439 lines
59 KiB
XML
439 lines
59 KiB
XML
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<mods:title id="DAA746CEB1353B76411C5D8BFBB71A40">Morphology and multilocus phylogeny of the Spiny-footed Lizard (Acanthodactylus erythrurus) complex reveal two new mountain species from the Moroccan Atlas</mods:title>
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<mods:namePart id="AD1EF214848D96E61F2F8219FDDE7807">Miralles, Aurélien</mods:namePart>
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<mods:namePart id="CC9D3594AF8A1E55DD999BF2B25AD7B0">Geniez, Philippe</mods:namePart>
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<mods:namePart id="0755A1E92F900515A370A2AF4CFAEB96">Beddek, Menad</mods:namePart>
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<mods:namePart id="5A9D9FCD39D215812827180B85927D05">Aranda, Daniel Mendez</mods:namePart>
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<mods:namePart id="0277C20D82688A2697DE2DE6D3678635">Brito, José Carlos</mods:namePart>
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<mods:namePart id="74D7DB9C7A5D86671C1A3293418CB6A4">Leblois, Raphaël</mods:namePart>
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<mods:namePart id="03795B97E4DBA2C0CC57C4D5AFACADED">Crochet, Pierre-André</mods:namePart>
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<mods:title id="62436B32E7B2954FEFAD4DBA78EC10A9">Zootaxa</mods:title>
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<mods:date id="5EC2FF6B64CC929A8B30F55FA09393EC">2020</mods:date>
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<mods:identifier id="F17EC2DD6618FEC37028175A29C163D3" type="DOI">10.11646/zootaxa.4747.2.4</mods:identifier>
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<treatment id="039DEF5B5F5B0C1962AB6024F737FF77" ID-DOI="http://doi.org/10.5281/zenodo.3704097" ID-GBIF-Taxon="162381193" ID-Zenodo-Dep="3704097" LSID="urn:lsid:plazi:treatment:039DEF5B5F5B0C1962AB6024F737FF77" httpUri="http://treatment.plazi.org/id/039DEF5B5F5B0C1962AB6024F737FF77" lastPageId="14" lastPageNumber="315" pageId="12" pageNumber="314">
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<subSubSection id="C32E0DC65F5B0C1B62AB6024F29CFEEC" pageId="12" pageNumber="314" type="nomenclature">
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<paragraph id="8B8B5E4D5F5B0C1B62AB6024F154FEC8" blockId="12.[151,572,297,359]" box="[151,572,297,324]" pageId="12" pageNumber="314">
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<heading id="D0C3E9215F5B0C1B62AB6024F154FEC8" bold="true" box="[151,572,297,324]" fontSize="11" level="1" pageId="12" pageNumber="314" reason="1">
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<emphasis id="B940825F5F5B0C1B62AB6024F154FEC8" bold="true" box="[151,572,297,324]" pageId="12" pageNumber="314">
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<taxonomicName id="4C3425CE5F5B0C1B62AB6024F2B0FECF" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[151,472,297,324]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="314" phylum="Chordata" rank="species" species="lacrymae" status="sp. nov.">
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<emphasis id="B940825F5F5B0C1B62AB6024F2B0FECF" bold="true" box="[151,472,297,324]" italics="true" pageId="12" pageNumber="314">Acanthodactylus lacrymae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2733F245F5B0C1B63E36024F154FEC8" box="[479,572,297,323]" pageId="12" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8B5E4D5F5B0C1B62AB6040F29CFEEC" blockId="12.[151,572,297,359]" box="[151,500,333,359]" pageId="12" pageNumber="314">
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||
(Appendixes I, II;
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||
<figureCitation id="130F42C85F5B0C1B635A6040F2DCFEEC" box="[358,436,333,359]" captionStart="FIGURE 6" captionStartId="15.[151,250,1657,1682]" captionTargetBox="[159,1433,202,1601]" captionTargetId="figure@15.[151,1436,196,1605]" captionTargetPageId="15" captionText="FIGURE 6. Phenotypic variation in Acanthodactylus montanus sp. nov. and A. lacrymae sp. nov. (compare with figure 6 showing the phenotypic variation in the rest of the Acanthodactylus erythrurus group): (A) BEV.11795*, (B) BEV.11793*, (C) MNHN-RA-2018.0026 (holotype of Acanthodactylus montanus), (D) BEV.11797, (E) BEV.11988, (F) BEV.11989, (G) MNHN- RA- 2018.0027 (holotype of A. lacrymae), (H) BEV.11992, (I) BEV.11998, (J) BEV.12000, (K) BEV.11997, (L) BEV.12006. Asterisks indicate symmetrically reversed pictures. Photo credits: A to D (PAC), E to L (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694836" httpUri="https://zenodo.org/record/3694836/files/figure.png" pageId="12" pageNumber="314">Figs. 6</figureCitation>
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,
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<figureCitation id="130F42C85F5B0C1B63FC6040F2A7FEEC" box="[448,463,333,359]" captionStart="FIGURE 8" captionStartId="17.[151,250,1935,1960]" captionTargetBox="[183,1406,522,1913]" captionTargetId="figure@17.[179,1409,517,1916]" captionTargetPageId="17" captionText="FIGURE 8. Holotypes of Acanthodactylus montanus sp. nov. (MNHN-RA-2018.0026, Tizi n’Tichka) and (B) Acanthodactylus lacrymae sp. nov. (MNHN-RA-2018.0027, Tislit) with a picture of the habitat of this species (Isli lake). Photo credits:A (PAC), B (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694844" httpUri="https://zenodo.org/record/3694844/files/figure.png" pageId="12" pageNumber="314">8</figureCitation>
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,
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<figureCitation id="130F42C85F5B0C1B63E06040F283FEEC" box="[476,491,333,359]" captionStart="FIGURE 9" captionStartId="18.[151,250,1859,1884]" captionTargetBox="[198,1394,184,1829]" captionTargetId="figure@18.[185,1402,174,1840]" captionTargetPageId="18" captionText="FIGURE 9. Phenotypic variation in juveniles. Pileus of juveniles in A lacrymae sp. nov. (A, BEV.11981 & B, BEV. 11990, both from Tislit) and A. montanus sp. nov. (C, JCB.A546, Jbel Siroua & D, BEV.11792, Tizi n’Tichka). Caudal and cloacal colour patterns in juveniles of A. lacrymae sp. nov. (E, BEV.11986 & F, BEV.11990, both from Tislit), A. montanus sp. nov. (G, BEV.11798, Tizi n’Tichka & H, JCB.A546, Jbel Siroua) and A. erythrurus (I, BEV.11926, Dayet Ifrah & J, BEV.11595, Marrakech). Photo credits: A, B, E, F, I (PG), D, G, J (PAC), C, H (JCB)." figureDoi="http://doi.org/10.5281/zenodo.3694846" httpUri="https://zenodo.org/record/3694846/files/figure.png" pageId="12" pageNumber="314">9</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32E0DC65F5B0C1B62AB6098F398FB4C" pageId="12" pageNumber="314" type="materials_examined">
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<paragraph id="8B8B5E4D5F5B0C1B62AB6098F0CBFD90" blockId="12.[151,1437,405,1980]" pageId="12" pageNumber="314">
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<materialsCitation id="3B5C54105F5B0C1B62AB6098F0CBFD90" ID-GBIF-Occurrence="2573218765" collectingDate="2012-05-29" country="Morocco" county="Imilchil" elevation="2260" latitude="32.1986" location="Museum" longLatPrecision="6" longitude="-5.6292" municipality="High Atlas" pageId="12" pageNumber="314" specimenCount="1" typeStatus="holotype">
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<emphasis id="B940825F5F5B0C1B62AB6098F263FE24" bold="true" box="[151,267,405,431]" pageId="12" pageNumber="314">
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<typeStatus id="548FE0EF5F5B0C1B62AB6098F26EFE24" box="[151,262,405,431]" pageId="12" pageNumber="314" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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||
<location id="8EEB08965F5B0C1B632E6098F21CFE24" LSID="urn:lsid:plazi:treatment:039DEF5B5F5B0C1962AB6024F737FF77:8EEB08965F5B0C1B632E6098F21CFE24" box="[274,372,405,431]" country="Morocco" county="Imilchil" latitude="32.1986" longLatPrecision="6" longitude="-5.6292" municipality="High Atlas" name="Museum" pageId="12" pageNumber="314">Muséum</location>
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national
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<location id="8EEB08965F5B0C1B63E76098F125FE24" LSID="urn:lsid:plazi:treatment:039DEF5B5F5B0C1962AB6024F737FF77:8EEB08965F5B0C1B63E76098F125FE24" box="[475,589,405,431]" country="Morocco" county="Imilchil" latitude="32.1986" longLatPrecision="6" longitude="-5.6292" municipality="High Atlas" name="d'Histoire" pageId="12" pageNumber="314">d’Histoire</location>
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naturelle (Paris), MNHN-RA-2018.0027 (formerly BEV.11991, tissue sample in the BEV tissue collection, code T5986), adult male, collected on
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<date id="FF8A788D5F5B0C1B61E960B4F7EDFE58" box="[981,1157,441,467]" pageId="12" pageNumber="314" value="2012-05-29">
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<collectingDate id="EFCE81655F5B0C1B61E960B4F7EDFE58" box="[981,1157,441,467]" pageId="12" pageNumber="314" value="2012-05-29">May 29th 2012</collectingDate>
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</date>
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by
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<location id="8EEB08965F5B0C1B668860B4F602FE58" LSID="urn:lsid:plazi:treatment:039DEF5B5F5B0C1962AB6024F737FF77:8EEB08965F5B0C1B668860B4F602FE58" box="[1204,1386,441,467]" country="Morocco" county="Imilchil" latitude="32.1986" longLatPrecision="6" longitude="-5.6292" municipality="High Atlas" name="Philippe Geniez" pageId="12" pageNumber="314">Philippe Geniez</location>
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and
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<location id="8EEB08965F5B0C1B62AB60D0F235FE7C" LSID="urn:lsid:plazi:treatment:039DEF5B5F5B0C1962AB6024F737FF77:8EEB08965F5B0C1B62AB60D0F235FE7C" box="[151,349,477,503]" country="Morocco" county="Imilchil" latitude="32.1986" longLatPrecision="6" longitude="-5.6292" municipality="High Atlas" name="Aurelien Miralles" pageId="12" pageNumber="314">Aurélien Miralles</location>
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(
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<figureCitation id="130F42C85F5B0C1B635060D0F2A9FE7C" box="[364,449,477,503]" captionStart="FIGURE 8" captionStartId="17.[151,250,1935,1960]" captionTargetBox="[183,1406,522,1913]" captionTargetId="figure@17.[179,1409,517,1916]" captionTargetPageId="17" captionText="FIGURE 8. Holotypes of Acanthodactylus montanus sp. nov. (MNHN-RA-2018.0026, Tizi n’Tichka) and (B) Acanthodactylus lacrymae sp. nov. (MNHN-RA-2018.0027, Tislit) with a picture of the habitat of this species (Isli lake). Photo credits:A (PAC), B (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694844" httpUri="https://zenodo.org/record/3694844/files/figure.png" pageId="12" pageNumber="314">Fig. 8B</figureCitation>
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) along the north-eastern shore of the lake of
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||
<location id="8EEB08965F5B0C1B618A60D0F09BFE7C" LSID="urn:lsid:plazi:treatment:039DEF5B5F5B0C1962AB6024F737FF77:8EEB08965F5B0C1B618A60D0F09BFE7C" box="[950,1011,477,503]" country="Morocco" county="Imilchil" latitude="32.1986" longLatPrecision="6" longitude="-5.6292" municipality="High Atlas" name="Tislit" pageId="12" pageNumber="314">Tislit</location>
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||
, north-eastern
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||
<collectingMunicipality id="6BEFC4375F5B0C1B66A460D0F678FE7C" box="[1176,1296,477,503]" pageId="12" pageNumber="314">High Atlas</collectingMunicipality>
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chain, north of
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<collectingCounty id="62EA26C15F5B0C1B628A630CF27AFD90" box="[182,274,513,539]" pageId="12" pageNumber="314">Imilchil</collectingCounty>
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,
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<collectingCountry id="F3231EDD5F5B0C1B6320630CF2ECFD90" box="[284,388,513,539]" name="Morocco" pageId="12" pageNumber="314">Morocco</collectingCountry>
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,
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<geoCoordinate id="EE00388A5F5B0C1B63B3630CF165FD90" box="[399,525,513,539]" degrees="32.1986" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="5" value="32.1986">32.1986°N</geoCoordinate>
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,
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<geoCoordinate id="EE00388A5F5B0C1B6024630DF1E2FD90" box="[536,650,512,539]" degrees="5.6292" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="5" value="-5.6292">5.6292°W</geoCoordinate>
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(WGS84),
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<elevation id="0019B97E5F5B0C1B6130630CF0CBFD90" box="[780,931,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.26" pageId="12" pageNumber="314" unit="m" value="2260.0">
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<quantity id="4CCCF3A85F5B0C1B6130630CF000FD90" box="[780,872,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.26" pageId="12" pageNumber="314" unit="m" value="2260.0">2,260 m</quantity>
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a.s.l.
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</elevation>
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8B5E4D5F5B0C1B62FB6328F08CFDE8" blockId="12.[151,1437,405,1980]" pageId="12" pageNumber="314">
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<emphasis id="B940825F5F5B0C1B62FB6328F22EFDB4" bold="true" box="[199,326,549,575]" pageId="12" pageNumber="314">
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<typeStatus id="548FE0EF5F5B0C1B62FB6328F229FDB4" box="[199,321,549,575]" pageId="12" pageNumber="314" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
Fifteen individuals, BEV.11980–11990, 11992–11995 (tissue sample codes T5975–5985, 5987– 5990), collected at the type locality at the same date by the same observers.
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||
</paragraph>
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<paragraph id="8B8B5E4D5F5B0C1B62FB6361F094FD44" blockId="12.[151,1437,405,1980]" pageId="12" pageNumber="314">
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<materialsCitation id="3B5C54105F5B0C1B62FB6361F090FD44" ID-GBIF-Occurrence="2573218763" collectingDate="2012-05-29" collectorName="P. Geniez & A. Miralles" country="Morocco" elevation="2276" latitude="32.2182" location="Twelve" longLatPrecision="6" longitude="-5.5501" municipality="Izely" pageId="12" pageNumber="314" specimenCount="2">
|
||
<emphasis id="B940825F5F5B0C1B62FB6361F2D1FD0C" bold="true" box="[199,441,620,647]" pageId="12" pageNumber="314">Additional material.</emphasis>
|
||
<location id="8EEB08965F5B0C1B63836362F165FD0C" LSID="urn:lsid:plazi:treatment:039DEF5B5F5B0C1962AB6024F737FF77:8EEB08965F5B0C1B63836362F165FD0C" box="[447,525,623,647]" country="Morocco" latitude="32.2182" longLatPrecision="6" longitude="-5.5501" municipality="Izely" name="Twelve" pageId="12" pageNumber="314">Twelve</location>
|
||
specimens: BEV.11996–12006 (tissue sample codes T5991–6001) from the western shore of the lake of
|
||
<collectingMunicipality id="6BEFC4375F5B0C1B638D639CF282FD20" box="[433,490,657,683]" pageId="12" pageNumber="314">Izely</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F3231EDD5F5B0C1B63CB639CF137FD20" box="[503,607,657,683]" name="Morocco" pageId="12" pageNumber="314">Morocco</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B6051639CF182FD20" box="[621,746,657,683]" degrees="32.2182" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="5" value="32.2182">32.2182°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B60C4639DF003FD20" box="[760,875,656,683]" degrees="5.5501" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="5" value="-5.5501">5.5501°W</geoCoordinate>
|
||
,
|
||
<quantity id="4CCCF3A85F5B0C1B6145639CF0BFFD20" box="[889,983,657,683]" metricMagnitude="3" metricUnit="m" metricValue="2.276" pageId="12" pageNumber="314" unit="m" value="2276.0">
|
||
<elevation id="0019B97E5F5B0C1B6145639CF0BFFD20" box="[889,983,657,683]" metricMagnitude="3" metricUnit="m" metricValue="2.276" pageId="12" pageNumber="314" unit="m" value="2276.0">2,276 m</elevation>
|
||
</quantity>
|
||
a.s.l, collected on
|
||
<date id="FF8A788D5F5B0C1B668E639CF632FD20" box="[1202,1370,656,683]" pageId="12" pageNumber="314" value="2012-05-29">
|
||
<collectingDate id="EFCE81655F5B0C1B668E639CF632FD20" box="[1202,1370,656,683]" pageId="12" pageNumber="314" value="2012-05-29">
|
||
May 29
|
||
<superScript id="7C41F3055F5B0C1B6737639DF670FD15" attach="left" box="[1291,1304,656,670]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
2012
|
||
</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26C13B9B5F5B0C1B67B5639CF38EFD44" pageId="12" pageNumber="314">P. Geniez</collectorName>
|
||
and
|
||
<collectorName id="26C13B9B5F5B0C1B632763B8F2F7FD44" box="[283,415,693,719]" pageId="12" pageNumber="314">A. Miralles</collectorName>
|
||
; photo PGe.1908, live subadult from the
|
||
<typeStatus id="548FE0EF5F5B0C1B615763B8F0F3FD44" box="[875,923,693,719]" pageId="12" pageNumber="314">type</typeStatus>
|
||
locality
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F5B0C1B62FB63D4F0D1FC64" blockId="12.[151,1437,405,1980]" pageId="12" pageNumber="314">
|
||
Moreover, we identified a number of specimens as
|
||
<taxonomicName id="4C3425CE5F5B0C1B612E63D4F0F2FD78" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[786,922,729,755]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="314" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F5B0C1B612E63D4F0F2FD78" box="[786,922,729,755]" italics="true" pageId="12" pageNumber="314">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
on the basis of morphology, distribution and ecology: BEV.4922–4923 from the Akkal Ouiad valley, north-west of Anefgou,
|
||
<geoCoordinate id="EE00388A5F5B0C1B661563F0F7E2FC9C" box="[1065,1162,765,791]" degrees="32.30" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="555" value="32.3">32.30°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B66AB63F1F786FC9C" box="[1175,1262,764,791]" degrees="5.42" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="555" value="-5.42">5.42°W</geoCoordinate>
|
||
,
|
||
<quantity id="4CCCF3A85F5B0C1B66C763F0F630FC9C" box="[1275,1368,765,791]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="314" unit="m" value="2200.0">2,200 m</quantity>
|
||
a.s.l., collected on
|
||
<date id="FF8A788D5F5B0C1B631F622CF2D5FCB0" box="[291,445,800,827]" pageId="12" pageNumber="314" value="1961-04-04">
|
||
April 4
|
||
<superScript id="7C41F3055F5B0C1B634E622DF217FCA5" attach="left" box="[370,383,800,814]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
1961
|
||
</date>
|
||
by Georges Pasteur; BEV.4924 from Agoudim, south of Tounfite,
|
||
<geoCoordinate id="EE00388A5F5B0C1B66A2622CF673FCB0" box="[1182,1307,801,827]" degrees="32.3763" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="5" value="32.3763">32.3763°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B6719622DF6F1FCB0" box="[1317,1433,800,827]" degrees="5.1797" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="5" value="-5.1797">5.1797°W</geoCoordinate>
|
||
,
|
||
<quantity id="4CCCF3A85F5B0C1B62AB6248F39AFCD4" box="[151,242,837,863]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="314" unit="m" value="2100.0">2,100 m</quantity>
|
||
a.s.l., collected on
|
||
<date id="FF8A788D5F5B0C1B63F86248F136FCD4" box="[452,606,836,863]" pageId="12" pageNumber="314" value="1961-04-04">
|
||
April 4
|
||
<superScript id="7C41F3055F5B0C1B602F6249F148FCD9" attach="left" box="[531,544,836,850]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
1961
|
||
</date>
|
||
by G. Pasteur; BEV.4927–4931 from a few km north of the Tizi n’Talremt,
|
||
<geoCoordinate id="EE00388A5F5B0C1B62AB6264F390FC08" box="[151,248,873,899]" degrees="32.61" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="555" value="32.61">32.61°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B63396265F233FC08" box="[261,347,872,899]" degrees="4.51" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="555" value="-4.51">4.51°W</geoCoordinate>
|
||
, no elevation reported, collected on
|
||
<date id="FF8A788D5F5B0C1B60C46264F0F5FC08" box="[760,925,872,899]" pageId="12" pageNumber="314" value="1962-05-28">
|
||
May 28
|
||
<superScript id="7C41F3055F5B0C1B616C6265F035FCFD" attach="left" box="[848,861,872,886]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
1962
|
||
</date>
|
||
by Jacques Bons; photo PGe.1541 from Jbel Aberdouz,
|
||
<quantity id="4CCCF3A85F5B0C1B63256280F212FC2C" box="[281,378,909,935]" metricMagnitude="3" metricUnit="m" metricValue="2.068" pageId="12" pageNumber="314" unit="m" value="2068.0">2,068 m</quantity>
|
||
a.s.l.,
|
||
<geoCoordinate id="EE00388A5F5B0C1B63F16280F130FC2C" box="[461,600,909,935]" degrees="32.18222" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="1" value="32.18222">32.18222°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B60556281F181FC2C" box="[617,745,908,935]" degrees="5.20532" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="1" value="-5.20532">5.20532°W</geoCoordinate>
|
||
, photographed on
|
||
<date id="FF8A788D5F5B0C1B61F46280F709FC2C" box="[968,1121,908,935]" pageId="12" pageNumber="314" value="2017-07-07">
|
||
July 7
|
||
<superScript id="7C41F3055F5B0C1B662D6281F776FC11" attach="left" box="[1041,1054,908,922]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
2017
|
||
</date>
|
||
by Michel-Ange Bouchet; BEV.4921, from Aït-Moualil, near Bou-Drâa de l’Oudrhès, around
|
||
<geoCoordinate id="EE00388A5F5B0C1B61B962BDF08FFC40" box="[901,999,944,971]" degrees="32.53" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="555" value="32.53">32.53°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B61CD62BDF720FC40" box="[1009,1096,944,971]" degrees="5.19" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="555" value="-5.19">5.19°W</geoCoordinate>
|
||
, at least
|
||
<quantity id="4CCCF3A85F5B0C1B669B62BCF66BFC40" box="[1191,1283,945,971]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="314" unit="m" value="1900.0">1,900 m</quantity>
|
||
a.s.l. (eastern High Atlas), collected on
|
||
<date id="FF8A788D5F5B0C1B638962D8F139FC64" box="[437,593,980,1007]" pageId="12" pageNumber="314" value="1961-04-03">
|
||
April 3
|
||
<superScript id="7C41F3055F5B0C1B603962D9F17BFC69" attach="left" box="[517,531,980,994]" fontSize="6" pageId="12" pageNumber="314">rd</superScript>
|
||
1961
|
||
</date>
|
||
by K. Klemmer and G. Pasteur.
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F5B0C1B62FB62F4F398FB4C" blockId="12.[151,1437,405,1980]" pageId="12" pageNumber="314">
|
||
In addition, two series of specimens exhibiting morphological features of
|
||
<taxonomicName id="4C3425CE5F5B0C1B662C62F4F7F0FB98" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[1040,1176,1017,1043]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="314" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F5B0C1B662C62F4F7F0FB98" box="[1040,1176,1017,1043]" italics="true" pageId="12" pageNumber="314">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
originate from outside the main distribution of the new species: we list them here but the identification is tentative only and they are represented by white diamonds in the map in
|
||
<figureCitation id="130F42C85F5B0C1B6061654CF1A6FBD0" box="[605,718,1089,1115]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="12" pageNumber="314">Figure 10</figureCitation>
|
||
: BEV.4964–4966 from Itzer (presumably the area around Itzer), approx.
|
||
<geoCoordinate id="EE00388A5F5B0C1B62C86568F23DFBF4" box="[244,341,1125,1151]" degrees="32.88" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="555" value="32.88">32.88°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B635D6569F2DFFBF4" box="[353,439,1124,1151]" degrees="5.05" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="555" value="-5.05">5.05°W</geoCoordinate>
|
||
, no elevation reported, collected on
|
||
<date id="FF8A788D5F5B0C1B616D6568F08EFBF4" box="[849,998,1124,1151]" pageId="12" pageNumber="314" value="1964-06-05">
|
||
June 5
|
||
<superScript id="7C41F3055F5B0C1B61A66569F0CFFBF9" attach="left" box="[922,935,1124,1138]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
1964
|
||
</date>
|
||
by Mr. Cauro; BEV.4920, from
|
||
<quantity id="4CCCF3A85F5B0C1B67696568F6F4FBF4" box="[1365,1436,1125,1151]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="314" unit="km" value="14.0">14 km</quantity>
|
||
after Boulemane going towards Midelt,
|
||
<geoCoordinate id="EE00388A5F5B0C1B60666585F1D3FB28" box="[602,699,1160,1187]" degrees="33.25" direction="north" orientation="latitude" pageId="12" pageNumber="314" precision="555" value="33.25">33.25°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE00388A5F5B0C1B60FB6584F076FB28" box="[711,798,1161,1187]" degrees="4.69" direction="west" orientation="longitude" pageId="12" pageNumber="314" precision="555" value="-4.69">4.69°W</geoCoordinate>
|
||
, no elevation reported, collected on
|
||
<date id="FF8A788D5F5B0C1B66896584F63EFB28" box="[1205,1366,1160,1187]" pageId="12" pageNumber="314" value="1961-04-02">
|
||
April 2
|
||
<superScript id="7C41F3055F5B0C1B673A6585F67EFB1D" attach="left" box="[1286,1302,1160,1174]" fontSize="6" pageId="12" pageNumber="314">nd</superScript>
|
||
1961
|
||
</date>
|
||
by G. Pasteur.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E0DC65F5B0C1B62FB65DCF190FAD3" pageId="12" pageNumber="314" type="etymology">
|
||
<paragraph id="8B8B5E4D5F5B0C1B62FB65DCF190FAD3" blockId="12.[151,1437,405,1980]" pageId="12" pageNumber="314">
|
||
<emphasis id="B940825F5F5B0C1B62FB65DCF226FB60" bold="true" box="[199,334,1233,1259]" pageId="12" pageNumber="314">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="4C3425CE5F5B0C1B607965DCF1C5FB60" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[581,685,1233,1259]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="314" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F5B0C1B607965DCF1C5FB60" box="[581,685,1233,1259]" italics="true" pageId="12" pageNumber="314">lacrymae</emphasis>
|
||
</taxonomicName>
|
||
”, noun in apposition in the plural form (“tears” in English), comes from a local Berber legend, according to which two young people from different tribes fell in love, but were forbidden to see each other by their families. The grief led them to cry themselves to death, their tears being at the origin of the neighbouring lakes of Tislit (hers) and Isli (his).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E0DC65F5B0C1A62FB646DF032FC23" lastPageId="13" lastPageNumber="315" pageId="12" pageNumber="314" type="diagnosis">
|
||
<paragraph id="8B8B5E4D5F5B0C1B62FB646DF61AF830" blockId="12.[151,1437,405,1980]" pageId="12" pageNumber="314">
|
||
<emphasis id="B940825F5F5B0C1B62FB646DF22AFAF0" bold="true" box="[199,322,1376,1403]" pageId="12" pageNumber="314">Diagnosis.</emphasis>
|
||
A new species of the
|
||
<taxonomicName id="4C3425CE5F5B0C1B6011646CF00BFAF0" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[557,867,1377,1403]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="314" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F5B0C1B6011646CF00BFAF0" box="[557,867,1377,1403]" italics="true" pageId="12" pageNumber="314">Acanthodactylus erythrurus</emphasis>
|
||
</taxonomicName>
|
||
species-group (small flat or carinated dorsal scales; three series of scales around the fingers; three entire supraoculars; 8–10, sometimes 12 straight longitudinal rows of ventrals; slightly pectinate toes; tail and underparts of the hind limbs red or reddish in juveniles, subadults and young adults) from the north-eastern part of the High Atlas Mountains of
|
||
<collectingCountry id="F3231EDD5F5B0C1B61BB64C0F084FA6C" box="[903,1004,1485,1511]" name="Morocco" pageId="12" pageNumber="314">Morocco</collectingCountry>
|
||
above
|
||
<quantity id="4CCCF3A85F5B0C1B660264C0F7F2FA6C" box="[1086,1178,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="314" unit="m" value="2000.0">2,000 m</quantity>
|
||
a.s.l., characterized by the combination of the following characters: (1) head scalation always of the
|
||
<emphasis id="B940825F5F5B0C1B61C764FCF73BF980" box="[1019,1107,1521,1547]" italics="true" pageId="12" pageNumber="314">
|
||
“
|
||
<taxonomicName id="4C3425CE5F5B0C1B663A64FCF720F980" authorityName="Gray" authorityYear="1845" box="[1030,1096,1521,1547]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="314" phylum="Chordata" rank="species" species="bellii">bellii</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
<typeStatus id="548FE0EF5F5B0C1B666664FCF7E2F980" box="[1114,1162,1521,1547]" pageId="12" pageNumber="314">type</typeStatus>
|
||
(subocular in large contact with the upper lip, wedged between the 4
|
||
<superScript id="7C41F3055F5B0C1B60B36719F1F4F9A9" attach="left" box="[655,668,1556,1570]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
and 5
|
||
<superScript id="7C41F3055F5B0C1B60E36719F184F9A9" attach="left" box="[735,748,1556,1570]" fontSize="6" pageId="12" pageNumber="314">th</superScript>
|
||
supralabials), one internasal plate, no scales inserted between the prefrontals; (2) a low number of scales, especially for the dorsal scales (44–55 longitudinal rows around the body, mean 50.7), the femoral pores (16–23 or 17–23 on each side, means 19.5 and 19.4) and the subdigital lamellae (19–23, mean 19.8); (3) always less than two rows of supraciliary granules on each side and a reduced number of scales around the supraoculars (15–29 on each side, mean 22.05); (4) smooth temporals; (5) smooth dorsal scales on the neck and back, sometimes slightly but distinctly carinated (tectiform) on the back; (6) stocky proportions, especially in adults, with a relatively short tail (45.6–58.5% of the total length, mean 53.9%); (7) base of the tail extremely enlarged and thick in adult males; (8) in juveniles, undertail coral red turning paler toward the base, even paler on the rear of the underside of the thighs, where it does not reach the rows of femoral pores; undertail white in adults of both sexes; (9) some marginal ventral plates yellow or yellowish in adult and subadult males, sometimes in juveniles (but not clearly so in females); (10) a series of small rounded ocelli on each flank (3–10 brightly coloured scales), usually yellowish in adults, rarely grey greenish, which tend to disappear completely in old specimens.
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F5A0C1A62FB6194F0BDFD90" blockId="13.[151,1437,153,2015]" pageId="13" pageNumber="315">
|
||
Body coloration is similar to the other members of the
|
||
<taxonomicName id="4C3425CE5F5A0C1A61006194F71EFF38" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[828,1142,153,179]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F5A0C1A61006194F71EFF38" box="[828,1142,153,179]" italics="true" pageId="13" pageNumber="315">Acanthodactylus erythrurus</emphasis>
|
||
</taxonomicName>
|
||
group. Juveniles are dark (black or blackish) to brown with three pale continuous lines on each side of the body (one on lower flanks, one at the junction between the dorsum and the flanks continuing along the tail, and one on the dorsum joining with the opposite one on the tail base) and pale mid-vertebral area starting on the nape where it is often entirely divided by a dark line and often disappearing on mid-body or before the tail; one or two series of pale ocelli are found in the dark interspaces between these stripes. Adults are medium brown with faint pale lines on the dorsum and upper flanks (corresponding with the juvenile pattern) and a better marked line on the lower flanks; irregular dark marks are present on the back between the pale lines, often leaving the area on the centre of the dorsum unmarked. The flanks typically appear darker due to the large dark areas surrounding each pale ocelli. In many specimens, the pale dorso-lateral lines continue on the outer edge of the pileus to the back of the eye and continue in a faded way along the tail. The pileus is the same colour as the back, usually with dark spots.
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F5A0C1A62FB6328F032FC23" blockId="13.[151,1437,153,2015]" pageId="13" pageNumber="315">
|
||
The combination of characters listed above separate most specimens from all other members of the
|
||
<taxonomicName id="4C3425CE5F5A0C1A671D6328F3D3FDE8" baseAuthorityName="Schinz" baseAuthorityYear="1833" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F5A0C1A671D6328F3D3FDE8" italics="true" pageId="13" pageNumber="315">A. erythrurus</emphasis>
|
||
</taxonomicName>
|
||
complex (except
|
||
<taxonomicName id="4C3425CE5F5A0C1A63BF6344F167FDE8" authority="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet, 2020" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[387,527,585,611]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="montanus" status="sp. nov.">
|
||
<emphasis id="B940825F5F5A0C1A63BF6344F167FDE8" box="[387,527,585,611]" italics="true" pageId="13" pageNumber="315">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2733F245F5A0C1A602A6344F104FDE8" box="[534,620,585,611]" pageId="13" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
|
||
which is extremely similar). Compared to
|
||
<taxonomicName id="4C3425CE5F5A0C1A66706344F7B0FDE8" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[1100,1240,585,611]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F5A0C1A66706344F7B0FDE8" box="[1100,1240,585,611]" italics="true" pageId="13" pageNumber="315">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3425CE5F5A0C1A66DA6344F605FDE8" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[1254,1389,585,611]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F5A0C1A66DA6344F605FDE8" box="[1254,1389,585,611]" italics="true" pageId="13" pageNumber="315">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
differs in (1) the parietals and sometimes supraoculars of juveniles with a more continuous and contrasting dark band along their outer edge, creating a more contrasted striped pattern with the pale dorsolateral band continuing through the supratemporals along the sides of the pileus (in
|
||
<taxonomicName id="4C3425CE5F5A0C1A60DE63B8F018FD44" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[738,880,693,719]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F5A0C1A60DE63B8F018FD44" box="[738,880,693,719]" italics="true" pageId="13" pageNumber="315">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
, parietals have a less distinctly striped pattern of dark marks and light reticulation, see
|
||
<figureCitation id="130F42C85F5A0C1A607E63D4F1EEFD78" box="[578,646,729,755]" captionStart="FIGURE 9" captionStartId="18.[151,250,1859,1884]" captionTargetBox="[198,1394,184,1829]" captionTargetId="figure@18.[185,1402,174,1840]" captionTargetPageId="18" captionText="FIGURE 9. Phenotypic variation in juveniles. Pileus of juveniles in A lacrymae sp. nov. (A, BEV.11981 & B, BEV. 11990, both from Tislit) and A. montanus sp. nov. (C, JCB.A546, Jbel Siroua & D, BEV.11792, Tizi n’Tichka). Caudal and cloacal colour patterns in juveniles of A. lacrymae sp. nov. (E, BEV.11986 & F, BEV.11990, both from Tislit), A. montanus sp. nov. (G, BEV.11798, Tizi n’Tichka & H, JCB.A546, Jbel Siroua) and A. erythrurus (I, BEV.11926, Dayet Ifrah & J, BEV.11595, Marrakech). Photo credits: A, B, E, F, I (PG), D, G, J (PAC), C, H (JCB)." figureDoi="http://doi.org/10.5281/zenodo.3694846" httpUri="https://zenodo.org/record/3694846/files/figure.png" pageId="13" pageNumber="315">Fig. 9</figureCitation>
|
||
); (2) the colour of the base of the undertail is paler and less extensive around the cloacal slit in juveniles (more intense orange red, more similar to the rest of the tail, and more extensive in juveniles
|
||
<taxonomicName id="4C3425CE5F5A0C1A6318622CF2DAFCB0" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[292,434,801,827]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F5A0C1A6318622CF2DAFCB0" box="[292,434,801,827]" italics="true" pageId="13" pageNumber="315">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="130F42C85F5A0C1A63FE622CF16FFCB0" box="[450,519,801,827]" captionStart="FIGURE 9" captionStartId="18.[151,250,1859,1884]" captionTargetBox="[198,1394,184,1829]" captionTargetId="figure@18.[185,1402,174,1840]" captionTargetPageId="18" captionText="FIGURE 9. Phenotypic variation in juveniles. Pileus of juveniles in A lacrymae sp. nov. (A, BEV.11981 & B, BEV. 11990, both from Tislit) and A. montanus sp. nov. (C, JCB.A546, Jbel Siroua & D, BEV.11792, Tizi n’Tichka). Caudal and cloacal colour patterns in juveniles of A. lacrymae sp. nov. (E, BEV.11986 & F, BEV.11990, both from Tislit), A. montanus sp. nov. (G, BEV.11798, Tizi n’Tichka & H, JCB.A546, Jbel Siroua) and A. erythrurus (I, BEV.11926, Dayet Ifrah & J, BEV.11595, Marrakech). Photo credits: A, B, E, F, I (PG), D, G, J (PAC), C, H (JCB)." figureDoi="http://doi.org/10.5281/zenodo.3694846" httpUri="https://zenodo.org/record/3694846/files/figure.png" pageId="13" pageNumber="315">Fig. 9</figureCitation>
|
||
); (3) the supralabials are unmarked, pale creamy even in juveniles (labials have diffuse dark stripes which persist in adults of
|
||
<taxonomicName id="4C3425CE5F5A0C1A60A16248F042FCD4" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[669,810,837,863]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F5A0C1A60A16248F042FCD4" box="[669,810,837,863]" italics="true" pageId="13" pageNumber="315">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
); (4) a sharper and better delimited border between the pale lower surface of the tail and a dark band running along the side of the tail, with a complex pattern in males (in
|
||
<taxonomicName id="4C3425CE5F5A0C1A62AB6280F24BFC2C" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[151,291,909,935]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="montanus">
|
||
<emphasis id="B940825F5F5A0C1A62AB6280F24BFC2C" box="[151,291,909,935]" italics="true" pageId="13" pageNumber="315">A. montanus</emphasis>
|
||
</taxonomicName>
|
||
the pattern is less contrasted, especially in males).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E0DC65F5A0C1A62FB62BDF6F4FA6C" pageId="13" pageNumber="315" type="description">
|
||
<paragraph id="8B8B5E4D5F5A0C1A62FB62BDF6F4FA6C" blockId="13.[151,1437,153,2015]" pageId="13" pageNumber="315">
|
||
<emphasis id="B940825F5F5A0C1A62FB62BDF17EFC40" bold="true" box="[199,534,944,971]" pageId="13" pageNumber="315">
|
||
Description of the
|
||
<typeStatus id="548FE0EF5F5A0C1A639462BCF179FC40" box="[424,529,945,971]" pageId="13" pageNumber="315" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
An adult male (
|
||
<figureCitation id="130F42C85F5A0C1A60F362BCF04FFC40" box="[719,807,945,971]" captionStart="FIGURE 8" captionStartId="17.[151,250,1935,1960]" captionTargetBox="[183,1406,522,1913]" captionTargetId="figure@17.[179,1409,517,1916]" captionTargetPageId="17" captionText="FIGURE 8. Holotypes of Acanthodactylus montanus sp. nov. (MNHN-RA-2018.0026, Tizi n’Tichka) and (B) Acanthodactylus lacrymae sp. nov. (MNHN-RA-2018.0027, Tislit) with a picture of the habitat of this species (Isli lake). Photo credits:A (PAC), B (PG)." figureDoi="http://doi.org/10.5281/zenodo.3694844" httpUri="https://zenodo.org/record/3694844/files/figure.png" pageId="13" pageNumber="315">Fig. 8B</figureCitation>
|
||
) measuring 54.0 mm of snout-vent length,
|
||
<quantity id="4CCCF3A85F5A0C1A672B62BCF615FC40" box="[1303,1405,945,971]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="13" pageNumber="315" unit="mm" value="13.8">13.8 mm</quantity>
|
||
of pileus length,
|
||
<quantity id="4CCCF3A85F5A0C1A630B62D8F2E7FC64" box="[311,399,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="13" pageNumber="315" unit="mm" value="7.9">7.9 mm</quantity>
|
||
of pileus width,
|
||
<quantity id="4CCCF3A85F5A0C1A606C62D9F1CFFC64" box="[592,679,980,1007]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="315" unit="mm" value="7.5">7.5 mm</quantity>
|
||
of head height,
|
||
<quantity id="4CCCF3A85F5A0C1A615D62D9F0AEFC64" box="[865,966,980,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="13" pageNumber="315" unit="mm" value="16.5">16.5 mm</quantity>
|
||
of forelimb length,
|
||
<quantity id="4CCCF3A85F5A0C1A669762D9F678FC64" box="[1195,1296,980,1007]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" pageId="13" pageNumber="315" unit="mm" value="25.6">25.6 mm</quantity>
|
||
of hindlimb length, with a relatively short non-regenerated tail (
|
||
<quantity id="4CCCF3A85F5A0C1A60E262F4F059FB98" box="[734,817,1017,1043]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="13" pageNumber="315" unit="mm" value="63.0">63 mm</quantity>
|
||
, representing only 54% of the total length), presenting the following scalation features: 54 longitudinal rows of smooth dorsal scales at mid-body, 23 gular scales counted along a line from the contact between the fourth pair of maxillary scales to the collar, 28 transversal rows of ventral plates from the collar to the anal plates, 21 femoral pores on each side, 20 subdigital lamellae beneath the fourth toe, 1 row of supraciliary granules on each side, with 29 (left) and 28 (right) granules, respectively, one entire internasal, no scale inserted between the two prefrontals, two supralabials in contact with the subocular, the latter in large contact with the upper lip on each side. Base of the tail extremely thick. Coloration in life: general ground colour light rufous brown with 6 longitudinal whitish to light grey stripes; the vertebral area between the two central light stripes is nearly uniform with only some vestigial black marks; a series of black transversal marks on the sides of the back; a series of 10 small light greenish ocelli, each surrounded by a large black spot, along the flanks; pileus of the same colour as the dorsum with some black and light marks corresponding approximately to the extension of the dorsal stripes; underparts pure white, a few marginal ventral plates pale yellow or with a black mark; colour delimitation between the side of the body and the underparts very strong and contrasted; a black longitudinal stripe runs along the side of the tail, the top of this stripe slightly wavy on the anterior part and straight on the rest of the side of the tail.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E0DC65F5A0C1962FB64FDF737FF77" lastPageId="14" lastPageNumber="316" pageId="13" pageNumber="315" type="distribution">
|
||
<paragraph id="8B8B5E4D5F5A0C1A62FB64FDF1B5F88C" blockId="13.[151,1437,153,2015]" pageId="13" pageNumber="315">
|
||
<emphasis id="B940825F5F5A0C1A62FB64FDF209F980" bold="true" box="[199,353,1520,1547]" pageId="13" pageNumber="315">Distribution.</emphasis>
|
||
Until now, known only with certainty from a handful of localities in a narrow stretch of the eastern High Atlas of
|
||
<collectingCountry id="F3231EDD5F5A0C1A63596718F2A6F9A4" box="[357,462,1557,1583]" name="Morocco" pageId="13" pageNumber="315">Morocco</collectingCountry>
|
||
; at high elevations (above
|
||
<quantity id="4CCCF3A85F5A0C1A613C6718F035F9A4" box="[768,861,1557,1583]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="315" unit="m" value="2000.0">2,000 m</quantity>
|
||
a.s.l. for all localities whose elevation is precisely known) between Tislit Lake to the west and the Tizi n’Talremt Lake to the east (
|
||
<figureCitation id="130F42C85F5A0C1A66276734F706F9D8" box="[1051,1134,1593,1619]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="13" pageNumber="315">Fig. 10</figureCitation>
|
||
). Based on the distribution of animals with the
|
||
<taxonomicName id="4C3425CE5F5A0C1A63456750F2DBF9FC" authorityName="Gray" authorityYear="1845" box="[377,435,1629,1655]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="bellii">
|
||
<emphasis id="B940825F5F5A0C1A63456750F2DBF9FC" box="[377,435,1629,1655]" italics="true" pageId="13" pageNumber="315">bellii</emphasis>
|
||
</taxonomicName>
|
||
-like subocular conformation in the eastern High Atlas, the potential distribution extends south to the upper Todra Valley (continuous white line in
|
||
<figureCitation id="130F42C85F5A0C1A6127678CF005F910" box="[795,877,1665,1691]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="13" pageNumber="315">Fig. 10</figureCitation>
|
||
). If the identity of the specimens originating from the surroundings of Itzer and the area south of Boulemane (white diamonds in
|
||
<figureCitation id="130F42C85F5A0C1A663D67A8F738F934" box="[1025,1104,1701,1727]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="13" pageNumber="315">Fig. 10</figureCitation>
|
||
) is confirmed by further data, the distribution would extend to the southern edge of the Middle Atlas, potentially in sympatry with the local populations of the IM clade (doted white line in
|
||
<figureCitation id="130F42C85F5A0C1A604167E0F1B8F88C" box="[637,720,1773,1799]" captionStart="FIGURE 10" captionStartId="19.[151,250,1622,1647]" captionTargetBox="[206,1380,802,1609]" captionTargetId="figure@19.[200,1387,798,1614]" captionTargetPageId="19" captionText="FIGURE 10. Distribution of Acanthodactylus montanus (squares) and Acanthodactylus lacrymae (diamonds) based on specimens examined in this study. Coloured dots represent specimens of the three main phenotypes (none is monophyletic in the mtDNA or nDNA trees) in Morocco based on Bons and Geniez 1995. Specimens with the “bellii” phenotype in the High Atlas are almost certainly montanus and lacrymae. White symbols represent specimens tentatively identified as A. montanus (square) or A. lacrymae (diamond, see text). The continuous white lines represent the currently known distribution of the new species while dotted white lines represent possible extensions of the distribution if tentative identifications are correct. The dotted dark blue line represents the southernmost extension of confirmed IM clade specimens based on our field sampling." figureDoi="http://doi.org/10.5281/zenodo.3694848" httpUri="https://zenodo.org/record/3694848/files/figure.png" pageId="13" pageNumber="315">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B5E4D5F5A0C1962FB661CF737FF77" blockId="13.[151,1437,153,2015]" lastBlockId="14.[151,1437,152,252]" lastPageId="14" lastPageNumber="316" pageId="13" pageNumber="315">
|
||
<emphasis id="B940825F5F5A0C1A62FB661CF2EEF8A0" bold="true" box="[199,390,1808,1835]" pageId="13" pageNumber="315">Natural history.</emphasis>
|
||
<taxonomicName id="4C3425CE5F5A0C1A63B2661CF1D1F8A0" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[398,697,1809,1835]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F5A0C1A63B2661CF1D1F8A0" box="[398,697,1809,1835]" italics="true" pageId="13" pageNumber="315">Acanthodactylus lacrymae</emphasis>
|
||
</taxonomicName>
|
||
lives on high plateaus above
|
||
<quantity id="4CCCF3A85F5A0C1A6630661CF702F8A0" box="[1036,1130,1809,1835]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="315" unit="m" value="2000.0">2,000 m</quantity>
|
||
a.s.l. In both of the visited localities the habitat was similar: open, earthy or gravel ground with short grazed vegetation interspaced with barren patches, numerous stones and low shrubs/patches of taller plants (
|
||
<taxonomicName id="4C3425CE5F5A0C1A618B6654F7C1F8F8" box="[951,1193,1881,1907]" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" pageId="13" pageNumber="315" phylum="Tracheophyta" rank="species" species="herba-alba">
|
||
<emphasis id="B940825F5F5A0C1A618B6654F7C1F8F8" box="[951,1193,1881,1907]" italics="true" pageId="13" pageNumber="315">Artemisia herba-alba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3425CE5F5A0C1A66DF6654F385F81C" class="Magnoliopsida" family="Brassicaceae" genus="Hormathophylla" kingdom="Plantae" order="Brassicales" pageId="13" pageNumber="315" phylum="Tracheophyta" rank="species" species="spinosa">
|
||
<emphasis id="B940825F5F5A0C1A66DF6654F385F81C" italics="true" pageId="13" pageNumber="315">Hormathophylla spinosa</emphasis>
|
||
</taxonomicName>
|
||
) that the lizards use as shelters. With its stocky proportions,
|
||
<taxonomicName id="4C3425CE5F5A0C1A619D6670F741F81C" authorityName="Miralles & Geniez & Beddek & Aranda & Brito & Leblois & Crochet" authorityYear="2020" box="[929,1065,1917,1943]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="lacrymae">
|
||
<emphasis id="B940825F5F5A0C1A619D6670F741F81C" box="[929,1065,1917,1943]" italics="true" pageId="13" pageNumber="315">A. lacrymae</emphasis>
|
||
</taxonomicName>
|
||
runs relatively slowly compared with other members of the
|
||
<taxonomicName id="4C3425CE5F5A0C1A63FF66ACF13FF830" baseAuthorityName="Schinz" baseAuthorityYear="1833" box="[451,599,1953,1979]" class="Reptilia" family="Lacertidae" genus="Acanthodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="315" phylum="Chordata" rank="species" species="erythrurus">
|
||
<emphasis id="B940825F5F5A0C1A63FF66ACF13FF830" box="[451,599,1953,1979]" italics="true" pageId="13" pageNumber="315">A. erythrurus</emphasis>
|
||
</taxonomicName>
|
||
complex, which makes them easier to catch. Both visited populations were dense, with numerous individuals in activity (respectively 17 and
|
||
<specimenCount id="9D3295C45F5A0C1A61A966C8F747F854" box="[917,1071,1989,2015]" pageId="13" pageNumber="315" type="generic">11 specimens</specimenCount>
|
||
caught and others seen but not captured in around
|
||
<quantity id="4CCCF3A85F590C19634C6194F2B8FF38" box="[368,464,152,179]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="316" unit="km" value="0.01">0.01 km</quantity>
|
||
²). These high densities may be explained by the well-preserved habitats typical of high altitudes areas in the region and the absence of direct competitors (the only other observed Lacertid was the North- African wall lizard
|
||
<emphasis id="B940825F5F590C19634E61ECF12DFF70" box="[370,581,225,251]" italics="true" pageId="14" pageNumber="316">
|
||
<taxonomicName id="4C3425CE5F590C19634E61ECF12AFF70" baseAuthorityName="Boulenger" baseAuthorityYear="1905" box="[370,578,225,251]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="316" phylum="Chordata" rank="species" species="vaucheri">Podarcis vaucheri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which prefers rocky areas or dense vegetation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |