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<document id="4DBF084DF2E428B473CD20C55297D039" ID-DOI="10.5281/zenodo.185641" ID-GBIF-Dataset="59926166-687d-44cd-b53d-ead63b598ac4" ID-ISSN="1175-5326" ID-Zenodo-Dep="185641" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459998664912" checkinUser="plazi" docAuthor="Evenhuis, Neal L." docDate="2009" docId="03D0879FA714FFE0FF27576DFB262FA1" docLanguage="en" docName="zt02004p048.pdf" docOrigin="Zootaxa 2004" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Campsicnemus gladiator" docType="treatment" docVersion="9" lastPageNumber="29" masterDocId="FFE9FFE7A717FFE4FFB05603FFD52D57" masterDocTitle="Review of Campsicnemus (Diptera: Dolichopodidae) of the Marquesas, French Polynesia, with description of four new species groups" masterLastPageNumber="48" masterPageNumber="25" pageNumber="28" updateTime="1698233059185" updateUser="plazi">
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<mods:title id="BD82E7238E331AD45DD4D672B1FFF3F3">Review of Campsicnemus (Diptera: Dolichopodidae) of the Marquesas, French Polynesia, with description of four new species groups</mods:title>
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<mods:namePart id="DD23644EBE0D28D1A2EE2CF2BB735250">Evenhuis, Neal L.</mods:namePart>
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<mods:date id="2414901F2D6BE7F35F950458AB1BD6BE">2009</mods:date>
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<treatment id="03D0879FA714FFE0FF27576DFB262FA1" ID-DOI="http://doi.org/10.5281/zenodo.6217119" ID-GBIF-Taxon="119380583" ID-Zenodo-Dep="6217119" LSID="urn:lsid:plazi:treatment:03D0879FA714FFE0FF27576DFB262FA1" httpUri="http://treatment.plazi.org/id/03D0879FA714FFE0FF27576DFB262FA1" lastPageId="4" lastPageNumber="29" pageId="3" pageNumber="28">
<subSubSection id="C3636502A714FFE7FF27576DFE032CFC" pageId="3" pageNumber="28" type="nomenclature">
<paragraph id="8BC63689A714FFE7FF27576DFE5B2CDF" blockId="3.[151,470,366,427]" box="[151,398,366,392]" pageId="3" pageNumber="28">
<heading id="D08E81E5A714FFE7FF27576DFE5B2CDF" bold="true" box="[151,398,366,392]" fontSize="11" level="1" pageId="3" pageNumber="28" reason="1">
<emphasis id="B90DEA9BA714FFE7FF27576DFE5B2CDF" bold="true" box="[151,398,366,392]" pageId="3" pageNumber="28">
The
<taxonomicName id="4C794D0AA714FFE7FF7F576DFEE92CDF" ID-CoL="8BBD8" box="[207,316,366,392]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B90DEA9BA714FFE7FF7F576DFEE92CDF" bold="true" box="[207,316,366,392]" italics="true" pageId="3" pageNumber="28">gladiator</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC63689A714FFE7FF275792FE032CFC" blockId="3.[151,470,366,427]" box="[151,470,401,427]" pageId="3" pageNumber="28">
(Marquesan
<taxonomicName id="4C794D0AA714FFE7FE9A5792FE472CFC" box="[298,402,401,427]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="gladiator">gladiator</taxonomicName>
flies)
</paragraph>
</subSubSection>
<subSubSection id="C3636502A714FFE7FF2757DDFF0C2F11" pageId="3" pageNumber="28" type="diagnosis">
<paragraph id="8BC63689A714FFE7FF2757DDFF0C2F11" blockId="3.[151,1436,478,582]" pageId="3" pageNumber="28">
<emphasis id="B90DEA9BA714FFE7FF2757DDFEC72CAF" bold="true" box="[151,274,478,504]" pageId="3" pageNumber="28">Diagnosis</emphasis>
. The group is characterized by the male fore coxa possessing a cluster of strong black setae that appear fused together resembling a thorn (
<figureCitation id="13422A0CA714FFE7FD205406FD0F2F48" box="[656,730,517,543]" captionStart="FIGURE 2" captionStartId="3.[151,254,1499,1523]" captionTargetBox="[360,1226,637,1476]" captionTargetId="figure@3.[360,1227,637,1476]" captionTargetPageId="3" captionText="FIGURE 2. Campsicnemus gladiator, n. sp., male fore coxa, lateral view, showing cluster of setae." httpUri="https://zenodo.org/record/185643/files/figure.png" pageId="3" pageNumber="28">Fig. 2</figureCitation>
). No other species of
<taxonomicName id="4C794D0AA714FFE7FC505406FB462F49" box="[992,1171,517,542]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA9BA714FFE7FC505406FB462F49" box="[992,1171,517,542]" italics="true" pageId="3" pageNumber="28">Campsicnemus</emphasis>
</taxonomicName>
possess these “fused” setae.
