263 lines
20 KiB
XML
263 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1071.66417" ID-GBIF-Dataset="6ced9566-cba1-4206-97aa-95939ce3444c" ID-PMC="PMC8616889" ID-Pensoft-Pub="1313-2970-1071-175" ID-Pensoft-UUID="B033EEBDF60E51B484556CA9E0F5E118" ID-PubMed="34887699" ID-ZooBank="0B9246161AC149B8BE5F531286EACE63" ModsDocID="1313-2970-1071-175" checkinTime="1637285687178" checkinUser="pensoft" docAuthor="Inaebnit, Thomas, Jochum, Adrienne, Slapnik, Raijko & Neubert, Eike" docDate="2021" docId="04D6F12E943D5710A5862DFCA1508456" docLanguage="en" docName="ZooKeys 1071: 175-193" docOrigin="ZooKeys 1071" docPubDate="2021-11-18" docSource="http://dx.doi.org/10.3897/zookeys.1071.66417" docTitle="Zospeum simplex Inäbnit, Jochum & Neubert, sp. nov." docType="treatment" docVersion="4" id="B033EEBDF60E51B484556CA9E0F5E118" lastPageNumber="175" masterDocId="B033EEBDF60E51B484556CA9E0F5E118" masterDocTitle="New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae)" masterLastPageNumber="193" masterPageNumber="175" pageNumber="175" updateTime="1668151133265" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inaebnit, Thomas</mods:namePart>
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<mods:affiliation>Institute for Biochemistry & Biology, University of Potsdam, Karl-Liebknecht-Strasse 24 - 25, House 26, 14476, Potsdam, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">inaebnit.thomas@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of the Burgergemeinde Bern, Bernastrasse 15, 3005, Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, 3012, Bern, Switzerland & Senckenberg Research Institute and Natural History Museum, Senckenberganlage 25, 60325 Frankfurt / M, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Slapnik, Raijko</mods:namePart>
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<mods:affiliation>Drnovskova pot 2, Mekinje, SI - 1240 Kamnik, Slovenia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neubert, Eike</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0277-2894</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of the Burgergemeinde Bern, Bernastrasse 15, 3005, Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, 3012, Bern, Switzerland</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1071</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>175</mods:start>
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<mods:end>193</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1071.66417</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1071.66417</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1071-175</mods:identifier>
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<mods:identifier type="ZooBank">0B9246161AC149B8BE5F531286EACE63</mods:identifier>
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<mods:identifier type="Pensoft-UUID">B033EEBDF60E51B484556CA9E0F5E118</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="190239463" LSID="urn:lsid:zoobank.org:act:0B924616-1AC1-49B8-BE5F-531286EACE63" httpUri="http://treatment.plazi.org/id/04D6F12E943D5710A5862DFCA1508456" lastPageNumber="175" pageId="0" pageNumber="175">
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<subSubSection pageId="0" pageNumber="175" type="nomenclature">
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<paragraph pageId="0" pageNumber="175">
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<taxonomicName LSID="http://zoobank.org/0B924616-1AC1-49B8-BE5F-531286EACE63" authority="Inaebnit, Jochum & Neubert, sp. nov." authorityName="Inäbnit, Jochum & Neubert, sp. nov." class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum simplex" order="Ellobiida" pageId="0" pageNumber="175" phylum="Mollusca" rank="species" species="simplex">
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Zospeum simplex
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<normalizedToken originalValue="Inäbnit">Inaebnit</normalizedToken>
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, Jochum & Neubert, sp. nov.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="description">
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<paragraph pageId="0" pageNumber="175">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map showing the distribution of the Zospeum pretneri group and the Zospeum alpestre group (except Z. isselianum). Austrian specimens from Kruckenhauser et al. (2019) are labelled as " Z. cf. amoenum "." figureDoi="10.3897/zookeys.1071.66417.figure1" httpUri="https://binary.pensoft.net/fig/612371" pageId="0" pageNumber="175">Figures 1</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Specimens sequenced in this study. Zospeum troglobalcanicum: NMBE 568052 & 568053 (both from Spilja Jezero); Zospeum simplex sp. nov.: NMBE 568054 (Spilja Dahna), NMBE 568055 - 568057 (Jama u kamenolomu), NMBE 568059 (Vranjaca), NMBE 568060 (Holotype, Jama Dobravljevac), NMBE 568061 - 568063 (Paratypes, Jama Dobravljevac)" figureDoi="10.3897/zookeys.1071.66417.figure3" httpUri="https://binary.pensoft.net/fig/612373" pageId="0" pageNumber="175">, 3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="type specimens">
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<paragraph pageId="0" pageNumber="175">Type specimens.</paragraph>
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<paragraph pageId="0" pageNumber="175">
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<emphasis bold="true" pageId="0" pageNumber="175">
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<emphasis italics="true" pageId="0" pageNumber="175">Holotype</emphasis>
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:
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</emphasis>
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NMBE 568060, Jama Dobravljevac, 25.08.2019, leg. R. Slapnik & J.
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<normalizedToken originalValue="Valentinčič">Valentincic</normalizedToken>
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;
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<emphasis bold="true" italics="true" pageId="0" pageNumber="175">Paratypes</emphasis>
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: NMBE 568061-568063; SMF 349425, 4 shells; RSC 3760, 6 shells; Jama Dobravljevac, 25.08.2019, leg. R. Slapnik & J.
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<normalizedToken originalValue="Valentinčič">Valentincic</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="materials_examined">
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<paragraph pageId="0" pageNumber="175">Specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="175">
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NMBE 568054,
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<normalizedToken originalValue="Špilja">Spilja</normalizedToken>
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Dahna,
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<date value="2009-09-03">03.09.2009</date>
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, leg. A. Schoenhoffer; NMBE 568055-568058, Jama u kamenolomu,
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<date value="2019-08-24">24.08.2019</date>
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, leg. R. Slapnik & J.
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<normalizedToken originalValue="Valentinčič">Valentincic</normalizedToken>
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; NMBE 568059,
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<normalizedToken originalValue="Vranjača">Vranjaca</normalizedToken>
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,
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<date value="2019-08-24">24.08.2019</date>
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, leg. R. Slapnik & J.
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<normalizedToken originalValue="Valentinčič">Valentincic</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="diagnosis">
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<paragraph pageId="0" pageNumber="175">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="175">Shell usually ca. 1.3 mm in height, transparent, conical, peristome thickened, roundish, with a differentiated parietal shield, lamellae not present.</paragraph>
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<paragraph pageId="0" pageNumber="175">
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Measurements (n = 9): Shell height: 1.26-1.42 mm (mean: 1.378
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.047 mm); shell width: 0.93-1.04 mm (mean: 0.976
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.035 mm); aperture height: 0.54-0.67 mm (mean: 0.6
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.037); aperture width: 0.54-0.65 mm (mean: 0.601
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.033 mm); number of whorls: 5-5.5.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="description">
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<paragraph pageId="0" pageNumber="175">Description.</paragraph>
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<paragraph pageId="0" pageNumber="175">Shell conical, translucent when fresh; suture deep; aperture somewhat roundish to reniform; parietal shield clearly differentiated from the rest of the lip, straight and thin; no lamellae present.</paragraph>
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<paragraph pageId="0" pageNumber="175">
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Differing from
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<taxonomicName lsidName="Z. pretneri" pageId="0" pageNumber="175" rank="species" species="pretneri">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. pretneri</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="Z. tholussum" pageId="0" pageNumber="175" rank="species" species="tholussum">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. tholussum</emphasis>
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</taxonomicName>
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by its broader shell and the differentiated parietal shield; differs from
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<taxonomicName lsidName="Z. manitaense" pageId="0" pageNumber="175" rank="species" species="manitaense">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. manitaense</emphasis>
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</taxonomicName>
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by the absence of a visible parietalis in the aperture; barely differs from
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<taxonomicName lsidName="Z. troglobalcanicum" pageId="0" pageNumber="175" rank="species" species="troglobalcanicum">Z. aff. troglobalcanicum</taxonomicName>
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morphologically, on average with reduced shell broadness and a slightly deeper suture (see Remarks).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="distribution">
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<paragraph pageId="0" pageNumber="175">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="175">
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Known from four caves (Jama Dobravljevac,
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<normalizedToken originalValue="Špilja">Spilja</normalizedToken>
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Dahna, Jama u kamenolomu,
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<normalizedToken originalValue="Vranjača">Vranjaca</normalizedToken>
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) in the municipality of Tomislavgrad in Bosnia-Herzegovina.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="etymology">
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<paragraph pageId="0" pageNumber="175">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="175">
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Named
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<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum simplex" order="Basommatophora" pageId="0" pageNumber="175" phylum="Mollusca" rank="species" species="simplex">
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<emphasis italics="true" pageId="0" pageNumber="175">Zospeum simplex</emphasis>
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</taxonomicName>
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(= simple, unsophisticated) due to the lack of any form of shell sculpture or lamellae.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="175" type="remarks">
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<paragraph pageId="0" pageNumber="175">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="175">
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Difficult to separate from
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<taxonomicName lsidName="Z. troglobalcanicum" pageId="0" pageNumber="175" rank="species" species="troglobalcanicum">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. troglobalcanicum</emphasis>
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</taxonomicName>
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without genetic data (which is not uncommon in
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<taxonomicName authorityName="Bourguignat" authorityYear="1856" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum" order="Basommatophora" pageId="0" pageNumber="175" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="175">Zospeum</emphasis>
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</taxonomicName>
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; see
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<bibRefCitation DOI="https://doi.org/10.1007/s13127-019-00400-8" author="Inaebnit, T" journalOrPublisher="Organisms Diversity & Evolution" pageId="0" pageNumber="175" pagination="135 - 177" refId="B16" refString="Inaebnit, T, Jochum, A, Kampschulte, M, Martels, G, Ruthensteiner, B, Slapnik, R, Nesselhauf, C, Neubert, E, 2019. An integrative taxonomic study reveals carychiid microsnails of the troglobitic genus Zospeum in the Eastern and Dinaric Alps (Gastropoda, Ellobioidea, Carychiinae). Organisms Diversity & Evolution 19: 135 - 177, DOI: https://doi.org/10.1007/s13127-019-00400-8" title="An integrative taxonomic study reveals carychiid microsnails of the troglobitic genus Zospeum in the Eastern and Dinaric Alps (Gastropoda, Ellobioidea, Carychiinae)." url="https://doi.org/10.1007/s13127-019-00400-8" volume="19" year="2019">
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<normalizedToken originalValue="Inäbnit">Inaebnit</normalizedToken>
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et al. 2019
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</bibRefCitation>
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). Both species have a nondescript shell without prominent shell sculpture or lamellae within the aperture.
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<bibRefCitation author="Absolon, K" journalOrPublisher="Zlata Praha" pageId="0" pageNumber="175" pagination="574 - 576" refId="B1" refString="Absolon, K, 1916. Z vyzkumnych cest po krasech Balkanu. O balkanske temnostni zvirene. Zlata Praha 33: 574 - 576" title="Z vyzkumnych cest po krasech Balkanu. O balkanske temnostni zvirene." volume="33" year="1916">
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<normalizedToken originalValue="Absolon’s">Absolon's</normalizedToken>
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(1916)
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</bibRefCitation>
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description of
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<taxonomicName lsidName="Z. troglobalcanicum" pageId="0" pageNumber="175" rank="species" species="troglobalcanicum">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. troglobalcanicum</emphasis>
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</taxonomicName>
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consisted out of a photograph depicting multiple specimens haphazardly clustered together in various positions and a legend that established the name and type locality. The lack of a written characterisation of the species in the original description and the fact that the specimens in the photograph
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<normalizedToken originalValue="weren’t">weren't</normalizedToken>
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depicted in any standardised position makes a characterisation of the species fairly challenging (putative syntype specimen, collected by K. Absolon from the type locality, was only located very recently by AJ in Vienna (NHMW Mol.Coll.Edlauer 32.749) and
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<normalizedToken originalValue="couldn’t">couldn't</normalizedToken>
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be studied yet). From the photograph in
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<bibRefCitation author="Absolon, K" journalOrPublisher="Zlata Praha" pageId="0" pageNumber="175" pagination="574 - 576" refId="B1" refString="Absolon, K, 1916. Z vyzkumnych cest po krasech Balkanu. O balkanske temnostni zvirene. Zlata Praha 33: 574 - 576" title="Z vyzkumnych cest po krasech Balkanu. O balkanske temnostni zvirene." volume="33" year="1916">Absolon (1916)</bibRefCitation>
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, the species can be characterised as similar to
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<taxonomicName lsidName="Z. manitaense" pageId="0" pageNumber="175" rank="species" species="manitaense">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. manitaense</emphasis>
|
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</taxonomicName>
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in shell shape, without any visible lamella in the aperture and with a comparatively large parietal shield. The larger parietal shield might serve as a distinguishing character between
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<taxonomicName lsidName="Z. simplex" pageId="0" pageNumber="175" rank="species" species="simplex">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. simplex</emphasis>
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||
</taxonomicName>
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and
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<taxonomicName lsidName="Z. troglobalcanicum" pageId="0" pageNumber="175" rank="species" species="troglobalcanicum">
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<emphasis italics="true" pageId="0" pageNumber="175">Z. troglobalcanicum</emphasis>
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</taxonomicName>
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, though the illustration of a topotypic specimen in
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<bibRefCitation author="Bole, J" journalOrPublisher="Razred za naravoslovne vede" pageId="0" pageNumber="175" pagination="249 - 291" refId="B3" refString="Bole, J, 1974. Rod Zospeum Bourguignat 1856 (Gastropoda, Ellobiidae) v Jugoslaviji. Die Gattung Zospeum Bourguignat 1886 (Gastropoda, Ellobiidae) in Jugoslawien. Razprave - Slovenska akademija znanosti in umetnosti. Razred za naravoslovne vede 17: 249 - 291" title="Rod Zospeum Bourguignat 1856 (Gastropoda, Ellobiidae) v Jugoslaviji. Die Gattung Zospeum Bourguignat 1886 (Gastropoda, Ellobiidae) in Jugoslawien. Razprave - Slovenska akademija znanosti in umetnosti." volume="17" year="1974">Bole (1974</bibRefCitation>
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; fig. 3h) might indicate that this character is variable within the population. The two specimens we preliminarily assigned to
|
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<taxonomicName lsidName="Z. troglobalcanicum" pageId="0" pageNumber="175" rank="species" species="troglobalcanicum">
|
||
<emphasis italics="true" pageId="0" pageNumber="175">Z. troglobalcanicum</emphasis>
|
||
</taxonomicName>
|
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(Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Specimens sequenced in this study. Zospeum troglobalcanicum: NMBE 568052 & 568053 (both from Spilja Jezero); Zospeum simplex sp. nov.: NMBE 568054 (Spilja Dahna), NMBE 568055 - 568057 (Jama u kamenolomu), NMBE 568059 (Vranjaca), NMBE 568060 (Holotype, Jama Dobravljevac), NMBE 568061 - 568063 (Paratypes, Jama Dobravljevac)" figureDoi="10.3897/zookeys.1071.66417.figure3" httpUri="https://binary.pensoft.net/fig/612373" pageId="0" pageNumber="175">3</figureCitation>
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, NMBE 568052;
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<bibRefCitation DOI="https://doi.org/10.1007/s13127-019-00400-8" author="Inaebnit, T" journalOrPublisher="Organisms Diversity & Evolution" pageId="0" pageNumber="175" pagination="135 - 177" refId="B16" refString="Inaebnit, T, Jochum, A, Kampschulte, M, Martels, G, Ruthensteiner, B, Slapnik, R, Nesselhauf, C, Neubert, E, 2019. An integrative taxonomic study reveals carychiid microsnails of the troglobitic genus Zospeum in the Eastern and Dinaric Alps (Gastropoda, Ellobioidea, Carychiinae). Organisms Diversity & Evolution 19: 135 - 177, DOI: https://doi.org/10.1007/s13127-019-00400-8" title="An integrative taxonomic study reveals carychiid microsnails of the troglobitic genus Zospeum in the Eastern and Dinaric Alps (Gastropoda, Ellobioidea, Carychiinae)." url="https://doi.org/10.1007/s13127-019-00400-8" volume="19" year="2019">
|
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<normalizedToken originalValue="Inäbnit">Inaebnit</normalizedToken>
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||
et al. 2019
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||
</bibRefCitation>
|
||
: fig. 7u) only have a small parietal shield. As of now, the shell height:shell width ratio seems to be the most effective way of separating the two specimens from
|
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<taxonomicName lsidName="Z. simplex" pageId="0" pageNumber="175" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="175">Z. simplex</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="Z. simplex" pageId="0" pageNumber="175" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="175">Z. simplex</emphasis>
|
||
</taxonomicName>
|
||
: generally higher than 1.3 (one exception);
|
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<taxonomicName lsidName="Z. troglobalcanicum" pageId="0" pageNumber="175" rank="species" species="troglobalcanicum">Z. aff. troglobalcanicum</taxonomicName>
|
||
: below 1.3), but that might just be due to the low sample sizes. Investigation of the inner aspects of the shells will be presented in a later work.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1071.66417.figure3" httpUri="https://binary.pensoft.net/fig/612373" pageId="0" pageNumber="175" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="175">
|
||
<emphasis bold="true" pageId="0" pageNumber="175">Figure 3.</emphasis>
|
||
Specimens sequenced in this study.
|
||
<taxonomicName authorityName="Absolon" authorityYear="1916" class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum troglobalcanicum" order="Basommatophora" pageId="0" pageNumber="175" phylum="Mollusca" rank="species" species="troglobalcanicum">
|
||
<emphasis italics="true" pageId="0" pageNumber="175">Zospeum troglobalcanicum</emphasis>
|
||
</taxonomicName>
|
||
: NMBE 568052 & 568053 (both from
|
||
<normalizedToken originalValue="Špilja">Spilja</normalizedToken>
|
||
Jezero);
|
||
<taxonomicName class="Gastropoda" family="Ellobiidae" genus="Zospeum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zospeum simplex" order="Basommatophora" pageId="0" pageNumber="175" phylum="Mollusca" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="175">Zospeum simplex</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.: NMBE 568054 (
|
||
<normalizedToken originalValue="Špilja">Spilja</normalizedToken>
|
||
Dahna), NMBE 568055-568057 (Jama u kamenolomu), NMBE 568059 (
|
||
<normalizedToken originalValue="Vranjača">Vranjaca</normalizedToken>
|
||
), NMBE 568060 (Holotype, Jama Dobravljevac), NMBE 568061-568063 (Paratypes, Jama Dobravljevac)
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |