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<document id="8B736F3B9CABE6EC23EE419E4B9A938C" ID-DOI="10.5281/zenodo.281089" ID-GBIF-Dataset="a24799b7-1a9a-461e-ae73-0591e74b4db5" ID-ISSN="1175-5326" ID-Zenodo-Dep="281089" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460481723120" checkinUser="plazi" docAuthor="Kury, Adriano B." docDate="2012" docId="03CE87D37D7AFFE8FF1DFC69FD25F96F" docLanguage="en" docName="zt03314p044.pdf" docOrigin="Zootaxa 3314" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Zannicranaus Kury, 2012, new genus" docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="FFF7FFAB7D7BFFE9FF8AFFA6FFD8FF90" masterDocTitle="A new genus of Cranaidae from Ecuador (Opiliones: Laniatores)" masterLastPageNumber="44" masterPageNumber="31" pageNumber="32" updateTime="1698307856339" updateUser="plazi">
<mods:mods id="E1FACB7C30D84F92EFDE31F44BAD3F03" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C8170D6CB7D04B2F57D97D196880063B">
<mods:title id="5F0FE84230FAB2494A4AC4F049C10AE3">A new genus of Cranaidae from Ecuador (Opiliones: Laniatores)</mods:title>
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<mods:name id="4D9CA0BC0A74ABE4B81CD76A9898175D" type="personal">
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<mods:namePart id="147885E8501459025E9E9E586A202E34">Kury, Adriano B.</mods:namePart>
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<mods:date id="3B8D70B1C2697061B39EDE3FC85CFEC3">2012</mods:date>
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<mods:classification id="FC253BFA947255A21D8FE16DCF0F853B">journal article</mods:classification>
<mods:identifier id="F56BFC266CCBB3A1BDD6E5DE47F4893D" type="DOI">10.5281/zenodo.281089</mods:identifier>
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<mods:identifier id="C8F95EE98A4F7711A744795829DA35ED" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CE87D37D7AFFE8FF1DFC69FD25F96F" ID-DOI="http://doi.org/10.5281/zenodo.5697370" ID-GBIF-Taxon="119517420" ID-Zenodo-Dep="5697370" LSID="urn:lsid:plazi:treatment:03CE87D37D7AFFE8FF1DFC69FD25F96F" httpUri="http://treatment.plazi.org/id/03CE87D37D7AFFE8FF1DFC69FD25F96F" lastPageNumber="32" pageId="1" pageNumber="32">
<subSubSection id="C37D654E7D7AFFE8FF1DFC69FE13FC79" box="[151,459,975,1001]" pageId="1" pageNumber="32" type="nomenclature">
<paragraph id="8BD836C57D7AFFE8FF1DFC69FE13FC79" blockId="1.[151,459,975,1001]" box="[151,459,975,1001]" pageId="1" pageNumber="32">
<heading id="D09081A97D7AFFE8FF1DFC69FE13FC79" bold="true" box="[151,459,975,1001]" fontSize="11" level="1" pageId="1" pageNumber="32" reason="1">
<emphasis id="B913EAD77D7AFFE8FF1DFC69FE13FC79" bold="true" box="[151,459,975,1001]" pageId="1" pageNumber="32">
<taxonomicName id="4C674D467D7AFFE8FF1DFC69FE9AFC79" ID-CoL="7Q6WR" box="[151,322,975,1001]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B913EAD77D7AFFE8FF1DFC69FE9AFC79" bold="true" box="[151,322,975,1001]" italics="true" pageId="1" pageNumber="32">Zannicranaus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057AC7D7AFFE8FEC3FC69FE13FC79" box="[329,459,975,1001]" pageId="1" pageNumber="32" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D654E7D7AFFE8FF1DFBB1FDC4FBE6" pageId="1" pageNumber="32" type="etymology">
<paragraph id="8BD836C57D7AFFE8FF1DFBB1FDC4FBE6" blockId="1.[151,1437,1046,1791]" pageId="1" pageNumber="32">
<emphasis id="B913EAD77D7AFFE8FF1DFBB1FEC6FBBF" bold="true" box="[151,286,1047,1071]" pageId="1" pageNumber="32">Etymology.</emphasis>
From Zanni, one of the masquerade stereotypes of Italian Commedia dell’arte, notable for having a big nose (which strongly resembles the spine in frontal hump which characterizes the genus) + pre-existing generic name
<taxonomicName id="4C674D467D7AFFE8FF50FBF9FEE2FBE6" box="[218,314,1119,1142]" class="Arachnida" family="Cranaidae" genus="Cranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAD77D7AFFE8FF50FBF9FEE2FBE6" box="[218,314,1119,1142]" italics="true" pageId="1" pageNumber="32">Cranaus</emphasis>
</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C37D654E7D7AFFE8FF43FB24FABAF94A" pageId="1" pageNumber="32" type="diagnosis">
<paragraph id="8BD836C57D7AFFE8FF43FB24FABAF94A" blockId="1.[151,1437,1046,1791]" pageId="1" pageNumber="32">
<emphasis id="B913EAD77D7AFFE8FF43FB24FE9BFB0B" bold="true" box="[201,323,1154,1179]" pageId="1" pageNumber="32">Diagnosis.</emphasis>
Outline of dorsal scutum in dorsal view guitar-shaped (widest part clearly displaced posteriorly) either with (
<taxonomicName id="4C674D467D7AFFE8FEAAFB0EFE7EFB2E" box="[288,422,1190,1215]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="morlacus">
<emphasis id="B913EAD77D7AFFE8FEAAFB0EFE7EFB2E" box="[288,422,1190,1215]" italics="true" pageId="1" pageNumber="32">Z. morlacus</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FE3EFB00FDDDFB2E" box="[436,517,1190,1214]" captionStart="FIGURES 21 – 23" captionStartId="8.[151,264,1945,1968]" captionTargetBox="[151,1436,465,1924]" captionTargetId="figure@8.[151,1436,465,1924]" captionTargetPageId="8" captionText="FIGURES 21 – 23. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. 21. Habitus, dorsal view; 22. Body, lateral view; 23. Body, dorsal view. Scale bars = 1 mm. Abbreviation: co = coda." httpUri="https://zenodo.org/record/281095/files/figure.png" pageId="1" pageNumber="32">Fig. 23</figureCitation>
) or without defined coda (
<taxonomicName id="4C674D467D7AFFE8FCBAFB0EFC04FB2E" box="[816,988,1190,1215]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monoclonius">
<emphasis id="B913EAD77D7AFFE8FCBAFB0EFC04FB2E" box="[816,988,1190,1215]" italics="true" pageId="1" pageNumber="32">Z. monoclonius</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FC61FB00FBF6FB2E" box="[1003,1070,1190,1214]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1775,1798]" captionTargetBox="[151,1436,195,1750]" captionTargetId="figure@3.[151,1436,195,1751]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Zannicranaus monoclonius sp. nov. 3 holotype, QCAZ AK 232, from Sibambe. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Body, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281090/files/figure.png" pageId="1" pageNumber="32">Fig. 3</figureCitation>
). Frontal hump of carapace with huge spine bent frontwards (
<figureCitation id="135C2A407D7AFFE8FE54FB6CFDF7FB72" box="[478,559,1226,1250]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1775,1798]" captionTargetBox="[151,1436,195,1750]" captionTargetId="figure@3.[151,1436,195,1751]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Zannicranaus monoclonius sp. nov. 3 holotype, QCAZ AK 232, from Sibambe. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Body, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281090/files/figure.png" pageId="1" pageNumber="32">Figs. 2</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FDB4FB6CFD86FB72" box="[574,606,1226,1250]" captionStart="FIGURES 21 – 23" captionStartId="8.[151,264,1945,1968]" captionTargetBox="[151,1436,465,1924]" captionTargetId="figure@8.[151,1436,465,1924]" captionTargetPageId="8" captionText="FIGURES 21 – 23. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. 21. Habitus, dorsal view; 22. Body, lateral view; 23. Body, dorsal view. Scale bars = 1 mm. Abbreviation: co = coda." httpUri="https://zenodo.org/record/281095/files/figure.png" pageId="1" pageNumber="32">22</figureCitation>
). Cheliceral bulla of male with mid-posterior dorsal erect spine (
<figureCitation id="135C2A407D7AFFE8FACFFB6CFA4FFB72" box="[1349,1431,1226,1250]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1775,1798]" captionTargetBox="[151,1436,195,1750]" captionTargetId="figure@3.[151,1436,195,1751]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Zannicranaus monoclonius sp. nov. 3 holotype, QCAZ AK 232, from Sibambe. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Body, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281090/files/figure.png" pageId="1" pageNumber="32">Figs. 2</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FF1DFB48FF6FFA96" box="[151,183,1262,1286]" captionStart="FIGURES 21 – 23" captionStartId="8.[151,264,1945,1968]" captionTargetBox="[151,1436,465,1924]" captionTargetId="figure@8.[151,1436,465,1924]" captionTargetPageId="8" captionText="FIGURES 21 – 23. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. 21. Habitus, dorsal view; 22. Body, lateral view; 23. Body, dorsal view. Scale bars = 1 mm. Abbreviation: co = coda." httpUri="https://zenodo.org/record/281095/files/figure.png" pageId="1" pageNumber="32">22</figureCitation>
). Tibia-tarsus of pedipalpus rotated ectally (
<figureCitation id="135C2A407D7AFFE8FD22FB48FCDDFA96" box="[680,773,1262,1286]" captionStart="FIGURES 11 – 13" captionStartId="5.[151,264,1420,1443]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@5.[151,1436,193,1399]" captionTargetPageId="5" captionText="FIGURES 11 – 13. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Left pedipalpus: 11. Trochanter-femur, ventral view; 12. Patella-tarsus, ventral view; 13. Trochanter-femur, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281093/files/figure.png" pageId="1" pageNumber="32">Figs. 11</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FC98FB48FCEAFA96" box="[786,818,1262,1286]" captionStart="FIGURES 30 – 32" captionStartId="10.[151,264,1239,1262]" captionTargetBox="[151,1436,219,1179]" captionTargetId="figure@10.[151,1436,219,1180]" captionTargetPageId="10" captionText="FIGURES 30 – 32. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Right pedipalpus: 30. Trochanter-femur, ventral view; 31. Patella-tarsus, ventral view; 32. Trochanter-femur, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281098/files/figure.png" pageId="1" pageNumber="32">30</figureCitation>
); femur-tibia with dorsal rounded tubercles (
<figureCitation id="135C2A407D7AFFE8FAA1FB48FA4FFA96" box="[1323,1431,1262,1286]" captionStart="FIGURES 7 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[151,1436,927,1882]" captionTargetId="figure@4.[151,1436,927,1900]" captionTargetPageId="4" captionText="FIGURES 7 – 10. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Left pedipalpus: 7. Dorsomesal view; 8. Mesal view; 9. Ectal view; 10. Dorso-ectal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281092/files/figure.png" pageId="1" pageNumber="32">Figs. 7–8</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FF1DFAB4FF6DFABA" box="[151,181,1298,1322]" captionStart="FIGURES 11 – 13" captionStartId="5.[151,264,1420,1443]" captionTargetBox="[151,1436,193,1398]" captionTargetId="figure@5.[151,1436,193,1399]" captionTargetPageId="5" captionText="FIGURES 11 – 13. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Left pedipalpus: 11. Trochanter-femur, ventral view; 12. Patella-tarsus, ventral view; 13. Trochanter-femur, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281093/files/figure.png" pageId="1" pageNumber="32">13</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FF49FAB4FED3FABA" box="[195,267,1298,1322]" captionStart="FIGURES 26 – 29" captionStartId="9.[151,264,1983,2006]" captionTargetBox="[151,1436,951,1961]" captionTargetId="figure@9.[151,1436,928,1962]" captionTargetPageId="9" captionText="FIGURES 26 – 29. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Right pedipalpus: 26. Dorso-mesal view; 27. Mesal view; 28. Ectal view; 29. Dorso-ectal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281097/files/figure.png" pageId="1" pageNumber="32">26–27</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FE93FAB4FEE1FABA" box="[281,313,1298,1322]" captionStart="FIGURES 30 – 32" captionStartId="10.[151,264,1239,1262]" captionTargetBox="[151,1436,219,1179]" captionTargetId="figure@10.[151,1436,219,1180]" captionTargetPageId="10" captionText="FIGURES 30 – 32. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Right pedipalpus: 30. Trochanter-femur, ventral view; 31. Patella-tarsus, ventral view; 32. Trochanter-femur, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281098/files/figure.png" pageId="1" pageNumber="32">32</figureCitation>
). Femur with dorso-apical short spine (
<figureCitation id="135C2A407D7AFFE8FD77FAB4FC96FABA" box="[765,846,1298,1322]" captionStart="FIGURES 7 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[151,1436,927,1882]" captionTargetId="figure@4.[151,1436,927,1900]" captionTargetPageId="4" captionText="FIGURES 7 – 10. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Left pedipalpus: 7. Dorsomesal view; 8. Mesal view; 9. Ectal view; 10. Dorso-ectal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281092/files/figure.png" pageId="1" pageNumber="32">Figs. 8</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FCD6FAB4FCA3FABA" box="[860,891,1298,1322]" captionStart="FIGURES 26 – 29" captionStartId="9.[151,264,1983,2006]" captionTargetBox="[151,1436,951,1961]" captionTargetId="figure@9.[151,1436,928,1962]" captionTargetPageId="9" captionText="FIGURES 26 – 29. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Right pedipalpus: 26. Dorso-mesal view; 27. Mesal view; 28. Ectal view; 29. Dorso-ectal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281097/files/figure.png" pageId="1" pageNumber="32">27</figureCitation>
), none of pedipalpal articles specially elongate. Male calcaneus I twice as long as astragalus and thickened. Coxa IV of male with pair of ventro-distal paramedian acuminate tubercles (
<figureCitation id="135C2A407D7AFFE8FE0FFAFCFE0CFAE2" box="[389,468,1370,1394]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1775,1798]" captionTargetBox="[151,1436,195,1750]" captionTargetId="figure@3.[151,1436,195,1751]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Zannicranaus monoclonius sp. nov. 3 holotype, QCAZ AK 232, from Sibambe. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Body, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281090/files/figure.png" pageId="1" pageNumber="32">Figs. 2</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FE6AFAFCFDD8FAE2" box="[480,512,1370,1394]" captionStart="FIGURES 21 – 23" captionStartId="8.[151,264,1945,1968]" captionTargetBox="[151,1436,465,1924]" captionTargetId="figure@8.[151,1436,465,1924]" captionTargetPageId="8" captionText="FIGURES 21 – 23. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. 21. Habitus, dorsal view; 22. Body, lateral view; 23. Body, dorsal view. Scale bars = 1 mm. Abbreviation: co = coda." httpUri="https://zenodo.org/record/281095/files/figure.png" pageId="1" pageNumber="32">22</figureCitation>
). Femur IV of male from substraight (
<figureCitation id="135C2A407D7AFFE8FC21FAFCFC24FAE2" box="[939,1020,1370,1394]" captionStart="FIGURES 21 – 23" captionStartId="8.[151,264,1945,1968]" captionTargetBox="[151,1436,465,1924]" captionTargetId="figure@8.[151,1436,465,1924]" captionTargetPageId="8" captionText="FIGURES 21 – 23. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. 21. Habitus, dorsal view; 22. Body, lateral view; 23. Body, dorsal view. Scale bars = 1 mm. Abbreviation: co = coda." httpUri="https://zenodo.org/record/281095/files/figure.png" pageId="1" pageNumber="32">Fig. 21</figureCitation>
) to slightly sinuous (
<figureCitation id="135C2A407D7AFFE8FB62FAFCFAF0FAE2" box="[1256,1320,1370,1394]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1775,1798]" captionTargetBox="[151,1436,195,1750]" captionTargetId="figure@3.[151,1436,195,1751]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Zannicranaus monoclonius sp. nov. 3 holotype, QCAZ AK 232, from Sibambe. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Body, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281090/files/figure.png" pageId="1" pageNumber="32">Fig. 1</figureCitation>
) and with few weak spines. Tarsi III–IV with weak scopula, tarsal claws unpectinated. Tarsal counts, male: 7(3)/12-16(3)/7-9/ 8-9, female: 6(3)/13(3)/8/9. Truncus penis either straight (
<taxonomicName id="4C674D467D7AFFE8FCAEFA02FC72FA2B" box="[804,938,1443,1467]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="morlacus">
<emphasis id="B913EAD77D7AFFE8FCAEFA02FC72FA2B" box="[804,938,1443,1467]" italics="true" pageId="1" pageNumber="32">Z. morlacus</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FC33FA04FBD2FA2A" box="[953,1034,1442,1466]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">Fig. 33</figureCitation>
) or clearly angled (
<taxonomicName id="4C674D467D7AFFE8FB62FA02FA4DFA2B" box="[1256,1429,1443,1467]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monoclonius">
<emphasis id="B913EAD77D7AFFE8FB62FA02FA4DFA2B" box="[1256,1429,1443,1467]" italics="true" pageId="1" pageNumber="32">Z. monoclonius</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FF1DFA60FF32FA4E" box="[151,234,1478,1502]" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Fig. 14</figureCitation>
). Ventral plate of penis strongly angled against truncus (
<figureCitation id="135C2A407D7AFFE8FCEEFA60FC19FA4E" box="[868,961,1478,1502]" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Figs. 14</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FC44FA60FC35FA4E" box="[974,1005,1478,1502]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">33</figureCitation>
), with wide and shallow V-cleft (
<figureCitation id="135C2A407D7AFFE8FAE9FA60FF07F992" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Figs. 15–16</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FF66FA4CFEEEF992" box="[236,310,1514,1538]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">34–35</figureCitation>
), either guitar-shaped (
<taxonomicName id="4C674D467D7AFFE8FDB4FA4AFD35F993" box="[574,749,1515,1539]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monoclonius">
<emphasis id="B913EAD77D7AFFE8FDB4FA4AFD35F993" box="[574,749,1515,1539]" italics="true" pageId="1" pageNumber="32">Z. monoclonius</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FD77FA4CFC96F992" box="[765,846,1514,1538]" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Fig. 15</figureCitation>
) or subrectangular (
<taxonomicName id="4C674D467D7AFFE8FBBDFA4AFB66F993" box="[1079,1214,1515,1539]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="morlacus">
<emphasis id="B913EAD77D7AFFE8FBBDFA4AFB66F993" box="[1079,1214,1515,1539]" italics="true" pageId="1" pageNumber="32">Z. morlacus</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FB44FA4CFAFBF992" box="[1230,1315,1514,1538]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">Fig. 34</figureCitation>
), and with two lateral patches of scale-setae on ventral surface (
<figureCitation id="135C2A407D7AFFE8FD6DF9A8FC9CF9B6" box="[743,836,1550,1574]" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Figs. 15</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FCDAF9A8FCA8F9B6" box="[848,880,1550,1574]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">34</figureCitation>
). Glans sac very long, fanfold, with small conical dorsal process (
<figureCitation id="135C2A407D7AFFE8FEC3F994FE0DF9DA" box="[329,469,1586,1610]" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Figs. 14, 16</figureCitation>
,
<figureCitation id="135C2A407D7AFFE8FE69F994FDF6F9DA" box="[483,558,1586,1610]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">33, 38</figureCitation>
). Stylus either straight, growing thinner distally (
<taxonomicName id="4C674D467D7AFFE8FBE8F992FAC9F9DB" box="[1122,1297,1587,1611]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monoclonius">
<emphasis id="B913EAD77D7AFFE8FBE8F992FAC9F9DB" box="[1122,1297,1587,1611]" italics="true" pageId="1" pageNumber="32">Z. monoclonius</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FAABF994FAABF9DA" box="[1313,1395,1586,1610]" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Fig. 20</figureCitation>
) or uniformly thin, sinuous (
<taxonomicName id="4C674D467D7AFFE8FE38F9FEFDE1F9FE" box="[434,569,1622,1647]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="morlacus">
<emphasis id="B913EAD77D7AFFE8FE38F9FEFDE1F9FE" box="[434,569,1622,1647]" italics="true" pageId="1" pageNumber="32">Z. morlacus</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FDC3F9F0FD44F9FE" box="[585,668,1622,1646]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">Fig. 38</figureCitation>
). Stylus head either simple, elongate-erythrocyte-shaped (
<taxonomicName id="4C674D467D7AFFE8FABBF9FEFF64F903" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="morlacus">
<emphasis id="B913EAD77D7AFFE8FABBF9FEFF64F903" italics="true" pageId="1" pageNumber="32">Z. morlacus</emphasis>
</taxonomicName>
,
<figureCitation id="135C2A407D7AFFE8FF41F9DCFE8AF902" box="[203,338,1658,1682]" captionStart="FIGURES 33 – 38" captionStartId="11.[151,264,1610,1633]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@11.[151,1436,193,1589]" captionTargetPageId="11" captionText="FIGURES 33 – 38. Zannicranaus morlacus sp. nov. 3 holotype, QCAZ AK 0 0 29, from Molleturo. Male genitalia: 33. Distal part of penis, lateral view; 34. Same, ventral view; 35. Same, dorsal view; 36. Stylar cap, ventral view; 37. Stylar cap, lateral view; 38. Glans, lateral view. Scale bars = 100 µm (Figs. 33 – 35), 10 µm (Figs. 36 – 37) and 50 µm (Fig. 38)." httpUri="https://zenodo.org/record/281099/files/figure.png" pageId="1" pageNumber="32">Figs. 36–37</figureCitation>
) or complex, with 2 stylar distally-villose caps (
<taxonomicName id="4C674D467D7AFFE8FCF3F9DAFBFEF903" box="[889,1062,1659,1683]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monoclonius">
<emphasis id="B913EAD77D7AFFE8FCF3F9DAFBFEF903" box="[889,1062,1659,1683]" italics="true" pageId="1" pageNumber="32">Z. monoclonius</emphasis>
</taxonomicName>
): dorsal horseshoe-shaped (
<figureCitation id="135C2A407D7AFFE8FAE9F9DCFF06F926" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Figs. 19–20</figureCitation>
) and ventral elongate tapering foliaceous (
<figureCitation id="135C2A407D7AFFE8FD30F938FCB4F926" box="[698,876,1694,1718]" captionStart="FIGURES 14 – 20" captionStartId="6.[151,264,1877,1900]" captionTargetBox="[151,1436,193,1855]" captionTargetId="figure@6.[151,1436,193,1856]" captionTargetPageId="6" captionText="FIGURES 14 – 20. Zannicranaus monoclonius sp. nov. 3 paratype, MNRJ 19322, from Sibambe. Male genitalia: 14. Distal part of penis, lateral view; 15. Same, ventral view; 16. Same, dorsal view. 17. Tip of stylus, lateral view; 18. Stylar cap, ventral view; 19. Stylar cap, dorsal view; 20. Stylus, dorso-apical view. Scale bars = 100 µm (Figs. 14 – 16), 10 µm (Figs. 17, 19) and 20 µm (Figs. 18, 20). Abbreviation: sc = stylar cap." httpUri="https://zenodo.org/record/281094/files/figure.png" pageId="1" pageNumber="32">Figs. 17–18, 20</figureCitation>
). Sexual dimorphism evident in shape of carapace and basichelicerite (compare
<figureCitation id="135C2A407D7AFFE8FE6AF964FDF4F94A" box="[480,556,1730,1754]" captionStart="FIGURES 1 – 3" captionStartId="3.[151,264,1775,1798]" captionTargetBox="[151,1436,195,1750]" captionTargetId="figure@3.[151,1436,195,1751]" captionTargetPageId="3" captionText="FIGURES 1 – 3. Zannicranaus monoclonius sp. nov. 3 holotype, QCAZ AK 232, from Sibambe. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Body, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281090/files/figure.png" pageId="1" pageNumber="32">Figs. 3</figureCitation>
and
<figureCitation id="135C2A407D7AFFE8FDE8F964FDAAF94A" box="[610,626,1730,1754]" captionStart="FIGURES 4 – 6" captionStartId="4.[151,264,824,847]" captionTargetBox="[151,1436,193,802]" captionTargetId="figure@4.[151,1436,193,803]" captionTargetPageId="4" captionText="FIGURES 4 – 6. Zannicranaus monoclonius sp. nov. 3 holotype, QCAZ AK 232, from Sibambe. 4. Body, frontal view; 5. Body, ventral view. 6. Ƥ allotype, QCAZ AK 232, from Sibambe, body, dorsal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/281091/files/figure.png" pageId="1" pageNumber="32">6</figureCitation>
), much stronger in male (females known only for
<taxonomicName id="4C674D467D7AFFE8FB2FF962FA89F94B" box="[1189,1361,1731,1755]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monoclonius">
<emphasis id="B913EAD77D7AFFE8FB2FF962FA89F94B" box="[1189,1361,1731,1755]" italics="true" pageId="1" pageNumber="32">Z. monoclonius</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D654E7D7AFFE8FF43F941FD25F96F" box="[201,765,1766,1791]" pageId="1" pageNumber="32" type="materials_examined">
<paragraph id="8BD836C57D7AFFE8FF43F941FD25F96F" blockId="1.[151,1437,1046,1791]" box="[201,765,1766,1791]" pageId="1" pageNumber="32">
<emphasis id="B913EAD77D7AFFE8FF43F941FEBCF96F" bold="true" box="[201,356,1766,1791]" pageId="1" pageNumber="32">
<typeStatus id="54DC88677D7AFFE8FF43F941FEDCF96F" box="[201,260,1767,1791]" pageId="1" pageNumber="32">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C674D467D7AFFE8FEE1F94EFD44F96E" box="[363,668,1766,1791]" class="Arachnida" family="Cranaidae" genus="Zannicranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="monoclonius" status="sp. nov.">
<emphasis id="B913EAD77D7AFFE8FEE1F94EFD44F96E" box="[363,668,1766,1791]" italics="true" pageId="1" pageNumber="32">Zannicranaus monoclonius</emphasis>
</taxonomicName>
<emphasis id="B913EAD77D7AFFE8FD2EF941FD25F96F" bold="true" box="[676,765,1767,1791]" pageId="1" pageNumber="32">
<taxonomicNameLabel id="A22057AC7D7AFFE8FD2EF941FD25F96F" box="[676,765,1767,1791]" pageId="1" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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