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<document id="FD09B4B76A12F14E0606934E862F9B79" ID-CLB-Dataset="28125" ID-DOI="10.11646/zootaxa.4568.3.12" ID-GBIF-Dataset="b84b1794-e082-40ba-9378-67ba33bf6e7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601586" ID-ZooBank="51088D66-B539-4C3B-9489-B246ACC8B95B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553172768616" checkinUser="plazi" docAuthor="Šumpich, Jan &amp; Jaroš, Josef" docDate="2019" docId="03D087BA3722AE5CFF04F98EFCA6FD01" docLanguage="en" docName="zootaxa.4568.3.12.pdf" docOrigin="Zootaxa 4568 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chrysoclista karsholti Šumpich &amp; Jaroš 2019, sp. n." docType="treatment" docUuid="DA8841CB-1832-4826-B4A0-0C5F6E05356F" docUuidSource="ZooBank" docVersion="9" lastPageNumber="591" masterDocId="FFE9FFC23723AE58FF93FFD4FFD0FFCC" masterDocTitle="Chrysoclista karsholti, sp. n., from Turkey, and a new record of C. germanica from central Europe (Lepidoptera: Elachistidae: Parametriotinae)" masterLastPageNumber="592" masterPageNumber="587" pageNumber="588" updateTime="1698703542916" updateUser="ExternalLinkService">
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<mods:title id="BEAC688722C4DE45AF1D69CEA452F954">Chrysoclista karsholti, sp. n., from Turkey, and a new record of C. germanica from central Europe (Lepidoptera: Elachistidae: Parametriotinae)</mods:title>
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<mods:namePart id="2AEAA7B4B62FF92ED6FEBB7ECE2BCEFE">Šumpich, Jan</mods:namePart>
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<mods:namePart id="2A9FB836C47E7DE89064FD5B7FE60C34">Jaroš, Josef</mods:namePart>
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<mods:date id="CDCAF2E8EBE2026060ED240E4824E8D3">2019</mods:date>
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<treatment id="03D087BA3722AE5CFF04F98EFCA6FD01" ID-DOI="http://doi.org/10.5281/zenodo.5930698" ID-GBIF-Taxon="154474964" ID-Zenodo-Dep="5930698" LSID="urn:lsid:zoobank.org:act:DA8841CB-1832-4826-B4A0-0C5F6E05356F" httpUri="http://treatment.plazi.org/id/03D087BA3722AE5CFF04F98EFCA6FD01" lastPageId="4" lastPageNumber="591" pageId="1" pageNumber="588">
<subSubSection id="C36365273722AE59FF04F98EFDB5F9B9" box="[151,613,1626,1653]" pageId="1" pageNumber="588" type="nomenclature">
<paragraph id="8BC636AC3722AE59FF04F98EFDB5F9B9" blockId="1.[151,613,1626,1686]" box="[151,613,1626,1653]" pageId="1" pageNumber="588">
<heading id="D08E81C03722AE59FF04F98EFDB5F9B9" bold="true" box="[151,613,1626,1653]" fontSize="11" level="1" pageId="1" pageNumber="588" reason="1">
<emphasis id="B90DEABE3722AE59FF04F98EFDB5F9B9" bold="true" box="[151,613,1626,1653]" pageId="1" pageNumber="588">
<taxonomicName id="4C794D2F3722AE59FF04F98EFDC6F9B9" authority="Sumpich" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[151,534,1626,1653]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="karsholti" status="sp. nov.">
<emphasis id="B90DEABE3722AE59FF04F98EFE70F9B8" bold="true" box="[151,416,1626,1652]" italics="true" pageId="1" pageNumber="588">Chrysoclista karsholti</emphasis>
Šumpich
</taxonomicName>
,
<taxonomicNameLabel id="A23E57C53722AE59FDB0F98FFDB5F9B9" box="[547,613,1627,1653]" pageId="1" pageNumber="588" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C36365273722AE59FF04F9A9FEC1F95A" box="[151,273,1661,1686]" pageId="1" pageNumber="588" type="description">
<paragraph id="8BC636AC3722AE59FF04F9A9FEC1F95A" blockId="1.[151,613,1626,1686]" box="[151,273,1661,1686]" pageId="1" pageNumber="588">
(
<figureCitation id="13422A293722AE59FF0CF9A9FED9F95A" box="[159,265,1661,1686]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[151,1435,245,1600]" captionTargetId="figure@2.[151,1435,245,1600]" captionTargetPageId="2" captionText="FIGURES 1–4. Chrysoclista species. 1. Holotype ♂, of Chrysoclista karsholti, Turkey. 2. Male genitalia of Chrysoclista karsholti. 3. Detail of the terminal part of phallus of Chrysoclista karsholti. 4. Chrysoclista germanica, ♂, Czech Republic." httpUri="https://zenodo.org/record/2601588/files/figure.png" pageId="1" pageNumber="588">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36365273722AE59FF04F911FAE0F8CD" pageId="1" pageNumber="588" type="materials_examined">
<paragraph id="8BC636AC3722AE59FF04F911FAE0F8CD" blockId="1.[151,1437,1733,1974]" pageId="1" pageNumber="588">
<emphasis id="B90DEABE3722AE59FF04F911FE99F912" bold="true" box="[151,329,1733,1758]" pageId="1" pageNumber="588">Type material.</emphasis>
<typeStatus id="54C2880E3722AE59FEC7F911FE6CF912" box="[340,444,1733,1758]" pageId="1" pageNumber="588" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7FFD253722AE59FE5BF912FE30F911" box="[456,480,1734,1757]" pageId="1" pageNumber="588" type="male">♂</specimenCount>
: ‘Anatolien | Kizilcahamam | leg. Pinker |
<date id="FFC7106C3722AE59FC4DF911FBB5F912" box="[990,1125,1733,1758]" pageId="1" pageNumber="588" value="1970-07-25">25.vii.1970</date>
’, ‘Photo / | Gen. prep. | J. Šumpich | 18/419’, ‘
<materialsCitation id="3B113CF13722AE59FEE4F93EFAFCF8CD" ID-GBIF-Occurrence="2013965654" box="[375,1324,1769,1794]" collectionCode="NHMW" pageId="1" pageNumber="588" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C2880E3722AE59FEE4F93EFDF3F8CE" box="[375,547,1770,1794]" pageId="1" pageNumber="588" type="holotype">HOLOTYPUS</typeStatus>
<specimenCount id="9D7FFD253722AE59FDB9F93EFDEFF8CD" box="[554,575,1770,1793]" pageId="1" pageNumber="588" type="male">♂</specimenCount>
|
<taxonomicName id="4C794D2F3722AE59FDC1F93DFC98F8CE" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[594,840,1769,1794]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="karsholti">Chrysoclista karsholti</taxonomicName>
| Šumpich, 2018’ [red label], coll.
<collectionCode id="ED68AE693722AE59FB5CF93EFAFCF8CD" box="[1231,1324,1770,1793]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="588" type="Museum">NHMW</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36365273722AE5AFF54F8D9FE7AF82A" lastPageId="2" lastPageNumber="589" pageId="1" pageNumber="588" type="description">
<paragraph id="8BC636AC3722AE59FF54F8D9FC3DF87A" blockId="1.[151,1437,1733,1974]" pageId="1" pageNumber="588">
<emphasis id="B90DEABE3722AE59FF54F8D9FE89F8EA" bold="true" box="[199,345,1805,1830]" pageId="1" pageNumber="588">Description.</emphasis>
Adult (
<figureCitation id="13422A293722AE59FE25F8D9FE2CF8EA" box="[438,508,1805,1830]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[151,1435,245,1600]" captionTargetId="figure@2.[151,1435,245,1600]" captionTargetPageId="2" captionText="FIGURES 1–4. Chrysoclista species. 1. Holotype ♂, of Chrysoclista karsholti, Turkey. 2. Male genitalia of Chrysoclista karsholti. 3. Detail of the terminal part of phallus of Chrysoclista karsholti. 4. Chrysoclista germanica, ♂, Czech Republic." httpUri="https://zenodo.org/record/2601588/files/figure.png" pageId="1" pageNumber="588">Fig. 1</figureCitation>
). Wingspan
<quantity id="4C819B493722AE59FD07F8DAFD39F8E9" box="[660,745,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="588" unit="mm" value="11.0">11 mm</quantity>
. Labial palpus narrow, short, slightly curved, covered with bronze scales on ventral side, and with silvery-grey on dorsal side, tip of third segment brown. Head and frons bronze with metallic shine. Forewing costa, dorsum, basal, apical and subapical areas brown, interspace in middle of wing light orange. Three dark spots with several silver scales in postbasal, tornal, and discal areas. Fringe and hindwing uniform chocolate brown, same shade as brown parts of forewing.
</paragraph>
<caption id="DF0666243721AE5AFF04F94CFAA7F901" httpUri="https://zenodo.org/record/2601588/files/figure.png" pageId="2" pageNumber="589" startId="2.[151,264,1688,1711]" targetBox="[151,1435,245,1600]" targetPageId="2">
<paragraph id="8BC636AC3721AE5AFF04F94CFAA7F901" blockId="2.[151,1436,1688,1741]" pageId="2" pageNumber="589">
<emphasis id="B90DEABE3721AE5AFF04F94CFEEEF962" bold="true" box="[151,318,1688,1711]" pageId="2" pageNumber="589">FIGURES 1–4.</emphasis>
<taxonomicName id="4C794D2F3721AE5AFEDBF94DFE18F962" authorityName="Stainton" authorityYear="1854" box="[328,456,1689,1710]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEABE3721AE5AFEDBF94DFE18F962" box="[328,456,1689,1710]" italics="true" pageId="2" pageNumber="589">Chrysoclista</emphasis>
</taxonomicName>
species. 1. Holotype ♂, of
<taxonomicName id="4C794D2F3721AE5AFD67F94DFC08F962" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[756,984,1689,1710]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3721AE5AFD67F94DFC08F962" box="[756,984,1689,1710]" italics="true" pageId="2" pageNumber="589">Chrysoclista karsholti</emphasis>
</taxonomicName>
, Turkey. 2. Male genitalia of
<taxonomicName id="4C794D2F3721AE5AFA8FF94DFF21F901" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3721AE5AFA8FF94DFF21F901" italics="true" pageId="2" pageNumber="589">Chrysoclista karsholti</emphasis>
</taxonomicName>
. 3. Detail of the terminal part of phallus of
<taxonomicName id="4C794D2F3721AE5AFD39F96CFC5BF901" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[682,907,1720,1741]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3721AE5AFD39F96CFC5BF901" box="[682,907,1720,1741]" italics="true" pageId="2" pageNumber="589">Chrysoclista karsholti</emphasis>
</taxonomicName>
. 4.
<taxonomicName id="4C794D2F3721AE5AFC23F96CFB72F901" authorityName="Sumpich &amp; Huemer" authorityYear="2016" box="[944,1186,1720,1741]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="B90DEABE3721AE5AFC23F96CFB72F901" box="[944,1186,1720,1741]" italics="true" pageId="2" pageNumber="589">Chrysoclista germanica</emphasis>
</taxonomicName>
, ♂, Czech Republic.
</paragraph>
</caption>
<paragraph id="8BC636AC3721AE5AFF54F920FB0FF8FD" blockId="2.[151,1437,1780,2022]" pageId="2" pageNumber="589">
Male genitalia (
<figureCitation id="13422A293721AE5AFEE8F920FE3AF8C1" box="[379,490,1780,1805]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1688,1711]" captionTargetBox="[151,1435,245,1600]" captionTargetId="figure@2.[151,1435,245,1600]" captionTargetPageId="2" captionText="FIGURES 1–4. Chrysoclista species. 1. Holotype ♂, of Chrysoclista karsholti, Turkey. 2. Male genitalia of Chrysoclista karsholti. 3. Detail of the terminal part of phallus of Chrysoclista karsholti. 4. Chrysoclista germanica, ♂, Czech Republic." httpUri="https://zenodo.org/record/2601588/files/figure.png" pageId="2" pageNumber="589">Figs. 2–3</figureCitation>
). Valva broad, slightly curved and elongated ventrally, rounded terminally. Phallus sligthly bent, without cornuti, and with group of approximately 15 smaller spines in terminal part.
</paragraph>
<paragraph id="8BC636AC3721AE5AFF54F8E8FDD5F899" blockId="2.[151,1437,1780,2022]" box="[199,517,1852,1877]" pageId="2" pageNumber="589">Female genitalia. Unknown.</paragraph>
<paragraph id="8BC636AC3721AE5AFF54F8B5FE7AF82A" blockId="2.[151,1437,1780,2022]" pageId="2" pageNumber="589">
<emphasis id="B90DEABE3721AE5AFF54F8B5FE58F8B6" bold="true" box="[199,392,1889,1914]" pageId="2" pageNumber="589">Molecular data.</emphasis>
BIN URI not stated previously. Process ID in BOLD: DEEUR1078-17. According to the phylogenetic tree,
<taxonomicName id="4C794D2F3721AE5AFEFFF851FE24F851" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[364,500,1925,1949]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3721AE5AFEFFF851FE24F851" box="[364,500,1925,1949]" italics="true" pageId="2" pageNumber="589">C. karsholti</emphasis>
</taxonomicName>
is sister to the genetically very close
<taxonomicName id="4C794D2F3721AE5AFC22F851FB9DF851" authorityName="Sumpich &amp; Huemer" authorityYear="2016" box="[945,1101,1925,1949]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="B90DEABE3721AE5AFC22F851FB9DF851" box="[945,1101,1925,1949]" italics="true" pageId="2" pageNumber="589">C. germanica</emphasis>
</taxonomicName>
with only 1,10 % difference between them (see
<figureCitation id="13422A293721AE5AFEE2F87DFE63F80E" box="[369,435,1961,1986]" captionStart="FIGURE 5" captionStartId="3.[151,250,1805,1827]" captionTargetBox="[161,1426,517,1771]" captionTargetId="figure@3.[151,1436,497,1783]" captionTargetPageId="3" captionText="FIGURE 5. Majority-rule consensus tree from Bayesian analysis based on the barcode fragment of all Chrysoclista samples. Posterior probabilities are provided only for splits indicating 0.95 or higher probability. Tip labels include Process IDs from BOLD and species names." httpUri="https://zenodo.org/record/2601592/files/figure.png" pageId="2" pageNumber="589">Fig. 5</figureCitation>
and
<tableCitation id="C6FB03173721AE5AFE78F87DFDEEF80D" box="[491,574,1961,1986]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. Mean average genetic distances between main phylogenetic clades defined in Fig. 5 which are corresponding with morphospecies in the genus Chrysoclista." pageId="2" pageNumber="589">Table 1</tableCitation>
). These two species are sister to
<taxonomicName id="4C794D2F3721AE5AFC24F87DFB93F80D" authorityName="Sumpich" authorityYear="2012" box="[951,1091,1961,1985]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="gabretica">
<emphasis id="B90DEABE3721AE5AFC24F87DFB93F80D" box="[951,1091,1961,1985]" italics="true" pageId="2" pageNumber="589">C. gabretica</emphasis>
</taxonomicName>
with genetic distances of 7.86 and 7.96 % respectively.
</paragraph>
</subSubSection>
<caption id="DF0666243720AE5BFF04FF4CFD59FF03" pageId="3" pageNumber="590">
<paragraph id="8BC636AC3720AE5BFF04FF4CFD59FF03" blockId="3.[151,1436,152,207]" pageId="3" pageNumber="590">
<emphasis id="B90DEABE3720AE5BFF04FF4CFEDAFF63" bold="true" box="[151,266,152,175]" pageId="3" pageNumber="590">TABLE 1.</emphasis>
Mean average genetic distances between main phylogenetic clades defined in Fig. 5 which are corresponding with morphospecies in the genus
<taxonomicName id="4C794D2F3720AE5BFE68FF63FD52FF03" authorityName="Stainton" authorityYear="1854" box="[507,642,183,207]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEABE3720AE5BFE68FF63FD52FF03" box="[507,642,183,207]" italics="true" pageId="3" pageNumber="590">Chrysoclista</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC636AC3720AE5BFE48FF3FFB10FE6A" pageId="3" pageNumber="590">
<table id="F979C40C372051A7FF0EFF3FFAE0FE6A" box="[157,1328,235,422]" gridcols="5" gridrows="5" pageId="3" pageNumber="590">
<tr id="354934EE372051A7FF0EFF3FFAE0FECC" box="[157,1328,235,256]" gridrow="0" pageId="3" pageNumber="590" rowspan-0="1">
<th id="76985D92372051A7FE48FF3FFD82FECC" box="[475,594,235,256]" gridcol="1" gridrow="0" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFE48FF3FFD82FECC" baseAuthorityName="Clerck" baseAuthorityYear="1759" box="[475,594,235,256]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="linneella">
<emphasis id="B90DEABE3720AE5BFE48FF3FFD82FECC" box="[475,594,235,256]" italics="true" pageId="3" pageNumber="590">C. linneella</emphasis>
</taxonomicName>
</th>
<th id="76985D92372051A7FD38FF3FFCE6FECC" box="[683,822,235,256]" gridcol="2" gridrow="0" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFD38FF3FFCE6FECC" authorityName="Fletcher" authorityYear="1936" box="[683,822,235,256]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="lathamella">
<emphasis id="B90DEABE3720AE5BFD38FF3FFCE6FECC" box="[683,822,235,256]" italics="true" pageId="3" pageNumber="590">C. lathamella</emphasis>
</taxonomicName>
</th>
<th id="76985D92372051A7FC01FF3FFBDFFECC" box="[914,1039,235,256]" gridcol="3" gridrow="0" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFC01FF3FFBDFFECC" authorityName="Sumpich" authorityYear="2012" box="[914,1039,235,256]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="gabretica">
<emphasis id="B90DEABE3720AE5BFC01FF3FFBDFFECC" box="[914,1039,235,256]" italics="true" pageId="3" pageNumber="590">C. gabretica</emphasis>
</taxonomicName>
</th>
<th id="76985D92372051A7FBEAFF3FFAE0FECC" box="[1145,1328,235,256]" gridcol="4" gridrow="0" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFBEAFF3FFB20FECC" authority="Šumpich &amp; Jaroš, 2019" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[1145,1264,235,256]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="karsholti" status="sp. nov.">
<emphasis id="B90DEABE3720AE5BFBEAFF3FFB20FECC" box="[1145,1264,235,256]" italics="true" pageId="3" pageNumber="590">C. karsholti</emphasis>
</taxonomicName>
<emphasis id="B90DEABE3720AE5BFB64FF3FFAE0FECC" bold="true" box="[1271,1328,235,256]" pageId="3" pageNumber="590">
<taxonomicNameLabel id="A23E57C53720AE5BFB64FF3FFAE0FECC" box="[1271,1328,235,256]" pageId="3" pageNumber="590" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="354934EE372051A7FF0EFEC0FAE0FEE6" box="[157,1328,276,298]" gridrow="1" pageId="3" pageNumber="590" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76985D92372051A7FF0EFEC0FE84FEE6" box="[157,340,276,298]" gridcol="0" gridrow="1" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFF0EFEC0FEF7FEE5" authorityName="Fletcher" authorityYear="1936" box="[157,295,276,297]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="lathamella">
<emphasis id="B90DEABE3720AE5BFF0EFEC0FEF7FEE5" box="[157,295,276,297]" italics="true" pageId="3" pageNumber="590">C. lathamella</emphasis>
</taxonomicName>
</th>
<td id="76985D92372051A7FE48FEC0FD82FEE6" box="[475,594,276,298]" gridcol="1" gridrow="1" pageId="3" pageNumber="590">8,94 %</td>
</tr>
<tr id="354934EE372051A7FF0EFEE9FAE0FE9F" box="[157,1328,317,339]" gridrow="2" pageId="3" pageNumber="590" rowspan-3="1" rowspan-4="1">
<th id="76985D92372051A7FF0EFEE9FE84FE9F" box="[157,340,317,339]" gridcol="0" gridrow="2" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFF0EFEE9FECBFE9E" authorityName="Sumpich" authorityYear="2012" box="[157,283,317,338]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="gabretica">
<emphasis id="B90DEABE3720AE5BFF0EFEE9FECBFE9E" box="[157,283,317,338]" italics="true" pageId="3" pageNumber="590">C. gabretica</emphasis>
</taxonomicName>
</th>
<td id="76985D92372051A7FE48FEE9FD82FE9F" box="[475,594,317,339]" gridcol="1" gridrow="2" pageId="3" pageNumber="590">8,19 %</td>
<td id="76985D92372051A7FD38FEE9FCE6FE9F" box="[683,822,317,339]" gridcol="2" gridrow="2" pageId="3" pageNumber="590">9,31 %</td>
</tr>
<tr id="354934EE372051A7FF0EFEB2FAE0FEB0" box="[157,1328,358,380]" gridrow="3" pageId="3" pageNumber="590" rowspan-4="1">
<th id="76985D92372051A7FF0EFEB2FE84FEB0" box="[157,340,358,380]" gridcol="0" gridrow="3" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFF0EFEB3FEC5FEB0" authority="Šumpich &amp; Jaroš, 2019" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[157,277,359,380]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="karsholti" status="sp. nov.">
<emphasis id="B90DEABE3720AE5BFF0EFEB3FEC5FEB0" box="[157,277,359,380]" italics="true" pageId="3" pageNumber="590">C. karsholti</emphasis>
</taxonomicName>
<emphasis id="B90DEABE3720AE5BFE8FFEB3FE84FEB0" bold="true" box="[284,340,359,380]" pageId="3" pageNumber="590">
<taxonomicNameLabel id="A23E57C53720AE5BFE8FFEB3FE84FEB0" box="[284,340,359,380]" pageId="3" pageNumber="590" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="76985D92372051A7FE48FEB2FD82FEB0" box="[475,594,358,380]" gridcol="1" gridrow="3" pageId="3" pageNumber="590">11,47 %</td>
<td id="76985D92372051A7FD38FEB2FCE6FEB0" box="[683,822,358,380]" gridcol="2" gridrow="3" pageId="3" pageNumber="590">11,34 %</td>
<td id="76985D92372051A7FC01FEB2FBDFFEB0" box="[914,1039,358,380]" gridcol="3" gridrow="3" pageId="3" pageNumber="590">7,96 %</td>
</tr>
<tr id="354934EE372051A7FF0EFE44FAE0FE6A" box="[157,1328,400,422]" gridrow="4" pageId="3" pageNumber="590">
<th id="76985D92372051A7FF0EFE44FE84FE6A" box="[157,340,400,422]" gridcol="0" gridrow="4" pageId="3" pageNumber="590">
<taxonomicName id="4C794D2F3720AE5BFF0EFE44FEF7FE6A" authorityName="Sumpich &amp; Huemer" authorityYear="2016" box="[157,295,400,422]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="B90DEABE3720AE5BFF0EFE44FEF7FE6A" box="[157,295,400,422]" italics="true" pageId="3" pageNumber="590">C. germanica</emphasis>
</taxonomicName>
</th>
<td id="76985D92372051A7FE48FE44FD82FE6A" box="[475,594,400,422]" gridcol="1" gridrow="4" pageId="3" pageNumber="590">11,86 %</td>
<td id="76985D92372051A7FD38FE44FCE6FE6A" box="[683,822,400,422]" gridcol="2" gridrow="4" pageId="3" pageNumber="590">11,56 %</td>
<td id="76985D92372051A7FC01FE44FBDFFE6A" box="[914,1039,400,422]" gridcol="3" gridrow="4" pageId="3" pageNumber="590">7,86 %</td>
<td id="76985D92372051A7FBEAFE44FAE0FE6A" box="[1145,1328,400,422]" gridcol="4" gridrow="4" pageId="3" pageNumber="590">1,10 %</td>
</tr>
</table>
</paragraph>
<caption id="DF0666243720AE5BFF04F8D9FE74F8AC" httpUri="https://zenodo.org/record/2601592/files/figure.png" pageId="3" pageNumber="590" startId="3.[151,250,1805,1827]" targetBox="[161,1426,517,1771]" targetPageId="3">
<paragraph id="8BC636AC3720AE5BFF04F8D9FE74F8AC" blockId="3.[151,1437,1805,1888]" pageId="3" pageNumber="590">
<emphasis id="B90DEABE3720AE5BFF04F8D9FEC4F8EE" bold="true" box="[151,276,1805,1827]" pageId="3" pageNumber="590">FIGURE 5.</emphasis>
Majority-rule consensus tree from Bayesian analysis based on the barcode fragment of all
<taxonomicName id="4C794D2F3720AE5BFB2FF8D9FAECF8EE" authorityName="Stainton" authorityYear="1854" box="[1212,1340,1805,1826]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEABE3720AE5BFB2FF8D9FAECF8EE" box="[1212,1340,1805,1826]" italics="true" pageId="3" pageNumber="590">Chrysoclista</emphasis>
</taxonomicName>
samples. Posterior probabilities are provided only for splits indicating 0.95 or higher probability. Tip labels include Process IDs from BOLD and species names.
</paragraph>
</caption>
<subSubSection id="C36365273720AE5CFF54F858FB85FDD4" lastPageId="4" lastPageNumber="591" pageId="3" pageNumber="590" type="diagnosis">
<paragraph id="8BC636AC3720AE5CFF54F858FB85FDD4" blockId="3.[151,1436,1932,2030]" lastBlockId="4.[151,1437,151,717]" lastPageId="4" lastPageNumber="591" pageId="3" pageNumber="590">
<emphasis id="B90DEABE3720AE5BFF54F858FEEAF869" bold="true" box="[199,314,1932,1957]" pageId="3" pageNumber="590">Diagnosis</emphasis>
.
<taxonomicName id="4C794D2F3720AE5BFEDBF859FDEFF869" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[328,575,1933,1957]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3720AE5BFEDBF859FDEFF869" box="[328,575,1933,1957]" italics="true" pageId="3" pageNumber="590">Chrysoclista karsholti</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C794D2F3720AE5BFD2DF859FC99F869" authorityName="Sumpich" authorityYear="2012" box="[702,841,1933,1957]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="gabretica">
<emphasis id="B90DEABE3720AE5BFD2DF859FC99F869" box="[702,841,1933,1957]" italics="true" pageId="3" pageNumber="590">C. gabretica</emphasis>
</taxonomicName>
; the orange scales of the forewing of the latter are of the same shade as those of
<taxonomicName id="4C794D2F3720AE5BFE5AF865FD9DF805" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[457,589,1969,1993]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3720AE5BFE5AF865FD9DF805" box="[457,589,1969,1993]" italics="true" pageId="3" pageNumber="590">C. karsholti</emphasis>
</taxonomicName>
, but in
<taxonomicName id="4C794D2F3720AE5BFD37F865FCFFF805" authorityName="Sumpich" authorityYear="2012" box="[676,815,1969,1993]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="gabretica">
<emphasis id="B90DEABE3720AE5BFD37F865FCFFF805" box="[676,815,1969,1993]" italics="true" pageId="3" pageNumber="590">C. gabretica</emphasis>
</taxonomicName>
the orange area reaches the dorsal margin (terminating before the margin in
<taxonomicName id="4C794D2F3720AE5BFE10F801FDD8F821" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[387,520,2005,2029]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3720AE5BFE10F801FDD8F821" box="[387,520,2005,2029]" italics="true" pageId="3" pageNumber="590">C. karsholti</emphasis>
</taxonomicName>
). Also, there is a conspicuous basal dark logitudinal streak in
<taxonomicName id="4C794D2F3720AE5BFB56F801FA80F821" authorityName="Sumpich" authorityYear="2012" box="[1221,1360,2005,2029]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="gabretica">
<emphasis id="B90DEABE3720AE5BFB56F801FA80F821" box="[1221,1360,2005,2029]" italics="true" pageId="3" pageNumber="590">C. gabretica</emphasis>
</taxonomicName>
that is indistinct in
<emphasis id="B90DEABE3727AE5CFEB9FF4CFE6AFF7C" box="[298,442,152,176]" italics="true" pageId="4" pageNumber="591">
<taxonomicName id="4C794D2F3727AE5CFEB9FF4CFE66FF7C" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[298,438,152,176]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="karsholti">C. karsholti</taxonomicName>
.
</emphasis>
The genitalia of these two species are very similar but the valvae are rounded in
<emphasis id="B90DEABE3727AE5CFA11FF4CFED8FF18" italics="true" pageId="4" pageNumber="591">
<taxonomicName id="4C794D2F3727AE5CFA11FF4CFED4FF18" authorityName="Sumpich" authorityYear="2012" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="gabretica">C. gabretica</taxonomicName>
,
</emphasis>
whereas they are distinctly elongated dorsoventrally in
<taxonomicName id="4C794D2F3727AE5CFC17FF68FBDAFF18" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[900,1034,188,212]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3727AE5CFC17FF68FBDAFF18" box="[900,1034,188,212]" italics="true" pageId="4" pageNumber="591">C. karsholti</emphasis>
</taxonomicName>
. The shape of the anellus lobes are distinct in both species.
<taxonomicName id="4C794D2F3727AE5CFE3AFF34FD73FF34" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[425,675,224,248]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3727AE5CFE3AFF34FD73FF34" box="[425,675,224,248]" italics="true" pageId="4" pageNumber="591">Chrysoclista karsholti</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="4C794D2F3727AE5CFCFDFF34FBD8FF3B" authorityName="Sumpich &amp; Huemer" authorityYear="2016" box="[878,1032,224,247]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="B90DEABE3727AE5CFCFDFF34FBD8FF3B" box="[878,1032,224,247]" italics="true" pageId="4" pageNumber="591">C. germanica</emphasis>
</taxonomicName>
, but in the latter species the orange area is red-orange, and the brown colour is darker. In the male genitalia, the main differences are the shapes of the valvae (much more elongated in
<taxonomicName id="4C794D2F3727AE5CFD85FEFCFD62FEF3" authorityName="Sumpich &amp; Huemer" authorityYear="2016" box="[534,690,296,319]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="B90DEABE3727AE5CFD85FEFCFD62FEF3" box="[534,690,296,319]" italics="true" pageId="4" pageNumber="591">C. germanica</emphasis>
</taxonomicName>
), and the size and number of spines in the terminal part of ther phallus (smaller and fewer in
<taxonomicName id="4C794D2F3727AE5CFE79FE98FDBFFEA8" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[490,623,332,356]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3727AE5CFE79FE98FDBFFEA8" box="[490,623,332,356]" italics="true" pageId="4" pageNumber="591">C. karsholti</emphasis>
</taxonomicName>
).
