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<document id="70DEEDB572AE991E6AD495D318D5EEC1" ID-CLB-Dataset="45917" ID-DOI="10.5281/zenodo.279260" ID-GBIF-Dataset="ffbcaa56-6a77-4df4-b944-508fb074171f" ID-ISSN="1175-5326" ID-Zenodo-Dep="279260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460413266680" checkinUser="plazi" docAuthor="Shear, William A." docDate="2011" docId="03C47A60FFF4726360C058357EC67A44" docLanguage="en" docName="zt03109p038.pdf" docOrigin="Zootaxa 3109" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pseudotremia johnholsingeri Shear, 2011, n. sp." docType="treatment" docVersion="9" lastPageNumber="34" masterDocId="FFFD0218FFD5724260575C087F167D21" masterDocTitle="Cave millipeds of the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="34" updateTime="1698301493022" updateUser="ExternalLinkService">
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<mods:title id="6E70A88DD14A12A88BA69AC3392D8075">Cave millipeds of the United States. X. New species and records of the genus Pseudotremia Cope. 2. Species from Virginia, USA (Diplopoda, Chordeumatida, Cleidogonidae)</mods:title>
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<mods:namePart id="EFB719C8802518BC8510E8F0B4D950AA">Shear, William A.</mods:namePart>
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<mods:date id="8B916CC0ED1E647D1661D36D81B9AB54">2011</mods:date>
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<subSubSection id="C37798FDFFF4726360C058357B74782E" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BD2CB76FFF4726360C058357D507976" blockId="33.[151,582,1085,1146]" box="[151,582,1085,1111]" pageId="33" pageNumber="34">
<heading id="D09A7C1AFFF4726360C058357D507976" bold="true" box="[151,582,1085,1111]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="B9191764FFF4726360C058357D507976" bold="true" box="[151,582,1085,1111]" pageId="33" pageNumber="34">
<taxonomicName id="4C6DB0F5FFF4726360C058357EE37976" box="[151,501,1085,1111]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="johnholsingeri" status="sp. nov.">
<emphasis id="B9191764FFF4726360C058357EE37976" bold="true" box="[151,501,1085,1111]" italics="true" pageId="33" pageNumber="34">Pseudotremia johnholsingeri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22AAA1FFFF47263625358357D507976" box="[516,582,1085,1111]" pageId="33" pageNumber="34" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD2CB76FFF4726360C0586A7E23795B" blockId="33.[151,582,1085,1146]" box="[151,309,1122,1146]" pageId="33" pageNumber="34">
<figureCitation id="1356D7F3FFF4726360C0586A7E23795B" box="[151,309,1122,1146]" captionStart="FIGURES 110 113" captionStartId="34.[151,264,1312,1335]" captionTargetBox="[151,1436,153,1289]" captionTargetId="figure@34.[151,1436,153,1289]" captionTargetPageId="34" captionText="FIGURES 110 113. Pseudotremia johnholsingeri, n. sp. Fig. 110, ocelli. Fig. 111, metazonite, dorsal view. Fig. 112. Metazonite, lateral view. Fig. 113, gonopods and ninth legpair, ventral view." httpUri="https://zenodo.org/record/279278/files/figure.png" pageId="33" pageNumber="34">Figs. 110113</figureCitation>
</paragraph>
<paragraph id="8BD2CB76FFF4726360C058A57B74782E" blockId="33.[151,1437,1197,1893]" pageId="33" pageNumber="34">
<emphasis id="B9191764FFF4726360C058A57FF579E4" bold="true" box="[151,227,1197,1221]" pageId="33" pageNumber="34">
<typeStatus id="54D675D4FFF4726360C058A57FCC79E4" box="[151,218,1197,1221]" pageId="33" pageNumber="34">Types</typeStatus>
:
</emphasis>
Male
<typeStatus id="54D675D4FFF47263617A58A57E9979E4" box="[301,399,1197,1221]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
and two female
<typeStatus id="54D675D4FFF47263621E58A67DA279E7" box="[585,692,1198,1222]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
from Carter Cave,
<quantity id="4C956693FFF4726363DA58A57CA979E4" box="[909,959,1197,1221]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="33" pageNumber="34" unit="mi" value="6.0">6 mi</quantity>
SW of Stickleyville,
<emphasis id="B9191764FFF4726364E758A67A1879E4" box="[1200,1294,1198,1221]" italics="true" pageId="33" pageNumber="34">Lee Co.,</emphasis>
VIRGINIA, collected
<date id="FFD3EDB6FFF47263615458DA7EBE79CB" box="[259,424,1234,1259]" pageId="33" pageNumber="34" value="1977-08-09">9 August 1977</date>
by John Holsinger. The gonopods, ninth legs of the male, segments and head of the
<typeStatus id="54D675D4FFF47263653758DA7FDC7831" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
, and genitalia of one of the female
<typeStatus id="54D675D4FFF47263620358F07DD57831" box="[596,707,1272,1296]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
, are mounted on SEM stub WS242.
</paragraph>
</subSubSection>
<subSubSection id="C37798FDFFF47263609059157DE4785E" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BD2CB76FFF47263609059157DE4785E" blockId="33.[151,1437,1197,1893]" pageId="33" pageNumber="34">
<emphasis id="B9191764FFF47263609059157E557817" bold="true" box="[199,323,1309,1334]" pageId="33" pageNumber="34">Diagnosis:</emphasis>
<taxonomicName id="4C6DB0F5FFF47263611D59157D987814" box="[330,654,1309,1333]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="johnholsingeri">
<emphasis id="B9191764FFF47263611D59157D987814" box="[330,654,1309,1333]" italics="true" pageId="33" pageNumber="34">Pseudotremia johnholsingeri</emphasis>
</taxonomicName>
is similar to
<emphasis id="B9191764FFF47263637759167CE67814" box="[800,1008,1309,1333]" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="4C6DB0F5FFF47263637759167CFA7814" box="[800,1004,1309,1333]" class="Diplopoda" family="Cleidogonidae" genus="Pseudotremia" kingdom="Animalia" order="Chordeumatida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="jaculohamatum">P. jaculohamatum</taxonomicName>
,
</emphasis>
but differs in that the VCP is a simple curved spine and the LAPs are longer and curved inward; the ninth legs of the male have a very large basal knob (fig. 113) distal to which is a smaller, ridge-like knob.
</paragraph>
</subSubSection>
<subSubSection id="C37798FDFFF47263609059857B5A78CE" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BD2CB76FFF47263609059857B5A78CE" blockId="33.[151,1437,1197,1893]" pageId="33" pageNumber="34">
<emphasis id="B9191764FFF47263609059857E477884" bold="true" box="[199,337,1421,1445]" pageId="33" pageNumber="34">Etymology:</emphasis>
The species epithet honors the collector, amphipod authority and speleobiologist John Holsinger, of Old Dominion University,
<collectingCountry id="F37A8BE6FFF4726361BE59BA7D5378EB" box="[489,581,1458,1482]" name="Norfolk Island" pageId="33" pageNumber="34">Norfolk</collectingCountry>
, Virginia. No one else has collected more animals from caves in Virginia, and likely no one ever will. Suggested vernacular name: Holsingers Cave Milliped.
</paragraph>
</subSubSection>
<subSubSection id="C37798FDFFF47263609059F57EC67A44" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD2CB76FFF47263609059F57A277BA4" blockId="33.[151,1437,1197,1893]" pageId="33" pageNumber="34">
<emphasis id="B9191764FFF47263609059F57E4D7B37" bold="true" box="[199,347,1533,1558]" pageId="33" pageNumber="34">Description:</emphasis>
Male
<typeStatus id="54D675D4FFF4726361F659F57D157B34" box="[417,515,1533,1557]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
about
<quantity id="4C956693FFF47263621959F57D8A7B34" box="[590,668,1533,1557]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="33" pageNumber="34" unit="mm" value="31.0">31 mm</quantity>
long,
<quantity id="4C956693FFF4726362B559F57C217B34" box="[738,823,1533,1557]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="33" pageNumber="34" unit="mm" value="2.9">2.9 mm</quantity>
wide, third antennal segment
<quantity id="4C956693FFF4726364D159F57BFF7B34" box="[1158,1257,1533,1557]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="33" pageNumber="34" unit="mm" value="1.75">1.75 mm</quantity>
long. Ocelli 19 on each side, pigmented. Segmental shoulders pronounced, both lobes with distinct rims; metazonites moderately rough with low nodules, two or three especially distinct nodules along posterior margin (fig. 111); lateral striae 10 12, mostly complete (fig. 112). Coloration medium purplish-brown, mottled darker especially anteriorly.
</paragraph>
<paragraph id="8BD2CB76FFF4726360905A9A7C557A1E" blockId="33.[151,1437,1197,1893]" pageId="33" pageNumber="34">Gonopods (fig. 113) with angiocoxites basally well-separated, converging then diverging distally; MAPs with long, anteriodorsally directed subapical spines, shorter, broader apical spines directed mesally; LAPs moderately long, apically curved inward. Colpocoxites not mitten-shaped, lobular; VCP moderately long, curved, acute. Ninth Legs (fig. 113) typical of pigmented, large species, coxoprefemora with very large basal knob under which the colpocoxites insert, smaller shelf-like knob more distally.</paragraph>
<paragraph id="8BD2CB76FFF4726360905B457EC67A44" blockId="33.[151,1437,1197,1893]" box="[199,464,1869,1893]" pageId="33" pageNumber="34">Female similar to male.</paragraph>
</subSubSection>
</treatment>
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