treatments-xml/data/03/D0/87/03D087DDFFF9FF999DFBDD31FD94E95B.xml

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<mods:title id="CCA35560799266BE24403C9BF94C2DC7">Leptalpheus pereirai sp. nov., a new alpheid shrimp from Panama and Venezuela (Decapoda: Caridea)</mods:title>
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<mods:namePart id="6EC041C7CFEF8908CCFA50D6815B25D4">Caripe, Jonathan Vera</mods:namePart>
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<treatment id="03D087DDFFF9FF999DFBDD31FD94E95B" ID-DOI="http://doi.org/10.5281/zenodo.3510458" ID-GBIF-Taxon="121061691" ID-Zenodo-Dep="3510458" LSID="urn:lsid:plazi:treatment:03D087DDFFF9FF999DFBDD31FD94E95B" httpUri="http://treatment.plazi.org/id/03D087DDFFF9FF999DFBDD31FD94E95B" lastPageId="6" lastPageNumber="190" pageId="1" pageNumber="186">
<subSubSection id="C3636540FFF9FF9E9DFBDD31FD60E9AF" pageId="1" pageNumber="186" type="nomenclature">
<paragraph id="8BC636CBFFF9FF9E9DFBDD31FE76E9C5" blockId="1.[151,501,359,419]" box="[151,501,359,386]" pageId="1" pageNumber="186">
<heading id="D08E81A7FFF9FF9E9DFBDD31FE76E9C5" bold="true" box="[151,501,359,386]" fontSize="11" level="1" pageId="1" pageNumber="186" reason="1">
<emphasis id="B90DEAD9FFF9FF9E9DFBDD31FE76E9C5" bold="true" box="[151,501,359,386]" pageId="1" pageNumber="186">
<taxonomicName id="4C794D48FFF9FF9E9DFBDD31FE13E9C6" ID-CoL="85TJS" box="[151,400,359,385]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFF9FF9E9DFBDD31FE13E9C6" bold="true" box="[151,400,359,385]" italics="true" pageId="1" pageNumber="186">Leptalpheus pereirai</emphasis>
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<taxonomicNameLabel id="A23E57A2FFF9FF9E9CF4DD3EFE76E9C5" box="[408,501,360,386]" pageId="1" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC636CBFFF9FF9E9DFBDDDCFE91E9E4" blockId="1.[151,501,359,419]" box="[151,274,394,419]" pageId="1" pageNumber="186">
(
<figureCitation id="13422A4EFFF9FF9E9DF3DDDCFE8AE9E4" box="[159,265,394,419]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1734,1756]" captionStartId-1="3.[151,250,1446,1468]" captionStartId-2="4.[151,250,1639,1661]" captionStartId-3="5.[151,250,1049,1071]" captionTargetBox-0="[158,1434,382,1703]" captionTargetBox-1="[155,1431,486,1420]" captionTargetBox-2="[169,1416,191,1620]" captionTargetBox-3="[151,1436,301,1026]" captionTargetId-0="figure@2.[151,1436,373,1713]" captionTargetId-1="figure@3.[151,1436,481,1425]" captionTargetId-2="figure@4.[162,1426,183,1625]" captionTargetId-3="figure@5.[151,1436,301,1027]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." captionText-1="FIGURE 2. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, major (right) cheliped, lateral view; b, same, mesial view; c, same, chela, mesial view; d, same, chela fingers, lateral view; e, detail of tooth on cutting edge of dactylus, lateral view." captionText-2="FIGURE 3. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, minor (left) cheliped, lateral view; b, same, chela, lateral view; c, same, chela fingers (open); d, second pereiopod, lateral view; e, third pereiopod, lateral view; f, fourth pereiopod, latreral view; g, fifth pereiopod, lateral view; h, same, distal portion of propodus and dactylus, mesial view." captionText-3="FIGURE 4. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): live shrimp in lateral view." httpUri-0="https://zenodo.org/record/267225/files/figure.png" httpUri-1="https://zenodo.org/record/267226/files/figure.png" httpUri-2="https://zenodo.org/record/267227/files/figure.png" httpUri-3="https://zenodo.org/record/267228/files/figure.png" pageId="1" pageNumber="186">Figs. 1–4</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="AB6911E5FFF9FF9E9DFBDD84FD60E9AF" box="[151,739,466,488]" pageId="1" pageNumber="186">
<taxonomicName id="4C794D48FFF9FF9E9DFBDD84FE90E9AF" ID-CoL="84PMC" box="[151,275,466,488]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEAD9FFF9FF9E9DFBDD84FE90E9AF" box="[151,275,466,488]" italics="true" pageId="1" pageNumber="186">Leptalpheus</emphasis>
</taxonomicName>
sp. 1— 
<treatmentCitation id="0AD810DAFFF9FF9E9C33DD84FD80E9AF" ID-DOI="http://doi.org/10.5281/zenodo.3505011" ID-Zenodo-Dep="3505011" author="Anker" box="[351,515,466,488]" httpUri="http://treatment.plazi.org/id/03BB87D5FF96FF8AFF7F207DFC399A17" page="24" pageId="1" pageNumber="186" year="2011">
<bibRefCitation id="EFE84B3AFFF9FF9E9C33DD84FE5EE9AF" author="Anker" box="[351,477,466,488]" pageId="1" pageNumber="186" refString="Anker, A. (2011) Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda). Zootaxa, 3041, 1 - 38." type="journal article" year="2011">Anker 2011</bibRefCitation>
: 24
</treatmentCitation>
, figs. 18A–E, 19A, B.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3636540FFF9FF9E9DFBDE43FEECEB99" pageId="1" pageNumber="186" type="materials_examined">
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<emphasis id="B90DEAD9FFF9FF9E9DFBDE43FEC2EA69" bold="true" box="[151,321,533,558]" pageId="1" pageNumber="186">
<typeStatus id="54C28869FFF9FF9E9DFBDE43FF51EA69" box="[151,210,533,558]" pageId="1" pageNumber="186">Type</typeStatus>
material
</emphasis>
. 
<collectingCountry id="F36E765BFFF9FF9E9C3FDE40FE2CEA6A" box="[339,431,534,557]" name="Panama" pageId="1" pageNumber="186">Panama</collectingCountry>
, Caribbean coast: 
<typeStatus id="54C28869FFF9FF9E9FEBDE43FD6FEA69" box="[647,748,533,558]" pageId="1" pageNumber="186" type="holotype">holotype</typeStatus>
, 
<specimenCount id="9D7FFD42FFF9FF9E9F97DE40FCC9EA69" box="[763,842,533,558]" pageId="1" pageNumber="186" type="male">1 male</specimenCount>
(cl 
<quantity id="4C819B2EFFF9FF9E9E11DE40FC58EA69" box="[893,987,534,558]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="186" unit="mm" value="7.2">7.2 mm</quantity>
), 
<collectionCode id="ED68AE0EFFF9FF9E9E9EDE40FBCCEA69" box="[1010,1103,534,558]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="186">MZUSP</collectionCode>
34067, Bocas del Toro, 
<collectingCountry id="F36E765BFFF9FF9E9818DE43FA1FEA69" box="[1396,1436,533,558]" name="Iceland" pageId="1" pageNumber="186">Isla</collectingCountry>
Colón, 
<collectionCode id="ED68AE0EFFF9FF9E9D80DE6FFEAAEA16" box="[236,297,569,593]" pageId="1" pageNumber="186">STRI</collectionCode>
Bay, near 
<collectionCode id="ED68AE0EFFF9FF9E9CC4DE6FFE67EA16" box="[424,484,569,593]" pageId="1" pageNumber="186">STRI</collectionCode>
dock, 9.351516 -82.257716, shallow subtidal sea grass flat near mangrove, depth: 
<quantity id="4C819B2EFFF9FF9E9DFBDE0BFE88EA32" box="[151,267,605,630]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="1" pageNumber="186" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 m</quantity>
, in burrows, suction pump, leg. A. Anker, 
<date id="FFC7100BFFF9FF9E9F9DDE0BFCE0EA31" box="[753,867,605,630]" pageId="1" pageNumber="186" value="2015-05-03">03.v.2015</date>
; 
<typeStatus id="54C28869FFF9FF9E9E03DE08FC5CEA31" box="[879,991,606,630]" pageId="1" pageNumber="186" type="paratype">paratypes</typeStatus>
: 
<specimenCount id="9D7FFD42FFF9FF9E9E86DE08FBCEEA31" box="[1002,1101,605,630]" pageId="1" pageNumber="186" type="female">1 female</specimenCount>
(cl 
<quantity id="4C819B2EFFF9FF9E9917DE0BFB55EA31" box="[1147,1238,605,630]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="186" unit="mm" value="5.5">5.5 mm</quantity>
), 
<collectionCode id="ED68AE0EFFF9FF9E9986DE08FAC4EA31" box="[1258,1351,606,630]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="186">MZUSP</collectionCode>
34068, between Colón and Portobelo, near Mechi, 9.491252 -79.709096, shallow pools on sand flat exposed at low tide, fringed by sea grass, near fossilised coral platform, depth: 
<quantity id="4C819B2EFFF9FF9E9E42DEF3FC21EAFA" box="[814,930,677,702]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="186" unit="m" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 m</quantity>
, in burrows, suction pump, coll. A. Anker 
<emphasis id="B90DEAD9FFF9FF9E98E4DEF0FF37EAA6" italics="true" pageId="1" pageNumber="186">et al.</emphasis>
, 
<date id="FFC7100BFFF9FF9E9DA8DE9FFEC2EAA5" box="[196,321,713,738]" pageId="1" pageNumber="186" value="2015-03-25">25.iii.2015</date>
; 
<specimenCount id="9D7FFD42FFF9FF9E9C3CDE9CFE30EAA5" box="[336,435,713,738]" pageId="1" pageNumber="186" type="female">1 female</specimenCount>
(cl 
<quantity id="4C819B2EFFF9FF9E9C89DE9CFDC1EAA5" box="[485,578,714,738]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="186" unit="mm" value="7.1">7.1 mm</quantity>
), 
<collectionCode id="ED68AE0EFFF9FF9E9F34DE9CFD4BEAA5" box="[600,712,714,738]" pageId="1" pageNumber="186">OUMNH</collectionCode>
.ZC. 2015-08-025, 
<collectingCountry id="F36E765BFFF9FF9E9EF3DE9FFC44EAA5" box="[927,967,713,738]" name="Iceland" pageId="1" pageNumber="186">Isla</collectingCountry>
Grande, southwestern shore, 9.628281 - 79.565538, silt-mud bottom, depth: 
<quantity id="4C819B2EFFF9FF9E9F47DEBBFD0BEB42" box="[555,648,749,774]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="186" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 m</quantity>
, in burrows, suction pump, coll. A. Anker 
<emphasis id="B90DEAD9FFF9FF9E990BDEB8FB1CEB42" box="[1127,1183,749,773]" italics="true" pageId="1" pageNumber="186">et al.</emphasis>
, 
<date id="FFC7100BFFF9FF9E99C1DEBBFAA9EB41" box="[1197,1322,749,774]" pageId="1" pageNumber="186" value="2015-03-27">27.iii.2015</date>
.
</paragraph>
<paragraph id="8BC636CBFFF9FF9E9DABDF47FEECEB99" blockId="1.[151,1437,533,2034]" pageId="1" pageNumber="186">
<emphasis id="B90DEAD9FFF9FF9E9DABDF47FDB0EB6D" bold="true" box="[199,563,785,810]" pageId="1" pageNumber="186">Additional material examined</emphasis>
. 
<collectingCountry id="F36E765BFFF9FF9E9F28DF44FD23EB6E" box="[580,672,786,809]" name="Panama" pageId="1" pageNumber="186">Panama</collectingCountry>
, Caribbean coast: 
<specimenCount id="9D7FFD42FFF9FF9E9E1BDF44FC46EB6D" box="[887,965,785,810]" pageId="1" pageNumber="186" type="male">1 male</specimenCount>
(cl 
<quantity id="4C819B2EFFF9FF9E9E94DF44FBD6EB6D" box="[1016,1109,786,810]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="186" unit="mm" value="3.9">3.9 mm</quantity>
), 
<specimenCount id="9D7FFD42FFF9FF9E9907DF44FB4CEB6D" box="[1131,1231,785,810]" pageId="1" pageNumber="186" type="female">1 female</specimenCount>
(cl 
<quantity id="4C819B2EFFF9FF9E986DDF47FADDEB6D" box="[1281,1374,785,810]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="186" unit="mm" value="5.7">5.7 mm</quantity>
), UP, between Colón and Portobelo, 9.491252 -79.709096, shallow pools on sand flat exposed at low tide, fringed by sea grass, near fossilised coral platform, depth: 
<quantity id="4C819B2EFFF9FF9E9FEEDF0FFD76EB36" box="[642,757,857,882]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="186" unit="m" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 m</quantity>
, in burrows, suction pump, coll. A. Anker 
<emphasis id="B90DEAD9FFF9FF9E99B9DF0CFA8DEB36" box="[1237,1294,857,881]" italics="true" pageId="1" pageNumber="186">et al.</emphasis>
, 
<date id="FFC7100BFFF9FF9E9870DF0FFA1BEB35" box="[1308,1432,857,882]" pageId="1" pageNumber="186" value="2015-03-25">25.iii.2015</date>
. 
<collectingCountry id="F36E765BFFF9FF9E9DFBDF2BFE8CEBD1" box="[151,271,893,918]" name="Venezuela" pageId="1" pageNumber="186">Venezuela</collectingCountry>
: 
<specimenCount id="9D7FFD42FFF9FF9E9C73DF28FEF2EBD1" box="[287,369,893,918]" pageId="1" pageNumber="186" type="male">1 male</specimenCount>
(cl 
<quantity id="4C819B2EFFF9FF9E9CC6DF28FD88EBD1" box="[426,523,894,918]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="1" pageNumber="186" unit="mm" value="9.2">9.2 mm</quantity>
), UDO-GIC653, Sucre, off Barcelona, 
<collectingCountry id="F36E765BFFF9FF9E9E8DDF2BFB89EBD1" box="[993,1034,893,918]" name="Iceland" pageId="1" pageNumber="186">Isla</collectingCountry>
Chimana Grande, La Cienaguita, 10.290833 -64.650278, sand-mud flat near mangrove, depth: 
<quantity id="4C819B2EFFF9FF9E9E34DFF4FC4DEBFE" box="[856,974,930,954]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="186" unit="m" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 m</quantity>
, in burrows, suction pump, coll. J. Vera Caripe, 
<date id="FFC7100BFFF9FF9E9D83DF93FEE9EB99" box="[239,362,965,990]" pageId="1" pageNumber="186" value="2010-06-05">05.vi.2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3636540FFF9FF9C9DABDFBFFBABEF71" lastPageId="3" lastPageNumber="188" pageId="1" pageNumber="186" type="description">
<paragraph id="8BC636CBFFF9FF9E9DABDFBFFDB2ED65" blockId="1.[151,1437,533,2034]" pageId="1" pageNumber="186">
<emphasis id="B90DEAD9FFF9FF9E9DABDFBFFED2EC45" bold="true" box="[199,337,1001,1026]" pageId="1" pageNumber="186">Description</emphasis>
. Carapace with frontal margin produced in broadly triangular rostral projection; pterygostomial angle rounded, not anteriorly produced; branchiostegial margin with pronounced lip; posterior margin with deep cardiac notch; carapace surface finely pitted (
<figureCitation id="13422A4EFFF9FF9E9FCCD867FD61EC0D" box="[672,738,1073,1098]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="1" pageNumber="186">Fig. 1</figureCitation>
a, b). Pleon with all pleomeres rounded; sixth pleomere with subtriangular, distally somewhat rounded articulated flap (
<figureCitation id="13422A4EFFF9FF9E9E2CD803FC06EC29" box="[832,901,1109,1134]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="1" pageNumber="186">Fig. 1</figureCitation>
c). Telson relatively narrow, tapering distally; dorsal surface with two pairs of strong spiniform setae situated at some distance from lateral margin; posterior margin broadly rounded, about 0.7 length of proximal width of telson, each posterolateral angle with two pairs of spiniform setae, lateral much shorter than mesial; margin between mesial spiniform setae with numerous (12–14) long plumose setae (
<figureCitation id="13422A4EFFF9FF9E9CE2D8B3FE50ECB9" box="[398,467,1253,1278]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="1" pageNumber="186">Fig. 1</figureCitation>
d). Eyestalks with anteromesial margin rounded, protruding, with minute tubercle anterodorsally (indicated in 
<figureCitation id="13422A4EFFF9FF9E9CBED95FFD93ED65" box="[466,528,1289,1314]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="1" pageNumber="186">Fig. 1</figureCitation>
b).
</paragraph>
<paragraph id="8BC636CBFFF9FF9E9DABD97BFCF1EE59" blockId="1.[151,1437,533,2034]" pageId="1" pageNumber="186">
Antennular peduncles fairly stout, flattened dorsoventrally; stylocerite appressed against first article, with subacute point not reaching distal margin of first article; ventromesial carina of first article with simple, anteriorly pointing tooth; second article about twice as long as wide; lateral flagellum with short secondary ramus partly fused to main ramus, furnished with several tufts of aesthetascs (
<figureCitation id="13422A4EFFF9FF9E9EE4D9CFFC48EDF5" box="[904,971,1433,1458]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="1" pageNumber="186">Fig. 1</figureCitation>
a, b, e, f). Antenna with large basicerite ending in stout sharp distoventral tooth; scaphocerite oval, with small subacute distolateral tooth, latter not reaching beyond anterior margin of broad blade; carpocerite stout, reaching far beyond scaphocerite and slightly beyond end of antennular peduncle; flagellum robust (
<figureCitation id="13422A4EFFF9FF9E9F9BDA53FCB4EE59" box="[759,823,1541,1566]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="1" pageNumber="186">Fig. 1</figureCitation>
a, b).
</paragraph>
<paragraph id="8BC636CBFFF9FF9E9DABDA7FFF7CEEE9" blockId="1.[151,1437,533,2034]" pageId="1" pageNumber="186">
Mouthparts (mandible, maxillule, maxilla, first and second maxillipeds) typical for genus in external view. Third maxilliped with lateral plate on coxa strongly projecting towards arthrobranch, distally subacute; ultimate article shorter than antepenultimate, with rings of thick serrulate setae and blunt tip, latter without spiniform setae (
<figureCitation id="13422A4EFFF9FF9E9DF3DAC3FF5CEEE9" box="[159,223,1685,1710]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="1" pageNumber="186">
Fig. 
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g).
</paragraph>
<paragraph id="8BC636CBFFF9FF9D9DABDAEFFB47E907" blockId="1.[151,1437,533,2034]" lastBlockId="2.[151,1437,151,320]" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186">
Chelipeds very asymmetrical in shape and unequal in size, with smooth surfaces, both folded when not in use. Major cheliped slender proximally, but with relatively stout chela; ischium very short; merus slender, slightly twisted, ventrally depressed, ventromesial margin rugose, ending in small triangular tooth; carpus short, cupshaped; chela elongate, moderately swollen, with palm depressed ventrally, fingers about half palm length, twisted, slightly gaping when closed; cutting edge of pollex with one small proximal tooth, one large broad subtriangular tooth at about 0.4 length of pollex, one stout, distally truncate subdistal tooth perpendicular to pollex axis, and one smaller distal tooth adjacent to truncate pollex; cutting edge of dactylus with one large, distally truncate and minutely serrated tooth, situated at about 0.3 pollex length, one broad bulge occupying cutting edge between 0.3 and 0.8 pollex length, and one small subtriangular subdistal tooth, dactylus tip strongly curved, truncate (
<figureCitation id="13422A4EFFF9FF9E9852DB8FFA02EFB5" box="[1342,1409,2009,2034]" captionStart="FIGURE 2" captionStartId="3.[151,250,1446,1468]" captionTargetBox="[155,1431,486,1420]" captionTargetId="figure@3.[151,1436,481,1425]" captionTargetPageId="3" captionText="FIGURE 2. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, major (right) cheliped, lateral view; b, same, mesial view; c, same, chela, mesial view; d, same, chela fingers, lateral view; e, detail of tooth on cutting edge of dactylus, lateral view." httpUri="https://zenodo.org/record/267226/files/figure.png" pageId="1" pageNumber="186">Fig. 2</figureCitation>
a– e). Minor cheliped much smaller than major cheliped; ischium short; merus moderately slender, ventrally depressed, with smooth margins; carpus very short, cup-shaped; chela moderately inflated, flattened mesially, with fingers slightly longer than palm, with crossing tips; cutting edge of pollex with about 14 small, widely spaced teeth evenly distributed from pollex base to about 0.7 pollex length; cutting edge of dactylus with seven small, widely spaced teeth, distributed approximately between 0.3 and 0.7 dactylus length (
<figureCitation id="13422A4EFFFAFF9D9927DD71FB0FE907" box="[1099,1164,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1639,1661]" captionTargetBox="[169,1416,191,1620]" captionTargetId="figure@4.[162,1426,183,1625]" captionTargetPageId="4" captionText="FIGURE 3. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, minor (left) cheliped, lateral view; b, same, chela, lateral view; c, same, chela fingers (open); d, second pereiopod, lateral view; e, third pereiopod, lateral view; f, fourth pereiopod, latreral view; g, fifth pereiopod, lateral view; h, same, distal portion of propodus and dactylus, mesial view." httpUri="https://zenodo.org/record/267227/files/figure.png" pageId="2" pageNumber="187">Fig. 3</figureCitation>
a–c).
</paragraph>
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<paragraph id="8BC636CBFFFAFF9D9DFBDA90FABBEF7F" blockId="2.[151,1436,1734,1848]" pageId="2" pageNumber="187">
<emphasis id="B90DEAD9FFFAFF9D9DFBDA90FE96EE9C" bold="true" box="[151,277,1734,1756]" pageId="2" pageNumber="187">FIGURE 1.</emphasis>
<taxonomicName id="4C794D48FFFAFF9D9C72DA90FE77EE9B" box="[286,500,1734,1756]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFAFF9D9C72DA90FE77EE9B" box="[286,500,1734,1756]" italics="true" pageId="2" pageNumber="187">Leptalpheus pereirai</emphasis>
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<emphasis id="B90DEAD9FFFAFF9D9C90DA90FDCCEE9C" bold="true" box="[508,591,1734,1755]" pageId="2" pageNumber="187">sp. nov.</emphasis>
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, holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC636CBFFFAFF9C9DABDB09FA9DE9EA" blockId="2.[151,1437,1887,2020]" lastBlockId="3.[151,1437,151,429]" lastPageId="3" lastPageNumber="188" pageId="2" pageNumber="187">
Second pereiopod slender, with merus distinctly longer than ischium; carpus five-articulated, with article ratio approximately equal to 5.0: 1.0: 1.5: 1.2: 3.4 (
<figureCitation id="13422A4EFFFAFF9D9FA7DBD5FC8CEFDB" box="[715,783,1923,1948]" captionStart="FIGURE 3" captionStartId="4.[151,250,1639,1661]" captionTargetBox="[169,1416,191,1620]" captionTargetId="figure@4.[162,1426,183,1625]" captionTargetPageId="4" captionText="FIGURE 3. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, minor (left) cheliped, lateral view; b, same, chela, lateral view; c, same, chela fingers (open); d, second pereiopod, lateral view; e, third pereiopod, lateral view; f, fourth pereiopod, latreral view; g, fifth pereiopod, lateral view; h, same, distal portion of propodus and dactylus, mesial view." httpUri="https://zenodo.org/record/267227/files/figure.png" pageId="2" pageNumber="187">Fig. 3</figureCitation>
d). Third pereiopod relatively stout, compressed; ischium without spiniform seta on ventrolateral surface; merus about four times as long as wide; carpus about 0.4 length of merus, with distoventral spiniform seta; propodus with two short spiniform setae on ventral margin and one pair of spiniform setae adjacent to dactylus; dactylus about half length of propodus, slender, conical, slightly curved, acute distally (
<figureCitation id="13422A4EFFFBFF9C9D95DCEAFEB8E893" box="[249,315,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1639,1661]" captionTargetBox="[169,1416,191,1620]" captionTargetId="figure@4.[162,1426,183,1625]" captionTargetPageId="4" captionText="FIGURE 3. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, minor (left) cheliped, lateral view; b, same, chela, lateral view; c, same, chela fingers (open); d, second pereiopod, lateral view; e, third pereiopod, lateral view; f, fourth pereiopod, latreral view; g, fifth pereiopod, lateral view; h, same, distal portion of propodus and dactylus, mesial view." httpUri="https://zenodo.org/record/267227/files/figure.png" pageId="3" pageNumber="188">Fig. 3</figureCitation>
e). Fourth pereiopod generally similar to third pereiopod, merus more slender, close to five times as long as wide; carpus with much smaller distoventral spiniform seta; propodus with three short spiniform setae on ventral margin and one pair of spiniform setae adjacent to dactylus; dactylus about 0.4 length of propodus, moderately slender, conical, slightly curved, acute distally (
<figureCitation id="13422A4EFFFBFF9C9E53DD71FC07E907" box="[831,900,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1639,1661]" captionTargetBox="[169,1416,191,1620]" captionTargetId="figure@4.[162,1426,183,1625]" captionTargetPageId="4" captionText="FIGURE 3. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, minor (left) cheliped, lateral view; b, same, chela, lateral view; c, same, chela fingers (open); d, second pereiopod, lateral view; e, third pereiopod, lateral view; f, fourth pereiopod, latreral view; g, fifth pereiopod, lateral view; h, same, distal portion of propodus and dactylus, mesial view." httpUri="https://zenodo.org/record/267227/files/figure.png" pageId="3" pageNumber="188">Fig. 3</figureCitation>
f). Fifth pereiopod distinctly more slender than third and fourth pereiopods, not compressed; carpus about 0.6 length of merus; propodus with well-developed grooming brush distolaterally, consisting of at least nine rows of serrulate setae, ventromesial margin with two short spiniform setae (invisible in lateral view); dactylus similar to that of fourth pereiopod (
<figureCitation id="13422A4EFFFBFF9C99CEDDC2FB67E9EB" box="[1186,1252,404,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1639,1661]" captionTargetBox="[169,1416,191,1620]" captionTargetId="figure@4.[162,1426,183,1625]" captionTargetPageId="4" captionText="FIGURE 3. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, minor (left) cheliped, lateral view; b, same, chela, lateral view; c, same, chela fingers (open); d, second pereiopod, lateral view; e, third pereiopod, lateral view; f, fourth pereiopod, latreral view; g, fifth pereiopod, lateral view; h, same, distal portion of propodus and dactylus, mesial view." httpUri="https://zenodo.org/record/267227/files/figure.png" pageId="3" pageNumber="188">
Fig. 
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</figureCitation>
g, h).
</paragraph>
<caption id="DF066643FFFBFF9C9DFBD9F0FD45EDBD" httpUri="https://zenodo.org/record/267226/files/figure.png" pageId="3" pageNumber="188" targetBox="[155,1431,486,1420]" targetPageId="3">
<paragraph id="8BC636CBFFFBFF9C9DFBD9F0FD45EDBD" blockId="3.[151,1436,1446,1530]" pageId="3" pageNumber="188">
<emphasis id="B90DEAD9FFFBFF9C9DFBD9F0FE96EDFB" bold="true" box="[151,277,1446,1468]" pageId="3" pageNumber="188">FIGURE 2.</emphasis>
<taxonomicName id="4C794D48FFFBFF9C9C72D9F0FE77EDFA" box="[286,500,1446,1469]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFBFF9C9C72D9F0FE77EDFA" box="[286,500,1446,1469]" italics="true" pageId="3" pageNumber="188">Leptalpheus pereirai</emphasis>
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<emphasis id="B90DEAD9FFFBFF9C9C90D9F1FDCCEDFB" bold="true" box="[508,591,1447,1468]" pageId="3" pageNumber="188">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, major (right) cheliped, lateral view; b, same, mesial view; c, same, chela, mesial view; d, same, chela fingers, lateral view; e, detail of tooth on cutting edge of dactylus, lateral view.
</paragraph>
</caption>
<paragraph id="8BC636CBFFFBFF9C9DABDA77FC8BEE8D" blockId="3.[151,1436,1569,2026]" pageId="3" pageNumber="188">
Male second pleopod with slender appendix masculina significantly exceeding appendix interna, with stiff setae on apex and margin opposed to endopod margin (
<figureCitation id="13422A4EFFFBFF9C9F90DA13FCB8EE19" box="[764,827,1605,1630]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="3" pageNumber="188">Fig. 1</figureCitation>
h). Uropod with lateral lobe of protopod ending in two small, sharp, somewhat spaced teeth; exopod with distal margin truncate, forming conspicuous lateral angle; diaeresis with straight lateral portion, broad and deep mesial incision flanked by strong projecting tooth near mesial margin (
<figureCitation id="13422A4EFFFBFF9C9D99DAE7FEB6EE8D" box="[245,309,1713,1738]" captionStart="FIGURE 1" captionStartId="2.[151,250,1734,1756]" captionTargetBox="[158,1434,382,1703]" captionTargetId="figure@2.[151,1436,373,1713]" captionTargetPageId="2" captionText="FIGURE 1. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, frontal region, dorsal view; b, same, lateral view; c, sixth pleomere, lateral view; d, telson, dorsal view; e, tooth on ventromesial carina of first article of antennular peduncle, lateral view; f, proximal portion of antennular flagella, lateral view; g, third maxilliped, lateral view; h, appendices masculina and interna of second pleopod, mesial view; i, uropod, dorsal view." httpUri="https://zenodo.org/record/267225/files/figure.png" pageId="3" pageNumber="188">
Fig. 
<date id="FFC7100BFFFBFF9C9C45DAE7FEB6EE8D" box="[297,309,1713,1738]" pageId="3" pageNumber="188">1</date>
</figureCitation>
i). Gill-exopod formula typical for genus.
</paragraph>
<paragraph id="8BC636CBFFFBFF9C9DABDA83FBABEF71" blockId="3.[151,1436,1569,2026]" pageId="3" pageNumber="188">
<emphasis id="B90DEAD9FFFBFF9C9DABDA83FEDBEEA9" bold="true" box="[199,344,1749,1774]" pageId="3" pageNumber="188">Colouration</emphasis>
. Body semitransparent pinkish, with numerous scattered red chromatophores, forming diffuse bands on the pleon; antennules and uropods sometimes brighter due to higher concentration of red chromatophores; major and minor chelae creamy white (
<figureCitation id="13422A4EFFFBFF9C9F22DB4BFD10EF71" box="[590,659,1821,1846]" captionStart="FIGURE 4" captionStartId="5.[151,250,1049,1071]" captionTargetBox="[151,1436,301,1026]" captionTargetId="figure@5.[151,1436,301,1027]" captionTargetPageId="5" captionText="FIGURE 4. Leptalpheus pereirai sp. nov., holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): live shrimp in lateral view." httpUri="https://zenodo.org/record/267228/files/figure.png" pageId="3" pageNumber="188">Fig. 4</figureCitation>
); see also 
<bibRefCitation id="EFE84B3AFFFBFF9C9E6BDB4BFC0CEF71" author="Anker" box="[775,911,1821,1846]" pageId="3" pageNumber="188" refString="Anker, A. (2011) Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda). Zootaxa, 3041, 1 - 38." type="journal article" year="2011">Anker 2011</bibRefCitation>
: fig. 19A, B).
</paragraph>
</subSubSection>
<subSubSection id="C3636540FFFBFF9C9DABDB17FCDFEF1E" box="[199,860,1857,1882]" pageId="3" pageNumber="188" type="materials_examined">
<paragraph id="8BC636CBFFFBFF9C9DABDB17FCDFEF1E" blockId="3.[151,1436,1569,2026]" box="[199,860,1857,1882]" pageId="3" pageNumber="188">
<emphasis id="B90DEAD9FFFBFF9C9DABDB17FEDEEF1D" bold="true" box="[199,349,1857,1882]" pageId="3" pageNumber="188">
<typeStatus id="54C28869FFFBFF9C9DABDB17FE81EF1D" box="[199,258,1857,1882]" pageId="3" pageNumber="188">Type</typeStatus>
locality
</emphasis>
. Bocas del Toro, Caribbean coast of 
<collectingCountry id="F36E765BFFFBFF9C9F90DB14FCDBEF1E" box="[764,856,1858,1881]" name="Panama" pageId="3" pageNumber="188">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3636540FFFBFF9C9DABDB33FD9AEFE5" pageId="3" pageNumber="188" type="distribution">
<paragraph id="8BC636CBFFFBFF9C9DABDB33FD9AEFE5" blockId="3.[151,1436,1569,2026]" pageId="3" pageNumber="188">
<emphasis id="B90DEAD9FFFBFF9C9DABDB33FEDAEF39" bold="true" box="[199,345,1893,1918]" pageId="3" pageNumber="188">Distribution</emphasis>
. Southern Caribbean Sea: 
<collectingCountry id="F36E765BFFFBFF9C9FF5DB30FD71EF3A" box="[665,754,1894,1917]" name="Panama" pageId="3" pageNumber="188">Panama</collectingCountry>
(Bocas del Toro, Colón-Portobelo area, 
<collectingCountry id="F36E765BFFFBFF9C99BBDB33FA83EF39" box="[1239,1280,1893,1918]" name="Iceland" pageId="3" pageNumber="188">Isla</collectingCountry>
Grande) and 
<collectingCountry id="F36E765BFFFBFF9C9DFBDBDFFE89EFE5" box="[151,266,1929,1954]" name="Venezuela" pageId="3" pageNumber="188">Venezuela</collectingCountry>
(
<collectingCountry id="F36E765BFFFBFF9C9C75DBDFFEC0EFE5" box="[281,323,1929,1954]" name="Iceland" pageId="3" pageNumber="188">Isla</collectingCountry>
Chimana Grande).
</paragraph>
</subSubSection>
<subSubSection id="C3636540FFFBFF9A9DABDBFBFA01E8BF" lastPageId="5" lastPageNumber="190" pageId="3" pageNumber="188" type="etymology">
<paragraph id="8BC636CBFFFBFF9C9DABDBFBFE06EFAD" blockId="3.[151,1436,1569,2026]" pageId="3" pageNumber="188">
<emphasis id="B90DEAD9FFFBFF9C9DABDBFBFEC5EF81" bold="true" box="[199,326,1965,1990]" pageId="3" pageNumber="188">Etymology</emphasis>
. The species is named after our late friend and colleague, the Venezuelan carcinologist Dr. Guido Pereira (
<date id="FFC7100BFFFBFF9C9D99DB87FEFAEFAD" box="[245,377,2001,2026]" pageId="3" pageNumber="188" value="1951" valueMax="2015">1951–2015</date>
).
</paragraph>
<caption id="DF066643FFFCFF9B9DFBDA31FD85EE9E" httpUri="https://zenodo.org/record/267227/files/figure.png" pageId="4" pageNumber="189" targetBox="[169,1416,191,1620]" targetPageId="4">
<paragraph id="8BC636CBFFFCFF9B9DFBDA31FD85EE9E" blockId="4.[151,1436,1639,1753]" pageId="4" pageNumber="189">
<emphasis id="B90DEAD9FFFCFF9B9DFBDA31FE96EE3B" bold="true" box="[151,277,1639,1661]" pageId="4" pageNumber="189">FIGURE 3.</emphasis>
<taxonomicName id="4C794D48FFFCFF9B9C72DA31FE77EE3A" box="[286,500,1639,1661]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFCFF9B9C72DA31FE77EE3A" box="[286,500,1639,1661]" italics="true" pageId="4" pageNumber="189">Leptalpheus pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFCFF9B9C90DA31FDCCEE3B" box="[508,591,1639,1660]" pageId="4" pageNumber="189" rank="species">
<emphasis id="B90DEAD9FFFCFF9B9C90DA31FDCCEE3B" bold="true" box="[508,591,1639,1660]" pageId="4" pageNumber="189">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): a, minor (left) cheliped, lateral view; b, same, chela, lateral view; c, same, chela fingers (open); d, second pereiopod, lateral view; e, third pereiopod, lateral view; f, fourth pereiopod, latreral view; g, fifth pereiopod, lateral view; h, same, distal portion of propodus and dactylus, mesial view.
</paragraph>
</caption>
<paragraph id="8BC636CBFFFCFF9A9DABDB56FA01E8BF" blockId="4.[151,1437,1792,2033]" lastBlockId="5.[151,1435,151,248]" lastPageId="5" lastPageNumber="190" pageId="4" pageNumber="189">
<emphasis id="B90DEAD9FFFCFF9B9DABDB56FEA7EF5E" bold="true" box="[199,292,1792,1817]" pageId="4" pageNumber="189">Ecology</emphasis>
. The Panamanian specimens of 
<taxonomicName id="4C794D48FFFCFF9B9FF0DB56FC0BEF5F" box="[668,904,1792,1816]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFCFF9B9FF0DB56FC0BEF5F" box="[668,904,1792,1816]" italics="true" pageId="4" pageNumber="189">Leptalpheus pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFCFF9B9EFDDB56FC6DEF5E" box="[913,1006,1792,1817]" pageId="4" pageNumber="189" rank="species">
<emphasis id="B90DEAD9FFFCFF9B9EFDDB56FC6DEF5E" bold="true" box="[913,1006,1792,1817]" pageId="4" pageNumber="189">sp. nov.</emphasis>
</taxonomicNameLabel>
were collected from burrows in silty sand or sandy mud substrate, usually close to or within sea grass beds, sometimes close to mangroves, in the lower intertidal or at shallow subtidal depths (
<quantity id="4C819B2EFFFCFF9B9F15DB1EFD75EF26" box="[633,758,1864,1889]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="4" pageNumber="189" unit="m" value="0.85" valueMax="1.5" valueMin="0.2">0.2–1.5 m</quantity>
). The callianassid ghost shrimp 
<taxonomicName id="4C794D48FFFCFF9B9912DB1EFECAEFC2" authority="Stimpson, 1866" authorityName="Stimpson" authorityYear="1866" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="acanthochirus">
<emphasis id="B90DEAD9FFFCFF9B9912DB1EFA1FEF27" box="[1150,1436,1864,1888]" italics="true" pageId="4" pageNumber="189">Glypturus acanthochirus</emphasis>
<bibRefCitation id="EFE84B3AFFFCFF9B9DFBDB3AFECAEFC2" author="Stimpson" box="[151,329,1900,1925]" pageId="4" pageNumber="189" refString="Stimpson, W. (1866) Descriptions of new genera and species of macrurous Crustacea from the coasts of North America. Proceedings of the Chicago Academy of Sciences, 1, 46 - 48." type="journal article" year="1866">Stimpson, 1866</bibRefCitation>
</taxonomicName>
was collected at all three Panamanian sites (Bocas del Toro, Mechi, and 
<collectingCountry id="F36E765BFFFCFF9B99F0DB3AFB47EFC2" box="[1180,1220,1900,1925]" name="Iceland" pageId="4" pageNumber="189">Isla</collectingCountry>
Grande), although never together with 
<taxonomicName id="4C794D48FFFCFF9B9C17DBC7FE70EFEF" box="[379,499,1937,1960]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFCFF9B9C17DBC7FE70EFEF" box="[379,499,1937,1960]" italics="true" pageId="4" pageNumber="189">L. pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFCFF9B9C97DBC6FDD5EFEE" box="[507,598,1936,1961]" pageId="4" pageNumber="189" rank="species">
<emphasis id="B90DEAD9FFFCFF9B9C97DBC6FDD5EFEE" bold="true" box="[507,598,1936,1961]" pageId="4" pageNumber="189">sp. nov.</emphasis>
</taxonomicNameLabel>
The single Venezuelan specimen was collected in similar conditions, in a depth of 
<quantity id="4C819B2EFFFCFF9B9D91DBE3FEF2EF8B" box="[253,369,1973,1997]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="4" pageNumber="189" unit="m" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 m</quantity>
, and only a few meters away from a mangrove, which are abundant on 
<collectingCountry id="F36E765BFFFCFF9B99C8DBE2FB4EEF8A" box="[1188,1229,1972,1997]" name="Iceland" pageId="4" pageNumber="189">Isla</collectingCountry>
Chimana Grande. The syntopic burrowing decapod fauna included the snapping shrimp 
<taxonomicName id="4C794D48FFFCFF9B9ED6DB8EFA1BEFB6" authority="Christoffersen, 1984" authorityName="Christoffersen" authorityYear="1984" box="[954,1432,2008,2033]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="estuariensis">
<emphasis id="B90DEAD9FFFCFF9B9ED6DB8EFB27EFB7" box="[954,1188,2008,2032]" italics="true" pageId="4" pageNumber="189">Alpheus estuariensis</emphasis>
<bibRefCitation id="EFE84B3AFFFCFF9B99C1DB8EFA1BEFB6" author="Christoffersen" box="[1197,1432,2008,2033]" pageId="4" pageNumber="189" refString="Christoffersen, M. L. (1984) The western Atlantic snapping shrimps related to Alpheus heterochaelis Say (Crustacea, Caridea) with the description of a new species. Papeis Avulsos de Zoologia, 35, 189 - 208." type="journal article" year="1984">Christoffersen, 1984</bibRefCitation>
</taxonomicName>
, and three ghost and mud shrimps, viz. 
<taxonomicName id="4C794D48FFFDFF9A9F1CDCCEFBC2E8F7" box="[624,1089,151,176]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="acantochirus">
<emphasis id="B90DEAD9FFFDFF9A9F1CDCCEFCFCE8F7" box="[624,895,152,176]" italics="true" pageId="5" pageNumber="190">Glypturus acantochirus</emphasis>
(Callianassidae)
</taxonomicName>
<taxonomicName id="4C794D48FFFDFF9A9921DCCEFE75E892" authority="Gomes Correa, 1968" authorityName="Gomes Correa" authorityYear="1968" class="Malacostraca" family="Upogebiidae" genus="Upogebia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="omissa">
<emphasis id="B90DEAD9FFFDFF9A9921DCCEFB38E8F7" box="[1101,1211,152,176]" italics="true" pageId="5" pageNumber="190">Upogebia</emphasis>
cf. 
<emphasis id="B90DEAD9FFFDFF9A999EDCCEFAC3E8E8" box="[1266,1344,152,175]" italics="true" pageId="5" pageNumber="190">omissa</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9820DCCEFEBAE892" author="Gomes" pageId="5" pageNumber="190" refString="Gomes Correa, M. M. (1968) Sobre as especies de &quot; Upogebia &quot; Leach do litoral brasileiro, com descricao de uma especie nova. Revista Brasileira de Biologia, 28, 97 - 109." type="journal article" year="1968">Gomes Corrêa, 1968</bibRefCitation>
(Upogebiidae)
</taxonomicName>
, and 
<taxonomicName id="4C794D48FFFDFF9A9F3DDCEAFB5EE892" authority="Schmitt, 1924" authorityName="Schmitt" authorityYear="1924" box="[593,1245,188,213]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B90DEAD9FFFDFF9A9F3DDCEAFCD7E893" box="[593,852,188,212]" italics="true" pageId="5" pageNumber="190">Axianassa intermedia</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9E00DCEAFB9AE893" author="Schmitt" box="[876,1049,188,213]" pageId="5" pageNumber="190" refString="Schmitt, W. L. (1924) Bidragen tot de kennis der fauna van Curacao. Resultaten eener reis van Dr. C. J. van der Horst in 1920. The macruran, anomuran and stomatopod Crustacea. Bijdragen tot de Dierkunde, 23, 61 - 81." type="journal article" year="1924">Schmitt, 1924</bibRefCitation>
(Axianassidae)
</taxonomicName>
. Therefore, 
<taxonomicName id="4C794D48FFFDFF9A98E8DCEAFEB4E8BF" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="acanthochirus">
<emphasis id="B90DEAD9FFFDFF9A98E8DCEAFEB4E8BF" italics="true" pageId="5" pageNumber="190">G. acanthochirus</emphasis>
</taxonomicName>
, being present at all collection sites, may well represent the burrowing host of 
<taxonomicName id="4C794D48FFFDFF9A99C6DCB7FAA2E8B0" box="[1194,1313,224,248]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A99C6DCB7FAA2E8B0" box="[1194,1313,224,248]" italics="true" pageId="5" pageNumber="190">L. pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A9844DC89FA01E8BF" box="[1320,1410,223,248]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A9844DC89FA01E8BF" bold="true" box="[1320,1410,223,248]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="DF066643FFFDFF9A9DFBD84FFEF9EC0A" httpUri="https://zenodo.org/record/267228/files/figure.png" pageId="5" pageNumber="190" targetBox="[151,1436,301,1026]" targetPageId="5">
<paragraph id="8BC636CBFFFDFF9A9DFBD84FFEF9EC0A" blockId="5.[151,1435,1049,1101]" pageId="5" pageNumber="190">
<emphasis id="B90DEAD9FFFDFF9A9DFBD84FFE97EC69" bold="true" box="[151,276,1049,1071]" pageId="5" pageNumber="190">FIGURE 4.</emphasis>
<taxonomicName id="4C794D48FFFDFF9A9C70D84FFE73EC68" box="[284,496,1049,1071]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A9C70D84FFE73EC68" box="[284,496,1049,1071]" italics="true" pageId="5" pageNumber="190">Leptalpheus pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A9C94D84FFDCAEC69" box="[504,585,1049,1070]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A9C94D84FFDCAEC69" bold="true" box="[504,585,1049,1070]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, male (cl 7.2 mm), from Bocas del Toro, Panama (MZUSP 34067): live shrimp in lateral view.
</paragraph>
</caption>
<subSubSection id="C3636540FFFDFF999DABD82CFD94E95B" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190" type="discussion">
<paragraph id="8BC636CBFFFDFF9A9DABD82CFAAEED00" blockId="5.[151,1436,1146,2035]" pageId="5" pageNumber="190">
<emphasis id="B90DEAD9FFFDFF9A9DABD82CFEB7ECD4" bold="true" box="[199,308,1146,1171]" pageId="5" pageNumber="190">Remarks</emphasis>
. 
<taxonomicName id="4C794D48FFFDFF9A9C2ED82CFDA9ECD5" box="[322,554,1146,1170]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A9C2ED82CFDA9ECD5" box="[322,554,1146,1170]" italics="true" pageId="5" pageNumber="190">Leptalpheus pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A9F5DD82CFD08ECD4" box="[561,651,1146,1171]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A9F5DD82CFD08ECD4" bold="true" box="[561,651,1146,1171]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to an informal group of species characterised by the presence of well-developed adhesive disks on the major chela. This group also includes the western Atlantic 
<taxonomicName id="4C794D48FFFDFF9A986DD8C9FE50EC9C" authority="Dworschak &amp; Coelho, 1999" authorityName="Dworschak &amp; Coelho" authorityYear="1999" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="axianassae">
<emphasis id="B90DEAD9FFFDFF9A986DD8C9FA1FECF1" box="[1281,1436,1183,1206]" italics="true" pageId="5" pageNumber="190">L. axianassae</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9DFBD894FE50EC9C" author="Dworschak" box="[151,467,1218,1243]" pageId="5" pageNumber="190" refString="Dworschak, P. C. &amp; Coelho, V. R. (1999) On two alpheids from Araca (Sao Paulo, Brazil) with a description of a new species of Leptalpheus (Decapoda: Caridea: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 101 B, 475 - 488." type="journal article" year="1999">Dworschak &amp; Coelho, 1999</bibRefCitation>
</taxonomicName>
, the eastern Pacific 
<taxonomicName id="4C794D48FFFDFF9A9FDED895FBF1EC9C" authority="Anker &amp; Lazarus, 2015" authorityName="Anker &amp; Lazarus" authorityYear="2015" box="[690,1138,1218,1243]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="canterakintzi">
<emphasis id="B90DEAD9FFFDFF9A9FDED895FCE1EC9D" box="[690,866,1218,1242]" italics="true" pageId="5" pageNumber="190">L. canterakintzi</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9E04D894FBF1EC9C" author="Anker" box="[872,1138,1218,1243]" pageId="5" pageNumber="190" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus, 2015</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C794D48FFFDFF9A9912D895FAF9EC9C" authority="Anker, 2011" authorityName="Anker" authorityYear="2011" box="[1150,1402,1218,1243]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azuero">
<emphasis id="B90DEAD9FFFDFF9A9912D895FB6BEC9D" box="[1150,1256,1219,1242]" italics="true" pageId="5" pageNumber="190">L. azuero</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9982D894FAF9EC9C" author="Anker" box="[1262,1402,1218,1243]" pageId="5" pageNumber="190" refString="Anker, A. (2011) Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda). Zootaxa, 3041, 1 - 38." type="journal article" year="2011">Anker, 2011</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C794D48FFFDFF9A98E9D895FCEAECB8" authority="Salgado-Barragan, Ayon-Parente &amp; Hendrickx, 2014" authorityName="Salgado-Barragan, Ayon-Parente &amp; Hendrickx" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="corderoae">
<emphasis id="B90DEAD9FFFDFF9A98E9D895FE8BECB9" italics="true" pageId="5" pageNumber="190">L. corderoae</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9C7CD8B0FCEAECB8" author="Salgado-Barragan" box="[272,873,1254,1279]" pageId="5" pageNumber="190" refString="Salgado-Barragan, J., Ayon-Parente, M. &amp; Hendrickx, M. E. (2014) A new species of Leptalpheus Williams, 1965 and new records of L. mexicanus Rios &amp; Carvacho, 1983 and L. hendrickxi Anker, 2011 (Crustacea: Decapoda: Alpheidae) from the Pacific coast of Mexico. Zootaxa, 3835 (4), 573 - 582. http: // dx. doi. org / 10.11646 / zootaxa. 3835.4.8" type="journal article" year="2014">Salgado-Barragán, Ayon-Parente &amp; Hendrickx, 2014</bibRefCitation>
</taxonomicName>
, and the Indo-West Pacific 
<taxonomicName id="4C794D48FFFDFF9A99CDD8B1FEB0ED64" authority="Banner &amp; Banner, 1974" authorityName="Banner &amp; Banner" authorityYear="1974" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B90DEAD9FFFDFF9A99CDD8B1FAA6ECB9" box="[1185,1317,1254,1278]" italics="true" pageId="5" pageNumber="190">L. pacificus</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9841D8B1FEB0ED64" author="Banner" pageId="5" pageNumber="190" refString="Banner, A. H. &amp; Banner, D. M. (1974) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part XVII: additional notes on the Hawaiian alpheids: new species, subspecies and some nomenclatorial changes. Pacific Science, 28, 423 - 437." type="journal article" year="1974">Banner &amp; Banner, 1974</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C794D48FFFDFF9A9C2ED95DFD7AED64" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[322,761,1290,1315]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B90DEAD9FFFDFF9A9C2ED95DFE6FED65" box="[322,492,1290,1314]" italics="true" pageId="5" pageNumber="190">L. denticulatus</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9C9BD95CFD7AED64" author="Anker" box="[503,761,1290,1315]" pageId="5" pageNumber="190" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin, 2009</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C794D48FFFDFF9A9E57D95DFB64ED64" authority="Anker &amp; Marin, 2009" authorityName="Anker &amp; Marin" authorityYear="2009" box="[827,1255,1290,1315]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="dworschaki">
<emphasis id="B90DEAD9FFFDFF9A9E57D95DFC5EED65" box="[827,989,1290,1314]" italics="true" pageId="5" pageNumber="190">L. dworschaki</emphasis>
<bibRefCitation id="EFE84B3AFFFDFF9A9E84D95CFB64ED64" author="Anker" box="[1000,1255,1290,1315]" pageId="5" pageNumber="190" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin, 2009</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFE84B3AFFFDFF9A9996D95CFEA9ED00" author="Dworschak" pageId="5" pageNumber="190" refString="Dworschak, P. C. &amp; Coelho, V. R. (1999) On two alpheids from Araca (Sao Paulo, Brazil) with a description of a new species of Leptalpheus (Decapoda: Caridea: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 101 B, 475 - 488." type="journal article" year="1999">Dworschak &amp; Coelho 1999</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A9C59D978FDA5ED00" author="Anker" box="[309,550,1326,1351]" pageId="5" pageNumber="190" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A9F5DD978FD3AED00" author="Anker" box="[561,697,1326,1351]" pageId="5" pageNumber="190" refString="Anker, A. (2011) Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda). Zootaxa, 3041, 1 - 38." type="journal article" year="2011">Anker 2011</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A9FA8D978FB8CED00" author="Salgado-Barragan" box="[708,1039,1326,1351]" pageId="5" pageNumber="190" refString="Salgado-Barragan, J., Ayon-Parente, M. &amp; Hendrickx, M. E. (2014) A new species of Leptalpheus Williams, 1965 and new records of L. mexicanus Rios &amp; Carvacho, 1983 and L. hendrickxi Anker, 2011 (Crustacea: Decapoda: Alpheidae) from the Pacific coast of Mexico. Zootaxa, 3835 (4), 573 - 582. http: // dx. doi. org / 10.11646 / zootaxa. 3835.4.8" type="journal article" year="2014">
Salgado-Barragán 
<emphasis id="B90DEAD9FFFDFF9A9EF9D979FC4EED01" box="[917,973,1326,1350]" italics="true" pageId="5" pageNumber="190">et al.</emphasis>
2014
</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A9977D978FAA2ED00" author="Anker" box="[1051,1313,1326,1351]" pageId="5" pageNumber="190" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
).
</paragraph>
<paragraph id="8BC636CBFFFDFF999DABD904FD94E95B" blockId="5.[151,1436,1146,2035]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190">
The new species can be easily separated from 
<taxonomicName id="4C794D48FFFDFF9A9FA2D905FCEAED2D" box="[718,873,1363,1386]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="axianassae">
<emphasis id="B90DEAD9FFFDFF9A9FA2D905FCEAED2D" box="[718,873,1363,1386]" italics="true" pageId="5" pageNumber="190">L. axianassae</emphasis>
</taxonomicName>
as well as its sister species 
<taxonomicName id="4C794D48FFFDFF9A99F3D905FACCED2D" box="[1183,1359,1362,1386]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="canterakintzi">
<emphasis id="B90DEAD9FFFDFF9A99F3D905FACCED2D" box="[1183,1359,1362,1386]" italics="true" pageId="5" pageNumber="190">L. canterakintzi</emphasis>
</taxonomicName>
by the bidentate tip of the pollex of the major chela; the presence of a stout proximal tooth on the cutting edge of the major chela dactylus; the absence of a conspicuously enlarged triangular tooth on the dactylus and pollex of the minor chela; the longer second article of the antennular peduncle (almost twice as long as wide in 
<taxonomicName id="4C794D48FFFDFF9A99F1D9E9FA97ED91" box="[1181,1300,1471,1494]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A99F1D9E9FA97ED91" box="[1181,1300,1471,1494]" italics="true" pageId="5" pageNumber="190">L. pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A9877D9E8FAF6ED90" box="[1307,1397,1470,1495]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A9877D9E8FAF6ED90" bold="true" box="[1307,1397,1470,1495]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
vs. 1.2–1.3 times as long as wide in 
<taxonomicName id="4C794D48FFFDFF9A9F76D9B5FD34EDBD" box="[538,695,1507,1530]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="axianassae">
<emphasis id="B90DEAD9FFFDFF9A9F76D9B5FD34EDBD" box="[538,695,1507,1530]" italics="true" pageId="5" pageNumber="190">L. axianassae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C794D48FFFDFF9A9F9FD9B5FC26EDBD" box="[755,933,1506,1530]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="canterakintzi">
<emphasis id="B90DEAD9FFFDFF9A9F9FD9B5FC26EDBD" box="[755,933,1506,1530]" italics="true" pageId="5" pageNumber="190">L. canterakintzi</emphasis>
</taxonomicName>
) (cf. 
<bibRefCitation id="EFE84B3AFFFDFF9A9E85D9B4FAABEDBC" author="Dworschak" box="[1001,1320,1506,1531]" pageId="5" pageNumber="190" refString="Dworschak, P. C. &amp; Coelho, V. R. (1999) On two alpheids from Araca (Sao Paulo, Brazil) with a description of a new species of Leptalpheus (Decapoda: Caridea: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 101 B, 475 - 488." type="journal article" year="1999">Dworschak &amp; Coelho 1999</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A985AD9B4FEBBEE58" author="Anker" pageId="5" pageNumber="190" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
). 
<taxonomicName id="4C794D48FFFDFF9A9C22DA50FDB8EE59" box="[334,571,1542,1566]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A9C22DA50FDB8EE59" box="[334,571,1542,1566]" italics="true" pageId="5" pageNumber="190">Leptalpheus pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A9F2ADA50FD27EE58" box="[582,676,1542,1567]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A9F2ADA50FD27EE58" bold="true" box="[582,676,1542,1567]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="4C794D48FFFDFF9A9E28DA51FC31EE59" box="[836,946,1543,1566]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azuero">
<emphasis id="B90DEAD9FFFDFF9A9E28DA51FC31EE59" box="[836,946,1543,1566]" italics="true" pageId="5" pageNumber="190">L. azuero</emphasis>
</taxonomicName>
by the much longer second article of the antennular peduncle (almost twice as long as wide in 
<taxonomicName id="4C794D48FFFDFF9A9E69DA7DFCFDEE05" box="[773,894,1579,1602]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A9E69DA7DFCFDEE05" box="[773,894,1579,1602]" italics="true" pageId="5" pageNumber="190">L. pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A9EE4DA7CFC67EE04" box="[904,996,1578,1603]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A9EE4DA7CFC67EE04" bold="true" box="[904,996,1578,1603]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
vs. as long as wide in 
<taxonomicName id="4C794D48FFFDFF9A9998DA7DFADCEE05" box="[1268,1375,1579,1602]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azuero">
<emphasis id="B90DEAD9FFFDFF9A9998DA7DFADCEE05" box="[1268,1375,1579,1602]" italics="true" pageId="5" pageNumber="190">L. azuero</emphasis>
</taxonomicName>
); the shorter, more appressed stylocerite, with the tip not exceeding the distal margin of the first article of the antennular peduncle (vs. exceeding this margin in 
<taxonomicName id="4C794D48FFFDFF9A9F20DA25FD35EECD" box="[588,694,1651,1674]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azuero">
<emphasis id="B90DEAD9FFFDFF9A9F20DA25FD35EECD" box="[588,694,1651,1674]" italics="true" pageId="5" pageNumber="190">L. azuero</emphasis>
</taxonomicName>
); and the absence of a spiniform seta on the ischia of the third and fourth pereiopods, which is present in 
<taxonomicName id="4C794D48FFFDFF9A9F38DAC1FD43EEE9" box="[596,704,1687,1710]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azuero">
<emphasis id="B90DEAD9FFFDFF9A9F38DAC1FD43EEE9" box="[596,704,1687,1710]" italics="true" pageId="5" pageNumber="190">L. azuero</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFE84B3AFFFDFF9A9F94DAC0FC07EEE8" author="Anker" box="[760,900,1686,1711]" pageId="5" pageNumber="190" refString="Anker, A. (2011) Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda). Zootaxa, 3041, 1 - 38." type="journal article" year="2011">Anker 2011</bibRefCitation>
). It may be rather easily distinguished from 
<taxonomicName id="4C794D48FFFDFF9A98E9DAC1FE8BEE95" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="corderoae">
<emphasis id="B90DEAD9FFFDFF9A98E9DAC1FE8BEE95" italics="true" pageId="5" pageNumber="190">L. corderoae</emphasis>
</taxonomicName>
, for instance, by the smooth ventral margin of the pollex and palm of the major chela, which is rugose or covered with tubercles in 
<taxonomicName id="4C794D48FFFDFF9A9CA1DA89FDE2EEB1" box="[461,609,1758,1782]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="corderoae">
<emphasis id="B90DEAD9FFFDFF9A9CA1DA89FDE2EEB1" box="[461,609,1758,1782]" italics="true" pageId="5" pageNumber="190">L. corderoae</emphasis>
</taxonomicName>
; and the absence of a conspicuously enlarged triangular tooth on the dactylus and pollex of the minor chela (cf. 
<bibRefCitation id="EFE84B3AFFFDFF9A9FEDDB54FC52EF5C" author="Salgado-Barragan" box="[641,977,1794,1819]" pageId="5" pageNumber="190" refString="Salgado-Barragan, J., Ayon-Parente, M. &amp; Hendrickx, M. E. (2014) A new species of Leptalpheus Williams, 1965 and new records of L. mexicanus Rios &amp; Carvacho, 1983 and L. hendrickxi Anker, 2011 (Crustacea: Decapoda: Alpheidae) from the Pacific coast of Mexico. Zootaxa, 3835 (4), 573 - 582. http: // dx. doi. org / 10.11646 / zootaxa. 3835.4.8" type="journal article" year="2014">
Salgado-Barragán 
<emphasis id="B90DEAD9FFFDFF9A9E38DB55FC0EEF5D" box="[852,909,1794,1818]" italics="true" pageId="5" pageNumber="190">et al.</emphasis>
2014
</bibRefCitation>
). In addition, 
<taxonomicName id="4C794D48FFFDFF9A991CDB55FB64EF5D" box="[1136,1255,1795,1818]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A991CDB55FB64EF5D" box="[1136,1255,1795,1818]" italics="true" pageId="5" pageNumber="190">L. pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A999CDB54FAC9EF5C" box="[1264,1354,1794,1819]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A999CDB54FAC9EF5C" bold="true" box="[1264,1354,1794,1819]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from 
<taxonomicName id="4C794D48FFFDFF9A9C19DB71FD92EF79" box="[373,529,1831,1854]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="axianassae">
<emphasis id="B90DEAD9FFFDFF9A9C19DB71FD92EF79" box="[373,529,1831,1854]" italics="true" pageId="5" pageNumber="190">L. axianassae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C794D48FFFDFF9A9F4EDB71FD57EF79" box="[546,724,1830,1854]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="canterakintzi">
<emphasis id="B90DEAD9FFFDFF9A9F4EDB71FD57EF79" box="[546,724,1830,1854]" italics="true" pageId="5" pageNumber="190">L. canterakintzi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C794D48FFFDFF9A9F89DB71FCD3EF79" box="[741,848,1831,1854]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="azuero">
<emphasis id="B90DEAD9FFFDFF9A9F89DB71FCD3EF79" box="[741,848,1831,1854]" italics="true" pageId="5" pageNumber="190">L. azuero</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C794D48FFFDFF9A9EE0DB71FB9EEF79" box="[908,1053,1830,1854]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="corderoae">
<emphasis id="B90DEAD9FFFDFF9A9EE0DB71FB9EEF79" box="[908,1053,1830,1854]" italics="true" pageId="5" pageNumber="190">L. corderoae</emphasis>
</taxonomicName>
by the tooth on the mesioventral carina of the first article of the antennular peduncle being simple and not having a protruding rounded lobe dorsally, as in the other four species (cf. 
<bibRefCitation id="EFE84B3AFFFDFF9A9F05DB38FC26EFC0" author="Dworschak" box="[617,933,1902,1927]" pageId="5" pageNumber="190" refString="Dworschak, P. C. &amp; Coelho, V. R. (1999) On two alpheids from Araca (Sao Paulo, Brazil) with a description of a new species of Leptalpheus (Decapoda: Caridea: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 101 B, 475 - 488." type="journal article" year="1999">Dworschak &amp; Coelho 1999</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A9EDDDB38FBB9EFC0" author="Anker" box="[945,1082,1902,1927]" pageId="5" pageNumber="190" refString="Anker, A. (2011) Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder &amp; Manning, 1986 (Crustacea, Decapoda). Zootaxa, 3041, 1 - 38." type="journal article" year="2011">Anker 2011</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A992BDB38FA14EFC0" author="Salgado-Barragan" box="[1095,1431,1902,1927]" pageId="5" pageNumber="190" refString="Salgado-Barragan, J., Ayon-Parente, M. &amp; Hendrickx, M. E. (2014) A new species of Leptalpheus Williams, 1965 and new records of L. mexicanus Rios &amp; Carvacho, 1983 and L. hendrickxi Anker, 2011 (Crustacea: Decapoda: Alpheidae) from the Pacific coast of Mexico. Zootaxa, 3835 (4), 573 - 582. http: // dx. doi. org / 10.11646 / zootaxa. 3835.4.8" type="journal article" year="2014">
Salgado-Barragán 
<emphasis id="B90DEAD9FFFDFF9A9876DB39FAD7EFC1" box="[1306,1364,1902,1926]" italics="true" pageId="5" pageNumber="190">et al.</emphasis>
2014
</bibRefCitation>
; 
<bibRefCitation id="EFE84B3AFFFDFF9A9DFBDBC4FE18EFEC" author="Anker" box="[151,411,1938,1963]" pageId="5" pageNumber="190" refString="Anker, A. &amp; Lazarus, J. F. (2015) Description of two new associated infaunal decapod crustaceans (Axianassidae and Alpheidae) from the tropical eastern Pacific. Papeis Avulsos de Zoologia, 55, 115 - 129. http: // dx. doi. org / 10.1590 / 0031 - 1049.2015.55.08" type="journal article" year="2015">Anker &amp; Lazarus 2015</bibRefCitation>
). Finally, 
<taxonomicName id="4C794D48FFFDFF9A9F64DBC5FDFDEFED" box="[520,638,1939,1962]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pereirai" status="sp. nov.">
<emphasis id="B90DEAD9FFFDFF9A9F64DBC5FDFDEFED" box="[520,638,1939,1962]" italics="true" pageId="5" pageNumber="190">L. pereirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57A2FFFDFF9A9FE8DBC4FD5DEFEC" box="[644,734,1938,1963]" pageId="5" pageNumber="190" rank="species">
<emphasis id="B90DEAD9FFFDFF9A9FE8DBC4FD5DEFEC" bold="true" box="[644,734,1938,1963]" pageId="5" pageNumber="190">sp. nov.</emphasis>
</taxonomicNameLabel>
can be separated from the three Indo-West Pacific species, viz. 
<taxonomicName id="4C794D48FFFDFF9A9DFBDBE1FEA3EF89" box="[151,288,1974,1998]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="pacificus">
<emphasis id="B90DEAD9FFFDFF9A9DFBDBE1FEA3EF89" box="[151,288,1974,1998]" italics="true" pageId="5" pageNumber="190">L. pacificus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C794D48FFFDFF9A9C58DBE1FE62EF89" box="[308,481,1974,1998]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B90DEAD9FFFDFF9A9C58DBE1FE62EF89" box="[308,481,1974,1998]" italics="true" pageId="5" pageNumber="190">L. denticulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C794D48FFFDFF9A9F4FDBE1FD4BEF89" box="[547,712,1974,1998]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="dworschaki">
<emphasis id="B90DEAD9FFFDFF9A9F4FDBE1FD4BEF89" box="[547,712,1974,1998]" italics="true" pageId="5" pageNumber="190">L. dworschaki</emphasis>
</taxonomicName>
, by the frontal margin having a triangular rostral projection (rounded in the Indo-West Pacific species); the presence of a stout proximal tooth on the cutting edge of the major chela dactylus; and the slightly rugose mesial margin of the major cheliped merus; specifically from 
<taxonomicName id="4C794D48FFFEFF99999ADCCFFA1FE8F7" box="[1270,1436,152,176]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B90DEAD9FFFEFF99999ADCCFFA1FE8F7" box="[1270,1436,152,176]" italics="true" pageId="6" pageNumber="191">L. denticulatus</emphasis>
</taxonomicName>
also by the absence of a small tooth on the mesial angle of the diaeresis of the uropodal exopod; and specifically from 
<taxonomicName id="4C794D48FFFEFF999DBBDCB7FEFAE8BF" box="[215,377,224,248]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="dworschaki">
<emphasis id="B90DEAD9FFFEFF999DBBDCB7FEFAE8BF" box="[215,377,224,248]" italics="true" pageId="6" pageNumber="191">L. dworschaki</emphasis>
</taxonomicName>
also by several obvious differences in the configuration of the armature of the major chela fingers (cf. 
<bibRefCitation id="EFE84B3AFFFEFF999C75DD52FD88E95B" author="Anker" box="[281,523,260,285]" pageId="6" pageNumber="191" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>