349 lines
45 KiB
XML
349 lines
45 KiB
XML
<document id="98B00C1CEA557182DD96E404789F3ED1" ID-DOI="10.11646/zootaxa.4996.3.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="5074540" ID-ZooBank="0F2F6614-08E3-46A9-AF21-14F7BB79A4CE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625555920056" checkinUser="plazi" docAuthor="Inoue, Shota & Nomura, Shûhei" docDate="2021" docId="03CE87A5FFD89465FF03AA26675723D6" docLanguage="en" docName="zootaxa.4996.3.11.pdf" docOrigin="Zootaxa 4996 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudophanias spinicornis Inoue & Nomura 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="595" masterDocId="FFF7FFDDFFD99461FF94AC2662362777" masterDocTitle="Two new species of Pseudophanias Raffray from Myanmar (Coleoptera Staphylinidae: Pselaphinae)" masterLastPageNumber="599" masterPageNumber="591" pageNumber="592" updateTime="1698992086504" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BF84720A851F5F76E5E681CC46C7ED16">
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<mods:title id="929DB5BF34B006B6A126448AA971BEE8">Two new species of Pseudophanias Raffray from Myanmar (Coleoptera Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="A02C2F7A077DF191B181ADDA081463CE" type="personal">
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<mods:roleTerm id="7BFF1056022F7E5DE4D1E3C039210B3C">Author</mods:roleTerm>
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<mods:namePart id="51195BC5D390386C81D694F71D8866F4">Inoue, Shota</mods:namePart>
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<mods:affiliation id="71713AEC7678DBE6E44025136A510924">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan. & The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:roleTerm id="112F7193494F23739277ED0AB95BAE7A">Author</mods:roleTerm>
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<mods:namePart id="0B76ABE9DC4ABFF15633B0BACBA3888C">Nomura, Shûhei</mods:namePart>
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<mods:nameIdentifier id="8868C9D2CF7FA67E7E9245086A0B84CD" type="ORCID">0000-0003-1086-5078</mods:nameIdentifier>
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<mods:affiliation id="A6E662BB5622508A9C6CEB884CD42942">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki, 305 - 0005 Japan. nomura @ kahaku. go. jp; https: // orcid. org / 0000 - 0003 - 1086 - 5078</mods:affiliation>
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<mods:nameIdentifier id="C1AC2100827CDC160EFAE59D73048153" type="email">nomura@kahaku.go.jp</mods:nameIdentifier>
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<mods:title id="CD8C5FF1C54348108D1EBA318F62A889">Zootaxa</mods:title>
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<mods:part id="D3C7BBA168DBC082A1B4866FC27225C0">
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<mods:date id="41A5ED56A7C36F492A1934C183A09781">2021</mods:date>
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<mods:number id="EB894A1D35058317FA963CD7855AA609">2021-07-06</mods:number>
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<mods:number id="033241791BB08D8A9B1FDEF8F5B85B97">4996</mods:number>
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<subSubSection id="C37D6538FFD89460FF03AA266077216C" box="[151,577,1536,1563]" pageId="1" pageNumber="592" type="nomenclature">
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<paragraph id="8BD836B3FFD89460FF03AA266077216C" blockId="1.[151,577,1536,1599]" box="[151,577,1536,1563]" pageId="1" pageNumber="592">
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<heading id="D09081DFFFD89460FF03AA266077216C" bold="true" box="[151,577,1536,1563]" fontSize="11" level="1" pageId="1" pageNumber="592" reason="1">
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<emphasis id="B913EAA1FFD89460FF03AA266077216C" bold="true" box="[151,577,1536,1563]" pageId="1" pageNumber="592">
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<taxonomicName id="4C674D30FFD89460FF03AA2663EB216D" ID-CoL="8Y8T3" authority="Inoue & Nomura, 2021" authorityName="Inoue & Nomura" authorityYear="2021" box="[151,477,1536,1563]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="spinicornis" status="sp. nov.">
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<emphasis id="B913EAA1FFD89460FF03AA2663EB216D" bold="true" box="[151,477,1536,1563]" italics="true" pageId="1" pageNumber="592">Pseudophanias spinicornis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22057DAFFD89460FE70AA276077216C" box="[484,577,1537,1563]" pageId="1" pageNumber="592" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6538FFD89460FF03AA02633D2148" box="[151,267,1572,1599]" pageId="1" pageNumber="592" type="description">
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<paragraph id="8BD836B3FFD89460FF03AA02633D2148" blockId="1.[151,577,1536,1599]" box="[151,267,1572,1599]" pageId="1" pageNumber="592">
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(
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<figureCitation id="135C2A36FFD89460FF0BAA0263342148" box="[159,258,1572,1599]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1369,1393]" captionStartId-1="2.[151,250,1649,1673]" captionStartId-2="3.[151,250,1782,1806]" captionTargetBox-0="[160,1430,461,1338]" captionTargetBox-1="[159,1435,187,1622]" captionTargetBox-2="[157,1427,389,1749]" captionTargetId-0="figure-155@1.[151,1436,453,1345]" captionTargetId-1="figure-27@2.[151,1436,181,1625]" captionTargetId-2="figure-106@3.[151,1436,381,1759]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Dorsal habitus of Pseudophanias spinicornis sp. nov. A. Male. B. Female. Scale bar: 1.0 mm." captionText-1="FIGURE 2. Morphological details of Pseudophanias spinicornis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Mesotibia. F. Metatibia. G. SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G." captionText-2="FIGURE 3. Male genitalia of Pseudophanias spinicornis sp. nov. A, B. Ventral view. C. Dorsal view. D. Lateral view. E. Another lateral view. F. Tergite VIII. G. Sternite VIII. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5074544" figureDoi-1="http://doi.org/10.5281/zenodo.5074546" figureDoi-2="http://doi.org/10.5281/zenodo.5074550" httpUri-0="https://zenodo.org/record/5074544/files/figure.png" httpUri-1="https://zenodo.org/record/5074546/files/figure.png" httpUri-2="https://zenodo.org/record/5074550/files/figure.png" pageId="1" pageNumber="592">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6538FFD89460FF03AA4B61F6204C" pageId="1" pageNumber="592" type="materials_examined">