<taxonomicName id="4C794D2F3727AE5CFD14FE98FC50FEA8" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[647,896,332,356]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3727AE5CFD14FE98FC50FEA8" box="[647,896,332,356]" italics="true" pageId="4" pageNumber="591">Chrysoclista karsholti</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C794D2F3727AE5CFBFFFE98FAD6FEA8" authorityName="Sumpich &amp; Huemer" authorityYear="2016" box="[1132,1286,332,356]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="B90DEABE3727AE5CFBFFFE98FAD6FEA8" box="[1132,1286,332,356]" italics="true" pageId="4" pageNumber="591">C. germanica</emphasis>
</taxonomicName>
based on the DNA barcode sequences. We cannot exclude the presence of cryptic diversity in this genus as was indicated by
<bibRefCitation id="EFE84B5D3727AE5CFF04FE40FE1BFE61" author="Sumpich, J. &amp; Huemer, P." box="[151,459,404,429]" pageId="4" pageNumber="591" pagination="785 - 788" refId="ref2914" refString="Sumpich, J. &amp; Huemer, P. (2016) Chrysoclista germanica sp. nov. and C. gabretica stat. rev., with an updated checklist of the genus (Lepidoptera, Elachistidae). Acta Entomologica Musei Nationalis Pragae, 54, 785 - 788." type="journal article" year="2016">Šumpich &amp; Huemer (2016)</bibRefCitation>
in which a similar dataset was interpreted in detail. For a clarification of this problem, additional material for genetic and morphological analysis is needed. Because the description of the new species is based on a single specimen, it is possible that
<taxonomicName id="4C794D2F3727AE5CFD47FE08FC8FFE38" authorityName="Šumpich &amp; Jaroš" authorityYear="2019" box="[724,863,476,500]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="karsholti">
<emphasis id="B90DEABE3727AE5CFD47FE08FC8FFE38" box="[724,863,476,500]" italics="true" pageId="4" pageNumber="591">C. karsholti</emphasis>
</taxonomicName>
merely represents variation with
<taxonomicName id="4C794D2F3727AE5CFB65FE08FA45FE38" authorityName="Sumpich &amp; Huemer" authorityYear="2016" box="[1270,1429,476,500]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="B90DEABE3727AE5CFB65FE08FA45FE38" box="[1270,1429,476,500]" italics="true" pageId="4" pageNumber="591">C. germanica</emphasis>
</taxonomicName>
. However, evidence from forewing pattern and genital morphology suggest otherwise.
</paragraph>
</subSubSection>
<subSubSection id="C36365273727AE5CFF54FDF0FAC3FD48" pageId="4" pageNumber="591" type="distribution">
<paragraph id="8BC636AC3727AE5CFF54FDF0FCFAFDF1" blockId="4.[151,1437,151,717]" box="[199,810,548,573]" pageId="4" pageNumber="591">
<emphasis id="B90DEABE3727AE5CFF54FDF0FE89FDF1" bold="true" box="[199,345,548,573]" pageId="4" pageNumber="591">Distribution</emphasis>
. The species is known only from
<collectingCountry id="F36E763C3727AE5CFD46FDF0FCF7FDF1" box="[725,807,548,573]" name="Turkey" pageId="4" pageNumber="591">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="8BC636AC3727AE5CFF54FD93FAC3FD48" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="591">
<emphasis id="B90DEABE3727AE5CFF54FD93FE9AFDAC" bold="true" box="[199,330,583,608]" pageId="4" pageNumber="591">Bionomics.</emphasis>
The early stages are unknown. The adult was collected in July. The exact collecting locality of the
<typeStatus id="54C2880E3727AE5CFF04FDB8FF29FD49" box="[151,249,620,645]" pageId="4" pageNumber="591" type="holotype">holotype</typeStatus>
is unknown - Kizilcahamam is located in an area with altitudes between
<quantity id="4C819B493727AE5CFBBFFDB8FB51FD48" box="[1068,1153,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="591" unit="m" value="1000.0">1000 m</quantity>
and
<quantity id="4C819B493727AE5CFB24FDB8FADFFD48" box="[1207,1295,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="591" unit="m" value="1400.0">1400 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36365273727AE5CFF54FD5BFCA6FD01" pageId="4" pageNumber="591" type="etymology">
<paragraph id="8BC636AC3727AE5CFF54FD5BFCA6FD01" blockId="4.[151,1437,151,717]" pageId="4" pageNumber="591">
<emphasis id="B90DEABE3727AE5CFF54FD5BFE9EFD64" bold="true" box="[199,334,655,680]" pageId="4" pageNumber="591">Etymology.</emphasis>
The name of the new species is a patronym for Ole Karsholt, who has contributed significantly to the research of gelechiioid moths, including
<taxonomicName id="4C794D2F3727AE5CFD1AFD60FCC6FD00" authorityName="Stainton" authorityYear="1854" box="[649,790,692,716]" class="Insecta" family="Agonoxenidae" genus="Chrysoclista" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEABE3727AE5CFD1AFD60FCC6FD00" box="[649,790,692,716]" italics="true" pageId="4" pageNumber="591">Chrysoclista</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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