</paragraph>
</subSubSection>
<caption id="DF066601A714FFE7FF2753D8FB6828A4" box="[151,1213,1499,1523]" httpUri="https://zenodo.org/record/185643/files/figure.png" pageId="3" pageNumber="28" targetBox="[360,1226,637,1476]" targetPageId="3">
<paragraph id="8BC63689A714FFE7FF2753D8FB6828A4" blockId="3.[151,1213,1499,1523]" box="[151,1213,1499,1523]" pageId="3" pageNumber="28">
<emphasis id="B90DEA9BA714FFE7FF2753D8FEC428A4" bold="true" box="[151,273,1499,1523]" pageId="3" pageNumber="28">FIGURE 2</emphasis>
.
<taxonomicName id="4C794D0AA714FFE7FEAF53DFFDFC28A4" box="[287,553,1500,1523]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="gladiator" status="sp. nov.">
<emphasis id="B90DEA9BA714FFE7FEAF53DFFDFC28A4" box="[287,553,1500,1523]" italics="true" pageId="3" pageNumber="28">Campsicnemus gladiator</emphasis>
</taxonomicName>
,
<emphasis id="B90DEA9BA714FFE7FD8753D8FDA628A4" bold="true" box="[567,627,1499,1523]" pageId="3" pageNumber="28">
<taxonomicNameLabel id="A23E57E0A714FFE7FD8753D8FDA628A4" box="[567,627,1499,1523]" pageId="3" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male fore coxa, lateral view, showing cluster of setae.
</paragraph>
</caption>
<caption id="DF066601A714FFE7FF2751B6FDAD2AA4" httpUri="https://zenodo.org/record/185644/files/figure.png" pageId="3" pageNumber="28" targetBox="[321,1285,1605,1965]" targetPageId="3">
<paragraph id="8BC63689A714FFE7FF2751B6FDAD2AA4" blockId="3.[151,1436,1973,2035]" pageId="3" pageNumber="28">
<emphasis id="B90DEA9BA714FFE7FF2751B6FEE92A9A" bold="true" box="[151,316,1973,1997]" pageId="3" pageNumber="28">FIGURES 36</emphasis>
.
<taxonomicName id="4C794D0AA714FFE7FEF951B5FD812A9A" box="[329,596,1974,1997]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B90DEA9BA714FFE7FEF951B5FD812A9A" box="[329,596,1974,1997]" italics="true" pageId="3" pageNumber="28">Campsicnemus gladiator</emphasis>
</taxonomicName>
group, male mid basitarsi.
<emphasis id="B90DEA9BA714FFE7FCC651B6FC562A9A" bold="true" box="[886,899,1973,1997]" pageId="3" pageNumber="28">3</emphasis>
.
<taxonomicName id="4C794D0AA714FFE7FC2151B5FBC72A9A" box="[913,1042,1974,1997]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="gladiator" status="sp. nov.">
<emphasis id="B90DEA9BA714FFE7FC2151B5FBC72A9A" box="[913,1042,1974,1997]" italics="true" pageId="3" pageNumber="28">C. gladiator</emphasis>
</taxonomicName>
,
<emphasis id="B90DEA9BA714FFE7FB9051B6FBBA2A9A" bold="true" box="[1056,1135,1973,1997]" pageId="3" pageNumber="28">
<taxonomicNameLabel id="A23E57E0A714FFE7FB9051B6FB8E2A9A" box="[1056,1115,1973,1997]" pageId="3" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
4
</emphasis>
.
<taxonomicName id="4C794D0AA714FFE7FBCD51B5FA572A9A" authority="Evenhuis." authorityName="Evenhuis." box="[1149,1410,1974,1997]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="limnobates">
<emphasis id="B90DEA9BA714FFE7FBCD51B5FAC42A9A" box="[1149,1297,1974,1997]" italics="true" pageId="3" pageNumber="28">C. limnobates</emphasis>
Evenhuis.
</taxonomicName>
<emphasis id="B90DEA9BA714FFE7FA3851B6FA402A9A" bold="true" box="[1416,1429,1973,1997]" pageId="3" pageNumber="28">5</emphasis>
.
<taxonomicName id="4C794D0AA714FFE7FF2751DFFE9B2AA4" authority="Parent." authorityName="Parent." box="[151,334,2012,2035]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="scurra">
<emphasis id="B90DEA9BA714FFE7FF2751DFFF292AA4" box="[151,252,2012,2035]" italics="true" pageId="3" pageNumber="28">C. scurra</emphasis>
Parent.
</taxonomicName>
<emphasis id="B90DEA9BA714FFE7FEE451D8FEB42AA4" bold="true" box="[340,353,2011,2035]" pageId="3" pageNumber="28">6</emphasis>
.
<taxonomicName id="4C794D0AA714FFE7FEDE51DFFDAD2AA4" authority="Evenhuis." authorityName="Evenhuis." box="[366,632,2012,2035]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="uncleremus">
<emphasis id="B90DEA9BA714FFE7FEDE51DFFDD22AA4" box="[366,519,2012,2035]" italics="true" pageId="3" pageNumber="28">C. uncleremus</emphasis>
Evenhuis.
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="C3636502A713FFE0FF765694FB492F63" pageId="4" pageNumber="29" type="discussion">
<paragraph id="8BC63689A713FFE0FF765694FC7F2C71" blockId="4.[151,1436,151,758]" pageId="4" pageNumber="29">
<emphasis id="B90DEA9BA713FFE0FF765694FEED2DE6" bold="true" box="[198,312,151,177]" pageId="4" pageNumber="29">Remarks</emphasis>
. This group is comprised of water-skating species of
<taxonomicName id="4C794D0AA713FFE0FC01569BFBB62DE6" box="[945,1123,152,177]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA9BA713FFE0FC01569BFBB62DE6" box="[945,1123,152,177]" italics="true" pageId="4" pageNumber="29">Campsicnemus</emphasis>
</taxonomicName>
in the Marquesas. Species appear to be island endemics with one species per island harboring swift running water. Although no species of
<taxonomicName id="4C794D0AA713FFE0FF0756E6FEBC2DA9" box="[183,361,229,254]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA9BA713FFE0FF0756E6FEBC2DA9" box="[183,361,229,254]" italics="true" pageId="4" pageNumber="29">Campsicnemus</emphasis>
</taxonomicName>
are currently known from there, it is most likely that another species of this group will be found on Tahuata when water sources there are searched diligently.
</paragraph>
<paragraph id="8BC63689A713FFE0FF765731FF362CB0" blockId="4.[151,1436,151,758]" pageId="4" pageNumber="29">
Species of the
<taxonomicName id="4C794D0AA713FFE0FECA5731FE352C1C" box="[378,480,306,331]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="englundi">
<emphasis id="B90DEA9BA713FFE0FECA5731FE352C1C" box="[378,480,306,331]" italics="true" pageId="4" pageNumber="29">englundi</emphasis>
</taxonomicName>
group another water-skating species group from Fatu Hiva and Hiva Oa in the Marquesas, are similar in appearance to species of the
<taxonomicName id="4C794D0AA713FFE0FC81575AFC742C25" box="[817,929,345,370]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B90DEA9BA713FFE0FC81575AFC742C25" box="[817,929,345,370]" italics="true" pageId="4" pageNumber="29">gladiator</emphasis>
</taxonomicName>
subgroup by virtue of the extremely long hairs on the hind femur, but the lack of the fore coxal thorn-like setal cluster (typical of the
<taxonomicName id="4C794D0AA713FFE0FB0C5783FAFC2CCE" box="[1212,1321,384,409]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B90DEA9BA713FFE0FB0C5783FAFC2CCE" box="[1212,1321,384,409]" italics="true" pageId="4" pageNumber="29">gladiator</emphasis>
</taxonomicName>
group) in combination with the bizarre shape of the mid femur with strong setae easily separate them from the
<taxonomicName id="4C794D0AA713FFE0FA9F57A4FA492C97" box="[1327,1436,423,448]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B90DEA9BA713FFE0FA9F57A4FA492C97" box="[1327,1436,423,448]" italics="true" pageId="4" pageNumber="29">gladiator</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8BC63689A713FFE0FF7657F0FB492F63" blockId="4.[151,1436,151,758]" pageId="4" pageNumber="29">
<emphasis id="B90DEA9BA713FFE0FF7657F0FE5D2F5A" bold="true" box="[198,392,499,525]" pageId="4" pageNumber="29">Common name</emphasis>
: The name “Marquesan
<taxonomicName id="4C794D0AA713FFE0FD0357F7FCCE2F59" box="[691,795,500,526]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="gladiator">gladiator</taxonomicName>
flies” refers to the characteristic thorn-like cluster of setae on the fore coxa that are reminiscent of the spiked weaponry of Roman gladiators.
</paragraph>
</subSubSection>
<subSubSection id="C3636502A713FFE0FF765443FB462FD5" pageId="4" pageNumber="29" type="description">
<paragraph id="8BC63689A713FFE0FF765443FE4E2F0C" blockId="4.[151,1436,151,758]" box="[198,411,576,603]" pageId="4" pageNumber="29">
<emphasis id="B90DEA9BA713FFE0FF765443FE472F0D" bold="true" box="[198,402,576,602]" pageId="4" pageNumber="29">Included species</emphasis>
:
</paragraph>
<paragraph id="8BC63689A713FFE0FF755464FB462FD5" blockId="4.[151,1436,151,758]" box="[197,1171,615,642]" pageId="4" pageNumber="29">
<taxonomicName id="4C794D0AA713FFE0FF755464FE732FD5" authority="Evenhuis" authorityName="Evenhuis" box="[197,422,615,642]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="gladiator" status="sp. nov.">
<emphasis id="B90DEA9BA713FFE0FF755464FEE72FD6" bold="true" box="[197,306,615,641]" italics="true" pageId="4" pageNumber="29">gladiator</emphasis>
Evenhuis
</taxonomicName>
,
<taxonomicNameLabel id="A23E57E0A713FFE0FE055464FE232FD5" box="[437,502,615,642]" pageId="4" pageNumber="29" rank="species">
<emphasis id="B90DEA9BA713FFE0FE055464FE3A2FD6" bold="true" box="[437,495,615,641]" pageId="4" pageNumber="29">n. sp</emphasis>
.
</taxonomicNameLabel>
(Marquesas [Ua Pou];
<typeStatus id="54C2882BA713FFE0FCB8546BFCBB2FD5" box="[776,878,616,642]" pageId="4" pageNumber="29" type="holotype">holotype</typeStatus>
ɗ in BPBM, examined).
</paragraph>
</subSubSection>
<subSubSection id="C3636502A713FFE0FF75548DFB262FA1" pageId="4" pageNumber="29" type="reference_group">
<paragraph id="8BC63689A713FFE0FF75548DFB262FFF" blockId="4.[151,1436,151,758]" box="[197,1267,654,680]" pageId="4" pageNumber="29">
<treatmentCitationGroup id="AB6911A7A713FFE0FF75548DFB262FFF" box="[197,1267,654,680]" pageId="4" pageNumber="29">
<taxonomicName id="4C794D0AA713FFE0FF75548DFD972FFF" ID-CoL="68Y8G" authority="Evenhuis, 2000: 293" authorityName="Evenhuis" authorityPageNumber="293" authorityYear="2000" box="[197,578,654,680]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="limnobates">
<emphasis id="B90DEA9BA713FFE0FF75548DFE9C2FFF" bold="true" box="[197,329,654,680]" italics="true" pageId="4" pageNumber="29">limnobates</emphasis>
<treatmentCitation id="0AD81098A713FFE0FEE0548DFD972FFF" author="Evenhuis" box="[336,578,654,680]" page="293" pageId="4" pageNumber="29" year="2000">
<bibRefCitation id="EFE84B78A713FFE0FEE0548DFDDE2FFF" author="Evenhuis" box="[336,523,654,680]" pageId="4" pageNumber="48" refString="Evenhuis, N. L. (2000) Water-skating Campsicnemus of the Marquesas Islands, including two new species (Diptera: Dolichopodidae). Journal of the New York Entomological Society, 107 (4) [1999], 289 - 296." type="journal article" year="2000">Evenhuis, 2000</bibRefCitation>
: 293
</treatmentCitation>
</taxonomicName>
(Marquesas [Ua Huka];
<typeStatus id="54C2882BA713FFE0FCD5548DFC192FFF" box="[869,972,654,680]" pageId="4" pageNumber="29" type="holotype">holotype</typeStatus>
ɗ in USNM, examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC63689A713FFE0FF7554B6FBE32F98" blockId="4.[151,1436,151,758]" box="[197,1078,692,719]" pageId="4" pageNumber="29">
<treatmentCitationGroup id="AB6911A7A713FFE0FF7554B6FBE32F98" box="[197,1078,692,719]" pageId="4" pageNumber="29">
<taxonomicName id="4C794D0AA713FFE0FF7554B6FE3F2F98" ID-CoL="699Y9" authority="Parent, 1934: 299" authorityName="Parent" authorityPageNumber="299" authorityYear="1934" box="[197,490,693,719]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="scurra">
<emphasis id="B90DEA9BA713FFE0FF7554B6FEC62F98" bold="true" box="[197,275,693,719]" italics="true" pageId="4" pageNumber="29">scurra</emphasis>
<treatmentCitation id="0AD81098A713FFE0FEAA54B6FE3F2F98" author="Parent" box="[282,490,693,719]" page="299" pageId="4" pageNumber="29" year="1934">
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: 299
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(Marquesas [Nuku Hiva];
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ɗ in BMNH).
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<taxonomicName id="4C794D0AA713FFE0FF7554DFFD9E2FA1" ID-CoL="5X9XZ" authority="Evenhuis, 2000: 294" authorityName="Evenhuis" authorityPageNumber="294" authorityYear="2000" box="[197,587,732,758]" class="Insecta" family="Dolichopodidae" genus="Campsicnemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="uncleremus">
<emphasis id="B90DEA9BA713FFE0FF7554DFFE862FA1" bold="true" box="[197,339,732,758]" italics="true" pageId="4" pageNumber="29">uncleremus</emphasis>
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: 294
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(Marquesas [Hiva Oa];
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ɗ in BPBM, examined).
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