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<document id="C098CB638B01401CDF0958001BAF9A46" ID-DOI="10.11646/zootaxa.3990.4.1" ID-GBIF-Dataset="24cc56ce-ad34-420d-8586-5949e370de07" ID-ISSN="1175-5326" ID-Zenodo-Dep="245656" ID-ZooBank="90BE4EAF-8594-4164-AE26-4F47E04A4A5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461270027784" checkinUser="plazi" docAuthor="Karanovic, Tomislav, Kim, Kichoon &amp; Lee, Wonchoel" docDate="2015" docId="03CE8781FFCAE67261802F28FAF1E2D5" docLanguage="en" docName="zt03990p496.pdf" docOrigin="Zootaxa 3990 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enhydrosoma kosmetron Karanovic, Kim &amp; Lee, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="486" masterDocId="FFF7FFF9FFD0E65161172924FFFEE170" masterDocTitle="Concordance between molecular and morphology-based phylogenies of Korean Enhydrosoma (Copepoda: Harpacticoida: Cletodidae) highlights important synapomorphies and homoplasies in this genus globally" masterLastPageNumber="496" masterPageNumber="451" pageNumber="477" updateTime="1698611457494" updateUser="plazi">
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<mods:title id="466F0C0D47E487C1CA3238B252CA80DB">Concordance between molecular and morphology-based phylogenies of Korean Enhydrosoma (Copepoda: Harpacticoida: Cletodidae) highlights important synapomorphies and homoplasies in this genus globally</mods:title>
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<mods:namePart id="6786A3D9AB8111A97A0B3AC0FEF16B43">Karanovic, Tomislav</mods:namePart>
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<mods:namePart id="3B1EB036DFD1077BEB55544FCAEEAEB3">Kim, Kichoon</mods:namePart>
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<mods:namePart id="E0B8DF2E25A5BD82320FC54A16768341">Lee, Wonchoel</mods:namePart>
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<mods:date id="6BC7E0A07F205DC184D6B1BC20F41D50">2015</mods:date>
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<treatment id="03CE8781FFCAE67261802F28FAF1E2D5" ID-DOI="http://doi.org/10.5281/zenodo.5616528" ID-GBIF-Taxon="119638708" ID-Zenodo-Dep="5616528" LSID="urn:lsid:plazi:treatment:03CE8781FFCAE67261802F28FAF1E2D5" httpUri="http://treatment.plazi.org/id/03CE8781FFCAE67261802F28FAF1E2D5" lastPageId="35" lastPageNumber="486" pageId="26" pageNumber="477">
<subSubSection id="C37D651CFFCAE64B61802F28FED0E737" pageId="26" pageNumber="477" type="nomenclature">
<paragraph id="8BD83697FFCAE64B61802F28FDD2E756" blockId="26.[151,556,1548,1607]" box="[151,556,1548,1574]" pageId="26" pageNumber="477">
<heading id="D09081FBFFCAE64B61802F28FDD2E756" bold="true" box="[151,556,1548,1574]" fontSize="11" level="1" pageId="26" pageNumber="477" reason="1">
<emphasis id="B913EA85FFCAE64B61802F28FDD2E756" bold="true" box="[151,556,1548,1574]" pageId="26" pageNumber="477">
<taxonomicName id="4C674D14FFCAE64B61802F28FE38E756" box="[151,454,1548,1574]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="kosmetron" status="sp. nov.">
<emphasis id="B913EA85FFCAE64B61802F28FE38E756" bold="true" box="[151,454,1548,1574]" italics="true" pageId="26" pageNumber="477">Enhydrosoma kosmetron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFCAE64B60D92F28FDD2E756" box="[462,556,1548,1574]" pageId="26" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD83697FFCAE64B61802F0AFED0E737" blockId="26.[151,556,1548,1607]" box="[151,302,1582,1607]" pageId="26" pageNumber="477">
(
<figureCitation id="135C2A12FFCAE64B61882F0AFED8E737" box="[159,294,1582,1607]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStart-4="FIGURE 20" captionStart-5="FIGURE 21" captionStartId-0="29.[151,250,1953,1975]" captionStartId-1="30.[151,250,1953,1975]" captionStartId-2="31.[151,250,1908,1930]" captionStartId-3="32.[151,250,1945,1967]" captionStartId-4="33.[151,250,1915,1937]" captionStartId-5="34.[151,250,1844,1866]" captionTargetBox-0="[174,1413,193,1931]" captionTargetBox-1="[170,1416,193,1931]" captionTargetBox-2="[190,1396,193,1885]" captionTargetBox-3="[163,1419,198,1917]" captionTargetBox-4="[151,1416,202,1871]" captionTargetBox-5="[151,1414,203,1820]" captionTargetId-0="figure@29.[174,1413,193,1932]" captionTargetId-1="figure@30.[170,1417,193,1932]" captionTargetId-2="figure@31.[190,1396,193,1887]" captionTargetId-3="figure@32.[163,1423,193,1927]" captionTargetId-4="figure@33.[151,1436,193,1894]" captionTargetId-5="figure@34.[151,1436,193,1823]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionTargetPageId-3="32" captionTargetPageId-4="33" captionTargetPageId-5="34" captionText-0="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." captionText-1="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." captionText-2="FIGURE 18. Enhydrosoma kosmetron sp. nov., SEM photographs, paratype ♂ 4: A, habitus, lateral; B, cephalic shield, lateral; C, postero-lateral corner of cephalic shield, lateral; D, sensillum on postero-lateral corner of cephalic shield; E, right antennula, lateral; F, transformed seta on first antennular segment, lateral; G, transformed seta on fifth antennular segment, lateral; H, anal somite and caudal rami, lateral." captionText-3="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." captionText-4="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." captionText-5="FIGURE 21. Enhydrosoma kosmetron sp. nov., line drawings, allotype ♂: A, right antennula, ventral; B, distal part of left antennula, antero-ventral; C, mandibular palp, posterior; D, cutting edge of mandibula, posterior; E, maxillular palp, dorsal; F, endopod of third leg, anterior; G, endopod of fourth leg, anterior; H, fifth leg, anterior." httpUri-0="https://zenodo.org/record/245672/files/figure.png" httpUri-1="https://zenodo.org/record/245673/files/figure.png" httpUri-2="https://zenodo.org/record/245674/files/figure.png" httpUri-3="https://zenodo.org/record/245675/files/figure.png" httpUri-4="https://zenodo.org/record/245676/files/figure.png" httpUri-5="https://zenodo.org/record/245677/files/figure.png" pageId="26" pageNumber="477">Figs. 1621</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D651CFFCAE64B61802F54FCE4E680" pageId="26" pageNumber="477" type="materials_examined">
<paragraph id="8BD83697FFCAE64B61802F54FDB4E7DD" blockId="26.[151,1436,1648,2033]" pageId="26" pageNumber="477">
<emphasis id="B913EA85FFCAE64B61802F54FEC9E7F9" bold="true" box="[151,311,1648,1673]" pageId="26" pageNumber="477">
<typeStatus id="54DC8835FFCAE64B61802F54FF2CE7F9" box="[151,210,1648,1673]" pageId="26" pageNumber="477">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3707607FFCAE64B60572F54FE2FE7F8" box="[320,465,1648,1673]" name="South Korea" pageId="26" pageNumber="477">South Korea</collectingCountry>
, South Sea, Gwangyang Bay, sampling station 3 (see Karanovic
<emphasis id="B913EA85FFCAE64B65AD2F55FB0AE7F8" box="[1210,1268,1648,1672]" italics="true" pageId="26" pageNumber="477">et al.</emphasis>
2014), muddy sediments, 3453'03.9&quot;
<accessionNumber id="9434AB74FFCAE64B609A2FB0FE0DE7DD" box="[397,499,1684,1709]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/N12739" pageId="26" pageNumber="477" type="EnaNcbi">N 12739</accessionNumber>
'50.5&quot;E.
</paragraph>
<paragraph id="8BD83697FFCAE64B61D02F9CFCE4E680" blockId="26.[151,1436,1648,2033]" pageId="26" pageNumber="477">
<emphasis id="B913EA85FFCAE64B61D02F9CFE41E7A1" bold="true" box="[199,447,1720,1745]" pageId="26" pageNumber="477">Specimens examined</emphasis>
.
<typeStatus id="54DC8835FFCAE64B60DA2F9CFDCBE7A1" box="[461,565,1720,1745]" pageId="26" pageNumber="477" type="holotype">Holotype</typeStatus>
female (NIBRIV 0000287224) dissected and mounted on one slide, collected from the
<typeStatus id="54DC8835FFCAE64B61EB2FF9FED2E785" box="[252,300,1757,1781]" pageId="26" pageNumber="477">type</typeStatus>
locality,
<date id="FFD91057FFCAE64B60842FF9FDA2E785" box="[403,604,1756,1781]" pageId="26" pageNumber="477" value="2013-02-17">17 February 2013</date>
, collected by K. Kim. Allotype male (NIBRIV 0000287225) dissected and mounted on one slide, collected from
<collectingCountry id="F3707607FFCAE64B632A2E25FD2EE669" box="[573,720,1792,1817]" name="South Korea" pageId="26" pageNumber="477">Korea, South</collectingCountry>
Sea, Gwangyang Bay, sampling station 10 (see Karanovic
<emphasis id="B913EA85FFCAE64B64732E25FA62E668" box="[1380,1436,1792,1816]" italics="true" pageId="26" pageNumber="477">et al.</emphasis>
2014),
<geoCoordinate id="EE535050FFCAE64B61F02E00FE7EE64D" box="[231,384,1828,1853]" direction="north" orientation="latitude" pageId="26" pageNumber="477" precision="1" value="34.920944">34°55'15.4&quot;N</geoCoordinate>
<geoCoordinate id="EE535050FFCAE64B609F2E01FDD1E64D" box="[392,559,1829,1853]" direction="east" orientation="longitude" pageId="26" pageNumber="477" precision="1" value="127.78553">127°47'07.9&quot;E</geoCoordinate>
,
<date id="FFD91057FFCAE64B632C2E01FCF6E64D" box="[571,776,1828,1853]" pageId="26" pageNumber="477" value="2012-02-18">18 February 2012</date>
, collected by K. Kim. Additional
<typeStatus id="54DC8835FFCAE64B659D2E01FB04E64D" box="[1162,1274,1829,1853]" pageId="26" pageNumber="477" type="paratype">paratypes</typeStatus>
: one dissected female (NIBRIV 0000287226) mounted on one slide; four males and two females together on one SEM stub (NIBRIV 0000287227); three males and three females together on another SEM stub (NIBRIV 0000287228); all from
<collectingCountry id="F3707607FFCAE64B61CB2EB5FE86E6D9" box="[220,376,1936,1961]" name="South Korea" pageId="26" pageNumber="477">Korea, South</collectingCountry>
Sea, Gwangyang Bay, sampling station 10 (see Karanovic
<emphasis id="B913EA85FFCAE64B654B2EB5FB62E6D8" box="[1116,1180,1936,1960]" italics="true" pageId="26" pageNumber="477">et al.</emphasis>
2014),
<geoCoordinate id="EE535050FFCAE64B64142EB4FA62E6D9" box="[1283,1436,1936,1961]" direction="north" orientation="latitude" pageId="26" pageNumber="477" precision="1" value="34.920944">34°55'15.4&quot;N</geoCoordinate>
<geoCoordinate id="EE535050FFCAE64B61802E91FEC3E6BD" box="[151,317,1973,1997]" direction="east" orientation="longitude" pageId="26" pageNumber="477" precision="1" value="127.78553">127°47'07.9&quot;E</geoCoordinate>
,
<date id="FFD91057FFCAE64B605C2E91FDE7E6BD" box="[331,537,1972,1997]" pageId="26" pageNumber="477" value="2012-02-18">18 February 2012</date>
, collected by K. Kim. One female used for molecular analysis collected by K. Kim from the
<typeStatus id="54DC8835FFCAE64B60212EFDFE98E681" box="[310,358,2009,2033]" pageId="26" pageNumber="477">type</typeStatus>
locality on
<date id="FFD91057FFCAE64B60FB2EFCFD82E680" box="[492,636,2008,2033]" pageId="26" pageNumber="477" value="2012-07-26">26 July 2012</date>
(see
<tableCitation id="C6E5032CFFCAE64B63A02EFCFCF7E680" box="[695,777,2008,2033]" captionStart="TABLE 1" captionStartId="4.[119,209,671,694]" captionTargetBox="[119,1909,817,1529]" captionTargetPageId="4" captionText="TABLE 1. List οf cοpepοd specimens fοr which mtCΟI fragment was successfully amplified; see text fοr authοrs οf the specific names. Nοte Stylicletodes sp. is an undescribed new species frοm Kοrea." httpUri="http://table.plazi.org/id/DF18661FFFD4E65561602BBBFDF7E394" pageId="26" pageNumber="477" tableUuid="DF18661FFFD4E65561602BBBFDF7E394">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D651CFFCBE64A61D029B3FDEDE187" pageId="27" pageNumber="478" type="etymology">
<paragraph id="8BD83697FFCBE64A61D029B3FDEDE187" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
<emphasis id="B913EA85FFCBE64A61D029B3FEB0E1C0" bold="true" box="[199,334,151,176]" pageId="27" pageNumber="478">Etymology.</emphasis>
This species is named after the Greek noun
<emphasis id="B913EA85FFCBE64A624A29BCFC26E1C0" box="[861,984,152,176]" italics="true" pageId="27" pageNumber="478">
<taxonomicName id="4C674D14FFCBE64A624A29BCFC2DE1C0" box="[861,979,152,176]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="kosmetron">kosmetron</taxonomicName>
,
</emphasis>
meaning “broom” and referring to the broom-like ornamentation of the male antennulae. The specific name is to be treated as a noun (gender masculine) in apposition to the generic name.
</paragraph>
</subSubSection>
<subSubSection id="C37D651CFFCBE67261D02820FAF1E2D5" lastPageId="35" lastPageNumber="486" pageId="27" pageNumber="478" type="description">
<paragraph id="8BD83697FFCBE64A61D02820FDEEE2F0" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
<emphasis id="B913EA85FFCBE64A61D02820FE32E06D" bold="true" box="[199,460,260,285]" pageId="27" pageNumber="478">Description of female.</emphasis>
Based on
<typeStatus id="54DC8835FFCBE64A63282820FD5FE06D" box="[575,673,260,285]" pageId="27" pageNumber="478" type="holotype">holotype</typeStatus>
and several
<typeStatus id="54DC8835FFCBE64A623B2821FC65E06D" box="[812,923,261,285]" pageId="27" pageNumber="478" type="paratype">paratypes</typeStatus>
. Total body length from 441 to 495 µm (mean = 483 µm, n = 7). Body segmentation, colour, nauplius eye, hyaline fringes, general integument thickness, angle between telescoped and non-telescoped parts of pleurons on most free somites, and most somite ornamentation as in
<taxonomicName id="4C674D14FFCBE64A61A32854FE43E0F8" box="[180,445,368,392]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon" status="sp. nov.">
<emphasis id="B913EA85FFCBE64A61A32854FE43E0F8" box="[180,445,368,392]" italics="true" pageId="27" pageNumber="478">Enhydrosoma apimelon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFCBE64A60D2284BFDE1E0F8" box="[453,543,367,392]" pageId="27" pageNumber="478" rank="species">
<emphasis id="B913EA85FFCBE64A60D2284BFDE1E0F8" bold="true" box="[453,543,367,392]" pageId="27" pageNumber="478">sp. nov.</emphasis>
</taxonomicNameLabel>
, however, surface relief on most somites slightly different. Habitus (
<figureCitation id="135C2A12FFCBE64A6434284BFA83E0F8" box="[1315,1405,367,392]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Figs. 16</figureCitation>
A) generally cylindrical in dorsal view, widest at posterior end of cephalothorax and tapering posteriorly, boundary between prosome and urosome inconspicuous; prosome/urosome length ratio about 1.1, and prosome only slightly more voluminous than urosome. Body length/width ratio about
<quantity id="4C9F9B72FFCBE64A627D28F8FC52E085" box="[874,940,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="9.398" pageId="27" pageNumber="478" unit="in" value="3.7">3.7 in</quantity>
dorsal view; cephalothorax about 1.2 times as wide as genital double-somite. Free pedigerous somites without lateral or dorsal expansions, heavily sculptured, pleurons only partly covering coxae of swimming legs in lateral view. Integumental relief more defined by ridges than depressions in dorsal and lateral view; ventro-lateral ridges on urosomites as in
<taxonomicName id="4C674D14FFCBE64A65962B6DFAF3E310" box="[1153,1293,584,608]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A65962B6DFAF3E310" box="[1153,1293,584,608]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
, but ventral surface with additional (mostly internal) ridges. Most cuticular depressions and posterior margin of prosomites without hair-like spinules but full of bacterial growth and detritus, making observation of cuticular pores and sensilla in those regions very difficult; posterior margin of urosomites with hair-like spinules, especially on ventral surface. Hyaline fringe of all somites narrow and rough, in some places nearly serrated, and in all but last two somites also with conical mound-looking protuberances with sensilla on tip, as in
<taxonomicName id="4C674D14FFCBE64A65252BD9FB44E264" box="[1074,1210,764,788]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A65252BD9FB44E264" box="[1074,1210,764,788]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
. In addition to hairlike spinules, surface ornamentation of somites and caudal rami consists of at least two different
<typeStatus id="54DC8835FFCBE64A65F22A04FADEE248" box="[1253,1312,800,824]" pageId="27" pageNumber="478">types</typeStatus>
of sensilla (slender and bottle-shaped; see
<figureCitation id="135C2A12FFCBE64A60E02A60FDB6E22D" box="[503,584,836,861]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Fig. 16</figureCitation>
C), simple cuticular pores, and few stronger spinules; exact number of pores and spinules difficult to establish.
</paragraph>
<paragraph id="8BD83697FFCBE64A61D02AA9FEEEE2B8" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
Rostrum (
<figureCitation id="135C2A12FFCBE64A60212AA8FE79E2D5" box="[310,391,908,933]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Fig. 16</figureCitation>
B, E) as in
<emphasis id="B913EA85FFCBE64A63152AA9FD6EE2D4" box="[514,656,908,932]" italics="true" pageId="27" pageNumber="478">
<taxonomicName id="4C674D14FFCBE64A63152AA9FD72E2D4" box="[514,652,908,932]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">E. apimelon</taxonomicName>
,
</emphasis>
except for space between sensilla smaller than width of one horn-like projection.
</paragraph>
<paragraph id="8BD83697FFCBE64A61D02AF0FAE5E50D" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
Cephalothorax (
<figureCitation id="135C2A12FFCBE64A606B2AF0FE30E29D" box="[380,462,980,1005]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Fig. 16</figureCitation>
B) as in
<taxonomicName id="4C674D14FFCBE64A63272AF1FD46E29C" box="[560,696,980,1004]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A63272AF1FD46E29C" box="[560,696,980,1004]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
, but with completely different relief in dorsal view, without oval depression and with more pronounced longitudinal ridges (compare
<figureCitation id="135C2A12FFCBE64A62A42AD3FBFDE560" box="[947,1027,1015,1040]" captionStart="FIGURE 3" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[189,1397,193,1886]" captionTargetId="figure@11.[189,1398,193,1887]" captionTargetPageId="11" captionText="FIGURE 3. Enhydrosoma apimelon sp. nov., SEM photographs, A &amp; B, paratype ♀ 7; C &amp; D, paratype ♀ 8; E &amp; F, paratype ♀ 9; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, tergites of third and fourth pedigerous somites, dorsal; D, anal operculum and anterior part of caudal rami, dorsal; E, rostrum and antennula, ventral; F, first swimming leg, anterior; G, habitus, dorsal. H, left antennula, dorsal." httpUri="https://zenodo.org/record/245659/files/figure.png" pageId="27" pageNumber="478">Figs. 3</figureCitation>
B and 16B), without comb of long setules in anterior part along inner surface of lateral margin (as observed in
<taxonomicName id="4C674D14FFCBE64A65002D39FB5CE544" box="[1047,1186,1052,1076]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A65002D39FB5CE544" box="[1047,1186,1052,1076]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
) or ventral flaps that protect first three pairs of mouth appendages (as observed in
<taxonomicName id="4C674D14FFCBE64A622B2D65FC3DE528" box="[828,963,1088,1112]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="robustum" status="sp. nov.">
<emphasis id="B913EA85FFCBE64A622B2D65FC3DE528" box="[828,963,1088,1112]" italics="true" pageId="27" pageNumber="478">E. robustum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFCBE64A62DE2D1BFBDCE528" box="[969,1058,1087,1112]" pageId="27" pageNumber="478" rank="species">
<emphasis id="B913EA85FFCBE64A62DE2D1BFBDCE528" bold="true" box="[969,1058,1087,1112]" pageId="27" pageNumber="478">sp. nov.</emphasis>
</taxonomicNameLabel>
); lateral relief similar to that in
<taxonomicName id="4C674D14FFCBE64A64932D65FF01E50C" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A64932D65FF01E50C" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
, except for posterior depression larger and more triangular; most sensilla easy to homologize.
</paragraph>
<paragraph id="8BD83697FFCBE64A61D02DA3FABDE47C" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
Pleuron of free prosomites (
<figureCitation id="135C2A12FFCBE64A63162DA3FDAFE5D0" box="[513,593,1159,1184]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Fig. 16</figureCitation>
A) very similar to each other, without triangular dorsal plates and posterior hair-like spinules, but with posterior conical mound-looking protuberances at base of sensilla, as well as dorsal and dorso-lateral ridges similar to those in
<taxonomicName id="4C674D14FFCBE64A634B2DF5FD19E598" box="[604,743,1232,1256]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A634B2DF5FD19E598" box="[604,743,1232,1256]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
but slightly smaller; all sensilla homologous to those in
<taxonomicName id="4C674D14FFCBE64A64932DF5FF01E47C" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A64932DF5FF01E47C" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
; angle between telescoped and non-telescoped parts of pleuron relatively wide as in
<taxonomicName id="4C674D14FFCBE64A65A32DD1FAC2E47C" box="[1204,1340,1268,1292]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A65A32DD1FAC2E47C" box="[1204,1340,1268,1292]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD83697FFCBE64A61D02C33FCABE408" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
First urosomite (
<figureCitation id="135C2A12FFCBE64A609A2C33FE1CE440" box="[397,482,1303,1328]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Fig. 16</figureCitation>
A) slightly narrower and significantly longer than fourth pedigerous somite and pleuron without free lateral margin, but relief very similar to that of other pedigerous somites, with pronounced dorso-lateral ridges resulting in angular shape in cross-section.
</paragraph>
<paragraph id="8BD83697FFCBE64A61D02CA0FD5CE705" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
Genital double-somite (
<figureCitation id="135C2A12FFCBE64A60C62CA0FDD3E4ED" box="[465,557,1412,1437]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Figs. 16</figureCitation>
A, 19A) 1.6 times as wide as long in ventral view; completely fused ventrally but with deep suture indicating original segmentation between genital and third urosomites dorsally, thus dividing double-somite into equally long and similarly wide halves; general shape and ornamentation as in
<taxonomicName id="4C674D14FFCBE64A641C2CE9FA6BE494" box="[1291,1429,1484,1508]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A641C2CE9FA6BE494" box="[1291,1429,1484,1508]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
, except for more pronounced ridges on ventral surface and additional pair of ventral pores. Female genital complex (
<figureCitation id="135C2A12FFCBE64A61882F30FF0CE75C" box="[159,242,1556,1581]" captionStart="FIGURE 19" captionStartId="32.[151,250,1945,1967]" captionTargetBox="[163,1419,198,1917]" captionTargetId="figure@32.[163,1423,193,1927]" captionTargetPageId="32" captionText="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="27" pageNumber="478">Fig. 19</figureCitation>
A) weakly sclerotized and hardly distinguishable from internal sutures and soft tissue, except for large copulatory pore near midlength of somite and wide copulatory duct; genital operculum as in
<taxonomicName id="4C674D14FFCBE64A65B82F1DFAC9E720" box="[1199,1335,1592,1616]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCBE64A65B82F1DFAC9E720" box="[1199,1335,1592,1616]" italics="true" pageId="27" pageNumber="478">E. apimelon</emphasis>
</taxonomicName>
but with longer armature elements and shorter spinules.
</paragraph>
<paragraph id="8BD83697FFCBE64A61D02F5BFEF4E790" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
Third urosomite (
<figureCitation id="135C2A12FFCBE64A609B2F5BFE16E7E8" box="[396,488,1663,1688]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Figs. 16</figureCitation>
A, 19A) very similar in shape and ornamentation to posterior part of genital doublesomite, but without ventral pores and with three or four large spinules on ventral surface at base of each ventral sensillum.
</paragraph>
<paragraph id="8BD83697FFCBE64A61D02FC8FB83E63C" blockId="27.[151,1437,151,2013]" pageId="27" pageNumber="478">
Fourth urosomite (
<figureCitation id="135C2A12FFCBE64A60B72FC8FE01E675" box="[416,511,1772,1797]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Figs. 16</figureCitation>
A, D, 19A) slightly shorter and narrower than third urosomite and without any sensilla and large spinules, but with well-developed dorsal, dorso-lateral, and ventro-lateral ridges (latter finely serrated), with posterior row of hair-like spinules, and with ventro-lateral cuticular pores.
</paragraph>
<paragraph id="8BD83697FFCBE64D61D02E73FABCE1A5" blockId="27.[151,1437,151,2013]" lastBlockId="28.[151,1437,151,1977]" lastPageId="28" lastPageNumber="479" pageId="27" pageNumber="478">
Anal somite (
<figureCitation id="135C2A12FFCBE64A60492E73FE47E600" box="[350,441,1879,1904]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Figs. 16</figureCitation>
A, D, 19A, B) only slightly clefted medially, with one pair of large dorsal sensilla at base of anal operculum, several minute spinules, small group of large ventral spinules on medial corners, and one pair of ventral simple pores; ventro-lateral corners produced as in preanal somite, but slightly longer and more flared out, also serrated; dorso-lateral corner also produced into small serrated flaps; anal operculum (
<figureCitation id="135C2A12FFCBE64A65892EE0FB05E6AD" box="[1182,1275,1988,2013]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="27" pageNumber="478">Figs. 16</figureCitation>
D, 19B) semicircular, short, not reaching posterior margin of somite (shorter than in
<taxonomicName id="4C674D14FFCCE64D62BB29BDFBCAE1C0" box="[940,1076,152,176]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D62BB29BDFBCAE1C0" box="[940,1076,152,176]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
but longer than in
<taxonomicName id="4C674D14FFCCE64D641329BDFA72E1C0" box="[1284,1420,152,176]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFCCE64D641329BDFA72E1C0" box="[1284,1420,152,176]" italics="true" pageId="28" pageNumber="479">E. robustum</emphasis>
</taxonomicName>
), serrated, representing 39% of somite's width; anal sinus widely open, with three rows of hair-like spinules.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D029FBFADDE0DC" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Caudal rami (
<figureCitation id="135C2A12FFCCE64D604829FBFE44E188" box="[351,442,223,248]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="28" pageNumber="479">Figs. 16</figureCitation>
A, D, 19A, B) spindle-shaped, widest at about proximal quarter of their length, about 1.4 times as long as anal somite, about 2.2 times as long as wide (ventral view), strongly divergent, with strong dorsal ridge, and with space between them about one ramus width. Armature as in
<taxonomicName id="4C674D14FFCCE64D62E7280DFB88E030" box="[1008,1142,296,320]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D62E7280DFB88E030" box="[1008,1142,296,320]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
but dorsal seta longer and inserted more posteriorly (at about midlength); ornamentation as in
<taxonomicName id="4C674D14FFCCE64D629B2869FBEAE014" box="[908,1044,332,356]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFCCE64D629B2869FBEAE014" box="[908,1044,332,356]" italics="true" pageId="28" pageNumber="479">E. robustum</emphasis>
</taxonomicName>
; no tubular pore at base of anterior lateral setae; outer distal process carrying posterior lateral seta even more produced than in
<taxonomicName id="4C674D14FFCCE64D65872855FAE6E0F8" box="[1168,1304,368,392]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFCCE64D65872855FAE6E0F8" box="[1168,1304,368,392]" italics="true" pageId="28" pageNumber="479">E. robustum</emphasis>
</taxonomicName>
; proportion of setae other than dorsal similar to that in
<taxonomicName id="4C674D14FFCCE64D636128B1FD00E0DC" box="[630,766,404,428]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D636128B1FD00E0DC" box="[630,766,404,428]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
; principal apical seta 1.3 times as long as ramus.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02893FDC5E310" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Antennula (
<figureCitation id="135C2A12FFCCE64D605A2893FE54E0A0" box="[333,426,439,464]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="28" pageNumber="479">Figs. 16</figureCitation>
F, 20A) as in
<taxonomicName id="4C674D14FFCCE64D635B289DFD2BE0A0" box="[588,725,440,464]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D635B289DFD2BE0A0" box="[588,725,440,464]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
, except for second segment with one seta less (thus armature formula being: 1.8.7+ae.1.11+ae), and five additional spiniform setae and with strong spinules (three setae on second segment, one on third segment, and stronger apical seta on fifth segment). Lnegth and ornamentation of segments as in
<taxonomicName id="4C674D14FFCCE64D605E2B01FE2DE34C" box="[329,467,548,572]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D605E2B01FE2DE34C" box="[329,467,548,572]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
; as in
<taxonomicName id="4C674D14FFCCE64D63322B01FD51E34C" box="[549,687,548,572]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D63322B01FD51E34C" box="[549,687,548,572]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D14FFCCE64D63FC2B01FC88E34C" box="[747,886,548,572]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFCCE64D63FC2B01FC88E34C" box="[747,886,548,572]" italics="true" pageId="28" pageNumber="479">E. robustum</emphasis>
</taxonomicName>
one unipinnate slender seta on second segment inserted into cone-shaped depression.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02B48FB89E380" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Antenna (
<figureCitation id="135C2A12FFCCE64D60212B48FE6AE3F4" box="[310,404,620,645]" captionStart="FIGURE 19" captionStartId="32.[151,250,1945,1967]" captionTargetBox="[163,1419,198,1917]" captionTargetId="figure@32.[163,1423,193,1927]" captionTargetPageId="32" captionText="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="28" pageNumber="479">Figs. 19</figureCitation>
C), labrum, paragnaths, general shape and armature of maxilla (
<figureCitation id="135C2A12FFCCE64D657E2B48FB40E3F4" box="[1129,1214,620,645]" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
B), maxilliped (
<figureCitation id="135C2A12FFCCE64D64782B48FF48E3D8" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
C), endopod of first swimming leg (
<figureCitation id="135C2A12FFCCE64D63502BABFD64E3D8" box="[583,666,655,680]" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
D), second swimming leg (
<figureCitation id="135C2A12FFCCE64D62DE2BABFBE5E3D8" box="[969,1051,655,680]" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
F), exopod of third swimming leg, and exopod of fourth swimming leg as in
<taxonomicName id="4C674D14FFCCE64D63672B91FD06E3BC" box="[624,760,692,716]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D63672B91FD06E3BC" box="[624,760,692,716]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
and
<emphasis id="B913EA85FFCCE64D62272B91FC41E3BC" box="[816,959,692,716]" italics="true" pageId="28" pageNumber="479">
<taxonomicName id="4C674D14FFCCE64D62272B91FC45E3BC" box="[816,955,692,716]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">E. robustum</taxonomicName>
,
</emphasis>
except for small differences in proportion of certain segments and armature elements and very minor differences in ornamentation.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02BD8FD21E248" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Mandibula (
<figureCitation id="135C2A12FFCCE64D60452BD8FE5AE264" box="[338,420,764,789]" captionStart="FIGURE 19" captionStartId="32.[151,250,1945,1967]" captionTargetBox="[163,1419,198,1917]" captionTargetId="figure@32.[163,1423,193,1927]" captionTargetPageId="32" captionText="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="28" pageNumber="479">Fig. 19</figureCitation>
D) size and segmentation as in
<taxonomicName id="4C674D14FFCCE64D62182BD9FC66E264" box="[783,920,764,788]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D62182BD9FC66E264" box="[783,920,764,788]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
, but palp with four setae and seta on cutting edge of coxal gnathobase much stronger and longer.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02A60FE7CE20F" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Maxillula (
<figureCitation id="135C2A12FFCCE64D60522A60FE6BE22C" box="[325,405,836,861]" captionStart="FIGURE 19" captionStartId="32.[151,250,1945,1967]" captionTargetBox="[163,1419,198,1917]" captionTargetId="figure@32.[163,1423,193,1927]" captionTargetPageId="32" captionText="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="28" pageNumber="479">Fig. 19</figureCitation>
E) size and segmentation as in
<taxonomicName id="4C674D14FFCCE64D63ED2A61FC7DE22C" box="[762,899,836,860]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D63ED2A61FC7DE22C" box="[762,899,836,860]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
, but both basis and praecoxal arthrite with one additional inner seta.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02AA8FC61E2B8" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Maxilliped (
<figureCitation id="135C2A12FFCCE64D60432AA8FE59E2D4" box="[340,423,908,933]" captionStart="FIGURE 19" captionStartId="32.[151,250,1945,1967]" captionTargetBox="[163,1419,198,1917]" captionTargetId="figure@32.[163,1423,193,1927]" captionTargetPageId="32" captionText="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="28" pageNumber="479">Fig. 19</figureCitation>
C) as in
<taxonomicName id="4C674D14FFCCE64D631A2AA9FD69E2D4" box="[525,663,908,932]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D631A2AA9FD69E2D4" box="[525,663,908,932]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
, except endopodal strong spine shorter and slender endopodal seta proportionately longer; coxa ornamented with three rows of spinules.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02AF0FBCCE29D" blockId="28.[151,1437,151,1977]" box="[199,1074,980,1005]" pageId="28" pageNumber="479">
Exopod of first swimming leg (
<figureCitation id="135C2A12FFCCE64D63312AF0FD86E29C" box="[550,632,980,1005]" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
E) with five elements on third segment.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02AD3FB8FE545" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Endopod of third swimming leg (
<figureCitation id="135C2A12FFCCE64D63512AD3FD64E560" box="[582,666,1015,1040]" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
G) and endopod of fourth swimming leg (
<figureCitation id="135C2A12FFCCE64D656C2AD3FB31E560" box="[1147,1231,1015,1040]" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">Fig. 20</figureCitation>
H) both with very short outer spines in addition to two slender and long terminal setae on second segment.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02D1BFCF2E4ED" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Fifth leg (
<figureCitation id="135C2A12FFCCE64D602B2D1BFE6EE528" box="[316,400,1087,1112]" captionStart="FIGURE 20" captionStartId="33.[151,250,1915,1937]" captionTargetBox="[151,1416,202,1871]" captionTargetId="figure@33.[151,1436,193,1894]" captionTargetPageId="33" captionText="FIGURE 20. Enhydrosoma kosmetron sp. nov., line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior." httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="28" pageNumber="479">
Fig.
<date id="FFD91057FFCCE64D60642D64FE6EE528" box="[371,400,1088,1112]" pageId="28" pageNumber="479" value="1920-01">20</date>
</figureCitation>
I) biramous as in
<taxonomicName id="4C674D14FFCCE64D63492D65FD16E528" box="[606,744,1088,1112]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D63492D65FD16E528" box="[606,744,1088,1112]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
, comprising conical exopod and baseoendopod, but exopod much more robust and endopodal lobe shorter; exopod also armed with three elements, but lateral elements inserted closer to distal margin (as in
<taxonomicName id="4C674D14FFCCE64D60CF2DADFDA1E5D0" box="[472,607,1160,1184]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFCCE64D60CF2DADFDA1E5D0" box="[472,607,1160,1184]" italics="true" pageId="28" pageNumber="479">E. robustum</emphasis>
</taxonomicName>
), with distal lateral element almost apical and difficult to homologize, and apical element very small and bare. Outer margin of exopod with numerous long hair-like spinules, inner margin smooth, without tubular pores; baseoendopod as in
<taxonomicName id="4C674D14FFCCE64D63CD2DF5FC9CE598" box="[730,866,1232,1256]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFCCE64D63CD2DF5FC9CE598" box="[730,866,1232,1256]" italics="true" pageId="28" pageNumber="479">E. robustum</emphasis>
</taxonomicName>
with robust distal seta, but with only one cuticular spiniform process on inner margin covered by small spinules; distal exopodal element about 0.3 times as long as proximal endopodal spine, 0.2 times as long as distal endopodal spine, 0.75 times as long as proximal exopodal element, 0.25 times as long as middle exopodal element, 0.3 times as long as distal endopodal element, and less than 0.2 times as long as entire exopod; exopod about 3.3 times as long as wide. Endopodal lobe without tubular pores, but with two simple pores at base of inner spines.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02C83FC72E495" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Sixth legs (
<figureCitation id="135C2A12FFCCE64D605E2C83FE64E4B0" box="[329,410,1447,1472]" captionStart="FIGURE 19" captionStartId="32.[151,250,1945,1967]" captionTargetBox="[163,1419,198,1917]" captionTargetId="figure@32.[163,1423,193,1927]" captionTargetPageId="32" captionText="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="28" pageNumber="479">Fig. 19</figureCitation>
A) as in
<taxonomicName id="4C674D14FFCCE64D60E82C8DFD76E4B0" box="[511,648,1448,1472]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFCCE64D60E82C8DFD76E4B0" box="[511,648,1448,1472]" italics="true" pageId="28" pageNumber="479">E. apimelon</emphasis>
</taxonomicName>
, but with proportionately smaller basal spinules and larger armature; inner smooth seta about 1.7 times as long as outer unipinnate spine.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02CCBFE73E63D" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
<emphasis id="B913EA85FFCCE64D61D02CCBFE45E778" bold="true" box="[199,443,1519,1544]" pageId="28" pageNumber="479">Description of male.</emphasis>
Based on allotype and four other
<typeStatus id="54DC8835FFCCE64D62522CD5FC4AE779" box="[837,948,1521,1545]" pageId="28" pageNumber="479" type="paratype">paratypes</typeStatus>
. Body length ranging from 279 to 312 Μm (mean = 302 µm, n = 5). Genital somite and third urosomite not fused (
<figureCitation id="135C2A12FFCCE64D62D72F30FBECE75C" box="[960,1042,1556,1581]" captionStart="FIGURE 19" captionStartId="32.[151,250,1945,1967]" captionTargetBox="[163,1419,198,1917]" captionTargetId="figure@32.[163,1423,193,1927]" captionTargetPageId="32" captionText="FIGURE 19. Enhydrosoma kosmetron sp. nov., line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral." httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="28" pageNumber="479">Fig. 19</figureCitation>
F). Habitus (
<figureCitation id="135C2A12FFCCE64D65B62F30FB02E75D" box="[1185,1276,1556,1581]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="28" pageNumber="479">Figs. 16</figureCitation>
G, 17A, 18A), colour, rostrum, shape and ornamentation of cephalothorax (
<figureCitation id="135C2A12FFCCE64D62402F13FC52E720" box="[855,940,1591,1616]" captionStart="FIGURE 18" captionStartId="31.[151,250,1908,1930]" captionTargetBox="[190,1396,193,1885]" captionTargetId="figure@31.[190,1396,193,1887]" captionTargetPageId="31" captionText="FIGURE 18. Enhydrosoma kosmetron sp. nov., SEM photographs, paratype ♂ 4: A, habitus, lateral; B, cephalic shield, lateral; C, postero-lateral corner of cephalic shield, lateral; D, sensillum on postero-lateral corner of cephalic shield; E, right antennula, lateral; F, transformed seta on first antennular segment, lateral; G, transformed seta on fifth antennular segment, lateral; H, anal somite and caudal rami, lateral." httpUri="https://zenodo.org/record/245674/files/figure.png" pageId="28" pageNumber="479">Fig. 18</figureCitation>
B, C, D), shape and ornamentation of free prosomites (
<figureCitation id="135C2A12FFCCE64D60322F78FE86E705" box="[293,376,1628,1653]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="28" pageNumber="479">Fig. 16</figureCitation>
G, 18A), shape and ornamentation of last three urosomites (
<figureCitation id="135C2A12FFCCE64D65232F78FB6DE705" box="[1076,1171,1628,1653]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="28" pageNumber="479">Figs. 16</figureCitation>
H, 17C, H, 18H, 19F), general shape, armature and ornamentation of caudal rami (
<figureCitation id="135C2A12FFCCE64D62252F5BFC70E7E8" box="[818,910,1663,1688]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="28" pageNumber="479">Figs. 16</figureCitation>
H, 17C, 18H, 19F), antenna (
<figureCitation id="135C2A12FFCCE64D65C32F5BFADAE7E8" box="[1236,1316,1663,1688]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
E), labrum (
<figureCitation id="135C2A12FFCCE64D61882F80FF0EE7CC" box="[159,240,1700,1725]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
F), paragnaths (
<figureCitation id="135C2A12FFCCE64D60B72F80FE0EE7CC" box="[416,496,1700,1725]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
F), mandibula (
<figureCitation id="135C2A12FFCCE64D638A2F80FD06E7CC" box="[669,760,1700,1725]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Figs. 17</figureCitation>
F, 21C, D), maxillula (
<figureCitation id="135C2A12FFCCE64D62E12F80FBB8E7CC" box="[1014,1094,1700,1725]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
F, 21E), maxilla (
<figureCitation id="135C2A12FFCCE64D641C2F80FAA2E7CC" box="[1291,1372,1700,1725]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
B, F), maxilliped (
<figureCitation id="135C2A12FFCCE64D60092FE3FE91E790" box="[286,367,1735,1760]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
F), first swimming leg (
<figureCitation id="135C2A12FFCCE64D636F2FE3FD36E790" box="[632,712,1735,1760]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
G), second swimming leg (
<figureCitation id="135C2A12FFCCE64D62E02FE3FBB9E790" box="[1015,1095,1735,1760]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
A), exopod of third swimming leg (
<figureCitation id="135C2A12FFCCE64D61DE2FC8FEE7E674" box="[201,281,1772,1797]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Fig. 17</figureCitation>
A), and fourth swimming leg (
<figureCitation id="135C2A12FFCCE64D63672FC8FD3EE674" box="[624,704,1772,1797]" captionStart="FIGURE 21" captionStartId="34.[151,250,1844,1866]" captionTargetBox="[151,1414,203,1820]" captionTargetId="figure@34.[151,1436,193,1823]" captionTargetPageId="34" captionText="FIGURE 21. Enhydrosoma kosmetron sp. nov., line drawings, allotype ♂: A, right antennula, ventral; B, distal part of left antennula, antero-ventral; C, mandibular palp, posterior; D, cutting edge of mandibula, posterior; E, maxillular palp, dorsal; F, endopod of third leg, anterior; G, endopod of fourth leg, anterior; H, fifth leg, anterior." httpUri="https://zenodo.org/record/245677/files/figure.png" pageId="28" pageNumber="479">Fig. 21</figureCitation>
G) as in female. Prosome/urosome ratio about 1.2, greatest width at posterior end of cephalothorax, body length/width ratio about 3.3; cephalothorax 1.6 times as wide as genital somite in dorsal view.
</paragraph>
<paragraph id="8BD83697FFCCE64D61D02E73FE81E6C9" blockId="28.[151,1437,151,1977]" pageId="28" pageNumber="479">
Genital somite (
<figureCitation id="135C2A12FFCCE64D606E2E73FE2AE600" box="[377,468,1879,1904]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="28" pageNumber="479">Figs. 17</figureCitation>
H, 19F) 1.8 times as wide as long in ventral view, similar in ornamentation to that in
<taxonomicName id="4C674D14FFCCE64D64932E7DFF01E6E4" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFCCE64D64932E7DFF01E6E4" italics="true" pageId="28" pageNumber="479">E. robustum</emphasis>
</taxonomicName>
, and also with left sixth leg functioning as genital operculum; no spermatophore visible inside any observed specimens.
</paragraph>
<caption id="DF18661FFFCDE64C61802E85FA62E684" httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="29" pageNumber="480" targetBox="[174,1413,193,1931]" targetPageId="29">
<paragraph id="8BD83697FFCDE64C61802E85FA62E684" blockId="29.[151,1436,1953,2036]" pageId="29" pageNumber="480">
<emphasis id="B913EA85FFCDE64C61802E85FEDEE6C6" bold="true" box="[151,288,1953,1975]" pageId="29" pageNumber="480">FIGURE 16.</emphasis>
<taxonomicName id="4C674D14FFCDE64C603F2E85FDDAE6C7" box="[296,548,1953,1975]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="kosmetron" status="sp. nov.">
<emphasis id="B913EA85FFCDE64C603F2E85FDDAE6C7" box="[296,548,1953,1975]" italics="true" pageId="29" pageNumber="480">Enhydrosoma kosmetron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFCDE64C633C2E86FD82E6C7" box="[555,636,1954,1975]" pageId="29" pageNumber="480" rank="species">
<emphasis id="B913EA85FFCDE64C633C2E86FD82E6C7" bold="true" box="[555,636,1954,1975]" pageId="29" pageNumber="480">sp. nov.</emphasis>
</taxonomicNameLabel>
, SEM photographs, A-D, paratype ♀1; E &amp; F, paratype ♀2; G &amp; H, paratype ♂1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal.
</paragraph>
</caption>
<caption id="DF18661FFFCEE64F61802E85FBCBE684" httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="30" pageNumber="481" targetBox="[170,1416,193,1931]" targetPageId="30">
<paragraph id="8BD83697FFCEE64F61802E85FBCBE684" blockId="30.[151,1436,1953,2036]" pageId="30" pageNumber="481">
<emphasis id="B913EA85FFCEE64F61802E85FEDEE6C6" bold="true" box="[151,288,1953,1975]" pageId="30" pageNumber="481">FIGURE 17.</emphasis>
<taxonomicName id="4C674D14FFCEE64F60302E85FDDFE6C7" box="[295,545,1953,1975]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="kosmetron" status="sp. nov.">
<emphasis id="B913EA85FFCEE64F60302E85FDDFE6C7" box="[295,545,1953,1975]" italics="true" pageId="30" pageNumber="481">Enhydrosoma kosmetron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFCEE64F633E2E86FD84E6C7" box="[553,634,1954,1975]" pageId="30" pageNumber="481" rank="species">
<emphasis id="B913EA85FFCEE64F633E2E86FD84E6C7" bold="true" box="[553,634,1954,1975]" pageId="30" pageNumber="481">sp. nov.</emphasis>
</taxonomicNameLabel>
, SEM photographs, A-C, paratype ♂2; D-H, paratype ♂3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior.
</paragraph>
</caption>
<caption id="DF18661FFFCFE64E61802E50FE2BE696" httpUri="https://zenodo.org/record/245674/files/figure.png" pageId="31" pageNumber="482" targetBox="[190,1396,193,1885]" targetPageId="31">
<paragraph id="8BD83697FFCFE64E61802E50FE2BE696" blockId="31.[151,1436,1908,2022]" pageId="31" pageNumber="482">
<emphasis id="B913EA85FFCFE64E61802E50FEE1E6F9" bold="true" box="[151,287,1908,1930]" pageId="31" pageNumber="482">FIGURE 18.</emphasis>
<taxonomicName id="4C674D14FFCFE64E60322E50FDDEE6FA" box="[293,544,1908,1930]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="kosmetron" status="sp. nov.">
<emphasis id="B913EA85FFCFE64E60322E50FDDEE6FA" box="[293,544,1908,1930]" italics="true" pageId="31" pageNumber="482">Enhydrosoma kosmetron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFCFE64E63312E51FD89E6FA" box="[550,631,1909,1930]" pageId="31" pageNumber="482" rank="species">
<emphasis id="B913EA85FFCFE64E63312E51FD89E6FA" bold="true" box="[550,631,1909,1930]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
, SEM photographs, paratype ♂4: A, habitus, lateral; B, cephalic shield, lateral; C, postero-lateral corner of cephalic shield, lateral; D, sensillum on postero-lateral corner of cephalic shield; E, right antennula, lateral; F, transformed seta on first antennular segment, lateral; G, transformed seta on fifth antennular segment, lateral; H, anal somite and caudal rami, lateral.
</paragraph>
</caption>
<caption id="DF18661FFFF0E67161802EBDFF1BE69D" httpUri="https://zenodo.org/record/245675/files/figure.png" pageId="32" pageNumber="483" targetBox="[163,1419,198,1917]" targetPageId="32">
<paragraph id="8BD83697FFF0E67161802EBDFF1BE69D" blockId="32.[151,1436,1945,2029]" pageId="32" pageNumber="483">
<emphasis id="B913EA85FFF0E67161802EBDFEDFE6DE" bold="true" box="[151,289,1945,1967]" pageId="32" pageNumber="483">FIGURE 19.</emphasis>
<taxonomicName id="4C674D14FFF0E671603F2EBDFDDBE6DF" box="[296,549,1945,1967]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="32" pageNumber="483" phylum="Arthropoda" rank="species" species="kosmetron" status="sp. nov.">
<emphasis id="B913EA85FFF0E671603F2EBDFDDBE6DF" box="[296,549,1945,1967]" italics="true" pageId="32" pageNumber="483">Enhydrosoma kosmetron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFF0E671633B2EBEFD80E6DF" box="[556,638,1946,1967]" pageId="32" pageNumber="483" rank="species">
<emphasis id="B913EA85FFF0E671633B2EBEFD80E6DF" bold="true" box="[556,638,1946,1967]" pageId="32" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
, line drawings, A-D, holotype ♀; F, allotype ♂: A, urosome, ventral; B, anal somite and caudal rami, dorsal; C, exopod of antenna, anterior; D, mandibula, posterior; E, maxillula, anterior; F, urosome, ventral.
</paragraph>
</caption>
<caption id="DF18661FFFF1E67061802E5FFA9CE6BE" httpUri="https://zenodo.org/record/245676/files/figure.png" pageId="33" pageNumber="484" targetBox="[151,1416,202,1871]" targetPageId="33">
<paragraph id="8BD83697FFF1E67061802E5FFA9CE6BE" blockId="33.[151,1436,1915,1998]" pageId="33" pageNumber="484">
<emphasis id="B913EA85FFF1E67061802E5FFEDCE6E0" bold="true" box="[151,290,1915,1937]" pageId="33" pageNumber="484">FIGURE 20.</emphasis>
<taxonomicName id="4C674D14FFF1E670603C2E5FFDD6E6E1" box="[299,552,1915,1937]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="kosmetron" status="sp. nov.">
<emphasis id="B913EA85FFF1E670603C2E5FFDD6E6E1" box="[299,552,1915,1937]" italics="true" pageId="33" pageNumber="484">Enhydrosoma kosmetron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFF1E67063252E58FD7AE6E1" box="[562,644,1916,1937]" pageId="33" pageNumber="484" rank="species">
<emphasis id="B913EA85FFF1E67063252E58FD7AE6E1" bold="true" box="[562,644,1916,1937]" pageId="33" pageNumber="484">sp. nov.</emphasis>
</taxonomicNameLabel>
, line drawings, holotype ♀: A, antennula, ventral; B, maxilla, posterior; C, maxilliped, posterior; D, second endopodal segment of first leg, anterior; E, third exopodal segment of first leg, anterior; F, endopod of second leg, anterior; G, endopod of third leg anterior; H, endopod of fourth leg, anterior; I, fifth leg, anterior.
</paragraph>
</caption>
<caption id="DF18661FFFF2E67361802E10FC01E6F7" httpUri="https://zenodo.org/record/245677/files/figure.png" pageId="34" pageNumber="485" targetBox="[151,1414,203,1820]" targetPageId="34">
<paragraph id="8BD83697FFF2E67361802E10FC01E6F7" blockId="34.[151,1436,1844,1927]" pageId="34" pageNumber="485">
<emphasis id="B913EA85FFF2E67361802E10FEDCE639" bold="true" box="[151,290,1844,1866]" pageId="34" pageNumber="485">FIGURE 21.</emphasis>
<taxonomicName id="4C674D14FFF2E673603C2E10FDD9E63A" box="[299,551,1844,1866]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="kosmetron" status="sp. nov.">
<emphasis id="B913EA85FFF2E673603C2E10FDD9E63A" box="[299,551,1844,1866]" italics="true" pageId="34" pageNumber="485">Enhydrosoma kosmetron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22057FEFFF2E67363272E11FD7DE63A" box="[560,643,1845,1866]" pageId="34" pageNumber="485" rank="species">
<emphasis id="B913EA85FFF2E67363272E11FD7DE63A" bold="true" box="[560,643,1845,1866]" pageId="34" pageNumber="485">sp. nov.</emphasis>
</taxonomicNameLabel>
, line drawings, allotype ♂: A, right antennula, ventral; B, distal part of left antennula, antero-ventral; C, mandibular palp, posterior; D, cutting edge of mandibula, posterior; E, maxillular palp, dorsal; F, endopod of third leg, anterior; G, endopod of fourth leg, anterior; H, fifth leg, anterior.
</paragraph>
</caption>
<paragraph id="8BD83697FFF3E67261D029B3FCC9E1A5" blockId="35.[151,1437,151,933]" pageId="35" pageNumber="486">
Third urosomite (
<figureCitation id="135C2A12FFF3E672608329B3FE0DE1C0" box="[404,499,151,176]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="35" pageNumber="486">Figs. 17</figureCitation>
H, 19F) as posterior part of genital double-somite in female but proportionately narrower, and with four or five large ventro-lateral spinules.
</paragraph>
<paragraph id="8BD83697FFF3E67261D029FBFE58E06D" blockId="35.[151,1437,151,933]" pageId="35" pageNumber="486">
Caudal rami (
<figureCitation id="135C2A12FFF3E672607229FBFE3DE188" box="[357,451,223,248]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="35" pageNumber="486">Figs. 16</figureCitation>
H, 17C, 19F) very similar to those in female but slightly longer in proportion to anal somite and more robust.
</paragraph>
<paragraph id="8BD83697FFF3E67261D02803FCD6E0A0" blockId="35.[151,1437,151,933]" pageId="35" pageNumber="486">
Antennula (
<figureCitation id="135C2A12FFF3E672605A2803FE54E030" box="[333,426,295,320]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="35" pageNumber="486">Figs. 17</figureCitation>
D, 18E, F, G, 21A, B) as in
<taxonomicName id="4C674D14FFF3E67263F9280DFC86E030" box="[750,888,296,320]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="apimelon">
<emphasis id="B913EA85FFF3E67263F9280DFC86E030" box="[750,888,296,320]" italics="true" pageId="35" pageNumber="486">E. apimelon</emphasis>
</taxonomicName>
, except for second segment with two spiniform setae, third segment with one spiniform seta, fourth segment with large broom-like structure instead of brush and with only six or seven large spinules, fifth segment with large spiniform process and three slender setae, and sixth segment with large characteristically shaped tube pore (
<figureCitation id="135C2A12FFF3E672623F28B0FC80E0DC" box="[808,894,404,429]" captionStart="FIGURE 18" captionStartId="31.[151,250,1908,1930]" captionTargetBox="[190,1396,193,1885]" captionTargetId="figure@31.[190,1396,193,1887]" captionTargetPageId="31" captionText="FIGURE 18. Enhydrosoma kosmetron sp. nov., SEM photographs, paratype ♂ 4: A, habitus, lateral; B, cephalic shield, lateral; C, postero-lateral corner of cephalic shield, lateral; D, sensillum on postero-lateral corner of cephalic shield; E, right antennula, lateral; F, transformed seta on first antennular segment, lateral; G, transformed seta on fifth antennular segment, lateral; H, anal somite and caudal rami, lateral." httpUri="https://zenodo.org/record/245674/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
G); second segment as in female with slender plumose seta inserted into cone-like depression (
<figureCitation id="135C2A12FFF3E67263AF2893FCF7E0A0" box="[696,777,439,464]" captionStart="FIGURE 18" captionStartId="31.[151,250,1908,1930]" captionTargetBox="[190,1396,193,1885]" captionTargetId="figure@31.[190,1396,193,1887]" captionTargetPageId="31" captionText="FIGURE 18. Enhydrosoma kosmetron sp. nov., SEM photographs, paratype ♂ 4: A, habitus, lateral; B, cephalic shield, lateral; C, postero-lateral corner of cephalic shield, lateral; D, sensillum on postero-lateral corner of cephalic shield; E, right antennula, lateral; F, transformed seta on first antennular segment, lateral; G, transformed seta on fifth antennular segment, lateral; H, anal somite and caudal rami, lateral." httpUri="https://zenodo.org/record/245674/files/figure.png" pageId="35" pageNumber="486">Fig. 18</figureCitation>
F).
</paragraph>
<paragraph id="8BD83697FFF3E67261D028F8FE86E34C" blockId="35.[151,1437,151,933]" pageId="35" pageNumber="486">
Endopod of third swimming leg (
<figureCitation id="135C2A12FFF3E672634528F8FD59E084" box="[594,679,476,501]" captionStart="FIGURE 21" captionStartId="34.[151,250,1844,1866]" captionTargetBox="[151,1414,203,1820]" captionTargetId="figure@34.[151,1436,193,1823]" captionTargetPageId="34" captionText="FIGURE 21. Enhydrosoma kosmetron sp. nov., line drawings, allotype ♂: A, right antennula, ventral; B, distal part of left antennula, antero-ventral; C, mandibular palp, posterior; D, cutting edge of mandibula, posterior; E, maxillular palp, dorsal; F, endopod of third leg, anterior; G, endopod of fourth leg, anterior; H, fifth leg, anterior." httpUri="https://zenodo.org/record/245677/files/figure.png" pageId="35" pageNumber="486">Fig. 21</figureCitation>
F) transformed and apparently three-segmented, with outer spine fused to segment and enlarged, secondary segmentation resulting in very small third segment bearing two long and slender apical setae.
</paragraph>
<paragraph id="8BD83697FFF3E67261D02B63FD7BE380" blockId="35.[151,1437,151,933]" pageId="35" pageNumber="486">
Fifth leg (
<figureCitation id="135C2A12FFF3E67260212B63FE6CE310" box="[310,402,583,608]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="35" pageNumber="486">Figs. 17</figureCitation>
H, 21H) somewhat smaller than in female but with similar basic structure; exopod about 2.3 times as long as wide, with all three elements present and of similar proportions as in female, except for apical one bipinnate; endopodal lobe without distal armature element or inner spiniform processes, its distal tip with bunch of hair-like spinules; proximal endopodal inner spine about 1.6 times as long as distal endopodal inner spine and 1.2 times as long as longest element on exopod.
</paragraph>
<paragraph id="8BD83697FFF3E67261D02BD8FEACE248" blockId="35.[151,1437,151,933]" pageId="35" pageNumber="486">
Sixth legs (
<figureCitation id="135C2A12FFF3E672605D2BD8FE56E264" box="[330,424,764,789]" captionStart="FIGURE 17" captionStartId="30.[151,250,1953,1975]" captionTargetBox="[170,1416,193,1931]" captionTargetId="figure@30.[170,1417,193,1932]" captionTargetPageId="30" captionText="FIGURE 17. Enhydrosoma kosmetron sp. nov., SEM photographs, A-C, paratype ♂ 2; D-H, paratype ♂ 3: A, habitus, ventral; B, maxilla, ventral; C, anal somite and caudal rami, ventral; D, rostrum and left antennula, ventral; E, antenna, ventral; F, mouth appendages, ventral; G, first swimming leg, anterior; H, fifth and sixth legs, anterior." httpUri="https://zenodo.org/record/245673/files/figure.png" pageId="35" pageNumber="486">Figs. 17</figureCitation>
H, 19F) as in
<taxonomicName id="4C674D14FFF3E672635D2BD9FD2BE264" box="[586,725,764,788]" class="Maxillopoda" family="Cletodidae" genus="Enhydrosoma" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="robustum">
<emphasis id="B913EA85FFF3E672635D2BD9FD2BE264" box="[586,725,764,788]" italics="true" pageId="35" pageNumber="486">E. robustum</emphasis>
</taxonomicName>
, withsimple cuticular plates, unarmed, with transverse row of slender spinules.
</paragraph>
<paragraph id="8BD83697FFF3E67261D02A60FAF1E2D5" blockId="35.[151,1437,151,933]" pageId="35" pageNumber="486">
<emphasis id="B913EA85FFF3E67261D02A60FEB3E22D" bold="true" box="[199,333,836,861]" pageId="35" pageNumber="486">Variability.</emphasis>
Despite numerous examined specimens and detailed examination using SEM (see
<figureCitation id="135C2A12FFF3E67264132A60FA9EE22D" box="[1284,1376,836,861]" captionStart="FIGURE 16" captionStartId="29.[151,250,1953,1975]" captionTargetBox="[174,1413,193,1931]" captionTargetId="figure@29.[174,1413,193,1932]" captionTargetPageId="29" captionText="FIGURE 16. Enhydrosoma kosmetron sp. nov., SEM photographs, A-D, paratype ♀ 1; E &amp; F, paratype ♀ 2; G &amp; H, paratype ♂ 1: A, habitus, dorsal; B, cephalic shield, dorsal; C, sensillum in central part of cephalic shield; D, anal somite and caudal rami, dorsal; E, rostrum, lateral; F, rostrum and left antennula, lateral; G, habitus, dorsal; H, anal somite and caudal rami, dorsal." httpUri="https://zenodo.org/record/245672/files/figure.png" pageId="35" pageNumber="486">Figs. 16</figureCitation>
A, G, 17A, 18A), we are not able to report on any significant morphological variability. The number of ventro-lateral spinules on fourth male and female urosomite and third male urosomite varies between three and five.
</paragraph>
</subSubSection>
</treatment>
</document>