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<paragraph id="8BD836B3FFD89460FF03AA4B60582184" blockId="1.[151,1437,1644,2030]" pageId="1" pageNumber="592">
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<emphasis id="B913EAA1FFD89460FF03AA4B637321F1" bold="true" box="[151,325,1644,1671]" pageId="1" pageNumber="592">Type material.</emphasis>
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<materialsCitation id="3B0F3CEEFFD89460FEDFAA4A60582184" collectingDate="2018-11-16" collectorName="S. Nomura" country="Myanmar" determinerName="S. Nomura" elevation="100" latitude="14.739445" location="Nature Reserve" longLatPrecision="21" longitude="98.195" pageId="1" pageNumber="592" specimenCount="2" specimenCount-male="2" stateProvince="Taninthayi" typeStatus="holotype">
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<emphasis id="B913EAA1FFD89460FEDFAA4A63C221F1" bold="true" box="[331,500,1644,1670]" pageId="1" pageNumber="592">
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<typeStatus id="54DC8811FFD89460FEDFAA4A63D821F1" box="[331,494,1644,1670]" pageId="1" pageNumber="592" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D61FD3AFFD89460FE6EAA4B602721F0" box="[506,529,1645,1671]" pageId="1" pageNumber="592" type="male">♂</specimenCount>
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, “
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<collectingRegion id="49A3F851FFD89460FDB2AA4A609C21F1" box="[550,682,1644,1670]" country="Myanmar" name="Taninthayi" pageId="1" pageNumber="592">Tanintharyi</collectingRegion>
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/ (by sifting leaf litter, D4, ca. /
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</quantity>
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),
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<collectingRegion id="49A3F851FFD89460FA88AA4A62DE21DD" country="Myanmar" name="Taninthayi" pageId="1" pageNumber="592">Tanintharyi Region</collectingRegion>
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/ [S
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<collectingCountry id="F3707623FFD89460FEB2AAB6638A21DC" box="[294,444,1680,1707]" name="Myanmar" pageId="1" pageNumber="592">MYANMAR</collectingCountry>
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] /
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<geoCoordinate id="EE535074FFD89460FDF8AAB460DB21DA" box="[620,749,1682,1709]" degrees="98" direction="east" minutes="11" orientation="longitude" pageId="1" pageNumber="592" precision="15" seconds="42" value="98.195">E98°11′42″</geoCoordinate>
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/
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<date id="FFD91073FFD89460FC9CAAB661B521DD" box="[776,899,1680,1706]" pageId="1" pageNumber="592" value="2018-11-16">
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<collectingDate id="EF9DE99BFFD89460FC9CAAB661B521DD" box="[776,899,1680,1706]" pageId="1" pageNumber="592" value="2018-11-16">16.xi.2018</collectingDate>
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</date>
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,
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<collectorName id="26925365FFD89460FC05AAB7663B21DC" box="[913,1037,1681,1707]" pageId="1" pageNumber="592">S. Nomura</collectorName>
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leg. // FAUNA MYANMARICA /
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<taxonomicName id="4C674D30FFD89460FF03AA92630221B9" box="[151,308,1716,1742]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
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,
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<taxonomicName id="4C674D30FFD89460FEABAA9263F321B9" authorityName="Latreille" authorityYear="1802" box="[319,453,1716,1742]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
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/
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<taxonomicName id="4C674D30FFD89460FE4FAA9260F421B8" box="[475,706,1716,1743]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="undefined-1">
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<emphasis id="B913EAA1FFD89460FE4FAA9260B021B9" box="[475,646,1716,1742]" italics="true" pageId="1" pageNumber="592">Pseudophanias</emphasis>
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sp. 1
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</taxonomicName>
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/
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<determinerName id="7E04DA77FFD89460FD4DAA93616421B8" box="[729,850,1717,1743]" pageId="1" pageNumber="592">S. Nomura</determinerName>
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det., 2019 //
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<typeStatus id="54DC8811FFD89460FC73AA93664D21B8" box="[999,1147,1717,1743]" pageId="1" pageNumber="592" type="holotype">HOLOTYPE</typeStatus>
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(red) /
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<specimenCount id="9D61FD3AFFD89460FB5AAA9366D321B8" box="[1230,1253,1717,1743]" pageId="1" pageNumber="592" type="male">♂</specimenCount>
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,
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<taxonomicName id="4C674D30FFD89460FB65AA9263252185" authorityName="Inoue & Nomura" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="spinicornis">
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<emphasis id="B913EAA1FFD89460FB65AA9263252185" italics="true" pageId="1" pageNumber="592">Pseudophanias spinicornis</emphasis>
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</taxonomicName>
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/ des. Inoue &
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<bibRefCitation id="EFF64B42FFD89460FE54AAFF60522184" author="Nomura, S. & Aung, M. M." box="[448,612,1753,1779]" pageId="1" pageNumber="592" pagination="19 - 29" refId="ref4067" refString="Nomura, S. & Aung, M. M. (2021) Inventory studies on the subfamily Pselaphinae (Coleoptera, Staphylinidae) of Myanmar Part 3: A list of collected species in Tanintharyi region in November 2018. Bulletin of the National Museum of Nature and Science, Series A, 47 (1), 19 - 29." type="journal article" year="2021">Nomura, 2021</bibRefCitation>
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”
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</materialsCitation>
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</paragraph>
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<paragraph id="8BD836B3FFD89460FF53AADA61F6204C" blockId="1.[151,1437,1644,2030]" pageId="1" pageNumber="592">
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<materialsCitation id="3B0F3CEEFFD89460FF53AADA618A204C" collectingDate="2018-11-16" collectorName="S. Nomura" country="Myanmar" determinerName="S. Nomura" elevation="100" latitude="14.739445" location="Nature Reserve" longLatPrecision="21" longitude="98.195" pageId="1" pageNumber="592" specimenCount="3" specimenCount-female="3" stateProvince="Taninthayi" typeStatus="paratype">
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<emphasis id="B913EAA1FFD89460FF53AADA63432061" bold="true" box="[199,373,1788,1814]" pageId="1" pageNumber="592">
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<typeStatus id="54DC8811FFD89460FF53AADA63592061" box="[199,367,1788,1814]" pageId="1" pageNumber="592" type="paratype">PARATYPES</typeStatus>
|
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:
|
||
</emphasis>
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<specimenCount id="9D61FD3AFFD89460FEE9AADB63802060" box="[381,438,1789,1815]" pageId="1" pageNumber="592" type="female">2♀♀</specimenCount>
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, same data as holotype. Each
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||
<typeStatus id="54DC8811FFD89460FC99AADB615B2060" box="[781,877,1789,1815]" pageId="1" pageNumber="592" type="paratype">paratype</typeStatus>
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is pinned with the following label: “
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||
<typeStatus id="54DC8811FFD89460FA9AAADB67AB2060" box="[1294,1437,1789,1815]" pageId="1" pageNumber="592" type="paratype">PARATYPE</typeStatus>
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(yellow) /
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||
<specimenCount id="9D61FD3AFFD89460FE98AB076317204C" box="[268,289,1825,1851]" pageId="1" pageNumber="592" type="female">♀</specimenCount>
|
||
,
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||
<taxonomicName id="4C674D30FFD89460FEBBAB06606B204D" authorityName="Inoue & Nomura" authorityYear="2021" box="[303,605,1824,1850]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="spinicornis">
|
||
<emphasis id="B913EAA1FFD89460FEBBAB06606B204D" box="[303,605,1824,1850]" italics="true" pageId="1" pageNumber="592">Pseudophanias spinicornis</emphasis>
|
||
</taxonomicName>
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||
/ des. Inoue &
|
||
<bibRefCitation id="EFF64B42FFD89460FC9EAB076187204C" author="Nomura, S. & Aung, M. M." box="[778,945,1825,1851]" pageId="1" pageNumber="592" pagination="19 - 29" refId="ref4067" refString="Nomura, S. & Aung, M. M. (2021) Inventory studies on the subfamily Pselaphinae (Coleoptera, Staphylinidae) of Myanmar Part 3: A list of collected species in Tanintharyi region in November 2018. Bulletin of the National Museum of Nature and Science, Series A, 47 (1), 19 - 29." type="journal article" year="2021">Nomura, 2021</bibRefCitation>
|
||
”
|
||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C37D6538FFD89460FF53AB6261312099" pageId="1" pageNumber="592" type="diagnosis">
|
||
<paragraph id="8BD836B3FFD89460FF53AB6261312099" blockId="1.[151,1437,1644,2030]" pageId="1" pageNumber="592">
|
||
<emphasis id="B913EAA1FFD89460FF53AB6263742029" bold="true" box="[199,322,1860,1886]" pageId="1" pageNumber="592">Diagnosis.</emphasis>
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The new species is readily distinguished from the other members of
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<taxonomicName id="4C674D30FFD89460FBBAAB6266EF2029" authorityName="Raffray" authorityYear="1890" box="[1070,1241,1860,1886]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B913EAA1FFD89460FBBAAB6266EF2029" box="[1070,1241,1860,1886]" italics="true" pageId="1" pageNumber="592">Pseudophanias</emphasis>
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||
</taxonomicName>
|
||
by a combination of the following characteristics: antennomeres 7 and 10 ventrally spinose in male; pronotum densely and coarsely punctate, with small conical spine anterior to median fovea; mesotibiae and metatibiae modified in male; tergite IV with pair of short discal carinae; aedeagus asymmetrical; median lobe strongly curved in lateral view, membranous laterally, narrowed posteriorly, then broadened apically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF18663BFFDB9463FF03AA5763612182" ID-DOI="http://doi.org/10.5281/zenodo.5074546" ID-Zenodo-Dep="5074546" httpUri="https://zenodo.org/record/5074546/files/figure.png" pageId="2" pageNumber="593" startId="2.[151,250,1649,1673]" targetBox="[159,1435,187,1622]" targetPageId="2">
|
||
<paragraph id="8BD836B3FFDB9463FF03AA5763612182" blockId="2.[151,1437,1649,1781]" pageId="2" pageNumber="593">
|
||
<emphasis id="B913EAA1FFDB9463FF03AA57632321FE" bold="true" box="[151,277,1649,1673]" pageId="2" pageNumber="593">FIGURE 2.</emphasis>
|
||
Morphological details of
|
||
<taxonomicName id="4C674D30FFDB9463FDB1AA57610C21FE" authority="Inoue & Nomura, 2021" authorityName="Inoue & Nomura" authorityYear="2021" box="[549,826,1649,1673]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="species" species="spinicornis" status="sp. nov.">
|
||
<emphasis id="B913EAA1FFDB9463FDB1AA57610C21FE" box="[549,826,1649,1673]" italics="true" pageId="2" pageNumber="593">Pseudophanias spinicornis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EAA1FFDB9463FCD6AA5761A321FE" bold="true" box="[834,917,1649,1673]" pageId="2" pageNumber="593">
|
||
<taxonomicNameLabel id="A22057DAFFDB9463FCD6AA5761A321FE" box="[834,917,1649,1673]" pageId="2" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male.
|
||
<emphasis id="B913EAA1FFDB9463FC70AA5761CB21FE" bold="true" box="[996,1021,1649,1673]" pageId="2" pageNumber="593">A.</emphasis>
|
||
Ventral habitus.
|
||
<emphasis id="B913EAA1FFDB9463FB3BAA5766F021FE" bold="true" box="[1199,1222,1649,1673]" pageId="2" pageNumber="593">B.</emphasis>
|
||
Head and pronotum in lateral view.
|
||
<emphasis id="B913EAA1FFDB9463FEAFAAB3636521DA" bold="true" box="[315,339,1685,1709]" pageId="2" pageNumber="593">C.</emphasis>
|
||
SEM image of antenna in ventral view.
|
||
<emphasis id="B913EAA1FFDB9463FD6BAAB3612E21DA" bold="true" box="[767,792,1685,1709]" pageId="2" pageNumber="593">D.</emphasis>
|
||
SEM image of metaventrite in ventral view.
|
||
<emphasis id="B913EAA1FFDB9463FB60AAB3673A21DA" bold="true" box="[1268,1292,1685,1709]" pageId="2" pageNumber="593">E.</emphasis>
|
||
Mesotibia.
|
||
<emphasis id="B913EAA1FFDB9463FA1DAAB367AA21DA" bold="true" box="[1417,1436,1685,1709]" pageId="2" pageNumber="593">F.</emphasis>
|
||
Metatibia.
|
||
<emphasis id="B913EAA1FFDB9463FE91AA9F632921A6" bold="true" box="[261,287,1721,1745]" pageId="2" pageNumber="593">G.</emphasis>
|
||
SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37D6538FFDB9465FF53AB0566EA2429" lastPageId="4" lastPageNumber="595" pageId="2" pageNumber="593" type="description">
|
||
<paragraph id="8BD836B3FFDB9463FF53AB0563392015" blockId="2.[151,1437,1827,2034]" pageId="2" pageNumber="593">
|
||
<emphasis id="B913EAA1FFDB9463FF53AB056367204A" bold="true" box="[199,337,1827,1853]" pageId="2" pageNumber="593">Description</emphasis>
|
||
. Male (
|
||
<figureCitation id="135C2A36FFDB9463FE31AB0563CB204A" box="[421,509,1827,1853]" captionStart="FIGURE 1" captionStartId="1.[151,250,1369,1393]" captionTargetBox="[160,1430,461,1338]" captionTargetId="figure-155@1.[151,1436,453,1345]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Pseudophanias spinicornis sp. nov. A. Male. B. Female. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5074544" httpUri="https://zenodo.org/record/5074544/files/figure.png" pageId="2" pageNumber="593">Figs 1A</figureCitation>
|
||
and
|
||
<figureCitation id="135C2A36FFDB9463FDA5AB0560432049" box="[561,629,1827,1854]" captionStart="FIGURE 2" captionStartId="2.[151,250,1649,1673]" captionTargetBox="[159,1435,187,1622]" captionTargetId="figure-27@2.[151,1436,181,1625]" captionTargetPageId="2" captionText="FIGURE 2. Morphological details of Pseudophanias spinicornis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Mesotibia. F. Metatibia. G. SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5074546" httpUri="https://zenodo.org/record/5074546/files/figure.png" pageId="2" pageNumber="593">2A, B</figureCitation>
|
||
). Body length
|
||
<quantity id="4C9F9B56FFDB9463FC83AB05614B2049" box="[791,893,1827,1854]" metricMagnitude="-3" metricUnit="m" metricValue="2.42" pageId="2" pageNumber="593" unit="mm" value="2.42">2.42 mm</quantity>
|
||
. Dorsal surface densely covered with moderately long setae.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDB9462FF53AB4D66E02612" blockId="2.[151,1437,1827,2034]" lastBlockId="3.[151,1437,150,357]" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593">
|
||
Head (HL/HW 0.94) slightly wider than long, densely and coarsely punctate; frons moderately short, with frontal rostrum about 3/4 as wide as HW, with longitudinal, shallow sulcus at middle including a glabrous fovea; vertex with pair of fovea; eyes prominent, moderately large; occipital margin in lateral view weakly concave, with dense tufts of setae; U-shaped setose sulci moderately broad, weakly connecting to eyes. Antennae elongate (
|
||
<figureCitation id="135C2A36FFDB9462FAFBABF1628D27C6" captionStart="FIGURE 2" captionStartId="2.[151,250,1649,1673]" captionTargetBox="[159,1435,187,1622]" captionTargetId="figure-27@2.[151,1436,181,1625]" captionTargetPageId="2" captionText="FIGURE 2. Morphological details of Pseudophanias spinicornis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Mesotibia. F. Metatibia. G. SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5074546" httpUri="https://zenodo.org/record/5074546/files/figure.png" lastPageId="3" lastPageNumber="594" pageId="2" pageNumber="593">Fig. 2C</figureCitation>
|
||
), length
|
||
<quantity id="4C9F9B56FFDA9462FE88ACB163B527C6" box="[284,387,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="594" unit="mm" value="1.31">1.31 mm</quantity>
|
||
; antennomeres 1 about as long as 2–3 combined; antennomeres 2–5 moniliform, successively widened towards apices; antennomeres 6–11 modified to form weak clasping in lateral view; antennomeres 6 enlarged, about as long as 1, spinose ventrally at apico-lateral; antennomeres 7–11 successively widened towards apices; antennomeres 7–10 each transverse, flattened on ventral surface; antennomeres 10 spinose ventrally at apico-lateral; antennomeres 11 largest, sub-oval. Maxillary palpi symmetrical; palpomeres 1 minute; palpomeres 2 elongate, narrowed in basal halves; palpomeres 3 small, widest at apices; palpomeres 4 fusiform.
|
||
</paragraph>
|
||
<caption id="DF18663BFFDA9462FF03AAD061BE2045" ID-DOI="http://doi.org/10.5281/zenodo.5074550" ID-Zenodo-Dep="5074550" httpUri="https://zenodo.org/record/5074550/files/figure.png" pageId="3" pageNumber="594" startId="3.[151,250,1782,1806]" targetBox="[157,1427,389,1749]" targetPageId="3">
|
||
<paragraph id="8BD836B3FFDA9462FF03AAD061BE2045" blockId="3.[151,1436,1782,1842]" pageId="3" pageNumber="594">
|
||
<emphasis id="B913EAA1FFDA9462FF03AAD063202079" bold="true" box="[151,278,1782,1806]" pageId="3" pageNumber="594">FIGURE 3.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C674D30FFDA9462FE4EAAD060D92079" authority="Inoue & Nomura, 2021" authorityName="Inoue & Nomura" authorityYear="2021" box="[474,751,1782,1806]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="594" phylum="Arthropoda" rank="species" species="spinicornis" status="sp. nov.">
|
||
<emphasis id="B913EAA1FFDA9462FE4EAAD060D92079" box="[474,751,1782,1806]" italics="true" pageId="3" pageNumber="594">Pseudophanias spinicornis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B913EAA1FFDA9462FD63AAD061BB2079" bold="true" box="[759,909,1782,1806]" pageId="3" pageNumber="594">
|
||
<taxonomicNameLabel id="A22057DAFFDA9462FD63AAD0617D2079" box="[759,843,1782,1806]" pageId="3" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A, B.
|
||
</emphasis>
|
||
Ventral view.
|
||
<emphasis id="B913EAA1FFDA9462FBB3AAD066092079" bold="true" box="[1063,1087,1782,1806]" pageId="3" pageNumber="594">C.</emphasis>
|
||
Dorsal view.
|
||
<emphasis id="B913EAA1FFDA9462FB40AAD066DA2079" bold="true" box="[1236,1260,1782,1806]" pageId="3" pageNumber="594">D.</emphasis>
|
||
Lateral view.
|
||
<emphasis id="B913EAA1FFDA9462FA11AAD067AA2079" bold="true" box="[1413,1436,1782,1806]" pageId="3" pageNumber="594">E.</emphasis>
|
||
Another lateral view.
|
||
<emphasis id="B913EAA1FFDA9462FEE6AB3C63B32045" bold="true" box="[370,389,1818,1842]" pageId="3" pageNumber="594">F.</emphasis>
|
||
Tergite VIII.
|
||
<emphasis id="B913EAA1FFDA9462FD85AB3C601D2045" bold="true" box="[529,555,1818,1842]" pageId="3" pageNumber="594">G.</emphasis>
|
||
Sternite VIII. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD836B3FFDA9462FF53AB4766BC20E9" blockId="3.[151,1436,1888,2023]" pageId="3" pageNumber="594">Pronotum (PL/PW 1.16) longer than wide, widest at anterior one-third, densely and coarsely punctate, with a median and pair of lateral foveae, with a small conical spine just anterior to median fovea.</paragraph>
|
||
<paragraph id="8BD836B3FFDA9465FF53AB8E614D26FE" blockId="3.[151,1436,1888,2023]" lastBlockId="4.[151,1437,151,1185]" lastPageId="4" lastPageNumber="595" pageId="3" pageNumber="594">
|
||
Elytra (EL/EW 0.69) much wider than long, nearly trapezoidal; each elytron with two basal foveae; discal stria distinct, extending to posterior third. Mesoventrite (
|
||
<figureCitation id="135C2A36FFDA9462FD77ABEA610B2091" box="[739,829,1996,2022]" captionStart="FIGURE 2" captionStartId="2.[151,250,1649,1673]" captionTargetBox="[159,1435,187,1622]" captionTargetId="figure-27@2.[151,1436,181,1625]" captionTargetPageId="2" captionText="FIGURE 2. Morphological details of Pseudophanias spinicornis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Mesotibia. F. Metatibia. G. SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5074546" httpUri="https://zenodo.org/record/5074546/files/figure.png" pageId="3" pageNumber="594">Fig. 2D</figureCitation>
|
||
) with median setose area wide; setose area about 1/3 as wide as mesoventral width; lateral mesoventral foveae setose; metaventrite finely punctate, convex, but area just above metaventral apex impressed; that impression nearly trapezoidal, half as long as metaventral length, occupying 1/3 metaventral width; anterior margin of that impression demarcated from other part of metaventrite. Legs with each femora broadest near middle; protibiae straight; mesotibiae (
|
||
<figureCitation id="135C2A36FFDD9465FCEAAD2561E5266A" box="[894,979,259,285]" captionStart="FIGURE 2" captionStartId="2.[151,250,1649,1673]" captionTargetBox="[159,1435,187,1622]" captionTargetId="figure-27@2.[151,1436,181,1625]" captionTargetPageId="2" captionText="FIGURE 2. Morphological details of Pseudophanias spinicornis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Mesotibia. F. Metatibia. G. SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5074546" httpUri="https://zenodo.org/record/5074546/files/figure.png" pageId="4" pageNumber="595">Fig. 2E</figureCitation>
|
||
) arcuate at apical 1/4, widened apically; apical part flattened posteriorly, with tufts of thick setae at apices; metatibiae (
|
||
<figureCitation id="135C2A36FFDD9465FC6FAD0166782636" box="[1019,1102,295,321]" captionStart="FIGURE 2" captionStartId="2.[151,250,1649,1673]" captionTargetBox="[159,1435,187,1622]" captionTargetId="figure-27@2.[151,1436,181,1625]" captionTargetPageId="2" captionText="FIGURE 2. Morphological details of Pseudophanias spinicornis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Mesotibia. F. Metatibia. G. SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5074546" httpUri="https://zenodo.org/record/5074546/files/figure.png" pageId="4" pageNumber="595">Fig. 2F</figureCitation>
|
||
) slightly arcuate at apical 1/3, broadly covered with thin, long setae; each tarsi (
|
||
<figureCitation id="135C2A36FFDD9465FD5AAD6D611E2612" box="[718,808,331,357]" captionStart="FIGURE 2" captionStartId="2.[151,250,1649,1673]" captionTargetBox="[159,1435,187,1622]" captionTargetId="figure-27@2.[151,1436,181,1625]" captionTargetPageId="2" captionText="FIGURE 2. Morphological details of Pseudophanias spinicornis sp. nov., male. A. Ventral habitus. B. Head and pronotum in lateral view. C. SEM image of antenna in ventral view. D. SEM image of metaventrite in ventral view. E. Mesotibia. F. Metatibia. G. SEM image of protarsus. Arrows indicate ventral spines of male antenna. Scale bars: 1.0 mm in A, B; 0.5 mm in C–F; 0.2 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5074546" httpUri="https://zenodo.org/record/5074546/files/figure.png" pageId="4" pageNumber="595">Fig. 2G</figureCitation>
|
||
) with tarsomeres 2 two-thirds as long as tarsomeres 3; tarsal claws each asymmetrical in size; posterior claws thin, short.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDD9465FF53ADB5618E26A6" blockId="4.[151,1437,151,1185]" pageId="4" pageNumber="595">
|
||
Abdomen (AL/AW 0.80) wider than long; tergite IV longest, twice as long as tergite V, with pair of short longitudinal discal carina; tergite VIII and sternite VIII as in
|
||
<figureCitation id="135C2A36FFDD9465FCD4AD91618226A6" box="[832,948,439,465]" captionStart="FIGURE 3" captionStartId="3.[151,250,1782,1806]" captionTargetBox="[157,1427,389,1749]" captionTargetId="figure-106@3.[151,1436,381,1759]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia of Pseudophanias spinicornis sp. nov. A, B. Ventral view. C. Dorsal view. D. Lateral view. E. Another lateral view. F. Tergite VIII. G. Sternite VIII. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074550" httpUri="https://zenodo.org/record/5074550/files/figure.png" pageId="4" pageNumber="595">Fig. 3F, G</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDD9465FF53ADFD6731254A" blockId="4.[151,1437,151,1185]" pageId="4" pageNumber="595">
|
||
Aedeagus (
|
||
<figureCitation id="135C2A36FFDD9465FED0ADFD638D2682" box="[324,443,475,501]" captionStart="FIGURE 3" captionStartId="3.[151,250,1782,1806]" captionTargetBox="[157,1427,389,1749]" captionTargetId="figure-106@3.[151,1436,381,1759]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia of Pseudophanias spinicornis sp. nov. A, B. Ventral view. C. Dorsal view. D. Lateral view. E. Another lateral view. F. Tergite VIII. G. Sternite VIII. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074550" httpUri="https://zenodo.org/record/5074550/files/figure.png" pageId="4" pageNumber="595">Fig. 3A–E</figureCitation>
|
||
)
|
||
<quantity id="4C9F9B56FFDD9465FE5EADFC601B2682" box="[458,557,474,501]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="595" unit="mm" value="0.5">0.50 mm</quantity>
|
||
in length, well-sclerotized, asymmetrical; median lobe nearly tubular, strongly curved in lateral view, membranous laterally, strongly narrowed posteriorly, then broadened apically, curved laterally; parameres moderately short, asymmetrical, with four setae at apices; endophallus indistinct.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDD9465FF53AE61611D25BA" blockId="4.[151,1437,151,1185]" pageId="4" pageNumber="595">
|
||
Female (
|
||
<figureCitation id="135C2A36FFDD9465FEB8AE6163B32516" box="[300,389,583,609]" captionStart="FIGURE 1" captionStartId="1.[151,250,1369,1393]" captionTargetBox="[160,1430,461,1338]" captionTargetId="figure-155@1.[151,1436,453,1345]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Pseudophanias spinicornis sp. nov. A. Male. B. Female. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5074544" httpUri="https://zenodo.org/record/5074544/files/figure.png" pageId="4" pageNumber="595">Fig. 1B</figureCitation>
|
||
). General appearance similar to male; antennae unmodified, successively widened towards apices, length
|
||
<quantity id="4C9F9B56FFDD9465FEAFAE4D63D425F2" box="[315,482,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.2" metricValueMin="1.18" pageId="4" pageNumber="595" unit="mm" value="1.19" valueMax="1.2" valueMin="1.18">1.18–1.20 mm</quantity>
|
||
; antennomeres 6 quadrate, slightly longer than 5 and 7; mesotibiae arcuate at apical 1/4; its width throughout equal; apical part simple; metatibiae simple, covered with slightly shorter and sparser setae than that of male; metaventrite simple, lacking impression.
|
||
</paragraph>
|
||
<paragraph id="8BD836B3FFDD9465FF53AEF166EA2429" blockId="4.[151,1437,151,1185]" pageId="4" pageNumber="595">
|
||
Measurements. Male (N = 1). BL
|
||
<quantity id="4C9F9B56FFDD9465FDD3AEF160992586" box="[583,687,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="2.42" pageId="4" pageNumber="595" unit="mm" value="2.42">2.42 mm</quantity>
|
||
; HL
|
||
<quantity id="4C9F9B56FFDD9465FD72AEF061782586" box="[742,846,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="595" unit="mm" value="0.5">0.50 mm</quantity>
|
||
; HW
|
||
<quantity id="4C9F9B56FFDD9465FC04AEF061C12586" box="[912,1015,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="595" unit="mm" value="0.53">0.53 mm</quantity>
|
||
; PL
|
||
<quantity id="4C9F9B56FFDD9465FBBEAEF066A42586" box="[1066,1170,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="595" unit="mm" value="0.51">0.51 mm</quantity>
|
||
; PW
|
||
<quantity id="4C9F9B56FFDD9465FB5BAEF167012586" box="[1231,1335,727,753]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="595" unit="mm" value="0.44">0.44 mm</quantity>
|
||
; EL
|
||
<quantity id="4C9F9B56FFDD9465FAFFAEF162F02462" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="4" pageNumber="595" unit="mm" value="0.66">0.66 mm</quantity>
|
||
; EW
|
||
<quantity id="4C9F9B56FFDD9465FE90AEDC635D2462" box="[260,363,762,789]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="595" unit="mm" value="0.95">0.95 mm</quantity>
|
||
; AL
|
||
<quantity id="4C9F9B56FFDD9465FE34AEDC60312462" box="[416,519,762,789]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="595" unit="mm" value="0.75">0.75 mm</quantity>
|
||
; AW
|
||
<quantity id="4C9F9B56FFDD9465FDD0AEDD609D2462" box="[580,683,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="4" pageNumber="595" unit="mm" value="0.94">0.94 mm</quantity>
|
||
. Female (N = 2). BL
|
||
<quantity id="4C9F9B56FFDD9465FC0EAEDD66762462" box="[922,1088,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.26" metricValueMin="2.24" pageId="4" pageNumber="595" unit="mm" value="2.25" valueMax="2.26" valueMin="2.24">2.24–2.26 mm</quantity>
|
||
; HL
|
||
<quantity id="4C9F9B56FFDD9465FBE2AEDD672B2462" box="[1142,1309,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="4.8" metricValueMin="4.6" pageId="4" pageNumber="595" unit="mm" value="0.47" valueMax="0.48" valueMin="0.46">0.46–0.48 mm</quantity>
|
||
; HW
|
||
<quantity id="4C9F9B56FFDD9465FAC9AEDD62CA244E" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="4.9" metricValueMin="4.8" pageId="4" pageNumber="595" unit="mm" value="0.485" valueMax="0.49" valueMin="0.48">0.48– 0.49 mm</quantity>
|
||
(HL/HW 0.95–0.97); PL
|
||
<quantity id="4C9F9B56FFDD9465FD8AAF3860F2244E" box="[542,708,798,825]" metricMagnitude="-4" metricUnit="m" metricValue="4.95" metricValueMax="5.1" metricValueMin="4.8" pageId="4" pageNumber="595" unit="mm" value="0.495" valueMax="0.51" valueMin="0.48">0.48–0.51 mm</quantity>
|
||
; PW
|
||
<quantity id="4C9F9B56FFDD9465FC95AF386192244E" box="[769,932,798,825]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="4" pageNumber="595" unit="mm" value="0.51" valueMax="0.52" valueMin="0.5">0.50–0.52 mm</quantity>
|
||
(PL/PW 0.96–0.98); EL
|
||
<quantity id="4C9F9B56FFDD9465FB2BAF396753244E" box="[1215,1381,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.3" metricValueMin="6.0" pageId="4" pageNumber="595" unit="mm" value="0.615" valueMax="0.63" valueMin="0.6">0.60–0.63 mm</quantity>
|
||
; EW
|
||
<quantity id="4C9F9B56FFDD9465FF03AF64630C242A" box="[151,314,834,861]" metricMagnitude="-4" metricUnit="m" metricValue="9.549999999999999" metricValueMax="9.6" metricValueMin="9.5" pageId="4" pageNumber="595" unit="mm" value="0.955" valueMax="0.96" valueMin="0.95">0.95–0.96 mm</quantity>
|
||
(EL/EW 0.625–0.66); AL
|
||
<quantity id="4C9F9B56FFDD9465FDF2AF64613A242A" box="[614,780,834,861]" metricMagnitude="-4" metricUnit="m" metricValue="8.049999999999999" metricValueMax="9.6" metricValueMin="6.5" pageId="4" pageNumber="595" unit="mm" value="0.8049999999999999" valueMax="0.96" valueMin="0.65">0.65–0.96 mm</quantity>
|
||
; AW
|
||
<quantity id="4C9F9B56FFDD9465FCDDAF6561DA242A" box="[841,1004,835,861]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" metricValueMax="9.8" metricValueMin="9.6" pageId="4" pageNumber="595" unit="mm" value="0.97" valueMax="0.98" valueMin="0.96">0.96–0.98 mm</quantity>
|
||
(AL/AW 0.66–0.72).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6538FFDD9465FF53AF4060F824F5" box="[199,718,870,898]" pageId="4" pageNumber="595" type="distribution">
|
||
<paragraph id="8BD836B3FFDD9465FF53AF4060F824F5" blockId="4.[151,1437,151,1185]" box="[199,718,870,898]" pageId="4" pageNumber="595">
|
||
<emphasis id="B913EAA1FFDD9465FF53AF40635724F7" bold="true" box="[199,353,870,896]" pageId="4" pageNumber="595">Distribution.</emphasis>
|
||
<collectingCountry id="F3707623FFDD9465FEF3AF4163E224F6" box="[359,468,871,897]" name="Myanmar" pageId="4" pageNumber="595">Myanmar</collectingCountry>
|
||
(
|
||
<collectingRegion id="49A3F851FFDD9465FE77AF4160F424F5" box="[483,706,871,898]" country="Myanmar" name="Taninthayi" pageId="4" pageNumber="595">Tanintharyi Region</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6538FFDD9465FF53AFAD60C624BE" pageId="4" pageNumber="595" type="etymology">
|
||
<paragraph id="8BD836B3FFDD9465FF53AFAD60C624BE" blockId="4.[151,1437,151,1185]" pageId="4" pageNumber="595">
|
||
<emphasis id="B913EAA1FFDD9465FF53AFAD637924D2" bold="true" box="[199,335,907,933]" pageId="4" pageNumber="595">Etymology.</emphasis>
|
||
The new specific epithet
|
||
<taxonomicName id="4C674D30FFDD9465FDE1AFAD60C724D3" authorityName="Inoue & Nomura" authorityYear="2021" box="[629,753,907,932]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="spinicornis">
|
||
<emphasis id="B913EAA1FFDD9465FDE1AFAD60C724D3" box="[629,753,907,932]" italics="true" pageId="4" pageNumber="595">spinicornis</emphasis>
|
||
</taxonomicName>
|
||
is a Latin adjective, composed of
|
||
<emphasis id="B913EAA1FFDD9465FBECAFAD668324D3" box="[1144,1205,907,932]" italics="true" pageId="4" pageNumber="595">spina</emphasis>
|
||
(= spine) and
|
||
<emphasis id="B913EAA1FFDD9465FACFAFAD67AA24D3" box="[1371,1436,907,932]" italics="true" pageId="4" pageNumber="595">cornu</emphasis>
|
||
(= horn), referring to the spinose antenna of the male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6538FFDD9465FF53AFF5675723D6" pageId="4" pageNumber="595" type="discussion">
|
||
<paragraph id="8BD836B3FFDD9465FF53AFF5675723D6" blockId="4.[151,1437,151,1185]" pageId="4" pageNumber="595">
|
||
<emphasis id="B913EAA1FFDD9465FF53AFF5630D249A" bold="true" box="[199,315,979,1005]" pageId="4" pageNumber="595">Remarks.</emphasis>
|
||
The new species can be distinguished from
|
||
<taxonomicName id="4C674D30FFDD9465FC8AAFF5664F249A" box="[798,1145,979,1005]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensi">
|
||
<emphasis id="B913EAA1FFDD9465FC8AAFF5664F249A" box="[798,1145,979,1005]" italics="true" pageId="4" pageNumber="595">Pseudophanias tanintharyiensi</emphasis>
|
||
</taxonomicName>
|
||
s
|
||
<emphasis id="B913EAA1FFDD9465FB1DAFF566D7249A" bold="true" box="[1161,1249,979,1005]" pageId="4" pageNumber="595">sp. nov.</emphasis>
|
||
by the following characteristics: U-shaped setose sulci connecting to eyes (separate from eyes in
|
||
<taxonomicName id="4C674D30FFDD9465FB9CAFD166E02366" authorityName="Inoue & Nomura" authorityYear="2021" box="[1032,1238,1015,1041]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensis">
|
||
<emphasis id="B913EAA1FFDD9465FB9CAFD166E02366" box="[1032,1238,1015,1041]" italics="true" pageId="4" pageNumber="595">P. tanintharyiensis</emphasis>
|
||
</taxonomicName>
|
||
); antennomeres 7 and 10 ventrally spinose in male; pronotum densely and coarsely punctate, with a small conical spine; mesoventrite with lateral mesoventral fovea setose (glabrous in
|
||
<taxonomicName id="4C674D30FFDD9465FD63A81961FB232E" authorityName="Inoue & Nomura" authorityYear="2021" box="[759,973,1087,1113]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensis">
|
||
<emphasis id="B913EAA1FFDD9465FD63A81961FB232E" box="[759,973,1087,1113]" italics="true" pageId="4" pageNumber="595">P. tanintharyiensis</emphasis>
|
||
</taxonomicName>
|
||
); mesotibiae with apical part flattened posteriorly in male; metatibiae broadly covered with thin, long setae in male (each metatibiae with slight laminate plate in the male of
|
||
<taxonomicName id="4C674D30FFDD9465FEE2A8A1607023D6" authorityName="Inoue & Nomura" authorityYear="2021" box="[374,582,1159,1185]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensis">
|
||
<emphasis id="B913EAA1FFDD9465FEE2A8A1607023D6" box="[374,582,1159,1185]" italics="true" pageId="4" pageNumber="595">P. tanintharyiensis</emphasis>
|
||
</taxonomicName>
|
||
); tergite IV with discal carinae present (lacking in
|
||
<taxonomicName id="4C674D30FFDD9465FB15A8A1676623D6" authorityName="Inoue & Nomura" authorityYear="2021" box="[1153,1360,1159,1185]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="595" phylum="Arthropoda" rank="species" species="tanintharyiensis">
|
||
<emphasis id="B913EAA1FFDD9465FB15A8A1676623D6" box="[1153,1360,1159,1185]" italics="true" pageId="4" pageNumber="595">P. tanintharyiensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |