312 lines
34 KiB
XML
312 lines
34 KiB
XML
<document id="A8BCA033B996C6D7F302592B1EC7EE47" ID-DOI="10.11646/zootaxa.3731.4.4" ID-GBIF-Dataset="26facfec-e615-4814-88c5-c966fcbc9efc" ID-ISSN="1175-5326" ID-Zenodo-Dep="248259" ID-ZooBank="2066C0D1-2ED6-41EB-AF3C-3DAB58EA1718" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460694621800" checkinUser="plazi" docAuthor="Krikken, Jan & Li, Chun-Lin" docDate="2013" docId="03ABBA06FFA5FFDAA181FDDBFCCBF83E" docLanguage="en" docName="zt03731p519.pdf" docOrigin="Zootaxa 3731 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bolbochromus (Bolbochromus) dumogensis Krikken" docType="treatment" docVersion="9" lastPageNumber="512" masterDocId="FF92C27EFFABFFCBA116FFB7FFCBFFE1" masterDocTitle="Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae)" masterLastPageNumber="519" masterPageNumber="495" pageNumber="509" updateTime="1698353700889" updateUser="plazi">
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<mods:mods id="6BC8815C35B8922FF3949261A598F4AD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4A1A86E4B1DD34276236B0BFBD74A9ED">
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<mods:title id="1922E756BFE358BE2E3138070DC999B9">Taxonomy of the Oriental genus Bolbochromus: a generic overview and descriptions of four new species (Coleoptera: Geotrupidae: Bolboceratinae)</mods:title>
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<mods:name id="254A77A6D4A019776C521697C1878D0F" type="personal">
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<mods:roleTerm id="5677DFCCB4CE496E8A053FD60FD58A0F">Author</mods:roleTerm>
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<mods:namePart id="729EF9E8B379A0E8339F015ED98190E1">Krikken, Jan</mods:namePart>
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<mods:name id="BCD5439321BCA559CC63877C5FEF60FF" type="personal">
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<mods:roleTerm id="CB7C35707C44BD37CBCC5F46669772EA">Author</mods:roleTerm>
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<mods:namePart id="D1EC759CA52EA01ABFB9FF4CE51AB4E5">Li, Chun-Lin</mods:namePart>
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<mods:typeOfResource id="8D5804EAC713DCF6672EEC43BAA63079">text</mods:typeOfResource>
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<mods:title id="E3015C5961D3EF08AF0F81367939E663">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="72D5D1B3C98A22C0A41020E9EF2ADEEF">
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<mods:date id="346A96427089751D3AC78E261E5EAA80">2013</mods:date>
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<mods:detail id="D61B4213633A0F3905F5ADB3CA03B6AD" type="volume">
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<mods:number id="A5DEC99B3F65D4D6345050E9ED42DF9A">3731</mods:number>
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<mods:detail id="2954E879C04836E00564358F99F8B4C9" type="issue">
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<mods:number id="EBF692B8442AEF9F1F71F49181721ABF">4</mods:number>
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<mods:start id="584C929F3E50EF9A63A4E3EF533BA30D">495</mods:start>
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<mods:end id="F3DC64C21540ADFD051DB81355B44045">519</mods:end>
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<mods:classification id="F64BC20E4FFCF49E9059D40B3E5F72C0">journal article</mods:classification>
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<mods:identifier id="77289687D99327CA715551D5C1127316" type="DOI">10.11646/zootaxa.3731.4.4</mods:identifier>
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<mods:identifier id="C498E79997BE45E173089DE060BDE32A" type="GBIF-Dataset">26facfec-e615-4814-88c5-c966fcbc9efc</mods:identifier>
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<mods:identifier id="486D147D52BDF0FC46602E9917524F58" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="580EF9B6DFF379411C19406035EE8490" type="Zenodo-Dep">248259</mods:identifier>
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<treatment id="03ABBA06FFA5FFDAA181FDDBFCCBF83E" ID-DOI="http://doi.org/10.5281/zenodo.5680765" ID-GBIF-Taxon="119581792" ID-Zenodo-Dep="5680765" LSID="urn:lsid:plazi:treatment:03ABBA06FFA5FFDAA181FDDBFCCBF83E" httpUri="http://treatment.plazi.org/id/03ABBA06FFA5FFDAA181FDDBFCCBF83E" lastPageId="17" lastPageNumber="512" pageId="14" pageNumber="509">
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<subSubSection id="C318589BFFA5FFC5A181FDDBFE71FD46" pageId="14" pageNumber="509" type="nomenclature">
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<paragraph id="8BBD0B10FFA5FFC5A181FDDBFC36FD67" blockId="14.[151,1021,620,679]" box="[151,1021,620,646]" pageId="14" pageNumber="509">
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<heading id="D0F5BC7CFFA5FFC5A181FDDBFC36FD67" bold="true" box="[151,1021,620,646]" fontSize="11" level="1" pageId="14" pageNumber="509" reason="1">
