312" httpUri="https://zenodo.org/record/5375312/files/figure.png" pageId="6" pageNumber="457">Fig. 5E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>