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<emphasis id="B976D702FFA5FFC5A181FDDBFC36FD67" bold="true" box="[151,1021,620,646]" pageId="14" pageNumber="509">
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<taxonomicName id="4C027093FFA5FFC5A181FDDBFCD6FD67" authority="Krikken" authorityName="Krikken" box="[151,797,620,646]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="509" phylum="Arthropoda" rank="species" species="dumogensis" subGenus="Bolbochromus">
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<emphasis id="B976D702FFA5FFC5A181FDDBFE82FD67" bold="true" box="[151,329,620,646]" italics="true" pageId="14" pageNumber="509">Bolbochromus</emphasis>
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(
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<emphasis id="B976D702FFA5FFC5A04FFDDBFDC1FD67" bold="true" box="[345,522,620,646]" italics="true" pageId="14" pageNumber="509">Bolbochromus</emphasis>
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)
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<emphasis id="B976D702FFA5FFC5A30AFDDBFD60FD67" bold="true" box="[540,683,620,646]" italics="true" pageId="14" pageNumber="509">dumogensis</emphasis>
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Krikken
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</taxonomicName>
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& Li, new species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A181FD39FE71FD46" blockId="14.[151,1021,620,679]" box="[151,442,654,679]" pageId="14" pageNumber="509">
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(
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<figureCitation id="13391795FFA5FFC5A189FD39FEC0FD46" box="[159,267,654,679]" captionStart="FIGURES 1 – 8" captionStartId="8.[151,264,1963,1985]" captionTargetBox="[189,1394,199,1948]" captionTargetId="figure@8.[183,1410,193,1949]" captionTargetPageId="8" captionText="FIGURES 1 – 8. Habitus of male Bolbochromus (oblique and lateral views). 1 – 2, B. nigriceps; 3 – 4, B. mindanaicus; 5 – 6, B. celebensis; 7 – 8, B. dumogensis. Scale lines 1 mm." httpUri="https://zenodo.org/record/248260/files/figure.png" pageId="14" pageNumber="509">Figs. 7–8</figureCitation>
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,
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<figureCitation id="13391795FFA5FFC5A000FD39FE94FD46" box="[278,351,654,679]" captionStart="FIGURES 25 – 30" captionStartId="13.[151,264,1766,1788]" captionTargetBox="[158,1436,205,1734]" captionTargetId="figure@13.[151,1436,195,1742]" captionTargetPageId="13" captionText="FIGURES 25 – 30. Bolbochromus holotypes, body parts, head (full-face), pronotum (dorsal), left elytron (dorsal). 25 – 27, B. dumogensis; 28 – 30, B. pumilus. Scale lines 1 mm." httpUri="https://zenodo.org/record/248263/files/figure.png" pageId="14" pageNumber="509">25–27</figureCitation>
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,
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<figureCitation id="13391795FFA5FFC5A07CFD39FE42FD46" box="[362,393,654,679]" captionStart="FIGURES 45 – 46" captionStartId="17.[151,264,1626,1648]" captionTargetBox="[152,1419,193,1586]" captionTargetId="figure@17.[151,1436,193,1605]" captionTargetPageId="17" captionText="FIGURES 45 – 46. Bolbochromus, anterolateral corner of pronotum (dorsolateral view). 45, B. celebensis; 46, B. dumogensis. Figs. 47 – 61. Aedeagus of Bolbochromus (various views). 47 – 49, B. laetus or near (47 upper side, 48 underside, 49 dextrolateral); 50, B. celebensis (upper side); 51, B. dumogensis (upper side); 52 – 53, B. nigriceps (52 upper side, 53 underside); 54, B. mindanaicus (upper side); 55, B. pumilus (upper side); 56 – 57, B. ludekingi (56 upper side, 57 underside); 58 – 59, B. sulcicollis (58 upper side, 59 dextro-lateral); 60 – 61, B. sinensis (60 dextro-lateral, 61 upper side of bent distal part). Scale lines 1 mm." httpUri="https://zenodo.org/record/248265/files/figure.png" pageId="14" pageNumber="509">46</figureCitation>
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, 51)
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</paragraph>
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</subSubSection>
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<subSubSection id="C318589BFFA5FFC5A181FD61FD77FBD3" pageId="14" pageNumber="509" type="materials_examined">
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<paragraph id="8BBD0B10FFA5FFC5A181FD61FE46FCBA" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">
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<emphasis id="B976D702FFA5FFC5A181FD61FE32FD0E" bold="true" box="[151,505,726,751]" pageId="14" pageNumber="509">
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Material examined.
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<typeStatus id="54B9B5B2FFA5FFC5A09AFD61FE32FD0E" box="[396,505,726,751]" pageId="14" pageNumber="509" type="holotype">Holotype</typeStatus>
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</emphasis>
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male (BMNH) from
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<collectingCountry id="F3154B80FFA5FFC5A3E6FD61FCA9FD0E" box="[752,866,726,751]" name="Indonesia" pageId="14" pageNumber="509">Indonesia</collectingCountry>
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: Sulawesi Utara [North Sulawesi]: Dumoga-Bone NP, labelled ”R. Ent. Soc. Lond.\ PROJECT WALLACE\ B.M. 1985-10”, “1.”.
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<emphasis id="B976D702FFA5FFC5A53DFD4DFB53FCF2" bold="true" box="[1067,1176,762,787]" pageId="14" pageNumber="509">
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<typeStatus id="54B9B5B2FFA5FFC5A53DFD4DFB53FCF2" box="[1067,1176,762,787]" pageId="14" pageNumber="509" type="paratype">Paratype</typeStatus>
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</emphasis>
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female (BMNH) from same region, with same label, and additional data:
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<date id="FFBC2DD0FFA5FFC5A3D8FCA9FCBEFCD6" box="[718,885,798,823]" pageId="14" pageNumber="509" value="1985-02">February 1985</date>
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, flight interception trap 3, plot D [?, illegible], ca
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<quantity id="4CFAA6F5FFA5FFC5A181FCF4FF11FCBA" box="[151,218,835,859]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="509" unit="m" value="400.0">400m</quantity>
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, lowland forest.
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</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A1D1FCD1FD77FBD3" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">
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<emphasis id="B976D702FFA5FFC5A1D1FCD1FEBBFC9E" bold="true" box="[199,368,870,895]" pageId="14" pageNumber="509">Identification.</emphasis>
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This species has 5 striae between the suture and the humeral umbone, like the preceding species. Compared to
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<taxonomicName id="4C027093FFA5FFC5A087FC3CFD64FC42" box="[401,687,907,931]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="509" phylum="Arthropoda" rank="species" species="celebensis">
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<emphasis id="B976D702FFA5FFC5A087FC3CFD64FC42" box="[401,687,907,931]" italics="true" pageId="14" pageNumber="509">Bolbochromus celebensis</emphasis>
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</taxonomicName>
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, however, it has (in full-face view) larger eye foramina, due to its very slender, tapering eye canthi. Interocular part of frons, between ends of paraocular ridge on either side, with fine transverse ridge over fine paramedian denticles, not a full tubercle. General colour brown, head black. Anterolateral section of pronotum (full-face view) fully rounded,
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<emphasis id="B976D702FFA5FFC5A26FFC40FC50FBEF" box="[889,923,1015,1038]" italics="true" pageId="14" pageNumber="509">i.e.</emphasis>
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, actually entire lateral border evenly rounded (non-angular). Body length roughly 8.0–
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<quantity id="4CFAA6F5FFA5FFC5A349FBADFD7CFBD3" box="[607,695,1050,1075]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="509" unit="mm" value="9.5">9.5 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C318589BFFA5FFDBA1D1FB89FC1DF839" lastPageId="16" lastPageNumber="511" pageId="14" pageNumber="509" type="description">
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<paragraph id="8BBD0B10FFA5FFC5A1D1FB89FB8BFB9A" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">
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<emphasis id="B976D702FFA5FFC5A1D1FB89FE99FBB6" bold="true" box="[199,338,1086,1111]" pageId="14" pageNumber="509">Description</emphasis>
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(
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<typeStatus id="54B9B5B2FFA5FFC5A074FB89FE02FBB6" box="[354,457,1086,1111]" pageId="14" pageNumber="509" type="holotype">holotype</typeStatus>
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, male). Body length approximately
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<quantity id="4CFAA6F5FFA5FFC5A275FB88FC7CFBB7" box="[867,951,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="509" unit="mm" value="10.0">10 mm</quantity>
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. Colour generally medium-brown, dorsum glossy, almost glabrous. Pilosity on underside and legs abundant, long, light brown.
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</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A1D1FB31FA57FAB2" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">Labrum with very vaguely emarginate anteromedian border, upper surface rugulate. Clypeal surface bordered by fine, widely arcuate perimarginal ridge, Clypeogenal angle hardly prominent. Clypeus and frons with abundant to dense, scattered, fine primary punctation, interspersed with minute secondary punctures. Eye canthus generally narrow, slender, anterior edge slightly, finely raised, widely arcuate from clypeogenal angle, tapering to temporal lobe; paraocular ridge (issuing from clypeogenal angle) fine, extending to lateral angle of low, transverse, interocular ridge; medial section of ridge with pair of very fine denticles (axial view). Eye foramen relatively large.</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A1D1FAE8FB08F9CA" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">Pronotum moderately convex, gradually declivous in front, outline of marginate (raised) anterior border evenly arcuate; pronotal midline with punctate sulcus from near base to apex; anterolateral edge fully rounded (full-face view), entire lateral edge widely rounded to base. Pronotal surface with double punctation, primary punctation (size slightly variable) distinct, abundant around lateral fovea, also lining anterior and lateral marginal ridge; secondary punctation sparse, minute. Pronotal base immarginate on section between elytral stria 5 on either side (outline medially slightly expanded- rounded). Scutellum with scattered, very sparse double punctation.</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A1D1F981FC9DF95B" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">Elytra with discal striae 1–5 reaching base between suture and humeral umbone; distinctly impressed, finely punctate; punctures separated by 4–7 puncture diameters, slightly crenulating interstriae. Elytral interstriae (on cross-section) moderately convex, interstria 1 distally slightly more convex; interstrial surface sparsely micropunctate. Punctures in juxtepipleural stria with long seta.</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A1D1F971FBDFF8AA" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">Protibia with 10 external denticles, proximally decreasing in size; apex unmodified, with robust, complanate, slightly tapering spur. Outer side of mesotibiae and metatibiae with 4+(1) and 3 arcuate anteapical fossorial elevations, respectively; their crest fringed with long, robust, acuminate spines; apical crest transversely subelliptic, with similar spines; terminal spurs long, apex blunt, slightly curved.</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A1D1F8E1FC26F88E" blockId="14.[151,1436,726,2011]" box="[199,1005,1878,1903]" pageId="14" pageNumber="509">Aedeagus, Fig. 51; slender-elongate, slightly transparent-membraneous.</paragraph>
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<paragraph id="8BBD0B10FFA5FFC5A1D1F8CCFBB4F83A" blockId="14.[151,1436,726,2011]" pageId="14" pageNumber="509">Measurements of respective parts in mm. Median length of head (full-face, excluding labrum and mandibles) 2.1, width 3.2. Median length of pronotum (dorsal) 3.6, maximum width 5.5. Median length of scutellum 1.0, maximum width 1.1. Sutural length of elytra (dorsal) 2.9, maximum width combined 5.9.</paragraph>
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<paragraph id="8BBD0B10FFA4FFC4A1D1FF20FB7AFF34" blockId="15.[151,1435,151,213]" pageId="15" pageNumber="510">
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<emphasis id="B976D702FFA4FFC4A1D1FF20FDA8FF51" bold="true" box="[199,611,151,176]" pageId="15" pageNumber="510">Variation and sexual dimorphism.</emphasis>
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Only two specimens available (male and female). Variation in size may well be larger than here indicated (see size range in
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<taxonomicName id="4C027093FFA4FFC4A3CAFF0AFCA5FF35" box="[732,878,188,212]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="510" phylum="Arthropoda" rank="species" species="celebensis">
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<emphasis id="B976D702FFA4FFC4A3CAFF0AFCA5FF35" box="[732,878,188,212]" italics="true" pageId="15" pageNumber="510">B. celebensis</emphasis>
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</taxonomicName>
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). The sexes are very similar.
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</paragraph>
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<caption id="DF7D5B98FFA4FFC4A181F881FD54F88A" httpUri="https://zenodo.org/record/248264/files/figure.png" pageId="15" pageNumber="510" targetBox="[151,1433,273,1814]" targetPageId="15">
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<paragraph id="8BBD0B10FFA4FFC4A181F881FD54F88A" blockId="15.[151,1436,1846,1899]" pageId="15" pageNumber="510">
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<emphasis id="B976D702FFA4FFC4A181F881FE84F8AD" bold="true" box="[151,335,1846,1868]" pageId="15" pageNumber="510">FIGURES 31–36</emphasis>
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.
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<taxonomicName id="4C027093FFA4FFC4A048F881FE3AF8AD" box="[350,497,1846,1868]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="510" phylum="Arthropoda" rank="genus">
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<emphasis id="B976D702FFA4FFC4A048F881FE3AF8AD" box="[350,497,1846,1868]" italics="true" pageId="15" pageNumber="510">Bolbochromus</emphasis>
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</taxonomicName>
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holotypes, body parts, head (full-face), pronotum (dorsal), left elytron (dorsal). 31–33,
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<taxonomicName id="4C027093FFA4FFC4A490F880FED3F88A" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="510" phylum="Arthropoda" rank="species" species="mindanaicus">
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<emphasis id="B976D702FFA4FFC4A490F880FED3F88A" italics="true" pageId="15" pageNumber="510">B. mindanaicus</emphasis>
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</taxonomicName>
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; 34–36,
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<taxonomicName id="4C027093FFA4FFC4A067F8E1FE16F88A" box="[369,477,1878,1899]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="510" phylum="Arthropoda" rank="species" species="sinensis">
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<emphasis id="B976D702FFA4FFC4A067F8E1FE4BF88A" box="[369,384,1878,1899]" italics="true" pageId="15" pageNumber="510">B</emphasis>
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.
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<emphasis id="B976D702FFA4FFC4A09BF8E1FE16F88A" box="[397,477,1878,1899]" italics="true" pageId="15" pageNumber="510">sinensis</emphasis>
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</taxonomicName>
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. Scale lines 1 mm.
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</paragraph>
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</caption>
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<paragraph id="8BBD0B10FFBBFFDBA181F814FC1DF839" blockId="16.[151,1435,1955,2008]" pageId="16" pageNumber="511">
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<emphasis id="B976D702FFBBFFDBA181F814FE9DF858" bold="true" box="[151,342,1955,1977]" pageId="16" pageNumber="511">FIGURES 37–44.</emphasis>
|
||
<taxonomicName id="4C027093FFBBFFDBA048F814FE3AF858" box="[350,497,1955,1977]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976D702FFBBFFDBA048F814FE3AF858" box="[350,497,1955,1977]" italics="true" pageId="16" pageNumber="511">Bolbochromus</emphasis>
|
||
</taxonomicName>
|
||
head (full-face). 37,
|
||
<taxonomicName id="4C027093FFBBFFDBA3C6F813FC87F858" box="[720,844,1956,1977]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="nigriceps">
|
||
<emphasis id="B976D702FFBBFFDBA3C6F813FC87F858" box="[720,844,1956,1977]" italics="true" pageId="16" pageNumber="511">B. nigriceps</emphasis>
|
||
</taxonomicName>
|
||
; 38,
|
||
<taxonomicName id="4C027093FFBBFFDBA292F813FBC0F858" box="[900,1035,1955,1977]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="celebensis">
|
||
<emphasis id="B976D702FFBBFFDBA292F813FC58F858" box="[900,915,1956,1977]" italics="true" pageId="16" pageNumber="511">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBBFFDBA2B5F814FBC0F858" box="[931,1035,1955,1977]" italics="true" pageId="16" pageNumber="511">celebensis</emphasis>
|
||
</taxonomicName>
|
||
; 39,
|
||
<taxonomicName id="4C027093FFBBFFDBA555F813FB09F858" box="[1091,1218,1956,1977]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="catenatus">
|
||
<emphasis id="B976D702FFBBFFDBA555F813FB99F858" box="[1091,1106,1956,1977]" italics="true" pageId="16" pageNumber="511">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBBFFDBA577F813FB09F858" box="[1121,1218,1956,1977]" italics="true" pageId="16" pageNumber="511">catenatus</emphasis>
|
||
</taxonomicName>
|
||
; 40,
|
||
<taxonomicName id="4C027093FFBBFFDBA5ECF813FA85F858" box="[1274,1358,1956,1977]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="B976D702FFBBFFDBA5ECF813FAC2F858" box="[1274,1289,1956,1977]" italics="true" pageId="16" pageNumber="511">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBBFFDBA40FF813FA85F858" box="[1305,1358,1956,1977]" italics="true" pageId="16" pageNumber="511">niger</emphasis>
|
||
</taxonomicName>
|
||
; 41,
|
||
<taxonomicName id="4C027093FFBBFFDBA490F813FF35F839" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="sulcicollis">
|
||
<emphasis id="B976D702FFBBFFDBA490F813FA5EF858" box="[1414,1429,1956,1977]" italics="true" pageId="16" pageNumber="511">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBBFFDBA181F875FF35F839" box="[151,254,1986,2008]" italics="true" pageId="16" pageNumber="511">sulcicollis</emphasis>
|
||
</taxonomicName>
|
||
; 42,
|
||
<taxonomicName id="4C027093FFBBFFDBA027F874FE60F839" box="[305,427,1986,2008]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="ludekingi">
|
||
<emphasis id="B976D702FFBBFFDBA027F874FE8BF839" box="[305,320,1987,2008]" italics="true" pageId="16" pageNumber="511">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBBFFDBA05BF875FE60F839" box="[333,427,1986,2008]" italics="true" pageId="16" pageNumber="511">ludekingi</emphasis>
|
||
</taxonomicName>
|
||
; 43,
|
||
<taxonomicName id="4C027093FFBBFFDBA0C8F874FDF6F839" box="[478,573,1986,2008]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="walshi">
|
||
<emphasis id="B976D702FFBBFFDBA0C8F874FE26F839" box="[478,493,1987,2008]" italics="true" pageId="16" pageNumber="511">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBBFFDBA0EDF875FDF6F839" box="[507,573,1986,2008]" italics="true" pageId="16" pageNumber="511">walshi</emphasis>
|
||
</taxonomicName>
|
||
; 44,
|
||
<taxonomicName id="4C027093FFBBFFDBA366F874FD03F839" box="[624,712,1986,2008]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="511" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B976D702FFBBFFDBA366F874FDB4F839" box="[624,639,1987,2008]" italics="true" pageId="16" pageNumber="511">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBBFFDBA39AF875FD03F839" box="[652,712,1986,2008]" italics="true" pageId="16" pageNumber="511">laetus</emphasis>
|
||
</taxonomicName>
|
||
or near. Scale lines
|
||
<quantity id="4CFAA6F5FFBBFFDBA282F875FC19F839" box="[916,978,1986,2008]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="511" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7D5B98FFBAFFDAA181F9EDFAB3F90A" httpUri="https://zenodo.org/record/248265/files/figure.png" pageId="17" pageNumber="512" targetBox="[152,1419,193,1586]" targetPageId="17">
|
||
<paragraph id="8BBD0B10FFBAFFDAA181F9EDFAB3F90A" blockId="17.[151,1436,1626,1771]" pageId="17" pageNumber="512">
|
||
<emphasis id="B976D702FFBAFFDAA181F9EDFE9EF991" bold="true" box="[151,341,1626,1648]" pageId="17" pageNumber="512">FIGURES 45–46.</emphasis>
|
||
<taxonomicName id="4C027093FFBAFFDAA04AF9EDFE24F991" box="[348,495,1626,1648]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976D702FFBAFFDAA04AF9EDFE24F991" box="[348,495,1626,1648]" italics="true" pageId="17" pageNumber="512">Bolbochromus</emphasis>
|
||
</taxonomicName>
|
||
, anterolateral corner of pronotum (dorsolateral view). 45,
|
||
<taxonomicName id="4C027093FFBAFFDAA550F9ECFB00F991" box="[1094,1227,1626,1648]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="celebensis">
|
||
<emphasis id="B976D702FFBAFFDAA550F9ECFB9EF991" box="[1094,1109,1627,1648]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA575F9EDFB00F991" box="[1123,1227,1626,1648]" italics="true" pageId="17" pageNumber="512">celebensis</emphasis>
|
||
</taxonomicName>
|
||
; 46,
|
||
<taxonomicName id="4C027093FFBAFFDAA417F9ECFA5EF991" box="[1281,1429,1626,1648]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="dumogensis">
|
||
<emphasis id="B976D702FFBAFFDAA417F9ECFADBF991" box="[1281,1296,1627,1648]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA408F9EDFA5EF991" box="[1310,1429,1626,1648]" italics="true" pageId="17" pageNumber="512">dumogensis</emphasis>
|
||
</taxonomicName>
|
||
. Figs. 47–61. Aedeagus of
|
||
<taxonomicName id="4C027093FFBAFFDAA0BCF9CEFDF6F96E" box="[426,573,1657,1679]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976D702FFBAFFDAA0BCF9CEFDF6F96E" box="[426,573,1657,1679]" italics="true" pageId="17" pageNumber="512">Bolbochromus</emphasis>
|
||
</taxonomicName>
|
||
(various views). 47–49,
|
||
<taxonomicName id="4C027093FFBAFFDAA254F9CDFC56F96E" box="[834,925,1657,1679]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B976D702FFBAFFDAA254F9CDFC56F96E" box="[834,925,1657,1679]" italics="true" pageId="17" pageNumber="512">B. laetus</emphasis>
|
||
</taxonomicName>
|
||
or near (47 upper side, 48 underside, 49 dextrolateral); 50,
|
||
<taxonomicName id="4C027093FFBAFFDAA007F92EFE5FF94C" box="[273,404,1687,1710]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="celebensis">
|
||
<emphasis id="B976D702FFBAFFDAA007F92EFEEBF94F" box="[273,288,1689,1710]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA03AF920FE5FF94C" box="[300,404,1687,1709]" italics="true" pageId="17" pageNumber="512">celebensis</emphasis>
|
||
</taxonomicName>
|
||
(upper side); 51,
|
||
<taxonomicName id="4C027093FFBAFFDAA353F92EFD13F94C" box="[581,728,1687,1710]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="dumogensis">
|
||
<emphasis id="B976D702FFBAFFDAA353F92EFD9FF94F" box="[581,596,1689,1710]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA377F920FD13F94C" box="[609,728,1687,1709]" italics="true" pageId="17" pageNumber="512">dumogensis</emphasis>
|
||
</taxonomicName>
|
||
(upper side); 52–53,
|
||
<taxonomicName id="4C027093FFBAFFDAA2B9F92EFBE3F94F" box="[943,1064,1688,1710]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="nigriceps">
|
||
<emphasis id="B976D702FFBAFFDAA2B9F92EFC75F94F" box="[943,958,1689,1710]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA2DCF92EFBE3F94F" box="[970,1064,1689,1710]" italics="true" pageId="17" pageNumber="512">nigriceps</emphasis>
|
||
</taxonomicName>
|
||
(52 upper side, 53 underside); 54,
|
||
<taxonomicName id="4C027093FFBAFFDAA490F92EFED2F92D" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="mindanaicus">
|
||
<emphasis id="B976D702FFBAFFDAA490F92EFA5EF94F" box="[1414,1429,1689,1710]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA181F901FED2F92D" box="[151,281,1718,1740]" italics="true" pageId="17" pageNumber="512">mindanaicus</emphasis>
|
||
</taxonomicName>
|
||
(upper side); 55,
|
||
<taxonomicName id="4C027093FFBAFFDAA0DDF900FDFCF92D" box="[459,567,1718,1740]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B976D702FFBAFFDAA0DDF900FE11F92D" box="[459,474,1719,1740]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA0F1F901FDFCF92D" box="[487,567,1718,1740]" italics="true" pageId="17" pageNumber="512">pumilus</emphasis>
|
||
</taxonomicName>
|
||
(upper side); 56–57,
|
||
<taxonomicName id="4C027093FFBAFFDAA207F900FC40F92D" box="[785,907,1718,1740]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="ludekingi">
|
||
<emphasis id="B976D702FFBAFFDAA207F900FCEBF92D" box="[785,800,1719,1740]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA23BF901FC40F92D" box="[813,907,1718,1740]" italics="true" pageId="17" pageNumber="512">ludekingi</emphasis>
|
||
</taxonomicName>
|
||
(56 upper side, 57 underside); 58–59,
|
||
<taxonomicName id="4C027093FFBAFFDAA405F900FA57F92D" box="[1299,1436,1718,1740]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="sulcicollis">
|
||
<emphasis id="B976D702FFBAFFDAA405F900FAE9F92D" box="[1299,1314,1719,1740]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA423F901FA57F92D" box="[1333,1436,1718,1740]" italics="true" pageId="17" pageNumber="512">sulcicollis</emphasis>
|
||
</taxonomicName>
|
||
(58 upper side, 59 dextro-lateral); 60–61,
|
||
<taxonomicName id="4C027093FFBAFFDAA328F961FD61F90A" box="[574,682,1750,1771]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B976D702FFBAFFDAA328F961FD86F90A" box="[574,589,1750,1771]" italics="true" pageId="17" pageNumber="512">B</emphasis>
|
||
.
|
||
<emphasis id="B976D702FFBAFFDAA34CF961FD61F90A" box="[602,682,1750,1771]" italics="true" pageId="17" pageNumber="512">sinensis</emphasis>
|
||
</taxonomicName>
|
||
(60 dextro-lateral, 61 upper side of bent distal part). Scale lines 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C318589BFFBAFFDAA1D1F8A5FDC1F876" pageId="17" pageNumber="512" type="discussion">
|
||
<paragraph id="8BBD0B10FFBAFFDAA1D1F8A5FDC1F876" blockId="17.[151,1436,1810,2015]" pageId="17" pageNumber="512">
|
||
<emphasis id="B976D702FFBAFFDAA1D1F8A5FE84F8CA" bold="true" box="[199,335,1810,1835]" pageId="17" pageNumber="512">Comments.</emphasis>
|
||
The apparently limited occurrence of this unusual
|
||
<taxonomicName id="4C027093FFBAFFDAA2D7F8A5FBA8F8CB" box="[961,1123,1810,1834]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976D702FFBAFFDAA2D7F8A5FBA8F8CB" box="[961,1123,1810,1834]" italics="true" pageId="17" pageNumber="512">Bolbochromus</emphasis>
|
||
</taxonomicName>
|
||
versus the widespread
|
||
<taxonomicName id="4C027093FFBAFFDAA492F8A4FEC1F8AF" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="species" species="celebensis">
|
||
<emphasis id="B976D702FFBAFFDAA492F8A4FEC1F8AF" italics="true" pageId="17" pageNumber="512">B. celebensis</emphasis>
|
||
</taxonomicName>
|
||
suggests that the exploration for
|
||
<taxonomicName id="4C027093FFBAFFDAA397F881FCE8F8AF" box="[641,803,1846,1870]" class="Insecta" family="Bolboceratidae" genus="Bolbochromus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B976D702FFBAFFDAA397F881FCE8F8AF" box="[641,803,1846,1870]" italics="true" pageId="17" pageNumber="512">Bolbochromus</emphasis>
|
||
</taxonomicName>
|
||
in the archipelago is far from complete, and might have more surprises in store. One would expect more specimens to be available, as the Dumoga-Bone National Park was the focal area of Project Wallace.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318589BFFBAFFDAA1D1F815FD59F85A" box="[199,658,1954,1979]" pageId="17" pageNumber="512" type="distribution">
|
||
<paragraph id="8BBD0B10FFBAFFDAA1D1F815FD59F85A" blockId="17.[151,1436,1810,2015]" box="[199,658,1954,1979]" pageId="17" pageNumber="512">
|
||
<emphasis id="B976D702FFBAFFDAA1D1F815FEABF85A" bold="true" box="[199,352,1954,1979]" pageId="17" pageNumber="512">Distribution.</emphasis>
|
||
<collectingCountry id="F3154B80FFBAFFDAA071F815FE13F85A" box="[359,472,1954,1979]" name="Indonesia" pageId="17" pageNumber="512">Indonesia</collectingCountry>
|
||
: north Sulawesi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318589BFFBAFFDAA1D1F871FCCBF83E" box="[199,768,1990,2015]" pageId="17" pageNumber="512" type="etymology">
|
||
<paragraph id="8BBD0B10FFBAFFDAA1D1F871FCCBF83E" blockId="17.[151,1436,1810,2015]" box="[199,768,1990,2015]" pageId="17" pageNumber="512">
|
||
<emphasis id="B976D702FFBAFFDAA1D1F871FE85F83E" bold="true" box="[199,334,1990,2015]" pageId="17" pageNumber="512">Etymology.</emphasis>
|
||
Name is derived from the
|
||
<typeStatus id="54B9B5B2FFBAFFDAA36FF870FD62F83E" box="[633,681,1991,2015]" pageId="17" pageNumber="512">type</typeStatus>
|
||
region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |