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<mods:title id="C7AC635E5271A5C947A86C9800F668D0">Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae)</mods:title>
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<taxonomicName id="6EF84D5D4C20B340FF6BFC00F6AFD68A" ID-CoL="8VRHV" authority="Nordmann 1837" authorityName="Nordmann" authorityYear="1837" box="[151,506,980,1006]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C20B340FF6BFC00F6AFD68A" bold="true" box="[151,506,980,1006]" pageId="8" pageNumber="9">
<emphasis id="9B8CEACC4C20B340FF6BFC00F67DD68A" bold="true" box="[151,296,980,1006]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
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<taxonomicName id="6EF84D5D4C20B340FF6BFBCEF6A3D154" ID-CoL="8VRHV" authority="Nordmann 1837: 129" authorityName="Nordmann" authorityPageNumber="129" authorityYear="1837" box="[151,502,1050,1073]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C20B340FF6BFBCEF641D155" bold="true" box="[151,276,1050,1073]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
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: 129
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. 
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species: 
<taxonomicName id="6EF84D5D4C20B340FD72FBCEF30CD154" ID-CoL="8Y9S2" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[654,1113,1050,1072]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="9B8CEACC4C20B340FD72FBCEF4E7D154" box="[654,946,1050,1072]" italics="true" pageId="8" pageNumber="9">Staphylinus haemorrhoidalis</emphasis>
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, by monotypy.
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= 
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<emphasis id="9B8CEACC4C20B340FF4CFBEDF666D12B" box="[176,307,1081,1103]" italics="true" pageId="8" pageNumber="9">Musicoderus</emphasis>
<treatmentCitation id="285910CF4C20B340FEC1FBEDF6A5D12B" author="" box="[317,496,1081,1103]" page="455" pageId="8" pageNumber="9">
<bibRefCitation id="CD694B2F4C20B340FEC1FBEDF6ECD12B" box="[317,441,1081,1103]" pageId="8" pageNumber="9" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 455
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. 
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species: 
<taxonomicName id="6EF84D5D4C20B340FD69FBEDF346D12B" ID-CoL="6S2R7" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[661,1043,1081,1103]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cephalotes">
<emphasis id="9B8CEACC4C20B340FD69FBEDF4DBD12B" box="[661,910,1081,1103]" italics="true" pageId="8" pageNumber="9">Musicoderus cephalotes</emphasis>
<bibRefCitation id="CD694B2F4C20B340FC65FBEDF346D12B" box="[921,1043,1081,1103]" pageId="8" pageNumber="9" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
, des. by 
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<bibRefCitation id="CD694B2F4C20B340FB84FBEEF3A0D12B" box="[1144,1269,1081,1103]" pageId="8" pageNumber="9" refId="ref68856" refString="Lucas, R. (1920) Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Archiv fur Naturgeschichte, 84 (A), i - xxxi, 1 - 696 pp. [1918]" type="book chapter">Lucas 1920</bibRefCitation>
: 425
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. Notes: as subgenus of 
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<emphasis id="9B8CEACC4C20B340FEBBFB8CF6E8D10A" box="[327,445,1112,1134]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
(
<treatmentCitation id="285910CF4C20B340FE36FB8CF5E5D10A" author="" box="[458,688,1112,1134]" page="266" pageId="8" pageNumber="9">
<bibRefCitation id="CD694B2F4C20B340FE36FB8CF5E5D10A" box="[458,688,1112,1134]" pageId="8" pageNumber="9" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c: 266</bibRefCitation>
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)
</taxonomicName>
; as synonym of 
<taxonomicName id="6EF84D5D4C20B340FCA0FB8CF38CD10A" ID-CoL="8VRHV" authority="(Blackwelder 1943: 421)" authorityName="Blackwelder" authorityPageNumber="421" authorityYear="1943" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" box="[860,1241,1112,1134]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C20B340FCA0FB8CF487D10A" box="[860,978,1112,1134]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
(
<treatmentCitation id="285910CF4C20B340FC23FB8CF384D10A" author="" box="[991,1233,1112,1134]" page="421" pageId="8" pageNumber="9">
<bibRefCitation id="CD694B2F4C20B340FC23FB8CF384D10A" box="[991,1233,1112,1134]" pageId="8" pageNumber="9" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943: 421</bibRefCitation>
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)
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.
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<paragraph id="A94736DE4C20B340FF6BFB4FF57CD003" blockId="8.[151,1437,1178,1383]" pageId="8" pageNumber="9">
Notes: There are 165 valid species of 
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<emphasis id="9B8CEACC4C20B340FDB5FB4EF599D1D7" box="[585,716,1178,1203]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
</taxonomicName>
distributed in CASA. A recent study (
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Chani-Posse 
<emphasis id="9B8CEACC4C20B340FAE5FB48F206D1D7" box="[1305,1363,1178,1203]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2018
</bibRefCitation>
) shows 
<taxonomicName id="6EF84D5D4C20B340FEE8FB6AF6C2D1B3" ID-CoL="8VRHV" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" box="[276,407,1214,1239]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C20B340FEE8FB6AF6C2D1B3" box="[276,407,1214,1239]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
</taxonomicName>
as a genus of Neotropical origin and generic limits so far undefined. The current concept of the genus is that of 
<bibRefCitation id="CD694B2F4C20B340FE47FB37F521D19F" box="[443,628,1250,1275]" pageId="8" pageNumber="9" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana (1995)</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C20B340FD79FB36F421D19F" box="[645,884,1250,1275]" pageId="8" pageNumber="9" refId="ref64382" refString="Chani-Posse, M. (2014 a) An illustrated key to the New World genera of Philonthina Kirby (Coleoptera: Staphylinidae), with morphological, taxonomical and distributional notes. Zootaxa, 3755 (1), 62 - 86. https: // doi. org / 10.11646 / zootaxa. 3755.1.3" type="journal article">Chani-Posse (2014a)</bibRefCitation>
and 
<bibRefCitation id="CD694B2F4C20B340FC48FB36F3B5D19F" box="[948,1248,1250,1275]" pageId="8" pageNumber="9" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C20B340FBB0FB30F3DFD19F" box="[1100,1162,1250,1275]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2018)
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but most of its Neotropical endemic species were never revised after being described and they do not fit into this concept. Species that may belong to the so called ‘ 
<taxonomicName id="6EF84D5D4C20B340FD51FAFEF4A1D027" ID-CoL="LC89" authorityName="" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[685,1012,1322,1347]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="9B8CEACC4C20B340FD51FAFEF4A1D027" box="[685,1012,1322,1347]" italics="true" pageId="8" pageNumber="9">Belonuchus haemorrhoidalis</emphasis>
</taxonomicName>
clade’ (
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Chani-Posse 
<emphasis id="9B8CEACC4C20B340FB16FAF8F270D027" box="[1258,1317,1322,1347]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2018
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) are here denoted with an asterisk *.
</paragraph>
<paragraph id="A94736DE4C20B340FF6BFA42F436D0CB" blockId="8.[151,1436,1430,1923]" box="[151,867,1430,1455]" pageId="8" pageNumber="9">
<emphasis id="9B8CEACC4C20B340FF6BFA42F7F6D0CB" bold="true" box="[151,163,1430,1455]" italics="true" pageId="8" pageNumber="9">*</emphasis>
<taxonomicName id="6EF84D5D4C20B340FF5EFA42F500D0CB" ID-CoL="LC4M" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,597,1430,1455]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abnormalis" status="comb. nov.">
<emphasis id="9B8CEACC4C20B340FF5EFA42F6ECD0CB" bold="true" box="[162,441,1430,1455]" italics="true" pageId="8" pageNumber="9">Belonuchus abnormalis</emphasis>
(
<bibRefCitation id="CD694B2F4C20B340FE34FA42F518D0CB" box="[456,589,1430,1455]" pageId="8" pageNumber="9" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C20B340FD9DFA42F5B4D0CB" box="[609,737,1430,1455]" pageId="8" pageNumber="9">
<emphasis id="9B8CEACC4C20B340FD9DFA42F5B4D0CB" bold="true" box="[609,737,1430,1455]" pageId="8" pageNumber="9">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C20B340FD0DFA42F40FD0CB" box="[753,858,1430,1455]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="8" pageNumber="9">Figs. 1–2</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C20B340FF6BFA6EF666D37F" blockId="8.[151,1436,1430,1923]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="89E811F04C20B340FF6BFA6EF666D37F" infered="true" pageId="8" pageNumber="9">
<taxonomicName id="6EF84D5D4C20B340FF6BFA6EF508D0B7" ID-CoL="6V9VC" authority="Sharp 1885: 397" authorityName="Sharp" authorityPageNumber="397" authorityYear="1885" box="[151,605,1466,1491]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abnormalis">
<emphasis id="9B8CEACC4C20B340FF6BFA6EF6CDD0B7" box="[151,408,1466,1491]" italics="true" pageId="8" pageNumber="9">Philonthus abnormalis</emphasis>
<treatmentCitation id="285910CF4C20B340FE63FA6EF508D0B7" author="" box="[415,605,1466,1491]" page="397" pageId="8" pageNumber="9">
<bibRefCitation id="CD694B2F4C20B340FE63FA6EF571D0B7" box="[415,548,1466,1491]" pageId="8" pageNumber="9" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 397
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C20B340FD94FA6FF5F5D0B7" box="[616,672,1467,1491]" pageId="8" pageNumber="9">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C20B340FCF7FA6EF430D0B7" box="[779,869,1466,1491]" country="Mexico" name="Mexico" pageId="8" pageNumber="9">
<collectingCountry id="D1EF764E4C20B340FCF7FA6EF430D0B7" box="[779,869,1466,1491]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
</collectingRegion>
: 
<collectingRegion id="6B3CF83C4C20B340FC8CFA6FF49DD0B7" box="[880,968,1467,1491]" country="Mexico" name="Oaxaca" pageId="8" pageNumber="9">Oaxaca</collectingRegion>
. 
<typeStatus id="7643887C4C20B340FC2FFA6FF35ED0B7" box="[979,1035,1467,1491]" pageId="8" pageNumber="9">Type</typeStatus>
depository: BMNH 2 ST: 
<specimenCount id="BFFEFD574C20B340FAC4FA6FF230D0B6" box="[1336,1381,1467,1491]" pageId="8" pageNumber="9" type="male">1 ♂</specimenCount>
, 
<specimenCount id="BFFEFD574C20B340FA8EFA6FF2C9D0B6" box="[1394,1436,1467,1491]" pageId="8" pageNumber="9" type="female">1 ♀</specimenCount>
[MCP seen]. Distribution: 
<collectingRegion id="6B3CF83C4C20B340FE04FA0AF505D093" box="[504,592,1502,1527]" country="Mexico" name="Mexico" pageId="8" pageNumber="9">
<collectingCountry id="D1EF764E4C20B340FE04FA0AF505D093" box="[504,592,1502,1527]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C20B340FDA1FA0AF58FD093" box="[605,730,1502,1527]" name="Guatemala" pageId="8" pageNumber="9">
<collectingRegion id="6B3CF83C4C20B340FDA1FA0AF58FD093" box="[605,730,1502,1527]" country="Guatemala" name="Guatemala" pageId="8" pageNumber="9">Guatemala</collectingRegion>
</collectingCountry>
. Note: distribution, also within 
<collectingRegion id="6B3CF83C4C20B340FBBBFA0AF3C9D093" box="[1095,1180,1502,1527]" country="Mexico" name="Mexico" pageId="8" pageNumber="9">
<collectingCountry id="D1EF764E4C20B340FBBBFA0AF3C9D093" box="[1095,1180,1502,1527]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C20B340FB50FA0AF67DD37F" pageId="8" pageNumber="9" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C20B340FA74FA0BF7B1D37F" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C20B340FF6BF9F2F454D3E3" pageId="8" pageNumber="9" type="discussion">
<paragraph id="A94736DE4C20B340FF6BF9F2F454D3E3" blockId="8.[151,1436,1430,1923]" pageId="8" pageNumber="9">
<emphasis id="9B8CEACC4C20B340FF6BF9F2F65ED35B" bold="true" box="[151,267,1574,1599]" pageId="8" pageNumber="9">Remarks.</emphasis>
Sharp explicitly mentioned two specimens (male and female) as part of the 
<typeStatus id="7643887C4C20B340FB55F9F3F38CD35B" box="[1193,1241,1575,1599]" pageId="8" pageNumber="9">type</typeStatus>
series. The two specimens listed below agree with the original description by 
<bibRefCitation id="CD694B2F4C20B340FC5BF99EF36AD307" box="[935,1087,1610,1635]" pageId="8" pageNumber="9" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp (1885)</bibRefCitation>
and considered to have been 
<typeStatus id="7643887C4C20B340FF3BF9BBF67CD3E3" box="[199,297,1647,1671]" pageId="8" pageNumber="9" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C20B340FDF7F9BAF527D3E3" box="[523,626,1646,1671]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C20B340FF6BF946F5BCD25F" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="A94736DE4C20B340FF6BF946F5BCD25F" blockId="8.[151,1436,1430,1923]" pageId="8" pageNumber="9">
<emphasis id="9B8CEACC4C20B340FF6BF946F6E8D3CF" bold="true" box="[151,445,1682,1707]" pageId="8" pageNumber="9">
<typeStatus id="7643887C4C20B340FF6BF946F787D3CF" box="[151,210,1682,1707]" pageId="8" pageNumber="9">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C20B340FE38F947F561D3CF" box="[452,564,1683,1707]" pageId="8" pageNumber="9" type="lectotype">Lectotype</typeStatus>
<specimenCount id="BFFEFD574C20B340FDC6F947F504D3CE" box="[570,593,1683,1706]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
, here designated (
<collectionCode id="CFE9AE1B4C20B340FCE5F947F424D3CF" box="[793,881,1683,1707]" pageId="8" pageNumber="9">BMNH</collectionCode>
, 
<figureCitation id="31C32A5B4C20B340FC80F946F4B2D3CF" box="[892,999,1682,1707]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="8" pageNumber="9">Figs. 1–2</figureCitation>
): card-mounted, “ 
<specimenCount id="BFFEFD574C20B340FB53F947F390D3CF" box="[1199,1221,1683,1707]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
(in ink) 
<taxonomicName id="6EF84D5D4C20B340FAD8F946F611D3AB" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abnormalis">Philonthus abnormalis</taxonomicName>
<typeStatus id="7643887C4C20B340FEB7F963F6D6D3AB" box="[331,387,1719,1743]" pageId="8" pageNumber="9">Type</typeStatus>
D.S. 
<collectingRegion id="6B3CF83C4C20B340FE38F963F549D3AB" box="[452,540,1719,1743]" country="Mexico" name="Oaxaca" pageId="8" pageNumber="9">Oaxaca</collectingRegion>
. 
<collectingRegion id="6B3CF83C4C20B340FDDBF962F5D5D3AB" box="[551,640,1718,1743]" country="Mexico" name="Mexico" pageId="8" pageNumber="9">
<collectingCountry id="D1EF764E4C20B340FDDBF962F5D5D3AB" box="[551,640,1718,1743]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
</collectingRegion>
. Höge [in Sharp’s hand] // 
<typeStatus id="7643887C4C20B340FC4DF963F4BDD3AB" box="[945,1000,1719,1743]" pageId="8" pageNumber="9">Type</typeStatus>
(round white label with red margin) // 
<collectingRegion id="6B3CF83C4C20B340FF3BF90FF64BD397" box="[199,286,1755,1779]" country="Mexico" name="Oaxaca" pageId="8" pageNumber="9">Oaxaca</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C20B340FED4F90EF6D4D397" box="[296,385,1754,1779]" country="Mexico" name="Mexico" pageId="8" pageNumber="9">
<collectingCountry id="D1EF764E4C20B340FED4F90EF6D4D397" box="[296,385,1754,1779]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
</collectingRegion>
. Hoege. (white label) // B. C. A. Col. 
<date id="DD46101E4C20B340FCDAF90FF407D397" box="[806,850,1755,1779]" pageId="8" pageNumber="9">I. 2.</date>
<taxonomicName id="6EF84D5D4C20B340FCA5F90EF3FFD397" authority=", Sharp." authorityName="Sharp." box="[857,1194,1754,1779]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abnormalis">Philonthus abnormalis, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”. 
<typeStatus id="7643887C4C20B340FDD3F92AF592D273" box="[559,711,1790,1815]" pageId="8" pageNumber="9" type="paralectotype">Paralectotype</typeStatus>
<specimenCount id="BFFEFD574C20B340FD32F92BF5B6D272" box="[718,739,1791,1814]" pageId="8" pageNumber="9" type="female">♀</specimenCount>
(
<collectionCode id="CFE9AE1B4C20B340FD0FF92BF41BD273" box="[755,846,1791,1815]" pageId="8" pageNumber="9">BMNH</collectionCode>
): with same locality and collection data as 
<typeStatus id="7643887C4C20B340FAD3F92AF2CDD273" box="[1327,1432,1790,1815]" pageId="8" pageNumber="9" type="lectotype">lectotype</typeStatus>
, and additional printed white label “Sp. Figured”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C20B340FF6BF892F545D2AF" pageId="8" pageNumber="9" type="discussion">
<paragraph id="A94736DE4C20B340FF6BF892F5ADD2E7" blockId="8.[151,1436,1430,1923]" pageId="8" pageNumber="9">
<emphasis id="9B8CEACC4C20B340FF6BF892F64AD23B" bold="true" box="[151,287,1862,1887]" pageId="8" pageNumber="9">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C20B340FCF4F892F38BD23B" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[776,1246,1862,1887]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C20B340FCF4F892F4DED23B" box="[776,907,1862,1887]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C20B340FC58F892F380D23B" box="[932,1237,1862,1887]" pageId="8" pageNumber="9" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C20B340FBB8F893F3DDD23B" box="[1092,1160,1862,1887]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
; determined as “ 
<taxonomicName id="6EF84D5D4C20B340FF2FF8BEF603D2E7" box="[211,342,1898,1923]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C20B340FF2FF8BEF603D2E7" box="[211,342,1898,1923]" italics="true" pageId="8" pageNumber="9">Belonuchus</emphasis>
</taxonomicName>
” by J. L. Navarrete-Heredia in 2002.
</paragraph>
<paragraph id="A94736DE4C20B340FF6BF866F545D2AF" blockId="8.[151,1437,1970,2031]" box="[151,528,1970,1995]" pageId="8" pageNumber="9">
<taxonomicName id="6EF84D5D4C20B340FF6BF866F545D2AF" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[151,528,1970,1995]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="9B8CEACC4C20B340FF6BF866F6DDD2AF" bold="true" box="[151,392,1970,1995]" italics="true" pageId="8" pageNumber="9">Belonuchus aequalis</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C20B340FF6BF802F2C8D28B" box="[151,1437,2006,2031]" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="A94736DE4C20B340FF6BF802F2C8D28B" blockId="8.[151,1437,1970,2031]" box="[151,1437,2006,2031]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="89E811F04C20B340FF6BF802F2C8D28B" box="[151,1437,2006,2031]" pageId="8" pageNumber="9">
<taxonomicName id="6EF84D5D4C20B340FF6BF802F51BD28B" ID-CoL="LC4S" authority="Sharp 1876 a: 161" authorityName="Sharp" authorityPageNumber="161" authorityYear="1876" box="[151,590,2006,2031]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="9B8CEACC4C20B340FF6BF802F62AD28B" box="[151,383,2006,2031]" italics="true" pageId="8" pageNumber="9">Belonuchus aequalis</emphasis>
<treatmentCitation id="285910CF4C20B340FE7AF802F51BD28B" author="" box="[390,590,2006,2031]" page="161" pageId="8" pageNumber="9">
<bibRefCitation id="CD694B2F4C20B340FE7AF802F543D28B" box="[390,534,2006,2031]" pageId="8" pageNumber="9" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 161
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C20B340FDA4F803F5C5D28B" box="[600,656,2007,2031]" pageId="8" pageNumber="9">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C20B340FD05F803F47BD28B" box="[761,814,2007,2031]" country="Brazil" name="Para" pageId="8" pageNumber="9">Pará</collectingRegion>
, Ega, St. Paulo [
<collectingRegion id="6B3CF83C4C20B340FC18F803F335D28B" box="[996,1120,2007,2031]" country="Venezuela" name="Amazonas" pageId="8" pageNumber="9">Amazonas</collectingRegion>
] &amp; Conceicão, Rio Mauhes.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C21B341FF3BFF4CF414D5B0" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="A94736DE4C21B341FF3BFF4CF414D5B0" blockId="9.[199,1437,151,213]" pageId="9" pageNumber="10">
<materialsCitation id="19903C834C21B341FF3BFF4CF351D5CB" ID-GBIF-Occurrence="1915497784" box="[199,1028,151,176]" collectionCode="MCP" location="St. Paulo" municipality="Ega" pageId="9" pageNumber="10" specimenCode="BMNH 6" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<typeStatus id="7643887C4C21B341FF3BFF4CF7AAD5D4" box="[199,255,152,176]" pageId="9" pageNumber="10">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C21B341FE6DFF4CF6ABD5D4" box="[401,510,151,176]" collectionCode="BMNH" pageId="9" pageNumber="10">BMNH 6</specimenCode>
ST: 4 
<collectingMunicipality id="4923ACA44C21B341FDAEFF4CF5D5D5D4" box="[594,640,152,176]" pageId="9" pageNumber="10">Ega</collectingMunicipality>
, 1 
<collectingRegion id="6B3CF83C4C21B341FD54FF4CF588D5D4" box="[680,733,152,176]" country="Brazil" name="Para" pageId="9" pageNumber="10">Pará</collectingRegion>
, 1 
<location id="AC2760054C21B341FCF8FF4CF439D5D4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C21B341FCF8FF4CF439D5D4" box="[772,876,151,176]" municipality="Ega" name="St. Paulo" pageId="9" pageNumber="10" stateProvince="Para">St. Paulo</location>
[
<collectionCode id="CFE9AE1B4C21B341FC81FF4CF4EED5D4" box="[893,955,152,176]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="9" pageNumber="10">MCP</collectionCode>
seen]
</materialsCitation>
. 
<materialsCitation id="19903C834C21B341FBEEFF4CF65AD5B1" ID-GBIF-Occurrence="1915497741" collectionCode="FMNH" country="Brazil" location="Brazil" pageId="9" pageNumber="10" specimenCode="FMNH 1" specimenCount="1" typeStatus="holotype">
<specimenCode id="F95E9EA54C21B341FBEEFF4CF32ED5D4" box="[1042,1147,152,176]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="10">FMNH 1</specimenCode>
ST: 1 Ega. Distribution: 
<collectingCountry id="D1EF764E4C21B341FF3BFF68F65AD5B1" box="[199,271,188,213]" name="Brazil" pageId="9" pageNumber="10">Brazil</collectingCountry>
</materialsCitation>
. Notes: distribution in Brazil (Asenjo 
<emphasis id="9B8CEACC4C21B341FD45FF69F5A7D5B1" box="[697,754,188,213]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2013).
</paragraph>
</subSubSection>
<caption id="FD8766564C21B341FF6BF8ECF3FCD2E8" httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,264,1848,1871]" targetBox="[227,1359,267,1824]" targetPageId="9">
<paragraph id="A94736DE4C21B341FF6BF8ECF3FCD2E8" blockId="9.[151,1436,1848,1932]" pageId="9" pageNumber="10">
<emphasis id="9B8CEACC4C21B341FF6BF8ECF615D22A" bold="true" box="[151,320,1848,1871]" pageId="9" pageNumber="10">FIGURES 1–12</emphasis>
[BMNH ©]. Type material (habitus and labels): 1, 2
<taxonomicName id="6EF84D5D4C21B341FCAFF8EDF3DCD22B" authority="Sharp" authorityName="Sharp" box="[851,1161,1849,1871]" pageId="9" pageNumber="10" rank="subSpecies" subSpecies="abnormalis">
, 
<emphasis id="9B8CEACC4C21B341FCA2F8EDF313D22B" box="[862,1094,1849,1871]" italics="true" pageId="9" pageNumber="10">Philonthus abnormalis</emphasis>
Sharp
</taxonomicName>
, lectotype; 3, 4
<taxonomicName id="6EF84D5D4C21B341FAD8F8EDF67CD209" authority="Sharp" authorityName="Sharp" pageId="9" pageNumber="10" rank="subSpecies" subSpecies="celatus">
, 
<emphasis id="9B8CEACC4C21B341FAD2F8EDF78AD209" italics="true" pageId="9" pageNumber="10">Philonthus celatus</emphasis>
Sharp
</taxonomicName>
, lectotype; 5, 6
<taxonomicName id="6EF84D5D4C21B341FE2AF883F45FD209" authority="Sharp" authorityName="Sharp" box="[470,778,1879,1901]" pageId="9" pageNumber="10" rank="subSpecies" subSpecies="corticalis">
, 
<emphasis id="9B8CEACC4C21B341FE1AF883F5EAD209" box="[486,703,1879,1901]" italics="true" pageId="9" pageNumber="10">Philonthus corticalis</emphasis>
Sharp
</taxonomicName>
, holotype; 7, 8
<taxonomicName id="6EF84D5D4C21B341FC4EF883F3B4D209" authority="Sharp" authorityName="Sharp" box="[946,1249,1879,1902]" pageId="9" pageNumber="10" rank="subSpecies" subSpecies="extremus">
, 
<emphasis id="9B8CEACC4C21B341FC38F883F3CDD20A" box="[964,1176,1879,1902]" italics="true" pageId="9" pageNumber="10">Philonthus extremus</emphasis>
Sharp
</taxonomicName>
, holotype; 9, 10
<taxonomicName id="6EF84D5D4C21B341FA6BF883F6E8D2E8" authority="Sharp" authorityName="Sharp" pageId="9" pageNumber="10" rank="subSpecies" subSpecies="latecinctus">
, 
<emphasis id="9B8CEACC4C21B341FF6BF8A2F62DD2E8" box="[151,376,1910,1932]" italics="true" pageId="9" pageNumber="10">Philonthus latecinctus</emphasis>
Sharp
</taxonomicName>
, lectotype; 11, 12
<taxonomicName id="6EF84D5D4C21B341FD8DF8A2F4D7D2E8" authority="Sharp" authorityName="Sharp" box="[625,898,1910,1932]" pageId="9" pageNumber="10" rank="subSpecies" subSpecies="muticus">
, 
<emphasis id="9B8CEACC4C21B341FD80F8A2F46AD2E8" box="[636,831,1910,1932]" italics="true" pageId="9" pageNumber="10">Philonthus muticus</emphasis>
Sharp
</taxonomicName>
, lectotype. Scale bar= 2 mm.
</paragraph>
</caption>
<subSubSection id="E1E265554C21B341FF6BF867F2C9D28B" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="A94736DE4C21B341FF6BF867F54BD2A8" blockId="9.[151,1436,1971,2032]" box="[151,542,1971,1996]" pageId="9" pageNumber="10">
<emphasis id="9B8CEACC4C21B341FF6BF867F7F6D2A8" bold="true" box="[151,163,1971,1996]" italics="true" pageId="9" pageNumber="10">*</emphasis>
<taxonomicName id="6EF84D5D4C21B341FF5EF867F54BD2A8" ID-CoL="LC4T" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[162,542,1971,1996]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="9B8CEACC4C21B341FF5EF867F621D2A8" bold="true" box="[162,372,1971,1996]" italics="true" pageId="9" pageNumber="10">Belonuchus agilis</emphasis>
<bibRefCitation id="CD694B2F4C21B341FE80F867F54BD2A8" box="[380,542,1971,1996]" pageId="9" pageNumber="10" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C21B341FF6BF803F2C9D28B" blockId="9.[151,1436,1971,2032]" box="[151,1436,2007,2032]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="89E811F04C21B341FF6BF803F2C9D28B" box="[151,1436,2007,2032]" pageId="9" pageNumber="10">
<taxonomicName id="6EF84D5D4C21B341FF6BF803F51ED294" ID-CoL="LC4T" authority="Erichson 1840: 423" authorityName="Erichson" authorityPageNumber="423" authorityYear="1840" box="[151,587,2007,2032]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="9B8CEACC4C21B341FF6BF803F635D294" box="[151,352,2007,2032]" italics="true" pageId="9" pageNumber="10">Belonuchus agilis</emphasis>
<treatmentCitation id="285910CF4C21B341FE95F803F51ED294" author="" box="[361,587,2007,2032]" page="423" pageId="9" pageNumber="10">
<bibRefCitation id="CD694B2F4C21B341FE95F803F544D294" box="[361,529,2007,2032]" pageId="9" pageNumber="10" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 423
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C21B341FDA4F80CF5DAD294" box="[600,655,2008,2032]" pageId="9" pageNumber="10">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C21B341FD00F803F46ED294" box="[764,827,2007,2032]" name="Cuba" pageId="9" pageNumber="10">Cuba</collectingCountry>
. 
<typeStatus id="7643887C4C21B341FCBBF80CF42AD294" box="[839,895,2008,2032]" pageId="9" pageNumber="10">Type</typeStatus>
depository: ZMHB 2 ST: 
<specimenCount id="BFFEFD574C21B341FB52F80CF386D294" box="[1198,1235,2008,2032]" pageId="9" pageNumber="10" type="male">1♂</specimenCount>
, 
<specimenCount id="BFFEFD574C21B341FB1DF80CF251D294" box="[1249,1284,2008,2032]" pageId="9" pageNumber="10" type="female">1♀</specimenCount>
[MCP seen].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C22B342FF3BFF43F35ED5B1" pageId="10" pageNumber="11" type="distribution">
<paragraph id="A94736DE4C22B342FF3BFF43F35ED5B1" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="11">
Distribution: 
<collectingCountry id="D1EF764E4C22B342FEA2FF4CF6CCD5D4" box="[350,409,152,176]" name="United States of America" pageId="10" pageNumber="11">USA</collectingCountry>
, Dominican Rep., 
<collectingCountry id="D1EF764E4C22B342FD92FF43F5F9D5D4" box="[622,684,151,176]" name="Cuba" pageId="10" pageNumber="11">Cuba</collectingCountry>
, 
<collectingCountry id="D1EF764E4C22B342FD44FF43F440D5D4" box="[696,789,151,176]" name="Jamaica" pageId="10" pageNumber="11">Jamaica</collectingCountry>
, 
<collectingCountry id="D1EF764E4C22B342FCDDFF43F409D5D4" box="[801,860,151,176]" name="Haiti" pageId="10" pageNumber="11">Haiti</collectingCountry>
, 
<collectingCountry id="D1EF764E4C22B342FC95FF43F48ED5D4" box="[873,987,151,176]" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
. Notes: distribution including 
<collectingCountry id="D1EF764E4C22B342FACFFF4CF23ED5D4" box="[1331,1387,152,176]" name="United States of America" pageId="10" pageNumber="11">USA</collectingCountry>
and Dominican Rep. (
<bibRefCitation id="CD694B2F4C22B342FE71FF68F541D5B0" box="[397,532,188,213]" pageId="10" pageNumber="11" refId="ref66436" refString="Frank, J. H. (2004) Belonuchus agilis, a fourth species of this genus (Coleoptera: Staphylinidae) reported from Florida. Florida Entomologist, 87 (1), 92. https: // doi. org / 10.1653 / 0015 - 4040 (2004) 087 [0092: BAAFSO] 2.0. CO; 2" type="journal article">Frank 2004</bibRefCitation>
); distribution, also within 
<collectingCountry id="D1EF764E4C22B342FCC5FF68F421D5B1" box="[825,884,188,213]" name="Cuba" pageId="10" pageNumber="11">Cuba</collectingCountry>
(
<bibRefCitation id="CD694B2F4C22B342FC7EFF68F4AAD5B1" box="[898,1023,188,213]" pageId="10" pageNumber="11" refId="ref70864" refString="Peck, S. B. (2005) A checklist of the beetles of Cuba: with data on distributions and bionomics (Insecta: Coleoptera). Arthropods of Florida and Neighboring Land Areas, 18, 1 - 241." type="journal article">Peck 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C22B342FF6BFF35F68BD704" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="A94736DE4C22B342FF6BFF35F417D478" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFF35F417D478" pageId="10" pageNumber="11">
<emphasis id="9B8CEACC4C22B342FF6BFF35F7FFD59C" box="[151,170,225,248]" italics="true" pageId="10" pageNumber="11">=</emphasis>
<taxonomicName id="6EF84D5D4C22B342FF4CFF0BF52CD59C" ID-CoL="LC6K" authority="Fauvel 1863: 432" authorityName="Fauvel" authorityPageNumber="432" authorityYear="1863" box="[176,633,223,248]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chevrolati">
<emphasis id="9B8CEACC4C22B342FF4CFF0BF6FFD59C" box="[176,426,223,248]" italics="true" pageId="10" pageNumber="11">Belonuchus chevrolati</emphasis>
<treatmentCitation id="285910CF4C22B342FE4DFF0BF52CD59C" author="" box="[433,633,223,248]" page="432" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE4DFF0BF515D59C" box="[433,576,223,248]" pageId="10" pageNumber="11" refId="ref65754" refString="Fauvel, A. (1863) Chevrolat, A. &amp; A. Fauvel. Coleopteres de l'ile de Cuba (suite). Notes, synonymies et descriptions d'especes nouvelles. Troisieme memoire. Famille des staphyliniens. Annales de la Societe Entomologique de France, Series 4, 3, 427 - 446." type="journal article">Fauvel 1863</bibRefCitation>
: 432
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C22B342FD7EFF34F5EFD59C" box="[642,698,224,248]" pageId="10" pageNumber="11">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C22B342FCDFFF34F4F5D593" box="[803,928,224,247]" country="Cuba" name="La Habana" pageId="10" pageNumber="11">La Havane</collectingRegion>
, 
<collectingCountry id="D1EF764E4C22B342FC57FF0BF4BFD59C" box="[939,1002,223,248]" name="Cuba" pageId="10" pageNumber="11">Cuba</collectingCountry>
. 
<typeStatus id="7643887C4C22B342FC08FF34F37ED59C" box="[1012,1067,224,248]" pageId="10" pageNumber="11">Type</typeStatus>
depository: IRSNB HT 
<specimenCount id="BFFEFD574C22B342FAC0FF34F204D593" box="[1340,1361,224,247]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
[MCP seen]. Note: as synonym of 
<taxonomicName id="6EF84D5D4C22B342FE03FED1F46BD478" ID-CoL="LC4T" authority="(Fauvel 1869: 489)" authorityName="Fauvel" authorityPageNumber="489" authorityYear="1869" baseAuthorityName="Fauvel" baseAuthorityYear="1869" box="[511,830,260,285]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="agilis">
<emphasis id="9B8CEACC4C22B342FE03FED1F509D479" box="[511,604,260,285]" italics="true" pageId="10" pageNumber="11">B. agilis</emphasis>
(
<treatmentCitation id="285910CF4C22B342FD97FED0F460D478" author="" box="[619,821,260,285]" page="489" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FD97FED0F460D478" box="[619,821,260,285]" pageId="10" pageNumber="11" refId="ref65845" refString="Fauvel, A. (1869) Remarques synonymiques sur les Staphylinides du Catalogus Coleopterorum de MM. v. Harold et Gemminger. Abeille, Memoires d'Entomologie, 5, 479 - 494." type="journal article">Fauvel 1869: 489</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFE9FF5D9D401" blockId="10.[151,1437,331,645]" box="[151,652,331,357]" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BFE9FF5D9D401" ID-CoL="68GGP" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,652,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="albovariegatus">
<emphasis id="9B8CEACC4C22B342FF6BFE9FF687D400" bold="true" box="[151,466,331,356]" italics="true" pageId="10" pageNumber="11">Belonuchus albovariegatus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFEBBF686D4B4" blockId="10.[151,1437,331,645]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFEBBF686D4B4" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BFEBBF59CD4EC" ID-CoL="68GGP" authority="Bernhauer 1916 c: 276" authorityName="Bernhauer" authorityPageNumber="276" authorityYear="1916" box="[151,713,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="albovariegatus">
<emphasis id="9B8CEACC4C22B342FF6BFEBBF69DD4EC" box="[151,456,367,392]" italics="true" pageId="10" pageNumber="11">Belonuchus albovariegatus</emphasis>
<treatmentCitation id="285910CF4C22B342FE32FEBBF59CD4EC" author="" box="[462,713,367,392]" page="276" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE32FEBBF5C6D4EC" box="[462,659,367,392]" pageId="10" pageNumber="11" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 276
</treatmentCitation>
</taxonomicName>
. Type locality: West-Columbien: Umgebung Cali am Rio Cauca, St. Antonio, 
<quantity id="6E009B3B4C22B342FE9EFE40F6EAD4C8" box="[354,447,404,428]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="2000.0">2000 m</quantity>
. Type depository: FMNH LT ♂, 2 PLT (des. 
<bibRefCitation id="CD694B2F4C22B342FC12FE40F2C4D4C8" box="[1006,1425,404,429]" pageId="10" pageNumber="11" refId="ref64722" refString="Chani-Posse, M. &amp; Solodovnikov, A. (2017) Tackling generic limits for the Neotropical Philonthina with a phylogenetic approach: revision of the genera Linoderus Sharp and Odontolinus Sharp (Coleoptera: Staphylinidae). Arthropod Systematics &amp; Phylogeny, 75 (1), 83 - 97." type="journal article">Chani-Posse &amp; Solodovnikov 2017</bibRefCitation>
). Distribution: Colombia.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFE09F68BD704" blockId="10.[151,1437,331,645]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFE09F68BD704" infered="true" pageId="10" pageNumber="11">
<materialsCitation id="19903C834C22B342FF6BFE09F206D773" ID-GBIF-Occurrence="1915497782" country="Colombia" county="Filandia" elevation="2000" latitude="4.7017775" location="Parque Regional Natural Barbas-Bremen" longLatPrecision="1" longitude="-75.637856" municipality="Cruces" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Quindio" typeStatus="holotype">
= 
<taxonomicName id="6EF84D5D4C22B342FF49FE08F4F0D491" ID-CoL="3V3RZ" authority="Lopez-Garcia &amp; Mendez-Rojas 2014: 86" authorityName="Lopez-Garcia &amp; Mendez-Rojas" authorityPageNumber="86" authorityYear="2014" box="[181,933,476,501]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="navarretei">
<emphasis id="9B8CEACC4C22B342FF49FE08F6FFD490" box="[181,426,476,501]" italics="true" pageId="10" pageNumber="11">Linoderus navarretei</emphasis>
<treatmentCitation id="285910CF4C22B342FE44FE08F4F0D491" author="" box="[440,933,476,501]" page="86" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE44FE08F426D490" box="[440,883,476,501]" pageId="10" pageNumber="11" refId="ref68711" refString="Lopez-Garcia, M. M. &amp; Mendez-Rojas, D. M. (2014) A new species of the genus Linoderus Sharp, 1885 (Coleoptera, Staphylinidae, Philonthina) from the Colombian Andes. Zootaxa, 3795 (1), 85 - 90. https: // doi. org / 10.11646 / zootaxa. 3795.1.9" type="journal article">López-García &amp; Méndez-Rojas 2014</bibRefCitation>
: 86
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C22B342FC44FE09F4A5D491" box="[952,1008,477,501]" pageId="10" pageNumber="11">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C22B342FB95FE08F38FD491" box="[1129,1242,476,501]" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C22B342FB11FE08F21CD491" box="[1261,1353,476,501]" country="Colombia" name="Quindio" pageId="10" pageNumber="11">Quindío</collectingRegion>
Dept., 
<collectingCounty id="40264E524C22B342FF3BFE2BF673D77C" box="[199,294,511,536]" pageId="10" pageNumber="11">Filandia</collectingCounty>
, 
<collectingMunicipality id="4923ACA44C22B342FEC4FDD4F6DCD77C" box="[312,393,512,536]" pageId="10" pageNumber="11">Cruces</collectingMunicipality>
, 
<location id="AC2760054C22B342FE67FDD4F42AD77C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C22B342FE67FDD4F42AD77C" box="[411,895,511,536]" country="Colombia" county="Filandia" latitude="4.7017775" longLatPrecision="1" longitude="-75.637856" municipality="Cruces" name="Parque Regional Natural Barbas-Bremen" pageId="10" pageNumber="11" stateProvince="Quindio">Parque Regional Natural Barbas-Bremen</location>
, 
<geoCoordinate id="CCCC50194C22B342FC6DFE2BF375D773" box="[913,1056,511,536]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="4.7017775">4°42’6.4” N</geoCoordinate>
<geoCoordinate id="CCCC50194C22B342FBD3FE2BF3B7D773" box="[1071,1250,511,536]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="-75.637856">75°38’16.3” W</geoCoordinate>
, 
<elevation id="22D5D1ED4C22B342FB08FDD4F206D773" box="[1268,1363,512,536]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="2000.0">
<quantity id="6E009B3B4C22B342FB08FDD4F206D773" box="[1268,1363,512,536]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="2000.0">2000 m</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C22B342FA98FDD4F68BD704" ID-GBIF-Occurrence="1915497737" collectionCode="UNCB" country="Colombia" location="Note" pageId="10" pageNumber="11" specimenCode="CIUQ 3" specimenCount="1" specimenCount-male="1" stateProvince="Quindio" typeStatus="holotype">
<typeStatus id="7643887C4C22B342FA98FDD4F2C9D77C" box="[1380,1436,512,536]" pageId="10" pageNumber="11">Type</typeStatus>
depositories: 
<collectionCode id="CFE9AE1B4C22B342FE9EFDF0F6E5D758" box="[354,432,548,572]" pageId="10" pageNumber="11">UNCB</collectionCode>
HT 
<specimenCount id="BFFEFD574C22B342FE11FDF1F551D758" box="[493,516,549,572]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
, 13 PT; 
<specimenCode id="F95E9EA54C22B342FD93FDF0F598D758" box="[623,717,548,572]" collectionCode="CIUQ" pageId="10" pageNumber="11">CIUQ 3</specimenCode>
PT. 
<location id="AC2760054C22B342FCF6FDF1F410D758" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C22B342FCF6FDF1F410D758" box="[778,837,549,572]" country="Colombia" name="Note" pageId="10" pageNumber="11" stateProvince="Quindio">Note</location>
: as synonym of 
<taxonomicName id="6EF84D5D4C22B342FBF1FDF1F68ED704" ID-CoL="68GGP" authority="(Chani-Posse &amp; Solodovnikov 2017: 90)" authorityName="Chani-Posse &amp; Solodovnikov" authorityPageNumber="90" authorityYear="2017" baseAuthorityName="Chani-Posse &amp; Solodovnikov" baseAuthorityYear="2017" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="albovariegatus">
<emphasis id="9B8CEACC4C22B342FBF1FDF1F38DD759" box="[1037,1240,548,573]" italics="true" pageId="10" pageNumber="11">B. albovariegatus</emphasis>
(
<treatmentCitation id="285910CF4C22B342FB10FDF0F687D704" author="" page="90" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FB10FDF0F687D704" pageId="10" pageNumber="11" refId="ref64722" refString="Chani-Posse, M. &amp; Solodovnikov, A. (2017) Tackling generic limits for the Neotropical Philonthina with a phylogenetic approach: revision of the genera Linoderus Sharp and Odontolinus Sharp (Coleoptera: Staphylinidae). Arthropod Systematics &amp; Phylogeny, 75 (1), 83 - 97." type="journal article">Chani-Posse &amp; Solodovnikov 2017: 90</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C22B342FF6BFDB8F465D7E1" box="[151,816,620,645]" pageId="10" pageNumber="11" type="discussion">
<paragraph id="A94736DE4C22B342FF6BFDB8F465D7E1" blockId="10.[151,1437,331,645]" box="[151,816,620,645]" pageId="10" pageNumber="11">
<emphasis id="9B8CEACC4C22B342FF6BFDB8F641D7E1" bold="true" box="[151,276,620,645]" pageId="10" pageNumber="11">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C22B342FEB0FDB8F69AD7E1" box="[332,463,620,645]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C22B342FEB0FDB8F69AD7E1" box="[332,463,620,645]" italics="true" pageId="10" pageNumber="11">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C22B342FF6BFD67F4AED054" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="A94736DE4C22B342FF6BFD67F568D7A9" blockId="10.[151,1436,691,824]" box="[151,573,691,717]" pageId="10" pageNumber="11">
<emphasis id="9B8CEACC4C22B342FF6BFD67F7F6D7A8" bold="true" box="[151,163,691,716]" italics="true" pageId="10" pageNumber="11">*</emphasis>
<taxonomicName id="6EF84D5D4C22B342FF5EFD67F568D7A9" ID-CoL="LC4V" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,573,691,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="9B8CEACC4C22B342FF5EFD67F6F6D7A8" bold="true" box="[162,419,691,716]" italics="true" pageId="10" pageNumber="11">Belonuchus alternans</emphasis>
(
<bibRefCitation id="CD694B2F4C22B342FE4EFD60F560D7A9" box="[434,565,692,717]" pageId="10" pageNumber="11" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFD03F5E2D65C" blockId="10.[151,1436,691,824]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFD03F5E2D65C" infered="true" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BFD03F501D794" ID-CoL="76XW4" authority="Sharp 1885: 418" authorityName="Sharp" authorityPageNumber="418" authorityYear="1885" box="[151,596,727,752]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="9B8CEACC4C22B342FF6BFD03F6D0D794" box="[151,389,727,752]" italics="true" pageId="10" pageNumber="11">Philonthus alternans</emphasis>
<treatmentCitation id="285910CF4C22B342FE6CFD03F501D794" author="" box="[400,596,727,752]" page="418" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE6CFD03F54DD794" box="[400,536,727,752]" pageId="10" pageNumber="11" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 418
</treatmentCitation>
</taxonomicName>
. Type locality: Mexico: Jalapa, Oaxaca. Type depository: BMNH 3 ST; FMNH 1ST. Distribution: Mexico. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C22B342FC9AFD28F2C2D670" ID-CoL="8VRYR" authority="(Navarrete-Heredia et al. 2002: 334)" authorityName="Navarrete-Heredia et al." authorityPageNumber="334" authorityYear="2002" box="[870,1431,764,789]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C22B342FC9AFD28F48AD671" box="[870,991,764,789]" italics="true" pageId="10" pageNumber="11">Philonthus</emphasis>
(
<treatmentCitation id="285910CF4C22B342FC0EFD28F2DAD670" author="" box="[1010,1423,764,789]" page="334" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FC0EFD28F207D670" box="[1010,1362,764,789]" pageId="10" pageNumber="11" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C22B342FB33FD29F259D671" box="[1231,1292,764,789]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2002
</bibRefCitation>
: 334
</treatmentCitation>
)
</taxonomicName>
; distribution within Mexico (
<bibRefCitation id="CD694B2F4C22B342FDFDFCF4F5FED65C" box="[513,683,799,824]" pageId="10" pageNumber="11" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFCB3F567D6E4" blockId="10.[151,1436,871,1040]" box="[151,562,871,896]" pageId="10" pageNumber="11">
<emphasis id="9B8CEACC4C22B342FF6BFCB3F7F6D6E4" bold="true" box="[151,163,871,896]" italics="true" pageId="10" pageNumber="11">*</emphasis>
<taxonomicName id="6EF84D5D4C22B342FF5EFCB3F567D6E4" ID-CoL="LC4W" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,562,871,896]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alternatus">
<emphasis id="9B8CEACC4C22B342FF5EFCB3F6FED6E4" bold="true" box="[162,427,871,896]" italics="true" pageId="10" pageNumber="11">Belonuchus alternatus</emphasis>
<bibRefCitation id="CD694B2F4C22B342FE4EFCB3F567D6E4" box="[434,562,871,896]" pageId="10" pageNumber="11" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFC58F3F1D174" blockId="10.[151,1436,871,1040]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFC58F3F1D174" infered="true" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BFC58F538D6C0" ID-CoL="LC4W" authority="Sharp 1885: 430" authorityName="Sharp" authorityPageNumber="430" authorityYear="1885" box="[151,621,908,933]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alternatus">
<emphasis id="9B8CEACC4C22B342FF6BFC58F6CDD6C1" box="[151,408,908,933]" italics="true" pageId="10" pageNumber="11">Belonuchus alternatus</emphasis>
<treatmentCitation id="285910CF4C22B342FE58FC58F538D6C0" author="" box="[420,621,908,933]" page="430" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE58FC58F57AD6C1" box="[420,559,908,933]" pageId="10" pageNumber="11" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 430
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C22B342FD81FC59F5E0D6C1" box="[637,693,909,933]" pageId="10" pageNumber="11">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C22B342FCD6FC58F4F3D6C1" box="[810,934,908,933]" country="Guatemala" name="Guatemala" pageId="10" pageNumber="11">
<collectingCountry id="D1EF764E4C22B342FCD6FC58F4F3D6C1" box="[810,934,908,933]" name="Guatemala" pageId="10" pageNumber="11">Guatemala</collectingCountry>
</collectingRegion>
, near the City, San Geronimo; 
<collectingCountry id="D1EF764E4C22B342FADEFC58F2CDD6C1" box="[1314,1432,908,933]" name="Nicaragua" pageId="10" pageNumber="11">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C22B342FF3BFC7BF66ED6AC" box="[199,315,943,968]" country="Nicaragua" name="Chontales" pageId="10" pageNumber="11">Chontales</collectingRegion>
; 
<collectingRegion id="6B3CF83C4C22B342FEB5FC64F6F0D6A3" box="[329,421,944,967]" country="Panama" name="Panama" pageId="10" pageNumber="11">
<collectingCountry id="D1EF764E4C22B342FEB5FC64F6F0D6A3" box="[329,421,944,967]" name="Panama" pageId="10" pageNumber="11">Panama</collectingCountry>
</collectingRegion>
, Bugaba. 
<typeStatus id="7643887C4C22B342FDE6FC64F507D6AC" box="[538,594,944,968]" pageId="10" pageNumber="11">Type</typeStatus>
depository: BMNH #? ST: several, from all 
<typeStatus id="7643887C4C22B342FB9DFC64F3C4D6AC" box="[1121,1169,944,968]" pageId="10" pageNumber="11">type</typeStatus>
localities [MCP seen]. FMNH 4 ST: 
<collectingRegion id="6B3CF83C4C22B342FE9EFC01F6EBD688" box="[354,446,981,1004]" country="Panama" name="Panama" pageId="10" pageNumber="11">
<collectingCountry id="D1EF764E4C22B342FE9EFC01F6EBD688" box="[354,446,981,1004]" name="Panama" pageId="10" pageNumber="11">Panama</collectingCountry>
</collectingRegion>
, Bugaba. Distribution: 
<collectingRegion id="6B3CF83C4C22B342FD3FFC01F44AD688" box="[707,799,981,1004]" country="Panama" name="Panama" pageId="10" pageNumber="11">
<collectingCountry id="D1EF764E4C22B342FD3FFC01F44AD688" box="[707,799,981,1004]" name="Panama" pageId="10" pageNumber="11">Panama</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C22B342FCD6FC00F4F5D689" box="[810,928,980,1005]" name="Nicaragua" pageId="10" pageNumber="11">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C22B342FC57FC00F37DD689" box="[939,1064,980,1005]" country="Guatemala" name="Guatemala" pageId="10" pageNumber="11">
<collectingCountry id="D1EF764E4C22B342FC57FC00F37DD689" box="[939,1064,980,1005]" name="Guatemala" pageId="10" pageNumber="11">Guatemala</collectingCountry>
</collectingRegion>
. Note: misspelled as 
<taxonomicName id="6EF84D5D4C22B342FAE9FC00F22AD689" ID-CoL="76XW4" authorityName="Sharp" authorityYear="1885" box="[1301,1407,980,1005]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="9B8CEACC4C22B342FAE9FC00F22AD689" box="[1301,1407,980,1005]" italics="true" pageId="10" pageNumber="11">alternans</emphasis>
</taxonomicName>
in error by 
<treatmentCitation id="285910CF4C22B342FEDBFC23F5FBD174" author="" box="[295,686,1015,1040]" page="369" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FEDBFC23F5FBD174" box="[295,686,1015,1040]" pageId="10" pageNumber="11" refId="ref62964" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus. Vol. 57. W. Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer &amp; Schubert (1914: 369)</bibRefCitation>
</treatmentCitation>
(not 
<taxonomicName id="6EF84D5D4C22B342FD15FC23F3CDD174" ID-CoL="76XW4" authority="Sharp 1885: 418" authorityName="Sharp" authorityPageNumber="418" authorityYear="1885" box="[745,1176,1015,1040]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="9B8CEACC4C22B342FD15FC23F486D174" box="[745,979,1015,1040]" italics="true" pageId="10" pageNumber="11">Philonthus alternans</emphasis>
<treatmentCitation id="285910CF4C22B342FC26FC23F3CDD174" author="" box="[986,1176,1015,1040]" page="418" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FC26FC23F30BD174" box="[986,1118,1015,1040]" pageId="10" pageNumber="11" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 418
</treatmentCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFB94F531D13C" blockId="10.[151,1436,1087,1184]" box="[151,612,1087,1112]" pageId="10" pageNumber="11">
* 
<taxonomicName id="6EF84D5D4C22B342FF59FBEBF531D13C" ID-CoL="LC4X" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[165,612,1087,1112]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="altissimus">
<emphasis id="9B8CEACC4C22B342FF59FBEBF6FCD13C" bold="true" box="[165,425,1087,1112]" italics="true" pageId="10" pageNumber="11">Belonuchus altissimus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFBB0F373D1C4" blockId="10.[151,1436,1087,1184]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFBB0F373D1C4" pageId="10" pageNumber="11">
<materialsCitation id="19903C834C22B342FF6BFBB0F377D1C4" ID-GBIF-Occurrence="1915497781" collectionCode="FMNH" country="Colombia" elevation="3800" location="Distribution" pageId="10" pageNumber="11" specimenCount="3" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C22B342FF6BFBB0F5CBD119" ID-CoL="LC4X" authority="Bernhauer 1916 c: 275" authorityName="Bernhauer" authorityPageNumber="275" authorityYear="1916" box="[151,670,1124,1149]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="altissimus">
<emphasis id="9B8CEACC4C22B342FF6BFBB0F6C6D119" box="[151,403,1124,1149]" italics="true" pageId="10" pageNumber="11">Belonuchus altissimus</emphasis>
<treatmentCitation id="285910CF4C22B342FE60FBB0F5CBD119" author="" box="[412,670,1124,1149]" page="275" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE60FBB0F531D119" box="[412,612,1124,1149]" pageId="10" pageNumber="11" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 275
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C22B342FD57FBB1F5B7D119" box="[683,738,1125,1149]" pageId="10" pageNumber="11">Type</typeStatus>
locality: Monte Socorro, West-Cordilleren, West-Columbien [
<elevation id="22D5D1ED4C22B342FF31FB5CF64BD1C4" box="[205,286,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="11" unit="m" value="3800.0">
<quantity id="6E009B3B4C22B342FF31FB5CF64BD1C4" box="[205,286,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="10" pageNumber="11" unit="m" value="3800.0">3800m</quantity>
</elevation>
on 
<typeStatus id="7643887C4C22B342FEB4FB5CF62DD1C4" box="[328,376,1160,1184]" pageId="10" pageNumber="11">type</typeStatus>
label]. 
<typeStatus id="7643887C4C22B342FE31FB5CF550D1C4" box="[461,517,1160,1184]" pageId="10" pageNumber="11">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C22B342FD6CFB5CF5B4D1FB" box="[656,737,1160,1183]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="11">FMNH</collectionCode>
HT. 
<location id="AC2760054C22B342FCE6FB53F4F3D1C4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C22B342FCE6FB53F4F3D1C4" box="[794,934,1159,1184]" country="Colombia" name="Distribution" pageId="10" pageNumber="11">Distribution</location>
: 
<collectingCountry id="D1EF764E4C22B342FC4CFB53F377D1C4" box="[944,1058,1159,1184]" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFB1BF50CD18C" blockId="10.[151,1436,1231,1365]" box="[151,601,1231,1256]" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BFB1BF50CD18C" ID-CoL="LC4Y" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,601,1231,1256]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="9B8CEACC4C22B342FF6BFB1BF6CBD18C" bold="true" box="[151,414,1231,1256]" italics="true" pageId="10" pageNumber="11">Belonuchus altivagans</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFB20F4AED054" blockId="10.[151,1436,1231,1365]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFB20F4AED054" pageId="10" pageNumber="11">
<materialsCitation id="19903C834C22B342FF6BFB20F4A2D054" ID-GBIF-Occurrence="1915497765" collectionCode="FMNH" country="Colombia" elevation="2000" location="ST. Distribution" pageId="10" pageNumber="11" specimenCode="FMNH 2" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C22B342FF6BFB20F5FDD068" ID-CoL="LC4Y" authority="Bernhauer 1916 c: 272" authorityName="Bernhauer" authorityPageNumber="272" authorityYear="1916" box="[151,680,1268,1293]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="9B8CEACC4C22B342FF6BFB20F6CCD069" box="[151,409,1268,1293]" italics="true" pageId="10" pageNumber="11">Belonuchus altivagans</emphasis>
<treatmentCitation id="285910CF4C22B342FE58FB20F5FDD068" author="" box="[420,680,1268,1293]" page="272" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE58FB20F538D069" box="[420,621,1268,1293]" pageId="10" pageNumber="11" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 272
</treatmentCitation>
</taxonomicName>
. 
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locality: West-Columbien: Pacho, 
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</elevation>
, Ost-Cordilleren, West-Columbien. 
<typeStatus id="7643887C4C22B342FE6EFACCF69CD054" box="[402,457,1304,1328]" pageId="10" pageNumber="11">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C22B342FDAAFACCF5EED054" box="[598,699,1304,1328]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="11">FMNH 2</specimenCode>
<location id="AC2760054C22B342FD3EFACCF42ED054" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C22B342FD3EFACCF42ED054" box="[706,891,1303,1328]" country="Colombia" name="ST. Distribution" pageId="10" pageNumber="11">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C22B342FC79FAC3F4A2D054" box="[901,1015,1303,1328]" name="Colombia" pageId="10" pageNumber="11">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C22B342FF6BFAE8F532D0F8" pageId="10" pageNumber="11" type="discussion">
<paragraph id="A94736DE4C22B342FF6BFAE8F465D031" blockId="10.[151,1436,1231,1365]" box="[151,816,1340,1365]" pageId="10" pageNumber="11">
<emphasis id="9B8CEACC4C22B342FF6BFAE8F641D031" bold="true" box="[151,276,1340,1365]" pageId="10" pageNumber="11">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C22B342FEB0FAE8F69AD031" box="[332,463,1340,1365]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C22B342FEB0FAE8F69AD031" box="[332,463,1340,1365]" italics="true" pageId="10" pageNumber="11">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
<paragraph id="A94736DE4C22B342FF6BFA57F532D0F8" blockId="10.[151,1436,1411,1544]" box="[151,615,1411,1437]" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BFA57F532D0F8" authority="(Bernhauer 1921)" authorityName="Bernhauer" authorityYear="1921" baseAuthorityName="Bernhauer" baseAuthorityYear="1921" box="[151,615,1411,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aluticollis">
<emphasis id="9B8CEACC4C22B342FF6BFA57F6CFD0F8" bold="true" box="[151,410,1411,1436]" italics="true" pageId="10" pageNumber="11">Belonuchus aluticollis</emphasis>
(Bernhauer 1921)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C22B343FF6BFA73F38CD59C" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="A94736DE4C22B342FF6BFA73F26FD36C" blockId="10.[151,1436,1411,1544]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BFA73F26FD36C" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BFA73F5C3D0A4" ID-CoL="76XVT" authority="Bernhauer 1921 c: 107" authorityName="Bernhauer" authorityPageNumber="107" authorityYear="1921" box="[151,662,1447,1472]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="aluticollis">
<emphasis id="9B8CEACC4C22B342FF6BFA73F6DFD0A4" box="[151,394,1447,1472]" italics="true" pageId="10" pageNumber="11">Philonthus aluticollis</emphasis>
<treatmentCitation id="285910CF4C22B342FE68FA73F5C3D0A4" author="" box="[404,662,1447,1472]" page="107" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE68FA73F50ED0A4" box="[404,603,1447,1472]" pageId="10" pageNumber="11" refId="ref62483" refString="Bernhauer, M. (1921 c) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article">Bernhauer 1921c</bibRefCitation>
: 107
</treatmentCitation>
</taxonomicName>
. Type locality: Cueva Iturbe, 
<quantity id="6E009B3B4C22B342FC0CFA7CF314D0A4" box="[1008,1089,1448,1472]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="10" pageNumber="11" unit="m" value="3700.0">3700m</quantity>
, Jujuy, Argentina [LT without &quot;Cueva&quot;]. Type depository: FMNH LT ♀, MACN 2 PLT. Distribution: Argentina. Note: comb. nov. ex 
<taxonomicName id="6EF84D5D4C22B342FF3BFA3BF615D36C" ID-CoL="8VRYR" box="[199,320,1519,1544]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C22B342FF3BFA3BF615D36C" box="[199,320,1519,1544]" italics="true" pageId="10" pageNumber="11">Philonthus</emphasis>
</taxonomicName>
and lectotype designation (
<treatmentCitation id="285910CF4C22B342FD88FA3BF40AD36C" author="" box="[628,863,1519,1544]" page="8" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FD88FA3BF40AD36C" box="[628,863,1519,1544]" pageId="10" pageNumber="11" refId="ref64208" refString="Chani-Posse, M. (2010) Revision of the southern South American species of Philonthus Stephens Coleoptera: Staphylinidae. Zootaxa, 2595, 1 - 70." type="journal article">Chani-Posse 2010: 8</bibRefCitation>
</treatmentCitation>
); redescription (
<treatmentCitation id="285910CF4C22B342FBEAFA3BF278D36C" author="" box="[1046,1325,1519,1544]" page="215" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FBEAFA3BF278D36C" box="[1046,1325,1519,1544]" pageId="10" pageNumber="11" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b: 215</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BF9E3F51DD334" blockId="10.[151,1436,1591,1688]" box="[151,584,1591,1616]" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BF9E3F51DD334" ID-CoL="LC52" authority="Fauvel 1891" authorityName="Fauvel" authorityYear="1891" box="[151,584,1591,1616]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="9B8CEACC4C22B342FF6BF9E3F6E0D334" bold="true" box="[151,437,1591,1616]" italics="true" pageId="10" pageNumber="11">Belonuchus amplipennis</emphasis>
<bibRefCitation id="CD694B2F4C22B342FE41F9E3F51DD334" box="[445,584,1591,1616]" pageId="10" pageNumber="11" refId="ref66096" refString="Fauvel, A. (1891) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 11 e memoire: Staphylinides. Revue d'Entomologie, 10, 87 - 127." type="journal article">Fauvel 1891</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BF988F4D0D3FC" blockId="10.[151,1436,1591,1688]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BF988F4D0D3FC" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BF988F5C9D310" ID-CoL="LC52" authority="Fauvel 1891: 117" authorityName="Fauvel" authorityPageNumber="117" authorityYear="1891" box="[151,668,1628,1653]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amplipennis">
<emphasis id="9B8CEACC4C22B342FF6BF988F6E6D311" box="[151,435,1628,1653]" italics="true" pageId="10" pageNumber="11">Belonuchus amplipennis</emphasis>
<treatmentCitation id="285910CF4C22B342FE3FF988F5C9D310" author="" box="[451,668,1628,1653]" page="117" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE3FF988F50ED310" box="[451,603,1628,1653]" pageId="10" pageNumber="11" refId="ref66096" refString="Fauvel, A. (1891) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 11 e memoire: Staphylinides. Revue d'Entomologie, 10, 87 - 127." type="journal article">Fauvel 1891</bibRefCitation>
: 117
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C22B342FD4CF989F5B2D311" box="[688,743,1629,1653]" pageId="10" pageNumber="11">Type</typeStatus>
locality: Colonie Tovar, San Esteban [
<collectingCountry id="D1EF764E4C22B342FB32F988F21ED311" box="[1230,1355,1628,1653]" name="Venezuela" pageId="10" pageNumber="11">Venezuela</collectingCountry>
]. 
<typeStatus id="7643887C4C22B342FA98F989F2C9D311" box="[1380,1436,1629,1653]" pageId="10" pageNumber="11">Type</typeStatus>
depository: IRSNB 2 ST [MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C22B342FCF0F9ABF4D5D3FC" box="[780,896,1663,1688]" name="Venezuela" pageId="10" pageNumber="11">Venezuela</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BF915F505D3BE" blockId="10.[151,1436,1729,1862]" box="[151,592,1729,1754]" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BF915F505D3BE" ID-CoL="LC53" authority="Blackwelder 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,592,1729,1754]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amplus">
<emphasis id="9B8CEACC4C22B342FF6BF915F629D3BE" bold="true" box="[151,380,1729,1754]" italics="true" pageId="10" pageNumber="11">Belonuchus amplus</emphasis>
<bibRefCitation id="CD694B2F4C22B342FE7FF915F505D3BE" box="[387,592,1729,1754]" pageId="10" pageNumber="11" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BF931F6F0D222" blockId="10.[151,1436,1729,1862]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BF931F6F0D222" infered="true" pageId="10" pageNumber="11">
<materialsCitation id="19903C834C22B342FF6BF931F596D246" ID-GBIF-Occurrence="1915497771" country="Saint Lucia" location="St. Remy" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Soufriere" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C22B342FF6BF931F5DCD39A" ID-CoL="LC53" authority="Blackwelder 1943: 425" authorityName="Blackwelder" authorityPageNumber="425" authorityYear="1943" box="[151,649,1765,1790]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amplus">
<emphasis id="9B8CEACC4C22B342FF6BF931F621D39A" box="[151,372,1765,1790]" italics="true" pageId="10" pageNumber="11">Belonuchus amplus</emphasis>
<treatmentCitation id="285910CF4C22B342FE80F931F5DCD39A" author="" box="[380,649,1765,1790]" page="425" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE80F931F51AD39A" box="[380,591,1765,1790]" pageId="10" pageNumber="11" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 425
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C22B342FD6AF932F59BD39A" box="[662,718,1766,1790]" pageId="10" pageNumber="11">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C22B342FCC0F932F4F2D39A" box="[828,935,1765,1790]" name="Saint Lucia" pageId="10" pageNumber="11">St. Lucia</collectingCountry>
: 
<quantity id="6E009B3B4C22B342FC48F931F347D39A" box="[948,1042,1765,1790]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="10" pageNumber="11" unit="mi" value="0.5">0.5 mile</quantity>
northwest of 
<location id="AC2760054C22B342FB4FF932F248D39A" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C22B342FB4FF932F248D39A" box="[1203,1309,1766,1790]" country="Saint Lucia" name="St. Remy" pageId="10" pageNumber="11" stateProvince="Soufriere">St. Remy</location>
on road to 
<collectingRegion id="6B3CF83C4C22B342FF3BF8DDF67AD246" box="[199,303,1801,1826]" country="Saint Lucia" name="Soufriere" pageId="10" pageNumber="11">Soufrière</collectingRegion>
(
<quantity id="6E009B3B4C22B342FEBEF8DDF6FBD246" box="[322,430,1801,1826]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" pageId="10" pageNumber="11" unit="mi" value="2.5">2.5 miles</quantity>
southeast of 
<collectingRegion id="6B3CF83C4C22B342FDB1F8DDF5EED246" box="[589,699,1801,1826]" country="Saint Lucia" name="Soufriere" pageId="10" pageNumber="11">Soufrière</collectingRegion>
)
</materialsCitation>
. 
<materialsCitation id="19903C834C22B342FD2DF8DEF6F4D222" ID-GBIF-Occurrence="1915497770" collectionCode="USNM" country="Saint Lucia" location="Lesser Antilles" municipality="Distribution" pageId="10" pageNumber="11" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<typeStatus id="7643887C4C22B342FD2DF8DEF45DD246" box="[721,776,1802,1826]" pageId="10" pageNumber="11">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C22B342FC67F8DEF4B9D246" box="[923,1004,1802,1826]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
HT 
<specimenCount id="BFFEFD574C22B342FBDBF8DEF369D245" box="[1063,1084,1802,1825]" pageId="10" pageNumber="11" type="female">♀</specimenCount>
, 1 PT 
<specimenCount id="BFFEFD574C22B342FB6EF8DEF3FDD245" box="[1170,1192,1802,1825]" pageId="10" pageNumber="11" type="female">♀</specimenCount>
. 
<collectingMunicipality id="4923ACA44C22B342FB45F8DDF210D246" box="[1209,1349,1801,1826]" pageId="10" pageNumber="11">Distribution</collectingMunicipality>
: 
<location id="AC2760054C22B342FAAFF8DEF675D222" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C22B342FAAFF8DEF675D222" country="Saint Lucia" municipality="Distribution" name="Lesser Antilles" pageId="10" pageNumber="11">Lesser Antilles</location>
(
<collectingCountry id="D1EF764E4C22B342FED3F8FAF6CCD222" box="[303,409,1837,1862]" name="Saint Lucia" pageId="10" pageNumber="11">St. Lucia</collectingCountry>
)
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BF8BAF515D2E3" blockId="10.[151,1436,1902,2035]" box="[151,576,1902,1927]" pageId="10" pageNumber="11">
<taxonomicName id="6EF84D5D4C22B342FF6BF8BAF515D2E3" ID-CoL="LC55" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,576,1902,1927]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="9B8CEACC4C22B342FF6BF8BAF6D0D2E3" bold="true" box="[151,389,1902,1927]" italics="true" pageId="10" pageNumber="11">Belonuchus andinus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C22B342FF6BF846F519D297" blockId="10.[151,1436,1902,2035]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="89E811F04C22B342FF6BF846F519D297" infered="true" pageId="10" pageNumber="11">
<materialsCitation id="19903C834C22B342FF6BF846F474D2AB" ID-GBIF-Occurrence="1915497717" collectionCode="FMNH, ZMUH" country="Bolivia" location="Bolivien" pageId="10" pageNumber="11" specimenCode="FMNH 1, ZMUH 1" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C22B342FF6BF846F52BD2CF" ID-CoL="LC55" authority="Bernhauer 1916 c: 278" authorityName="Bernhauer" authorityPageNumber="278" authorityYear="1916" box="[151,638,1938,1963]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="9B8CEACC4C22B342FF6BF846F62ED2CF" box="[151,379,1938,1963]" italics="true" pageId="10" pageNumber="11">Belonuchus andinus</emphasis>
<treatmentCitation id="285910CF4C22B342FE7FF846F52BD2CF" author="" box="[387,638,1938,1963]" page="278" pageId="10" pageNumber="11">
<bibRefCitation id="CD694B2F4C22B342FE7FF846F51CD2CF" box="[387,585,1938,1963]" pageId="10" pageNumber="11" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 278
</treatmentCitation>
</taxonomicName>
(attributed to Fauvel 
<emphasis id="9B8CEACC4C22B342FC8AF840F4EFD2CF" box="[886,954,1938,1963]" italics="true" pageId="10" pageNumber="11">in litt.</emphasis>
). 
<typeStatus id="7643887C4C22B342FC2FF847F35ED2CF" box="[979,1035,1939,1963]" pageId="10" pageNumber="11">Type</typeStatus>
locality: Yuracarès, 
<collectingCountry id="D1EF764E4C22B342FB09F846F20CD2CF" box="[1269,1369,1938,1963]" name="Bolivia" pageId="10" pageNumber="11">Bolivien</collectingCountry>
. 
<typeStatus id="7643887C4C22B342FA98F847F2C9D2CF" box="[1380,1436,1939,1963]" pageId="10" pageNumber="11">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C22B342FEB0F863F6E7D2AB" box="[332,434,1975,1999]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="11">FMNH 1</specimenCode>
ST; 
<specimenCode id="F95E9EA54C22B342FE1AF863F51CD2AB" box="[486,585,1975,1999]" collectionCode="ZMUH" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="11">ZMUH 1</specimenCode>
+ST? [destroyed?]
</materialsCitation>
; 
<materialsCitation id="19903C834C22B342FCD0F863F519D297" ID-GBIF-Occurrence="1915497790" collectionCode="MCP" country="Bolivia" location="Distribution" pageId="10" pageNumber="11" specimenCode="IRSNB 2" specimenCount="1" typeStatus="holotype">
<specimenCode id="F95E9EA54C22B342FCD0F863F4C1D2AB" box="[812,916,1975,1999]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="11">IRSNB 2</specimenCode>
+3? ST: only two of them in Bernhauer´s hand [
<collectionCode id="CFE9AE1B4C22B342FF31F80FF65ED297" box="[205,267,2011,2035]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="10" pageNumber="11">MCP</collectionCode>
seen]. 
<location id="AC2760054C22B342FEA7F80EF6B2D297" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C22B342FEA7F80EF6B2D297" box="[347,487,2010,2035]" country="Bolivia" name="Distribution" pageId="10" pageNumber="11">Distribution</location>
: 
<collectingCountry id="D1EF764E4C22B342FE0DF80EF512D297" box="[497,583,2010,2035]" name="Bolivia" pageId="10" pageNumber="11">Bolivia</collectingCountry>
.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFF43F52DD5D4" blockId="11.[151,1436,151,285]" box="[151,632,151,176]" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BFF43F52DD5D4" ID-CoL="LC56" authority="Scheerpeltz 1960" authorityName="Scheerpeltz" authorityYear="1960" box="[151,632,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="9B8CEACC4C23B343FF6BFF43F6FAD5D4" bold="true" box="[151,431,151,176]" italics="true" pageId="11" pageNumber="12">Belonuchus angusticeps</emphasis>
<bibRefCitation id="CD694B2F4C23B343FE4AFF43F52DD5D4" box="[438,632,151,176]" pageId="11" pageNumber="12" refId="ref71794" refString="Scheerpeltz, O. (1960) Zur Kenntnis neotropischer Staphyliniden (Col.). Beitrage zur Neotropischen Fauna, 2 (2), 65 - 138. https: // doi. org / 10.1080 / 01650526009380624" type="journal article">Scheerpeltz 1960</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFF68F38CD59C" blockId="11.[151,1436,151,285]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BFF68F38CD59C" pageId="11" pageNumber="12">
<materialsCitation id="19903C834C23B343FF6BFF68F6BAD59C" ID-GBIF-Occurrence="1915497756" collectorName="in den Yungas de Palmar" country="Bolivia" elevation="2000" location="Yungas de Palmar" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Cochabamba" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C23B343FF6BFF68F43BD5B0" ID-CoL="LC56" authority="Scheerpeltz 1960: 117" authorityName="Scheerpeltz" authorityPageNumber="117" authorityYear="1960" box="[151,878,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angusticeps" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C23B343FF6BFF68F50FD5B0" box="[151,602,188,213]" italics="true" pageId="11" pageNumber="12">Belonuchus (Musicoderus) angusticeps</emphasis>
<treatmentCitation id="285910CF4C23B343FD9BFF68F43BD5B0" author="" box="[615,878,188,213]" page="117" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FD9BFF68F467D5B1" box="[615,818,188,213]" pageId="11" pageNumber="12" refId="ref71794" refString="Scheerpeltz, O. (1960) Zur Kenntnis neotropischer Staphyliniden (Col.). Beitrage zur Neotropischen Fauna, 2 (2), 65 - 138. https: // doi. org / 10.1080 / 01650526009380624" type="journal article">Scheerpeltz 1960</bibRefCitation>
: 117
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FC83FF69F4E3D5B1" box="[895,950,189,213]" pageId="11" pageNumber="12">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C23B343FBD0FF68F3EBD5B1" box="[1068,1214,188,213]" country="Bolivia" name="Cochabamba" pageId="11" pageNumber="12">Cochabamba</collectingRegion>
(
<collectingCountry id="D1EF764E4C23B343FB2EFF68F26DD5B1" box="[1234,1336,188,213]" name="Bolivia" pageId="11" pageNumber="12">Bolivien</collectingCountry>
), 
<collectorName id="040D53084C23B343FAADFF68F6C6D59C" pageId="11" pageNumber="12">
in den 
<location id="AC2760054C23B343FF3BFF34F6C6D59C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FF3BFF34F6C6D59C" box="[199,403,223,248]" country="Bolivia" name="Yungas de Palmar" pageId="11" pageNumber="12" stateProvince="Cochabamba">Yungas de Palmar</location>
</collectorName>
, 
<elevation id="22D5D1ED4C23B343FE62FF34F6BAD59C" box="[414,495,224,248]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="2000.0">
<quantity id="6E009B3B4C23B343FE62FF34F6BAD59C" box="[414,495,224,248]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="2000.0">2000m</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C23B343FE06FF34F381D59C" ID-GBIF-Occurrence="1915497778" box="[506,1236,223,248]" collectionCode="NMW, FMNH" country="Bolivia" location="ST. Distribution" pageId="11" pageNumber="12" specimenCode="NMW 2, FMNH 2" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C23B343FE06FF34F567D59C" box="[506,562,224,248]" pageId="11" pageNumber="12">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C23B343FD41FF34F44FD59C" box="[701,794,224,248]" collectionCode="NMW" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW 2</specimenCode>
ST; 
<specimenCode id="F95E9EA54C23B343FCB3FF34F4E0D59C" box="[847,949,224,248]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH 2</specimenCode>
<location id="AC2760054C23B343FC40FF34F321D59C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FC40FF34F321D59C" box="[956,1140,223,248]" country="Bolivia" name="ST. Distribution" pageId="11" pageNumber="12">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C23B343FB82FF0BF381D59C" box="[1150,1236,223,248]" name="Bolivia" pageId="11" pageNumber="12">Bolivia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C23B343FF6BFED0F465D479" box="[151,816,260,285]" pageId="11" pageNumber="12" type="discussion">
<paragraph id="A94736DE4C23B343FF6BFED0F465D479" blockId="11.[151,1436,151,285]" box="[151,816,260,285]" pageId="11" pageNumber="12">
<emphasis id="9B8CEACC4C23B343FF6BFED0F641D479" bold="true" box="[151,276,260,285]" pageId="11" pageNumber="12">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C23B343FEB0FED0F69AD479" box="[332,463,260,285]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C23B343FEB0FED0F69AD479" box="[332,463,260,285]" italics="true" pageId="11" pageNumber="12">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C23B344FF6BFE9FF3D0D759" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="A94736DE4C23B343FF6BFE9FF5D6D401" blockId="11.[151,1437,331,429]" box="[151,643,331,357]" pageId="11" pageNumber="12">
<emphasis id="9B8CEACC4C23B343FF6BFE9FF7F6D400" bold="true" box="[151,163,331,356]" italics="true" pageId="11" pageNumber="12">*</emphasis>
<taxonomicName id="6EF84D5D4C23B343FF5EFE9FF5D6D401" ID-CoL="LC57" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[162,643,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="9B8CEACC4C23B343FF5EFE9FF69DD400" bold="true" box="[162,456,331,356]" italics="true" pageId="11" pageNumber="12">Belonuchus angusticollis</emphasis>
<bibRefCitation id="CD694B2F4C23B343FE33FE98F5D6D401" box="[463,643,332,357]" pageId="11" pageNumber="12" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFEBBF593D4C8" blockId="11.[151,1437,331,429]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BFEBBF593D4C8" pageId="11" pageNumber="12">
<materialsCitation id="19903C834C23B343FF6BFEBBF310D4E3" ID-GBIF-Occurrence="1915497794" box="[151,1093,367,392]" country="Peru" location="Chanchamayo" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C23B343FF6BFEBBF5F8D4EC" ID-CoL="LC57" authority="Bernhauer 1906: 330" authorityName="Bernhauer" authorityPageNumber="330" authorityYear="1906" box="[151,685,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="9B8CEACC4C23B343FF6BFEBBF6E4D4EC" box="[151,433,367,392]" italics="true" pageId="11" pageNumber="12">Belonuchus angusticollis</emphasis>
<treatmentCitation id="285910CF4C23B343FE46FEBBF5F8D4EC" author="" box="[442,685,367,392]" page="330" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE46FEBBF526D4EC" box="[442,627,367,392]" pageId="11" pageNumber="12" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 330
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD45FEA4F5A4D4EC" box="[697,753,368,392]" pageId="11" pageNumber="12">Type</typeStatus>
locality: 
<location id="AC2760054C23B343FCA2FEBBF354D4EC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FCA2FEBBF354D4EC" box="[862,1025,367,392]" country="Peru" name="Chanchamayo" pageId="11" pageNumber="12">Chanchamayo</location>
, 
<collectingCountry id="D1EF764E4C23B343FBF2FEA4F310D4E3" box="[1038,1093,368,391]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C23B343FBADFEA4F6C1D4C8" ID-GBIF-Occurrence="1915497755" collectionCode="FMNH" country="Peru" location="Distribution" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C23B343FBADFEA4F3DCD4EC" box="[1105,1161,368,392]" pageId="11" pageNumber="12">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C23B343FAE4FEA4F23CD4E3" box="[1304,1385,368,391]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH</collectionCode>
HT. 
<location id="AC2760054C23B343FF3BFE40F606D4C9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FF3BFE40F606D4C9" box="[199,339,404,429]" country="Peru" name="Distribution" pageId="11" pageNumber="12">Distribution</location>
: 
<collectingCountry id="D1EF764E4C23B343FEA1FE41F6C1D4C8" box="[349,404,405,428]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
</materialsCitation>
. Note: Peru (
<bibRefCitation id="CD694B2F4C23B343FDD9FE40F5EED4C8" box="[549,699,404,429]" pageId="11" pageNumber="12" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFE0FF534D490" blockId="11.[151,1437,475,645]" box="[151,609,475,501]" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BFE0FF534D490" ID-CoL="LC59" authority="Blackwelder 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,609,475,501]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antiguae">
<emphasis id="9B8CEACC4C23B343FF6BFE0FF6D8D490" bold="true" box="[151,397,475,500]" italics="true" pageId="11" pageNumber="12">Belonuchus antiguae</emphasis>
<bibRefCitation id="CD694B2F4C23B343FE68FE08F534D490" box="[404,609,476,501]" pageId="11" pageNumber="12" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFE2BF64AD7E1" blockId="11.[151,1437,475,645]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BFE2BF64AD7E1" infered="true" pageId="11" pageNumber="12">
<materialsCitation id="19903C834C23B343FF6BFE2BF6A1D759" ID-GBIF-Occurrence="1915497748" country="Antigua and Barbuda" elevation="396" location="Boggy Peak" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Mary" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C23B343FF6BFE2BF5C8D77C" ID-CoL="LC59" authority="Blackwelder 1943: 434" authorityName="Blackwelder" authorityPageNumber="434" authorityYear="1943" box="[151,669,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="antiguae">
<emphasis id="9B8CEACC4C23B343FF6BFE2BF6D0D77C" box="[151,389,511,536]" italics="true" pageId="11" pageNumber="12">Belonuchus antiguae</emphasis>
<treatmentCitation id="285910CF4C23B343FE72FE2BF5C8D77C" author="" box="[398,669,511,536]" page="434" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE72FE2BF534D77C" box="[398,609,511,536]" pageId="11" pageNumber="12" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 434
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD55FDD4F5B4D77C" box="[681,737,512,536]" pageId="11" pageNumber="12">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C23B343FCACFE2BF4FAD77C" box="[848,943,511,536]" name="Antigua and Barbuda" pageId="11" pageNumber="12">Antigua</collectingCountry>
: near summit of 
<location id="AC2760054C23B343FB88FDD4F254D77C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FB88FDD4F254D77C" box="[1140,1281,511,536]" country="Antigua and Barbuda" name="Boggy Peak" pageId="11" pageNumber="12" stateProvince="Mary">Boggy Peak</location>
, parish of St. 
<collectingRegion id="6B3CF83C4C23B343FF3BFDF1F650D759" box="[199,261,549,573]" country="Turkmenistan" name="Mary Welayaty" pageId="11" pageNumber="12">Mary</collectingRegion>
, 
<elevation id="22D5D1ED4C23B343FEEFFDF0F6A1D759" box="[275,500,548,573]" metricMagnitude="2" metricUnit="m" metricValue="3.9624" pageId="11" pageNumber="12" unit="ft" value="1300.0">
elevation 
<quantity id="6E009B3B4C23B343FE7FFDF0F6A1D759" box="[387,500,548,573]" metricMagnitude="2" metricUnit="m" metricValue="3.9624" pageId="11" pageNumber="12" unit="ft" value="1300.0">1300 feet</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C23B343FDFCFDF1F64AD7E1" ID-GBIF-Occurrence="1915497753" collectionCode="USNM" country="Antigua and Barbuda" location="Lesser Antilles" municipality="Distribution" pageId="11" pageNumber="12" specimenCode="BMNH 1, FMNH 1" specimenCount="1" specimenCount-male="1" stateProvince="Northern" typeStatus="holotype">
<typeStatus id="7643887C4C23B343FDFCFDF1F56DD759" box="[512,568,549,573]" pageId="11" pageNumber="12">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C23B343FD36FDF0F44ED758" box="[714,795,548,572]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12">USNM</collectionCode>
HT 
<specimenCount id="BFFEFD574C23B343FCA8FDF1F43ED758" box="[852,875,549,572]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
, 5 PT; 
<specimenCode id="F95E9EA54C23B343FC3FFDF1F365D758" box="[963,1072,548,572]" collectionCode="BMNH" pageId="11" pageNumber="12">BMNH 1</specimenCode>
PT; 
<specimenCode id="F95E9EA54C23B343FB96FDF1F386D758" box="[1130,1235,548,572]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH 1</specimenCode>
PT. 
<collectingMunicipality id="4923ACA44C23B343FAF1FDF0F2CCD759" box="[1293,1433,548,573]" pageId="11" pageNumber="12">Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C23B343FF3BFD93F670D704" box="[199,293,583,608]" name="Antigua and Barbuda" pageId="11" pageNumber="12">Antigua</collectingCountry>
. Notes: distribution within 
<location id="AC2760054C23B343FD9DFD9CF45BD704" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FD9DFD9CF45BD704" box="[609,782,583,608]" country="Antigua and Barbuda" municipality="Distribution" name="Lesser Antilles" pageId="11" pageNumber="12" stateProvince="Northern">Lesser Antilles</location>
(
<collectingRegion id="6B3CF83C4C23B343FCE2FD93F4D0D704" box="[798,901,583,608]" country="Sri Lanka" name="Northern" pageId="11" pageNumber="12">Northern</collectingRegion>
Leeward Is.) (
<bibRefCitation id="CD694B2F4C23B343FBCDFD93F3EED704" box="[1073,1211,583,608]" pageId="11" pageNumber="12" refId="ref71064" refString="Peck, S. B. (2011 a) The diversity and distributions of the beetles (Insecta: Coleoptera) of the northern Leeward Islands, Lesser Antilles (Anguilla, Antigua, Barbuda, Nevis, Saba, St. Barthelemy, St. Eustatius, St. Kitts, and St. Martin). Insecta Mundi, 0 159, 1 - 54." type="journal article">Peck 2011a</bibRefCitation>
); generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFD67F536D7A9" blockId="11.[151,1437,691,824]" box="[151,611,691,717]" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BFD67F536D7A9" ID-CoL="LC5B" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,611,691,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apicicornis">
<emphasis id="9B8CEACC4C23B343FF6BFD67F6FDD7A8" bold="true" box="[151,424,691,716]" italics="true" pageId="11" pageNumber="12">Belonuchus apicicornis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFD03F686D65C" blockId="11.[151,1437,691,824]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BFD03F686D65C" infered="true" pageId="11" pageNumber="12">
<materialsCitation id="19903C834C23B343FF6BFD03F685D65C" ID-GBIF-Occurrence="1915497763" collectionCode="FMNH" collectorName="West-Columbien &amp; S. Antonio" country="Colombia" county="ST. Distribution" elevation="1800" location="Rio Aquatal" municipality="Rio Vitaco" pageId="11" pageNumber="12" specimenCode="FMNH 2" specimenCount="2" stateProvince="Cauca" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C23B343FF6BFD03F418D794" ID-CoL="LC5B" authority="Bernhauer 1916 c: 270" authorityName="Bernhauer" authorityPageNumber="270" authorityYear="1916" box="[151,845,727,752]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apicicornis" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C23B343FF6BFD03F51CD794" box="[151,585,727,752]" italics="true" pageId="11" pageNumber="12">Belonuchus (Musicoderus) apicicornis</emphasis>
<treatmentCitation id="285910CF4C23B343FDADFD03F418D794" author="" box="[593,845,727,752]" page="270" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FDADFD03F443D794" box="[593,790,727,752]" pageId="11" pageNumber="12" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 270
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FCA4FD0CF4C5D794" box="[856,912,728,752]" pageId="11" pageNumber="12">Type</typeStatus>
locality: 
<collectorName id="040D53084C23B343FC07FD03F3E8D794" box="[1019,1213,727,752]" pageId="11" pageNumber="12">West-Columbien</collectorName>
: Cali am Rio 
<collectingRegion id="6B3CF83C4C23B343FAAAFD0CF2C9D794" box="[1366,1436,728,752]" country="Colombia" name="Cauca" pageId="11" pageNumber="12">Cauca</collectingRegion>
(
<location id="AC2760054C23B343FF32FD28F634D671" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FF32FD28F634D671" box="[206,353,764,789]" country="Colombia" county="ST. Distribution" municipality="Rio Vitaco" name="Rio Aquatal" pageId="11" pageNumber="12" stateProvince="Cauca">Rio Aquatal</location>
, 
<elevation id="22D5D1ED4C23B343FE8EFD28F696D670" box="[370,451,764,788]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="12" unit="m" value="1800.0">
<quantity id="6E009B3B4C23B343FE8EFD28F696D670" box="[370,451,764,788]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="12" unit="m" value="1800.0">1800m</quantity>
</elevation>
; 
<collectorName id="040D53084C23B343FE28FD28F502D671" box="[468,599,764,789]" pageId="11" pageNumber="12">S. Antonio</collectorName>
, 
<elevation id="22D5D1ED4C23B343FD9BFD28F5ECD670" box="[615,697,764,788]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="2000.0">
<quantity id="6E009B3B4C23B343FD9BFD28F5ECD670" box="[615,697,764,788]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="2000.0">2000m</quantity>
</elevation>
; 
<collectingMunicipality id="4923ACA44C23B343FD36FD28F41ED671" box="[714,843,764,789]" pageId="11" pageNumber="12">Rio Vitaco</collectingMunicipality>
, 
<elevation id="22D5D1ED4C23B343FCA0FD28F4FAD670" box="[860,943,764,788]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="12" unit="m" value="2200.0">
<quantity id="6E009B3B4C23B343FCA0FD28F4FAD670" box="[860,943,764,788]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="12" unit="m" value="2200.0">2200m</quantity>
</elevation>
). 
<typeStatus id="7643887C4C23B343FC3BFD29F4ABD671" box="[967,1022,765,789]" pageId="11" pageNumber="12">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C23B343FB6AFD29F257D670" box="[1174,1282,764,788]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH 2</specimenCode>
ST;? 1 
<collectingCounty id="40264E524C23B343FA8BFD28F606D65C" pageId="11" pageNumber="12">ST. Distribution</collectingCounty>
: 
<collectingCountry id="D1EF764E4C23B343FEA1FCCBF685D65C" box="[349,464,799,824]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFCB3F503D6E4" blockId="11.[151,1437,871,1077]" box="[151,598,871,896]" pageId="11" pageNumber="12">
<emphasis id="9B8CEACC4C23B343FF6BFCB3F7F6D6E4" bold="true" box="[151,163,871,896]" italics="true" pageId="11" pageNumber="12">*</emphasis>
<taxonomicName id="6EF84D5D4C23B343FF5EFCB3F503D6E4" ID-CoL="LC5C" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,598,871,896]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apiciventris">
<emphasis id="9B8CEACC4C23B343FF5EFCB3F6E9D6E4" bold="true" box="[162,444,871,896]" italics="true" pageId="11" pageNumber="12">Belonuchus apiciventris</emphasis>
(
<bibRefCitation id="CD694B2F4C23B343FE36FCB3F518D6E4" box="[458,589,871,896]" pageId="11" pageNumber="12" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFC58F62FD151" blockId="11.[151,1437,871,1077]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BFC58F62FD151" infered="true" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BFC58F52BD6C0" ID-CoL="76XHQ" authority="Sharp 1885: 419" authorityName="Sharp" authorityPageNumber="419" authorityYear="1885" box="[151,638,908,933]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="apiciventris">
<emphasis id="9B8CEACC4C23B343FF6BFC58F6F6D6C0" box="[151,419,908,933]" italics="true" pageId="11" pageNumber="12">Philonthus apiciventris</emphasis>
<treatmentCitation id="285910CF4C23B343FE4EFC58F52BD6C0" author="" box="[434,638,908,933]" page="419" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE4EFC58F56AD6C1" box="[434,575,908,933]" pageId="11" pageNumber="12" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 419
</treatmentCitation>
</taxonomicName>
. Type locality: Mexico: Juquila. Type depository: BMNH HT ♂. Distribution: Mexico. Notes: in cave collections (
<bibRefCitation id="CD694B2F4C23B343FCDAFC7BF274D6AC" box="[806,1313,943,968]" pageId="11" pageNumber="12" refId="ref70079" refString="Navarrete-Heredia, J. L. &amp; Marquez Luna, J. (1993) Notas sobre los Staphylinidae (Insecta: Coleoptera) de cuevas en Mexico. Mundos Subterraneos, 4, 43 - 52." type="journal article">Navarrete-Heredia &amp; Márquez-Luna 1993</bibRefCitation>
); cited as 
<emphasis id="9B8CEACC4C23B343FF3BFC01F607D688" box="[199,338,981,1004]" italics="true" pageId="11" pageNumber="12">apaciventris</emphasis>
, characters. habits, natural history notes, host mushrooms (Navarrete-Heredia 1995); comb. nov. ex 
<taxonomicName id="6EF84D5D4C23B343FF11FC23F58CD174" ID-CoL="8VRYR" authority="(Jimenez-Sanchez et al. 2000)" authorityName="Jimenez-Sanchez et al." authorityYear="2000" box="[237,729,1015,1040]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C23B343FF11FC23F633D174" box="[237,358,1015,1040]" italics="true" pageId="11" pageNumber="12">Philonthus</emphasis>
(
<bibRefCitation id="CD694B2F4C23B343FE87FC23F587D174" box="[379,722,1015,1040]" pageId="11" pageNumber="12" refId="ref67947" refString="Jimenez-Sanchez, E., Navarrete-Heredia, J. L. &amp; Padilla-Ramirez, J. R. (2000) Estafilinidos (Coleoptera: Staphylinidae) necrofilos de la Sierra de Nanchititla, Estado de Mexico, Mexico. Folia Entomologica Mexicana, 108, 53 - 78." type="journal article">
Jiménez-Sánchez 
<emphasis id="9B8CEACC4C23B343FDB7FC2DF5DCD174" box="[587,649,1015,1040]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2000
</bibRefCitation>
)
</taxonomicName>
; distribution within Mexico (
<bibRefCitation id="CD694B2F4C23B343FBC9FC23F2CDD174" box="[1077,1432,1015,1040]" pageId="11" pageNumber="12" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C23B343FAEFFC2DF204D174" box="[1299,1361,1015,1040]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2002
</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C23B343FF3BFBC8F63BD151" box="[199,366,1052,1077]" pageId="11" pageNumber="12" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFBB7F50AD119" blockId="11.[151,1436,1123,1328]" box="[151,607,1123,1149]" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BFBB7F50AD119" ID-CoL="68GGN" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,607,1123,1149]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinus">
<emphasis id="9B8CEACC4C23B343FF6BFBB7F6F1D118" bold="true" box="[151,420,1123,1148]" italics="true" pageId="11" pageNumber="12">Belonuchus argentinus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFB53F6C1D054" blockId="11.[151,1436,1123,1328]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BFB53F6C1D054" infered="true" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BFB53F5CBD1C4" ID-CoL="68GGN" authority="Bernhauer 1916 c: 278" authorityName="Bernhauer" authorityPageNumber="278" authorityYear="1916" box="[151,670,1159,1184]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="argentinus">
<emphasis id="9B8CEACC4C23B343FF6BFB53F6CDD1C4" box="[151,408,1159,1184]" italics="true" pageId="11" pageNumber="12">Belonuchus argentinus</emphasis>
<treatmentCitation id="285910CF4C23B343FE5CFB53F5CBD1C4" author="" box="[416,670,1159,1184]" page="278" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE5CFB53F530D1C4" box="[416,613,1159,1184]" pageId="11" pageNumber="12" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 278
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD55FB5CF5B4D1C4" box="[681,737,1160,1184]" pageId="11" pageNumber="12">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C23B343FCB0FB53F4E3D1C4" box="[844,950,1159,1184]" country="Paraguay" name="Misiones" pageId="11" pageNumber="12">Misiones</collectingRegion>
, 
<collectingCountry id="D1EF764E4C23B343FC3DFB53F361D1C4" box="[961,1076,1159,1184]" name="Argentina" pageId="11" pageNumber="12">Argentina</collectingCountry>
. 
<typeStatus id="7643887C4C23B343FBBCFB5CF322D1C4" box="[1088,1143,1160,1184]" pageId="11" pageNumber="12">Type</typeStatus>
depositary: FMNH HT, 2 PT; MLPA 8 PT (as ST in error, 
<bibRefCitation id="CD694B2F4C23B343FDCEFB78F47CD1A0" box="[562,809,1196,1221]" pageId="11" pageNumber="12" refId="ref66201" refString="Fernandez, L. A., Cabrera, N., Chani-Posse, M. R. &amp; Gabellone, C. (2007) Catalog of the types of Staphylinidae Insecta, Coleoptera deposited at Museo de La Plata Argentina. Zootaxa, 1611, 25 - 44." type="journal article">
Fernandez 
<emphasis id="9B8CEACC4C23B343FD51FB79F5B0D1A1" box="[685,741,1196,1221]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2007
</bibRefCitation>
). Distribution: 
<collectingCountry id="D1EF764E4C23B343FC2DFB78F316D1A1" box="[977,1091,1196,1221]" name="Argentina" pageId="11" pageNumber="12">Argentina</collectingCountry>
, 
<collectingCountry id="D1EF764E4C23B343FBB2FB78F3C0D1A1" box="[1102,1173,1196,1221]" name="Brazil" pageId="11" pageNumber="12">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C23B343FB5CFB78F3A3D1A1" box="[1184,1270,1196,1221]" name="Bolivia" pageId="11" pageNumber="12">Bolivia</collectingCountry>
. Notes: MLPA &quot;ST&quot;, 
<bibRefCitation id="CD694B2F4C23B343FEF2FB1BF55FD18C" box="[270,522,1231,1256]" pageId="11" pageNumber="12" refId="ref66201" refString="Fernandez, L. A., Cabrera, N., Chani-Posse, M. R. &amp; Gabellone, C. (2007) Catalog of the types of Staphylinidae Insecta, Coleoptera deposited at Museo de La Plata Argentina. Zootaxa, 1611, 25 - 44." type="journal article">
Fernandez 
<emphasis id="9B8CEACC4C23B343FE70FB05F693D18C" box="[396,454,1231,1256]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2007
</bibRefCitation>
, but original reference to &quot;Das typische Stück …&quot;; name cited by 
<treatmentCitation id="285910CF4C23B343FAFAFB1BF7AFD069" author="" page="503" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FAFAFB1BF7AFD069" pageId="11" pageNumber="12" refId="ref63589" refString="Bruch, C. (1915) Catalogo sistematico de los coleopteros de la Republica Argentina. II. Revista del Museo de La Plata, 19, 471 - 526." type="journal article">Bruch (1915: 503)</bibRefCitation>
</treatmentCitation>
as nomen nudum, attributed to Bernhauer 
<emphasis id="9B8CEACC4C23B343FD19FB21F418D069" box="[741,845,1268,1293]" italics="true" pageId="11" pageNumber="12">in litteris</emphasis>
; distribution in 
<collectingCountry id="D1EF764E4C23B343FBFAFB20F31FD069" box="[1030,1098,1268,1293]" name="Brazil" pageId="11" pageNumber="12">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C23B343FB49FB21F3BAD069" box="[1205,1263,1268,1293]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2013); generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFA8BF549D01C" blockId="11.[151,1437,1375,1472]" box="[151,540,1375,1400]" pageId="11" pageNumber="12">
<emphasis id="9B8CEACC4C23B343FF6BFA8BF7F6D01C" bold="true" box="[151,163,1375,1400]" italics="true" pageId="11" pageNumber="12">*</emphasis>
<taxonomicName id="6EF84D5D4C23B343FF5EFA8BF549D01C" ID-CoL="LC5G" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[162,540,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="9B8CEACC4C23B343FF5EFA8BF6C0D013" bold="true" box="[162,405,1375,1400]" italics="true" pageId="11" pageNumber="12">Belonuchus armatus</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFA50F493D0A4" blockId="11.[151,1437,1375,1472]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BFA50F493D0A4" pageId="11" pageNumber="12">
<materialsCitation id="19903C834C23B343FF6BFA50F6CAD0A4" ID-GBIF-Occurrence="1915497704" collectionCode="BMNH" country="Brazil" location="Distribution" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C23B343FF6BFA50F50ED0F9" ID-CoL="LC5G" authority="Sharp 1876 a: 163" authorityName="Sharp" authorityPageNumber="163" authorityYear="1876" box="[151,603,1412,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="9B8CEACC4C23B343FF6BFA50F6D5D0F8" box="[151,384,1412,1437]" italics="true" pageId="11" pageNumber="12">Belonuchus armatus</emphasis>
<treatmentCitation id="285910CF4C23B343FE77FA50F50ED0F9" author="" box="[395,603,1412,1437]" page="163" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE77FA50F54AD0F9" box="[395,543,1412,1437]" pageId="11" pageNumber="12" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 163
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD95FA51F5F4D0F9" box="[617,673,1413,1437]" pageId="11" pageNumber="12">Type</typeStatus>
locality: Abacaxis. 
<typeStatus id="7643887C4C23B343FC70FA51F491D0F9" box="[908,964,1413,1437]" pageId="11" pageNumber="12">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C23B343FBA4FA51F3F9D0F8" box="[1112,1196,1413,1436]" pageId="11" pageNumber="12">BMNH</collectionCode>
HT 
<specimenCount id="BFFEFD574C23B343FB1AFA51F3ABD0F8" box="[1254,1278,1413,1436]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
. 
<location id="AC2760054C23B343FAF1FA50F2CCD0F9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FAF1FA50F2CCD0F9" box="[1293,1433,1412,1437]" country="Brazil" name="Distribution" pageId="11" pageNumber="12" stateProvince="Amazonas">Distribution</location>
: 
<collectingCountry id="D1EF764E4C23B343FF3BFA73F65ED0A4" box="[199,267,1447,1472]" name="Brazil" pageId="11" pageNumber="12">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C23B343FEE6FA7CF6C2D0A4" box="[282,407,1448,1472]" country="Venezuela" name="Amazonas" pageId="11" pageNumber="12">Amazonas</collectingRegion>
)
</materialsCitation>
. Note: distribution in Brazil (Asenjo 
<emphasis id="9B8CEACC4C23B343FCC2FA7DF423D0A4" box="[830,886,1447,1472]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BFA3BF51ED36C" blockId="11.[151,1436,1519,1616]" box="[151,587,1519,1544]" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BFA3BF51ED36C" ID-CoL="LC5H" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,587,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="arrogans">
<emphasis id="9B8CEACC4C23B343FF6BFA3BF6C5D363" bold="true" box="[151,400,1519,1544]" italics="true" pageId="11" pageNumber="12">Belonuchus arrogans</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BF9C0F3E5D334" blockId="11.[151,1436,1519,1616]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BF9C0F3E5D334" pageId="11" pageNumber="12">
<materialsCitation id="19903C834C23B343FF6BF9C0F2CDD348" ID-GBIF-Occurrence="1915497752" box="[151,1432,1556,1581]" county="Pacho" elevation="2000" location="West-Columbien" municipality="Ost-Cordilleren" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C23B343FF6BF9C0F5CFD348" ID-CoL="LC5H" authority="Bernhauer 1916 c: 274" authorityName="Bernhauer" authorityPageNumber="274" authorityYear="1916" box="[151,666,1556,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="arrogans">
<emphasis id="9B8CEACC4C23B343FF6BF9C0F6DFD348" box="[151,394,1556,1581]" italics="true" pageId="11" pageNumber="12">Belonuchus arrogans</emphasis>
<treatmentCitation id="285910CF4C23B343FE69F9C0F5CFD348" author="" box="[405,666,1556,1581]" page="274" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE69F9C0F50AD349" box="[405,607,1556,1581]" pageId="11" pageNumber="12" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 274
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD55F9C1F5B5D349" box="[681,736,1557,1581]" pageId="11" pageNumber="12">Type</typeStatus>
locality: 
<location id="AC2760054C23B343FCAFF9C0F340D349" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FCAFF9C0F340D349" box="[851,1045,1556,1581]" county="Pacho" municipality="Ost-Cordilleren" name="West-Columbien" pageId="11" pageNumber="12">West-Columbien</location>
: 
<collectingMunicipality id="4923ACA44C23B343FBDFF9C0F383D349" box="[1059,1238,1556,1581]" pageId="11" pageNumber="12">Ost-Cordilleren</collectingMunicipality>
, 
<collectingCounty id="40264E524C23B343FB19F9C0F279D349" box="[1253,1324,1556,1581]" pageId="11" pageNumber="12">Pacho</collectingCounty>
, 
<elevation id="22D5D1ED4C23B343FAC0F9C0F2CDD348" box="[1340,1432,1556,1580]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="2000.0">
<quantity id="6E009B3B4C23B343FAC0F9C0F2CDD348" box="[1340,1432,1556,1580]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="2000.0">2000 m</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C23B343FF3BF9ECF3E5D334" ID-GBIF-Occurrence="1915497697" box="[199,1200,1591,1616]" collectionCode="FMNH" country="Colombia" location="Distribution" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C23B343FF3BF9ECF7AAD334" box="[199,255,1592,1616]" pageId="11" pageNumber="12">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C23B343FE76F9ECF68ED32B" box="[394,475,1592,1615]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH</collectionCode>
HT 
<specimenCount id="BFFEFD574C23B343FDF2F9ECF573D32B" box="[526,550,1592,1615]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
. 
<location id="AC2760054C23B343FDCEF9E3F5EBD334" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C23B343FDCEF9E3F5EBD334" box="[562,702,1591,1616]" country="Colombia" name="Distribution" pageId="11" pageNumber="12">Distribution</location>
: 
<collectingCountry id="D1EF764E4C23B343FD34F9E3F46FD334" box="[712,826,1591,1616]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BF9ABF50AD3FC" blockId="11.[151,1436,1663,1760]" box="[151,607,1663,1688]" pageId="11" pageNumber="12">
<emphasis id="9B8CEACC4C23B343FF6BF9ABF7F6D3FC" bold="true" box="[151,163,1663,1688]" italics="true" pageId="11" pageNumber="12">*</emphasis>
<taxonomicName id="6EF84D5D4C23B343FF5EF9ABF50AD3FC" ID-CoL="LC5J" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[162,607,1663,1688]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="9B8CEACC4C23B343FF5EF9ABF6F1D3F3" bold="true" box="[162,420,1663,1688]" italics="true" pageId="11" pageNumber="12">Belonuchus asperatus</emphasis>
<bibRefCitation id="CD694B2F4C23B343FE50F9ABF50AD3FC" box="[428,607,1663,1688]" pageId="11" pageNumber="12" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BF970F225D384" blockId="11.[151,1436,1663,1760]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BF970F225D384" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BF970F5CAD3D9" ID-CoL="LC5J" authority="Bernhauer 1906: 336" authorityName="Bernhauer" authorityPageNumber="336" authorityYear="1906" box="[151,671,1700,1725]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="9B8CEACC4C23B343FF6BF970F6C1D3D8" box="[151,404,1700,1725]" italics="true" pageId="11" pageNumber="12">Belonuchus asperatus</emphasis>
<treatmentCitation id="285910CF4C23B343FE5EF970F5CAD3D9" author="" box="[418,671,1700,1725]" page="336" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE5EF970F535D3D9" box="[418,608,1700,1725]" pageId="11" pageNumber="12" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 336
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD4DF971F5BCD3D9" box="[689,745,1701,1725]" pageId="11" pageNumber="12">Type</typeStatus>
locality: Pachitea, 
<collectingCountry id="D1EF764E4C23B343FC28F971F35ED3D8" box="[980,1035,1701,1724]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
. 
<typeStatus id="7643887C4C23B343FBE0F971F301D3D9" box="[1052,1108,1701,1725]" pageId="11" pageNumber="12">Type</typeStatus>
depository: FMNH HT 
<specimenCount id="BFFEFD574C23B343FA7CF971F2CDD3D8" box="[1408,1432,1701,1724]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
. Distribution: 
<collectingCountry id="D1EF764E4C23B343FEA1F913F69BD384" box="[349,462,1735,1760]" name="Colombia" pageId="11" pageNumber="12">Colombia</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C23B343FE25F913F56DD384" box="[473,568,1735,1760]" country="Democratic Republic of the Congo" name="Equateur" pageId="11" pageNumber="12">
<collectingCountry id="D1EF764E4C23B343FE25F913F56DD384" box="[473,568,1735,1760]" name="Ecuador" pageId="11" pageNumber="12">Ecuador</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C23B343FDB8F91CF52FD3BB" box="[580,634,1736,1759]" name="Peru" pageId="11" pageNumber="12">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C23B343FD79F913F58ED384" box="[645,731,1735,1760]" name="Bolivia" pageId="11" pageNumber="12">Bolivia</collectingCountry>
. Notes: distribution (
<treatmentCitation id="285910CF4C23B343FC39F913F391D384" author="" box="[965,1220,1735,1760]" page="120" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FC39F913F3D8D384" box="[965,1165,1735,1760]" pageId="11" pageNumber="12" refId="ref71794" refString="Scheerpeltz, O. (1960) Zur Kenntnis neotropischer Staphyliniden (Col.). Beitrage zur Neotropischen Fauna, 2 (2), 65 - 138. https: // doi. org / 10.1080 / 01650526009380624" type="journal article">Scheerpeltz 1960</bibRefCitation>
: 120
</treatmentCitation>
; 
<bibRefCitation id="CD694B2F4C23B343FB2CF913F231D384" box="[1232,1380,1735,1760]" pageId="11" pageNumber="12" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BF8DBF511D24C" blockId="11.[151,1437,1807,2013]" box="[151,580,1807,1832]" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BF8DBF511D24C" ID-CoL="LC5M" authority="(Wasmann 1925)" authorityName="Wasmann" authorityYear="1925" baseAuthorityName="Wasmann" baseAuthorityYear="1925" box="[151,580,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="atticida">
<emphasis id="9B8CEACC4C23B343FF6BF8DBF628D24C" bold="true" box="[151,381,1807,1832]" italics="true" pageId="11" pageNumber="12">Belonuchus atticida</emphasis>
(
<bibRefCitation id="CD694B2F4C23B343FE70F8C4F569D24C" box="[396,572,1807,1832]" pageId="11" pageNumber="12" refId="ref73915" refString="Wasmann, E. (1925) Die Ameisenmimikry. Ein exakter Beitrag zum Mimikryproblem und zur Theorie der Anpassung. Abhandlungen zur theoretischen Biologie, 19, i - xii + 1 - 164." type="journal article">Wasmann 1925</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C23B343FF6BF8E0F6B2D2DC" blockId="11.[151,1437,1807,2013]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B343FF6BF8E0F6B2D2DC" infered="true" pageId="11" pageNumber="12">
<taxonomicName id="6EF84D5D4C23B343FF6BF8E0F521D228" ID-CoL="6W3MH" authority="Wasmann 1925: 71" authorityName="Wasmann" authorityPageNumber="71" authorityYear="1925" box="[151,628,1844,1869]" class="Insecta" family="Staphylinidae" genus="Plociopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="atticida">
<emphasis id="9B8CEACC4C23B343FF6BF8E0F6D3D229" box="[151,390,1844,1869]" italics="true" pageId="11" pageNumber="12">Plociopterus atticida</emphasis>
<treatmentCitation id="285910CF4C23B343FE6EF8E1F521D228" author="" box="[402,628,1844,1869]" page="71" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE6EF8E1F512D229" box="[402,583,1844,1869]" pageId="11" pageNumber="12" refId="ref73915" refString="Wasmann, E. (1925) Die Ameisenmimikry. Ein exakter Beitrag zum Mimikryproblem und zur Theorie der Anpassung. Abhandlungen zur theoretischen Biologie, 19, i - xii + 1 - 164." type="journal article">Wasmann 1925</bibRefCitation>
: 71
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD78F8E1F5EED229" box="[644,699,1845,1869]" pageId="11" pageNumber="12">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C23B343FCD1F8E0F4D3D229" box="[813,902,1844,1869]" country="Uruguay" name="Colonia" pageId="11" pageNumber="12">Colonia</collectingRegion>
alpina, Rio d. Janeiro [LT]. 
<typeStatus id="7643887C4C23B343FB27F8E1F246D229" box="[1243,1299,1845,1869]" pageId="11" pageNumber="12">Type</typeStatus>
depository: NHME LT 
<specimenCount id="BFFEFD574C23B343FEA9F88CF639D20B" box="[341,364,1880,1903]" pageId="11" pageNumber="12" type="male">♂</specimenCount>
, #?PLT. Distribution: 
<collectingCountry id="D1EF764E4C23B343FD86F883F594D214" box="[634,705,1879,1904]" name="Brazil" pageId="11" pageNumber="12">Brazil</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C23B343FD28F88CF46BD214" box="[724,830,1880,1904]" country="Paraguay" name="Asuncion" pageId="11" pageNumber="12">
<collectingCountry id="D1EF764E4C23B343FD28F88CF46BD214" box="[724,830,1880,1904]" name="Paraguay" pageId="11" pageNumber="12">Paraguay</collectingCountry>
</collectingRegion>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C23B343FB9DF883F3A5D214" ID-CoL="8X7QS" box="[1121,1264,1879,1904]" class="Insecta" family="Staphylinidae" genus="Plociopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C23B343FB9DF883F3A5D214" box="[1121,1264,1879,1904]" italics="true" pageId="11" pageNumber="12">Plociopterus</emphasis>
</taxonomicName>
and 
<typeStatus id="7643887C4C23B343FAC9F883F2C9D214" box="[1333,1436,1879,1904]" pageId="11" pageNumber="12" type="lectotype">lectotype</typeStatus>
designation 
<emphasis id="9B8CEACC4C23B343FEA5F8A8F637D2F7" box="[345,354,1916,1939]" italics="true" pageId="11" pageNumber="12">(</emphasis>
<treatmentCitation id="285910CF4C23B343FE9FF8A8F40FD2F0" author="" box="[355,858,1916,1941]" page="214" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE9FF8A8F40FD2F0" box="[355,858,1916,1941]" pageId="11" pageNumber="12" refId="ref64817" refString="Chatzimanolis, S. &amp; Chani Posse, M. (2016) On the myrmecophilous species of Plociopterus (Coleoptera: Staphylinidae: Staphylinini) Kraatz described by E. Wasmann. The Coleopterists Bulletin, 70 (2), 214 - 216. https: // doi. org / 10.1649 / 0010 - 065 X- 70.2.214" type="journal article">Chatzimanolis &amp; Chani-Posse 2016: 214</bibRefCitation>
</treatmentCitation>
); with 
<taxonomicName id="6EF84D5D4C23B343FC40F8A8F3C3D2F0" ID-CoL="9N7M" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[956,1174,1916,1940]" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="9B8CEACC4C23B343FC40F8A8F3C3D2F0" box="[956,1174,1916,1940]" italics="true" pageId="11" pageNumber="12">Acromyrmex niger</emphasis>
</taxonomicName>
, 
<taxonomicName id="6EF84D5D4C23B343FB52F8A8F210D2F1" ID-CoL="JMC9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1198,1349,1916,1941]" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sexdens">
<emphasis id="9B8CEACC4C23B343FB52F8A8F210D2F1" box="[1198,1349,1916,1941]" italics="true" pageId="11" pageNumber="12">Atta sexdens</emphasis>
</taxonomicName>
[orig.; 
<bibRefCitation id="CD694B2F4C23B343FF3BF874F689D2DC" box="[199,476,1952,1977]" pageId="11" pageNumber="12" refId="ref69955" refString="Navarrete-Heredia, J. L. (2001) Beetles associated with Atta and Acromyrmex ants (Hymenoptera: Formicidae: Attini). Transactions of the American Entomological Society, 127, 381 - 429." type="journal article">Navarrete-Heredia 2001</bibRefCitation>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C23B344FF6BF811F4B9D478" blockId="11.[151,1437,1807,2013]" lastBlockId="12.[199,1436,151,285]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<treatmentCitationGroup id="89E811F04C23B344FF6BF811F4B9D478" infered="true" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="9B8CEACC4C23B343FF6BF811F7FFD2B8" box="[151,170,1989,2012]" italics="true" pageId="11" pageNumber="12">=</emphasis>
<taxonomicName id="6EF84D5D4C23B343FF4DF811F5C7D2B8" ID-CoL="6M2W9" authority="Scheerpeltz 1936: 484" authorityName="Scheerpeltz" authorityPageNumber="484" authorityYear="1936" box="[177,658,1988,2013]" class="Insecta" family="Staphylinidae" genus="Hesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="eidmanni">
<emphasis id="9B8CEACC4C23B343FF4DF811F6DED2B9" box="[177,395,1988,2013]" italics="true" pageId="11" pageNumber="12">Hesperus eidmanni</emphasis>
<treatmentCitation id="285910CF4C23B343FE6EF810F5C7D2B8" author="" box="[402,658,1988,2013]" page="484" pageId="11" pageNumber="12">
<bibRefCitation id="CD694B2F4C23B343FE6EF810F50CD2B9" box="[402,601,1988,2013]" pageId="11" pageNumber="12" refId="ref71733" refString="Scheerpeltz, O. (1936) Die von Prof. Dr. H. Eidmann gelegentlich seiner im Jahre 1933 nach Brasilien unternommenen Studeinreise aufgesammelten Staphyliniden. I. Die in den Nestern von Atta sexdens L. aufgefundenen Staphyliniden, nebst einigen Bemerkungen + Scariphaeus. Archiv fur Naturgeschichte, Neue Folge, 5, 483 - 540." type="journal article">Scheerpeltz 1936</bibRefCitation>
: 484
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C23B343FD61F811F580D2B9" box="[669,725,1989,2013]" pageId="11" pageNumber="12">Type</typeStatus>
locality: Villa Rica in 
<collectingRegion id="6B3CF83C4C23B343FC29F811F368D2B9" box="[981,1085,1989,2013]" country="Paraguay" name="Asuncion" pageId="11" pageNumber="12">
<collectingCountry id="D1EF764E4C23B343FC29F811F368D2B9" box="[981,1085,1989,2013]" name="Paraguay" pageId="11" pageNumber="12">Paraguay</collectingCountry>
</collectingRegion>
[LT]; Umgebung von Mendes, E. d. Rio [4PLT; 
<collectingCountry id="D1EF764E4C24B344FE73FF43F68FD5D4" box="[399,474,151,176]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
]. 
<typeStatus id="7643887C4C24B344FE12FF4CF573D5D4" box="[494,550,152,176]" pageId="12" pageNumber="13">Type</typeStatus>
depository: NMW (Scheerpeltz) LT 
<specimenCount id="BFFEFD574C24B344FC2CFF4CF4B2D5CB" box="[976,999,152,175]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
, 4 PLT (des. Chatzimanolis &amp; Chani- 
<treatmentCitation id="285910CF4C24B344FF3BFF69F6DFD5B0" author="" box="[199,394,188,213]" page="214" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FF3BFF69F61BD5B1" box="[199,334,188,213]" pageId="12" pageNumber="13" refId="ref64817" refString="Chatzimanolis, S. &amp; Chani Posse, M. (2016) On the myrmecophilous species of Plociopterus (Coleoptera: Staphylinidae: Staphylinini) Kraatz described by E. Wasmann. The Coleopterists Bulletin, 70 (2), 214 - 216. https: // doi. org / 10.1649 / 0010 - 065 X- 70.2.214" type="journal article">Posse 2016</bibRefCitation>
: 214
</treatmentCitation>
). Distribution: 
<collectingCountry id="D1EF764E4C24B344FDC6FF68F5D4D5B1" box="[570,641,188,213]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C24B344FD6CFF69F5AFD5B1" box="[656,762,189,213]" country="Paraguay" name="Asuncion" pageId="12" pageNumber="13">
<collectingCountry id="D1EF764E4C24B344FD6CFF69F5AFD5B1" box="[656,762,189,213]" name="Paraguay" pageId="12" pageNumber="13">Paraguay</collectingCountry>
</collectingRegion>
. Notes: distribution within 
<collectingCountry id="D1EF764E4C24B344FBC5FF68F328D5B1" box="[1081,1149,188,213]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C24B344FB17FF69F272D5B1" box="[1259,1319,188,213]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2013); as synonym of 
<taxonomicName id="6EF84D5D4C24B344FEA5FF35F490D59C" ID-CoL="6W3MH" authority="(Chatzimanolis &amp; Chani-Posse 2016: 214)" authorityName="Chatzimanolis &amp; Chani-Posse" authorityPageNumber="214" authorityYear="2016" baseAuthorityName="Chatzimanolis &amp; Chani-Posse" baseAuthorityYear="2016" box="[345,965,223,248]" class="Insecta" family="Staphylinidae" genus="Plociopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="atticida">
<emphasis id="9B8CEACC4C24B344FEA5FF35F684D59C" box="[345,465,223,248]" italics="true" pageId="12" pageNumber="13">B. atticida</emphasis>
(
<treatmentCitation id="285910CF4C24B344FE1EFF0BF4E8D59C" author="" box="[482,957,223,248]" page="214" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE1EFF0BF4E8D59C" box="[482,957,223,248]" pageId="12" pageNumber="13" refId="ref64817" refString="Chatzimanolis, S. &amp; Chani Posse, M. (2016) On the myrmecophilous species of Plociopterus (Coleoptera: Staphylinidae: Staphylinini) Kraatz described by E. Wasmann. The Coleopterists Bulletin, 70 (2), 214 - 216. https: // doi. org / 10.1649 / 0010 - 065 X- 70.2.214" type="journal article">Chatzimanolis &amp; Chani-Posse 2016: 214</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; in 
<taxonomicName id="6EF84D5D4C24B344FC0FFF34F3D6D59C" ID-CoL="JMC9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1011,1155,223,248]" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sexdens">
<emphasis id="9B8CEACC4C24B344FC0FFF34F3D6D59C" box="[1011,1155,223,248]" italics="true" pageId="12" pageNumber="13">Atta sexdens</emphasis>
</taxonomicName>
fungus chamber [orig.]; also 
<taxonomicName id="6EF84D5D4C24B344FF07FED0F554D479" ID-CoL="9N7K" baseAuthorityName="Emery" baseAuthorityYear="1888" box="[251,513,260,285]" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lobicornis">
<emphasis id="9B8CEACC4C24B344FF07FED0F554D479" box="[251,513,260,285]" italics="true" pageId="12" pageNumber="13">Acromyrmex lobicornis</emphasis>
</taxonomicName>
fungus chamber (
<bibRefCitation id="CD694B2F4C24B344FD36FED0F4B5D478" box="[714,992,260,285]" pageId="12" pageNumber="13" refId="ref69955" refString="Navarrete-Heredia, J. L. (2001) Beetles associated with Atta and Acromyrmex ants (Hymenoptera: Formicidae: Attini). Transactions of the American Entomological Society, 127, 381 - 429." type="journal article">Navarrete-Heredia 2001</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFE9FF549D400" blockId="12.[151,1437,331,429]" box="[151,540,331,357]" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFE9FF549D400" ID-CoL="LC5N" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[151,540,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="9B8CEACC4C24B344FF6BFE9FF626D400" bold="true" box="[151,371,331,356]" italics="true" pageId="12" pageNumber="13">Belonuchus avidus</emphasis>
<bibRefCitation id="CD694B2F4C24B344FE86FE98F549D400" box="[378,540,332,357]" pageId="12" pageNumber="13" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFEBBF200D4C9" blockId="12.[151,1437,331,429]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFEBBF200D4C9" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFEBBF509D4EC" ID-CoL="LC5N" authority="Erichson 1840: 423" authorityName="Erichson" authorityPageNumber="423" authorityYear="1840" box="[151,604,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="9B8CEACC4C24B344FF6BFEBBF638D4EC" box="[151,365,367,392]" italics="true" pageId="12" pageNumber="13">Belonuchus avidus</emphasis>
<treatmentCitation id="285910CF4C24B344FE8BFEBBF509D4EC" author="" box="[375,604,367,392]" page="423" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE8BFEBBF574D4EC" box="[375,545,367,392]" pageId="12" pageNumber="13" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 423
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FD96FEA4F5F7D4EC" box="[618,674,368,392]" pageId="12" pageNumber="13">Type</typeStatus>
locality: Novae Valenciae in Columbia [
<collectingCountry id="D1EF764E4C24B344FB80FEBBF3A0D4EC" box="[1148,1269,367,392]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
, teste 
<treatmentCitation id="285910CF4C24B344FABEFEA4F60FD4C9" author="" page="1594" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FABEFEA4F644D4C9" pageId="12" pageNumber="13" refId="ref67417" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article">Herman 2001d</bibRefCitation>
: 1594
</treatmentCitation>
]. 
<typeStatus id="7643887C4C24B344FE97FE41F6F6D4C9" box="[363,419,405,429]" pageId="12" pageNumber="13">Type</typeStatus>
depository: ZMHB 2 ST: 
<specimenCount id="BFFEFD574C24B344FD35FE40F5A0D4C8" box="[713,757,404,428]" pageId="12" pageNumber="13" type="male">1 ♂</specimenCount>
, 
<specimenCount id="BFFEFD574C24B344FCFDFE40F47ED4C8" box="[769,811,404,428]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
[MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C24B344FBA1FE40F39BD4C9" box="[1117,1230,404,429]" name="Colombia" pageId="12" pageNumber="13">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C24B344FB25FE40F205D4C9" box="[1241,1360,404,429]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFE0FF532D491" blockId="12.[151,1436,475,608]" box="[151,615,475,501]" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFE0FF532D491" ID-CoL="LC5R" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,615,475,501]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="barbicornis">
<emphasis id="9B8CEACC4C24B344FF6BFE0FF6F9D490" bold="true" box="[151,428,475,500]" italics="true" pageId="12" pageNumber="13">Belonuchus barbicornis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFE2BF3D0D759" blockId="12.[151,1436,475,608]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFE2BF3D0D759" pageId="12" pageNumber="13">
<materialsCitation id="19903C834C24B344FF6BFE2BF3D4D759" ID-GBIF-Occurrence="1915497800" collectionCode="FMNH" collectorName="West-Columbien &amp; Alto &amp; S. Antonio" country="Colombia" elevation="2200" location="ST. Distribution" pageId="12" pageNumber="13" specimenCode="FMNH 5" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C24B344FF6BFE2BF407D77C" ID-CoL="LC5R" authority="Bernhauer 1916 c: 271" authorityName="Bernhauer" authorityPageNumber="271" authorityYear="1916" box="[151,850,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="barbicornis" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C24B344FF6BFE2BF51AD77C" box="[151,591,511,536]" italics="true" pageId="12" pageNumber="13">Belonuchus (Musicoderus) barbicornis</emphasis>
<treatmentCitation id="285910CF4C24B344FDA9FE2BF407D77C" author="" box="[597,850,511,536]" page="271" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FDA9FE2BF44ED77C" box="[597,795,511,536]" pageId="12" pageNumber="13" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 271
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FCA1FDD4F4C0D77C" box="[861,917,512,536]" pageId="12" pageNumber="13">Type</typeStatus>
locality: 
<collectorName id="040D53084C24B344FBFCFE2BF397D77C" box="[1024,1218,511,536]" pageId="12" pageNumber="13">West-Columbien</collectorName>
: 
<collectorName id="040D53084C24B344FB30FE2BF3ABD77C" box="[1228,1278,511,536]" pageId="12" pageNumber="13">Alto</collectorName>
de las cruces, 
<elevation id="22D5D1ED4C24B344FF3BFDF0F640D758" box="[199,277,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="13" unit="m" value="2200.0">
<quantity id="6E009B3B4C24B344FF3BFDF0F640D758" box="[199,277,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="13" unit="m" value="2200.0">2200m</quantity>
</elevation>
&amp; 
<collectorName id="040D53084C24B344FEC4FDF0F6E0D759" box="[312,437,548,573]" pageId="12" pageNumber="13">S. Antonio</collectorName>
, 
<elevation id="22D5D1ED4C24B344FE3CFDF0F544D758" box="[448,529,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="13" unit="m" value="2200.0">
<quantity id="6E009B3B4C24B344FE3CFDF0F544D758" box="[448,529,548,572]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="12" pageNumber="13" unit="m" value="2200.0">2200m</quantity>
</elevation>
. 
<typeStatus id="7643887C4C24B344FDE0FDF1F506D759" box="[540,595,549,573]" pageId="12" pageNumber="13">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C24B344FD23FDF1F410D759" box="[735,837,548,573]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="13">FMNH 5</specimenCode>
<location id="AC2760054C24B344FCB0FDF0F351D759" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C24B344FCB0FDF0F351D759" box="[844,1028,548,573]" country="Colombia" name="ST. Distribution" pageId="12" pageNumber="13">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C24B344FBF3FDF0F3D4D759" box="[1039,1153,548,573]" name="Colombia" pageId="12" pageNumber="13">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C24B344FF6BFD93F465D704" box="[151,816,583,608]" pageId="12" pageNumber="13" type="discussion">
<paragraph id="A94736DE4C24B344FF6BFD93F465D704" blockId="12.[151,1436,475,608]" box="[151,816,583,608]" pageId="12" pageNumber="13">
<emphasis id="9B8CEACC4C24B344FF6BFD93F641D704" bold="true" box="[151,276,583,608]" pageId="12" pageNumber="13">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C24B344FEB0FD93F69AD704" box="[332,463,583,608]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C24B344FEB0FD93F69AD704" box="[332,463,583,608]" italics="true" pageId="12" pageNumber="13">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C24B345FF6BFD5BF645D6E4" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="A94736DE4C24B344FF6BFD5BF519D7CC" blockId="12.[151,1436,655,789]" box="[151,588,655,680]" pageId="12" pageNumber="13">
<emphasis id="9B8CEACC4C24B344FF6BFD5BF7F6D7CC" bold="true" box="[151,163,655,680]" italics="true" pageId="12" pageNumber="13">*</emphasis>
<taxonomicName id="6EF84D5D4C24B344FF5EFD5BF519D7CC" ID-CoL="LC5S" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,588,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="basiventris">
<emphasis id="9B8CEACC4C24B344FF5EFD5BF6E7D7CC" bold="true" box="[162,434,655,680]" italics="true" pageId="12" pageNumber="13">Belonuchus basiventris</emphasis>
(
<bibRefCitation id="CD694B2F4C24B344FE3DFD5BF511D7CC" box="[449,580,655,680]" pageId="12" pageNumber="13" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFD60F234D670" blockId="12.[151,1436,655,789]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFD60F234D670" infered="true" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFD60F539D7A8" ID-CoL="76XHQ" authority="Sharp 1885: 418" authorityName="Sharp" authorityPageNumber="418" authorityYear="1885" box="[151,620,692,717]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="apiciventris">
<emphasis id="9B8CEACC4C24B344FF6BFD60F6CBD7A8" box="[151,414,692,717]" italics="true" pageId="12" pageNumber="13">Philonthus apiciventris</emphasis>
<treatmentCitation id="285910CF4C24B344FE55FD60F539D7A8" author="" box="[425,620,692,717]" page="418" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE55FD60F565D7A9" box="[425,560,692,717]" pageId="12" pageNumber="13" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 418
</treatmentCitation>
</taxonomicName>
. Type locality: Mexico: Cordova, Puebla, Yolos, Oaxaca, Jalapa. Type depository: BMNH 2 ST. Distribution: Mexico. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C24B344FBFFFD03F659D670" ID-CoL="8VRYR" authority="(Jimenez-Sanchez et al. 2000)" authorityName="Jimenez-Sanchez et al." authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C24B344FBFFFD03F329D794" box="[1027,1148,727,752]" italics="true" pageId="12" pageNumber="13">Philonthus</emphasis>
(
<bibRefCitation id="CD694B2F4C24B344FB73FD03F651D670" pageId="12" pageNumber="13" refId="ref67947" refString="Jimenez-Sanchez, E., Navarrete-Heredia, J. L. &amp; Padilla-Ramirez, J. R. (2000) Estafilinidos (Coleoptera: Staphylinidae) necrofilos de la Sierra de Nanchititla, Estado de Mexico, Mexico. Folia Entomologica Mexicana, 108, 53 - 78." type="journal article">
Jiménez-Sánchez 
<emphasis id="9B8CEACC4C24B344FAA3FD0DF2C9D794" box="[1375,1436,727,752]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2000
</bibRefCitation>
)
</taxonomicName>
; in 
<taxonomicName id="6EF84D5D4C24B344FEC8FD28F680D670" ID-CoL="JMC4" baseAuthorityName="Kistner" baseAuthorityYear="1982" box="[308,469,764,788]" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="9B8CEACC4C24B344FEC8FD28F680D670" box="[308,469,764,788]" italics="true" pageId="12" pageNumber="13">Atta mexicana</emphasis>
</taxonomicName>
external debris [
<bibRefCitation id="CD694B2F4C24B344FD72FD28F4F1D670" box="[654,932,764,789]" pageId="12" pageNumber="13" refId="ref69955" refString="Navarrete-Heredia, J. L. (2001) Beetles associated with Atta and Acromyrmex ants (Hymenoptera: Formicidae: Attini). Transactions of the American Entomological Society, 127, 381 - 429." type="journal article">Navarrete-Heredia 2001</bibRefCitation>
; distribution (
<bibRefCitation id="CD694B2F4C24B344FBC3FD28F200D670" box="[1087,1365,764,789]" pageId="12" pageNumber="13" refId="ref69307" refString="Marquez, J. &amp; Asiain, J. (2012) Primeros registros estatales de especies mexicanas de Staphylinidae (Insecta: Coleoptera). Acta Zoologica Mexicana, Nueva Serie, 28, 118 - 132. https: // doi. org / 10.21829 / azm. 2012.281820" type="journal article">Márquez &amp; Asiain 2012</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFC97F555D639" blockId="12.[151,1436,835,933]" box="[151,512,835,861]" pageId="12" pageNumber="13">
<emphasis id="9B8CEACC4C24B344FF6BFC97F7F6D638" bold="true" box="[151,163,835,860]" italics="true" pageId="12" pageNumber="13">*</emphasis>
<taxonomicName id="6EF84D5D4C24B344FF5EFC97F555D639" ID-CoL="LC5T" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[162,512,835,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="9B8CEACC4C24B344FF5EFC97F62CD638" bold="true" box="[162,377,835,860]" italics="true" pageId="12" pageNumber="13">Belonuchus batesi</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFCB3F42FD6C0" blockId="12.[151,1436,835,933]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFCB3F42FD6C0" pageId="12" pageNumber="13">
<materialsCitation id="19903C834C24B344FF6BFCB3F601D6C0" ID-GBIF-Occurrence="1915497743" collectionCode="BMNH" country="Brazil" location="Distribution" pageId="12" pageNumber="13" specimenCount="2" stateProvince="Amazonas" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C24B344FF6BFCB3F51BD6E4" ID-CoL="LC5T" authority="Sharp 1876 a: 157" authorityName="Sharp" authorityPageNumber="157" authorityYear="1876" box="[151,590,871,896]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="batesi">
<emphasis id="9B8CEACC4C24B344FF6BFCB3F63FD6E4" box="[151,362,871,896]" italics="true" pageId="12" pageNumber="13">Belonuchus batesi</emphasis>
<treatmentCitation id="285910CF4C24B344FE84FCB3F51BD6E4" author="" box="[376,590,871,896]" page="157" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE84FCB3F545D6E4" box="[376,528,871,896]" pageId="12" pageNumber="13" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 157
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FD9CFCBCF5CDD6E4" box="[608,664,872,896]" pageId="12" pageNumber="13">Type</typeStatus>
locality: Ega. 
<typeStatus id="7643887C4C24B344FCB3FCBCF4D2D6E4" box="[847,903,872,896]" pageId="12" pageNumber="13">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C24B344FBDDFCBCF320D61B" box="[1057,1141,872,895]" pageId="12" pageNumber="13">BMNH</collectionCode>
HT. 
<location id="AC2760054C24B344FB47FCB3F212D6E4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C24B344FB47FCB3F212D6E4" box="[1211,1351,871,896]" country="Brazil" name="Distribution" pageId="12" pageNumber="13" stateProvince="Amazonas">Distribution</location>
: 
<collectingCountry id="D1EF764E4C24B344FAA4FCB3F2C9D6E4" box="[1368,1436,871,896]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C24B344FF33FC58F619D6C0" box="[207,332,908,932]" country="Venezuela" name="Amazonas" pageId="12" pageNumber="13">Amazonas</collectingRegion>
)
</materialsCitation>
. Note: distribution in Brazil (Asenjo 
<emphasis id="9B8CEACC4C24B344FD0FFC59F47ED6C1" box="[755,811,908,933]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFC07F55DD689" blockId="12.[151,1436,979,1149]" box="[151,520,979,1005]" pageId="12" pageNumber="13">
<emphasis id="9B8CEACC4C24B344FF6BFC07F7F6D688" bold="true" box="[151,163,979,1004]" italics="true" pageId="12" pageNumber="13">*</emphasis>
<taxonomicName id="6EF84D5D4C24B344FF5EFC07F55DD689" ID-CoL="LC5W" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,520,979,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="9B8CEACC4C24B344FF5EFC07F6D4D688" bold="true" box="[162,385,979,1004]" italics="true" pageId="12" pageNumber="13">Belonuchus bidens</emphasis>
<bibRefCitation id="CD694B2F4C24B344FE74FC00F55DD689" box="[392,520,980,1005]" pageId="12" pageNumber="13" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFC23F5E9D119" blockId="12.[151,1436,979,1149]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFC23F5E9D119" infered="true" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFC23F506D174" ID-CoL="LC5W" authority="Sharp 1885: 430" authorityName="Sharp" authorityPageNumber="430" authorityYear="1885" box="[151,595,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="9B8CEACC4C24B344FF6BFC23F626D174" box="[151,371,1015,1040]" italics="true" pageId="12" pageNumber="13">Belonuchus bidens</emphasis>
<treatmentCitation id="285910CF4C24B344FE7FFC23F506D174" author="" box="[387,595,1015,1040]" page="430" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE7FFC23F544D174" box="[387,529,1015,1040]" pageId="12" pageNumber="13" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 430
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FD9AFC2CF5C8D174" box="[614,669,1016,1040]" pageId="12" pageNumber="13">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C24B344FCE6FC23F427D174" box="[794,882,1015,1040]" country="Mexico" name="Mexico" pageId="12" pageNumber="13">
<collectingCountry id="D1EF764E4C24B344FCE6FC23F427D174" box="[794,882,1015,1040]" name="Mexico" pageId="12" pageNumber="13">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C24B344FC7BFC23F485D174" box="[903,976,1015,1040]" country="Guatemala" name="Jalapa" pageId="12" pageNumber="13">Jalapa</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C24B344FC19FC2CF369D174" box="[997,1084,1016,1040]" country="Mexico" name="Oaxaca" pageId="12" pageNumber="13">Oaxaca</collectingRegion>
, Mirador, Tuxtla; 
<collectingCountry id="D1EF764E4C24B344FADEFC23F2CDD174" box="[1314,1432,1015,1040]" name="Nicaragua" pageId="12" pageNumber="13">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C24B344FF3BFBC8F66ED151" box="[199,315,1052,1077]" country="Nicaragua" name="Chontales" pageId="12" pageNumber="13">Chontales</collectingRegion>
. 
<typeStatus id="7643887C4C24B344FEB7FBC9F6D6D151" box="[331,387,1053,1077]" pageId="12" pageNumber="13">Type</typeStatus>
depository: BMNH 5 ST: from all 
<typeStatus id="7643887C4C24B344FCCAFBC9F433D151" box="[822,870,1053,1077]" pageId="12" pageNumber="13">type</typeStatus>
localities [MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C24B344FADEFBC8F2CDD151" box="[1314,1432,1052,1077]" name="Nicaragua" pageId="12" pageNumber="13">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C24B344FF3BFBEBF675D13C" box="[199,288,1087,1112]" country="Mexico" name="Mexico" pageId="12" pageNumber="13">
<collectingCountry id="D1EF764E4C24B344FF3BFBEBF675D13C" box="[199,288,1087,1112]" name="Mexico" pageId="12" pageNumber="13">Mexico</collectingCountry>
</collectingRegion>
. Notes: distribution, also within 
<collectingRegion id="6B3CF83C4C24B344FD48FBEBF45CD13C" box="[692,777,1087,1112]" country="Mexico" name="Mexico" pageId="12" pageNumber="13">
<collectingCountry id="D1EF764E4C24B344FD48FBEBF45CD13C" box="[692,777,1087,1112]" name="Mexico" pageId="12" pageNumber="13">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C24B344FCE3FBEBF3DFD13C" box="[799,1162,1087,1112]" pageId="12" pageNumber="13" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C24B344FBFCFB95F315D13C" box="[1024,1088,1087,1112]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2002
</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C24B344FB61FB94F21FD13C" box="[1181,1354,1088,1112]" pageId="12" pageNumber="13" refId="ref69122" refString="Marquez, J. (2004) Primeros registros estatales de especies mexicanas de Staphylininae (Coleoptera). Acta Zoologica Mexicana, 20, 91 - 97." type="journal article">Márquez 2004</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C24B344FAA1FBEBF2CDD13C" box="[1373,1432,1087,1112]" pageId="12" pageNumber="13" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">2006</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C24B344FF3BFBB0F68CD118" box="[199,473,1124,1149]" pageId="12" pageNumber="13" refId="ref69307" refString="Marquez, J. &amp; Asiain, J. (2012) Primeros registros estatales de especies mexicanas de Staphylinidae (Insecta: Coleoptera). Acta Zoologica Mexicana, Nueva Serie, 28, 118 - 132. https: // doi. org / 10.21829 / azm. 2012.281820" type="journal article">Márquez &amp; Asiain 2012</bibRefCitation>
, Asiain 
<emphasis id="9B8CEACC4C24B344FDC8FBB1F539D119" box="[564,620,1124,1149]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2015).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFB7FF53FD1A1" blockId="12.[151,1436,1195,1293]" box="[151,618,1195,1221]" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFB7FF53FD1A1" ID-CoL="LC5Y" authority="Bernhauer 1935" authorityName="Bernhauer" authorityYear="1935" box="[151,618,1195,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="9B8CEACC4C24B344FF6BFB7FF6FAD1A0" bold="true" box="[151,431,1195,1220]" italics="true" pageId="12" pageNumber="13">Belonuchus bipunctatus</emphasis>
<bibRefCitation id="CD694B2F4C24B344FE4AFB78F53FD1A1" box="[438,618,1196,1221]" pageId="12" pageNumber="13" refId="ref62766" refString="Bernhauer, M. (1935) Neue Staphyliniden (Col.) aus Sudamerika. Stylops, a Journal of Taxonomic Entomology, 4, 90 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00565. x" type="journal article">Bernhauer 1935</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFB1BF4AAD069" blockId="12.[151,1436,1195,1293]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFB1BF4AAD069" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFB1BF5E5D18C" ID-CoL="LC5Y" authority="Bernhauer 1935: 90" authorityName="Bernhauer" authorityPageNumber="90" authorityYear="1935" box="[151,688,1231,1256]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="9B8CEACC4C24B344FF6BFB1BF6F9D18C" box="[151,428,1231,1256]" italics="true" pageId="12" pageNumber="13">Belonuchus bipunctatus</emphasis>
<treatmentCitation id="285910CF4C24B344FE40FB1BF5E5D18C" author="" box="[444,688,1231,1256]" page="90" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE40FB1BF528D18C" box="[444,637,1231,1256]" pageId="12" pageNumber="13" refId="ref62766" refString="Bernhauer, M. (1935) Neue Staphyliniden (Col.) aus Sudamerika. Stylops, a Journal of Taxonomic Entomology, 4, 90 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00565. x" type="journal article">Bernhauer 1935</bibRefCitation>
: 90
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FD38FB04F5AED18C" box="[708,763,1232,1256]" pageId="12" pageNumber="13">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C24B344FC8BFB1BF4ACD18C" box="[887,1017,1231,1256]" name="Colombia" pageId="12" pageNumber="13">Kolumbien</collectingCountry>
; Cali (Villa Elvira, 
<quantity id="6E009B3B4C24B344FB0AFB04F21CD18C" box="[1270,1353,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="m" value="1800.0">1800m</quantity>
). 
<typeStatus id="7643887C4C24B344FA98FB04F2C9D18C" box="[1380,1436,1232,1256]" pageId="12" pageNumber="13">Type</typeStatus>
depository: FMNH 2 ST; BMNH #?. Distribution: 
<collectingCountry id="D1EF764E4C24B344FCFCFB20F424D069" box="[768,881,1268,1293]" name="Colombia" pageId="12" pageNumber="13">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C24B344FC81FB20F4AFD069" box="[893,1018,1268,1293]" name="Costa Rica" pageId="12" pageNumber="13">
<collectingRegion id="6B3CF83C4C24B344FC81FB20F4E8D068" box="[893,957,1268,1292]" country="Tanzania" name="Pwani" pageId="12" pageNumber="13">Costa</collectingRegion>
Rica
</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFAEFF555D031" blockId="12.[151,1436,1339,1437]" box="[151,512,1339,1365]" pageId="12" pageNumber="13">
<emphasis id="9B8CEACC4C24B344FF6BFAEFF7F6D030" bold="true" box="[151,163,1339,1364]" italics="true" pageId="12" pageNumber="13">*</emphasis>
<taxonomicName id="6EF84D5D4C24B344FF5EFAEFF555D031" ID-CoL="LC62" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,512,1339,1365]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="9B8CEACC4C24B344FF5EFAEFF62CD030" bold="true" box="[162,377,1339,1364]" italics="true" pageId="12" pageNumber="13">Belonuchus boops</emphasis>
<bibRefCitation id="CD694B2F4C24B344FE7CFAE8F555D031" box="[384,512,1340,1365]" pageId="12" pageNumber="13" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFA8BF507D0F8" blockId="12.[151,1436,1339,1437]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFA8BF507D0F8" pageId="12" pageNumber="13">
<materialsCitation id="19903C834C24B344FF6BFA8BF4FAD01C" ID-GBIF-Occurrence="1915497731" box="[151,943,1375,1400]" country="Panama" location="Bugaba" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C24B344FF6BFA8BF561D01C" ID-CoL="LC62" authority="Sharp 1885: 435" authorityName="Sharp" authorityPageNumber="435" authorityYear="1885" box="[151,564,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="boops">
<emphasis id="9B8CEACC4C24B344FF6BFA8BF632D01C" box="[151,359,1375,1400]" italics="true" pageId="12" pageNumber="13">Belonuchus boops</emphasis>
<treatmentCitation id="285910CF4C24B344FE8DFA8BF561D01C" author="" box="[369,564,1375,1400]" page="435" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE8DFA8BF6ACD01C" box="[369,505,1375,1400]" pageId="12" pageNumber="13" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 435
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FDBFFAB4F52ED01C" box="[579,635,1376,1400]" pageId="12" pageNumber="13">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C24B344FD17FAB4F412D013" box="[747,839,1376,1399]" name="Panama" pageId="12" pageNumber="13">Panama</collectingCountry>
, 
<location id="AC2760054C24B344FCAAFA8BF4FAD01C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C24B344FCAAFA8BF4FAD01C" box="[854,943,1375,1400]" country="Panama" name="Bugaba" pageId="12" pageNumber="13">Bugaba</location>
</materialsCitation>
. 
<materialsCitation id="19903C834C24B344FC41FAB4F518D0F8" ID-GBIF-Occurrence="1915497780" collectionCode="MCP" country="Panama" location="Distribution" pageId="12" pageNumber="13" specimenCode="BMNH 2" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C24B344FC41FAB4F4A1D01C" box="[957,1012,1376,1400]" pageId="12" pageNumber="13">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C24B344FB7BFAB4F3A1D01C" box="[1159,1268,1376,1400]" collectionCode="BMNH" pageId="12" pageNumber="13">BMNH 2</specimenCode>
ST: 
<specimenCount id="BFFEFD574C24B344FACCFAB4F20AD013" box="[1328,1375,1376,1400]" pageId="12" pageNumber="13" type="male">1 ♂</specimenCount>
, 
<specimenCount id="BFFEFD574C24B344FA93FAB4F2C9D013" box="[1391,1436,1376,1400]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
[
<collectionCode id="CFE9AE1B4C24B344FF31FA50F65ED0F8" box="[205,267,1412,1436]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="12" pageNumber="13">MCP</collectionCode>
seen]. 
<location id="AC2760054C24B344FEA7FA50F6B2D0F9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C24B344FEA7FA50F6B2D0F9" box="[347,487,1412,1437]" country="Panama" name="Distribution" pageId="12" pageNumber="13">Distribution</location>
: 
<collectingCountry id="D1EF764E4C24B344FE0DFA51F518D0F8" box="[497,589,1413,1436]" name="Panama" pageId="12" pageNumber="13">Panama</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFA1FF541D080" blockId="12.[151,1437,1483,1581]" box="[151,532,1483,1509]" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BFA1FF541D080" ID-CoL="LC64" authority="Bernhauer 1934" authorityName="Bernhauer" authorityYear="1934" box="[151,532,1483,1509]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="boxi">
<emphasis id="9B8CEACC4C24B344FF6BFA1FF60FD080" bold="true" box="[151,346,1483,1508]" italics="true" pageId="12" pageNumber="13">Belonuchus boxi</emphasis>
Bernhauer 1934
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BFA3BF5E4D349" blockId="12.[151,1437,1483,1581]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BFA3BF5E4D349" pageId="12" pageNumber="13">
<materialsCitation id="19903C834C24B344FF6BFA3BF42FD36C" ID-GBIF-Occurrence="1915497709" box="[151,890,1519,1544]" location="Tucuman" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Tucuman" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C24B344FF6BFA3BF509D36C" ID-CoL="LC64" authority="Bernhauer 1934 b: 212" authorityName="Bernhauer" authorityPageNumber="212" authorityYear="1934" box="[151,604,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="boxi">
<emphasis id="9B8CEACC4C24B344FF6BFA3BF607D36C" box="[151,338,1519,1544]" italics="true" pageId="12" pageNumber="13">Belonuchus boxi</emphasis>
<treatmentCitation id="285910CF4C24B344FEA7FA3BF509D36C" author="" box="[347,604,1519,1544]" page="212" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FEA7FA3BF576D36C" box="[347,547,1519,1544]" pageId="12" pageNumber="13" refId="ref62723" refString="Bernhauer, M. (1934 b) Neue Staphyliniden aus Argentinien (IV) (Col.) (34. Beitrag zur Kenntnis der sudamerikanischen Fauna). Revista de Entomologia, 4 (2), 212 - 221." type="journal article">Bernhauer 1934b</bibRefCitation>
: 212
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FD95FA24F5F5D36C" box="[617,672,1520,1544]" pageId="12" pageNumber="13">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C24B344FCF2FA24F42FD36C" box="[782,890,1520,1544]" country="Argentina" name="Tucuman" pageId="12" pageNumber="13">Tucumán</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C24B344FC7BFA24F5E4D349" ID-GBIF-Occurrence="1915497803" collectionCode="FMNH" country="Argentina" location="Distribution" pageId="12" pageNumber="13" specimenCode="FMNH 1" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C24B344FC7BFA24F4EAD36C" box="[903,959,1520,1544]" pageId="12" pageNumber="13">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C24B344FBB2FA24F3E3D36C" box="[1102,1206,1520,1544]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="13">FMNH 1</specimenCode>
ST 
<specimenCount id="BFFEFD574C24B344FB14FA24F255D363" box="[1256,1280,1520,1543]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
. 
<location id="AC2760054C24B344FAF1FA3BF2CCD36C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C24B344FAF1FA3BF2CCD36C" box="[1293,1433,1519,1544]" country="Argentina" name="Distribution" pageId="12" pageNumber="13">Distribution</location>
: 
<collectingCountry id="D1EF764E4C24B344FF3BF9C0F66FD349" box="[199,314,1556,1581]" name="Argentina" pageId="12" pageNumber="13">Argentina</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BF98FF531D310" blockId="12.[151,1437,1627,1760]" box="[151,612,1627,1653]" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BF98FF531D310" ID-CoL="LC65" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,612,1627,1653]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brasilianus">
<emphasis id="9B8CEACC4C24B344FF6BF98FF6FCD310" bold="true" box="[151,425,1627,1652]" italics="true" pageId="12" pageNumber="13">Belonuchus brasilianus</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BF9ABF64AD384" blockId="12.[151,1437,1627,1760]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BF9ABF64AD384" pageId="12" pageNumber="13">
<materialsCitation id="19903C834C24B344FF6BF9ABF31CD3FC" ID-GBIF-Occurrence="1915497806" box="[151,1097,1663,1688]" collectorName="S. Catharina" country="Brazil" location="Brasilien" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C24B344FF6BF9ABF5CDD3FC" ID-CoL="LC65" authority="Bernhauer 1917 b: 19" authorityName="Bernhauer" authorityPageNumber="19" authorityYear="1917" box="[151,664,1663,1688]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brasilianus">
<emphasis id="9B8CEACC4C24B344FF6BF9ABF6CBD3FC" box="[151,414,1663,1688]" italics="true" pageId="12" pageNumber="13">Belonuchus brasilianus</emphasis>
<treatmentCitation id="285910CF4C24B344FE5BF9ABF5CDD3FC" author="" box="[423,664,1663,1688]" page="19" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE5BF9ABF538D3FC" box="[423,621,1663,1688]" pageId="12" pageNumber="13" refId="ref62221" refString="Bernhauer, M. (1917 b) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (17. Beitrag - Fortsetzung). Entomologische Blatter, 13 (1 - 3), 17 - 24." type="journal article">Bernhauer 1917b</bibRefCitation>
: 19
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FD58F954F589D3FC" box="[676,732,1664,1688]" pageId="12" pageNumber="13">Type</typeStatus>
locality: 
<collectorName id="040D53084C24B344FCBBF954F480D3FC" box="[839,981,1663,1688]" pageId="12" pageNumber="13">S. Catharina</collectorName>
, 
<collectingCountry id="D1EF764E4C24B344FC1CF9ABF31CD3FC" box="[992,1097,1663,1688]" name="Brazil" pageId="12" pageNumber="13">Brasilien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C24B344FBA8F954F527D3D9" ID-GBIF-Occurrence="1915497768" collectionCode="FMNH" country="Brazil" location="Distribution" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Santa Catarina" typeStatus="holotype">
<typeStatus id="7643887C4C24B344FBA8F954F3D9D3FC" box="[1108,1164,1664,1688]" pageId="12" pageNumber="13">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C24B344FAE5F954F23FD3F3" box="[1305,1386,1664,1687]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="13">FMNH</collectionCode>
HT. 
<location id="AC2760054C24B344FF3BF970F606D3D9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C24B344FF3BF970F606D3D9" box="[199,339,1700,1725]" country="Brazil" name="Distribution" pageId="12" pageNumber="13" stateProvince="Santa Catarina">Distribution</location>
: 
<collectingCountry id="D1EF764E4C24B344FE9EF970F6F3D3D9" box="[354,422,1700,1725]" name="Brazil" pageId="12" pageNumber="13">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C24B344FE46F970F53FD3D9" box="[442,618,1700,1725]" country="Brazil" name="Santa Catarina" pageId="12" pageNumber="13">Santa Catarina</collectingRegion>
)
</materialsCitation>
. Note: distribution in Brazil (Asenjo 
<emphasis id="9B8CEACC4C24B344FBCDF971F33BD3D9" box="[1073,1134,1700,1725]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2013); generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BF8DEF573D247" blockId="12.[151,1436,1802,1899]" box="[151,550,1802,1827]" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BF8DEF573D247" ID-CoL="LC66" authority="Fauvel 1891" authorityName="Fauvel" authorityYear="1891" box="[151,550,1802,1827]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="9B8CEACC4C24B344FF6BF8DEF6C6D247" bold="true" box="[151,403,1802,1827]" italics="true" pageId="12" pageNumber="13">Belonuchus breviceps</emphasis>
<bibRefCitation id="CD694B2F4C24B344FE67F8DEF573D247" box="[411,550,1802,1827]" pageId="12" pageNumber="13" refId="ref66096" refString="Fauvel, A. (1891) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 11 e memoire: Staphylinides. Revue d'Entomologie, 10, 87 - 127." type="journal article">Fauvel 1891</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BF8FAF48AD20F" blockId="12.[151,1436,1802,1899]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BF8FAF48AD20F" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BF8FAF50FD223" ID-CoL="LC66" authority="Fauvel 1891: 117" authorityName="Fauvel" authorityPageNumber="117" authorityYear="1891" box="[151,602,1838,1863]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis id="9B8CEACC4C24B344FF6BF8FAF6DED223" box="[151,395,1838,1863]" italics="true" pageId="12" pageNumber="13">Belonuchus breviceps</emphasis>
<treatmentCitation id="285910CF4C24B344FE6FF8FAF50FD223" author="" box="[403,602,1838,1863]" page="117" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE6FF8FAF576D223" box="[403,547,1838,1863]" pageId="12" pageNumber="13" refId="ref66096" refString="Fauvel, A. (1891) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 11 e memoire: Staphylinides. Revue d'Entomologie, 10, 87 - 127." type="journal article">Fauvel 1891</bibRefCitation>
: 117
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C24B344FD9AF8FBF5CBD223" box="[614,670,1839,1863]" pageId="12" pageNumber="13">Type</typeStatus>
locality: Colonie Tovar [
<collectingCountry id="D1EF764E4C24B344FC44F8FAF360D223" box="[952,1077,1838,1863]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
]. 
<typeStatus id="7643887C4C24B344FBBAF8FBF328D223" box="[1094,1149,1839,1863]" pageId="12" pageNumber="13">Type</typeStatus>
depository: IRSNB 2 ST [MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C24B344FE0DF886F53DD20F" box="[497,616,1874,1899]" name="Venezuela" pageId="12" pageNumber="13">Venezuela</collectingCountry>
. Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BF840F57AD2CA" blockId="12.[151,1436,1940,2038]" box="[151,559,1940,1966]" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BF840F57AD2CA" ID-CoL="LC68" authority="Bernhauer 1912" authorityName="Bernhauer" authorityYear="1912" box="[151,559,1940,1966]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bruchi">
<emphasis id="9B8CEACC4C24B344FF6BF840F621D2C9" bold="true" box="[151,372,1940,1965]" italics="true" pageId="12" pageNumber="13">Belonuchus bruchi</emphasis>
Bernhauer 1912
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C24B344FF6BF86DF318D292" blockId="12.[151,1436,1940,2038]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="89E811F04C24B344FF6BF86DF318D292" pageId="12" pageNumber="13">
<taxonomicName id="6EF84D5D4C24B344FF6BF86DF524D2B6" ID-CoL="LC68" authority="Bernhauer 1912 b: 175" authorityName="Bernhauer" authorityPageNumber="175" authorityYear="1912" box="[151,625,1977,2002]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bruchi">
<emphasis id="9B8CEACC4C24B344FF6BF86DF63FD2B6" box="[151,362,1977,2002]" italics="true" pageId="12" pageNumber="13">Belonuchus bruchi</emphasis>
<treatmentCitation id="285910CF4C24B344FE8EF86DF524D2B6" author="" box="[370,625,1977,2002]" page="175" pageId="12" pageNumber="13">
<bibRefCitation id="CD694B2F4C24B344FE8EF86DF56CD2B6" box="[370,569,1977,2002]" pageId="12" pageNumber="13" refId="ref62005" refString="Bernhauer, M. (1912 b) Zur Staphylinidenfauna von Sudamerika (8. Beitrag). Entomologische Blatter, 8 (6 - 7), 167 - 179." type="journal article">Bernhauer 1912b</bibRefCitation>
: 175
</treatmentCitation>
</taxonomicName>
. Type locality: Buenos Aires prov., Argentina. Type depository: FMNH HT. Distribution: Argentina. Note: not 
<taxonomicName id="6EF84D5D4C24B344FD80F809F4D8D292" ID-CoL="LC68" authority="Bernhauer 1934 b" authorityName="Bernhauer" authorityYear="1934" box="[636,909,2013,2038]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bruchi">
<emphasis id="9B8CEACC4C24B344FD80F809F590D292" box="[636,709,2013,2038]" italics="true" pageId="12" pageNumber="13">bruchi</emphasis>
<bibRefCitation id="CD694B2F4C24B344FD30F809F4D8D292" box="[716,909,2013,2038]" pageId="12" pageNumber="13" refId="ref62723" refString="Bernhauer, M. (1934 b) Neue Staphyliniden aus Argentinien (IV) (Col.) (34. Beitrag zur Kenntnis der sudamerikanischen Fauna). Revista de Entomologia, 4 (2), 212 - 221." type="journal article">Bernhauer 1934b</bibRefCitation>
</taxonomicName>
(see 
<taxonomicName id="6EF84D5D4C24B344FC34F809F369D292" ID-CoL="LC7D" authorityName="Blackwelder" authorityYear="1944" box="[968,1084,2013,2038]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="duplicatus">
<emphasis id="9B8CEACC4C24B344FC34F809F369D292" box="[968,1084,2013,2038]" italics="true" pageId="12" pageNumber="13">duplicatus</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFF43F521D5D4" blockId="13.[151,1436,151,248]" box="[151,628,151,176]" pageId="13" pageNumber="14">
<emphasis id="9B8CEACC4C25B345FF6BFF43F7F6D5D4" bold="true" box="[151,163,151,176]" italics="true" pageId="13" pageNumber="14">*</emphasis>
<taxonomicName id="6EF84D5D4C25B345FF5EFF43F521D5D4" ID-CoL="LC69" authority="Bernhauer 1911" authorityName="Bernhauer" authorityYear="1911" box="[162,628,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bruchianus">
<emphasis id="9B8CEACC4C25B345FF5EFF43F6EFD5D4" bold="true" box="[162,442,151,176]" italics="true" pageId="13" pageNumber="14">Belonuchus bruchianus</emphasis>
<bibRefCitation id="CD694B2F4C25B345FE3EFF43F521D5D4" box="[450,628,151,176]" pageId="13" pageNumber="14" refId="ref61942" refString="Bernhauer, M. (1911) Zur Staphylinidenfauna von Sud-Amerika (Col.) (7. Beitrag). Deutsche Entomologische Zeitschrift, 4, 403 - 422." type="journal article">Bernhauer 1911</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFF68F576D59C" blockId="13.[151,1436,151,248]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFF68F576D59C" pageId="13" pageNumber="14">
<materialsCitation id="19903C834C25B345FF6BFF68F32DD5B1" ID-GBIF-Occurrence="1915497786" box="[151,1144,188,213]" country="Argentina" location="Misiones" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Misiones" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C25B345FF6BFF68F5CED5B0" ID-CoL="LC69" authority="Bernhauer 1911: 413" authorityName="Bernhauer" authorityPageNumber="413" authorityYear="1911" box="[151,667,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bruchianus">
<emphasis id="9B8CEACC4C25B345FF6BFF68F6F5D5B1" box="[151,416,188,213]" italics="true" pageId="13" pageNumber="14">Belonuchus bruchianus</emphasis>
<treatmentCitation id="285910CF4C25B345FE55FF68F5CED5B0" author="" box="[425,667,188,213]" page="413" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE55FF68F534D5B0" box="[425,609,188,213]" pageId="13" pageNumber="14" refId="ref61942" refString="Bernhauer, M. (1911) Zur Staphylinidenfauna von Sud-Amerika (Col.) (7. Beitrag). Deutsche Entomologische Zeitschrift, 4, 403 - 422." type="journal article">Bernhauer 1911</bibRefCitation>
: 413
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C25B345FD54FF69F5B5D5B1" box="[680,736,189,213]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C25B345FCB1FF68F4E1D5B1" box="[845,948,188,213]" country="Paraguay" name="Misiones" pageId="13" pageNumber="14">Misiones</collectingRegion>
prov., 
<collectingCountry id="D1EF764E4C25B345FBF9FF68F32DD5B1" box="[1029,1144,188,213]" name="Argentina" pageId="13" pageNumber="14">Argentina</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C25B345FB78FF69F54AD59C" ID-GBIF-Occurrence="1915497767" collectionCode="FMNH" country="Argentina" location="Distribution" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="7643887C4C25B345FB78FF69F3E9D5B1" box="[1156,1212,189,213]" pageId="13" pageNumber="14">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C25B345FAB7FF69F2C9D5B0" box="[1355,1436,189,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14">FMNH</collectionCode>
HT 
<specimenCount id="BFFEFD574C25B345FF0FFF34F65CD593" box="[243,265,224,247]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
. 
<location id="AC2760054C25B345FEEAFF0BF6F7D59C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FEEAFF0BF6F7D59C" box="[278,418,223,248]" country="Argentina" name="Distribution" pageId="13" pageNumber="14">Distribution</location>
: 
<collectingCountry id="D1EF764E4C25B345FE50FF0BF54AD59C" box="[428,543,223,248]" name="Argentina" pageId="13" pageNumber="14">Argentina</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFEF3F576D424" blockId="13.[151,1437,295,501]" box="[151,547,295,320]" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFEF3F576D424" ID-CoL="68GHV" authority="Fauvel 1901" authorityName="Fauvel" authorityYear="1901" box="[151,547,295,320]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bugnioni">
<emphasis id="9B8CEACC4C25B345FF6BFEF3F6C5D424" bold="true" box="[151,400,295,320]" italics="true" pageId="13" pageNumber="14">Belonuchus bugnioni</emphasis>
<bibRefCitation id="CD694B2F4C25B345FE6BFEF3F576D424" box="[407,547,295,320]" pageId="13" pageNumber="14" refId="ref66160" refString="Fauvel, A. (1901) Voyage de M. le Dr. Ed. Bugnion au Venezuela, en Colombie et aux Antilles. Revue d'Entomologie, 20, 69 - 91, pls. 1 - 2." type="journal article">Fauvel 1901</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFE98F3E4D4C9" blockId="13.[151,1437,295,501]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFE98F3E4D4C9" infered="true" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFE98F504D401" ID-CoL="68GHV" authority="Fauvel 1901: 86" authorityName="Fauvel" authorityPageNumber="86" authorityYear="1901" box="[151,593,332,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bugnioni">
<emphasis id="9B8CEACC4C25B345FF6BFE98F6DDD401" box="[151,392,332,357]" italics="true" pageId="13" pageNumber="14">Belonuchus bugnioni</emphasis>
<treatmentCitation id="285910CF4C25B345FE6EFE98F504D401" author="" box="[402,593,332,357]" page="86" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE6EFE98F571D400" box="[402,548,332,357]" pageId="13" pageNumber="14" refId="ref66160" refString="Fauvel, A. (1901) Voyage de M. le Dr. Ed. Bugnion au Venezuela, en Colombie et aux Antilles. Revue d'Entomologie, 20, 69 - 91, pls. 1 - 2." type="journal article">Fauvel 1901</bibRefCitation>
: 86
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C25B345FDA3FE99F5C2D401" box="[607,663,333,357]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C25B345FCFBFE98F4D2D401" box="[775,903,332,357]" country="France" name="Martinique" pageId="13" pageNumber="14">
<collectingCountry id="D1EF764E4C25B345FCFBFE98F4D2D401" box="[775,903,332,357]" name="Martinique" pageId="13" pageNumber="14">Martinique</collectingCountry>
</collectingRegion>
. 
<typeStatus id="7643887C4C25B345FC69FE99F498D401" box="[917,973,333,357]" pageId="13" pageNumber="14">Type</typeStatus>
depository: IRSNB HT 
<specimenCount id="BFFEFD574C25B345FB10FE99F254D400" box="[1260,1281,333,356]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
[MCP seen]. Distribution: 
<collectingRegion id="6B3CF83C4C25B345FEA1FEBBF689D4EC" box="[349,476,367,392]" country="France" name="Martinique" pageId="13" pageNumber="14">
<collectingCountry id="D1EF764E4C25B345FEA1FEBBF689D4EC" box="[349,476,367,392]" name="Martinique" pageId="13" pageNumber="14">Martinique</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C25B345FE1BFEBBF527D4EC" box="[487,626,367,392]" country="France" name="Guadeloupe" pageId="13" pageNumber="14">
<collectingCountry id="D1EF764E4C25B345FE1BFEBBF527D4EC" box="[487,626,367,392]" name="Guadeloupe" pageId="13" pageNumber="14">Guadeloupe</collectingCountry>
</collectingRegion>
. Note: distribution within (and endemic to) Lesser Antilles (
<bibRefCitation id="CD694B2F4C25B345FAECFEBBF2CCD4EC" box="[1296,1433,367,392]" pageId="13" pageNumber="14" refId="ref71133" refString="Peck, S. B. (2011 b) The beetles of Martinique, Lesser Antilles (Insecta: Coleoptera); diversity and distributions. Insecta Mundi, 0 178, 1 - 57." type="journal article">Peck 2011b</bibRefCitation>
: 
<collectingRegion id="6B3CF83C4C25B345FF3BFE40F613D4C9" box="[199,326,404,429]" country="France" name="Martinique" pageId="13" pageNumber="14">
<collectingCountry id="D1EF764E4C25B345FF3BFE40F613D4C9" box="[199,326,404,429]" name="Martinique" pageId="13" pageNumber="14">Martinique</collectingCountry>
</collectingRegion>
, 
<bibRefCitation id="CD694B2F4C25B345FEADFE40F55FD4C8" box="[337,522,404,429]" pageId="13" pageNumber="14" refId="ref71205" refString="Peck, S. B., Thomas, M. C. &amp; Thurnbow, R. H. (2014) The diversity and distributions of the beetles (Insecta: Coleoptera) of the Guadeloupe Archipelago (Grande-Terre, Basse-Terre, La Desirade, Marie-Galante, Les Saintes, and Petite-Terre), Lesser Antilles. Insecta Mundi, 0 352, 1 - 156." type="journal article">
Peck 
<emphasis id="9B8CEACC4C25B345FE73FE41F692D4C9" box="[399,455,404,429]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2014
</bibRefCitation>
: 
<collectingRegion id="6B3CF83C4C25B345FDE8FE40F5CED4C9" box="[532,667,404,429]" country="France" name="Guadeloupe" pageId="13" pageNumber="14">
<collectingCountry id="D1EF764E4C25B345FDE8FE40F5CED4C9" box="[532,667,404,429]" name="Guadeloupe" pageId="13" pageNumber="14">Guadeloupe</collectingCountry>
</collectingRegion>
Archipelago). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFE6DF3DED490" blockId="13.[151,1437,295,501]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFE6DF3DED490" pageId="13" pageNumber="14">
<materialsCitation id="19903C834C25B345FF6BFE6DF345D4B4" ID-GBIF-Occurrence="1915497759" box="[151,1040,439,464]" country="Guadeloupe" location="Guadeloupe" pageId="13" pageNumber="14" specimenCount="1" typeStatus="holotype">
<emphasis id="9B8CEACC4C25B345FF6BFE6DF7FFD4B4" box="[151,170,441,464]" italics="true" pageId="13" pageNumber="14">=</emphasis>
<taxonomicName id="6EF84D5D4C25B345FF45FE63F5EBD4B4" ID-CoL="LC7K" authority="Bernhauer 1917 c: 105" authorityName="Bernhauer" authorityPageNumber="105" authorityYear="1917" box="[185,702,439,464]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="eximius">
<emphasis id="9B8CEACC4C25B345FF45FE63F6F4D4B4" box="[185,417,439,464]" italics="true" pageId="13" pageNumber="14">Belonuchus eximius</emphasis>
<treatmentCitation id="285910CF4C25B345FE4DFE63F5EBD4B4" author="" box="[433,702,439,464]" page="105" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE4DFE63F52BD4B4" box="[433,638,439,464]" pageId="13" pageNumber="14" refId="ref62261" refString="Bernhauer, M. (1917 c) Neue sudamerikanische Staphyliniden (18. Beitrag). Wiener Entomologische Zeitung, 36 (3 - 5), 102 - 116. https: // doi. org / 10.5962 / bhl. part. 12925" type="journal article">Bernhauer 1917c</bibRefCitation>
: 105
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C25B345FD2EFE6CF45FD4B4" box="[722,778,440,464]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C25B345FC79FE63F345D4B4" box="[901,1040,439,464]" name="Guadeloupe" pageId="13" pageNumber="14">Guadeloupe</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C25B345FBDFFE6CF3DED490" ID-GBIF-Occurrence="1915497719" collectionCode="FMNH" country="Guadeloupe" location="ST. Distribution" pageId="13" pageNumber="14" specimenCode="FMNH 2" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C25B345FBDFFE6CF30ED4B4" box="[1059,1115,440,464]" pageId="13" pageNumber="14">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C25B345FB04FE6CF232D4B4" box="[1272,1383,440,464]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14">FMNH 2</specimenCode>
<location id="AC2760054C25B345FA8BFE6CF606D491" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FA8BFE6CF606D491" country="Guadeloupe" name="ST. Distribution" pageId="13" pageNumber="14">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C25B345FEA1FE08F6BDD491" box="[349,488,476,501]" name="Guadeloupe" pageId="13" pageNumber="14">Guadeloupe</collectingCountry>
. Note: as synonym of 
<taxonomicName id="6EF84D5D4C25B345FD1EFE09F3D3D490" ID-CoL="68GHV" authority="(Blackwelder 1943: 428)" authorityName="Blackwelder" authorityPageNumber="428" authorityYear="1943" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" box="[738,1158,476,501]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bugnioni">
<emphasis id="9B8CEACC4C25B345FD1EFE09F430D491" box="[738,869,476,501]" italics="true" pageId="13" pageNumber="14">B. bugnioni</emphasis>
(
<treatmentCitation id="285910CF4C25B345FC8FFE08F32BD490" author="" box="[883,1150,476,501]" page="428" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FC8FFE08F32BD490" box="[883,1150,476,501]" pageId="13" pageNumber="14" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943: 428</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFDF7F56BD758" blockId="13.[151,1437,547,680]" box="[151,574,547,573]" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFDF7F56BD758" ID-CoL="LC6B" authority="Blackwelder 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,574,547,573]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cacao">
<emphasis id="9B8CEACC4C25B345FF6BFDF7F63ED75F" bold="true" box="[151,363,547,572]" italics="true" pageId="13" pageNumber="14">Belonuchus cacao</emphasis>
<bibRefCitation id="CD694B2F4C25B345FE8EFDF0F56BD758" box="[370,574,548,573]" pageId="13" pageNumber="14" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFD93F4CDD7CC" blockId="13.[151,1437,547,680]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFD93F4CDD7CC" infered="true" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFD93F529D704" ID-CoL="LC6B" authority="Blackwelder 1943: 434" authorityName="Blackwelder" authorityPageNumber="434" authorityYear="1943" box="[151,636,583,608]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cacao">
<emphasis id="9B8CEACC4C25B345FF6BFD93F633D704" box="[151,358,583,608]" italics="true" pageId="13" pageNumber="14">Belonuchus cacao</emphasis>
<treatmentCitation id="285910CF4C25B345FE92FD93F529D704" author="" box="[366,636,583,608]" page="434" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE92FD93F514D704" box="[366,577,583,608]" pageId="13" pageNumber="14" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 434
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C25B345FD74FD9CF595D704" box="[648,704,584,608]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C25B345FCD1FD93F4C4D704" box="[813,913,583,608]" name="Trinidad and Tobago" pageId="13" pageNumber="14">Trinidad</collectingCountry>
: one-half mile south-southwest of Manzanilla junction on the Plum Road. 
<typeStatus id="7643887C4C25B345FDF9FDB9F568D7E1" box="[517,573,621,645]" pageId="13" pageNumber="14">Type</typeStatus>
depository: USNM HT 
<specimenCount id="BFFEFD574C25B345FCACFDB9F432D7E0" box="[848,871,621,644]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
, 1 PT 
<specimenCount id="BFFEFD574C25B345FC4FFDB9F49CD7E0" box="[947,969,621,644]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
. Distribution: 
<collectingCountry id="D1EF764E4C25B345FB92FDB8F3B1D7E1" box="[1134,1252,620,645]" name="Venezuela" pageId="13" pageNumber="14">Venezuela</collectingCountry>
, 
<collectingCountry id="D1EF764E4C25B345FB0CFDB8F206D7E1" box="[1264,1363,620,645]" name="Trinidad and Tobago" pageId="13" pageNumber="14">Trinidad</collectingCountry>
. Note: 
<collectingCountry id="D1EF764E4C25B345FF3BFD5BF668D7CC" box="[199,317,655,680]" name="Venezuela" pageId="13" pageNumber="14">Venezuela</collectingCountry>
, biology, in 
<taxonomicName id="6EF84D5D4C25B345FE3AFD5BF561D7CC" ID-CoL="92B8T" box="[454,564,655,680]" class="Insecta" family="Nymphalidae" genus="Heliconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C25B345FE3AFD5BF561D7CC" box="[454,564,655,680]" italics="true" pageId="13" pageNumber="14">Heliconia</emphasis>
</taxonomicName>
bracts (
<bibRefCitation id="CD694B2F4C25B345FD71FD5BF4D9D7CC" box="[653,908,655,680]" pageId="13" pageNumber="14" refId="ref66551" refString="Frank, J. H. &amp; Barrera, R. (2010) Natural history of Belonuchus Nordmann spp. and allies (Coleoptera: Staphylinidae) in Heliconia L. (Zingiberales: Heliconiaceae) flower bracts. Insecta Mundi, 110, 1 - 12." type="journal article">Frank &amp; Barrera 2010</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFD03F516D794" blockId="13.[151,1436,727,933]" box="[151,579,727,752]" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFD03F516D794" ID-CoL="LC6C" authority="(Erichson 1840)" authorityName="Erichson" authorityYear="1840" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[151,579,727,752]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="candens">
<emphasis id="9B8CEACC4C25B345FF6BFD03F6D2D794" bold="true" box="[151,391,727,752]" italics="true" pageId="13" pageNumber="14">Belonuchus candens</emphasis>
(
<bibRefCitation id="CD694B2F4C25B345FE6AFD03F56ED794" box="[406,571,727,752]" pageId="13" pageNumber="14" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFD28F645D6E4" blockId="13.[151,1436,727,933]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFD28F645D6E4" infered="true" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFD28F537D671" ID-CoL="779JZ" authority="Erichson 1840: 460" authorityName="Erichson" authorityPageNumber="460" authorityYear="1840" box="[151,610,764,789]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="candens">
<emphasis id="9B8CEACC4C25B345FF6BFD28F623D671" box="[151,374,764,789]" italics="true" pageId="13" pageNumber="14">Philonthus candens</emphasis>
<treatmentCitation id="285910CF4C25B345FE82FD28F537D671" author="" box="[382,610,764,789]" page="460" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE82FD28F572D670" box="[382,551,764,789]" pageId="13" pageNumber="14" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 460
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C25B345FD93FD29F5F3D671" box="[623,678,765,789]" pageId="13" pageNumber="14">Type</typeStatus>
locality: Bogotae in Columbia interiore. 
<typeStatus id="7643887C4C25B345FB7EFD29F3EFD671" box="[1154,1210,765,789]" pageId="13" pageNumber="14">Type</typeStatus>
depository: ZMHB 1+3 ST: 
<specimenCount id="BFFEFD574C25B345FED5FCF4F606D653" box="[297,339,800,824]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
in Erichson´s hand, 
<specimenCount id="BFFEFD574C25B345FDC1FCF4F532D653" box="[573,615,800,824]" pageId="13" pageNumber="14" type="female">3 ♀</specimenCount>
[MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C25B345FC67FCCBF359D65C" box="[923,1036,799,824]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C25B345FBE4FCCBF30AD65C" box="[1048,1119,799,824]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C25B345FB97FCF4F3F7D653" box="[1131,1186,800,823]" name="Peru" pageId="13" pageNumber="14">Peru</collectingCountry>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C25B345FF3BFC90F588D639" ID-CoL="8VRYR" authority="(Bernhauer &amp; Schubert 1914: 369)" authorityName="Bernhauer &amp; Schubert" authorityPageNumber="369" authorityYear="1914" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[199,733,836,861]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C25B345FF3BFC90F615D639" box="[199,320,836,861]" italics="true" pageId="13" pageNumber="14">Philonthus</emphasis>
(
<treatmentCitation id="285910CF4C25B345FEADFC90F580D639" author="" box="[337,725,836,861]" page="369" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FEADFC90F580D639" box="[337,725,836,861]" pageId="13" pageNumber="14" refId="ref62964" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus. Vol. 57. W. Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer &amp; Schubert 1914: 369</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; distribution including 
<collectingCountry id="D1EF764E4C25B345FC16FC91F375D638" box="[1002,1056,837,860]" name="Peru" pageId="13" pageNumber="14">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C25B345FBD2FC90F327D639" box="[1070,1138,836,861]" name="Brazil" pageId="13" pageNumber="14">Brazil</collectingCountry>
questioned (Asenjo 
<emphasis id="9B8CEACC4C25B345FA9DFC91F2C9D639" box="[1377,1436,836,861]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C25B345FF6BFC58F465D6C1" box="[151,816,908,933]" pageId="13" pageNumber="14" type="discussion">
<paragraph id="A94736DE4C25B345FF6BFC58F465D6C1" blockId="13.[151,1436,727,933]" box="[151,816,908,933]" pageId="13" pageNumber="14">
<emphasis id="9B8CEACC4C25B345FF6BFC58F641D6C1" bold="true" box="[151,276,908,933]" pageId="13" pageNumber="14">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C25B345FEB0FC58F69AD6C1" box="[332,463,908,933]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C25B345FEB0FC58F69AD6C1" box="[332,463,908,933]" italics="true" pageId="13" pageNumber="14">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C25B345FF6BFC07F5D9D3FC" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="A94736DE4C25B345FF6BFC07F520D688" blockId="13.[151,1436,979,1112]" box="[151,629,979,1005]" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFC07F520D688" ID-CoL="LC6D" authority="Bernhauer 1918" authorityName="Bernhauer" authorityYear="1918" box="[151,629,979,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cariniventris">
<emphasis id="9B8CEACC4C25B345FF6BFC07F6EFD688" bold="true" box="[151,442,979,1004]" italics="true" pageId="13" pageNumber="14">Belonuchus cariniventris</emphasis>
<bibRefCitation id="CD694B2F4C25B345FE3DFC00F520D688" box="[449,629,980,1005]" pageId="13" pageNumber="14" refId="ref62367" refString="Bernhauer, M. (1918) Vier neue Belonuchus aus Mexiko [cont.]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 67, 225 - 226." type="journal article">Bernhauer 1918</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFC23F56BD13C" blockId="13.[151,1436,979,1112]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFC23F56BD13C" pageId="13" pageNumber="14">
<materialsCitation id="19903C834C25B345FF6BFC23F205D16B" ID-GBIF-Occurrence="1915497809" box="[151,1360,1015,1040]" country="Mexico" location="Jalappa" municipality="Xalapa" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C25B345FF6BFC23F59AD174" ID-CoL="LC6D" authority="Bernhauer 1918: 225" authorityName="Bernhauer" authorityPageNumber="225" authorityYear="1918" box="[151,719,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cariniventris">
<emphasis id="9B8CEACC4C25B345FF6BFC23F6EFD174" box="[151,442,1015,1040]" italics="true" pageId="13" pageNumber="14">Belonuchus cariniventris</emphasis>
<treatmentCitation id="285910CF4C25B345FE37FC23F59AD174" author="" box="[459,719,1015,1040]" page="225" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE37FC23F5D8D174" box="[459,653,1015,1040]" pageId="13" pageNumber="14" refId="ref62367" refString="Bernhauer, M. (1918) Vier neue Belonuchus aus Mexiko [cont.]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 67, 225 - 226." type="journal article">Bernhauer 1918</bibRefCitation>
: 225
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C25B345FD18FC2CF449D174" box="[740,796,1016,1040]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C25B345FC66FC23F4A1D174" box="[922,1012,1015,1040]" name="Mexico" pageId="13" pageNumber="14">Mexico</collectingCountry>
: 
<location id="AC2760054C25B345FBF5FC23F308D174" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FBF5FC23F308D174" box="[1033,1117,1015,1040]" country="Mexico" municipality="Xalapa" name="Jalappa" pageId="13" pageNumber="14" stateProvince="Veracruz">Jalappa</location>
[
<collectingMunicipality id="4923ACA44C25B345FB89FC23F39FD174" box="[1141,1226,1015,1040]" pageId="13" pageNumber="14">Xalapa</collectingMunicipality>
, 
<collectingRegion id="6B3CF83C4C25B345FB1CFC2CF21FD16B" box="[1248,1354,1016,1039]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="13" pageNumber="14">Veracruz</collectingRegion>
]
</materialsCitation>
. 
<materialsCitation id="19903C834C25B345FA98FC2CF5B2D151" ID-GBIF-Occurrence="1915497817" collectionCode="FMNH" country="Mexico" location="ST. Distribution" pageId="13" pageNumber="14" specimenCode="FMNH 3" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C25B345FA98FC2CF2C9D174" box="[1380,1436,1016,1040]" pageId="13" pageNumber="14">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C25B345FEB3FBC9F6ECD150" box="[335,441,1052,1076]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14">FMNH 3</specimenCode>
<location id="AC2760054C25B345FE38FBC8F5D5D151" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FE38FBC8F5D5D151" box="[452,640,1052,1077]" country="Mexico" name="ST. Distribution" pageId="13" pageNumber="14">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C25B345FD72FBC8F5B2D151" box="[654,743,1052,1077]" name="Mexico" pageId="13" pageNumber="14">Mexico</collectingCountry>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C25B345FB7FFBC8F651D13C" pageId="13" pageNumber="14" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C25B345FA9CFBC9F2C9D151" box="[1376,1436,1052,1077]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2002
</bibRefCitation>
); generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFB53F539D1C4" blockId="13.[151,1436,1159,1328]" box="[151,620,1159,1184]" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFB53F539D1C4" ID-CoL="5WGHT" authority="Chani-Posse 2006" authorityName="Chani-Posse" authorityYear="2006" box="[151,620,1159,1184]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="9B8CEACC4C25B345FF6BFB53F6CCD1FB" bold="true" box="[151,409,1159,1184]" italics="true" pageId="13" pageNumber="14">Belonuchus castaneus</emphasis>
<bibRefCitation id="CD694B2F4C25B345FE5CFB53F539D1C4" box="[416,620,1159,1184]" pageId="13" pageNumber="14" refId="ref64063" refString="Chani-Posse, M. (2006) Systematic revision and cladistic analysis of the Neotropical genus Chroaptomus Sharp (Coleoptera: Staphylinidae), with descriptions of two new species. Insect Systematics &amp; Evolution, 37, 361 - 383. https: // doi. org / 10.1163 / 187631206788831452" type="journal article">Chani-Posse 2006</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFB78F24BD1A1" blockId="13.[151,1436,1159,1328]" box="[151,1310,1196,1221]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFB78F24BD1A1" box="[151,1310,1196,1221]" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFB78F5CFD1A1" ID-CoL="5WGHT" authority="Chani-Posse 2006: 366" authorityName="Chani-Posse" authorityPageNumber="366" authorityYear="2006" box="[151,666,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="9B8CEACC4C25B345FF6BFB78F6DBD1A0" box="[151,398,1196,1221]" italics="true" pageId="13" pageNumber="14">Belonuchus castaneus</emphasis>
<treatmentCitation id="285910CF4C25B345FE6AFB78F5CFD1A1" author="" box="[406,666,1196,1221]" page="366" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE6AFB78F530D1A1" box="[406,613,1196,1221]" pageId="13" pageNumber="14" refId="ref64063" refString="Chani-Posse, M. (2006) Systematic revision and cladistic analysis of the Neotropical genus Chroaptomus Sharp (Coleoptera: Staphylinidae), with descriptions of two new species. Insect Systematics &amp; Evolution, 37, 361 - 383. https: // doi. org / 10.1163 / 187631206788831452" type="journal article">Chani-Posse 2006</bibRefCitation>
: 366
</treatmentCitation>
</taxonomicName>
(new name for 
<taxonomicName id="6EF84D5D4C25B345FCB7FB78F244D1A1" ID-CoL="LCB2" authority="Bernhauer 1922: 15" authorityName="Bernhauer" authorityPageNumber="15" authorityYear="1922" box="[843,1297,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richteri">
<emphasis id="9B8CEACC4C25B345FCB7FB78F37DD1A1" box="[843,1064,1196,1221]" italics="true" pageId="13" pageNumber="14">Belonuchus richteri</emphasis>
<treatmentCitation id="285910CF4C25B345FBD3FB78F244D1A1" author="" box="[1071,1297,1196,1221]" page="15" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FBD3FB78F3B2D1A0" box="[1071,1255,1196,1221]" pageId="13" pageNumber="14" refId="ref62514" refString="Bernhauer, M. (1922) Neue Staphyliniden aus Sudamerika (23. Beitrag). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 71 (1921), 1 - 23." type="journal article">Bernhauer 1922</bibRefCitation>
: 15
</treatmentCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFB05F5A9D054" blockId="13.[151,1436,1159,1328]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFB05F5A9D054" pageId="13" pageNumber="14">
<emphasis id="9B8CEACC4C25B345FF6BFB05F7FFD18C" box="[151,170,1233,1256]" italics="true" pageId="13" pageNumber="14">=</emphasis>
<taxonomicName id="6EF84D5D4C25B345FF4DFB1BF522D18C" ID-CoL="LCB2" authority="Bernhauer 1922: 15" authorityName="Bernhauer" authorityPageNumber="15" authorityYear="1922" box="[177,631,1231,1256]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richteri">
<emphasis id="9B8CEACC4C25B345FF4DFB1BF6C5D18C" box="[177,400,1231,1256]" italics="true" pageId="13" pageNumber="14">Belonuchus richteri</emphasis>
<treatmentCitation id="285910CF4C25B345FE6BFB1BF522D18C" author="" box="[407,631,1231,1256]" page="15" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE6BFB1BF505D18C" box="[407,592,1231,1256]" pageId="13" pageNumber="14" refId="ref62514" refString="Bernhauer, M. (1922) Neue Staphyliniden aus Sudamerika (23. Beitrag). Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 71 (1921), 1 - 23." type="journal article">Bernhauer 1922</bibRefCitation>
: 15
</treatmentCitation>
</taxonomicName>
(preoccupied, junior secondary homonym in 
<taxonomicName id="6EF84D5D4C25B345FB85FB1BF3AED18C" ID-CoL="8VRHV" box="[1145,1275,1231,1256]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C25B345FB85FB1BF3AED18C" box="[1145,1275,1231,1256]" italics="true" pageId="13" pageNumber="14">Belonuchus</emphasis>
</taxonomicName>
of 
<taxonomicName id="6EF84D5D4C25B345FADFFB1BF545D068" ID-CoL="4G9MH" authority="Bernhauer 1912 a: 44" authorityName="Bernhauer" authorityPageNumber="44" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richteri">
<emphasis id="9B8CEACC4C25B345FADFFB1BF64ED069" italics="true" pageId="13" pageNumber="14">Philonthus richteri</emphasis>
<treatmentCitation id="285910CF4C25B345FEDEFB20F545D068" author="" box="[290,528,1268,1293]" page="44" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FEDEFB20F6B3D068" box="[290,486,1268,1293]" pageId="13" pageNumber="14" refId="ref61973" refString="Bernhauer, M. (1912 a) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article">Bernhauer 1912a</bibRefCitation>
: 44
</treatmentCitation>
</taxonomicName>
). 
<typeStatus id="7643887C4C25B345FDD8FB21F509D069" box="[548,604,1269,1293]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C25B345FD39FB20F47BD069" box="[709,814,1268,1293]" country="Paraguay" name="Misiones" pageId="13" pageNumber="14">Misiones</collectingRegion>
, 
<collectingCountry id="D1EF764E4C25B345FCC5FB20F4F9D069" box="[825,940,1268,1293]" name="Argentina" pageId="13" pageNumber="14">Argentina</collectingCountry>
. 
<typeStatus id="7643887C4C25B345FC4BFB21F4BBD069" box="[951,1006,1269,1293]" pageId="13" pageNumber="14">Type</typeStatus>
depository: FMNH LT 
<specimenCount id="BFFEFD574C25B345FB04FB21F25AD068" box="[1272,1295,1269,1292]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
, 2 PLT (des. 
<treatmentCitation id="285910CF4C25B345FF3BFAC3F68ED054" author="" box="[199,475,1303,1328]" page="219" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FF3BFAC3F6F6D054" box="[199,419,1303,1328]" pageId="13" pageNumber="14" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b</bibRefCitation>
: 219
</treatmentCitation>
). Distribution: 
<collectingCountry id="D1EF764E4C25B345FD79FAC3F5ADD054" box="[645,760,1303,1328]" name="Argentina" pageId="13" pageNumber="14">Argentina</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFA8BF50BD01C" blockId="13.[151,1436,1375,1509]" box="[151,606,1375,1400]" pageId="13" pageNumber="14">
<taxonomicName id="6EF84D5D4C25B345FF6BFA8BF50BD01C" ID-CoL="LC6F" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,606,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cavipennis">
<emphasis id="9B8CEACC4C25B345FF6BFA8BF6F6D01C" bold="true" box="[151,419,1375,1400]" italics="true" pageId="13" pageNumber="14">Belonuchus cavipennis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BFA50F6C1D081" blockId="13.[151,1436,1375,1509]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BFA50F6C1D081" pageId="13" pageNumber="14">
<materialsCitation id="19903C834C25B345FF6BFA50F6F8D0A4" ID-GBIF-Occurrence="1915497814" country="Bolivia" location="Yuracares" pageId="13" pageNumber="14" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C25B345FF6BFA50F43ED0F9" ID-CoL="LC6F" authority="Bernhauer 1916 c: 267" authorityName="Bernhauer" authorityPageNumber="267" authorityYear="1916" box="[151,875,1412,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cavipennis" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C25B345FF6BFA50F507D0F8" box="[151,594,1412,1437]" italics="true" pageId="13" pageNumber="14">Belonuchus (Musicoderus) cavipennis</emphasis>
<treatmentCitation id="285910CF4C25B345FD9CFA50F43ED0F9" author="" box="[608,875,1412,1437]" page="267" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FD9CFA50F478D0F9" box="[608,813,1412,1437]" pageId="13" pageNumber="14" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 267
</treatmentCitation>
</taxonomicName>
, attributed to Fauvel 
<emphasis id="9B8CEACC4C25B345FB8BFA51F3B1D0F9" box="[1143,1252,1412,1437]" italics="true" pageId="13" pageNumber="14">in litteris</emphasis>
. 
<typeStatus id="7643887C4C25B345FB06FA51F267D0F9" box="[1274,1330,1413,1437]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<location id="AC2760054C25B345FF3BFA7CF66DD0A4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FF3BFA7CF66DD0A4" box="[199,312,1448,1472]" country="Bolivia" name="Yuracares" pageId="13" pageNumber="14">Yuracarès</location>
, 
<collectingCountry id="D1EF764E4C25B345FEB5FA73F6F8D0A4" box="[329,429,1447,1472]" name="Bolivia" pageId="13" pageNumber="14">Bolivien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C25B345FE42FA7CF6C1D081" ID-GBIF-Occurrence="1915497810" collectionCode="FMNH, ZMUH" country="Bolivia" location="Distribution" pageId="13" pageNumber="14" specimenCode="FMNH 1, ZMUH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C25B345FE42FA7CF6A3D0A4" box="[446,502,1448,1472]" pageId="13" pageNumber="14">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C25B345FD72FA7CF5AED0A4" box="[654,763,1448,1472]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14">FMNH 1</specimenCode>
ST; 
<specimenCode id="F95E9EA54C25B345FCC0FA7CF4F3D0A4" box="[828,934,1448,1472]" collectionCode="ZMUH" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="13" pageNumber="14">ZMUH 1</specimenCode>
+ST?. 
<location id="AC2760054C25B345FC07FA73F3D2D0A4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FC07FA73F3D2D0A4" box="[1019,1159,1447,1472]" country="Bolivia" name="Distribution" pageId="13" pageNumber="14">Distribution</location>
: 
<collectingCountry id="D1EF764E4C25B345FB6BFA73F3B8D0A4" box="[1175,1261,1447,1472]" name="Bolivia" pageId="13" pageNumber="14">Bolivia</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C25B345FF6BF9C7F464D348" blockId="13.[151,1437,1555,2013]" box="[151,817,1555,1581]" pageId="13" pageNumber="14">
<emphasis id="9B8CEACC4C25B345FF6BF9C7F7F6D348" bold="true" box="[151,163,1555,1580]" italics="true" pageId="13" pageNumber="14">*</emphasis>
<taxonomicName id="6EF84D5D4C25B345FF5EF9C7F576D349" ID-CoL="6V9KL" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,547,1555,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="celatus" status="comb. nov.">
<emphasis id="9B8CEACC4C25B345FF5EF9C7F6D2D348" bold="true" box="[162,391,1555,1580]" italics="true" pageId="13" pageNumber="14">Belonuchus celatus</emphasis>
(
<bibRefCitation id="CD694B2F4C25B345FE6AF9C0F54ED349" box="[406,539,1556,1581]" pageId="13" pageNumber="14" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C25B345FDD3F9C0F5FAD349" box="[559,687,1556,1581]" pageId="13" pageNumber="14">
<emphasis id="9B8CEACC4C25B345FDD3F9C0F5FAD349" bold="true" box="[559,687,1556,1581]" pageId="13" pageNumber="14">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C25B345FD43F9C0F47CD348" box="[703,809,1556,1581]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="13" pageNumber="14">Figs. 3–4</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C25B345FF6BF9E3F5D9D3FC" blockId="13.[151,1437,1555,2013]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="89E811F04C25B345FF6BF9E3F5D9D3FC" infered="true" pageId="13" pageNumber="14">
<materialsCitation id="19903C834C25B345FF6BF9E3F4C9D334" ID-GBIF-Occurrence="1915497793" box="[151,924,1591,1616]" country="Mexico" location="Mirador" pageId="13" pageNumber="14" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C25B345FF6BF9E3F579D334" ID-CoL="6V9KL" authority="Sharp 1885: 420" authorityName="Sharp" authorityPageNumber="420" authorityYear="1885" box="[151,556,1591,1616]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="celatus">
<emphasis id="9B8CEACC4C25B345FF6BF9E3F632D334" box="[151,359,1591,1616]" italics="true" pageId="13" pageNumber="14">Philonthus celatus</emphasis>
<treatmentCitation id="285910CF4C25B345FE92F9E3F579D334" author="" box="[366,556,1591,1616]" page="420" pageId="13" pageNumber="14">
<bibRefCitation id="CD694B2F4C25B345FE92F9E3F6A6D334" box="[366,499,1591,1616]" pageId="13" pageNumber="14" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 420
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C25B345FDCBF9ECF53AD334" box="[567,623,1592,1616]" pageId="13" pageNumber="14">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C25B345FD25F9E3F466D334" box="[729,819,1591,1616]" name="Mexico" pageId="13" pageNumber="14">Mexico</collectingCountry>
: 
<location id="AC2760054C25B345FCC2F9E3F4C9D334" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FCC2F9E3F4C9D334" box="[830,924,1591,1616]" country="Mexico" name="Mirador" pageId="13" pageNumber="14">Mirador</location>
</materialsCitation>
, 
<materialsCitation id="19903C834C25B345FC5BF9ECF328D311" ID-GBIF-Occurrence="1915497792" collectionCode="MCP" country="Mexico" location="Distribution" pageId="13" pageNumber="14" specimenCode="BMNH 10" specimenCount="2" stateProvince="Oaxaca" typeStatus="holotype">
<collectingRegion id="6B3CF83C4C25B345FC5BF9ECF4ABD334" box="[935,1022,1592,1616]" country="Mexico" name="Oaxaca" pageId="13" pageNumber="14">Oaxaca</collectingRegion>
, Tehuacan, Yolos, Cuernavaca. 
<typeStatus id="7643887C4C25B345FA98F9ECF2C9D334" box="[1380,1436,1592,1616]" pageId="13" pageNumber="14">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C25B345FEB0F989F697D310" box="[332,450,1628,1652]" collectionCode="BMNH" pageId="13" pageNumber="14">BMNH 10</specimenCode>
ST: from all 
<typeStatus id="7643887C4C25B345FDABF989F5D2D311" box="[599,647,1629,1653]" pageId="13" pageNumber="14">type</typeStatus>
localities [
<collectionCode id="CFE9AE1B4C25B345FCFCF988F46BD310" box="[768,830,1628,1652]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="13" pageNumber="14">MCP</collectionCode>
seen]. 
<location id="AC2760054C25B345FC72F988F34FD311" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C25B345FC72F988F34FD311" box="[910,1050,1628,1653]" country="Mexico" name="Distribution" pageId="13" pageNumber="14" stateProvince="Oaxaca">Distribution</location>
: 
<collectingCountry id="D1EF764E4C25B345FBD8F988F328D311" box="[1060,1149,1628,1653]" name="Mexico" pageId="13" pageNumber="14">Mexico</collectingCountry>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C25B345FED7F9ABF5D5D3FC" box="[299,640,1663,1688]" pageId="13" pageNumber="14" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C25B345FDF8F955F569D3FC" box="[516,572,1663,1688]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C25B345FF6BF970F33AD24C" pageId="13" pageNumber="14" type="discussion">
<paragraph id="A94736DE4C25B345FF6BF970F33AD24C" blockId="13.[151,1437,1555,2013]" pageId="13" pageNumber="14">
<emphasis id="9B8CEACC4C25B345FF6BF970F65ED3D9" bold="true" box="[151,267,1700,1725]" pageId="13" pageNumber="14">Remarks.</emphasis>
Sharp did not specify the number of specimens he examined for naming this species, but it seems likely that he based his description on at least two specimens of different sex. Ten specimens were located in the BMNH under this name. All the ten specimens listed below agree with the original description by 
<bibRefCitation id="CD694B2F4C25B345FAF6F938F2C9D261" box="[1290,1436,1772,1797]" pageId="13" pageNumber="14" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp (1885)</bibRefCitation>
and are considered to have been 
<typeStatus id="7643887C4C25B345FDC8F8C4F5C3D24C" box="[564,662,1808,1832]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C25B345FC84F8DBF48AD24C" box="[888,991,1807,1832]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C25B346FF6BF8E0F5C6D424" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="A94736DE4C25B346FF6BF8E0F5C6D424" blockId="13.[151,1437,1555,2013]" lastBlockId="14.[151,1436,151,357]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="9B8CEACC4C25B345FF6BF8E0F696D229" bold="true" box="[151,451,1844,1869]" pageId="13" pageNumber="14">Type material examined.</emphasis>
Lectotype ♂, here designated (BMNH, 
<figureCitation id="31C32A5B4C25B345FC6DF8E0F355D228" box="[913,1024,1844,1869]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="13" pageNumber="14">Figs. 3–4</figureCitation>
): card-mounted, “ 
<taxonomicName id="6EF84D5D4C25B345FB37F8E0F2C9D229" box="[1227,1436,1844,1869]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="celatus">Philonthus celatus</taxonomicName>
Type D.S. Oaxaca. Höge” [in Sharp’s hand] // Type (round white label with red margin) // Oaxaca, Mexico. Hoege (white label) // B. C. A. Col. 
<date id="DD46101E4C25B345FD89F8A9F5F1D2F0" box="[629,676,1916,1940]" pageId="13" pageNumber="14">I. 2.</date>
<taxonomicName id="6EF84D5D4C25B345FD52F8A8F48DD2F1" authority=", Sharp." authorityName="Sharp." box="[686,984,1916,1941]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="celatus">Philonthus celatus, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 2 ♀, one of them “Cuernavaca” (white label) [in Sharp’s hand], the other one “Yolos” (white label) [in Sharp’s hand], both of them with the following additional labels, “Mexico. Salle. Coll (white label) // B. C. A. Col. 
<date id="DD46101E4C26B346FD7BFF4CF5E6D5D4" box="[647,691,152,176]" pageId="14" pageNumber="15">I. 2.</date>
<taxonomicName id="6EF84D5D4C26B346FD45FF43F489D5D4" authority=", Sharp." authorityName="Sharp." box="[697,988,151,176]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="celatus">Philonthus celatus, Sharp.</taxonomicName>
(white label)”; 1 ♂, 4 ♀, card-mounted, ♂ with “ 
<taxonomicName id="6EF84D5D4C26B346FEDAFF68F5F6D5B0" authority="Mirador. Hoge" authorityName="Mirador. Hoge" box="[294,675,188,213]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="celatus">Philonthus celatus Mirador. Höge</taxonomicName>
” [in Sharp’s hand]), “Mirador, Mexico. Hoege (white label) // B. C. A. Col. 
<date id="DD46101E4C26B346FEE2FF34F61FD59C" box="[286,330,224,248]" pageId="14" pageNumber="15">I. 2.</date>
<taxonomicName id="6EF84D5D4C26B346FEADFF0BF521D59C" authority=", Sharp." authorityName="Sharp." box="[337,628,223,248]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="celatus">Philonthus celatus, Sharp.</taxonomicName>
(white label)”; 2 ♀, card-mounted (one of them also “ 
<taxonomicName id="6EF84D5D4C26B346FB32FF0BF2C9D59C" box="[1230,1436,223,248]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="celatus">Philonthus celatus</taxonomicName>
Type D.S. Tehuacan. Höge” [in Sharp’s hand]), “Tehuacan, Mexico. Hoege (white label) // B. C. A. Col. 
<date id="DD46101E4C26B346FA93FED1F2C9D478" box="[1391,1436,260,284]" pageId="14" pageNumber="15">I. 2.</date>
<taxonomicName id="6EF84D5D4C26B346FF3BFEF3F6BFD424" authority=", Sharp." authorityName="Sharp." box="[199,490,295,320]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="celatus">Philonthus celatus, Sharp.</taxonomicName>
(white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C26B346FF6BFE98F33DD400" box="[151,1128,332,357]" pageId="14" pageNumber="15" type="discussion">
<paragraph id="A94736DE4C26B346FF6BFE98F33DD400" blockId="14.[151,1436,151,357]" box="[151,1128,332,357]" pageId="14" pageNumber="15">
<emphasis id="9B8CEACC4C26B346FF6BFE98F64AD401" bold="true" box="[151,287,332,357]" pageId="14" pageNumber="15">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C26B346FD46FE98F331D400" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,332,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C26B346FD46FE98F468D401" box="[698,829,332,357]" italics="true" pageId="14" pageNumber="15">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C26B346FCB7FE98F309D400" box="[843,1116,332,357]" pageId="14" pageNumber="15" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C26B346FC1CFE99F34DD401" box="[992,1048,332,357]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C26B346FF6BFE47F37DD0F9" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="A94736DE4C26B346FF6BFE47F56ED4C9" blockId="14.[151,1437,403,573]" box="[151,571,403,429]" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BFE47F56ED4C9" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[151,571,403,429]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cephalotes">
<emphasis id="9B8CEACC4C26B346FF6BFE47F6F4D4C8" bold="true" box="[151,417,403,428]" italics="true" pageId="14" pageNumber="15">Belonuchus cephalotes</emphasis>
(
<bibRefCitation id="CD694B2F4C26B346FE4CFE40F566D4C9" box="[432,563,404,429]" pageId="14" pageNumber="15" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFE63F393D759" blockId="14.[151,1437,403,573]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BFE63F393D759" infered="true" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BFE63F529D4B4" ID-CoL="6S2R7" authority="Sharp 1885: 455" authorityName="Sharp" authorityPageNumber="455" authorityYear="1885" box="[151,636,439,464]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cephalotes">
<emphasis id="9B8CEACC4C26B346FF6BFE63F6FED4B4" box="[151,427,439,464]" italics="true" pageId="14" pageNumber="15">Musicoderus cephalotes</emphasis>
<treatmentCitation id="285910CF4C26B346FE4AFE63F529D4B4" author="" box="[438,636,439,464]" page="455" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE4AFE63F56AD4B4" box="[438,575,439,464]" pageId="14" pageNumber="15" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 455
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C26B346FD70FE6CF596D4B4" box="[652,707,440,464]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C26B346FCCAFE6CF4C7D4AB" box="[822,914,440,463]" country="Panama" name="Panama" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FCCAFE6CF4C7D4AB" box="[822,914,440,463]" name="Panama" pageId="14" pageNumber="15">Panama</collectingCountry>
</collectingRegion>
, Bugaba 
<quantity id="6E009B3B4C26B346FBFFFE6CF399D4B4" box="[1027,1228,439,464]" metricMagnitude="2" metricUnit="m" metricValue="3.5052" metricValueMax="4.572" metricValueMin="2.4384" pageId="14" pageNumber="15" unit="ft" value="1150.0" valueMax="1500.0" valueMin="800.0">800 to 1500 feet</quantity>
. 
<typeStatus id="7643887C4C26B346FB26FE6CF247D4B4" box="[1242,1298,440,464]" pageId="14" pageNumber="15">Type</typeStatus>
depository: BMNH 12 ST: all of them with labels as &quot;Bugaba&quot; but only two of them with the elevation data [MCP seen]; FMNH 3 ST. Distribution: 
<collectingRegion id="6B3CF83C4C26B346FE07FDD4F502D773" box="[507,599,512,535]" country="Panama" name="Panama" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FE07FDD4F502D773" box="[507,599,512,535]" name="Panama" pageId="14" pageNumber="15">Panama</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C26B346FD9FFE2BF58FD77C" box="[611,730,511,536]" name="Venezuela" pageId="14" pageNumber="15">Venezuela</collectingCountry>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C26B346FC22FE2BF2C2D77C" ID-CoL="8XMY8" authority="(Scheerpeltz 1933: 1377)" authorityName="Scheerpeltz" authorityPageNumber="1377" authorityYear="1933" baseAuthorityName="Scheerpeltz 1933:" baseAuthorityYear="1377" box="[990,1431,511,536]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C26B346FC22FE2BF33AD77C" box="[990,1135,511,536]" italics="true" pageId="14" pageNumber="15">Musicoderus</emphasis>
(
<treatmentCitation id="285910CF4C26B346FB82FE2BF2DAD77C" author="" box="[1150,1423,511,536]" page="1377" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FB82FE2BF212D77C" box="[1150,1351,511,536]" pageId="14" pageNumber="15" refId="ref71693" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. 129. W. Junk, Berlin, pp. 989 - 1500." type="book chapter">Scheerpeltz 1933</bibRefCitation>
: 1377
</treatmentCitation>
)
</taxonomicName>
; 
<collectingCountry id="D1EF764E4C26B346FF3BFDF0F668D759" box="[199,317,548,573]" name="Venezuela" pageId="14" pageNumber="15">Venezuela</collectingCountry>
, biology, in 
<taxonomicName id="6EF84D5D4C26B346FE3AFDF0F561D759" ID-CoL="92B8T" box="[454,564,548,573]" class="Insecta" family="Nymphalidae" genus="Heliconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C26B346FE3AFDF0F561D759" box="[454,564,548,573]" italics="true" pageId="14" pageNumber="15">Heliconia</emphasis>
</taxonomicName>
bracts (
<bibRefCitation id="CD694B2F4C26B346FD71FDF0F4D9D758" box="[653,908,548,573]" pageId="14" pageNumber="15" refId="ref66551" refString="Frank, J. H. &amp; Barrera, R. (2010) Natural history of Belonuchus Nordmann spp. and allies (Coleoptera: Staphylinidae) in Heliconia L. (Zingiberales: Heliconiaceae) flower bracts. Insecta Mundi, 110, 1 - 12." type="journal article">Frank &amp; Barrera 2010</bibRefCitation>
); generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFDBFF400D7E1" blockId="14.[151,1437,619,752]" box="[151,853,619,645]" pageId="14" pageNumber="15">
<emphasis id="9B8CEACC4C26B346FF6BFDBFF7F6D7E0" bold="true" box="[151,163,619,644]" italics="true" pageId="14" pageNumber="15">*</emphasis>
<taxonomicName id="6EF84D5D4C26B346FF5EFDBFF400D7E1" ID-CoL="LC6L" authority="Rodriguez &amp; Navarrete-Heredia 2016" authorityName="Rodriguez &amp; Navarrete-Heredia" authorityYear="2016" box="[162,853,619,645]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cifuentesi">
<emphasis id="9B8CEACC4C26B346FF5EFDBFF6F3D7E0" bold="true" box="[162,422,619,644]" italics="true" pageId="14" pageNumber="15">Belonuchus cifuentesi</emphasis>
<bibRefCitation id="CD694B2F4C26B346FE51FDB8F400D7E1" box="[429,853,620,645]" pageId="14" pageNumber="15" refId="ref71443" refString="Rodriguez, W. D. &amp; Navarrete-Heredia, J. L. (2016) A new Mexican species of Belonuchus Nordmann (Coleoptera, Staphylinidae). Z ootaxa, 4083 (3), 444 - 450. https: // doi. org / 10.11646 / zootaxa. 4083.3.8" type="journal article">Rodríguez &amp; Navarrete-Heredia 2016</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFD5BF3CBD794" blockId="14.[151,1437,619,752]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BFD5BF3CBD794" infered="true" pageId="14" pageNumber="15">
<materialsCitation id="19903C834C26B346FF6BFD5BF3DDD7CC" ID-GBIF-Occurrence="1915497819" box="[151,1160,655,680]" country="Mexico" location="Mexico" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C26B346FF6BFD5BF42BD7CC" ID-CoL="LC6L" authority="Rodriguez &amp; Navarrete-Heredia 2016: 445" authorityName="Rodriguez &amp; Navarrete-Heredia" authorityPageNumber="445" authorityYear="2016" box="[151,894,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cifuentesi">
<emphasis id="9B8CEACC4C26B346FF6BFD5BF6DBD7CC" box="[151,398,655,680]" italics="true" pageId="14" pageNumber="15">Belonuchus cifuentesi</emphasis>
<treatmentCitation id="285910CF4C26B346FE69FD5BF42BD7CC" author="" box="[405,894,655,680]" page="445" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE69FD5BF410D7CC" box="[405,837,655,680]" pageId="14" pageNumber="15" refId="ref71443" refString="Rodriguez, W. D. &amp; Navarrete-Heredia, J. L. (2016) A new Mexican species of Belonuchus Nordmann (Coleoptera, Staphylinidae). Z ootaxa, 4083 (3), 444 - 450. https: // doi. org / 10.11646 / zootaxa. 4083.3.8" type="journal article">Rodríguez &amp; Navarrete-Heredia 2016</bibRefCitation>
: 445
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C26B346FC76FD44F494D7CC" box="[906,961,656,680]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C26B346FBD2FD5BF3DDD7CC" box="[1070,1160,655,680]" name="Mexico" pageId="14" pageNumber="15">Mexico</collectingCountry>
</materialsCitation>
: 
<materialsCitation id="19903C834C26B346FB68FD5BF64DD794" ID-GBIF-Occurrence="1915497815" collectionCode="CZUG, MUD, CNIN" collectorName="de Garcia" country="Mexico" elevation="2314" latitude="20.15" location="Cerro de Garcia" longLatPrecision="1268" longitude="-103.316666" municipality="Teocuitatlan de Corona" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="holotype">
<collectingRegion id="6B3CF83C4C26B346FB68FD5BF3B6D7CC" box="[1172,1251,655,680]" country="Mexico" name="Jalisco" pageId="14" pageNumber="15">Jalisco</collectingRegion>
, 
<collectingMunicipality id="4923ACA44C26B346FB13FD5BF649D7A8" pageId="14" pageNumber="15">Teocuitatlán de Corona</collectingMunicipality>
, 
<location id="AC2760054C26B346FEDBFD60F68ED7A8" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C26B346FEDBFD60F68ED7A8" box="[295,475,692,717]" country="Mexico" latitude="20.15" longLatPrecision="1268" longitude="-103.316666" municipality="Teocuitatlan de Corona" name="Cerro de Garcia" pageId="14" pageNumber="15" stateProvince="Jalisco">
Cerro 
<collectorName id="040D53084C26B346FE91FD60F68ED7A8" box="[365,475,692,717]" pageId="14" pageNumber="15">de García</collectorName>
</location>
, 
<geoCoordinate id="CCCC50194C26B346FE1AFD60F519D7A8" box="[486,588,692,716]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="20.15">20°09´N</geoCoordinate>
<geoCoordinate id="CCCC50194C26B346FDAFFD60F59AD7A8" box="[595,719,692,716]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="-103.316666">103°19´W</geoCoordinate>
, 
<elevation id="22D5D1ED4C26B346FD25FD60F465D7A8" box="[729,816,692,716]" metricMagnitude="3" metricUnit="m" metricValue="2.314" pageId="14" pageNumber="15" unit="m" value="2314.0">
<quantity id="6E009B3B4C26B346FD25FD60F465D7A8" box="[729,816,692,716]" metricMagnitude="3" metricUnit="m" metricValue="2.314" pageId="14" pageNumber="15" unit="m" value="2314.0">2314 m</quantity>
</elevation>
. 
<typeStatus id="7643887C4C26B346FCC7FD61F426D7A9" box="[827,883,693,717]" pageId="14" pageNumber="15">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C26B346FC01FD60F31CD7A8" box="[1021,1097,692,716]" country="Mexico" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="14" pageNumber="15">CZUG</collectionCode>
HT 
<specimenCount id="BFFEFD574C26B346FB87FD61F3C7D7A8" box="[1147,1170,693,716]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
, 117 PT (
<collectionCode id="CFE9AE1B4C26B346FB00FD60F21FD7A8" box="[1276,1354,692,716]" country="Mexico" httpUri="http://grbio.org/cool/ntgr-f7vv" name="Universidad de Guadalajara,Centro de Estudios en Zoologia, Entomologia" pageId="14" pageNumber="15">CZUG</collectionCode>
, 
<collectionCode id="CFE9AE1B4C26B346FAA8FD61F2CDD7A8" box="[1364,1432,693,716]" pageId="14" pageNumber="15">MUD</collectionCode>
, 
<collectionCode id="CFE9AE1B4C26B346FF3BFD0CF644D794" box="[199,273,728,752]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="14" pageNumber="15">CNIN</collectionCode>
)
</materialsCitation>
. Distribution: Mexico. Note: in decomposing agave (
<emphasis id="9B8CEACC4C26B346FC90FD0CF315D794" box="[876,1088,727,752]" italics="true" pageId="14" pageNumber="15">Agave inaequidens</emphasis>
) [orig.].
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFCCBF57CD65C" blockId="14.[151,1436,799,896]" box="[151,553,799,824]" pageId="14" pageNumber="15">
<emphasis id="9B8CEACC4C26B346FF6BFCCBF7F6D65C" bold="true" box="[151,163,799,824]" italics="true" pageId="14" pageNumber="15">*</emphasis>
<taxonomicName id="6EF84D5D4C26B346FF5EFCCBF57CD65C" ID-CoL="LC6M" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[162,553,799,824]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="9B8CEACC4C26B346FF5EFCCBF6F4D65C" bold="true" box="[162,417,799,824]" italics="true" pageId="14" pageNumber="15">Belonuchus clypeatus</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFC90F3B4D6E4" blockId="14.[151,1436,799,896]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BFC90F3B4D6E4" pageId="14" pageNumber="15">
<materialsCitation id="19903C834C26B346FF6BFC90F405D639" ID-GBIF-Occurrence="1915497761" box="[151,848,836,861]" location="Ega" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C26B346FF6BFC90F53CD639" ID-CoL="LC6M" authority="Sharp 1876 a: 160" authorityName="Sharp" authorityPageNumber="160" authorityYear="1876" box="[151,617,836,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="clypeatus">
<emphasis id="9B8CEACC4C26B346FF6BFC90F6DBD639" box="[151,398,836,861]" italics="true" pageId="14" pageNumber="15">Belonuchus clypeatus</emphasis>
<treatmentCitation id="285910CF4C26B346FE65FC90F53CD639" author="" box="[409,617,836,861]" page="160" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE65FC90F578D639" box="[409,557,836,861]" pageId="14" pageNumber="15" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 160
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C26B346FD84FC91F5FAD639" box="[632,687,837,861]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<location id="AC2760054C26B346FCDDFC91F405D639" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C26B346FCDDFC91F405D639" box="[801,848,837,861]" name="Ega" pageId="14" pageNumber="15">Ega</location>
</materialsCitation>
. 
<materialsCitation id="19903C834C26B346FCA2FC91F606D6E4" ID-GBIF-Occurrence="1915497777" collectionCode="BMNH" country="Brazil" location="Distribution" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<typeStatus id="7643887C4C26B346FCA2FC91F4C0D639" box="[862,917,837,861]" pageId="14" pageNumber="15">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C26B346FBD5FC91F328D638" box="[1065,1149,837,860]" pageId="14" pageNumber="15">BMNH</collectionCode>
HT. 
<location id="AC2760054C26B346FB41FC90F21CD639" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C26B346FB41FC90F21CD639" box="[1213,1353,836,861]" country="Brazil" name="Distribution" pageId="14" pageNumber="15" stateProvince="Amazonas">Distribution</location>
: 
<collectingCountry id="D1EF764E4C26B346FAABFC90F2C9D639" box="[1367,1436,836,861]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C26B346FF33FCBCF61ED6E4" box="[207,331,872,896]" country="Venezuela" name="Amazonas" pageId="14" pageNumber="15">Amazonas</collectingRegion>
)
</materialsCitation>
, Peru. Note: Peru (Newton 2015); distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C26B346FBA5FCBDF3C7D6E4" box="[1113,1170,871,896]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFC7BF536D6AC" blockId="14.[151,1437,943,1293]" box="[151,611,943,968]" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BFC7BF536D6AC" ID-CoL="LC6N" authority="Bernhauer 1908" authorityName="Bernhauer" authorityYear="1908" box="[151,611,943,968]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coelestinus">
<emphasis id="9B8CEACC4C26B346FF6BFC7BF6F2D6AC" bold="true" box="[151,423,943,968]" italics="true" pageId="14" pageNumber="15">Belonuchus coelestinus</emphasis>
<bibRefCitation id="CD694B2F4C26B346FE53FC7BF536D6AC" box="[431,611,943,968]" pageId="14" pageNumber="15" refId="ref61850" refString="Bernhauer, M. (1908) Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte, 74 (1), 283 - 372." type="journal article">Bernhauer 1908</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFC00F464D151" blockId="14.[151,1437,943,1293]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BFC00F464D151" infered="true" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BFC00F5CAD688" ID-CoL="LC6N" authority="Bernhauer 1908: 330" authorityName="Bernhauer" authorityPageNumber="330" authorityYear="1908" box="[151,671,980,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coelestinus">
<emphasis id="9B8CEACC4C26B346FF6BFC00F6F5D689" box="[151,416,980,1005]" italics="true" pageId="14" pageNumber="15">Belonuchus coelestinus</emphasis>
<treatmentCitation id="285910CF4C26B346FE56FC00F5CAD688" author="" box="[426,671,980,1005]" page="330" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE56FC00F531D688" box="[426,612,980,1005]" pageId="14" pageNumber="15" refId="ref61850" refString="Bernhauer, M. (1908) Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte, 74 (1), 283 - 372." type="journal article">Bernhauer 1908</bibRefCitation>
: 330
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C26B346FD51FC01F5B0D689" box="[685,741,981,1005]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C26B346FCA8FC00F483D689" box="[852,982,980,1005]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
. 
<typeStatus id="7643887C4C26B346FC1FFC01F34FD689" box="[995,1050,981,1005]" pageId="14" pageNumber="15">Type</typeStatus>
depository: NMW 1+ST, FMNH 1ST. Distribution: Lesser Antilles (
<collectingCountry id="D1EF764E4C26B346FD92FC2CF5A1D174" box="[622,756,1015,1040]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FCFBFC23F43ED174" box="[775,875,1015,1040]" name="Grenada" pageId="14" pageNumber="15">Grenada</collectingCountry>
). Note: distribution within Lesser Antilles (
<collectingCountry id="D1EF764E4C26B346FA82FC2CF675D151" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
) (
<bibRefCitation id="CD694B2F4C26B346FEC9FBC8F6E7D150" box="[309,434,1052,1077]" pageId="14" pageNumber="15" refId="ref71023" refString="Peck, S. B. (2010) The beetles of the island of St. Vincent, Lesser Antilles (Insecta: Coleoptera); diversity and distributions. Insecta Mundi, 0 144, 1 - 77." type="journal article">Peck 2010</bibRefCitation>
). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFB94F657D1C4" blockId="14.[151,1437,943,1293]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BFB94F657D1C4" pageId="14" pageNumber="15">
= 
<taxonomicName id="6EF84D5D4C26B346FF44FBEBF595D13C" ID-CoL="LC6R" authority="Cameron 1922: 122" authorityName="Cameron" authorityPageNumber="122" authorityYear="1922" box="[184,704,1087,1112]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coeruleus">
<emphasis id="9B8CEACC4C26B346FF44FBEBF6ECD13C" box="[184,441,1087,1112]" italics="true" pageId="14" pageNumber="15">Belonuchus coeruleus</emphasis>
<treatmentCitation id="285910CF4C26B346FE36FB94F595D13C" author="" box="[458,704,1088,1112]" page="122" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE36FB94F52BD13C" box="[458,638,1088,1112]" pageId="14" pageNumber="15" refId="ref63795" refString="Cameron, M. (1922) Descriptions of new species of Staphylinidae from the West Indies. Part II (continued). Annals and Magazine of Natural History, Series 9, 9, 113 - 128, 633 - 652. https: // doi. org / 10.1080 / 00222932208632721" type="journal article">Cameron 1922</bibRefCitation>
: 122
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C26B346FD29FB94F458D13C" box="[725,781,1088,1112]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C26B346FC70FB94F343D13C" box="[908,1046,1087,1112]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
. 
<typeStatus id="7643887C4C26B346FBD6FB94F337D13C" box="[1066,1122,1088,1112]" pageId="14" pageNumber="15">Type</typeStatus>
depository: BMNH 2ST. Distribution: Lesser Antilles (
<collectingCountry id="D1EF764E4C26B346FDE4FBB0F5CDD119" box="[536,664,1124,1149]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FD58FBB0F45DD119" box="[676,776,1124,1149]" name="Grenada" pageId="14" pageNumber="15">Grenada</collectingCountry>
). Notes: as synonym of 
<taxonomicName id="6EF84D5D4C26B346FBE5FBB1F7A8D1C4" ID-CoL="LC6N" authority="(Blackwelder 1943: 426)" authorityName="Blackwelder" authorityPageNumber="426" authorityYear="1943" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coelestinus">
<emphasis id="9B8CEACC4C26B346FBE5FBB1F3E3D119" box="[1049,1206,1124,1149]" italics="true" pageId="14" pageNumber="15">B. coelestinus</emphasis>
(
<treatmentCitation id="285910CF4C26B346FB39FBB0F7A3D1C4" author="" page="426" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FB39FBB0F7A3D1C4" pageId="14" pageNumber="15" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943: 426</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFB79F380D069" blockId="14.[151,1437,943,1293]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BFB79F380D069" infered="true" pageId="14" pageNumber="15">
= 
<taxonomicName id="6EF84D5D4C26B346FF4FFB78F52FD1A0" ID-CoL="LCBQ" authority="Cameron 1922: 122" authorityName="Cameron" authorityPageNumber="122" authorityYear="1922" box="[179,634,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="9B8CEACC4C26B346FF4FFB78F6D0D1A1" box="[179,389,1196,1221]" italics="true" pageId="14" pageNumber="15">Belonuchus smithi</emphasis>
<treatmentCitation id="285910CF4C26B346FE6EFB78F52FD1A0" author="" box="[402,634,1196,1220]" page="122" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE6EFB78F515D1A0" box="[402,576,1196,1220]" pageId="14" pageNumber="15" refId="ref63795" refString="Cameron, M. (1922) Descriptions of new species of Staphylinidae from the West Indies. Part II (continued). Annals and Magazine of Natural History, Series 9, 9, 113 - 128, 633 - 652. https: // doi. org / 10.1080 / 00222932208632721" type="journal article">Cameron 1922</bibRefCitation>
: 122
</treatmentCitation>
</taxonomicName>
(as synonym of 
<emphasis id="9B8CEACC4C26B346FCB4FB79F4B7D1A1" box="[840,994,1196,1221]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FCB4FB79F48FD1A1" ID-CoL="LC6R" authorityName="Cameron" authorityYear="1922" box="[840,986,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coeruleus">B. coeruleus</taxonomicName>
)
</emphasis>
, attributed to Fauvel 
<emphasis id="9B8CEACC4C26B346FB1AFB79F204D1A1" box="[1254,1361,1196,1221]" italics="true" pageId="14" pageNumber="15">in litteris</emphasis>
. 
<typeStatus id="7643887C4C26B346FA98FB79F2C9D1A1" box="[1380,1436,1197,1221]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C26B346FED5FB04F6FCD18C" box="[297,425,1231,1256]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
. 
<typeStatus id="7643887C4C26B346FE4EFB04F6BFD18C" box="[434,490,1232,1256]" pageId="14" pageNumber="15">Type</typeStatus>
depository: BMNH 2 ST. Distribution: Lesser Antilles (
<collectingCountry id="D1EF764E4C26B346FBA2FB04F388D18C" box="[1118,1245,1231,1256]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FB14FB1BF219D18C" box="[1256,1356,1231,1256]" name="Grenada" pageId="14" pageNumber="15">Grenada</collectingCountry>
). Note: as synonym of 
<taxonomicName id="6EF84D5D4C26B346FE8DFB21F460D069" ID-CoL="LC6N" authority="(Blackwelder, 1943: 426)" authorityName="Blackwelder" authorityPageNumber="426" authorityYear="1943" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" box="[369,821,1268,1293]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coelestinus">
<emphasis id="9B8CEACC4C26B346FE8DFB21F559D069" box="[369,524,1268,1293]" italics="true" pageId="14" pageNumber="15">B. coelestinus</emphasis>
(
<treatmentCitation id="285910CF4C26B346FDE7FB20F478D069" author="" box="[539,813,1268,1293]" page="426" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FDE7FB20F478D069" box="[539,813,1268,1293]" pageId="14" pageNumber="15" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder, 1943: 426</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; unavailable (
<bibRefCitation id="CD694B2F4C26B346FC2CFB20F39DD069" box="[976,1224,1268,1293]" pageId="14" pageNumber="15" refId="ref67781" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition, adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp. [http: // www. nhm. ac. uk / hosted-sites / iczn / code /]" type="book">ICZN 1999: Art. 11.6</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFAEFF516D031" blockId="14.[151,1436,1339,1472]" box="[151,579,1339,1365]" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BFAEFF516D031" ID-CoL="LC6P" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,579,1339,1365]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coelestis">
<emphasis id="9B8CEACC4C26B346FF6BFAEFF6DDD030" bold="true" box="[151,392,1339,1364]" italics="true" pageId="14" pageNumber="15">Belonuchus coelestis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BFA8BF37DD0F9" blockId="14.[151,1436,1339,1472]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BFA8BF37DD0F9" pageId="14" pageNumber="15">
<materialsCitation id="19903C834C26B346FF6BFA8BF67ED0F9" ID-GBIF-Occurrence="1915497825" country="Bolivia" location="Yuracares" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C26B346FF6BFA8BF465D01C" ID-CoL="LC6P" authority="Bernhauer 1916 c: 266" authorityName="Bernhauer" authorityPageNumber="266" authorityYear="1916" box="[151,816,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coelestis" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C26B346FF6BFA8BF578D01C" box="[151,557,1375,1400]" italics="true" pageId="14" pageNumber="15">Belonuchus (Musicoderus) coelestis</emphasis>
<treatmentCitation id="285910CF4C26B346FDC9FA8BF465D01C" author="" box="[565,816,1375,1400]" page="266" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FDC9FA8BF5AED01C" box="[565,763,1375,1400]" pageId="14" pageNumber="15" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 266
</treatmentCitation>
</taxonomicName>
(attributed to Fauvel 
<emphasis id="9B8CEACC4C26B346FBDBFAB5F33ED01C" box="[1063,1131,1375,1400]" italics="true" pageId="14" pageNumber="15">in litt.</emphasis>
). 
<typeStatus id="7643887C4C26B346FB7FFAB4F3EFD01C" box="[1155,1210,1376,1400]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<location id="AC2760054C26B346FADAFAB4F2CDD01C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C26B346FADAFAB4F2CDD01C" box="[1318,1432,1376,1400]" country="Bolivia" name="Yuracares" pageId="14" pageNumber="15">Yuracarès</location>
, 
<collectingCountry id="D1EF764E4C26B346FF3BFA50F67ED0F9" box="[199,299,1412,1437]" name="Bolivia" pageId="14" pageNumber="15">Bolivien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C26B346FEC9FA51F371D0F9" ID-GBIF-Occurrence="1915497805" box="[309,1060,1412,1437]" collectionCode="FMNH, ZMUH" country="Bolivia" location="Distribution" pageId="14" pageNumber="15" specimenCode="FMNH 1, ZMUH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C26B346FEC9FA51F638D0F9" box="[309,365,1413,1437]" pageId="14" pageNumber="15">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C26B346FE05FA51F50ED0F8" box="[505,603,1412,1436]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="14" pageNumber="15">FMNH 1</specimenCode>
ST; 
<specimenCode id="F95E9EA54C26B346FD71FA51F5A5D0F8" box="[653,752,1412,1436]" collectionCode="ZMUH" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="14" pageNumber="15">ZMUH 1</specimenCode>
+ST? 
<location id="AC2760054C26B346FCC4FA50F491D0F9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C26B346FCC4FA50F491D0F9" box="[824,964,1412,1437]" country="Bolivia" name="Distribution" pageId="14" pageNumber="15">Distribution</location>
: 
<collectingCountry id="D1EF764E4C26B346FC32FA50F371D0F9" box="[974,1060,1412,1437]" name="Bolivia" pageId="14" pageNumber="15">Bolivia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C26B346FF6BFA73F465D0A4" box="[151,816,1447,1472]" pageId="14" pageNumber="15" type="discussion">
<paragraph id="A94736DE4C26B346FF6BFA73F465D0A4" blockId="14.[151,1436,1339,1472]" box="[151,816,1447,1472]" pageId="14" pageNumber="15">
<emphasis id="9B8CEACC4C26B346FF6BFA73F641D0A4" bold="true" box="[151,276,1447,1472]" pageId="14" pageNumber="15">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C26B346FEB0FA73F69AD0A4" box="[332,463,1447,1472]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C26B346FEB0FA73F69AD0A4" box="[332,463,1447,1472]" italics="true" pageId="14" pageNumber="15">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C26B347FF6BFA3BF3ECD4C9" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="A94736DE4C26B346FF6BFA3BF5E6D36C" blockId="14.[151,1436,1519,1688]" box="[151,691,1519,1544]" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BFA3BF5E6D36C" ID-CoL="LC6Q" authority="Motschulsky 1858" authorityName="Motschulsky" authorityYear="1858" box="[151,691,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coeruleispennis">
<emphasis id="9B8CEACC4C26B346FF6BFA3BF689D36C" bold="true" box="[151,476,1519,1544]" italics="true" pageId="14" pageNumber="15">Belonuchus coeruleispennis</emphasis>
<bibRefCitation id="CD694B2F4C26B346FE18FA3BF5E6D36C" box="[484,691,1519,1544]" pageId="14" pageNumber="15" refId="ref69748" refString="Motschulsky, V. (1858) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. 2 - d. Article (continuation). Bulletin de la Societe Imperiale des Naturalistes de Moscou, 31 (2), 634 - 670." type="journal article">Motschulsky 1858</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BF9C0F5EDD3FC" blockId="14.[151,1436,1519,1688]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BF9C0F5EDD3FC" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BF9C0F5A1D349" ID-CoL="LC6Q" authority="Motschulsky 1858: 668" authorityName="Motschulsky" authorityPageNumber="668" authorityYear="1858" box="[151,756,1556,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coeruleispennis">
<emphasis id="9B8CEACC4C26B346FF6BF9C0F680D349" box="[151,469,1556,1581]" italics="true" pageId="14" pageNumber="15">Belonuchus coeruleispennis</emphasis>
<treatmentCitation id="285910CF4C26B346FE1CF9C0F5A1D349" author="" box="[480,756,1556,1581]" page="668" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE1CF9C0F5EDD349" box="[480,696,1556,1581]" pageId="14" pageNumber="15" refId="ref69748" refString="Motschulsky, V. (1858) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. 2 - d. Article (continuation). Bulletin de la Societe Imperiale des Naturalistes de Moscou, 31 (2), 634 - 670." type="journal article">Motschulsky 1858</bibRefCitation>
: 668
</treatmentCitation>
</taxonomicName>
. Type locality: Brésil intérieur. Type depository: ZMUM? Distribution: Brazil. Note: subsequently spelled 
<emphasis id="9B8CEACC4C26B346FD07F9E3F4F5D334" box="[763,928,1591,1616]" italics="true" pageId="14" pageNumber="15">coeruleipennis</emphasis>
(e.g., 
<treatmentCitation id="285910CF4C26B346FC13F9E3F3A3D334" author="" box="[1007,1270,1591,1616]" page="49" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FC13F9E3F39DD334" box="[1007,1224,1591,1616]" pageId="14" pageNumber="15" refId="ref69794" refString="Motschulsky, V. (1870) Genres et especes d'insectes, publies dans differents ouvrages par Victor Motschoulsky. Horae Societatis Entomologicae Rossicae, 6 (Supplement), 47 - 118." type="journal article">Motschulsky 1870</bibRefCitation>
: 49
</treatmentCitation>
; 
<treatmentCitation id="285910CF4C26B346FAFAF9E3F6F0D311" author="" page="369" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FAFAF9E3F638D310" pageId="14" pageNumber="15" refId="ref62964" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus. Vol. 57. W. Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer &amp; Schubert 1914</bibRefCitation>
: 369
</treatmentCitation>
), but 
<taxonomicName id="6EF84D5D4C26B346FE1EF988F5C7D311" ID-CoL="LC6Q" authorityName="Motschulsky" authorityYear="1858" box="[482,658,1628,1653]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coeruleispennis">
<emphasis id="9B8CEACC4C26B346FE1EF988F5C7D311" box="[482,658,1628,1653]" italics="true" pageId="14" pageNumber="15">coeruleispennis</emphasis>
</taxonomicName>
maintained as correct spelling by 
<treatmentCitation id="285910CF4C26B346FBF2F989F25DD311" author="" box="[1038,1288,1628,1653]" page="2522" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FBF2F989F397D311" box="[1038,1218,1628,1653]" pageId="14" pageNumber="15" refId="ref67417" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article">Herman (2001d</bibRefCitation>
: 2522
</treatmentCitation>
); distribution within Brazil unknown (Asenjo 
<emphasis id="9B8CEACC4C26B346FDCCF955F53DD3FC" box="[560,616,1663,1688]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BF913F573D384" blockId="14.[151,1436,1735,1977]" box="[151,550,1735,1760]" pageId="14" pageNumber="15">
<emphasis id="9B8CEACC4C26B346FF6BF913F7F6D384" bold="true" box="[151,163,1735,1760]" italics="true" pageId="14" pageNumber="15">*</emphasis>
<taxonomicName id="6EF84D5D4C26B346FF5EF913F573D384" ID-CoL="LC6S" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,550,1735,1760]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9B8CEACC4C26B346FF5EF913F6CBD3BB" bold="true" box="[162,414,1735,1760]" italics="true" pageId="14" pageNumber="15">Belonuchus cognatus</emphasis>
<bibRefCitation id="CD694B2F4C26B346FE59F913F573D384" box="[421,550,1735,1760]" pageId="14" pageNumber="15" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C26B346FF6BF938F3D2D2DC" blockId="14.[151,1436,1735,1977]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="89E811F04C26B346FF6BF938F3D2D2DC" infered="true" pageId="14" pageNumber="15">
<taxonomicName id="6EF84D5D4C26B346FF6BF938F537D260" ID-CoL="LC6S" authority="Sharp 1885: 429" authorityName="Sharp" authorityPageNumber="429" authorityYear="1885" box="[151,610,1772,1797]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="9B8CEACC4C26B346FF6BF938F6D9D260" box="[151,396,1772,1797]" italics="true" pageId="14" pageNumber="15">Belonuchus cognatus</emphasis>
<treatmentCitation id="285910CF4C26B346FE65F938F537D260" author="" box="[409,610,1772,1797]" page="429" pageId="14" pageNumber="15">
<bibRefCitation id="CD694B2F4C26B346FE65F938F576D261" box="[409,547,1772,1797]" pageId="14" pageNumber="15" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 429
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C26B346FD8FF939F5FFD261" box="[627,682,1773,1797]" pageId="14" pageNumber="15">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C26B346FCDDF938F4BFD261" box="[801,1002,1772,1797]" name="Belize" pageId="14" pageNumber="15">British Honduras</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C26B346FC07F938F313D261" box="[1019,1094,1772,1797]" country="Belize" name="Belize" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FC07F938F313D261" box="[1019,1094,1772,1797]" name="Belize" pageId="14" pageNumber="15">Belize</collectingCountry>
</collectingRegion>
; 
<collectingRegion id="6B3CF83C4C26B346FBA4F938F381D261" box="[1112,1236,1772,1797]" country="Guatemala" name="Guatemala" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FBA4F938F381D261" box="[1112,1236,1772,1797]" name="Guatemala" pageId="14" pageNumber="15">Guatemala</collectingCountry>
</collectingRegion>
, El Reposo, Las Mercedes, San Isidro, Mirandilla; 
<collectingCountry id="D1EF764E4C26B346FDAFF8DBF59CD24C" box="[595,713,1807,1832]" name="Nicaragua" pageId="14" pageNumber="15">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C26B346FD2BF8DBF41ED24C" box="[727,843,1807,1832]" country="Nicaragua" name="Chontales" pageId="14" pageNumber="15">Chontales</collectingRegion>
; 
<collectingRegion id="6B3CF83C4C26B346FCA5F8C4F4E0D243" box="[857,949,1808,1831]" country="Panama" name="Panama" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FCA5F8C4F4E0D243" box="[857,949,1808,1831]" name="Panama" pageId="14" pageNumber="15">Panama</collectingCountry>
</collectingRegion>
, Bugaba. 
<typeStatus id="7643887C4C26B346FBD5F8C4F335D24C" box="[1065,1120,1808,1832]" pageId="14" pageNumber="15">Type</typeStatus>
depository: BMNH #? ST: several, from all 
<typeStatus id="7643887C4C26B346FE78F8E1F6E1D229" box="[388,436,1845,1869]" pageId="14" pageNumber="15">type</typeStatus>
localities [MCP seen]; IRSNB 5 ST: 4 Bugaba, two of them labelled as “Ext-ypis” [MCP seen]; 1 
<collectingRegion id="6B3CF83C4C26B346FED7F883F6CED214" box="[299,411,1879,1904]" country="Nicaragua" name="Chontales" pageId="14" pageNumber="15">Chontales</collectingRegion>
[MCP seen]; FMNH 3 ST: 3 Bugaba. Distribution: 
<collectingRegion id="6B3CF83C4C26B346FC0BF883F314D214" box="[1015,1089,1879,1904]" country="Belize" name="Belize" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FC0BF883F314D214" box="[1015,1089,1879,1904]" name="Belize" pageId="14" pageNumber="15">Belize</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C26B346FBB3F883F39ED214" box="[1103,1227,1879,1904]" country="Guatemala" name="Guatemala" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FBB3F883F39ED214" box="[1103,1227,1879,1904]" name="Guatemala" pageId="14" pageNumber="15">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C26B346FB25F883F21AD214" box="[1241,1359,1879,1904]" name="Nicaragua" pageId="14" pageNumber="15">Nicaragua</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FAA0F88CF7A8D2F1" name="Costa Rica" pageId="14" pageNumber="15">
<collectingRegion id="6B3CF83C4C26B346FAA0F88CF2C9D214" box="[1372,1436,1880,1904]" country="Tanzania" name="Pwani" pageId="14" pageNumber="15">Costa</collectingRegion>
Rica
</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C26B346FEFBF8A9F636D2F0" box="[263,355,1917,1940]" country="Panama" name="Panama" pageId="14" pageNumber="15">
<collectingCountry id="D1EF764E4C26B346FEFBF8A9F636D2F0" box="[263,355,1917,1940]" name="Panama" pageId="14" pageNumber="15">Panama</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C26B346FE92F8A8F68AD2F1" box="[366,479,1916,1941]" name="Colombia" pageId="14" pageNumber="15">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FE15F8A8F50AD2F1" box="[489,607,1916,1941]" name="Venezuela" pageId="14" pageNumber="15">Venezuela</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FD96F8A8F599D2F1" box="[618,716,1916,1941]" name="Trinidad and Tobago" pageId="14" pageNumber="15">Trinidad</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FD2BF8A8F44BD2F1" box="[727,798,1916,1941]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FCD4F8A8F4CED2F1" box="[808,923,1916,1941]" name="Argentina" pageId="14" pageNumber="15">Argentina</collectingCountry>
, 
<collectingCountry id="D1EF764E4C26B346FC59F8A8F370D2F1" box="[933,1061,1916,1941]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
. Notes: distribution within Lesser Antilles (
<collectingCountry id="D1EF764E4C26B346FED3F874F6F8D2DD" box="[303,429,1952,1977]" name="Saint Vincent and the Grenadines" pageId="14" pageNumber="15">St. Vincent</collectingCountry>
) (
<bibRefCitation id="CD694B2F4C26B346FE3FF874F515D2DC" box="[451,576,1952,1977]" pageId="14" pageNumber="15" refId="ref71023" refString="Peck, S. B. (2010) The beetles of the island of St. Vincent, Lesser Antilles (Insecta: Coleoptera); diversity and distributions. Insecta Mundi, 0 144, 1 - 77." type="journal article">Peck 2010</bibRefCitation>
); within (and including) 
<collectingCountry id="D1EF764E4C26B346FCA8F874F4CDD2DD" box="[852,920,1952,1977]" name="Brazil" pageId="14" pageNumber="15">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C26B346FC03F875F36DD2DD" box="[1023,1080,1952,1977]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFF43F545D5D4" blockId="15.[151,1437,151,285]" box="[151,528,151,176]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BFF43F7F6D5D4" bold="true" box="[151,163,151,176]" italics="true" pageId="15" pageNumber="16">*</emphasis>
<taxonomicName id="6EF84D5D4C27B347FF5EFF43F545D5D4" ID-CoL="LC6T" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,528,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="colon">
<emphasis id="9B8CEACC4C27B347FF5EFF43F623D5D4" bold="true" box="[162,374,151,176]" italics="true" pageId="15" pageNumber="16">Belonuchus colon</emphasis>
(
<bibRefCitation id="CD694B2F4C27B347FE79FF43F55DD5D4" box="[389,520,151,176]" pageId="15" pageNumber="16" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFF68F57AD479" blockId="15.[151,1437,151,285]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BFF68F57AD479" infered="true" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFF68F54AD5B0" ID-CoL="779JJ" authority="Sharp 1885: 419" authorityName="Sharp" authorityPageNumber="419" authorityYear="1885" box="[151,543,188,213]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="colon">
<emphasis id="9B8CEACC4C27B347FF6BFF68F602D5B1" box="[151,343,188,213]" italics="true" pageId="15" pageNumber="16">Philonthus colon</emphasis>
<treatmentCitation id="285910CF4C27B347FEA3FF68F54AD5B0" author="" box="[351,543,188,213]" page="419" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FEA3FF68F6B0D5B1" box="[351,485,188,213]" pageId="15" pageNumber="16" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 419
</treatmentCitation>
</taxonomicName>
. Type locality: Mexico: Cordova. Type depository: BMNH 2 ST. Distribution: Mexico. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C27B347FDC7FF0BF5E1D59C" ID-CoL="8VRYR" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[571,692,223,248]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C27B347FDC7FF0BF5E1D59C" box="[571,692,223,248]" italics="true" pageId="15" pageNumber="16">Philonthus</emphasis>
</taxonomicName>
, distribution (
<treatmentCitation id="285910CF4C27B347FCA1FF0BF254D59C" author="" box="[861,1281,223,248]" page="334" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FCA1FF0BF397D59C" box="[861,1218,223,248]" pageId="15" pageNumber="16" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C27B347FBC0FF35F32FD59C" box="[1084,1146,223,248]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2002
</bibRefCitation>
: 334
</treatmentCitation>
); distribution within Mexico (
<bibRefCitation id="CD694B2F4C27B347FE85FED0F576D479" box="[377,547,260,285]" pageId="15" pageNumber="16" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFE9FF532D401" blockId="15.[151,1436,331,464]" box="[151,615,331,357]" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFE9FF532D401" ID-CoL="LC6V" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,615,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="columbicus">
<emphasis id="9B8CEACC4C27B347FF6BFE9FF6F9D400" bold="true" box="[151,428,331,356]" italics="true" pageId="15" pageNumber="16">Belonuchus columbicus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFEBBF3ECD4C9" blockId="15.[151,1436,331,464]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BFEBBF3ECD4C9" pageId="15" pageNumber="16">
<materialsCitation id="19903C834C27B347FF6BFEBBF3E0D4C9" ID-GBIF-Occurrence="1915497808" collectionCode="FMNH" collectorName="West-Columbien &amp; Umgebung von Cali" country="Colombia" elevation="2000" location="St. Antonio" municipality="ST. Distribution" pageId="15" pageNumber="16" specimenCode="FMNH 2" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C27B347FF6BFEBBF437D4EC" ID-CoL="LC6V" authority="Bernhauer 1916 c: 271" authorityName="Bernhauer" authorityPageNumber="271" authorityYear="1916" box="[151,866,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="columbicus" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C27B347FF6BFEBBF506D4EC" box="[151,595,367,392]" italics="true" pageId="15" pageNumber="16">Belonuchus (Musicoderus) columbicus</emphasis>
<treatmentCitation id="285910CF4C27B347FDA1FEBBF437D4EC" author="" box="[605,866,367,392]" page="271" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FDA1FEBBF472D4EC" box="[605,807,367,392]" pageId="15" pageNumber="16" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 271
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FC8DFEA4F4FCD4EC" box="[881,937,368,392]" pageId="15" pageNumber="16">Type</typeStatus>
locality: 
<collectorName id="040D53084C27B347FBE6FEBBF389D4EC" box="[1050,1244,367,392]" pageId="15" pageNumber="16">West-Columbien</collectorName>
: 
<collectorName id="040D53084C27B347FB16FEBBF7A3D4C9" pageId="15" pageNumber="16">Umgebung von Cali</collectorName>
(
<location id="AC2760054C27B347FEF9FE40F6DFD4C9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C27B347FEF9FE40F6DFD4C9" box="[261,394,404,429]" country="Colombia" municipality="ST. Distribution" name="St. Antonio" pageId="15" pageNumber="16">St. Antonio</location>
, 
<elevation id="22D5D1ED4C27B347FE69FE40F6B2D4C8" box="[405,487,404,428]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="m" value="2000.0">
<quantity id="6E009B3B4C27B347FE69FE40F6B2D4C8" box="[405,487,404,428]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="m" value="2000.0">2000m</quantity>
</elevation>
). 
<typeStatus id="7643887C4C27B347FE06FE41F564D4C9" box="[506,561,405,429]" pageId="15" pageNumber="16">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C27B347FD41FE41F476D4C8" box="[701,803,404,428]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="16">FMNH 2</specimenCode>
ST;? 1 
<collectingMunicipality id="4923ACA44C27B347FC7CFE40F36CD4C9" box="[896,1081,404,429]" pageId="15" pageNumber="16">ST. Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C27B347FBBFFE40F3E0D4C9" box="[1091,1205,404,429]" name="Colombia" pageId="15" pageNumber="16">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C27B347FF6BFE63F465D4B4" box="[151,816,439,464]" pageId="15" pageNumber="16" type="discussion">
<paragraph id="A94736DE4C27B347FF6BFE63F465D4B4" blockId="15.[151,1436,331,464]" box="[151,816,439,464]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BFE63F641D4B4" bold="true" box="[151,276,439,464]" pageId="15" pageNumber="16">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C27B347FEB0FE63F69AD4B4" box="[332,463,439,464]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C27B347FEB0FE63F69AD4B4" box="[332,463,439,464]" italics="true" pageId="15" pageNumber="16">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C27B347FF6BFE2BF53DD133" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="A94736DE4C27B347FF6BFE2BF54CD77C" blockId="15.[151,1437,511,645]" box="[151,537,511,536]" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFE2BF54CD77C" ID-CoL="LC6W" authority="(Say 1830)" authorityName="Say" authorityYear="1830" baseAuthorityName="Say" baseAuthorityYear="1830" box="[151,537,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="connexus">
<emphasis id="9B8CEACC4C27B347FF6BFE2BF6C3D773" bold="true" box="[151,406,511,536]" italics="true" pageId="15" pageNumber="16">Belonuchus connexus</emphasis>
(
<bibRefCitation id="CD694B2F4C27B347FE5AFDD4F544D77C" box="[422,529,512,536]" pageId="15" pageNumber="16" refId="ref71623" refString="Say, T. (1830) Descriptions of new species of North American insects, and observations on some already described. New Harmony, Indiana, pp. 1 - 41." type="book chapter">Say 1830</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFDF0F474D7E1" blockId="15.[151,1437,511,645]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BFDF0F474D7E1" infered="true" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFDF0F573D759" ID-CoL="4ZJW9" authority="Say 1830: 35" authorityName="Say" authorityPageNumber="35" authorityYear="1830" box="[151,550,548,573]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="connexus">
<emphasis id="9B8CEACC4C27B347FF6BFDF0F6DDD758" box="[151,392,548,573]" italics="true" pageId="15" pageNumber="16">Staphylinus connexus</emphasis>
<treatmentCitation id="285910CF4C27B347FE73FDF0F573D759" author="" box="[399,550,548,573]" page="35" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FE73FDF0F6A8D758" box="[399,509,548,572]" pageId="15" pageNumber="16" refId="ref71623" refString="Say, T. (1830) Descriptions of new species of North American insects, and observations on some already described. New Harmony, Indiana, pp. 1 - 41." type="book chapter">Say 1830</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
, also 1834: 448. 
<typeStatus id="7643887C4C27B347FD1BFDF1F44AD759" box="[743,799,549,573]" pageId="15" pageNumber="16">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C27B347FC75FDF0F4B7D759" box="[905,994,548,573]" country="Mexico" name="Mexico" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FC75FDF0F4B7D759" box="[905,994,548,573]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
</collectingRegion>
. 
<typeStatus id="7643887C4C27B347FC11FDF1F371D759" box="[1005,1060,549,573]" pageId="15" pageNumber="16">Type</typeStatus>
depository:? #?ST. Distribution: 
<collectingRegion id="6B3CF83C4C27B347FF3BFD93F675D704" box="[199,288,583,608]" country="Mexico" name="Mexico" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FF3BFD93F675D704" box="[199,288,583,608]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
</collectingRegion>
. Note: comb. nov. ex 
<taxonomicName id="6EF84D5D4C27B347FDC8FD93F494D704" ID-CoL="8VS43" authority="(Erichson 1840: 927)" authorityName="Erichson" authorityPageNumber="927" authorityYear="1840" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[564,961,583,608]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C27B347FDC8FD93F5E3D704" box="[564,694,583,608]" italics="true" pageId="15" pageNumber="16">Staphylinus</emphasis>
(
<treatmentCitation id="285910CF4C27B347FD37FD93F4ECD704" author="" box="[715,953,583,608]" page="927" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FD37FD93F4ECD704" box="[715,953,583,608]" pageId="15" pageNumber="16" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840: 927</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; distribution within 
<collectingRegion id="6B3CF83C4C27B347FB44FD93F25BD704" box="[1208,1294,583,608]" country="Mexico" name="Mexico" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FB44FD93F25BD704" box="[1208,1294,583,608]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
</collectingRegion>
(Navarrete- 
<collectingRegion id="6B3CF83C4C27B347FF3BFDB8F675D7E1" box="[199,288,620,645]" country="Costa Rica" name="Heredia" pageId="15" pageNumber="16">Heredia</collectingRegion>
<emphasis id="9B8CEACC4C27B347FEDAFDB9F60AD7E1" box="[294,351,620,645]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2002). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFD67F534D7A8" blockId="15.[151,1436,691,824]" box="[151,609,691,717]" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFD67F534D7A8" ID-CoL="LC6X" authority="(Bernhauer 1908)" authorityName="Bernhauer" authorityYear="1908" baseAuthorityName="Bernhauer" baseAuthorityYear="1908" box="[151,609,691,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="9B8CEACC4C27B347FF6BFD67F6C6D7AF" bold="true" box="[151,403,691,716]" italics="true" pageId="15" pageNumber="16">Belonuchus convexus</emphasis>
(
<bibRefCitation id="CD694B2F4C27B347FE5EFD60F50DD7A8" box="[418,600,692,717]" pageId="15" pageNumber="16" refId="ref61850" refString="Bernhauer, M. (1908) Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte, 74 (1), 283 - 372." type="journal article">Bernhauer 1908</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFD03F6AFD65C" blockId="15.[151,1436,691,824]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BFD03F6AFD65C" infered="true" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFD03F5F1D794" ID-CoL="6RQR7" authority="Bernhauer 1908: 331" authorityName="Bernhauer" authorityPageNumber="331" authorityYear="1908" box="[151,676,727,752]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="9B8CEACC4C27B347FF6BFD03F6CED794" box="[151,411,727,752]" italics="true" pageId="15" pageNumber="16">Musicoderus convexus</emphasis>
<treatmentCitation id="285910CF4C27B347FE55FD03F5F1D794" author="" box="[425,676,727,752]" page="331" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FE55FD03F532D794" box="[425,615,727,752]" pageId="15" pageNumber="16" refId="ref61850" refString="Bernhauer, M. (1908) Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte, 74 (1), 283 - 372." type="journal article">Bernhauer 1908</bibRefCitation>
: 331
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FD4AFD0CF5BBD794" box="[694,750,728,752]" pageId="15" pageNumber="16">Type</typeStatus>
locality: Pozuzo, 
<collectingCountry id="D1EF764E4C27B347FC36FD0CF354D78B" box="[970,1025,728,751]" name="Peru" pageId="15" pageNumber="16">Peru</collectingCountry>
. 
<typeStatus id="7643887C4C27B347FBEEFD0CF31CD794" box="[1042,1097,728,752]" pageId="15" pageNumber="16">Type</typeStatus>
depository: FMNH HT 
<specimenCount id="BFFEFD574C27B347FA8FFD0CF2D2D794" box="[1395,1415,728,752]" pageId="15" pageNumber="16" type="female">♀</specimenCount>
?: Distribution: 
<collectingCountry id="D1EF764E4C27B347FEA3FD29F6C0D670" box="[351,405,765,788]" name="Peru" pageId="15" pageNumber="16">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C27B347FE5EFD28F54CD671" box="[418,537,764,789]" name="Nicaragua" pageId="15" pageNumber="16">Nicaragua</collectingCountry>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C27B347FCDDFD28F4E7D671" ID-CoL="8XMY8" baseAuthorityName="Scheerpeltz 1933:" baseAuthorityYear="1377" box="[801,946,764,789]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C27B347FCDDFD28F4E7D671" box="[801,946,764,789]" italics="true" pageId="15" pageNumber="16">Musicoderus</emphasis>
</taxonomicName>
, distribution 
<collectingCountry id="D1EF764E4C27B347FBB7FD28F3EBD671" box="[1099,1214,764,789]" name="Nicaragua" pageId="15" pageNumber="16">Nicaragua</collectingCountry>
(
<treatmentCitation id="285910CF4C27B347FB33FD28F656D65C" author="" page="1377" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FB33FD28F2CDD670" box="[1231,1432,764,789]" pageId="15" pageNumber="16" refId="ref71693" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Vol. 129. W. Junk, Berlin, pp. 989 - 1500." type="book chapter">Scheerpeltz 1933</bibRefCitation>
: 1377
</treatmentCitation>
); 
<collectingCountry id="D1EF764E4C27B347FEEBFCF4F61FD653" box="[279,330,800,823]" name="Peru" pageId="15" pageNumber="16">Peru</collectingCountry>
(
<bibRefCitation id="CD694B2F4C27B347FEA5FCCBF6BBD65C" box="[345,494,799,824]" pageId="15" pageNumber="16" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFCB3F528D6E4" blockId="15.[151,1436,871,968]" box="[151,637,871,896]" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFCB3F528D6E4" ID-CoL="LC6Z" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,637,871,896]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cordilleranus">
<emphasis id="9B8CEACC4C27B347FF6BFCB3F697D6E4" bold="true" box="[151,450,871,896]" italics="true" pageId="15" pageNumber="16">Belonuchus cordilleranus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFC58F3BFD6AC" blockId="15.[151,1436,871,968]" pageId="15" pageNumber="16">
<materialsCitation id="19903C834C27B347FF6BFC58F3BFD6AC" ID-GBIF-Occurrence="1915497799" collectionCode="FMNH" country="Colombia" elevation="675" location="ST. Distribution" pageId="15" pageNumber="16" specimenCode="FMNH 3" specimenCount="2" typeStatus="holotype">
<treatmentCitationGroup id="89E811F04C27B347FF6BFC58F3BFD6AC" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFC58F43CD6C0" ID-CoL="LC6Z" authority="Bernhauer 1916 c: 270" authorityName="Bernhauer" authorityPageNumber="270" authorityYear="1916" box="[151,873,908,933]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cordilleranus" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C27B347FF6BFC58F536D6C1" box="[151,611,908,933]" italics="true" pageId="15" pageNumber="16">Belonuchus (Musicoderus) cordilleranus</emphasis>
<treatmentCitation id="285910CF4C27B347FD97FC58F43CD6C0" author="" box="[619,873,908,933]" page="270" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FD97FC58F464D6C1" box="[619,817,908,933]" pageId="15" pageNumber="16" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 270
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FC89FC59F4F8D6C1" box="[885,941,909,933]" pageId="15" pageNumber="16">Type</typeStatus>
locality: Columbien: Ost-Cordilleren, 
<elevation id="22D5D1ED4C27B347FA98FC58F65FD6AC" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="9.0" metricValueMin="4.5" pageId="15" pageNumber="16" unit="m" value="675.0" valueMax="900.0" valueMin="450.0">
<quantity id="6E009B3B4C27B347FA98FC58F65FD6AC" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="9.0" metricValueMin="4.5" pageId="15" pageNumber="16" unit="m" value="675.0" valueMax="900.0" valueMin="450.0">450– 900m</quantity>
</elevation>
. 
<typeStatus id="7643887C4C27B347FEE9FC64F618D6AC" box="[277,333,944,968]" pageId="15" pageNumber="16">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C27B347FE24FC64F568D6AC" box="[472,573,944,968]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="16">FMNH 3</specimenCode>
<location id="AC2760054C27B347FDC1FC64F5A2D6AC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C27B347FDC1FC64F5A2D6AC" box="[573,759,943,968]" country="Colombia" name="ST. Distribution" pageId="15" pageNumber="16">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C27B347FCFDFC7BF426D6AC" box="[769,883,943,968]" name="Colombia" pageId="15" pageNumber="16">Colombia</collectingCountry>
. Note: generic affiliation unclear.
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFC23F41FD174" blockId="15.[151,1437,1015,1328]" box="[151,842,1015,1040]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BFC23F7F6D174" bold="true" box="[151,163,1015,1040]" italics="true" pageId="15" pageNumber="16">*</emphasis>
<taxonomicName id="6EF84D5D4C27B347FF5EFC23F569D174" ID-CoL="LC72" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,572,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="corticalis" status="comb. nov.">
<emphasis id="9B8CEACC4C27B347FF5EFC23F6F5D174" bold="true" box="[162,416,1015,1040]" italics="true" pageId="15" pageNumber="16">Belonuchus corticalis</emphasis>
(
<bibRefCitation id="CD694B2F4C27B347FE53FC23F561D174" box="[431,564,1015,1040]" pageId="15" pageNumber="16" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C27B347FDB4FC23F59DD174" box="[584,712,1015,1040]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FDB4FC23F59DD174" bold="true" box="[584,712,1015,1040]" pageId="15" pageNumber="16">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C27B347FD24FC23F417D174" box="[728,834,1015,1040]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="15" pageNumber="16">Figs. 5–6</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFBC8F53DD133" blockId="15.[151,1437,1015,1328]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BFBC8F53DD133" pageId="15" pageNumber="16">
<materialsCitation id="19903C834C27B347FF6BFBC8F386D151" ID-GBIF-Occurrence="1915497775" box="[151,1235,1052,1077]" country="Panama" elevation="762" location="Chiriqui" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C27B347FF6BFBC8F512D150" ID-CoL="6V9M7" authority="Sharp 1885: 413" authorityName="Sharp" authorityPageNumber="413" authorityYear="1885" box="[151,583,1052,1077]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="corticalis">
<emphasis id="9B8CEACC4C27B347FF6BFBC8F6D7D151" box="[151,386,1052,1077]" italics="true" pageId="15" pageNumber="16">Philonthus corticalis</emphasis>
<treatmentCitation id="285910CF4C27B347FE75FBC8F512D150" author="" box="[393,583,1052,1077]" page="413" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FE75FBC8F55AD151" box="[393,527,1052,1077]" pageId="15" pageNumber="16" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 413
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FDAFFBC9F5DED151" box="[595,651,1053,1077]" pageId="15" pageNumber="16">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C27B347FD0AFBC9F407D150" box="[758,850,1053,1076]" name="Panama" pageId="15" pageNumber="16">Panama</collectingCountry>
, Volcan de 
<collectingRegion id="6B3CF83C4C27B347FC28FBC8F364D151" box="[980,1073,1052,1077]" country="Panama" name="Chiriqui" pageId="15" pageNumber="16">Chiriqui</collectingRegion>
<elevation id="22D5D1ED4C27B347FBC5FBC8F386D151" box="[1081,1235,1052,1077]" metricMagnitude="2" metricUnit="m" metricValue="7.62" metricValueMax="9.144" metricValueMin="6.096" pageId="15" pageNumber="16" unit="ft" value="2500.0" valueMax="3000.0" valueMin="2000.0">
<quantity id="6E009B3B4C27B347FBC5FBC8F386D151" box="[1081,1235,1052,1077]" metricMagnitude="2" metricUnit="m" metricValue="7.62" metricValueMax="9.144" metricValueMin="6.096" pageId="15" pageNumber="16" unit="ft" value="2500.0" valueMax="3000.0" valueMin="2000.0">2000–3000 ft</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C27B347FB23FBC9F536D133" ID-GBIF-Occurrence="1915497732" collectionCode="BMNH" country="Panama" location="Distribution" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C27B347FB23FBC9F243D151" box="[1247,1302,1053,1077]" pageId="15" pageNumber="16">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C27B347FF3BFB94F64ED133" box="[199,283,1088,1111]" pageId="15" pageNumber="16">BMNH</collectionCode>
HT 
<specimenCount id="BFFEFD574C27B347FEB2FB94F633D133" box="[334,358,1088,1111]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
. 
<location id="AC2760054C27B347FE8DFBEBF6A8D13C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C27B347FE8DFBEBF6A8D13C" box="[369,509,1087,1112]" country="Panama" name="Distribution" pageId="15" pageNumber="16">Distribution</location>
: 
<collectingCountry id="D1EF764E4C27B347FDFBFB94F536D133" box="[519,611,1088,1111]" name="Panama" pageId="15" pageNumber="16">Panama</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C27B347FF6BFBB0F48BD119" box="[151,990,1124,1149]" pageId="15" pageNumber="16" type="discussion">
<paragraph id="A94736DE4C27B347FF6BFBB0F48BD119" blockId="15.[151,1437,1015,1328]" box="[151,990,1124,1149]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BFBB0F65ED119" bold="true" box="[151,267,1124,1149]" pageId="15" pageNumber="16">Remarks.</emphasis>
The “unique” specimen in BMNH is confirmed as the 
<typeStatus id="7643887C4C27B347FC88FBB0F48FD119" box="[884,986,1124,1149]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C27B347FF6BFB53F525D069" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="A94736DE4C27B347FF6BFB53F525D069" blockId="15.[151,1437,1015,1328]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BFB53F694D1C4" bold="true" box="[151,449,1159,1184]" pageId="15" pageNumber="16">
<typeStatus id="7643887C4C27B347FF6BFB53F787D1C4" box="[151,210,1159,1184]" pageId="15" pageNumber="16">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C27B347FE35FB53F564D1C4" box="[457,561,1159,1184]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
<specimenCount id="BFFEFD574C27B347FDC5FB5CF51BD1FB" box="[569,590,1160,1183]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
(
<collectionCode id="CFE9AE1B4C27B347FDA2FB5CF5E2D1C4" box="[606,695,1160,1184]" pageId="15" pageNumber="16">BMNH</collectionCode>
, 
<figureCitation id="31C32A5B4C27B347FD3EFB53F465D1C4" box="[706,816,1159,1184]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="15" pageNumber="16">Figs. 5–6</figureCitation>
): card-mounted, “ 
<specimenCount id="BFFEFD574C27B347FC07FB5CF344D1C4" box="[1019,1041,1160,1184]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
(in ink) 
<taxonomicName id="6EF84D5D4C27B347FB8AFB53F209D1C4" box="[1142,1372,1159,1184]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="corticalis">Philonthus corticalis</taxonomicName>
<typeStatus id="7643887C4C27B347FA98FB5CF2C9D1C4" box="[1380,1436,1160,1184]" pageId="15" pageNumber="16">Type</typeStatus>
D.S. V. de 
<collectingRegion id="6B3CF83C4C27B347FEC2FB78F6CED1A1" box="[318,411,1196,1221]" country="Panama" name="Chiriqui" pageId="15" pageNumber="16">Chiriqui</collectingRegion>
<quantity id="6E009B3B4C27B347FE5EFB78F545D1A1" box="[418,528,1196,1221]" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" pageId="15" pageNumber="16" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">2–3000 ft</quantity>
. [in Sharp’s hand] // 
<typeStatus id="7643887C4C27B347FD0AFB79F478D1A1" box="[758,813,1197,1221]" pageId="15" pageNumber="16">Type</typeStatus>
(round white label with red margin) // V. de 
<collectingRegion id="6B3CF83C4C27B347FAE0FB78F22CD1A1" box="[1308,1401,1196,1221]" country="Panama" name="Chiriqui" pageId="15" pageNumber="16">Chiriqui</collectingRegion>
<quantity id="6E009B3B4C27B347FA7CFB78F648D18C" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" pageId="15" pageNumber="16" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">2– 3000 ft</quantity>
., 
<quantity id="6E009B3B4C27B347FED3FB04F6CAD18C" box="[303,415,1231,1256]" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" pageId="15" pageNumber="16" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">2–3000 ft</quantity>
. Champion (white label) // B. C. A. Col. 
<date id="DD46101E4C27B347FC84FB04F4F0D18C" box="[888,933,1232,1256]" pageId="15" pageNumber="16">I. 2.</date>
<taxonomicName id="6EF84D5D4C27B347FC52FB1BF3B8D18C" authority=", Sharp." authorityName="Sharp." box="[942,1261,1231,1256]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="corticalis">Philonthus corticalis, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C27B347FF6BFAC3F501D01C" pageId="15" pageNumber="16" type="discussion">
<paragraph id="A94736DE4C27B347FF6BFAC3F33DD054" blockId="15.[151,1437,1015,1328]" box="[151,1128,1303,1328]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BFAC3F64AD054" bold="true" box="[151,287,1303,1328]" pageId="15" pageNumber="16">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C27B347FD46FAC3F331D054" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1303,1328]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C27B347FD46FAC3F468D054" box="[698,829,1303,1328]" italics="true" pageId="15" pageNumber="16">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C27B347FCB7FAC3F309D054" box="[843,1116,1303,1328]" pageId="15" pageNumber="16" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C27B347FC1CFACDF34DD054" box="[992,1048,1303,1328]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="A94736DE4C27B347FF6BFA8BF501D01C" blockId="15.[151,1437,1375,1509]" box="[151,596,1375,1400]" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFA8BF501D01C" authority="Bernhauer 1921" authorityName="Bernhauer" authorityYear="1921" box="[151,596,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cribriceps">
<emphasis id="9B8CEACC4C27B347FF6BFA8BF6CFD01C" bold="true" box="[151,410,1375,1400]" italics="true" pageId="15" pageNumber="16">Belonuchus cribriceps</emphasis>
Bernhauer 1921
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C27B359FF6BFA50F68BD4EC" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="A94736DE4C27B347FF6BFA50F5D7D081" blockId="15.[151,1437,1375,1509]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BFA50F5D7D081" infered="true" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BFA50F5CDD0F8" ID-CoL="LC73" authority="Bernhauer 1921 c: 172" authorityName="Bernhauer" authorityPageNumber="172" authorityYear="1921" box="[151,664,1412,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cribriceps">
<emphasis id="9B8CEACC4C27B347FF6BFA50F6C6D0F9" box="[151,403,1412,1437]" italics="true" pageId="15" pageNumber="16">Belonuchus cribriceps</emphasis>
<treatmentCitation id="285910CF4C27B347FE67FA50F5CDD0F8" author="" box="[411,664,1412,1437]" page="172" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FE67FA50F535D0F8" box="[411,608,1412,1437]" pageId="15" pageNumber="16" refId="ref62483" refString="Bernhauer, M. (1921 c) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article">Bernhauer 1921c</bibRefCitation>
: 172
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FD58FA51F58ED0F9" box="[676,731,1413,1437]" pageId="15" pageNumber="16">Type</typeStatus>
locality: San Roque, 
<collectingRegion id="6B3CF83C4C27B347FC2DFA50F313D0F9" box="[977,1094,1412,1437]" country="Argentina" name="Corrientes" pageId="15" pageNumber="16">Corrientes</collectingRegion>
[
<collectingCountry id="D1EF764E4C27B347FBA8FA50F398D0F9" box="[1108,1229,1412,1437]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
]. 
<typeStatus id="7643887C4C27B347FB22FA51F243D0F9" box="[1246,1302,1413,1437]" pageId="15" pageNumber="16">Type</typeStatus>
depository: FMNH 1 ST. Distribution: 
<collectingCountry id="D1EF764E4C27B347FDFDFA73F526D0A4" box="[513,627,1447,1472]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
, 
<collectingCountry id="D1EF764E4C27B347FD7DFA73F59CD0A4" box="[641,713,1447,1472]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
. Note: distribution within (and including) 
<collectingCountry id="D1EF764E4C27B347FB4EFA73F3A3D0A4" box="[1202,1270,1447,1472]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C27B347FA9DFA7DF2C9D0A4" box="[1377,1436,1447,1472]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2013). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BF9C7F53CD348" blockId="15.[151,1436,1555,1653]" box="[151,617,1555,1581]" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BF9C7F53CD348" ID-CoL="LC74" authority="Blackwelder 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,617,1555,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="danforthi">
<emphasis id="9B8CEACC4C27B347FF6BF9C7F6C0D348" bold="true" box="[151,405,1555,1580]" italics="true" pageId="15" pageNumber="16">Belonuchus danforthi</emphasis>
<bibRefCitation id="CD694B2F4C27B347FE60F9C0F53CD348" box="[412,617,1556,1581]" pageId="15" pageNumber="16" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BF9E3F3A4D311" blockId="15.[151,1436,1555,1653]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BF9E3F3A4D311" pageId="15" pageNumber="16">
<materialsCitation id="19903C834C27B347FF6BF9E3F50CD311" ID-GBIF-Occurrence="1915497764" country="Puerto Rico" elevation="914" location="Maricao" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Maricao" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C27B347FF6BF9E3F5C3D334" ID-CoL="LC74" authority="Blackwelder 1943: 429" authorityName="Blackwelder" authorityPageNumber="429" authorityYear="1943" box="[151,662,1591,1616]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="danforthi">
<emphasis id="9B8CEACC4C27B347FF6BF9E3F6DDD334" box="[151,392,1591,1616]" italics="true" pageId="15" pageNumber="16">Belonuchus danforthi</emphasis>
<treatmentCitation id="285910CF4C27B347FE73F9E3F5C3D334" author="" box="[399,662,1591,1616]" page="429" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FE73F9E3F50AD334" box="[399,607,1591,1616]" pageId="15" pageNumber="16" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 429
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FD5DF9ECF58DD334" box="[673,728,1592,1616]" pageId="15" pageNumber="16">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C27B347FCBDF9ECF49FD334" box="[833,970,1591,1616]" name="Puerto Rico" pageId="15" pageNumber="16">Puerto Rico</collectingCountry>
: about 
<quantity id="6E009B3B4C27B347FBE5F9ECF32DD334" box="[1049,1144,1591,1616]" metricMagnitude="4" metricUnit="m" metricValue="2.0921472" pageId="15" pageNumber="16" unit="mi" value="13.0">13 miles</quantity>
east of 
<collectingRegion id="6B3CF83C4C27B347FB32F9E3F27ED334" box="[1230,1323,1591,1616]" country="Puerto Rico" name="Maricao" pageId="15" pageNumber="16">Maricao</collectingRegion>
and north of 
<collectingRegion id="6B3CF83C4C27B347FF19F989F67BD310" box="[229,302,1629,1652]" country="Puerto Rico" name="Yauco" pageId="15" pageNumber="16">Yauco</collectingRegion>
, elevation about 
<elevation id="22D5D1ED4C27B347FE10F988F50CD311" box="[492,601,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="15" pageNumber="16" unit="ft" value="3000.0">
<quantity id="6E009B3B4C27B347FE10F988F50CD311" box="[492,601,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="15" pageNumber="16" unit="ft" value="3000.0">3000 feet</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C27B347FD9FF989F3B9D311" ID-GBIF-Occurrence="1915497811" box="[611,1260,1628,1653]" collectionCode="USNM" country="Puerto Rico" location="Distribution" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C27B347FD9FF989F5CED311" box="[611,667,1629,1653]" pageId="15" pageNumber="16">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C27B347FCDBF988F42DD310" box="[807,888,1628,1652]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="16">USNM</collectionCode>
HT 
<specimenCount id="BFFEFD574C27B347FC57F989F496D310" box="[939,963,1629,1652]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
. 
<location id="AC2760054C27B347FC32F988F30FD311" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C27B347FC32F988F30FD311" box="[974,1114,1628,1653]" country="Puerto Rico" name="Distribution" pageId="15" pageNumber="16">Distribution</location>
: 
<collectingCountry id="D1EF764E4C27B347FB98F989F3B9D311" box="[1124,1260,1628,1653]" name="Puerto Rico" pageId="15" pageNumber="16">Puerto Rico</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BF94AF57CD3D3" blockId="15.[151,1437,1694,1863]" box="[151,553,1694,1719]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BF94AF7F6D3D3" bold="true" box="[151,163,1694,1719]" italics="true" pageId="15" pageNumber="16">*</emphasis>
<taxonomicName id="6EF84D5D4C27B347FF5EF94AF57CD3D3" ID-CoL="LC75" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[162,553,1694,1719]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="9B8CEACC4C27B347FF5EF94AF6F4D3D3" bold="true" box="[162,417,1694,1719]" italics="true" pageId="15" pageNumber="16">Belonuchus decipiens</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BF916F61AD223" blockId="15.[151,1437,1694,1863]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BF916F61AD223" infered="true" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BF916F523D3BF" ID-CoL="LC75" authority="Sharp 1876 a: 159" authorityName="Sharp" authorityPageNumber="159" authorityYear="1876" box="[151,630,1730,1755]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="9B8CEACC4C27B347FF6BF916F6C4D3BF" box="[151,401,1730,1755]" italics="true" pageId="15" pageNumber="16">Belonuchus decipiens</emphasis>
<treatmentCitation id="285910CF4C27B347FE63F916F523D3BF" author="" box="[415,630,1730,1755]" page="159" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FE63F916F562D3BF" box="[415,567,1730,1755]" pageId="15" pageNumber="16" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 159
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FD74F917F5EAD3BF" box="[648,703,1731,1755]" pageId="15" pageNumber="16">Type</typeStatus>
locality: Ega. 
<typeStatus id="7643887C4C27B347FC84F917F4FAD3BF" box="[888,943,1731,1755]" pageId="15" pageNumber="16">Type</typeStatus>
depository: BMNH 6 ST, FMNH 1 ST. Distribution: 
<collectingCountry id="D1EF764E4C27B347FEA2F932F6F0D39B" box="[350,421,1766,1791]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C27B347FE4EF932F571D39B" box="[434,548,1766,1791]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C27B347FDCDF933F5C8D39B" box="[561,669,1767,1791]" country="Paraguay" name="Asuncion" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FDCDF933F5C8D39B" box="[561,669,1767,1791]" name="Paraguay" pageId="15" pageNumber="16">Paraguay</collectingCountry>
</collectingRegion>
; 
<collectingCountry id="D1EF764E4C27B347FD56F932F462D39B" box="[682,823,1766,1791]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
?; 
<collectingCountry id="D1EF764E4C27B347FCB2F932F4EED39B" box="[846,955,1766,1791]" name="Indonesia" pageId="15" pageNumber="16">Indonesia</collectingCountry>
(Lombok)? Notes: redescription (
<bibRefCitation id="CD694B2F4C27B347FAC1F932F656D247" pageId="15" pageNumber="16" refId="ref72647" refString="Silvestri, F. (1945) Descrizione e biologia del coleottero stafilinide Belonuchus formosus Grav. introdotto in Italia per la lotta contro Ditteri Tripaneidi. Bollettino del R. Laboratorio di Entomologia Agraria di Portici, 5, 312 - 328." type="journal article">Silvestri 1945</bibRefCitation>
); distribution, including 
<collectingCountry id="D1EF764E4C27B347FDE4F8DEF5D2D247" box="[536,647,1802,1827]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
and 
<collectingRegion id="6B3CF83C4C27B347FD42F8DFF473D247" box="[702,806,1803,1827]" country="Paraguay" name="Asuncion" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FD42F8DFF473D247" box="[702,806,1803,1827]" name="Paraguay" pageId="15" pageNumber="16">Paraguay</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C27B347FCC9F8DEF341D247" box="[821,1044,1802,1827]" pageId="15" pageNumber="16" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b</bibRefCitation>
); records from 
<collectingCountry id="D1EF764E4C27B347FB42F8DEF257D247" box="[1214,1282,1802,1827]" name="Brazil" pageId="15" pageNumber="16">Brazil</collectingCountry>
only (Asenjo 
<emphasis id="9B8CEACC4C27B347FF3BF8FBF7AAD223" box="[199,255,1838,1863]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BF8A5F510D2EE" blockId="15.[151,1437,1905,2038]" box="[151,581,1905,1930]" pageId="15" pageNumber="16">
<emphasis id="9B8CEACC4C27B347FF6BF8A5F7F6D2EE" bold="true" box="[151,163,1905,1930]" italics="true" pageId="15" pageNumber="16">*</emphasis>
<taxonomicName id="6EF84D5D4C27B347FF5EF8A5F510D2EE" ID-CoL="LC77" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[162,581,1905,1930]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dichrous">
<emphasis id="9B8CEACC4C27B347FF5EF8A5F6CED2EE" bold="true" box="[162,411,1905,1930]" italics="true" pageId="15" pageNumber="16">Belonuchus dichrous</emphasis>
<bibRefCitation id="CD694B2F4C27B347FE5EF8A5F510D2EE" box="[418,581,1905,1930]" pageId="15" pageNumber="16" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C27B347FF6BF841F3CBD292" blockId="15.[151,1437,1905,2038]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="89E811F04C27B347FF6BF841F3CBD292" infered="true" pageId="15" pageNumber="16">
<taxonomicName id="6EF84D5D4C27B347FF6BF841F53DD2CA" ID-CoL="LC77" authority="Erichson 1840: 420" authorityName="Erichson" authorityPageNumber="420" authorityYear="1840" box="[151,616,1941,1966]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dichrous">
<emphasis id="9B8CEACC4C27B347FF6BF841F6D7D2CA" box="[151,386,1941,1966]" italics="true" pageId="15" pageNumber="16">Belonuchus dichrous</emphasis>
<treatmentCitation id="285910CF4C27B347FE75F841F53DD2CA" author="" box="[393,616,1941,1966]" page="420" pageId="15" pageNumber="16">
<bibRefCitation id="CD694B2F4C27B347FE75F841F57AD2CA" box="[393,559,1941,1966]" pageId="15" pageNumber="16" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 420
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C27B347FD8EF842F5FCD2CA" box="[626,681,1942,1966]" pageId="15" pageNumber="16">Type</typeStatus>
locality: Monte Orizaba in 
<collectingRegion id="6B3CF83C4C27B347FC1CF841F36CD2CA" box="[992,1081,1941,1966]" country="Mexico" name="Mexico" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FC1CF841F36CD2CA" box="[992,1081,1941,1966]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
</collectingRegion>
. 
<typeStatus id="7643887C4C27B347FBB8F842F32ED2CA" box="[1092,1147,1942,1966]" pageId="15" pageNumber="16">Type</typeStatus>
depository: ZMHB 4 ST: 
<quantity id="6E009B3B4C27B347FF3BF86EF7A7D2B6" box="[199,242,1977,2002]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="16" unit="in" value="2.0">2 in</quantity>
Bernhauer´s hand &quot; 
<taxonomicName id="6EF84D5D4C27B347FE32F86DF563D2B6" ID-CoL="LC77" authorityName="Erichson" authorityYear="1840" box="[462,566,1977,2002]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dichrous">dichrous</taxonomicName>
&quot;, 2 with &quot;Orizaba&quot; [MCP seen]. Distribution: 
<collectingRegion id="6B3CF83C4C27B347FBCBF86DF3C5D2B6" box="[1079,1168,1977,2002]" country="Mexico" name="Mexico" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FBCBF86DF3C5D2B6" box="[1079,1168,1977,2002]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C27B347FB67F86DF242D2B6" box="[1179,1303,1977,2002]" country="Guatemala" name="Guatemala" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FB67F86DF242D2B6" box="[1179,1303,1977,2002]" name="Guatemala" pageId="15" pageNumber="16">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C27B347FADEF86DF2CCD2B6" box="[1314,1433,1977,2002]" name="Venezuela" pageId="15" pageNumber="16">Venezuela</collectingCountry>
. Notes: distribution, also within 
<collectingRegion id="6B3CF83C4C27B347FDD5F809F52BD292" box="[553,638,2013,2038]" country="Mexico" name="Mexico" pageId="15" pageNumber="16">
<collectingCountry id="D1EF764E4C27B347FDD5F809F52BD292" box="[553,638,2013,2038]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C27B347FD71F809F4B5D292" box="[653,992,2013,2038]" pageId="15" pageNumber="16" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C27B347FC9AF80AF4CBD292" box="[870,926,2013,2038]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
2002
</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C27B347FC17F80AF3C7D292" box="[1003,1170,2013,2038]" pageId="15" pageNumber="16" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFF43F5D1D5D4" blockId="16.[151,1436,151,248]" box="[151,644,151,176]" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BFF43F7F6D5D4" bold="true" box="[151,163,151,176]" italics="true" pageId="16" pageNumber="17">*</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF5EFF43F5D1D5D4" ID-CoL="LC78" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[162,644,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diversicornis">
<emphasis id="9B8CEACC4C38B358FF5EFF43F69FD5D4" bold="true" box="[162,458,151,176]" italics="true" pageId="16" pageNumber="17">Belonuchus diversicornis</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFF68F3BDD59C" blockId="16.[151,1436,151,248]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFF68F3BDD59C" pageId="16" pageNumber="17">
<materialsCitation id="19903C834C38B358FF6BFF68F329D5B1" ID-GBIF-Occurrence="1915497848" box="[151,1148,188,213]" country="Brazil" location="Espirito Santo" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C38B358FF6BFF68F5E2D5B0" ID-CoL="LC78" authority="Bernhauer 1917 c: 104" authorityName="Bernhauer" authorityPageNumber="104" authorityYear="1917" box="[151,695,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diversicornis">
<emphasis id="9B8CEACC4C38B358FF6BFF68F6E6D5B1" box="[151,435,188,213]" italics="true" pageId="16" pageNumber="17">Belonuchus diversicornis</emphasis>
<treatmentCitation id="285910CF4C38B358FE47FF68F5E2D5B0" author="" box="[443,695,188,213]" page="104" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE47FF68F52AD5B0" box="[443,639,188,213]" pageId="16" pageNumber="17" refId="ref62261" refString="Bernhauer, M. (1917 c) Neue sudamerikanische Staphyliniden (18. Beitrag). Wiener Entomologische Zeitung, 36 (3 - 5), 102 - 116. https: // doi. org / 10.5962 / bhl. part. 12925" type="journal article">Bernhauer 1917c</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C38B358FD3FFF69F5AED5B1" box="[707,763,189,213]" pageId="16" pageNumber="17">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C38B358FC99FF68F352D5B0" box="[869,1031,188,213]" country="Brazil" name="Espirito Santo" pageId="16" pageNumber="17">Espirito Santo</collectingRegion>
, 
<collectingCountry id="D1EF764E4C38B358FBEFFF68F329D5B1" box="[1043,1148,188,213]" name="Brazil" pageId="16" pageNumber="17">Brasilien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C38B358FB7BFF69F5DAD59C" ID-GBIF-Occurrence="1915497828" collectionCode="FMNH" country="Brazil" location="Distribution" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<typeStatus id="7643887C4C38B358FB7BFF69F3EBD5B1" box="[1159,1214,189,213]" pageId="16" pageNumber="17">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C38B358FAB7FF69F2C9D5B0" box="[1355,1436,189,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="17">FMNH</collectionCode>
HT. 
<location id="AC2760054C38B358FF04FF0BF6D1D59C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C38B358FF04FF0BF6D1D59C" box="[248,388,223,248]" country="Brazil" name="Distribution" pageId="16" pageNumber="17" stateProvince="Espirito Santo">Distribution</location>
: 
<collectingCountry id="D1EF764E4C38B358FE72FF0BF687D59C" box="[398,466,223,248]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C38B358FE1DFF0BF5D2D59C" box="[481,647,223,248]" country="Brazil" name="Espirito Santo" pageId="16" pageNumber="17">Espírito Santo</collectingRegion>
)
</materialsCitation>
. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C38B358FB9DFF35F3CCD59C" box="[1121,1177,223,248]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFEF3F55DD424" blockId="16.[151,1437,295,429]" box="[151,520,295,320]" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BFEF3F7F6D424" bold="true" box="[151,163,295,320]" italics="true" pageId="16" pageNumber="17">*</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF5EFEF3F55DD424" ID-CoL="5WGHS" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,520,295,320]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="docilis">
<emphasis id="9B8CEACC4C38B358FF5EFEF3F6D4D424" bold="true" box="[162,385,295,320]" italics="true" pageId="16" pageNumber="17">Belonuchus docilis</emphasis>
<bibRefCitation id="CD694B2F4C38B358FE74FEF3F55DD424" box="[392,520,295,320]" pageId="16" pageNumber="17" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFE98F2C9D4C8" blockId="16.[151,1437,295,429]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFE98F2C9D4C8" infered="true" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BFE98F50DD401" ID-CoL="5WGHS" authority="Sharp 1885: 435" authorityName="Sharp" authorityPageNumber="435" authorityYear="1885" box="[151,600,332,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="docilis">
<emphasis id="9B8CEACC4C38B358FF6BFE98F623D401" box="[151,374,332,357]" italics="true" pageId="16" pageNumber="17">Belonuchus docilis</emphasis>
<treatmentCitation id="285910CF4C38B358FE7BFE98F50DD401" author="" box="[391,600,332,357]" page="435" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE7BFE98F540D401" box="[391,533,332,357]" pageId="16" pageNumber="17" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 435
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C38B358FD91FE99F5F0D401" box="[621,677,333,357]" pageId="16" pageNumber="17">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C38B358FCDFFE98F4CAD401" box="[803,927,332,357]" country="Guatemala" name="Guatemala" pageId="16" pageNumber="17">
<collectingCountry id="D1EF764E4C38B358FCDFFE98F4CAD401" box="[803,927,332,357]" name="Guatemala" pageId="16" pageNumber="17">Guatemala</collectingCountry>
</collectingRegion>
, Pantaleon; 
<collectingRegion id="6B3CF83C4C38B358FBC0FE99F3CDD400" box="[1084,1176,333,356]" country="Panama" name="Panama" pageId="16" pageNumber="17">
<collectingCountry id="D1EF764E4C38B358FBC0FE99F3CDD400" box="[1084,1176,333,356]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
</collectingRegion>
, Bugaba, Tolé. 
<typeStatus id="7643887C4C38B358FA98FE99F2C9D401" box="[1380,1436,333,357]" pageId="16" pageNumber="17">Type</typeStatus>
depository: BMNH 8 ST: 2 Pantaleón, 3 Bugaba, 3 Tolé [MCP seen]. FMNH 2 ST: 2 Bugaba. Distribution: 
<collectingRegion id="6B3CF83C4C38B358FF3BFE40F616D4C9" box="[199,323,404,429]" country="Guatemala" name="Guatemala" pageId="16" pageNumber="17">
<collectingCountry id="D1EF764E4C38B358FF3BFE40F616D4C9" box="[199,323,404,429]" name="Guatemala" pageId="16" pageNumber="17">Guatemala</collectingCountry>
</collectingRegion>
,? 
<collectingCountry id="D1EF764E4C38B358FE9CFE40F688D4C9" box="[352,477,404,429]" name="Costa Rica" pageId="16" pageNumber="17">
<collectingRegion id="6B3CF83C4C38B358FE9CFE40F6F5D4C8" box="[352,416,404,428]" country="Tanzania" name="Pwani" pageId="16" pageNumber="17">Costa</collectingRegion>
Rica
</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C38B358FE1BFE41F516D4C8" box="[487,579,405,428]" country="Panama" name="Panama" pageId="16" pageNumber="17">
<collectingCountry id="D1EF764E4C38B358FE1BFE41F516D4C8" box="[487,579,405,428]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C38B358FDB2FE40F5C3D4C9" box="[590,662,404,429]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
. Note: distribution within (and including) 
<collectingCountry id="D1EF764E4C38B358FB90FE40F3E5D4C9" box="[1132,1200,404,429]" name="Brazil" pageId="16" pageNumber="17">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C38B358FAE9FE41F218D4C9" box="[1301,1357,404,429]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFE0FF521D490" blockId="16.[151,1437,475,608]" box="[151,628,475,501]" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BFE0FF521D490" ID-CoL="LC7C" authority="Blackwelder 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,628,475,501]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dominicus">
<emphasis id="9B8CEACC4C38B358FF6BFE0FF6F5D490" bold="true" box="[151,416,475,500]" italics="true" pageId="16" pageNumber="17">Belonuchus dominicus</emphasis>
<bibRefCitation id="CD694B2F4C38B358FE5BFE08F521D490" box="[423,628,476,501]" pageId="16" pageNumber="17" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFE2BF3D9D704" blockId="16.[151,1437,475,608]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFE2BF3D9D704" infered="true" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BFE2BF5FCD77C" ID-CoL="LC7C" authority="Blackwelder 1943: 431" authorityName="Blackwelder" authorityPageNumber="431" authorityYear="1943" box="[151,681,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dominicus">
<emphasis id="9B8CEACC4C38B358FF6BFE2BF6C0D77C" box="[151,405,511,536]" italics="true" pageId="16" pageNumber="17">Belonuchus dominicus</emphasis>
<treatmentCitation id="285910CF4C38B358FE61FE2BF5FCD77C" author="" box="[413,681,511,536]" page="431" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE61FE2BF53AD77C" box="[413,623,511,536]" pageId="16" pageNumber="17" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 431
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C38B358FD49FDD4F5B9D77C" box="[693,748,512,536]" pageId="16" pageNumber="17">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C38B358FCA5FE2BF499D77C" box="[857,972,511,536]" name="Dominica" pageId="16" pageNumber="17">Dominica</collectingCountry>
: Hillsboro Estate, at mouth of the Layou River, Leeward side. 
<typeStatus id="7643887C4C38B358FE3CFDF1F6ADD759" box="[448,504,549,573]" pageId="16" pageNumber="17">Type</typeStatus>
depository: USNM HT 
<specimenCount id="BFFEFD574C38B358FCEBFDF1F47BD758" box="[791,814,549,572]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
, 4 PT; Blackwelder 1 PT; FMNH 1 PT. Distribution: 
<collectingCountry id="D1EF764E4C38B358FF3BFD93F660D704" box="[199,309,583,608]" name="Dominica" pageId="16" pageNumber="17">Dominica</collectingCountry>
Note: distribution within 
<collectingCountry id="D1EF764E4C38B358FDA4FD93F593D704" box="[600,710,583,608]" name="Dominica" pageId="16" pageNumber="17">Dominica</collectingCountry>
(Peck 2006); generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFD5BF521D7CC" blockId="16.[151,1437,655,824]" box="[151,628,655,680]" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BFD5BF521D7CC" ID-CoL="LC7D" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,628,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="duplicatus">
<emphasis id="9B8CEACC4C38B358FF6BFD5BF6F5D7CC" bold="true" box="[151,416,655,680]" italics="true" pageId="16" pageNumber="17">Belonuchus duplicatus</emphasis>
<bibRefCitation id="CD694B2F4C38B358FE5BFD5BF521D7CC" box="[423,628,655,680]" pageId="16" pageNumber="17" refId="ref63204" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum, 185, 1 - 188." type="journal article">Blackwelder 1944</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFD60F27CD7A8" blockId="16.[151,1437,655,824]" box="[151,1321,692,717]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFD60F27CD7A8" box="[151,1321,692,717]" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BFD60F5F4D7A8" ID-CoL="LC7D" authority="Blackwelder 1944: 137" authorityName="Blackwelder" authorityPageNumber="137" authorityYear="1944" box="[151,673,692,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="duplicatus">
<emphasis id="9B8CEACC4C38B358FF6BFD60F6C0D7A9" box="[151,405,692,717]" italics="true" pageId="16" pageNumber="17">Belonuchus duplicatus</emphasis>
<treatmentCitation id="285910CF4C38B358FE60FD60F5F4D7A8" author="" box="[412,673,692,717]" page="137" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE60FD60F538D7A8" box="[412,621,692,717]" pageId="16" pageNumber="17" refId="ref63204" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum, 185, 1 - 188." type="journal article">Blackwelder 1944</bibRefCitation>
: 137
</treatmentCitation>
</taxonomicName>
(new name for 
<taxonomicName id="6EF84D5D4C38B358FCAFFD60F248D7A8" ID-CoL="LC68" authority="Bernhauer 1934: 313" authorityName="Bernhauer" authorityPageNumber="313" authorityYear="1934" box="[851,1309,692,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bruchi">
<emphasis id="9B8CEACC4C38B358FCAFFD60F370D7A9" box="[851,1061,692,717]" italics="true" pageId="16" pageNumber="17">Belonuchus bruchi</emphasis>
Bernhauer 1934: 313
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFD0CF5E4D65C" blockId="16.[151,1437,655,824]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFD0CF5E4D65C" pageId="16" pageNumber="17">
= 
<taxonomicName id="6EF84D5D4C38B358FF49FD03F5FFD794" ID-CoL="LC68" authority="Bernhauer 1934 b: 213" authorityName="Bernhauer" authorityPageNumber="213" authorityYear="1934" box="[181,682,727,752]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bruchi">
<emphasis id="9B8CEACC4C38B358FF49FD03F6C5D794" box="[181,400,727,752]" italics="true" pageId="16" pageNumber="17">Belonuchus bruchi</emphasis>
<treatmentCitation id="285910CF4C38B358FE63FD03F5FFD794" author="" box="[415,682,727,752]" page="213" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE63FD03F538D794" box="[415,621,727,752]" pageId="16" pageNumber="17" refId="ref62723" refString="Bernhauer, M. (1934 b) Neue Staphyliniden aus Argentinien (IV) (Col.) (34. Beitrag zur Kenntnis der sudamerikanischen Fauna). Revista de Entomologia, 4 (2), 212 - 221." type="journal article">Bernhauer 1934b</bibRefCitation>
: 213
</treatmentCitation>
</taxonomicName>
(preoccupied, junior primary homonym of 
<taxonomicName id="6EF84D5D4C38B358FB3DFD03F69CD671" ID-CoL="LC68" authority="Bernhauer 1912: 175" authorityName="Bernhauer" authorityPageNumber="175" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bruchi">
<emphasis id="9B8CEACC4C38B358FB3DFD03F2C9D794" box="[1217,1436,727,752]" italics="true" pageId="16" pageNumber="17">Belonuchus bruchi</emphasis>
Bernhauer 1912: 175
</taxonomicName>
). Type locality: Tucuman, Argentina. Type depository: FMNH HT. Distribution: Argentina. Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFCB3F52FD6E4" blockId="16.[151,1436,871,968]" box="[151,634,871,896]" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BFCB3F52FD6E4" ID-CoL="68GHS" authority="Bernhauer 1935" authorityName="Bernhauer" authorityYear="1935" box="[151,634,871,896]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="9B8CEACC4C38B358FF6BFCB3F6EAD6E4" bold="true" box="[151,447,871,896]" italics="true" pageId="16" pageNumber="17">Belonuchus ecuadorensis</emphasis>
<bibRefCitation id="CD694B2F4C38B358FE3AFCB3F52FD6E4" box="[454,634,871,896]" pageId="16" pageNumber="17" refId="ref62766" refString="Bernhauer, M. (1935) Neue Staphyliniden (Col.) aus Sudamerika. Stylops, a Journal of Taxonomic Entomology, 4, 90 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00565. x" type="journal article">Bernhauer 1935</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFC58F5F7D6AC" blockId="16.[151,1436,871,968]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFC58F5F7D6AC" pageId="16" pageNumber="17">
<materialsCitation id="19903C834C38B358FF6BFC58F33AD6C1" ID-GBIF-Occurrence="1915497835" box="[151,1135,908,933]" country="Ecuador" location="Ecuador" pageId="16" pageNumber="17" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C38B358FF6BFC58F40DD6C0" ID-CoL="68GHS" authority="Bernhauer 1935: 91" authorityName="Bernhauer" authorityPageNumber="91" authorityYear="1935" box="[151,856,908,933]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ecuadorensis" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C38B358FF6BFC58F530D6C1" box="[151,613,908,933]" italics="true" pageId="16" pageNumber="17">Belonuchus (Musicoderus) ecuadorensis</emphasis>
<treatmentCitation id="285910CF4C38B358FD93FC58F40DD6C0" author="" box="[623,856,908,933]" page="91" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FD93FC58F47ED6C1" box="[623,811,908,933]" pageId="16" pageNumber="17" refId="ref62766" refString="Bernhauer, M. (1935) Neue Staphyliniden (Col.) aus Sudamerika. Stylops, a Journal of Taxonomic Entomology, 4, 90 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00565. x" type="journal article">Bernhauer 1935</bibRefCitation>
: 91
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C38B358FC9BFC59F4CAD6C1" box="[871,927,909,933]" pageId="16" pageNumber="17">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C38B358FBECFC58F33AD6C1" box="[1040,1135,908,933]" name="Ecuador" pageId="16" pageNumber="17">Ecuador</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C38B358FB81FC59F5CAD6AC" ID-GBIF-Occurrence="1915497838" collectionCode="BMNH" country="Ecuador" location="ST. Distribution" pageId="16" pageNumber="17" specimenCode="FMNH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C38B358FB81FC59F3E1D6C1" box="[1149,1204,909,933]" pageId="16" pageNumber="17">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C38B358FAB4FC59F2C9D6C0" box="[1352,1436,909,932]" pageId="16" pageNumber="17">BMNH</collectionCode>
#?ST; 
<specimenCode id="F95E9EA54C38B358FEF3FC64F620D6AC" box="[271,373,944,968]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="16" pageNumber="17">FMNH 1</specimenCode>
<location id="AC2760054C38B358FE80FC64F560D6AC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C38B358FE80FC64F560D6AC" box="[380,565,943,968]" country="Ecuador" name="ST. Distribution" pageId="16" pageNumber="17">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C38B358FDC3FC7BF5CAD6AC" box="[575,671,943,968]" name="Ecuador" pageId="16" pageNumber="17">Ecuador</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFC23F56AD174" blockId="16.[151,1436,1015,1581]" box="[151,575,1015,1040]" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BFC23F7F6D174" bold="true" box="[151,163,1015,1040]" italics="true" pageId="16" pageNumber="17">*</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF5EFC23F56AD174" ID-CoL="LC7F" authority="(Say 1830)" authorityName="Say" authorityYear="1830" baseAuthorityName="Say" baseAuthorityYear="1830" box="[162,575,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="9B8CEACC4C38B358FF5EFC23F6E9D174" bold="true" box="[162,444,1015,1040]" italics="true" pageId="16" pageNumber="17">Belonuchus ephippiatus</emphasis>
(
<bibRefCitation id="CD694B2F4C38B358FE37FC2CF563D174" box="[459,566,1016,1040]" pageId="16" pageNumber="17" refId="ref71623" refString="Say, T. (1830) Descriptions of new species of North American insects, and observations on some already described. New Harmony, Indiana, pp. 1 - 41." type="book chapter">Say 1830</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFBC8F469D1C4" blockId="16.[151,1436,1015,1581]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFBC8F469D1C4" infered="true" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BFBC8F507D151" ID-CoL="4ZJY4" authority="Say 1830: 35" authorityName="Say" authorityPageNumber="35" authorityYear="1830" box="[151,594,1052,1077]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="9B8CEACC4C38B358FF6BFBC8F6FDD151" box="[151,424,1052,1077]" italics="true" pageId="16" pageNumber="17">Staphylinus ephippiatus</emphasis>
<treatmentCitation id="285910CF4C38B358FE4FFBC8F507D151" author="" box="[435,594,1052,1077]" page="35" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE4FFBC8F570D150" box="[435,549,1052,1076]" pageId="16" pageNumber="17" refId="ref71623" refString="Say, T. (1830) Descriptions of new species of North American insects, and observations on some already described. New Harmony, Indiana, pp. 1 - 41." type="book chapter">Say 1830</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
, also 1834: 448. 
<typeStatus id="7643887C4C38B358FCD9FBC9F408D151" box="[805,861,1053,1077]" pageId="16" pageNumber="17">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C38B358FC2CFBC8F37CD151" box="[976,1065,1052,1077]" country="Mexico" name="Mexico" pageId="16" pageNumber="17">
<collectingCountry id="D1EF764E4C38B358FC2CFBC8F37CD151" box="[976,1065,1052,1077]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
</collectingRegion>
. 
<typeStatus id="7643887C4C38B358FBC5FBC9F324D151" box="[1081,1137,1053,1077]" pageId="16" pageNumber="17">Type</typeStatus>
depository: MCZ NT 
<specimenCount id="BFFEFD574C38B358FA7CFBC9F2CDD150" box="[1408,1432,1053,1076]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
. Distribution: 
<collectingCountry id="D1EF764E4C38B358FE9DFB94F6C9D13C" box="[353,412,1088,1112]" name="United States of America" pageId="16" pageNumber="17">USA</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C38B358FE57FBEBF556D13C" box="[427,515,1087,1112]" country="Mexico" name="Mexico" pageId="16" pageNumber="17">
<collectingCountry id="D1EF764E4C38B358FE57FBEBF556D13C" box="[427,515,1087,1112]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C38B358FDEFFBEBF5D1D13C" box="[531,644,1087,1112]" name="Colombia" pageId="16" pageNumber="17">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C38B358FD68FBEBF45ED13C" box="[660,779,1087,1112]" name="Venezuela" pageId="16" pageNumber="17">Venezuela</collectingCountry>
; adventive in 
<collectingRegion id="6B3CF83C4C38B358FC4FFBEBF352D13C" box="[947,1031,1087,1112]" country="United States of America" name="Hawaii" pageId="16" pageNumber="17">Hawaii</collectingRegion>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C38B358FAE6FBEBF690D118" ID-CoL="8VS43" authority="(Erichson 1840: 927)" authorityName="Erichson" authorityPageNumber="927" authorityYear="1840" baseAuthorityName="Erichson" baseAuthorityYear="1840" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C38B358FAE6FBEBF2C9D13C" box="[1306,1436,1087,1112]" italics="true" pageId="16" pageNumber="17">Staphylinus</emphasis>
(
<treatmentCitation id="285910CF4C38B358FF32FBB0F6E8D118" author="" box="[206,445,1124,1149]" page="927" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FF32FBB0F6E8D118" box="[206,445,1124,1149]" pageId="16" pageNumber="17" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840: 927</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; 
<typeStatus id="7643887C4C38B358FE24FBB1F564D119" box="[472,561,1125,1149]" pageId="16" pageNumber="17" type="neotype">neotype</typeStatus>
designation, redescription (
<treatmentCitation id="285910CF4C38B358FC80FBB0F33CD119" author="" box="[892,1129,1124,1149]" page="746" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FC80FBB0F33CD119" box="[892,1129,1124,1149]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 746</bibRefCitation>
</treatmentCitation>
); distribution, also within 
<collectingRegion id="6B3CF83C4C38B358FF3BFB53F649D1C4" box="[199,284,1159,1184]" country="Mexico" name="Mexico" pageId="16" pageNumber="17">
<collectingCountry id="D1EF764E4C38B358FF3BFB53F649D1C4" box="[199,284,1159,1184]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C38B358FED7FB53F52BD1C4" box="[299,638,1159,1184]" pageId="16" pageNumber="17" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C38B358FDF8FB5DF569D1C4" box="[516,572,1159,1184]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2002
</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C38B358FD75FB5CF464D1C4" box="[649,817,1159,1184]" pageId="16" pageNumber="17" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFB79F657D068" blockId="16.[151,1436,1015,1581]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFB79F657D068" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BFB79F7FFD1A0" box="[151,170,1197,1220]" italics="true" pageId="16" pageNumber="17">=</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF4FFB78F50AD1A0" ID-CoL="LCBN" authority="Sharp 1885: 432" authorityName="Sharp" authorityPageNumber="432" authorityYear="1885" box="[179,607,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="9B8CEACC4C38B358FF4FFB78F6C1D1A1" box="[179,404,1196,1221]" italics="true" pageId="16" pageNumber="17">Belonuchus simplex</emphasis>
<treatmentCitation id="285910CF4C38B358FE61FB78F50AD1A0" author="" box="[413,607,1196,1221]" page="432" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE61FB78F571D1A1" box="[413,548,1196,1221]" pageId="16" pageNumber="17" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 432
</treatmentCitation>
</taxonomicName>
. Type locality: Mexico: Oaxaca. Type depository: BMNH LT ♂, 2 PLT; FMNH 1 PLT?. Notes: as synonym of 
<taxonomicName id="6EF84D5D4C38B358FD8BFB05F4BED18C" ID-CoL="4ZJY4" authority="(Fauvel 1901: 86)" authorityName="Fauvel" authorityPageNumber="86" authorityYear="1901" baseAuthorityName="Fauvel" baseAuthorityYear="1901" box="[631,1003,1231,1256]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="9B8CEACC4C38B358FD8BFB05F44CD18C" box="[631,793,1231,1256]" italics="true" pageId="16" pageNumber="17">B. ephippiatus</emphasis>
(
<treatmentCitation id="285910CF4C38B358FCDBFB1BF4B4D18C" author="" box="[807,993,1231,1256]" page="86" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FCDBFB1BF4B4D18C" box="[807,993,1231,1256]" pageId="16" pageNumber="17" refId="ref66160" refString="Fauvel, A. (1901) Voyage de M. le Dr. Ed. Bugnion au Venezuela, en Colombie et aux Antilles. Revue d'Entomologie, 20, 69 - 91, pls. 1 - 2." type="journal article">Fauvel 1901: 86</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; lectotype designation (
<treatmentCitation id="285910CF4C38B358FB0FFB04F7A3D068" author="" page="748" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FB0FFB04F7A3D068" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 748</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFACDF23CD030" blockId="16.[151,1436,1015,1581]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFACDF23CD030" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BFACDF7FFD054" box="[151,170,1305,1328]" italics="true" pageId="16" pageNumber="17">=</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF4EFAC3F5CFD054" ID-CoL="5WGJ8" authority="Casey 1915: 446" authorityName="Casey" authorityPageNumber="446" authorityYear="1915" box="[178,666,1303,1328]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="9B8CEACC4C38B358FF4EFAC3F69AD054" box="[178,463,1303,1328]" italics="true" pageId="16" pageNumber="17">Belonuchus punctiventris</emphasis>
<treatmentCitation id="285910CF4C38B358FE2BFACCF5CFD054" author="" box="[471,666,1303,1328]" page="446" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE2BFACCF535D054" box="[471,608,1303,1328]" pageId="16" pageNumber="17" refId="ref63944" refString="Casey, T. L. (1915) Studies in some staphylinid genera of North America. In: Memoirs on the Coleoptera. Vol. 6. New Era Printing Co., Lancaster, Pennsylvania, pp. 395 - 450." type="book chapter">Casey 1915</bibRefCitation>
: 446
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C38B358FD59FACCF589D054" box="[677,732,1304,1328]" pageId="16" pageNumber="17">Type</typeStatus>
locality: Benson, 
<collectingRegion id="6B3CF83C4C38B358FC50FAC3F35FD054" box="[940,1034,1303,1328]" country="United States of America" name="Arizona" pageId="16" pageNumber="17">Arizona</collectingRegion>
. 
<typeStatus id="7643887C4C38B358FBE9FACCF318D054" box="[1045,1101,1304,1328]" pageId="16" pageNumber="17">Type</typeStatus>
depository: USNM LT 
<specimenCount id="BFFEFD574C38B358FAA0FACCF226D04B" box="[1372,1395,1304,1327]" pageId="16" pageNumber="17" type="male">♂</specimenCount>
, 13 PLT. Notes: synonym of 
<taxonomicName id="6EF84D5D4C38B358FE22FAE9F420D031" ID-CoL="4ZJY4" authority="(Smetana 1995: 746)" authorityName="Smetana" authorityPageNumber="746" authorityYear="1995" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[478,885,1340,1365]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="9B8CEACC4C38B358FE22FAE9F5D5D031" box="[478,640,1340,1365]" italics="true" pageId="16" pageNumber="17">B. ephippiatus</emphasis>
(
<treatmentCitation id="285910CF4C38B358FD72FAE8F439D031" author="" box="[654,876,1340,1365]" page="746" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FD72FAE8F439D031" box="[654,876,1340,1365]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 746</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; 
<typeStatus id="7643887C4C38B358FC7CFAE8F4B2D031" box="[896,999,1340,1365]" pageId="16" pageNumber="17" type="lectotype">lectotype</typeStatus>
designation (
<treatmentCitation id="285910CF4C38B358FB82FAE8F208D030" author="" box="[1150,1373,1340,1365]" page="748" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FB82FAE8F208D030" box="[1150,1373,1340,1365]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 748</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFAB5F278D0F8" blockId="16.[151,1436,1015,1581]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFAB5F278D0F8" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BFAB5F7FFD01C" box="[151,170,1377,1400]" italics="true" pageId="16" pageNumber="17">=</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF48FA8BF539D01C" ID-CoL="LC8T" authority="Casey 1915: 446" authorityName="Casey" authorityPageNumber="446" authorityYear="1915" box="[180,620,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="laticeps">
<emphasis id="9B8CEACC4C38B358FF48FA8BF6CCD01C" box="[180,409,1375,1400]" italics="true" pageId="16" pageNumber="17">Belonuchus laticeps</emphasis>
<treatmentCitation id="285910CF4C38B358FE58FAB4F539D01C" author="" box="[420,620,1375,1400]" page="446" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE58FAB4F57AD01C" box="[420,559,1375,1400]" pageId="16" pageNumber="17" refId="ref63944" refString="Casey, T. L. (1915) Studies in some staphylinid genera of North America. In: Memoirs on the Coleoptera. Vol. 6. New Era Printing Co., Lancaster, Pennsylvania, pp. 395 - 450." type="book chapter">Casey 1915</bibRefCitation>
: 446
</treatmentCitation>
</taxonomicName>
. Type locality: near Indio, California. Type depository: USNM LT ♂. Notes: synonym of 
<taxonomicName id="6EF84D5D4C38B358FE5FFA51F46FD0F9" ID-CoL="4ZJY4" authority="(Smetana 1995: 746)" authorityName="Smetana" authorityPageNumber="746" authorityYear="1995" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[419,826,1412,1437]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="9B8CEACC4C38B358FE5FFA51F510D0F9" box="[419,581,1412,1437]" italics="true" pageId="16" pageNumber="17">B. ephippiatus</emphasis>
(
<treatmentCitation id="285910CF4C38B358FDAFFA50F464D0F9" author="" box="[595,817,1412,1437]" page="746" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FDAFFA50F464D0F9" box="[595,817,1412,1437]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 746</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; lectotype designation (
<treatmentCitation id="285910CF4C38B358FBBFFA50F274D0F8" author="" box="[1091,1313,1412,1437]" page="749" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FBBFFA50F274D0F8" box="[1091,1313,1412,1437]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 749</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFA7DF278D080" blockId="16.[151,1436,1015,1581]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFA7DF278D080" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BFA7DF7FFD0A4" box="[151,170,1449,1472]" italics="true" pageId="16" pageNumber="17">=</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF4EFA73F5D7D0A4" ID-CoL="LC8M" authority="Casey 1915: 447" authorityName="Casey" authorityPageNumber="447" authorityYear="1915" box="[178,642,1447,1472]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="jacobianus">
<emphasis id="9B8CEACC4C38B358FF4EFA73F6EDD0A4" box="[178,440,1447,1472]" italics="true" pageId="16" pageNumber="17">Belonuchus jacobianus</emphasis>
<treatmentCitation id="285910CF4C38B358FE3CFA7CF5D7D0A4" author="" box="[448,642,1447,1472]" page="447" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE3CFA7CF51CD0A4" box="[448,585,1447,1472]" pageId="16" pageNumber="17" refId="ref63944" refString="Casey, T. L. (1915) Studies in some staphylinid genera of North America. In: Memoirs on the Coleoptera. Vol. 6. New Era Printing Co., Lancaster, Pennsylvania, pp. 395 - 450." type="book chapter">Casey 1915</bibRefCitation>
: 447
</treatmentCitation>
</taxonomicName>
. Type locality: San Diego, California. Type depository: USNM LT ♂. Notes: synonym of 
<taxonomicName id="6EF84D5D4C38B358FE5FFA19F46FD081" ID-CoL="4ZJY4" authority="(Smetana 1995: 746)" authorityName="Smetana" authorityPageNumber="746" authorityYear="1995" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[419,826,1484,1509]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="9B8CEACC4C38B358FE5FFA19F510D081" box="[419,581,1484,1509]" italics="true" pageId="16" pageNumber="17">B. ephippiatus</emphasis>
(
<treatmentCitation id="285910CF4C38B358FDAFFA18F464D081" author="" box="[595,817,1484,1509]" page="746" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FDAFFA18F464D081" box="[595,817,1484,1509]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 746</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; lectotype designation (
<treatmentCitation id="285910CF4C38B358FBBFFA18F274D080" author="" box="[1091,1313,1484,1509]" page="749" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FBBFFA18F274D080" box="[1091,1313,1484,1509]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 749</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BFA25F484D349" blockId="16.[151,1436,1015,1581]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BFA25F484D349" pageId="16" pageNumber="17">
<materialsCitation id="19903C834C38B358FF6BFA25F372D363" ID-GBIF-Occurrence="1915497822" box="[151,1063,1519,1544]" location="Brownsville" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
<emphasis id="9B8CEACC4C38B358FF6BFA25F7FFD36C" box="[151,170,1521,1544]" italics="true" pageId="16" pageNumber="17">=</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF48FA3BF5C5D36C" ID-CoL="LCBD" authority="Cooper 1933: 545" authorityName="Cooper" authorityPageNumber="545" authorityYear="1933" box="[180,656,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="schaefferi">
<emphasis id="9B8CEACC4C38B358FF48FA3BF6E5D36C" box="[180,432,1519,1544]" italics="true" pageId="16" pageNumber="17">Belonuchus schaefferi</emphasis>
<treatmentCitation id="285910CF4C38B358FE47FA24F5C5D36C" author="" box="[443,656,1519,1544]" page="545" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE47FA24F501D36C" box="[443,596,1520,1544]" pageId="16" pageNumber="17" refId="ref65034" refString="Cooper, K. W. (1933) A new species of Belonuchus Nordm. (Col. Staphylinidae). Journal of the New York Entomological Society, 41, 545 - 546." type="journal article">Cooper 1933</bibRefCitation>
: 545
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C38B358FD63FA24F582D36C" box="[671,727,1520,1544]" pageId="16" pageNumber="17">Type</typeStatus>
locality: 
<location id="AC2760054C38B358FCB4FA3BF481D36C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C38B358FCB4FA3BF481D36C" box="[840,980,1519,1544]" name="Brownsville" pageId="16" pageNumber="17" stateProvince="Texas">Brownsville</location>
, 
<collectingRegion id="6B3CF83C4C38B358FC1FFA24F372D363" box="[995,1063,1520,1543]" country="United States of America" name="Texas" pageId="16" pageNumber="17">Texas</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C38B358FBCAFA24F484D349" ID-GBIF-Occurrence="1915497909" collectionCode="CUIC" location="Note" pageId="16" pageNumber="17" specimenCode="AMNH 1" specimenCount="1" specimenCount-female="1" stateProvince="Texas" typeStatus="holotype">
<typeStatus id="7643887C4C38B358FBCAFA24F33BD36C" box="[1078,1134,1520,1544]" pageId="16" pageNumber="17">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C38B358FAFDFA24F211D36C" box="[1281,1348,1520,1544]" country="USA" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="16" pageNumber="17">CUIC</collectionCode>
HT 
<specimenCount id="BFFEFD574C38B358FA83FA24F2C0D363" box="[1407,1429,1520,1543]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
; 
<specimenCode id="F95E9EA54C38B358FF3BF9C0F664D348" box="[199,305,1556,1580]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="17">AMNH 1</specimenCode>
PT. 
<location id="AC2760054C38B358FE99F9C1F6F5D348" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C38B358FE99F9C1F6F5D348" box="[357,416,1557,1580]" name="Note" pageId="16" pageNumber="17" stateProvince="Texas">Note</location>
: synonym of 
<taxonomicName id="6EF84D5D4C38B358FDCAF9C1F498D349" ID-CoL="4ZJY4" authority="(Smetana 1995: 746)" authorityName="Smetana" authorityPageNumber="746" authorityYear="1995" baseAuthorityName="Smetana" baseAuthorityYear="1995" box="[566,973,1556,1581]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ephippiatus">
<emphasis id="9B8CEACC4C38B358FDCAF9C1F58DD349" box="[566,728,1556,1581]" italics="true" pageId="16" pageNumber="17">B. ephippiatus</emphasis>
(
<treatmentCitation id="285910CF4C38B358FD1AF9C0F491D349" author="" box="[742,964,1556,1581]" page="746" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FD1AF9C0F491D349" box="[742,964,1556,1581]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 746</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BF98FF509D310" blockId="16.[151,1436,1627,1725]" box="[151,604,1627,1653]" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BF98FF7F6D310" bold="true" box="[151,163,1627,1652]" italics="true" pageId="16" pageNumber="17">*</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF5EF98FF509D310" ID-CoL="LC7G" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[162,604,1627,1653]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="erichsoni">
<emphasis id="9B8CEACC4C38B358FF5EF98FF6F4D310" bold="true" box="[162,417,1627,1652]" italics="true" pageId="16" pageNumber="17">Belonuchus erichsoni</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BF9ABF207D3D9" blockId="16.[151,1436,1627,1725]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BF9ABF207D3D9" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BF9ABF5EFD3FC" ID-CoL="LC7G" authority="Bernhauer 1917 d: 223" authorityName="Bernhauer" authorityPageNumber="223" authorityYear="1917" box="[151,698,1663,1688]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="erichsoni">
<emphasis id="9B8CEACC4C38B358FF6BF9ABF6C1D3FC" box="[151,404,1663,1688]" italics="true" pageId="16" pageNumber="17">Belonuchus erichsoni</emphasis>
<treatmentCitation id="285910CF4C38B358FE5AF9ABF5EFD3FC" author="" box="[422,698,1663,1688]" page="223" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE5AF9ABF522D3FC" box="[422,631,1663,1688]" pageId="16" pageNumber="17" refId="ref62307" refString="Bernhauer, M. (1917 d) Vier neue Belonuchus aus Mexiko. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 67, 223 - 224." type="journal article">Bernhauer 1917d</bibRefCitation>
: 223
</treatmentCitation>
</taxonomicName>
. Type locality: [Mexiko]. Type depository: FMNH HT ♂. Distribution: Mexico. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C38B358FCBCF970F3C6D3D8" box="[832,1171,1700,1725]" pageId="16" pageNumber="17" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C38B358FBE5F971F304D3D9" box="[1049,1105,1700,1725]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2002
</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C38B358FB62F970F213D3D9" box="[1182,1350,1700,1725]" pageId="16" pageNumber="17" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BF93FF531D261" blockId="16.[151,1437,1771,1941]" box="[151,612,1771,1797]" pageId="16" pageNumber="17">
<emphasis id="9B8CEACC4C38B358FF6BF93FF7F6D260" bold="true" box="[151,163,1771,1796]" italics="true" pageId="16" pageNumber="17">*</emphasis>
<taxonomicName id="6EF84D5D4C38B358FF5EF93FF531D261" ID-CoL="LC7H" authority="Solsky 1868" authorityName="Solsky" authorityYear="1868" box="[162,612,1771,1797]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="erythropterus">
<emphasis id="9B8CEACC4C38B358FF5EF93FF687D260" bold="true" box="[162,466,1771,1796]" italics="true" pageId="16" pageNumber="17">Belonuchus erythropterus</emphasis>
<bibRefCitation id="CD694B2F4C38B358FE25F938F531D261" box="[473,612,1772,1797]" pageId="16" pageNumber="17" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C38B358FF6BF8DBF62FD2F1" blockId="16.[151,1437,1771,1941]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="89E811F04C38B358FF6BF8DBF62FD2F1" infered="true" pageId="16" pageNumber="17">
<taxonomicName id="6EF84D5D4C38B358FF6BF8DBF5C5D24C" ID-CoL="LC7H" authority="Solsky 1868: 137" authorityName="Solsky" authorityPageNumber="137" authorityYear="1868" box="[151,656,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="erythropterus">
<emphasis id="9B8CEACC4C38B358FF6BF8DBF6EFD24C" box="[151,442,1807,1832]" italics="true" pageId="16" pageNumber="17">Belonuchus erythropterus</emphasis>
<treatmentCitation id="285910CF4C38B358FE38F8DBF5C5D24C" author="" box="[452,656,1807,1832]" page="137" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE38F8DBF503D24C" box="[452,598,1807,1832]" pageId="16" pageNumber="17" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
: 137
</treatmentCitation>
</taxonomicName>
. Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #? ST, not seen by 
<bibRefCitation id="CD694B2F4C38B358FD06F8E0F4E3D229" box="[762,950,1844,1869]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana (1995)</bibRefCitation>
. Distribution: USA (TX), Mexico. Note: redescription, distribution (
<treatmentCitation id="285910CF4C38B358FE01F88CF5B4D214" author="" box="[509,737,1879,1904]" page="752" pageId="16" pageNumber="17">
<bibRefCitation id="CD694B2F4C38B358FE01F88CF5B4D214" box="[509,737,1879,1904]" pageId="16" pageNumber="17" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 752</bibRefCitation>
</treatmentCitation>
); distribution within Mexico (
<bibRefCitation id="CD694B2F4C38B358FBC0F883F2CDD214" box="[1084,1432,1879,1904]" pageId="16" pageNumber="17" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C38B358FAE4F88DF206D214" box="[1304,1363,1879,1904]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2002
</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C38B358FF3BF8A8F63BD2F1" box="[199,366,1916,1941]" pageId="16" pageNumber="17" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFF43F516D5D4" blockId="17.[151,1435,151,248]" box="[151,579,151,176]" pageId="17" pageNumber="18">
<taxonomicName id="6EF84D5D4C39B359FF6BFF43F516D5D4" ID-CoL="LC7J" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,579,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsus">
<emphasis id="9B8CEACC4C39B359FF6BFF43F6DDD5D4" bold="true" box="[151,392,151,176]" italics="true" pageId="17" pageNumber="18">Belonuchus excelsus</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFF68F4FED59C" blockId="17.[151,1435,151,248]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFF68F4FED59C" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BFF68F4F2D59C" ID-GBIF-Occurrence="1915497892" collectionCode="FMNH" collectorName="Umgebung von Cali" country="Colombia" elevation="2000" location="St. Antonio" municipality="Distribution" pageId="17" pageNumber="18" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BFF68F5C4D5B0" ID-CoL="LC7J" authority="Bernhauer 1917 b: 19" authorityName="Bernhauer" authorityPageNumber="19" authorityYear="1917" box="[151,657,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="excelsus">
<emphasis id="9B8CEACC4C39B359FF6BFF68F6D0D5B1" box="[151,389,188,213]" italics="true" pageId="17" pageNumber="18">Belonuchus excelsus</emphasis>
<treatmentCitation id="285910CF4C39B359FE6FFF68F5C4D5B0" author="" box="[403,657,188,213]" page="19" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE6FFF68F535D5B1" box="[403,608,188,213]" pageId="17" pageNumber="18" refId="ref62221" refString="Bernhauer, M. (1917 b) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (17. Beitrag - Fortsetzung). Entomologische Blatter, 13 (1 - 3), 17 - 24." type="journal article">Bernhauer 1917b</bibRefCitation>
: 19
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C39B359FD58FF69F58ED5B1" box="[676,731,189,213]" pageId="17" pageNumber="18">Type</typeStatus>
locality: 
<location id="AC2760054C39B359FCA8FF68F48AD5B1" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FCA8FF68F48AD5B1" box="[852,991,188,213]" country="Colombia" municipality="Distribution" name="St. Antonio" pageId="17" pageNumber="18">St. Antonio</location>
, 
<elevation id="22D5D1ED4C39B359FC0DFF68F317D5B0" box="[1009,1090,188,212]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="m" value="2000.0">
<quantity id="6E009B3B4C39B359FC0DFF68F317D5B0" box="[1009,1090,188,212]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="m" value="2000.0">2000m</quantity>
</elevation>
, 
<collectorName id="040D53084C39B359FBA8FF68F21DD5B1" box="[1108,1352,188,213]" pageId="17" pageNumber="18">Umgebung von Cali</collectorName>
, West- Columbien. 
<typeStatus id="7643887C4C39B359FEADFF34F6DCD59C" box="[337,393,224,248]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C39B359FDE9FF34F533D593" box="[533,614,224,247]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
HT. 
<collectingMunicipality id="4923ACA44C39B359FD62FF0BF47FD59C" box="[670,810,223,248]" pageId="17" pageNumber="18">Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C39B359FCC8FF0BF4F2D59C" box="[820,935,223,248]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFEF3F41FD424" blockId="17.[151,1437,295,608]" box="[151,842,295,320]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFEF3F7F6D424" bold="true" box="[151,163,295,320]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="6EF84D5D4C39B359FF5EFEF3F569D424" ID-CoL="LC7L" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,572,295,320]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="extremus" status="comb. nov.">
<emphasis id="9B8CEACC4C39B359FF5EFEF3F6F5D45B" bold="true" box="[162,416,295,320]" italics="true" pageId="17" pageNumber="18">Belonuchus extremus</emphasis>
(
<bibRefCitation id="CD694B2F4C39B359FE53FEF3F561D424" box="[431,564,295,320]" pageId="17" pageNumber="18" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C39B359FDB4FEF3F59DD424" box="[584,712,295,320]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FDB4FEF3F59DD424" bold="true" box="[584,712,295,320]" pageId="17" pageNumber="18">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C39B359FD24FEF3F417D424" box="[728,834,295,320]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="17" pageNumber="18">Figs. 7–8</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFE98F68BD4EC" blockId="17.[151,1437,295,608]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFE98F68BD4EC" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BFE98F68FD4EC" ID-GBIF-Occurrence="1915497853" collectionCode="BMNH" country="Guatemala" location="Distribution" pageId="17" pageNumber="18" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BFE98F518D401" ID-CoL="76XP4" authority="Sharp 1885: 415" authorityName="Sharp" authorityPageNumber="415" authorityYear="1885" box="[151,589,332,357]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="extremus">
<emphasis id="9B8CEACC4C39B359FF6BFE98F62AD400" box="[151,383,332,357]" italics="true" pageId="17" pageNumber="18">Philonthus extremus</emphasis>
<treatmentCitation id="285910CF4C39B359FE76FE98F518D401" author="" box="[394,589,332,357]" page="415" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE76FE98F547D401" box="[394,530,332,357]" pageId="17" pageNumber="18" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 415
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C39B359FDA0FE99F5C1D401" box="[604,660,333,357]" pageId="17" pageNumber="18">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C39B359FCF9FE98F4D4D401" box="[773,897,332,357]" name="Guatemala" pageId="17" pageNumber="18">Guatemala</collectingCountry>
, San Gerónimo. 
<typeStatus id="7643887C4C39B359FBB4FE99F32AD401" box="[1096,1151,333,357]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C39B359FAEFFE99F232D400" box="[1299,1383,333,356]" pageId="17" pageNumber="18">BMNH</collectionCode>
HT. 
<location id="AC2760054C39B359FF3BFEBBF606D4EC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FF3BFEBBF606D4EC" box="[199,339,367,392]" country="Guatemala" name="Distribution" pageId="17" pageNumber="18">Distribution</location>
: 
<collectingCountry id="D1EF764E4C39B359FEA1FEBBF68FD4EC" box="[349,474,367,392]" name="Guatemala" pageId="17" pageNumber="18">Guatemala</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C39B359FF6BFE40F48BD4C9" box="[151,990,404,429]" pageId="17" pageNumber="18" type="discussion">
<paragraph id="A94736DE4C39B359FF6BFE40F48BD4C9" blockId="17.[151,1437,295,608]" box="[151,990,404,429]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFE40F65ED4C9" bold="true" box="[151,267,404,429]" pageId="17" pageNumber="18">Remarks.</emphasis>
The “unique” specimen in BMNH is confirmed as the 
<typeStatus id="7643887C4C39B359FC88FE40F48FD4C9" box="[884,986,404,429]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C39B359FF6BFE63F525D759" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="A94736DE4C39B359FF6BFE63F525D759" blockId="17.[151,1437,295,608]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFE63F696D4B4" bold="true" box="[151,451,439,464]" pageId="17" pageNumber="18">Type material examined.</emphasis>
Holotype ♂ (BMNH, 
<figureCitation id="31C32A5B4C39B359FD35FE63F46DD4B4" box="[713,824,439,464]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="17" pageNumber="18">Figs. 7–8</figureCitation>
): card-mounted, “ 
<taxonomicName id="6EF84D5D4C39B359FBFFFE63F3BBD4AB" box="[1027,1262,439,464]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="extremus">Philonthus extremus</taxonomicName>
Type D.S. San Geronimo, Guatemala. Champion. [in Sharp’s hand] // Type (round white label with red margin) // San Geronimo, Verapaz. Champion. (white label) // B. C. A. Col. 
<date id="DD46101E4C39B359FC81FDD4F4FFD77C" box="[893,938,512,536]" pageId="17" pageNumber="18">I. 2.</date>
<taxonomicName id="6EF84D5D4C39B359FC4DFE2BF3BAD77C" authority=", Sharp." authorityName="Sharp." box="[945,1263,511,536]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="extremus">Philonthus extremus, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C39B359FF6BFD93F560D7CC" pageId="17" pageNumber="18" type="discussion">
<paragraph id="A94736DE4C39B359FF6BFD93F33DD704" blockId="17.[151,1437,295,608]" box="[151,1128,583,608]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFD93F64AD704" bold="true" box="[151,287,583,608]" pageId="17" pageNumber="18">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C39B359FD46FD93F331D704" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,583,608]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C39B359FD46FD93F468D704" box="[698,829,583,608]" italics="true" pageId="17" pageNumber="18">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C39B359FCB7FD93F309D704" box="[843,1116,583,608]" pageId="17" pageNumber="18" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C39B359FC1CFD9DF34DD704" box="[992,1048,583,608]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFD5BF560D7CC" blockId="17.[151,1436,655,752]" box="[151,565,655,680]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFD5BF7F6D7CC" bold="true" box="[151,163,655,680]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="6EF84D5D4C39B359FF5EFD5BF560D7CC" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[162,565,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="9B8CEACC4C39B359FF5EFD5BF62FD7CC" bold="true" box="[162,378,655,680]" italics="true" pageId="17" pageNumber="18">Belonuchus falsus</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C39B359FF6BFD60F383D081" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="A94736DE4C39B359FF6BFD60F68DD794" blockId="17.[151,1436,655,752]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFD60F68DD794" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BFD60F4D9D7A9" ID-GBIF-Occurrence="1915497872" box="[151,908,692,717]" country="Venezuela" location="Venezuela" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BFD60F532D7A9" ID-CoL="LC7M" authority="Bernhauer 1917 c: 105" authorityName="Bernhauer" authorityPageNumber="105" authorityYear="1917" box="[151,615,692,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="falsus">
<emphasis id="9B8CEACC4C39B359FF6BFD60F637D7A9" box="[151,354,692,717]" italics="true" pageId="17" pageNumber="18">Belonuchus falsus</emphasis>
<treatmentCitation id="285910CF4C39B359FE96FD60F532D7A9" author="" box="[362,615,692,717]" page="105" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE96FD60F57AD7A8" box="[362,559,692,717]" pageId="17" pageNumber="18" refId="ref62261" refString="Bernhauer, M. (1917 c) Neue sudamerikanische Staphyliniden (18. Beitrag). Wiener Entomologische Zeitung, 36 (3 - 5), 102 - 116. https: // doi. org / 10.5962 / bhl. part. 12925" type="journal article">Bernhauer 1917c</bibRefCitation>
: 105
</treatmentCitation>
</taxonomicName>
. 
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locality: 
<collectingCountry id="D1EF764E4C39B359FCE9FD60F4D9D7A9" box="[789,908,692,717]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C39B359FC6AFD61F681D794" ID-GBIF-Occurrence="1915497845" collectionCode="FMNH, ZMUH" country="Venezuela" location="Distribution" pageId="17" pageNumber="18" specimenCode="FMNH 1, ZMUH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C39B359FC6AFD61F49BD7A9" box="[918,974,693,717]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C39B359FBA7FD61F3EBD7A8" box="[1115,1214,692,716]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH 1</specimenCode>
ST; 
<specimenCode id="F95E9EA54C39B359FB0CFD61F201D7A8" box="[1264,1364,692,716]" collectionCode="ZMUH" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="17" pageNumber="18">ZMUH 1</specimenCode>
+ST?. 
<location id="AC2760054C39B359FF3BFD03F606D794" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FF3BFD03F606D794" box="[199,339,727,752]" country="Venezuela" name="Distribution" pageId="17" pageNumber="18">Distribution</location>
: 
<collectingCountry id="D1EF764E4C39B359FEA1FD03F681D794" box="[349,468,727,752]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
</materialsCitation>
.
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</paragraph>
<paragraph id="A94736DE4C39B359FF6BFCCBF578D65C" blockId="17.[151,1436,799,896]" box="[151,557,799,824]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFCCBF7F6D65C" bold="true" box="[151,163,799,824]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="6EF84D5D4C39B359FF5EFCCBF578D65C" ID-CoL="LC7N" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[162,557,799,824]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fassli">
<emphasis id="9B8CEACC4C39B359FF5EFCCBF627D65C" bold="true" box="[162,370,799,824]" italics="true" pageId="17" pageNumber="18">Belonuchus fassli</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFC90F3CCD6E4" blockId="17.[151,1436,799,896]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFC90F3CCD6E4" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BFC90F3C1D6E4" ID-GBIF-Occurrence="1915497893" collectionCode="FMNH" collectorName="West-Columbien &amp; Umgebung von Cali &amp; Canon" country="Colombia" elevation="1700" location="Cauca" pageId="17" pageNumber="18" specimenCount="2" stateProvince="Cauca" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BFC90F532D638" ID-CoL="LC7N" authority="Bernhauer 1917 a: 37" authorityName="Bernhauer" authorityPageNumber="37" authorityYear="1917" box="[151,615,836,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fassli">
<emphasis id="9B8CEACC4C39B359FF6BFC90F634D639" box="[151,353,836,861]" italics="true" pageId="17" pageNumber="18">Belonuchus fassli</emphasis>
<treatmentCitation id="285910CF4C39B359FE92FC90F532D638" author="" box="[366,615,836,861]" page="37" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE92FC90F56DD638" box="[366,568,836,861]" pageId="17" pageNumber="18" refId="ref62187" refString="Bernhauer, M. (1917 a) Beitrag zur Staphyliniden-Fauna Sudamerikas. (17. Beitrag). Neue Beitrage zur Systematischen Insektenkunde, 1 (5), 35 - 38." type="journal article">Bernhauer 1917a</bibRefCitation>
: 37
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C39B359FD84FC91F5E5D639" box="[632,688,837,861]" pageId="17" pageNumber="18">Type</typeStatus>
locality: 
<collectorName id="040D53084C39B359FCD9FC90F4B2D639" box="[805,999,836,861]" pageId="17" pageNumber="18">West-Columbien</collectorName>
: 
<collectorName id="040D53084C39B359FC0BFC90F3B3D639" box="[1015,1254,836,861]" pageId="17" pageNumber="18">Umgebung von Cali</collectorName>
am Rio 
<collectingRegion id="6B3CF83C4C39B359FAAAFC90F2C9D638" box="[1366,1436,836,860]" country="Colombia" name="Cauca" pageId="17" pageNumber="18">Cauca</collectingRegion>
(
<collectorName id="040D53084C39B359FF32FCBCF64CD6E4" box="[206,281,872,896]" pageId="17" pageNumber="18">Cañon</collectorName>
del Mte 
<collectingRegion id="6B3CF83C4C39B359FE81FCB3F69AD6E4" box="[381,463,871,896]" country="Colombia" name="Tolima" pageId="17" pageNumber="18">Tolima</collectingRegion>
, 
<elevation id="22D5D1ED4C39B359FE27FCBCF578D6E4" box="[475,557,872,896]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="m" value="1700.0">
<quantity id="6E009B3B4C39B359FE27FCBCF578D6E4" box="[475,557,872,896]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="m" value="1700.0">1700m</quantity>
</elevation>
). 
<typeStatus id="7643887C4C39B359FDBCFCBCF522D6E4" box="[576,631,872,896]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C39B359FCFFFCBCF401D61B" box="[771,852,872,895]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
HT. Distribution: 
<collectingCountry id="D1EF764E4C39B359FBDEFCB3F3C1D6E4" box="[1058,1172,871,896]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFC7BF531D6AC" blockId="17.[151,1436,943,1077]" box="[151,612,943,968]" pageId="17" pageNumber="18">
<taxonomicName id="6EF84D5D4C39B359FF6BFC7BF531D6AC" ID-CoL="LC7Q" authority="(Bernhauer 1912)" authorityName="Bernhauer" authorityYear="1912" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[151,612,943,968]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavicoxis">
<emphasis id="9B8CEACC4C39B359FF6BFC7BF6C3D6AC" bold="true" box="[151,406,943,968]" italics="true" pageId="17" pageNumber="18">Belonuchus flavicoxis</emphasis>
(Bernhauer 1912)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFC00F4AAD150" blockId="17.[151,1436,943,1077]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFC00F4AAD150" pageId="17" pageNumber="18">
<taxonomicName id="6EF84D5D4C39B359FF6BFC00F52FD688" ID-CoL="76XNF" authority="Bernhauer 1912 a: 43" authorityName="Bernhauer" authorityPageNumber="43" authorityYear="1912" box="[151,634,980,1005]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavicoxis">
<emphasis id="9B8CEACC4C39B359FF6BFC00F6D6D689" box="[151,387,980,1005]" italics="true" pageId="17" pageNumber="18">Philonthus flavicoxis</emphasis>
<treatmentCitation id="285910CF4C39B359FE77FC00F52FD688" author="" box="[395,634,980,1005]" page="43" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE77FC00F505D688" box="[395,592,980,1005]" pageId="17" pageNumber="18" refId="ref61973" refString="Bernhauer, M. (1912 a) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article">Bernhauer 1912a</bibRefCitation>
: 43
</treatmentCitation>
</taxonomicName>
. Type locality: Tucuman prov., Argentina. Type depository: FMNH LT ♀. Distribution: Argentina. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C39B359FD0FFC23F439D174" ID-CoL="8VRYR" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[755,876,1015,1040]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C39B359FD0FFC23F439D174" box="[755,876,1015,1040]" italics="true" pageId="17" pageNumber="18">Philonthus</emphasis>
</taxonomicName>
and lectotype designation (
<treatmentCitation id="285910CF4C39B359FB5CFC23F2D8D174" author="" box="[1184,1421,1015,1040]" page="8" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FB5CFC23F2D8D174" box="[1184,1421,1015,1040]" pageId="17" pageNumber="18" refId="ref64208" refString="Chani-Posse, M. (2010) Revision of the southern South American species of Philonthus Stephens Coleoptera: Staphylinidae. Zootaxa, 2595, 1 - 70." type="journal article">Chani-Posse 2010: 8</bibRefCitation>
</treatmentCitation>
); distribution (Chani-Posse 2004); redescription (
<treatmentCitation id="285910CF4C39B359FD20FBC8F4A6D150" author="" box="[732,1011,1052,1077]" page="221" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FD20FBC8F4A6D150" box="[732,1011,1052,1077]" pageId="17" pageNumber="18" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b: 221</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFBB7F51DD118" blockId="17.[151,1437,1123,1221]" box="[151,584,1123,1149]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFBB7F7F6D118" bold="true" box="[151,163,1123,1148]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="6EF84D5D4C39B359FF5EFBB7F51DD118" ID-CoL="LC7R" authority="Solsky 1870" authorityName="Solsky" authorityYear="1870" box="[162,584,1123,1149]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavipennis">
<emphasis id="9B8CEACC4C39B359FF5EFBB7F6E0D118" bold="true" box="[162,437,1123,1148]" italics="true" pageId="17" pageNumber="18">Belonuchus flavipennis</emphasis>
<bibRefCitation id="CD694B2F4C39B359FE41FBB0F51DD118" box="[445,584,1124,1149]" pageId="17" pageNumber="18" refId="ref73086" refString="Solsky, S. (1870) Staphylins de l'Amerique meridionale et du Mexique. II. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 42 (4), 257 - 267. [1869]" type="journal article">Solsky 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFB53F21AD1A0" blockId="17.[151,1437,1123,1221]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFB53F21AD1A0" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BFB53F4F7D1C4" ID-GBIF-Occurrence="1915497889" box="[151,930,1159,1184]" country="Mexico" location="Mexique" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BFB53F524D1C4" ID-CoL="LC7R" authority="Solsky 1870: 263" authorityName="Solsky" authorityPageNumber="263" authorityYear="1870" box="[151,625,1159,1184]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavipennis">
<emphasis id="9B8CEACC4C39B359FF6BFB53F6CBD1C4" box="[151,414,1159,1184]" italics="true" pageId="17" pageNumber="18">Belonuchus flavipennis</emphasis>
<treatmentCitation id="285910CF4C39B359FE5AFB53F524D1C4" author="" box="[422,625,1159,1184]" page="263" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE5AFB53F56DD1C4" box="[422,568,1159,1184]" pageId="17" pageNumber="18" refId="ref73086" refString="Solsky, S. (1870) Staphylins de l'Amerique meridionale et du Mexique. II. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 42 (4), 257 - 267. [1869]" type="journal article">Solsky 1870</bibRefCitation>
: 263
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C39B359FD82FB5CF5E3D1C4" box="[638,694,1160,1184]" pageId="17" pageNumber="18">Type</typeStatus>
locality: &quot; 
<location id="AC2760054C39B359FCD0FB53F4CDD1C4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FCD0FB53F4CDD1C4" box="[812,920,1159,1184]" country="Mexico" name="Mexique" pageId="17" pageNumber="18">
<collectingCountry id="D1EF764E4C39B359FCD0FB53F4CDD1C4" box="[812,920,1159,1184]" name="Mexico" pageId="17" pageNumber="18">Mexique</collectingCountry>
</location>
&quot;
</materialsCitation>
. 
<materialsCitation id="19903C834C39B359FC52FB5CF675D1A1" ID-GBIF-Occurrence="1915497858" collectionCode="ZIN" country="Mexico" location="Distribution" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Mexico" typeStatus="holotype">
<typeStatus id="7643887C4C39B359FC52FB5CF4B3D1C4" box="[942,998,1160,1184]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C39B359FB8AFB5CF3F2D1C4" box="[1142,1191,1160,1184]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="17" pageNumber="18">ZIN</collectionCode>
? HT 
<specimenCount id="BFFEFD574C39B359FB14FB5CF3ABD1FB" box="[1256,1278,1160,1183]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
. 
<location id="AC2760054C39B359FAF1FB53F2CCD1C4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FAF1FB53F2CCD1C4" box="[1293,1433,1159,1184]" country="Mexico" name="Distribution" pageId="17" pageNumber="18" stateProvince="Mexico">Distribution</location>
: 
<collectingRegion id="6B3CF83C4C39B359FF3BFB78F675D1A1" box="[199,288,1196,1221]" country="Mexico" name="Mexico" pageId="17" pageNumber="18">
<collectingCountry id="D1EF764E4C39B359FF3BFB78F675D1A1" box="[199,288,1196,1221]" name="Mexico" pageId="17" pageNumber="18">Mexico</collectingCountry>
</collectingRegion>
</materialsCitation>
; 
<materialsCitation id="19903C834C39B359FED7FB78F6E0D1A1" ID-GBIF-Occurrence="1915497883" box="[299,437,1196,1221]" country="Costa Rica" location="Costa" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Costa" typeStatus="holotype">
<collectingCountry id="D1EF764E4C39B359FED7FB78F6FFD1A1" box="[299,426,1196,1221]" name="Costa Rica" pageId="17" pageNumber="18">
<collectingRegion id="6B3CF83C4C39B359FED7FB78F63ED1A0" box="[299,363,1196,1220]" country="Tanzania" name="Pwani" pageId="17" pageNumber="18">Costa</collectingRegion>
Rica
</collectingCountry>
?
</materialsCitation>
, 
<materialsCitation id="19903C834C39B359FE3CFB78F57DD1A0" ID-GBIF-Occurrence="1915497861" box="[448,552,1196,1220]" country="Panama" location="Panama" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Panama" typeStatus="holotype">
<collectingRegion id="6B3CF83C4C39B359FE3CFB78F54BD1A0" box="[448,542,1196,1220]" country="Panama" name="Panama" pageId="17" pageNumber="18">
<collectingCountry id="D1EF764E4C39B359FE3CFB78F54BD1A0" box="[448,542,1196,1220]" name="Panama" pageId="17" pageNumber="18">Panama</collectingCountry>
</collectingRegion>
?
</materialsCitation>
. Note: distribution, also within Mexico (
<bibRefCitation id="CD694B2F4C39B359FC12FB78F216D1A0" box="[1006,1347,1196,1221]" pageId="17" pageNumber="18" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C39B359FB3BFB79F3AAD1A1" box="[1223,1279,1196,1221]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFB27F5C6D068" blockId="17.[151,1436,1267,1365]" box="[151,659,1267,1293]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFB27F7F6D068" bold="true" box="[151,163,1267,1292]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="6EF84D5D4C39B359FF5EFB27F5C6D068" ID-CoL="LC7S" authority="Bernhauer 1921" authorityName="Bernhauer" authorityYear="1921" box="[162,659,1267,1293]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavocaudatus">
<emphasis id="9B8CEACC4C39B359FF5EFB27F68DD068" bold="true" box="[162,472,1267,1292]" italics="true" pageId="17" pageNumber="18">Belonuchus flavocaudatus</emphasis>
Bernhauer 1921
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFAC3F4D6D031" blockId="17.[151,1436,1267,1365]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFAC3F4D6D031" pageId="17" pageNumber="18">
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<taxonomicName id="6EF84D5D4C39B359FF6BFAC3F5B5D054" ID-CoL="LC7S" authority="Bernhauer 1921 c: 170" authorityName="Bernhauer" authorityPageNumber="170" authorityYear="1921" box="[151,736,1303,1328]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="flavocaudatus">
<emphasis id="9B8CEACC4C39B359FF6BFAC3F690D054" box="[151,453,1303,1328]" italics="true" pageId="17" pageNumber="18">Belonuchus flavocaudatus</emphasis>
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: 170
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. 
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locality: 
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, 
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, 
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, 
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</materialsCitation>
. 
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depository: 
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ST; 
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<location id="AC2760054C39B359FDB8FAE8F454D031" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FDB8FAE8F454D031" box="[580,769,1340,1365]" country="Argentina" name="ST. Distribution" pageId="17" pageNumber="18">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C39B359FCF7FAE8F42BD031" box="[779,894,1340,1365]" name="Argentina" pageId="17" pageNumber="18">Argentina</collectingCountry>
</materialsCitation>
.
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</paragraph>
<paragraph id="A94736DE4C39B359FF6BFA57F510D0F9" blockId="17.[151,1436,1411,1544]" box="[151,581,1411,1437]" pageId="17" pageNumber="18">
<taxonomicName id="6EF84D5D4C39B359FF6BFA57F510D0F9" ID-CoL="LC7T" authority="Scheerpeltz 1960" authorityName="Scheerpeltz" authorityYear="1960" box="[151,581,1411,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="forsteri">
<emphasis id="9B8CEACC4C39B359FF6BFA57F629D0F8" bold="true" box="[151,380,1411,1436]" italics="true" pageId="17" pageNumber="18">Belonuchus forsteri</emphasis>
<bibRefCitation id="CD694B2F4C39B359FE7FFA50F510D0F9" box="[387,581,1412,1437]" pageId="17" pageNumber="18" refId="ref71794" refString="Scheerpeltz, O. (1960) Zur Kenntnis neotropischer Staphyliniden (Col.). Beitrage zur Neotropischen Fauna, 2 (2), 65 - 138. https: // doi. org / 10.1080 / 01650526009380624" type="journal article">Scheerpeltz 1960</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BFA73F383D081" blockId="17.[151,1436,1411,1544]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BFA73F383D081" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BFA73F688D081" ID-GBIF-Occurrence="1915497867" country="Bolivia" elevation="2350" location="Yungas de Arepucho" municipality="Sihuencas" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BFA73F401D0A4" ID-CoL="LC7T" authority="Scheerpeltz 1960: 115" authorityName="Scheerpeltz" authorityPageNumber="115" authorityYear="1960" box="[151,852,1447,1472]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="forsteri" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C39B359FF6BFA73F567D0A4" box="[151,562,1447,1472]" italics="true" pageId="17" pageNumber="18">Belonuchus (Musicoderus) forsteri</emphasis>
<treatmentCitation id="285910CF4C39B359FDBFFA73F401D0A4" author="" box="[579,852,1447,1472]" page="115" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FDBFFA73F446D0A4" box="[579,787,1447,1472]" pageId="17" pageNumber="18" refId="ref71794" refString="Scheerpeltz, O. (1960) Zur Kenntnis neotropischer Staphyliniden (Col.). Beitrage zur Neotropischen Fauna, 2 (2), 65 - 138. https: // doi. org / 10.1080 / 01650526009380624" type="journal article">Scheerpeltz 1960</bibRefCitation>
: 115
</treatmentCitation>
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. 
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locality: 
<collectingCountry id="D1EF764E4C39B359FBE2FA73F3D7D0A4" box="[1054,1154,1447,1472]" name="Bolivia" pageId="17" pageNumber="18">Bolivien</collectingCountry>
, 
<location id="AC2760054C39B359FB6BFA7CF2C2D0A4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FB6BFA7CF2C2D0A4" box="[1175,1431,1447,1472]" country="Bolivia" municipality="Sihuencas" name="Yungas de Arepucho" pageId="17" pageNumber="18">Yungas de Arepucho</location>
, 
<collectingMunicipality id="4923ACA44C39B359FF3BFA18F669D081" box="[199,316,1484,1509]" pageId="17" pageNumber="18">Sihuencas</collectingMunicipality>
, 
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</materialsCitation>
. 
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<typeStatus id="7643887C4C39B359FE14FA19F575D081" box="[488,544,1485,1509]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C39B359FD50FA19F5A1D080" box="[684,756,1485,1508]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18">NMW</collectionCode>
(
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) HT 
<specimenCount id="BFFEFD574C39B359FC3FFA19F48CD080" box="[963,985,1485,1508]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
. 
<collectingMunicipality id="4923ACA44C39B359FC1AFA18F327D081" box="[998,1138,1484,1509]" pageId="17" pageNumber="18">Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C39B359FB80FA18F387D081" box="[1148,1234,1484,1509]" name="Bolivia" pageId="17" pageNumber="18">Bolivia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C39B359FF6BFA3BF465D36C" box="[151,816,1519,1544]" pageId="17" pageNumber="18" type="discussion">
<paragraph id="A94736DE4C39B359FF6BFA3BF465D36C" blockId="17.[151,1436,1411,1544]" box="[151,816,1519,1544]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BFA3BF641D36C" bold="true" box="[151,276,1519,1544]" pageId="17" pageNumber="18">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C39B359FEB0FA3BF69AD36C" box="[332,463,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C39B359FEB0FA3BF69AD36C" box="[332,463,1519,1544]" italics="true" pageId="17" pageNumber="18">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C39B35AFF6BF9E3F365D689" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="A94736DE4C39B359FF6BF9E3F50AD334" blockId="17.[151,1436,1591,1725]" box="[151,607,1591,1616]" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BF9E3F7F6D334" bold="true" box="[151,163,1591,1616]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="6EF84D5D4C39B359FF5EF9E3F50AD334" ID-CoL="5WGHZ" authority="Wasmann 1887" authorityName="Wasmann" authorityYear="1887" box="[162,607,1591,1616]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fossulatus">
<emphasis id="9B8CEACC4C39B359FF5EF9E3F6FFD334" bold="true" box="[162,426,1591,1616]" italics="true" pageId="17" pageNumber="18">Belonuchus fossulatus</emphasis>
<bibRefCitation id="CD694B2F4C39B359FE4EF9ECF50AD334" box="[434,607,1592,1616]" pageId="17" pageNumber="18" refId="ref73770" refString="Wasmann, E. (1887) Neue Brasilianische Staphyliniden, bei Eciton hamatum gesammelt von Dr. W. Muller. Deutsche Entomologische Zeitschrift, 31 (2), 403 - 416, pl. 5." type="journal article">Wasmann 1887</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BF988F6FBD3D8" blockId="17.[151,1436,1591,1725]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BF988F6FBD3D8" infered="true" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BF988F205D311" ID-GBIF-Occurrence="1915497899" box="[151,1360,1628,1653]" country="Brazil" county="Blumenau" location="St. Catharina" municipality="Prov. St." pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BF988F5FCD310" ID-CoL="5WGHZ" authority="Wasmann 1887: 414" authorityName="Wasmann" authorityPageNumber="414" authorityYear="1887" box="[151,681,1628,1653]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fossulatus">
<emphasis id="9B8CEACC4C39B359FF6BF988F6CED311" box="[151,411,1628,1653]" italics="true" pageId="17" pageNumber="18">Belonuchus fossulatus</emphasis>
<treatmentCitation id="285910CF4C39B359FE50F989F5FCD310" author="" box="[428,681,1628,1652]" page="414" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE50F989F533D310" box="[428,614,1628,1652]" pageId="17" pageNumber="18" refId="ref73770" refString="Wasmann, E. (1887) Neue Brasilianische Staphyliniden, bei Eciton hamatum gesammelt von Dr. W. Muller. Deutsche Entomologische Zeitschrift, 31 (2), 403 - 416, pl. 5." type="journal article">Wasmann 1887</bibRefCitation>
: 414
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C39B359FD42F989F5A0D311" box="[702,757,1629,1653]" pageId="17" pageNumber="18">Type</typeStatus>
locality: 
<collectingCounty id="40264E524C39B359FC8FF988F4B3D311" box="[883,998,1628,1653]" pageId="17" pageNumber="18">Blumenau</collectingCounty>
(
<collectingMunicipality id="4923ACA44C39B359FC03F988F33FD310" box="[1023,1130,1628,1652]" pageId="17" pageNumber="18">
Prov. 
<location id="AC2760054C39B359FBB0F988F33FD310" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FBB0F988F33FD310" box="[1100,1130,1628,1652]" country="Brazil" county="Blumenau" municipality="Prov. St." name="St. Catharina" pageId="17" pageNumber="18">St.</location>
</collectingMunicipality>
Catharina, 
<collectingCountry id="D1EF764E4C39B359FAFCF988F21DD311" box="[1280,1352,1628,1653]" name="Brazil" pageId="17" pageNumber="18">Brasil</collectingCountry>
)
</materialsCitation>
. 
<materialsCitation id="19903C834C39B359FA98F989F379D3FC" ID-GBIF-Occurrence="1915497851" collectionCode="NHME" country="Brazil" location="Distribution" municipality="Wasmann" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Santa Catarina" typeStatus="holotype">
<typeStatus id="7643887C4C39B359FA98F989F2C9D311" box="[1380,1436,1629,1653]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C39B359FEB1F954F6F5D3F3" box="[333,416,1664,1687]" country="Netherlands" httpUri="http://grbio.org/cool/6ssp-k6rj" name="Natuurhistorisch Museum" pageId="17" pageNumber="18">NHME</collectionCode>
(
<collectingMunicipality id="4923ACA44C39B359FE53F954F575D3FC" box="[431,544,1664,1688]" pageId="17" pageNumber="18">Wasmann</collectingMunicipality>
) 1ST 
<specimenCount id="BFFEFD574C39B359FD94F954F52BD3F3" box="[616,638,1664,1687]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
. 
<location id="AC2760054C39B359FD70F9ABF44DD3FC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FD70F9ABF44DD3FC" box="[652,792,1663,1688]" country="Brazil" municipality="Wasmann" name="Distribution" pageId="17" pageNumber="18" stateProvince="Santa Catarina">Distribution</location>
: 
<collectingCountry id="D1EF764E4C39B359FCD8F9ABF43DD3FC" box="[804,872,1663,1688]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C39B359FC84F954F371D3FC" box="[888,1060,1663,1688]" country="Brazil" name="Santa Catarina" pageId="17" pageNumber="18">Santa Catarina</collectingRegion>
)
</materialsCitation>
. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C39B359FEDAF971F60AD3D9" box="[294,351,1700,1725]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BF93FF571D261" blockId="17.[151,1437,1771,1869]" box="[151,548,1771,1797]" pageId="17" pageNumber="18">
<taxonomicName id="6EF84D5D4C39B359FF6BF93FF571D261" ID-CoL="LC7V" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[151,548,1771,1797]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="9B8CEACC4C39B359FF6BF93FF63CD260" bold="true" box="[151,361,1771,1796]" italics="true" pageId="17" pageNumber="18">Belonuchus frater</emphasis>
<bibRefCitation id="CD694B2F4C39B359FE8DF938F571D261" box="[369,548,1772,1797]" pageId="17" pageNumber="18" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BF8DBF434D229" blockId="17.[151,1437,1771,1869]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BF8DBF434D229" pageId="17" pageNumber="18">
<materialsCitation id="19903C834C39B359FF6BF8DBF400D243" ID-GBIF-Occurrence="1915497846" box="[151,853,1807,1832]" country="Peru" location="Peru" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C39B359FF6BF8DBF533D24C" ID-CoL="LC7V" authority="Bernhauer 1906: 333" authorityName="Bernhauer" authorityPageNumber="333" authorityYear="1906" box="[151,614,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="9B8CEACC4C39B359FF6BF8DBF631D24C" box="[151,356,1807,1832]" italics="true" pageId="17" pageNumber="18">Belonuchus frater</emphasis>
<treatmentCitation id="285910CF4C39B359FE93F8DBF533D24C" author="" box="[367,614,1807,1832]" page="333" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE93F8DBF57FD24C" box="[367,554,1807,1832]" pageId="17" pageNumber="18" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 333
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C39B359FD89F8C4F5F8D24C" box="[629,685,1808,1832]" pageId="17" pageNumber="18">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C39B359FCE2F8C4F400D243" box="[798,853,1808,1831]" name="Peru" pageId="17" pageNumber="18">Peru</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C39B359FC9FF8C4F2CCD243" ID-GBIF-Occurrence="1915497785" box="[867,1433,1807,1832]" collectionCode="FMNH" country="Peru" location="ST. Distribution" pageId="17" pageNumber="18" specimenCode="FMNH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C39B359FC9FF8C4F4CFD24C" box="[867,922,1808,1832]" pageId="17" pageNumber="18">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C39B359FBD2F8C4F3C6D24C" box="[1070,1171,1808,1832]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH 1</specimenCode>
<location id="AC2760054C39B359FB6FF8C4F201D24C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C39B359FB6FF8C4F201D24C" box="[1171,1364,1807,1832]" country="Peru" name="ST. Distribution" pageId="17" pageNumber="18">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C39B359FA9EF8C4F2CCD243" box="[1378,1433,1808,1831]" name="Peru" pageId="17" pageNumber="18">Peru</collectingCountry>
</materialsCitation>
. Note: Peru (
<bibRefCitation id="CD694B2F4C39B359FEB2F8E0F6B6D228" box="[334,483,1844,1869]" pageId="17" pageNumber="18" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B359FF6BF8AFF2C9D2DC" blockId="17.[151,1436,1915,2013]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="89E811F04C39B359FF6BF8AFF2C9D2DC" pageId="17" pageNumber="18">
<emphasis id="9B8CEACC4C39B359FF6BF8AFF7F6D2F0" bold="true" box="[151,163,1915,1940]" italics="true" pageId="17" pageNumber="18">*</emphasis>
<taxonomicName id="6EF84D5D4C39B359FF5EF8AFF563D2F0" ID-CoL="LC7X" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[162,566,1915,1941]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gagates">
<emphasis id="9B8CEACC4C39B359FF5EF8AFF6D8D2F7" bold="true" box="[162,397,1915,1940]" italics="true" pageId="17" pageNumber="18">Belonuchus gagates</emphasis>
<bibRefCitation id="CD694B2F4C39B359FE68F8A8F563D2F0" box="[404,566,1916,1941]" pageId="17" pageNumber="18" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
<taxonomicName id="6EF84D5D4C39B359FF6BF874F53CD2DC" ID-CoL="LC7X" authority="Erichson 1840: 424" authorityName="Erichson" authorityPageNumber="424" authorityYear="1840" box="[151,617,1952,1977]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gagates">
<emphasis id="9B8CEACC4C39B359FF6BF874F62FD2DC" box="[151,378,1952,1977]" italics="true" pageId="17" pageNumber="18">Belonuchus gagates</emphasis>
<treatmentCitation id="285910CF4C39B359FE78F874F53CD2DC" author="" box="[388,617,1952,1977]" page="424" pageId="17" pageNumber="18">
<bibRefCitation id="CD694B2F4C39B359FE78F874F57BD2DC" box="[388,558,1952,1977]" pageId="17" pageNumber="18" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 424
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C39B359FD8AF875F5FBD2DD" box="[630,686,1953,1977]" pageId="17" pageNumber="18">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C39B359FCE1F875F4F0D2DD" box="[797,933,1952,1977]" name="Puerto Rico" pageId="17" pageNumber="18">Puerto Rico</collectingCountry>
&amp; 
<collectingCountry id="D1EF764E4C39B359FC32F874F358D2DD" box="[974,1037,1952,1977]" name="Cuba" pageId="17" pageNumber="18">Cuba</collectingCountry>
. 
<typeStatus id="7643887C4C39B359FBE7F875F307D2DD" box="[1051,1106,1953,1977]" pageId="17" pageNumber="18">Type</typeStatus>
depository: ZMHB LT 
<specimenCount id="BFFEFD574C39B359FA94F875F22AD2DC" box="[1384,1407,1953,1976]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
, 7
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C39B35AFF3BF811F2CFD5B0" blockId="17.[151,1436,1915,2013]" lastBlockId="18.[151,1437,151,645]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
PLT. Distribution: 
<collectingCountry id="D1EF764E4C39B359FE5DF810F68CD2B8" box="[417,473,1988,2012]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
(FL), 
<collectingCountry id="D1EF764E4C39B359FDD6F810F53DD2B9" box="[554,616,1988,2013]" name="Cuba" pageId="17" pageNumber="18">Cuba</collectingCountry>
, 
<collectingCountry id="D1EF764E4C39B359FD8BF810F581D2B9" box="[631,724,1988,2013]" name="Jamaica" pageId="17" pageNumber="18">Jamaica</collectingCountry>
, Hispaniola, 
<collectingCountry id="D1EF764E4C39B359FC93F811F4A9D2B9" box="[879,1020,1988,2013]" name="Puerto Rico" pageId="17" pageNumber="18">Puerto Rico</collectingCountry>
, Cayman Is., St. Thomas, St. John, 
<collectingRegion id="6B3CF83C4C3AB35AFF3BFF4CF610D5CB" box="[199,325,152,175]" country="Montserrat" name="Saint Peter" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFF3BFF4CF610D5CB" box="[199,325,152,175]" name="Montserrat" pageId="18" pageNumber="19">Montserrat</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C3AB35AFEA4FF43F697D5D4" box="[344,450,151,176]" name="Bahamas" pageId="18" pageNumber="19">Bahamas</collectingCountry>
. Note: 
<typeStatus id="7643887C4C3AB35AFDDEFF43F5DCD5D4" box="[546,649,151,176]" pageId="18" pageNumber="19" type="lectotype">lectotype</typeStatus>
designation, redescription and distribution (
<bibRefCitation id="CD694B2F4C3AB35AFB5CFF4CF2DAD5D4" box="[1184,1423,151,176]" pageId="18" pageNumber="19" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 737</bibRefCitation>
); distribution, also within 
<collectingCountry id="D1EF764E4C3AB35AFE25FF68F541D5B1" box="[473,532,188,213]" name="Cuba" pageId="18" pageNumber="19">Cuba</collectingCountry>
(
<bibRefCitation id="CD694B2F4C3AB35AFDDEFF68F5CBD5B1" box="[546,670,188,213]" pageId="18" pageNumber="19" refId="ref70864" refString="Peck, S. B. (2005) A checklist of the beetles of Cuba: with data on distributions and bionomics (Insecta: Coleoptera). Arthropods of Florida and Neighboring Land Areas, 18, 1 - 241." type="journal article">Peck 2005</bibRefCitation>
); distribution within (and including) Cayman Is. (Frank 
<emphasis id="9B8CEACC4C3AB35AFAEFFF69F219D5B1" box="[1299,1356,188,213]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2011).
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFF34F511D424" blockId="18.[151,1437,151,645]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFF34F511D424" pageId="18" pageNumber="19">
= 
<taxonomicName id="6EF84D5D4C3AB35AFF53FF0BF5D2D59C" ID-CoL="LC85" authority="Bernhauer 1935: 91" authorityName="Bernhauer" authorityPageNumber="91" authorityYear="1935" box="[175,647,223,248]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gowdeyi">
<emphasis id="9B8CEACC4C3AB35AFF53FF0BF6CDD59C" box="[175,408,223,248]" italics="true" pageId="18" pageNumber="19">Belonuchus gowdeyi</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE5DFF0BF5D2D59C" author="" box="[417,647,223,248]" page="91" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE5DFF0BF50ED59C" box="[417,603,223,248]" pageId="18" pageNumber="19" refId="ref62766" refString="Bernhauer, M. (1935) Neue Staphyliniden (Col.) aus Sudamerika. Stylops, a Journal of Taxonomic Entomology, 4, 90 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00565. x" type="journal article">Bernhauer 1935</bibRefCitation>
: 91
</treatmentCitation>
</taxonomicName>
. Type locality: Hope Gardens, Jamaica. Type depository: FMNH LT, BMNH PLT?. Distribution: Jamaica. Note: lectotype designation (
<treatmentCitation id="285910CF4C3AB35AFC30FED0F3E6D478" author="" box="[972,1203,260,285]" page="739" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFC30FED0F3E6D478" box="[972,1203,260,285]" pageId="18" pageNumber="19" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 739</bibRefCitation>
</treatmentCitation>
); as synonym of 
<taxonomicName id="6EF84D5D4C3AB35AFA78FED1F515D424" ID-CoL="LC7X" authority="(Blackwelder 1943: 424)" authorityName="Blackwelder" authorityPageNumber="424" authorityYear="1943" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gagates">
<emphasis id="9B8CEACC4C3AB35AFA78FED1F64BD424" italics="true" pageId="18" pageNumber="19">B. gagates</emphasis>
(
<treatmentCitation id="285910CF4C3AB35AFED0FEF3F56CD424" author="" box="[300,569,295,320]" page="424" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFED0FEF3F56CD424" box="[300,569,295,320]" pageId="18" pageNumber="19" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943: 424</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFE99F516D4C8" blockId="18.[151,1437,151,645]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFE99F516D4C8" pageId="18" pageNumber="19">
<emphasis id="9B8CEACC4C3AB35AFF6BFE99F7FFD400" box="[151,170,333,356]" italics="true" pageId="18" pageNumber="19">=</emphasis>
<taxonomicName id="6EF84D5D4C3AB35AFF4CFE98F536D400" ID-CoL="LC98" authority="Erichson 1840: 424" authorityName="Erichson" authorityPageNumber="424" authorityYear="1840" box="[176,611,332,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="9B8CEACC4C3AB35AFF4CFE98F62AD400" box="[176,383,332,357]" italics="true" pageId="18" pageNumber="19">Belonuchus minax</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE79FE98F536D400" author="" box="[389,611,332,357]" page="424" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE79FE98F57ED400" box="[389,555,332,357]" pageId="18" pageNumber="19" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 424
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3AB35AFD91FE99F5F1D401" box="[621,676,333,357]" pageId="18" pageNumber="19">Type</typeStatus>
locality: ins. St. Thomae &amp; St. Johannis. 
<typeStatus id="7643887C4C3AB35AFB88FE99F3FED401" box="[1140,1195,333,357]" pageId="18" pageNumber="19">Type</typeStatus>
depository: ZMHB 4 ST [MCP seen]. Distribution: Lesser Antilles (St. Thomas). Note: synonym of 
<taxonomicName id="6EF84D5D4C3AB35AFBC3FEA5F7A8D4C8" ID-CoL="LC7X" authority="(Blackwelder 1943: 424)" authorityName="Blackwelder" authorityPageNumber="424" authorityYear="1943" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gagates">
<emphasis id="9B8CEACC4C3AB35AFBC3FEA5F3E3D4EC" box="[1087,1206,369,392]" italics="true" pageId="18" pageNumber="19">B. gagates</emphasis>
(
<treatmentCitation id="285910CF4C3AB35AFB39FEBBF7A3D4C8" author="" page="424" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFB39FEBBF7A3D4C8" pageId="18" pageNumber="19" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943: 424</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; not in 
<treatmentCitation id="285910CF4C3AB35AFEACFE40F56BD4C8" author="" box="[336,574,404,429]" page="737" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFEACFE40F56BD4C8" box="[336,574,404,429]" pageId="18" pageNumber="19" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana (1995: 737)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFE6CF43ED7E1" blockId="18.[151,1437,151,645]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFE6CF43ED7E1" infered="true" pageId="18" pageNumber="19">
= 
<taxonomicName id="6EF84D5D4C3AB35AFF53FE63F58BD4B4" ID-CoL="6RR2V" authority="Cameron 1913: 175" authorityName="Cameron" authorityPageNumber="175" authorityYear="1913" box="[175,734,439,464]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nigrocaeruleus">
<emphasis id="9B8CEACC4C3AB35AFF53FE63F6A5D4B4" box="[175,496,439,464]" italics="true" pageId="18" pageNumber="19">Musicoderus nigrocaeruleus</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE05FE6CF58BD4B4" author="" box="[505,734,439,464]" page="175" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE05FE6CF5F1D4B4" box="[505,676,440,464]" pageId="18" pageNumber="19" refId="ref63767" refString="Cameron, M. (1913) Description of a new species of Musicoderus (Staphylinidae) from Jamaica. Entomologist's Monthly Magazine, 49, 175." type="journal article">Cameron 1913</bibRefCitation>
: 175
</treatmentCitation>
</taxonomicName>
. Type locality: Jamaica, Newcastle district, 
<quantity id="6E009B3B4C3AB35AFB2FFE6CF271D4B4" box="[1235,1316,439,464]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="18" pageNumber="19" unit="ft" value="3000.0">3000 ft</quantity>
above sea level. Type depository: BMNH (Cameron) HT ♂. Distribution: Jamaica. Notes: as synonym of 
<taxonomicName id="6EF84D5D4C3AB35AFADEFE09F6BCD77C" ID-CoL="LC7X" authority="(Blackwelder 1943: 424)" authorityName="Blackwelder" authorityPageNumber="424" authorityYear="1943" baseAuthorityName="Blackwelder" baseAuthorityYear="1943" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gagates">
<emphasis id="9B8CEACC4C3AB35AFADEFE09F2C9D490" box="[1314,1436,477,500]" italics="true" pageId="18" pageNumber="19">B. gagates</emphasis>
(
<treatmentCitation id="285910CF4C3AB35AFF32FE2BF6B7D77C" author="" box="[206,482,511,536]" page="424" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFF32FE2BF6B7D77C" box="[206,482,511,536]" pageId="18" pageNumber="19" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943: 424</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; although both male and female characters were used in the original description, 
<bibRefCitation id="CD694B2F4C3AB35AFF3BFDF0F6D6D758" box="[199,387,548,572]" pageId="18" pageNumber="19" refId="ref63767" refString="Cameron, M. (1913) Description of a new species of Musicoderus (Staphylinidae) from Jamaica. Entomologist's Monthly Magazine, 49, 175." type="journal article">Cameron (1913)</bibRefCitation>
only refers to a “type” deposited in his collection. 
<treatmentCitation id="285910CF4C3AB35AFC10FDF0F3B5D758" author="" box="[1004,1248,548,573]" page="738" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFC10FDF0F3B5D758" box="[1004,1248,548,573]" pageId="18" pageNumber="19" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana (1995: 738)</bibRefCitation>
</treatmentCitation>
examined eight conspecific specimens under this name in the Cameron collection at the BMNH, one of them being the specimen chosen as “Type” by the author, thus the holotype.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFD67F53AD7A9" blockId="18.[151,1437,691,789]" box="[151,623,691,717]" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFD67F53AD7A9" ID-CoL="LC7Y" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[151,623,691,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="9B8CEACC4C3AB35AFF6BFD67F6E1D7A8" bold="true" box="[151,436,691,716]" italics="true" pageId="18" pageNumber="19">Belonuchus ganglbaueri</emphasis>
<bibRefCitation id="CD694B2F4C3AB35AFE47FD60F53AD7A9" box="[443,623,692,717]" pageId="18" pageNumber="19" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFD03F47FD671" blockId="18.[151,1437,691,789]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFD03F47FD671" pageId="18" pageNumber="19">
<materialsCitation id="19903C834C3AB35AFF6BFD03F4FBD794" ID-GBIF-Occurrence="1915497897" box="[151,942,727,752]" country="Bolivia" location="Bolivien" pageId="18" pageNumber="19" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFD03F5CAD794" ID-CoL="LC7Y" authority="Bernhauer 1906: 331" authorityName="Bernhauer" authorityPageNumber="331" authorityYear="1906" box="[151,671,727,752]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="9B8CEACC4C3AB35AFF6BFD03F6FCD794" box="[151,425,727,752]" italics="true" pageId="18" pageNumber="19">Belonuchus ganglbaueri</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE4CFD03F5CAD794" author="" box="[432,671,727,752]" page="331" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE4CFD03F532D794" box="[432,615,727,752]" pageId="18" pageNumber="19" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 331
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3AB35AFD55FD0CF5B5D794" box="[681,736,728,752]" pageId="18" pageNumber="19">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3AB35AFCB6FD03F4FBD794" box="[842,942,727,752]" name="Bolivia" pageId="18" pageNumber="19">Bolivien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C3AB35AFC44FD0CF47FD671" ID-GBIF-Occurrence="1915497854" collectionCode="NMW" country="Bolivia" location="ST. Distribution" pageId="18" pageNumber="19" specimenCode="FMNH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3AB35AFC44FD0CF4BAD794" box="[952,1007,728,752]" pageId="18" pageNumber="19">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3AB35AFB87FD0CF388D794" box="[1147,1245,728,752]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="18" pageNumber="19">FMNH 1</specimenCode>
ST, 
<collectionCode id="CFE9AE1B4C3AB35AFAF1FD0CF200D78B" box="[1293,1365,728,751]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="19">NMW</collectionCode>
#? 
<location id="AC2760054C3AB35AFA8AFD0CF606D671" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3AB35AFA8AFD0CF606D671" country="Bolivia" name="ST. Distribution" pageId="18" pageNumber="19">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3AB35AFEA1FD28F6E6D671" box="[349,435,764,789]" name="Bolivia" pageId="18" pageNumber="19">Bolivia</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFC97F572D639" blockId="18.[151,1436,835,1040]" box="[151,551,835,861]" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFC97F572D639" ID-CoL="LC83" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[151,551,835,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="godmani">
<emphasis id="9B8CEACC4C3AB35AFF6BFC97F6D8D638" bold="true" box="[151,397,835,860]" italics="true" pageId="18" pageNumber="19">Belonuchus godmani</emphasis>
(
<bibRefCitation id="CD694B2F4C3AB35AFE60FC90F54AD639" box="[412,543,836,861]" pageId="18" pageNumber="19" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFCB3F365D689" blockId="18.[151,1436,835,1040]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFCB3F365D689" infered="true" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFCB3F56AD6E4" ID-CoL="76ZG5" authority="Sharp 1885: 413" authorityName="Sharp" authorityPageNumber="413" authorityYear="1885" box="[151,575,871,896]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="godmani">
<emphasis id="9B8CEACC4C3AB35AFF6BFCB3F62FD6E4" box="[151,378,871,896]" italics="true" pageId="18" pageNumber="19">Philonthus godmani</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE7DFCB3F56AD6E4" author="" box="[385,575,871,896]" page="413" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE7DFCB3F553D6E4" box="[385,518,871,896]" pageId="18" pageNumber="19" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 413
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3AB35AFDB6FCBCF5D7D6E4" box="[586,642,872,896]" pageId="18" pageNumber="19">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C3AB35AFD10FCB3F413D6E4" box="[748,838,871,896]" country="Mexico" name="Mexico" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFD10FCB3F413D6E4" box="[748,838,871,896]" name="Mexico" pageId="18" pageNumber="19">Mexico</collectingCountry>
</collectingRegion>
: 
<collectingRegion id="6B3CF83C4C3AB35AFCADFCB3F49AD6E4" box="[849,975,871,896]" country="Guatemala" name="Guatemala" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFCADFCB3F49AD6E4" box="[849,975,871,896]" name="Guatemala" pageId="18" pageNumber="19">Guatemala</collectingCountry>
</collectingRegion>
: Calderas [LT]. 
<typeStatus id="7643887C4C3AB35AFB78FCBCF3EED6E4" box="[1156,1211,872,896]" pageId="18" pageNumber="19">Type</typeStatus>
depository: BMNH LT 
<specimenCount id="BFFEFD574C3AB35AFF0FFC59F65DD6C0" box="[243,264,909,932]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
, 7+ PLT. Distribution: 
<collectingRegion id="6B3CF83C4C3AB35AFDDFFC58F52ED6C1" box="[547,635,908,933]" country="Mexico" name="Mexico" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFDDFFC58F52ED6C1" box="[547,635,908,933]" name="Mexico" pageId="18" pageNumber="19">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C3AB35AFD70FC58F45DD6C1" box="[652,776,908,933]" country="Guatemala" name="Guatemala" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFD70FC58F45DD6C1" box="[652,776,908,933]" name="Guatemala" pageId="18" pageNumber="19">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C3AB35AFCE5FC58F4CED6C1" box="[793,923,908,933]" name="Costa Rica" pageId="18" pageNumber="19">
<collectingRegion id="6B3CF83C4C3AB35AFCE5FC58F40CD6C0" box="[793,857,908,932]" country="Tanzania" name="Pwani" pageId="18" pageNumber="19">Costa</collectingRegion>
Rica
</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C3AB35AFC50FC59F35DD6C0" box="[940,1032,909,932]" country="Panama" name="Panama" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFC50FC59F35DD6C0" box="[940,1032,909,932]" name="Panama" pageId="18" pageNumber="19">Panama</collectingCountry>
</collectingRegion>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C3AB35AFADFFC58F563D6AC" ID-CoL="8VRYR" authority="(Navarrete-Heredia et al. 2002)" authorityName="Navarrete-Heredia et al." authorityYear="2002" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3AB35AFADFFC58F2C9D6C1" box="[1315,1436,908,933]" italics="true" pageId="18" pageNumber="19">Philonthus</emphasis>
(
<bibRefCitation id="CD694B2F4C3AB35AFF32FC7BF57BD6AC" box="[206,558,943,968]" pageId="18" pageNumber="19" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3AB35AFE57FC65F6B2D6AC" box="[427,487,943,968]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2002
</bibRefCitation>
)
</taxonomicName>
; distribution, also within 
<collectingRegion id="6B3CF83C4C3AB35AFC9DFC7BF4E2D6AC" box="[865,951,943,968]" country="Mexico" name="Mexico" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFC9DFC7BF4E2D6AC" box="[865,951,943,968]" name="Mexico" pageId="18" pageNumber="19">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C3AB35AFC35FC7BF3B4D6AC" box="[969,1249,943,968]" pageId="18" pageNumber="19" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">Navarrete-Heredia 2002</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C3AB35AFB13FC64F2CDD6AC" box="[1263,1432,944,968]" pageId="18" pageNumber="19" refId="ref69122" refString="Marquez, J. (2004) Primeros registros estatales de especies mexicanas de Staphylininae (Coleoptera). Acta Zoologica Mexicana, 20, 91 - 97." type="journal article">Márquez 2004</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C3AB35AFF3BFC00F651D689" box="[199,260,980,1005]" pageId="18" pageNumber="19" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">2006</bibRefCitation>
); 
<typeStatus id="7643887C4C3AB35AFEEBFC00F628D689" box="[279,381,980,1005]" pageId="18" pageNumber="19" type="lectotype">lectotype</typeStatus>
designation, redescription, distribution (Márquez 2011: 206).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3AB35AFF6BFC23F465D174" box="[151,816,1015,1040]" pageId="18" pageNumber="19" type="discussion">
<paragraph id="A94736DE4C3AB35AFF6BFC23F465D174" blockId="18.[151,1436,835,1040]" box="[151,816,1015,1040]" pageId="18" pageNumber="19">
<emphasis id="9B8CEACC4C3AB35AFF6BFC23F641D174" bold="true" box="[151,276,1015,1040]" pageId="18" pageNumber="19">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C3AB35AFEB0FC23F69AD174" box="[332,463,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3AB35AFEB0FC23F69AD174" box="[332,463,1015,1040]" italics="true" pageId="18" pageNumber="19">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3AB35BFF6BFBEBF645D424" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="A94736DE4C3AB35AFF6BFBEBF569D13C" blockId="18.[151,1437,1087,1256]" box="[151,572,1087,1112]" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFBEBF569D13C" ID-CoL="LC84" authority="(Wasmann 1925)" authorityName="Wasmann" authorityYear="1925" baseAuthorityName="Wasmann" baseAuthorityYear="1925" box="[151,572,1087,1112]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="goeldii">
<emphasis id="9B8CEACC4C3AB35AFF6BFBEBF623D13C" bold="true" box="[151,374,1087,1112]" italics="true" pageId="18" pageNumber="19">Belonuchus goeldii</emphasis>
(
<bibRefCitation id="CD694B2F4C3AB35AFE78FB94F561D13C" box="[388,564,1087,1112]" pageId="18" pageNumber="19" refId="ref73915" refString="Wasmann, E. (1925) Die Ameisenmimikry. Ein exakter Beitrag zum Mimikryproblem und zur Theorie der Anpassung. Abhandlungen zur theoretischen Biologie, 19, i - xii + 1 - 164." type="journal article">Wasmann 1925</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFBB0F65AD18C" blockId="18.[151,1437,1087,1256]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFBB0F65AD18C" infered="true" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFBB0F525D118" authority="Wasmann 1925: 72" authorityName="Wasmann" authorityPageNumber="72" authorityYear="1925" box="[151,624,1124,1149]" class="Insecta" family="Staphylinidae" genus="Plociopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="goeldii">
<emphasis id="9B8CEACC4C3AB35AFF6BFBB0F6D5D119" box="[151,384,1124,1149]" italics="true" pageId="18" pageNumber="19">Plociopterus goeldii</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE77FBB1F525D118" author="" box="[395,624,1124,1149]" page="72" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE77FBB1F514D119" box="[395,577,1124,1149]" pageId="18" pageNumber="19" refId="ref73915" refString="Wasmann, E. (1925) Die Ameisenmimikry. Ein exakter Beitrag zum Mimikryproblem und zur Theorie der Anpassung. Abhandlungen zur theoretischen Biologie, 19, i - xii + 1 - 164." type="journal article">Wasmann 1925</bibRefCitation>
: 72
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3AB35AFD82FBB1F5E3D119" box="[638,694,1125,1149]" pageId="18" pageNumber="19">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C3AB35AFCD5FBB0F4D7D119" box="[809,898,1124,1149]" country="Uruguay" name="Colonia" pageId="18" pageNumber="19">Colonia</collectingRegion>
alpina, Rio d. Janeiro [LT]. 
<typeStatus id="7643887C4C3AB35AFB27FBB1F246D119" box="[1243,1299,1125,1149]" pageId="18" pageNumber="19">Type</typeStatus>
depository: NHME LT 
<specimenCount id="BFFEFD574C3AB35AFEBAFB5CF60ED1FB" box="[326,347,1160,1183]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
, #? PLT. Distribution: 
<collectingCountry id="D1EF764E4C3AB35AFDA5FB53F5F5D1C4" box="[601,672,1159,1184]" name="Brazil" pageId="18" pageNumber="19">Brazil</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C3AB35AFD57FB5CF441D1C4" box="[683,788,1160,1184]" country="Paraguay" name="Asuncion" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFD57FB5CF441D1C4" box="[683,788,1160,1184]" name="Paraguay" pageId="18" pageNumber="19">Paraguay</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C3AB35AFCE3FB53F420D1C4" box="[799,885,1159,1184]" name="Bolivia" pageId="18" pageNumber="19">Bolivia</collectingCountry>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C3AB35AFB8DFB53F255D1C4" ID-CoL="8X7QS" box="[1137,1280,1159,1184]" class="Insecta" family="Staphylinidae" genus="Plociopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3AB35AFB8DFB53F255D1C4" box="[1137,1280,1159,1184]" italics="true" pageId="18" pageNumber="19">Plociopterus</emphasis>
</taxonomicName>
and 
<typeStatus id="7643887C4C3AB35AFAC9FB53F2C9D1C4" box="[1333,1436,1159,1184]" pageId="18" pageNumber="19" type="lectotype">lectotype</typeStatus>
designation (
<treatmentCitation id="285910CF4C3AB35AFEA0FB78F414D1A1" author="" box="[348,833,1196,1221]" page="215" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFEA0FB78F414D1A1" box="[348,833,1196,1221]" pageId="18" pageNumber="19" refId="ref64817" refString="Chatzimanolis, S. &amp; Chani Posse, M. (2016) On the myrmecophilous species of Plociopterus (Coleoptera: Staphylinidae: Staphylinini) Kraatz described by E. Wasmann. The Coleopterists Bulletin, 70 (2), 214 - 216. https: // doi. org / 10.1649 / 0010 - 065 X- 70.2.214" type="journal article">Chatzimanolis &amp; Chani-Posse 2016: 215</bibRefCitation>
</treatmentCitation>
); with 
<taxonomicName id="6EF84D5D4C3AB35AFC64FB78F33BD1A0" ID-CoL="9N7M" baseAuthorityName="Smith" baseAuthorityYear="1858" box="[920,1134,1196,1220]" class="Insecta" family="Formicidae" genus="Acromyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="9B8CEACC4C3AB35AFC64FB78F33BD1A0" box="[920,1134,1196,1220]" italics="true" pageId="18" pageNumber="19">Acromyrmex niger</emphasis>
</taxonomicName>
[orig.; 
<bibRefCitation id="CD694B2F4C3AB35AFB37FB78F650D18C" pageId="18" pageNumber="19" refId="ref69955" refString="Navarrete-Heredia, J. L. (2001) Beetles associated with Atta and Acromyrmex ants (Hymenoptera: Formicidae: Attini). Transactions of the American Entomological Society, 127, 381 - 429." type="journal article">Navarrete-Heredia 2001</bibRefCitation>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFAC3F54DD054" blockId="18.[151,1436,1303,1400]" box="[151,536,1303,1328]" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFAC3F54DD054" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[151,536,1303,1328]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="9B8CEACC4C3AB35AFF6BFAC3F62BD054" bold="true" box="[151,382,1303,1328]" italics="true" pageId="18" pageNumber="19">Belonuchus gracilis</emphasis>
(
<bibRefCitation id="CD694B2F4C3AB35AFE70FAC3F55AD054" box="[396,527,1303,1328]" pageId="18" pageNumber="19" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFAE8F68DD01C" blockId="18.[151,1436,1303,1400]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFAE8F68DD01C" pageId="18" pageNumber="19">
<materialsCitation id="19903C834C3AB35AFF6BFAE8F344D031" ID-GBIF-Occurrence="1915497841" box="[151,1041,1340,1365]" country="Nicaragua" location="Chontales" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Chontales" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFAE8F509D031" ID-CoL="6RQQV" authority="Sharp 1885: 456" authorityName="Sharp" authorityPageNumber="456" authorityYear="1885" box="[151,604,1340,1365]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="9B8CEACC4C3AB35AFF6BFAE8F6DCD031" box="[151,393,1340,1365]" italics="true" pageId="18" pageNumber="19">Musicoderus gracilis</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE69FAE8F509D031" author="" box="[405,604,1340,1365]" page="456" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE69FAE8F54AD031" box="[405,543,1340,1365]" pageId="18" pageNumber="19" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 456
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3AB35AFD90FAE9F5F1D031" box="[620,676,1341,1365]" pageId="18" pageNumber="19">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3AB35AFCEBFAE8F4D8D031" box="[791,909,1340,1365]" name="Nicaragua" pageId="18" pageNumber="19">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C3AB35AFC61FAE8F344D031" box="[925,1041,1340,1365]" country="Nicaragua" name="Chontales" pageId="18" pageNumber="19">Chontales</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C3AB35AFBDDFAE9F681D01C" ID-GBIF-Occurrence="1915497906" collectionCode="BMNH" country="Nicaragua" location="Distribution" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="7643887C4C3AB35AFBDDFAE9F30DD031" box="[1057,1112,1341,1365]" pageId="18" pageNumber="19">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3AB35AFB12FAE9F217D030" box="[1262,1346,1341,1364]" pageId="18" pageNumber="19">BMNH</collectionCode>
HT 
<specimenCount id="BFFEFD574C3AB35AFA7CFAE9F2C3D030" box="[1408,1430,1341,1364]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
. 
<location id="AC2760054C3AB35AFF3BFA8BF606D01C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3AB35AFF3BFA8BF606D01C" box="[199,339,1375,1400]" country="Nicaragua" name="Distribution" pageId="18" pageNumber="19">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3AB35AFEA1FA8BF681D01C" box="[349,468,1375,1400]" name="Nicaragua" pageId="18" pageNumber="19">Nicaragua</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFA73F5E3D0A4" blockId="18.[151,1436,1447,1760]" box="[151,694,1447,1472]" pageId="18" pageNumber="19">
<emphasis id="9B8CEACC4C3AB35AFF6BFA73F7F6D0A4" bold="true" box="[151,163,1447,1472]" italics="true" pageId="18" pageNumber="19">*</emphasis>
<taxonomicName id="6EF84D5D4C3AB35AFF5EFA73F5E3D0A4" authority="(Fabricius 1801)" authorityName="Fabricius" authorityYear="1801" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[162,694,1447,1472]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="haemohrroidalis">
<emphasis id="9B8CEACC4C3AB35AFF5EFA73F6A0D0A4" bold="true" box="[162,501,1447,1472]" italics="true" pageId="18" pageNumber="19">Belonuchus haemohrroidalis</emphasis>
(
<bibRefCitation id="CD694B2F4C3AB35AFDF8FA73F5FBD0A4" box="[516,686,1447,1472]" pageId="18" pageNumber="19" refId="ref65632" refString="Fabricius, J. C. (1801) Systema eleutheratorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Vol. 2. Bibliopolii Academici Novi, Kiliae (Kiel), 687 pp." type="book">Fabricius 1801</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BFA18F6AFD384" blockId="18.[151,1436,1447,1760]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BFA18F6AFD384" infered="true" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BFA18F585D081" ID-CoL="8Y9S2" authority="Fabricius 1801: 596" authorityName="Fabricius" authorityPageNumber="596" authorityYear="1801" box="[151,720,1484,1509]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="9B8CEACC4C3AB35AFF6BFA18F689D081" box="[151,476,1484,1509]" italics="true" pageId="18" pageNumber="19">Staphylinus haemorrhoidalis</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE1AFA18F585D081" author="" box="[486,720,1484,1509]" page="596" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE1AFA18F5C1D080" box="[486,660,1484,1509]" pageId="18" pageNumber="19" refId="ref65632" refString="Fabricius, J. C. (1801) Systema eleutheratorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Vol. 2. Bibliopolii Academici Novi, Kiliae (Kiel), 687 pp." type="book">Fabricius 1801</bibRefCitation>
: 596
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3AB35AFD21FA19F440D081" box="[733,789,1485,1509]" pageId="18" pageNumber="19">Type</typeStatus>
locality: America meridionali. 
<typeStatus id="7643887C4C3AB35AFB7CFA19F3E2D081" box="[1152,1207,1485,1509]" pageId="18" pageNumber="19">Type</typeStatus>
depository: ZMUK 4ST (until recently at ZMUC, 
<bibRefCitation id="CD694B2F4C3AB35AFDD0FA3BF59DD36C" box="[556,712,1519,1544]" pageId="18" pageNumber="19" refId="ref74168" refString="Zimsen, E. (1964) The type material of I. C. Fabricius. Munksgaard, Copenhagen, 656 pp." type="book">Zimsen 1964</bibRefCitation>
): 3ST (former Sehested-Tønder Lund collection), 1ST (former Fabricius´s personal collection, seen AFN 2010). Distribution: 
<collectingCountry id="D1EF764E4C3AB35AFC5EF9C0F346D349" box="[930,1043,1556,1581]" name="Colombia" pageId="18" pageNumber="19">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3AB35AFBDEF9C0F3CDD349" box="[1058,1176,1556,1581]" name="Venezuela" pageId="18" pageNumber="19">Venezuela</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C3AB35AFB5BF9C0F206D349" box="[1191,1363,1556,1581]" country="France" name="Guyane" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFB5BF9C0F206D349" box="[1191,1363,1556,1581]" name="French Guiana" pageId="18" pageNumber="19">French Guiana</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C3AB35AFA9EF9C1F2CDD348" box="[1378,1432,1557,1580]" name="Peru" pageId="18" pageNumber="19">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3AB35AFF3BF9E3F649D334" box="[199,284,1591,1616]" name="Bolivia" pageId="18" pageNumber="19">Bolivia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3AB35AFED7F9E3F627D334" box="[299,370,1591,1616]" name="Brazil" pageId="18" pageNumber="19">Brazil</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C3AB35AFE7CF9ECF6BCD334" box="[384,489,1592,1616]" country="Paraguay" name="Asuncion" pageId="18" pageNumber="19">
<collectingCountry id="D1EF764E4C3AB35AFE7CF9ECF6BCD334" box="[384,489,1592,1616]" name="Paraguay" pageId="18" pageNumber="19">Paraguay</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C3AB35AFE04F9E3F53ED334" box="[504,619,1591,1616]" name="Argentina" pageId="18" pageNumber="19">Argentina</collectingCountry>
. Notes: comb. nov. ex 
<emphasis id="9B8CEACC4C3AB35AFC86F9E3F356D334" box="[890,1027,1591,1616]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFC86F9E3F4AAD334" ID-CoL="8VS43" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[890,1023,1591,1616]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Staphylinus</taxonomicName>
,
</emphasis>
cited as 
<emphasis id="9B8CEACC4C3AB35AFB8CF9E3F244D334" box="[1136,1297,1591,1616]" italics="true" pageId="18" pageNumber="19">haemoroidalis</emphasis>
(
<treatmentCitation id="285910CF4C3AB35AFADEF9E3F66BD310" author="" page="130" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFADEF9E3F66BD310" pageId="18" pageNumber="19" refId="ref70688" refString="Nordmann, A. (1837) Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum, 4, 1 - 167. https: // doi. org / 10.5962 / bhl. title. 144129" type="journal article">Nordmann 1837: 130</bibRefCitation>
</treatmentCitation>
); preoccupied, junior primary homonym of 
<taxonomicName id="6EF84D5D4C3AB35AFCC7F988F23FD311" ID-CoL="8Y9S2" authority="Gmelin 1790: 2036" authorityName="Gmelin" authorityPageNumber="2036" authorityYear="1790" box="[827,1386,1628,1653]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="9B8CEACC4C3AB35AFCC7F988F3D5D311" box="[827,1152,1628,1653]" italics="true" pageId="18" pageNumber="19">Staphylinus haemorrhoidalis</emphasis>
<treatmentCitation id="285910CF4C3AB35AFB76F988F23FD311" author="" box="[1162,1386,1628,1653]" page="2036" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFB76F988F271D310" box="[1162,1316,1628,1653]" pageId="18" pageNumber="19" refId="ref66804" refString="Gmelin, J. F. (1790) Caroli a Linne, … Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima tertia, aucta, reformata. Vol. 1. Part 4. G. E. Beer, Lipsiae, 708 pp. [pp. 1517 - 2224]" type="book chapter">Gmelin 1790</bibRefCitation>
: 2036
</treatmentCitation>
</taxonomicName>
and junior primary homonym of 
<taxonomicName id="6EF84D5D4C3AB35AFDF4F9ABF4B0D3FC" ID-CoL="8Y9S2" authority="Olivier 1795" authorityName="Olivier" authorityYear="1795" box="[520,997,1663,1688]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="9B8CEACC4C3AB35AFDF4F9ABF41ED3FC" box="[520,843,1663,1688]" italics="true" pageId="18" pageNumber="19">Staphylinus haemorrhoidalis</emphasis>
<bibRefCitation id="CD694B2F4C3AB35AFCAEF9ABF4B0D3FC" box="[850,997,1663,1688]" pageId="18" pageNumber="19" refId="ref70725" refString="Olivier, A. G. (1795) Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee. Coleopteres. Vol. 3. Nos. 35 - 65. Lanneau, Paris, 557 pp." type="book">Olivier 1795</bibRefCitation>
</taxonomicName>
: (42) 11; ICZN application to conserve name (
<treatmentCitation id="285910CF4C3AB35AFEEFF970F6B9D3D8" author="" box="[275,492,1700,1724]" page="42" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFEEFF970F6B9D3D8" box="[275,492,1700,1724]" pageId="18" pageNumber="19" refId="ref67257" refString="Herman, L. H. (2001 a) Nomenclatural changes in the Staphylinidae (Insecta: Coleoptera). Bulletin of the American Museum of Natural History, 264, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 (2001) 264 % 3 C 0003: NCITSI % 3 E 2.0. CO; 2" type="journal article">Herman 2001a: 42</bibRefCitation>
</treatmentCitation>
), conserved in Opinion 2053 (
<bibRefCitation id="CD694B2F4C3AB35AFCB8F970F499D3D8" box="[836,972,1700,1724]" pageId="18" pageNumber="19" refId="ref67836" refString="ICZN (2003) Opinion 2053 (Case 3207). Staphylinidae Latreille, 1804 (Insecta, Coleoptera): 65 specific names conserved. Bulletin of Zoological Nomenclature, 60 (3), 237 - 246." type="journal article">ICZN 2003</bibRefCitation>
); distribution including 
<collectingCountry id="D1EF764E4C3AB35AFB21F971F246D3D8" box="[1245,1299,1701,1724]" name="Peru" pageId="18" pageNumber="19">Peru</collectingCountry>
, also within 
<collectingCountry id="D1EF764E4C3AB35AFF3BF913F65ED384" box="[199,267,1735,1760]" name="Brazil" pageId="18" pageNumber="19">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C3AB35AFE8EF91DF6FFD384" box="[370,426,1735,1760]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BF8DBF5D7D24C" blockId="18.[151,1437,1807,1940]" box="[151,642,1807,1832]" pageId="18" pageNumber="19">
<taxonomicName id="6EF84D5D4C3AB35AFF6BF8DBF5D7D24C" ID-CoL="68GHZ" authority="Blackwelder 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,642,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hispaniolus">
<emphasis id="9B8CEACC4C3AB35AFF6BF8DBF6FBD24C" bold="true" box="[151,430,1807,1832]" italics="true" pageId="18" pageNumber="19">Belonuchus hispaniolus</emphasis>
<bibRefCitation id="CD694B2F4C3AB35AFE49F8DBF5D7D24C" box="[437,642,1807,1832]" pageId="18" pageNumber="19" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3AB35AFF6BF8E0F645D2F0" blockId="18.[151,1437,1807,1940]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="89E811F04C3AB35AFF6BF8E0F645D2F0" pageId="18" pageNumber="19">
<materialsCitation id="19903C834C3AB35AFF6BF8E0F2CDD229" ID-GBIF-Occurrence="1915497881" box="[151,1432,1844,1869]" country="Dominican Republic" location="San Francisco Mountains" pageId="18" pageNumber="19" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3AB35AFF6BF8E0F5EAD228" ID-CoL="68GHZ" authority="Blackwelder 1943: 422" authorityName="Blackwelder" authorityPageNumber="422" authorityYear="1943" box="[151,703,1844,1869]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hispaniolus">
<emphasis id="9B8CEACC4C3AB35AFF6BF8E0F6F0D229" box="[151,421,1844,1869]" italics="true" pageId="18" pageNumber="19">Belonuchus hispaniolus</emphasis>
<treatmentCitation id="285910CF4C3AB35AFE53F8E0F5EAD228" author="" box="[431,703,1844,1869]" page="422" pageId="18" pageNumber="19">
<bibRefCitation id="CD694B2F4C3AB35AFE53F8E0F5D6D228" box="[431,643,1844,1869]" pageId="18" pageNumber="19" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 422
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3AB35AFD30F8E1F456D229" box="[716,771,1845,1869]" pageId="18" pageNumber="19">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3AB35AFC88F8E0F330D229" box="[884,1125,1844,1869]" name="Dominican Republic" pageId="18" pageNumber="19">Dominican Republic</collectingCountry>
: 
<location id="AC2760054C3AB35AFB8EF8E0F2CDD229" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3AB35AFB8EF8E0F2CDD229" box="[1138,1432,1844,1869]" country="Dominican Republic" name="San Francisco Mountains" pageId="18" pageNumber="19">San Francisco Mountains</location>
</materialsCitation>
. 
<materialsCitation id="19903C834C3AB35AFF3BF88CF645D2F0" ID-GBIF-Occurrence="1915497919" collectionCode="USNM" country="Haiti" location="Distribution" pageId="18" pageNumber="19" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<typeStatus id="7643887C4C3AB35AFF3BF88CF7AAD214" box="[199,255,1880,1904]" pageId="18" pageNumber="19">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3AB35AFE6CF88CF6B4D214" box="[400,481,1880,1904]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">USNM</collectionCode>
HT 
<specimenCount id="BFFEFD574C3AB35AFDE6F88CF57AD20B" box="[538,559,1880,1903]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
, 1PT 
<specimenCount id="BFFEFD574C3AB35AFD84F88CF5DBD20B" box="[632,654,1880,1903]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
. 
<location id="AC2760054C3AB35AFD62F883F47FD214" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3AB35AFD62F883F47FD214" box="[670,810,1879,1904]" country="Haiti" name="Distribution" pageId="18" pageNumber="19">Distribution</location>
: Dominican Rep., 
<collectingCountry id="D1EF764E4C3AB35AFBFEF883F36BD214" box="[1026,1086,1879,1904]" name="Haiti" pageId="18" pageNumber="19">Haiti</collectingCountry>
. Note: 
<collectingCountry id="D1EF764E4C3AB35AFB6EF883F39FD214" box="[1170,1226,1879,1904]" name="Haiti" pageId="18" pageNumber="19">Haiti</collectingCountry>
(
<bibRefCitation id="CD694B2F4C3AB35AFB27F883F651D2F0" pageId="18" pageNumber="19" refId="ref66551" refString="Frank, J. H. &amp; Barrera, R. (2010) Natural history of Belonuchus Nordmann spp. and allies (Coleoptera: Staphylinidae) in Heliconia L. (Zingiberales: Heliconiaceae) flower bracts. Insecta Mundi, 110, 1 - 12." type="journal article">Frank &amp; Barrera 2010</bibRefCitation>
).
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFF43F576D5D4" blockId="19.[151,1436,151,357]" box="[151,547,151,176]" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFF43F576D5D4" ID-CoL="LC8C" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[151,547,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="holisoides">
<emphasis id="9B8CEACC4C3BB35BFF6BFF43F6CED5D4" bold="true" box="[151,411,151,176]" italics="true" pageId="19" pageNumber="20">Belonuchus holisoides</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFF68F645D424" blockId="19.[151,1436,151,357]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BFF68F645D424" infered="true" pageId="19" pageNumber="20">
<materialsCitation id="19903C834C3BB35BFF6BFF68F4DFD5B1" ID-GBIF-Occurrence="1915497894" box="[151,906,188,213]" country="Brazil" location="Ega" pageId="19" pageNumber="20" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFF68F50ED5B1" ID-CoL="LC8C" authority="Sharp 1876 a: 161" authorityName="Sharp" authorityPageNumber="161" authorityYear="1876" box="[151,603,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="holisoides">
<emphasis id="9B8CEACC4C3BB35BFF6BFF68F6C6D5B1" box="[151,403,188,213]" italics="true" pageId="19" pageNumber="20">Belonuchus holisoides</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE66FF68F50ED5B1" author="" box="[410,603,188,213]" page="161" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE66FF68F576D5B1" box="[410,547,188,213]" pageId="19" pageNumber="20" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp1876a</bibRefCitation>
: 161
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3BB35BFD9BFF69F5CBD5B1" box="[615,670,189,213]" pageId="19" pageNumber="20">Type</typeStatus>
locality: 
<location id="AC2760054C3BB35BFCF5FF69F462D5B1" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3BB35BFCF5FF69F462D5B1" box="[777,823,189,213]" country="Brazil" name="Ega" pageId="19" pageNumber="20">Ega</location>
, 
<collectingCountry id="D1EF764E4C3BB35BFCBEFF68F4DFD5B1" box="[834,906,188,213]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C3BB35BFC69FF69F446D478" ID-GBIF-Occurrence="1915497797" collectionCode="MCP" country="Brazil" county="Ega" location="Distribution" municipality="Metoponcus" pageId="19" pageNumber="20" specimenCode="BMNH 2" specimenCount="4" stateProvince="Amazonas" typeStatus="holotype">
<typeStatus id="7643887C4C3BB35BFC69FF69F499D5B1" box="[917,972,189,213]" pageId="19" pageNumber="20">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3BB35BFBA5FF69F396D5B0" box="[1113,1219,188,212]" collectionCode="BMNH" pageId="19" pageNumber="20">BMNH 2</specimenCode>
ST: 1 
<collectingCounty id="40264E524C3BB35BFAF2FF69F26FD5B1" box="[1294,1338,189,213]" pageId="19" pageNumber="20">Ega</collectingCounty>
together with other 
<specimenCount id="BFFEFD574C3BB35BFEBDFF34F69ED59C" box="[321,459,223,248]" pageId="19" pageNumber="20" type="generic">4 specimens</specimenCount>
from probably a different species, all of them under the name &quot; 
<collectingMunicipality id="4923ACA44C3BB35BFB6AFF34F276D59C" box="[1174,1315,224,248]" pageId="19" pageNumber="20">Metoponcus</collectingMunicipality>
<taxonomicName id="6EF84D5D4C3BB35BFAD6FF0BF65ED479" ID-CoL="LC8C" authority="Sharp" authorityName="Sharp" authorityYear="1876" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="holisoides">holisoides Sharp</taxonomicName>
&quot;[
<collectionCode id="CFE9AE1B4C3BB35BFEE5FED0F602D479" box="[281,343,260,285]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="19" pageNumber="20">MCP</collectionCode>
seen]. 
<location id="AC2760054C3BB35BFE5BFED0F566D479" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3BB35BFE5BFED0F566D479" box="[423,563,260,285]" country="Brazil" county="Ega" municipality="Metoponcus" name="Distribution" pageId="19" pageNumber="20" stateProvince="Amazonas">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3BB35BFDC0FED0F5D5D479" box="[572,640,260,285]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C3BB35BFD73FED0F45ED478" box="[655,779,260,284]" country="Venezuela" name="Amazonas" pageId="19" pageNumber="20">Amazonas</collectingRegion>
)
</materialsCitation>
, Argentina. Note: distribution only in Brazil (Asenjo 
<emphasis id="9B8CEACC4C3BB35BFA98FED1F2C9D479" box="[1380,1436,260,285]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3BB35BFF6BFE98F5B5D400" box="[151,736,332,357]" pageId="19" pageNumber="20" type="discussion">
<paragraph id="A94736DE4C3BB35BFF6BFE98F5B5D400" blockId="19.[151,1436,151,357]" box="[151,736,332,357]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BFE98F641D401" bold="true" box="[151,276,332,357]" pageId="19" pageNumber="20">Comment.</emphasis>
It probably belongs to 
<taxonomicName id="6EF84D5D4C3BB35BFDE4FE98F5B5D400" authority="Erichson." authorityName="Erichson." box="[536,736,332,357]" class="Insecta" family="Staphylinidae" genus="Holisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3BB35BFDE4FE98F53BD401" box="[536,622,332,357]" italics="true" pageId="19" pageNumber="20">Holisus</emphasis>
Erichson.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3BB35BFF6BFE47F6FAD16B" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="A94736DE4C3BB35BFF6BFE47F407D4C9" blockId="19.[151,1437,403,536]" box="[151,850,403,429]" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFE47F5CFD4C8" authority="Plavilstschikov 1929" authorityName="Plavilstschikov" authorityYear="1929" box="[151,666,403,429]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="iconnicoffi">
<emphasis id="9B8CEACC4C3BB35BFF6BFE47F6F0D4C8" bold="true" box="[151,421,403,428]" italics="true" pageId="19" pageNumber="20">Belonuchus iconnicoffi</emphasis>
<bibRefCitation id="CD694B2F4C3BB35BFE51FE40F5CFD4C8" box="[429,666,404,429]" pageId="19" pageNumber="20" refId="ref71306" refString="Plavilstschikov, N. N. (1929) Les Staphylinides recoltes par Mr. Nicolas Iconnicov dans le Perou (Coleoptera, Staphylinidae). Entomologisches Nachrichtsblatt, Troppau, 3 (1), 29 - 30." type="journal article">Plavilstschikov 1929</bibRefCitation>
</taxonomicName>
, 
<emphasis id="9B8CEACC4C3BB35BFD59FE40F407D4C9" bold="true" box="[677,850,404,429]" pageId="19" pageNumber="20">nomen nudum</emphasis>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFE63F6D4D77C" blockId="19.[151,1437,403,536]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BFE63F6D4D77C" infered="true" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFE63F5ECD4B4" authority="Plavilstschikov 1929: 30" authorityName="Plavilstschikov" authorityPageNumber="30" authorityYear="1929" box="[151,697,439,464]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="iconnicoffi">
<emphasis id="9B8CEACC4C3BB35BFF6BFE63F6CCD4B4" box="[151,409,439,464]" italics="true" pageId="19" pageNumber="20">Belonuchus iconnicoffi</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE5CFE63F5ECD4B4" author="" box="[416,697,439,464]" page="30" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE5CFE63F5DAD4B4" box="[416,655,439,464]" pageId="19" pageNumber="20" refId="ref71306" refString="Plavilstschikov, N. N. (1929) Les Staphylinides recoltes par Mr. Nicolas Iconnicov dans le Perou (Coleoptera, Staphylinidae). Entomologisches Nachrichtsblatt, Troppau, 3 (1), 29 - 30." type="journal article">Plavilstschikov 1929</bibRefCitation>
: 30
</treatmentCitation>
</taxonomicName>
, as &quot;n. sp. (Bernhauer 
<emphasis id="9B8CEACC4C3BB35BFC46FE6DF375D4B4" box="[954,1056,439,464]" italics="true" pageId="19" pageNumber="20">in litteris</emphasis>
)&quot;; nomen nudum, no description. Distribution: 
<collectingCountry id="D1EF764E4C3BB35BFE9DFE09F6CDD490" box="[353,408,477,500]" name="Peru" pageId="19" pageNumber="20">Peru</collectingCountry>
. Note: listed in error as valid species in 
<bibRefCitation id="CD694B2F4C3BB35BFC81FE08F335D490" box="[893,1120,476,501]" pageId="19" pageNumber="20" refId="ref63204" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum, 185, 1 - 188." type="journal article">Blackwelder (1944)</bibRefCitation>
and 
<bibRefCitation id="CD694B2F4C3BB35BFB63FE08F212D490" box="[1183,1351,476,501]" pageId="19" pageNumber="20" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo (2004)</bibRefCitation>
; not in 
<bibRefCitation id="CD694B2F4C3BB35BFF3BFDD4F6D4D77C" box="[199,385,511,536]" pageId="19" pageNumber="20" refId="ref67417" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article">Herman (2001d)</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFD93F544D704" blockId="19.[151,1436,583,717]" box="[151,529,583,608]" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFD93F544D704" ID-CoL="LC8D" authority="(Solsky 1872)" authorityName="Solsky" authorityYear="1872" baseAuthorityName="Solsky" baseAuthorityYear="1872" box="[151,529,583,608]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ielskii">
<emphasis id="9B8CEACC4C3BB35BFF6BFD93F639D704" bold="true" box="[151,364,583,608]" italics="true" pageId="19" pageNumber="20">Belonuchus ielskii</emphasis>
(
<bibRefCitation id="CD694B2F4C3BB35BFE87FD93F55CD704" box="[379,521,583,608]" pageId="19" pageNumber="20" refId="ref73125" refString="Solsky, S. (1872) Enumeration et description des Coleopteres de la famille des Staphylinides recueillis par Mrs. C. Ielsky et le Baron de Nolken pendant leurs voyages dans l'Amerique du Sud en 1870 et 1871. Premiere partie. Horae Societatis Entomologicae Rossicae, 8, 289 - 314." type="journal article">Solsky 1872</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFDB8F307D7A9" blockId="19.[151,1436,583,717]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BFDB8F307D7A9" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFDB8F517D7E1" ID-CoL="76ZKD" authority="Solsky 1872: 296" authorityName="Solsky" authorityPageNumber="296" authorityYear="1872" box="[151,578,620,645]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ielskii">
<emphasis id="9B8CEACC4C3BB35BFF6BFDB8F635D7E1" box="[151,352,620,645]" italics="true" pageId="19" pageNumber="20">Philonthus ielskii</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE92FDB8F517D7E1" author="" box="[366,578,620,645]" page="296" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE92FDB8F551D7E0" box="[366,516,620,645]" pageId="19" pageNumber="20" refId="ref73125" refString="Solsky, S. (1872) Enumeration et description des Coleopteres de la famille des Staphylinides recueillis par Mrs. C. Ielsky et le Baron de Nolken pendant leurs voyages dans l'Amerique du Sud en 1870 et 1871. Premiere partie. Horae Societatis Entomologicae Rossicae, 8, 289 - 314." type="journal article">Solsky 1872</bibRefCitation>
: 296
</treatmentCitation>
</taxonomicName>
. Type locality: Puna, Lima [Amérique du Sud]. Type depository: ZIN? (Solsky) #?ST. Distribution: Peru. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C3BB35BFCA7FD5BF220D7CC" ID-CoL="8VRYR" authority="(Bernhauer &amp; Schubert 1914: 370)" authorityName="Bernhauer &amp; Schubert" authorityPageNumber="370" authorityYear="1914" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[859,1397,655,680]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3BB35BFCA7FD5BF481D7CC" box="[859,980,655,680]" italics="true" pageId="19" pageNumber="20">Philonthus</emphasis>
(
<treatmentCitation id="285910CF4C3BB35BFC1AFD5BF238D7CC" author="" box="[998,1389,655,680]" page="370" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFC1AFD5BF238D7CC" box="[998,1389,655,680]" pageId="19" pageNumber="20" refId="ref62964" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus. Vol. 57. W. Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer &amp; Schubert 1914: 370</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; as 
<emphasis id="9B8CEACC4C3BB35BFF3BFD60F65FD7A9" box="[199,266,692,717]" italics="true" pageId="19" pageNumber="20">jelskii</emphasis>
(
<treatmentCitation id="285910CF4C3BB35BFEE4FD60F55ED7A9" author="" box="[280,523,692,717]" page="2527" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFEE4FD60F693D7A9" box="[280,454,692,717]" pageId="19" pageNumber="20" refId="ref67417" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article">Herman 2001d</bibRefCitation>
: 2527
</treatmentCitation>
, 
<treatmentCitation id="285910CF4C3BB35BFDEAFD60F587D7A9" author="" box="[534,722,692,717]" page="61" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFDEAFD60F5FCD7A8" box="[534,681,692,717]" pageId="19" pageNumber="20" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
: 61
</treatmentCitation>
). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFD2FF50ED671" blockId="19.[151,1436,763,861]" box="[151,603,763,789]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BFD2FF7F6D670" bold="true" box="[151,163,763,788]" italics="true" pageId="19" pageNumber="20">*</emphasis>
<taxonomicName id="6EF84D5D4C3BB35BFF5EFD2FF50ED671" ID-CoL="LC8G" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[162,603,763,789]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="impressifrons">
<emphasis id="9B8CEACC4C3BB35BFF5EFD2FF686D670" bold="true" box="[162,467,763,788]" italics="true" pageId="19" pageNumber="20">Belonuchus impressifrons</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFCCBF37ED638" blockId="19.[151,1436,763,861]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BFCCBF37ED638" pageId="19" pageNumber="20">
<materialsCitation id="19903C834C3BB35BFF6BFCCBF687D639" ID-GBIF-Occurrence="1915497885" collectionCode="BMNH" country="Brazil" location="Amazonas" municipality="Distribution" pageId="19" pageNumber="20" specimenCount="2" stateProvince="Sao Paulo" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFCCBF5DDD65C" ID-CoL="LC8G" authority="Sharp 1876 a: 162" authorityName="Sharp" authorityPageNumber="162" authorityYear="1876" box="[151,648,799,824]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="impressifrons">
<emphasis id="9B8CEACC4C3BB35BFF6BFCCBF6E2D65C" box="[151,439,799,824]" italics="true" pageId="19" pageNumber="20">Belonuchus impressifrons</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE42FCCBF5DDD65C" author="" box="[446,648,799,824]" page="162" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE42FCCBF505D65C" box="[446,592,799,824]" pageId="19" pageNumber="20" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 162
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3BB35BFD6FFCF4F59FD65C" box="[659,714,800,824]" pageId="19" pageNumber="20">Type</typeStatus>
locality: Ega. 
<typeStatus id="7643887C4C3BB35BFC93FCF4F4F2D65C" box="[879,935,800,824]" pageId="19" pageNumber="20">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3BB35BFBC8FCF4F3DDD653" box="[1076,1160,800,823]" pageId="19" pageNumber="20">BMNH</collectionCode>
HT. 
<collectingMunicipality id="4923ACA44C3BB35BFB3DFCCBF218D65C" box="[1217,1357,799,824]" pageId="19" pageNumber="20">Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C3BB35BFAABFCCBF2C9D65C" box="[1367,1436,799,824]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
(
<location id="AC2760054C3BB35BFF32FC90F61CD638" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3BB35BFF32FC90F61CD638" box="[206,329,836,860]" country="Brazil" municipality="Distribution" name="Amazonas" pageId="19" pageNumber="20" stateProvince="Sao Paulo">Amazonas</location>
, 
<collectingRegion id="6B3CF83C4C3BB35BFEA8FC90F69FD639" box="[340,458,836,861]" country="Brazil" name="Sao Paulo" pageId="19" pageNumber="20">São Paulo</collectingRegion>
)
</materialsCitation>
. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C3BB35BFC5FFC91F489D639" box="[931,988,836,861]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFC5FF5E2D6C1" blockId="19.[151,1437,907,1544]" box="[151,695,907,933]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BFC5FF7F6D6C0" bold="true" box="[151,163,907,932]" italics="true" pageId="19" pageNumber="20">*</emphasis>
<taxonomicName id="6EF84D5D4C3BB35BFF5EFC5FF57ED6C1" ID-CoL="LC8H" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,555,907,933]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus" status="comb. nov.">
<emphasis id="9B8CEACC4C3BB35BFF5EFC5FF6DAD6C0" bold="true" box="[162,399,907,932]" italics="true" pageId="19" pageNumber="20">Belonuchus infimus</emphasis>
(
<bibRefCitation id="CD694B2F4C3BB35BFE61FC58F577D6C1" box="[413,546,908,933]" pageId="19" pageNumber="20" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C3BB35BFDCBFC58F5E2D6C1" box="[567,695,908,933]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFDCBFC58F5E2D6C1" bold="true" box="[567,695,908,933]" pageId="19" pageNumber="20">comb. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BFC7BF6FAD16B" blockId="19.[151,1437,907,1544]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BFC7BF6FAD16B" infered="true" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BFC7BF566D6AC" ID-CoL="76YZF" authority="Sharp 1885: 414" authorityName="Sharp" authorityPageNumber="414" authorityYear="1885" box="[151,563,943,968]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus">
<emphasis id="9B8CEACC4C3BB35BFF6BFC7BF63ED6AC" box="[151,363,943,968]" italics="true" pageId="19" pageNumber="20">Philonthus infimus</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE88FC7BF566D6AC" author="" box="[372,563,943,968]" page="414" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE88FC7BF6ACD6AC" box="[372,505,943,968]" pageId="19" pageNumber="20" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 414
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3BB35BFDC3FC64F522D6AC" box="[575,631,944,968]" pageId="19" pageNumber="20">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C3BB35BFD1FFC7BF40AD6AC" box="[739,863,943,968]" country="Guatemala" name="Guatemala" pageId="19" pageNumber="20">
<collectingCountry id="D1EF764E4C3BB35BFD1FFC7BF40AD6AC" box="[739,863,943,968]" name="Guatemala" pageId="19" pageNumber="20">Guatemala</collectingCountry>
</collectingRegion>
, Las Mercedes 
<quantity id="6E009B3B4C3BB35BFBEEFC64F333D6AC" box="[1042,1126,943,968]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="19" pageNumber="20" unit="ft" value="3000.0">3000 ft</quantity>
, Chiacam; 
<collectingRegion id="6B3CF83C4C3BB35BFB1BFC64F217D6A3" box="[1255,1346,944,967]" country="Panama" name="Panama" pageId="19" pageNumber="20">
<collectingCountry id="D1EF764E4C3BB35BFB1BFC64F217D6A3" box="[1255,1346,944,967]" name="Panama" pageId="19" pageNumber="20">Panama</collectingCountry>
</collectingRegion>
, Volcan de 
<collectingRegion id="6B3CF83C4C3BB35BFF14FC00F61CD689" box="[232,329,980,1005]" country="Panama" name="Chiriqui" pageId="19" pageNumber="20">Chiriqui</collectingRegion>
. 
<typeStatus id="7643887C4C3BB35BFEA8FC01F6DED689" box="[340,395,981,1005]" pageId="19" pageNumber="20">Type</typeStatus>
depository: BMNH 8 ST: 5 Las Mercedes, 2 
<collectingRegion id="6B3CF83C4C3BB35BFC70FC00F4B9D689" box="[908,1004,980,1005]" country="Panama" name="Chiriqui" pageId="19" pageNumber="20">Chiriqui</collectingRegion>
, 1 Chiacam [MCP seen]. Distribution: 
<collectingRegion id="6B3CF83C4C3BB35BFF3BFC23F616D174" box="[199,323,1015,1040]" country="Guatemala" name="Guatemala" pageId="19" pageNumber="20">
<collectingCountry id="D1EF764E4C3BB35BFF3BFC23F616D174" box="[199,323,1015,1040]" name="Guatemala" pageId="19" pageNumber="20">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C3BB35BFEB2FC2CF6FFD16B" box="[334,426,1016,1039]" country="Panama" name="Panama" pageId="19" pageNumber="20">
<collectingCountry id="D1EF764E4C3BB35BFEB2FC2CF6FFD16B" box="[334,426,1016,1039]" name="Panama" pageId="19" pageNumber="20">Panama</collectingCountry>
</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3BB35BFF6BFBC8F3CDD1C4" pageId="19" pageNumber="20" type="discussion">
<paragraph id="A94736DE4C3BB35BFF6BFBC8F3CDD1C4" blockId="19.[151,1437,907,1544]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BFBC8F65ED151" bold="true" box="[151,267,1052,1077]" pageId="19" pageNumber="20">Remarks.</emphasis>
Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on more than two specimens, including male/s and female/s. Eight specimens were located in the BMNH under this name. The specimens below agree with the original description by 
<bibRefCitation id="CD694B2F4C3BB35BFAA7FBB0F644D1C4" pageId="19" pageNumber="20" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp (1885)</bibRefCitation>
and considered to have been 
<typeStatus id="7643887C4C3BB35BFDA1FB5CF5EAD1C4" box="[605,703,1160,1184]" pageId="19" pageNumber="20" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C3BB35BFC5DFB53F352D1C4" box="[929,1031,1159,1184]" pageId="19" pageNumber="20" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3BB35BFF6BFB78F549D081" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="A94736DE4C3BB35BFF6BFB78F549D081" blockId="19.[151,1437,907,1544]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BFB78F69FD1A1" bold="true" box="[151,458,1196,1221]" pageId="19" pageNumber="20">Type material examined.</emphasis>
Lectotype ♂, here designated (BMNH): card-mounted (with additional label “Sp. figured”), “ 
<taxonomicName id="6EF84D5D4C3BB35BFEB9FB1BF5EBD18C" authority="Las Mercedes" authorityName="Las Mercedes" box="[325,702,1231,1256]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus">Philonthus infimus Las Mercedes</taxonomicName>
<quantity id="6E009B3B4C3BB35BFD38FB04F442D18C" box="[708,791,1231,1256]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="19" pageNumber="20" unit="ft" value="3000.0">3000 ft</quantity>
. Guatemala Champion. [in Sharp’s hand] // Las Mercedes 
<quantity id="6E009B3B4C3BB35BFF3BFB20F64ED069" box="[199,283,1268,1293]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="19" pageNumber="20" unit="ft" value="3000.0">3000 ft</quantity>
. Champion. (white label) // B. C. A. Col. 
<date id="DD46101E4C3BB35BFD01FB21F47ED068" box="[765,811,1268,1292]" pageId="19" pageNumber="20">I. 2.</date>
<taxonomicName id="6EF84D5D4C3BB35BFCC8FB20F337D069" authority=", Sharp." authorityName="Sharp." box="[820,1122,1268,1293]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus">Philonthus infimus, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 2 ♂, 2 ♀, card-mounted (one ♀ with additional “Type” round white label with red margin and “Type D.S” on the mounting card [in Sharp’s hand]), “ 
<taxonomicName id="6EF84D5D4C3BB35BFADEFAE8F694D01C" authority="Las Mercedes" authorityName="Las Mercedes" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus">Philonthus infimus Las Mercedes</taxonomicName>
<quantity id="6E009B3B4C3BB35BFE34FAB4F549D01C" box="[456,540,1375,1400]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="19" pageNumber="20" unit="ft" value="3000.0">3000 ft</quantity>
. Guat. Champion. [in Sharp’s hand] // Las Mercedes 
<quantity id="6E009B3B4C3BB35BFB8FFAB4F393D01C" box="[1139,1222,1375,1400]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="19" pageNumber="20" unit="ft" value="3000.0">3000 ft</quantity>
. Champion. (white label) // B. C. A. Col. 
<date id="DD46101E4C3BB35BFE3FFA51F6A5D0F8" box="[451,496,1412,1436]" pageId="19" pageNumber="20">I. 2.</date>
<taxonomicName id="6EF84D5D4C3BB35BFE04FA50F470D0F9" authority=", Sharp." authorityName="Sharp." box="[504,805,1412,1437]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus">Philonthus infimus, Sharp.</taxonomicName>
(white label)”; 2 ♀, card-mounted, “ 
<taxonomicName id="6EF84D5D4C3BB35BFB38FA50F2C9D0F9" box="[1220,1436,1412,1437]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus">Philonthus infimus</taxonomicName>
D.S. [in Sharp’s hand] // V. de Chiriqui 
<quantity id="6E009B3B4C3BB35BFD66FA7CF45DD0A4" box="[666,776,1447,1472]" metricMagnitude="2" metricUnit="m" metricValue="4.578096" metricValueMax="9.144" metricValueMin="0.012192" pageId="19" pageNumber="20" unit="ft" value="1502.0" valueMax="3000.0" valueMin="4.0">4–3000 ft</quantity>
Champion (white label) // B. C. A. Col. 
<date id="DD46101E4C3BB35BFB17FA7CF24FD0A4" box="[1259,1306,1448,1472]" pageId="19" pageNumber="20">I. 2.</date>
<taxonomicName id="6EF84D5D4C3BB35BFAD8FA73F626D081" authority=", Sharp." authorityName="Sharp." class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="infimus">Philonthus infimus, Sharp.</taxonomicName>
(white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3BB35BFF6BFA3BF33DD36C" box="[151,1128,1519,1544]" pageId="19" pageNumber="20" type="discussion">
<paragraph id="A94736DE4C3BB35BFF6BFA3BF33DD36C" blockId="19.[151,1437,907,1544]" box="[151,1128,1519,1544]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BFA3BF64AD36C" bold="true" box="[151,287,1519,1544]" pageId="19" pageNumber="20">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C3BB35BFD46FA3BF331D36C" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3BB35BFD46FA3BF468D36C" box="[698,829,1519,1544]" italics="true" pageId="19" pageNumber="20">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C3BB35BFCB7FA3BF309D36C" box="[843,1116,1519,1544]" pageId="19" pageNumber="20" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C3BB35BFC1CFA25F34DD36C" box="[992,1048,1519,1544]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3BB35BFF6BF9E3F3F4D2B8" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="A94736DE4C3BB35BFF6BF9E3F569D334" blockId="19.[151,1436,1591,1725]" box="[151,572,1591,1616]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BF9E3F7F6D334" bold="true" box="[151,163,1591,1616]" italics="true" pageId="19" pageNumber="20">*</emphasis>
<taxonomicName id="6EF84D5D4C3BB35BFF5EF9E3F569D334" ID-CoL="LC8J" authority="Solsky 1868" authorityName="Solsky" authorityYear="1868" box="[162,572,1591,1616]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="iridescens">
<emphasis id="9B8CEACC4C3BB35BFF5EF9E3F6FCD334" bold="true" box="[162,425,1591,1616]" italics="true" pageId="19" pageNumber="20">Belonuchus iridescens</emphasis>
<bibRefCitation id="CD694B2F4C3BB35BFE4CF9E3F569D334" box="[432,572,1591,1616]" pageId="19" pageNumber="20" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BF988F666D3D8" blockId="19.[151,1436,1591,1725]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BF988F666D3D8" pageId="19" pageNumber="20">
<materialsCitation id="19903C834C3BB35BFF6BF988F201D311" ID-GBIF-Occurrence="1915497856" box="[151,1364,1628,1653]" collectorName="de l'Etat d'Oaxaca" country="Mexico" location="Mexique" pageId="19" pageNumber="20" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3BB35BFF6BF988F523D310" ID-CoL="LC8J" authority="Solsky 1868: 140" authorityName="Solsky" authorityPageNumber="140" authorityYear="1868" box="[151,630,1628,1653]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="iridescens">
<emphasis id="9B8CEACC4C3BB35BFF6BF988F6CDD311" box="[151,408,1628,1653]" italics="true" pageId="19" pageNumber="20">Belonuchus iridescens</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE58F988F523D310" author="" box="[420,630,1628,1653]" page="140" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE58F988F56CD311" box="[420,569,1628,1653]" pageId="19" pageNumber="20" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
: 140
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3BB35BFD7AF989F5EBD311" box="[646,702,1629,1653]" pageId="19" pageNumber="20">Type</typeStatus>
locality: [
<collectingCountry id="D1EF764E4C3BB35BFCC5F988F4F5D311" box="[825,928,1628,1653]" name="Mexico" pageId="19" pageNumber="20">Mexique</collectingCountry>
et probablement 
<collectorName id="040D53084C3BB35BFB8FF988F21BD311" box="[1139,1358,1628,1653]" pageId="19" pageNumber="20">de l'Etat d'Oaxaca</collectorName>
]
</materialsCitation>
. 
<materialsCitation id="19903C834C3BB35BFA98F989F472D3FC" ID-GBIF-Occurrence="1915497908" collectionCode="ZIN" country="Mexico" location="Solsky" municipality="ST. Distribution" pageId="19" pageNumber="20" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3BB35BFA98F989F2C9D311" box="[1380,1436,1629,1653]" pageId="19" pageNumber="20">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3BB35BFEB7F954F629D3FC" box="[331,380,1664,1688]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="19" pageNumber="20">ZIN</collectionCode>
? (
<location id="AC2760054C3BB35BFE69F9ABF6B1D3FC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3BB35BFE69F9ABF6B1D3FC" box="[405,484,1663,1688]" country="Mexico" municipality="ST. Distribution" name="Solsky" pageId="19" pageNumber="20">Solsky</location>
) #? 
<collectingMunicipality id="4923ACA44C3BB35BFDF1F954F590D3FC" box="[525,709,1663,1688]" pageId="19" pageNumber="20">ST. Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C3BB35BFD32F9ABF472D3FC" box="[718,807,1663,1688]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C3BB35BFB53F9ABF67DD3D8" pageId="19" pageNumber="20" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3BB35BFA74F955F7B1D3D9" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BF93FF50BD261" blockId="19.[151,1436,1771,1869]" box="[151,606,1771,1797]" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BF93FF50BD261" ID-CoL="LC8K" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,606,1771,1797]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="irregularis">
<emphasis id="9B8CEACC4C3BB35BFF6BF93FF6F6D260" bold="true" box="[151,419,1771,1796]" italics="true" pageId="19" pageNumber="20">Belonuchus irregularis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BF8DBF208D229" blockId="19.[151,1436,1771,1869]" pageId="19" pageNumber="20">
<materialsCitation id="19903C834C3BB35BFF6BF8DBF208D229" ID-GBIF-Occurrence="1915497864" collectionCode="FMNH" country="Colombia" elevation="2000" location="Distribution" pageId="19" pageNumber="20" specimenCount="2" typeStatus="holotype">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BF8DBF208D229" pageId="19" pageNumber="20">
<taxonomicName id="6EF84D5D4C3BB35BFF6BF8DBF5FED24C" ID-CoL="LC8K" authority="Bernhauer 1916 c: 275" authorityName="Bernhauer" authorityPageNumber="275" authorityYear="1916" box="[151,683,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="irregularis">
<emphasis id="9B8CEACC4C3BB35BFF6BF8DBF6C9D24C" box="[151,412,1807,1832]" italics="true" pageId="19" pageNumber="20">Belonuchus irregularis</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE5BF8DBF5FED24C" author="" box="[423,683,1807,1832]" page="275" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE5BF8DBF53AD24C" box="[423,623,1807,1832]" pageId="19" pageNumber="20" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 275
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3BB35BFD44F8C4F5A5D24C" box="[696,752,1808,1832]" pageId="19" pageNumber="20">Type</typeStatus>
locality: West-Columbien: Pacho, 
<elevation id="22D5D1ED4C3BB35BFB7AF8C4F382D24C" box="[1158,1239,1808,1832]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="2000.0">
<quantity id="6E009B3B4C3BB35BFB7AF8C4F382D24C" box="[1158,1239,1808,1832]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="2000.0">2000m</quantity>
</elevation>
, Ost-Cordilleren, West-Columbien. 
<typeStatus id="7643887C4C3BB35BFE6EF8E1F69CD229" box="[402,457,1845,1869]" pageId="19" pageNumber="20">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3BB35BFDA9F8E1F5F3D228" box="[597,678,1845,1868]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="19" pageNumber="20">FMNH</collectionCode>
HT. 
<location id="AC2760054C3BB35BFD22F8E0F43FD229" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3BB35BFD22F8E0F43FD229" box="[734,874,1844,1869]" country="Colombia" name="Distribution" pageId="19" pageNumber="20">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3BB35BFC89F8E0F4B2D229" box="[885,999,1844,1869]" name="Colombia" pageId="19" pageNumber="20">Colombia</collectingCountry>
. Note: generic affiliation unclear.
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BF8AFF5E0D2F1" blockId="19.[151,1436,1915,2013]" box="[151,693,1915,1941]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF6BF8AFF7F6D2F0" bold="true" box="[151,163,1915,1940]" italics="true" pageId="19" pageNumber="20">*</emphasis>
<taxonomicName id="6EF84D5D4C3BB35BFF5EF8AFF57FD2F0" ID-CoL="4GB2F" authority="(Sharp 1887)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[162,554,1915,1941]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="iteratus" status="comb. nov.">
<emphasis id="9B8CEACC4C3BB35BFF5EF8AFF6C8D2F1" bold="true" box="[162,413,1915,1941]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFF5EF8AFF6D8D2F0" bold="true" box="[162,397,1915,1940]" italics="true" pageId="19" pageNumber="20">Belonuchus iteratus</emphasis>
(
</emphasis>
<bibRefCitation id="CD694B2F4C3BB35BFE62F8A8F577D2F0" box="[414,546,1916,1941]" pageId="19" pageNumber="20" refId="ref72534" refString="Sharp, D. (1887) Introduction, Supplement, Index. In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. v - x + 748 - 802 + 803 - 824. [http: // www. sil. si. edu / digitalcollections / bca /]" type="book chapter">Sharp 1887</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C3BB35BFDC9F8A8F5E0D2F1" box="[565,693,1916,1941]" pageId="19" pageNumber="20">
<emphasis id="9B8CEACC4C3BB35BFDC9F8A8F5E0D2F1" bold="true" box="[565,693,1916,1941]" pageId="19" pageNumber="20">comb. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="A94736DE4C3BB35BFF6BF874F3F4D2B8" blockId="19.[151,1436,1915,2013]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="89E811F04C3BB35BFF6BF874F3F4D2B8" pageId="19" pageNumber="20">
<materialsCitation id="19903C834C3BB35BFF6BF874F6E3D2B9" ID-GBIF-Occurrence="1915497836" collectionCode="BMNH" country="Mexico" location="ST. Distribution" pageId="19" pageNumber="20" specimenCode="BMNH 23" specimenCount="2" stateProvince="Durango" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3BB35BFF6BF874F513D2DC" ID-CoL="4GB2F" authority="Sharp 1887: 788" authorityName="Sharp" authorityPageNumber="788" authorityYear="1887" box="[151,582,1952,1977]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="iteratus">
<emphasis id="9B8CEACC4C3BB35BFF6BF874F627D2DC" box="[151,370,1952,1977]" italics="true" pageId="19" pageNumber="20">Philonthus iteratus</emphasis>
<treatmentCitation id="285910CF4C3BB35BFE83F874F513D2DC" author="" box="[383,582,1952,1977]" page="788" pageId="19" pageNumber="20">
<bibRefCitation id="CD694B2F4C3BB35BFE83F874F55CD2DC" box="[383,521,1952,1977]" pageId="19" pageNumber="20" refId="ref72534" refString="Sharp, D. (1887) Introduction, Supplement, Index. In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. v - x + 748 - 802 + 803 - 824. [http: // www. sil. si. edu / digitalcollections / bca /]" type="book chapter">Sharp 1887</bibRefCitation>
: 788
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3BB35BFDABF875F5DBD2DD" box="[599,654,1953,1977]" pageId="19" pageNumber="20">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C3BB35BFCF8F875F433D2DD" box="[772,870,1953,1977]" country="Mexico" name="Durango" pageId="19" pageNumber="20">Durango</collectingRegion>
(Villa Lerdo). 
<typeStatus id="7643887C4C3BB35BFBE2F875F303D2DD" box="[1054,1110,1953,1977]" pageId="19" pageNumber="20">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3BB35BFB11F875F23FD2DC" box="[1261,1386,1952,1976]" collectionCode="BMNH" pageId="19" pageNumber="20">BMNH 23</specimenCode>
<location id="AC2760054C3BB35BFA8AF874F606D2B9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3BB35BFA8AF874F606D2B9" country="Mexico" name="ST. Distribution" pageId="19" pageNumber="20" stateProvince="Durango">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3BB35BFEA1F810F6E3D2B9" box="[349,438,1988,2013]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C3BB35BFCBCF810F3C0D2B8" box="[832,1173,1988,2013]" pageId="19" pageNumber="20" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3BB35BFBE5F811F304D2B9" box="[1049,1105,1988,2013]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BFF43F527D400" pageId="20" pageNumber="21" type="discussion">
<paragraph id="A94736DE4C3CB35CFF6BFF43F52FD593" blockId="20.[151,1436,151,285]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFF43F65ED5D4" bold="true" box="[151,267,151,176]" pageId="20" pageNumber="21">Remarks.</emphasis>
Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on several specimens, including male/s and female/s. Twenty-three specimens were located in the BMNH under this name.
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFED0F33DD478" blockId="20.[151,1436,151,285]" box="[151,1128,260,285]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFED0F64AD479" bold="true" box="[151,287,260,285]" pageId="20" pageNumber="21">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C3CB35CFD46FED0F331D478" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,260,285]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3CB35CFD46FED0F468D479" box="[698,829,260,285]" italics="true" pageId="20" pageNumber="21">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C3CB35CFCB7FED0F309D478" box="[843,1116,260,285]" pageId="20" pageNumber="21" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C3CB35CFC1CFED1F34DD479" box="[992,1048,260,285]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFE9FF527D400" blockId="20.[151,1436,331,464]" box="[151,626,331,357]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFE9FF7F6D400" bold="true" box="[151,163,331,356]" italics="true" pageId="20" pageNumber="21">*</emphasis>
<taxonomicName id="6EF84D5D4C3CB35CFF5EFE9FF527D400" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[162,626,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="jalappensis">
<emphasis id="9B8CEACC4C3CB35CFF5EFE9FF6E2D400" bold="true" box="[162,439,331,356]" italics="true" pageId="20" pageNumber="21">Belonuchus jalappensis</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BFEBBF3AFD794" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="A94736DE4C3CB35CFF6BFEBBF645D4B4" blockId="20.[151,1436,331,464]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="89E811F04C3CB35CFF6BFEBBF645D4B4" pageId="20" pageNumber="21">
<materialsCitation id="19903C834C3CB35CFF6BFEBBF21AD4E3" ID-GBIF-Occurrence="1915497924" box="[151,1359,367,392]" country="Mexico" location="Jalappa" municipality="Xalapa" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3CB35CFF6BFEBBF59BD4EC" ID-CoL="LC8N" authority="Bernhauer 1917 d: 224" authorityName="Bernhauer" authorityPageNumber="224" authorityYear="1917" box="[151,718,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="jalappensis">
<emphasis id="9B8CEACC4C3CB35CFF6BFEBBF6FED4EC" box="[151,427,367,392]" italics="true" pageId="20" pageNumber="21">Belonuchus jalappensis</emphasis>
<treatmentCitation id="285910CF4C3CB35CFE40FEBBF59BD4EC" author="" box="[444,718,367,392]" page="224" pageId="20" pageNumber="21">
<bibRefCitation id="CD694B2F4C3CB35CFE40FEBBF5D9D4EC" box="[444,652,367,392]" pageId="20" pageNumber="21" refId="ref62307" refString="Bernhauer, M. (1917 d) Vier neue Belonuchus aus Mexiko. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 67, 223 - 224." type="journal article">Bernhauer 1917d</bibRefCitation>
: 224
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3CB35CFD18FEA4F449D4EC" box="[740,796,368,392]" pageId="20" pageNumber="21">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3CB35CFC66FEBBF4A1D4EC" box="[922,1012,367,392]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
: 
<location id="AC2760054C3CB35CFBF5FEBBF308D4EC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFBF5FEBBF308D4EC" box="[1033,1117,367,392]" country="Mexico" municipality="Xalapa" name="Jalappa" pageId="20" pageNumber="21" stateProvince="Veracruz">Jalappa</location>
[
<collectingMunicipality id="4923ACA44C3CB35CFB89FEBBF39FD4EC" box="[1141,1226,367,392]" pageId="20" pageNumber="21">Xalapa</collectingMunicipality>
, 
<collectingRegion id="6B3CF83C4C3CB35CFB23FEA4F21CD4E3" box="[1247,1353,368,391]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="20" pageNumber="21">Veracruz</collectingRegion>
]
</materialsCitation>
. 
<materialsCitation id="19903C834C3CB35CFA98FEA4F5B2D4C9" ID-GBIF-Occurrence="1915497863" collectionCode="FMNH" country="Mexico" location="ST. Distribution" pageId="20" pageNumber="21" specimenCode="FMNH 2" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3CB35CFA98FEA4F2C9D4EC" box="[1380,1436,368,392]" pageId="20" pageNumber="21">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3CB35CFEB3FE41F6ECD4C8" box="[335,441,404,428]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21">FMNH 2</specimenCode>
<location id="AC2760054C3CB35CFE38FE40F5D5D4C9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFE38FE40F5D5D4C9" box="[452,640,404,429]" country="Mexico" name="ST. Distribution" pageId="20" pageNumber="21">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3CB35CFD72FE40F5B2D4C9" box="[654,743,404,429]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C3CB35CFB7FFE40F651D4B4" pageId="20" pageNumber="21" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3CB35CFA9CFE41F2C9D4C9" box="[1376,1436,404,429]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFE2BF530D77C" blockId="20.[151,1437,511,608]" box="[151,613,511,536]" pageId="20" pageNumber="21">
<taxonomicName id="6EF84D5D4C3CB35CFF6BFE2BF530D77C" ID-CoL="LC8R" authority="Bernhauer 1942" authorityName="Bernhauer" authorityYear="1942" box="[151,613,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="9B8CEACC4C3CB35CFF6BFE2BF6FED77C" bold="true" box="[151,427,511,536]" italics="true" pageId="20" pageNumber="21">Belonuchus laevipennis</emphasis>
<bibRefCitation id="CD694B2F4C3CB35CFE4EFE2BF530D77C" box="[434,613,511,536]" pageId="20" pageNumber="21" refId="ref62877" refString="Bernhauer, M. (1942) Neue Staphyliniden aus Kostarika. Zoologischer Anzeiger, 138, 1 - 27." type="journal article">Bernhauer 1942</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFDF0F4B1D704" blockId="20.[151,1437,511,608]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="89E811F04C3CB35CFF6BFDF0F4B1D704" pageId="20" pageNumber="21">
<materialsCitation id="19903C834C3CB35CFF6BFDF0F381D759" ID-GBIF-Occurrence="1915497910" box="[151,1236,548,573]" country="Costa Rica" elevation="1000" location="Pino" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Cartago" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3CB35CFF6BFDF0F5DFD759" ID-CoL="LC8R" authority="Bernhauer 1942: 25" authorityName="Bernhauer" authorityPageNumber="25" authorityYear="1942" box="[151,650,548,573]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="9B8CEACC4C3CB35CFF6BFDF0F6F4D759" box="[151,417,548,573]" italics="true" pageId="20" pageNumber="21">Belonuchus laevipennis</emphasis>
<treatmentCitation id="285910CF4C3CB35CFE54FDF0F5DFD759" author="" box="[424,650,548,573]" page="25" pageId="20" pageNumber="21">
<bibRefCitation id="CD694B2F4C3CB35CFE54FDF0F535D758" box="[424,608,548,573]" pageId="20" pageNumber="21" refId="ref62877" refString="Bernhauer, M. (1942) Neue Staphyliniden aus Kostarika. Zoologischer Anzeiger, 138, 1 - 27." type="journal article">Bernhauer 1942</bibRefCitation>
: 25
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3CB35CFD6AFDF1F59BD759" box="[662,718,549,573]" pageId="20" pageNumber="21">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C3CB35CFCC4FDF0F4C4D758" box="[824,913,548,572]" country="Costa Rica" name="Cartago" pageId="20" pageNumber="21">Cartago</collectingRegion>
(
<location id="AC2760054C3CB35CFC5DFDF0F48CD759" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFC5DFDF0F48CD759" box="[929,985,548,573]" country="Costa Rica" name="Pino" pageId="20" pageNumber="21" stateProvince="Cartago">Pino</location>
), 
<elevation id="22D5D1ED4C3CB35CFC11FDF0F36ED758" box="[1005,1083,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1000.0">
<quantity id="6E009B3B4C3CB35CFC11FDF0F36ED758" box="[1005,1083,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="m" value="1000.0">1000m</quantity>
</elevation>
[
<collectingCountry id="D1EF764E4C3CB35CFBB5FDF0F399D759" box="[1097,1228,548,573]" name="Costa Rica" pageId="20" pageNumber="21">Costa Rica</collectingCountry>
]
</materialsCitation>
. 
<materialsCitation id="19903C834C3CB35CFB23FDF1F4B1D704" ID-GBIF-Occurrence="1915497903" collectionCode="FMNH" country="Costa Rica" location="ST. Distribution" pageId="20" pageNumber="21" specimenCode="FMNH 3" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3CB35CFB23FDF1F242D759" box="[1247,1303,549,573]" pageId="20" pageNumber="21">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3CB35CFF3BFD9CF679D704" box="[199,300,584,608]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21">FMNH 3</specimenCode>
<location id="AC2760054C3CB35CFED7FD9CF6B0D704" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFED7FD9CF6B0D704" box="[299,485,583,608]" country="Costa Rica" name="ST. Distribution" pageId="20" pageNumber="21">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3CB35CFE13FD9CF538D704" box="[495,621,583,608]" name="Costa Rica" pageId="20" pageNumber="21">Costa Rica</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFD5BF43ED7CC" blockId="20.[151,1437,655,1149]" box="[151,875,655,680]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFD5BF7F6D7CC" bold="true" box="[151,163,655,680]" italics="true" pageId="20" pageNumber="21">*</emphasis>
<taxonomicName id="6EF84D5D4C3CB35CFF5EFD5BF51AD7CC" ID-CoL="4GB53" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,591,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latecinctus" status="comb. nov.">
<emphasis id="9B8CEACC4C3CB35CFF5EFD5BF6E7D7CC" bold="true" box="[162,434,655,680]" italics="true" pageId="20" pageNumber="21">Belonuchus latecinctus</emphasis>
(
<bibRefCitation id="CD694B2F4C3CB35CFE3DFD5BF513D7CC" box="[449,582,655,680]" pageId="20" pageNumber="21" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C3CB35CFDA7FD5BF58ED7CC" box="[603,731,655,680]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFDA7FD5BF58ED7CC" bold="true" box="[603,731,655,680]" pageId="20" pageNumber="21">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C3CB35CFD16FD5BF436D7CC" box="[746,867,655,680]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="20" pageNumber="21">Figs. 9–10</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFD60F3AFD794" blockId="20.[151,1437,655,1149]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="89E811F04C3CB35CFF6BFD60F3AFD794" pageId="20" pageNumber="21">
<materialsCitation id="19903C834C3CB35CFF6BFD60F3A3D794" ID-GBIF-Occurrence="1915497891" collectionCode="MCP" country="Guatemala" county="Distribution" elevation="1372" location="Cerro Zunil" municipality="San Geronimo" pageId="20" pageNumber="21" specimenCode="BMNH 3" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3CB35CFF6BFD60F506D7A9" ID-CoL="4GB53" authority="Sharp 1885: 415" authorityName="Sharp" authorityPageNumber="415" authorityYear="1885" box="[151,595,692,717]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latecinctus">
<emphasis id="9B8CEACC4C3CB35CFF6BFD60F6C5D7A9" box="[151,400,692,717]" italics="true" pageId="20" pageNumber="21">Philonthus latecinctus</emphasis>
<treatmentCitation id="285910CF4C3CB35CFE6BFD60F506D7A9" author="" box="[407,595,692,717]" page="415" pageId="20" pageNumber="21">
<bibRefCitation id="CD694B2F4C3CB35CFE6BFD60F54ED7A9" box="[407,539,692,717]" pageId="20" pageNumber="21" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 415
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3CB35CFDA2FD61F5C3D7A9" box="[606,662,693,717]" pageId="20" pageNumber="21">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3CB35CFD03FD60F42ED7A9" box="[767,891,692,717]" name="Guatemala" pageId="20" pageNumber="21">Guatemala</collectingCountry>
, Cerro Zunil 
<elevation id="22D5D1ED4C3CB35CFBEDFD60F3FCD7A9" box="[1041,1193,692,717]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" metricValueMax="1.524" metricValueMin="1.2192" pageId="20" pageNumber="21" unit="ft" value="4500.0" valueMax="5000.0" valueMin="4000.0">
<quantity id="6E009B3B4C3CB35CFBEDFD60F3FCD7A9" box="[1041,1193,692,717]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" metricValueMax="1.524" metricValueMin="1.2192" pageId="20" pageNumber="21" unit="ft" value="4500.0" valueMax="5000.0" valueMin="4000.0">4000–5000 ft</quantity>
</elevation>
, San Gerónimo. 
<typeStatus id="7643887C4C3CB35CFA98FD61F2C9D7A9" box="[1380,1436,693,717]" pageId="20" pageNumber="21">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3CB35CFEB0FD0CF6E0D794" box="[332,437,728,752]" collectionCode="BMNH" pageId="20" pageNumber="21">BMNH 3</specimenCode>
ST: 2 
<location id="AC2760054C3CB35CFE03FD0CF5D0D794" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFE03FD0CF5D0D794" box="[511,645,727,752]" country="Guatemala" county="Distribution" municipality="San Geronimo" name="Cerro Zunil" pageId="20" pageNumber="21">Cerro Zunil</location>
, 1 
<collectingMunicipality id="4923ACA44C3CB35CFD59FD0CF41DD794" box="[677,840,727,752]" pageId="20" pageNumber="21">San Gerónimo</collectingMunicipality>
[
<collectionCode id="CFE9AE1B4C3CB35CFCA9FD0CF4C6D794" box="[853,915,728,752]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="20" pageNumber="21">MCP</collectionCode>
seen]. 
<collectingCounty id="40264E524C3CB35CFC1FFD03F33AD794" box="[995,1135,727,752]" pageId="20" pageNumber="21">Distribution</collectingCounty>
: 
<collectingCountry id="D1EF764E4C3CB35CFB85FD03F3A3D794" box="[1145,1270,727,752]" name="Guatemala" pageId="20" pageNumber="21">Guatemala</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BFD28F5CDD639" pageId="20" pageNumber="21" type="discussion">
<paragraph id="A94736DE4C3CB35CFF6BFD28F5CDD639" blockId="20.[151,1437,655,1149]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFD28F65ED671" bold="true" box="[151,267,764,789]" pageId="20" pageNumber="21">Remarks.</emphasis>
Sharp based his description on three specimens which were located in the BMNH under this name. The specimens below agree with the original description by 
<bibRefCitation id="CD694B2F4C3CB35CFCB7FCCBF4B5D65C" box="[843,992,799,824]" pageId="20" pageNumber="21" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp (1885)</bibRefCitation>
and considered to have been 
<typeStatus id="7643887C4C3CB35CFAC6FCF4F2C9D65C" box="[1338,1436,800,824]" pageId="20" pageNumber="21" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C3CB35CFE5EFC90F55CD639" box="[418,521,836,861]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BFCB3F215D13C" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="A94736DE4C3CB35CFF6BFCB3F215D13C" blockId="20.[151,1437,655,1149]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFCB3F685D6E4" bold="true" box="[151,464,871,896]" pageId="20" pageNumber="21">Type material examined.</emphasis>
Lectotype ♂, here designated (BMNH, 
<figureCitation id="31C32A5B4C3CB35CFC3AFCB3F31CD6E4" box="[966,1097,871,896]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="20" pageNumber="21">Figs. 9–10</figureCitation>
): card-mounted, “ 
<taxonomicName id="6EF84D5D4C3CB35CFADEFCB3F66AD6C1" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latecinctus">Philonthus latecinctus</taxonomicName>
Type D.S. Cerro Zunil 
<quantity id="6E009B3B4C3CB35CFDB6FC58F5EFD6C1" box="[586,698,908,933]" metricMagnitude="2" metricUnit="m" metricValue="9.150096" metricValueMax="18.288" metricValueMin="0.012192" pageId="20" pageNumber="21" unit="ft" value="3002.0" valueMax="6000.0" valueMin="4.0">4–6000 ft</quantity>
. Guatem. Champion. [in Sharp’s hand] // Type (round white label with red margin) // Cerro Zunil, Guatemala. Champion. (white label) // B. C. A. Col. 
<date id="DD46101E4C3CB35CFB18FC64F242D6AC" box="[1252,1303,944,968]" pageId="20" pageNumber="21">I. 2.</date>
<taxonomicName id="6EF84D5D4C3CB35CFAD8FC7BF6CDD689" authority=", Sharp." authorityName="Sharp." class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latecinctus">Philonthus latecinctus, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”. Paralectotypes (BMNH): 1 ♂ with same locality and collection data as lectotype and additional printed white label “Sp. Figured”; 1 ♀, cardmounted, “ 
<taxonomicName id="6EF84D5D4C3CB35CFEB7FBC8F51AD151" box="[331,591,1052,1077]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latecinctus">Philonthus latecinctus</taxonomicName>
D.S. San Gerónimo Champion. [in Sharp’s hand] //San Gerónimo, Guatemala. Champion. (white label) // B. C. A. Col. 
<date id="DD46101E4C3CB35CFCEBFB94F416D13C" box="[791,835,1088,1112]" pageId="20" pageNumber="21">I. 2.</date>
<taxonomicName id="6EF84D5D4C3CB35CFCB6FBEBF3CDD13C" authority=", Sharp." authorityName="Sharp." box="[842,1176,1087,1112]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latecinctus">Philonthus latecinctus, Sharp.</taxonomicName>
(white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BFBB0F33DD118" box="[151,1128,1124,1149]" pageId="20" pageNumber="21" type="discussion">
<paragraph id="A94736DE4C3CB35CFF6BFBB0F33DD118" blockId="20.[151,1437,655,1149]" box="[151,1128,1124,1149]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFBB0F64AD119" bold="true" box="[151,287,1124,1149]" pageId="20" pageNumber="21">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C3CB35CFD46FBB0F331D118" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1124,1149]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3CB35CFD46FBB0F468D119" box="[698,829,1124,1149]" italics="true" pageId="20" pageNumber="21">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C3CB35CFCB7FBB0F309D118" box="[843,1116,1124,1149]" pageId="20" pageNumber="21" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C3CB35CFC1CFBB1F34DD119" box="[992,1048,1124,1149]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BFB7FF51CD32B" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="A94736DE4C3CB35CFF6BFB7FF543D1A0" blockId="20.[151,1435,1195,1328]" box="[151,534,1195,1221]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFB7FF7F6D1A0" bold="true" box="[151,163,1195,1220]" italics="true" pageId="20" pageNumber="21">*</emphasis>
<taxonomicName id="6EF84D5D4C3CB35CFF5EFB7FF543D1A0" ID-CoL="68GHY" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[162,534,1195,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="9B8CEACC4C3CB35CFF5EFB7FF639D1A0" bold="true" box="[162,364,1195,1220]" italics="true" pageId="20" pageNumber="21">Belonuchus latro</emphasis>
<bibRefCitation id="CD694B2F4C3CB35CFE88FB78F543D1A0" box="[372,534,1196,1221]" pageId="20" pageNumber="21" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFB1BF6FBD054" blockId="20.[151,1435,1195,1328]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="89E811F04C3CB35CFF6BFB1BF6FBD054" infered="true" pageId="20" pageNumber="21">
<materialsCitation id="19903C834C3CB35CFF6BFB1BF5C5D069" ID-GBIF-Occurrence="1915497900" collectionCode="MCP" country="Brazil" location="Distribution" pageId="20" pageNumber="21" specimenCode="ZMHB 1" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3CB35CFF6BFB1BF568D18C" ID-CoL="68GHY" authority="Erichson 1840: 422" authorityName="Erichson" authorityPageNumber="422" authorityYear="1840" box="[151,573,1231,1256]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="9B8CEACC4C3CB35CFF6BFB1BF603D18C" box="[151,342,1231,1256]" italics="true" pageId="20" pageNumber="21">Belonuchus latro</emphasis>
<treatmentCitation id="285910CF4C3CB35CFEA1FB1BF568D18C" author="" box="[349,573,1231,1256]" page="422" pageId="20" pageNumber="21">
<bibRefCitation id="CD694B2F4C3CB35CFEA1FB1BF551D18C" box="[349,516,1231,1256]" pageId="20" pageNumber="21" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 422
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3CB35CFDB4FB04F5D5D18C" box="[584,640,1232,1256]" pageId="20" pageNumber="21">Type</typeStatus>
locality: [no specific locality]. 
<typeStatus id="7643887C4C3CB35CFC1DFB04F34CD18C" box="[993,1049,1232,1256]" pageId="20" pageNumber="21">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3CB35CFB59FB04F259D18C" box="[1189,1292,1232,1256]" collectionCode="ZMHB" pageId="20" pageNumber="21">ZMHB 1</specimenCode>
ST 
<specimenCount id="BFFEFD574C3CB35CFAC7FB04F205D183" box="[1339,1360,1232,1255]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
[
<collectionCode id="CFE9AE1B4C3CB35CFAA1FB04F2CED18C" box="[1373,1435,1232,1256]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="20" pageNumber="21">MCP</collectionCode>
seen]. 
<location id="AC2760054C3CB35CFEEAFB20F6F7D069" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFEEAFB20F6F7D069" box="[278,418,1268,1293]" country="Brazil" name="Distribution" pageId="20" pageNumber="21" stateProvince="Sao Paulo">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3CB35CFE4EFB20F6A3D069" box="[434,502,1268,1293]" name="Brazil" pageId="20" pageNumber="21">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C3CB35CFDF7FB20F5DDD069" box="[523,648,1268,1293]" country="Brazil" name="Sao Paulo" pageId="20" pageNumber="21">São Paulo</collectingRegion>
)
</materialsCitation>
; 
<materialsCitation id="19903C834C3CB35CFD5EFB20F6FBD054" ID-GBIF-Occurrence="1915497915" country="Brazil" location="South" pageId="20" pageNumber="21" specimenCount="1" stateProvince="South" typeStatus="holotype">
<collectingRegion id="6B3CF83C4C3CB35CFD5EFB20F5B6D069" box="[674,739,1268,1293]" country="Tanzania" name="Kusini Unguja" pageId="20" pageNumber="21">South</collectingRegion>
America? Note: distribution within (and including) 
<collectingCountry id="D1EF764E4C3CB35CFAABFB20F2CED069" box="[1367,1435,1268,1293]" name="Brazil" pageId="20" pageNumber="21">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C3CB35CFEDAFACDF60AD054" box="[294,351,1303,1328]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2013).
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFA8BF505D01C" blockId="20.[151,1436,1375,1472]" box="[151,592,1375,1400]" pageId="20" pageNumber="21">
<taxonomicName id="6EF84D5D4C3CB35CFF6BFA8BF505D01C" ID-CoL="LC8X" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[151,592,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="9B8CEACC4C3CB35CFF6BFA8BF6C0D01C" bold="true" box="[151,405,1375,1400]" italics="true" pageId="20" pageNumber="21">Belonuchus longiceps</emphasis>
<bibRefCitation id="CD694B2F4C3CB35CFE60FA8BF505D01C" box="[412,592,1375,1400]" pageId="20" pageNumber="21" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFA50F497D0A4" blockId="20.[151,1436,1375,1472]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="89E811F04C3CB35CFF6BFA50F497D0A4" pageId="20" pageNumber="21">
<taxonomicName id="6EF84D5D4C3CB35CFF6BFA50F5D2D0F8" ID-CoL="LC8X" authority="Bernhauer 1906: 332" authorityName="Bernhauer" authorityPageNumber="332" authorityYear="1906" box="[151,647,1412,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="9B8CEACC4C3CB35CFF6BFA50F6D8D0F9" box="[151,397,1412,1437]" italics="true" pageId="20" pageNumber="21">Belonuchus longiceps</emphasis>
<treatmentCitation id="285910CF4C3CB35CFE69FA50F5D2D0F8" author="" box="[405,647,1412,1437]" page="332" pageId="20" pageNumber="21">
<bibRefCitation id="CD694B2F4C3CB35CFE69FA50F51BD0F9" box="[405,590,1412,1437]" pageId="20" pageNumber="21" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 332
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3CB35CFD6FFA51F59ED0F9" box="[659,715,1413,1437]" pageId="20" pageNumber="21">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3CB35CFCCBFA51F43BD0F8" box="[823,878,1413,1436]" name="Peru" pageId="20" pageNumber="21">Peru</collectingCountry>
. 
<typeStatus id="7643887C4C3CB35CFC85FA51F4E4D0F9" box="[889,945,1413,1437]" pageId="20" pageNumber="21">Type</typeStatus>
depository: FMNH HT. Distribution: 
<collectingCountry id="D1EF764E4C3CB35CFA9EFA51F2CDD0F8" box="[1378,1432,1413,1436]" name="Peru" pageId="20" pageNumber="21">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3CB35CFF3BFA73F648D0A4" box="[199,285,1447,1472]" name="Bolivia" pageId="20" pageNumber="21">Bolivia</collectingCountry>
. Note: 
<collectingCountry id="D1EF764E4C3CB35CFE91FA7CF6F5D0DB" box="[365,416,1448,1471]" name="Peru" pageId="20" pageNumber="21">Peru</collectingCountry>
(
<bibRefCitation id="CD694B2F4C3CB35CFE52FA73F516D0A4" box="[430,579,1447,1472]" pageId="20" pageNumber="21" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BFA3BF5E1D36C" blockId="20.[151,1437,1519,1832]" box="[151,692,1519,1544]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BFA3BF7F6D36C" bold="true" box="[151,163,1519,1544]" italics="true" pageId="20" pageNumber="21">*</emphasis>
<taxonomicName id="6EF84D5D4C3CB35CFF5EFA3BF57DD36C" ID-CoL="LC8Y" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,552,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lucilius" status="comb. nov.">
<emphasis id="9B8CEACC4C3CB35CFF5EFA3BF6DED36C" bold="true" box="[162,395,1519,1544]" italics="true" pageId="20" pageNumber="21">Belonuchus lucilius</emphasis>
(
<bibRefCitation id="CD694B2F4C3CB35CFE66FA3BF54AD36C" box="[410,543,1519,1544]" pageId="20" pageNumber="21" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C3CB35CFDC8FA3BF5E1D36C" box="[564,692,1519,1544]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFDC8FA3BF5E1D36C" bold="true" box="[564,692,1519,1544]" pageId="20" pageNumber="21">comb. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BF9C0F51CD32B" blockId="20.[151,1437,1519,1832]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="89E811F04C3CB35CFF6BF9C0F51CD32B" pageId="20" pageNumber="21">
<materialsCitation id="19903C834C3CB35CFF6BF9C0F385D349" ID-GBIF-Occurrence="1915497868" box="[151,1232,1556,1581]" country="Panama" elevation="762" location="Chiriqui" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3CB35CFF6BF9C0F563D349" ID-CoL="4GB7C" authority="Sharp 1885: 415" authorityName="Sharp" authorityPageNumber="415" authorityYear="1885" box="[151,566,1556,1581]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lucilius">
<emphasis id="9B8CEACC4C3CB35CFF6BF9C0F639D349" box="[151,364,1556,1581]" italics="true" pageId="20" pageNumber="21">Philonthus lucilius</emphasis>
<treatmentCitation id="285910CF4C3CB35CFE89F9C0F563D349" author="" box="[373,566,1556,1581]" page="415" pageId="20" pageNumber="21">
<bibRefCitation id="CD694B2F4C3CB35CFE89F9C0F6A9D349" box="[373,508,1556,1581]" pageId="20" pageNumber="21" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 415
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3CB35CFDB8F9C1F529D349" box="[580,636,1557,1581]" pageId="20" pageNumber="21">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3CB35CFD16F9C1F410D348" box="[746,837,1557,1580]" name="Panama" pageId="20" pageNumber="21">Panama</collectingCountry>
, Volcan de 
<collectingRegion id="6B3CF83C4C3CB35CFC32F9C0F37ED349" box="[974,1067,1556,1581]" country="Panama" name="Chiriqui" pageId="20" pageNumber="21">Chiriqui</collectingRegion>
<elevation id="22D5D1ED4C3CB35CFBC8F9C0F385D349" box="[1076,1232,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="7.62" metricValueMax="9.144" metricValueMin="6.096" pageId="20" pageNumber="21" unit="ft" value="2500.0" valueMax="3000.0" valueMin="2000.0">
<quantity id="6E009B3B4C3CB35CFBC8F9C0F385D349" box="[1076,1232,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="7.62" metricValueMax="9.144" metricValueMin="6.096" pageId="20" pageNumber="21" unit="ft" value="2500.0" valueMax="3000.0" valueMin="2000.0">2000–3000 ft</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C3CB35CFB21F9C1F510D32B" ID-GBIF-Occurrence="1915497898" collectionCode="BMNH" country="Panama" location="Distribution" pageId="20" pageNumber="21" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3CB35CFB21F9C1F240D349" box="[1245,1301,1557,1581]" pageId="20" pageNumber="21">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3CB35CFF3BF9ECF64ED32B" box="[199,283,1592,1615]" pageId="20" pageNumber="21">BMNH</collectionCode>
HT. 
<location id="AC2760054C3CB35CFEAFF9E3F68AD334" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFEAFF9E3F68AD334" box="[339,479,1591,1616]" country="Panama" name="Distribution" pageId="20" pageNumber="21">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3CB35CFE15F9ECF510D32B" box="[489,581,1592,1615]" name="Panama" pageId="20" pageNumber="21">Panama</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BF988F48BD311" box="[151,990,1628,1653]" pageId="20" pageNumber="21" type="discussion">
<paragraph id="A94736DE4C3CB35CFF6BF988F48BD311" blockId="20.[151,1437,1519,1832]" box="[151,990,1628,1653]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BF988F65ED311" bold="true" box="[151,267,1628,1653]" pageId="20" pageNumber="21">Remarks.</emphasis>
The “unique” specimen in BMNH is confirmed as the 
<typeStatus id="7643887C4C3CB35CFC88F988F48FD311" box="[884,986,1628,1653]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BF9ABF63CD261" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="A94736DE4C3CB35CFF6BF9ABF63CD261" blockId="20.[151,1437,1519,1832]" pageId="20" pageNumber="21">
<materialsCitation id="19903C834C3CB35CFF6BF9ABF64ED3D9" ID-GBIF-Occurrence="1915497880" collectionCode="BMNH" collectorName="Type D. S. V." elevation="458" location="Chiriqui" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="holotype">
<emphasis id="9B8CEACC4C3CB35CFF6BF9ABF694D3FC" bold="true" box="[151,449,1663,1688]" pageId="20" pageNumber="21">
<typeStatus id="7643887C4C3CB35CFF6BF9ABF787D3FC" box="[151,210,1663,1688]" pageId="20" pageNumber="21">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C3CB35CFE35F9ABF564D3FC" box="[457,561,1663,1688]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
<specimenCount id="BFFEFD574C3CB35CFDC5F954F51BD3F3" box="[569,590,1664,1687]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
(
<collectionCode id="CFE9AE1B4C3CB35CFDA2F954F5ECD3FC" box="[606,697,1664,1688]" pageId="20" pageNumber="21">BMNH</collectionCode>
): card-mounted, “ 
<taxonomicName id="6EF84D5D4C3CB35CFC7EF9ABF302D3FC" box="[898,1111,1663,1688]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lucilius">Philonthus lucilius</taxonomicName>
<collectorName id="040D53084C3CB35CFBA3F954F3A4D3F3" box="[1119,1265,1664,1688]" pageId="20" pageNumber="21">
<typeStatus id="7643887C4C3CB35CFBA3F954F3C2D3FC" box="[1119,1175,1664,1688]" pageId="20" pageNumber="21">Type</typeStatus>
D.S. V.
</collectorName>
de 
<collectingRegion id="6B3CF83C4C3CB35CFAE7F9ABF22DD3FC" box="[1307,1400,1663,1688]" country="Panama" name="Chiriqui" pageId="20" pageNumber="21">Chiriqui</collectingRegion>
<elevation id="22D5D1ED4C3CB35CFA7CF954F64ED3D9" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" pageId="20" pageNumber="21" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">
<quantity id="6E009B3B4C3CB35CFA7CF954F64ED3D9" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" pageId="20" pageNumber="21" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">2– 3000 ft</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C3CB35CFEDBF970F2CDD3D9" ID-GBIF-Occurrence="1915497916" box="[295,1432,1700,1725]" elevation="458" location="Type" municipality="Sharp's" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
Champion. [in 
<collectingMunicipality id="4923ACA44C3CB35CFE2EF970F573D3D9" box="[466,550,1700,1725]" pageId="20" pageNumber="21">Sharp’s</collectingMunicipality>
hand] // 
<location id="AC2760054C3CB35CFD71F971F591D3D9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFD71F971F591D3D9" box="[653,708,1701,1725]" municipality="Sharp's" name="Type" pageId="20" pageNumber="21" stateProvince="Chiriqui">
<typeStatus id="7643887C4C3CB35CFD71F971F591D3D9" box="[653,708,1701,1725]" pageId="20" pageNumber="21">Type</typeStatus>
</location>
(round white label with red margin) // V. de 
<collectingRegion id="6B3CF83C4C3CB35CFB3FF970F275D3D9" box="[1219,1312,1700,1725]" country="Panama" name="Chiriqui" pageId="20" pageNumber="21">Chiriqui</collectingRegion>
<elevation id="22D5D1ED4C3CB35CFAD4F970F2CDD3D9" box="[1320,1432,1700,1725]" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" pageId="20" pageNumber="21" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">
<quantity id="6E009B3B4C3CB35CFAD4F970F2CDD3D9" box="[1320,1432,1700,1725]" metricMagnitude="2" metricUnit="m" metricValue="4.575048" metricValueMax="9.144" metricValueMin="0.006096" pageId="20" pageNumber="21" unit="ft" value="1501.0" valueMax="3000.0" valueMin="2.0">2–3000 ft</quantity>
</elevation>
</materialsCitation>
. Champion. (white label) // B. C. A. Col. 
<date id="DD46101E4C3CB35CFD73F91CF5EED384" box="[655,699,1736,1760]" pageId="20" pageNumber="21">I. 2.</date>
<taxonomicName id="6EF84D5D4C3CB35CFD3EF913F4BFD384" authority=", Sharp." authorityName="Sharp." box="[706,1002,1735,1760]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="lucilius">Philonthus lucilius, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35CFF6BF8DBF531D214" pageId="20" pageNumber="21" type="discussion">
<paragraph id="A94736DE4C3CB35CFF6BF8DBF33DD24C" blockId="20.[151,1437,1519,1832]" box="[151,1128,1807,1832]" pageId="20" pageNumber="21">
<emphasis id="9B8CEACC4C3CB35CFF6BF8DBF64AD24C" bold="true" box="[151,287,1807,1832]" pageId="20" pageNumber="21">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C3CB35CFD46F8DBF331D24C" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3CB35CFD46F8DBF468D24C" box="[698,829,1807,1832]" italics="true" pageId="20" pageNumber="21">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C3CB35CFCB7F8DBF309D24C" box="[843,1116,1807,1832]" pageId="20" pageNumber="21" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C3CB35CFC1CF8C5F34DD24C" box="[992,1048,1807,1832]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="A94736DE4C3CB35CFF6BF883F531D214" blockId="20.[151,1436,1879,1977]" box="[151,612,1879,1904]" pageId="20" pageNumber="21">
<taxonomicName id="6EF84D5D4C3CB35CFF6BF883F531D214" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,612,1879,1904]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis id="9B8CEACC4C3CB35CFF6BF883F6FCD214" bold="true" box="[151,425,1879,1904]" italics="true" pageId="20" pageNumber="21">Belonuchus magnificus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3CB35EFF6BF8A8F380D7E0" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="A94736DE4C3CB35CFF6BF8A8F253D2DD" blockId="20.[151,1436,1879,1977]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="89E811F04C3CB35CFF6BF8A8F253D2DD" pageId="20" pageNumber="21">
<taxonomicName id="6EF84D5D4C3CB35CFF6BF8A8F5F1D2F0" ID-CoL="LC92" authority="Bernhauer 1916 c: 277" authorityName="Bernhauer" authorityPageNumber="277" authorityYear="1916" box="[151,676,1916,1941]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="magnificus">
<emphasis id="9B8CEACC4C3CB35CFF6BF8A8F6C8D2F0" box="[151,413,1916,1941]" italics="true" pageId="20" pageNumber="21">Belonuchus magnificus</emphasis>
<treatmentCitation id="285910CF4C3CB35CFE59F8A8F5F1D2F0" author="" box="[421,676,1916,1941]" page="277" pageId="20" pageNumber="21">
<bibRefCitation id="CD694B2F4C3CB35CFE59F8A8F53ED2F1" box="[421,619,1916,1941]" pageId="20" pageNumber="21" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 277
</treatmentCitation>
</taxonomicName>
. Type locality: West-Kolumbien: Umgebung von Cali (S. Antonio, 
<quantity id="6E009B3B4C3CB35CFF39F874F642D2DC" box="[197,279,1952,1976]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="m" value="2000.0">2000m</quantity>
). 
<materialsCitation id="19903C834C3CB35CFED6F875F253D2DD" ID-GBIF-Occurrence="1915497862" box="[298,1286,1952,1977]" collectionCode="FMNH" country="Colombia" location="ST. Distribution" pageId="20" pageNumber="21" specimenCode="FMNH 3" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3CB35CFED6F875F637D2DD" box="[298,354,1953,1977]" pageId="20" pageNumber="21">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3CB35CFE11F875F501D2DC" box="[493,596,1952,1976]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="20" pageNumber="21">FMNH 3</specimenCode>
<location id="AC2760054C3CB35CFDA6F874F446D2DD" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3CB35CFDA6F874F446D2DD" box="[602,787,1952,1977]" country="Colombia" name="ST. Distribution" pageId="20" pageNumber="21">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3CB35CFCE1F874F4DAD2DD" box="[797,911,1952,1977]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFF43F521D5D4" blockId="21.[151,1436,151,248]" box="[151,628,151,176]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFF43F521D5D4" ID-CoL="LC94" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,628,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="9B8CEACC4C3DB35DFF6BFF43F6ECD5D4" bold="true" box="[151,441,151,176]" italics="true" pageId="21" pageNumber="22">Belonuchus meridionalis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFF68F232D59C" blockId="21.[151,1436,151,248]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFF68F232D59C" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFF68F599D5B1" ID-CoL="LC94" authority="Bernhauer 1916 c: 269" authorityName="Bernhauer" authorityPageNumber="269" authorityYear="1916" box="[151,716,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="9B8CEACC4C3DB35DFF6BFF68F6E1D5B1" box="[151,436,188,213]" italics="true" pageId="21" pageNumber="22">Belonuchus meridionalis</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE3EFF68F599D5B1" author="" box="[450,716,188,213]" page="269" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE3EFF68F5DBD5B1" box="[450,654,188,213]" pageId="21" pageNumber="22" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 269
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD21FF69F440D5B1" box="[733,789,189,213]" pageId="21" pageNumber="22">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3DB35DFC70FF68F344D5B1" box="[908,1041,188,213]" name="Argentina" pageId="21" pageNumber="22">Argentinien</collectingCountry>
[illeg. locality] &amp; 
<collectingCountry id="D1EF764E4C3DB35DFB01FF68F206D5B1" box="[1277,1363,188,213]" name="Bolivia" pageId="21" pageNumber="22">Bolivia</collectingCountry>
. 
<typeStatus id="7643887C4C3DB35DFA98FF69F2C9D5B1" box="[1380,1436,189,213]" pageId="21" pageNumber="22">Type</typeStatus>
depository: FMNH 2 ST, ZMUH 1+ST. Distribution: 
<collectingCountry id="D1EF764E4C3DB35DFCE1FF0BF4DAD59C" box="[797,911,223,248]" name="Argentina" pageId="21" pageNumber="22">Argentina</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3DB35DFC67FF0BF4A4D59C" box="[923,1009,223,248]" name="Bolivia" pageId="21" pageNumber="22">Bolivia</collectingCountry>
. Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFEF3F562D424" blockId="21.[151,1437,295,536]" box="[151,567,295,320]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFEF3F562D424" ID-CoL="LC95" authority="Solsky 1864" authorityName="Solsky" authorityYear="1864" box="[151,567,295,320]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="9B8CEACC4C3DB35DFF6BFEF3F6F1D424" bold="true" box="[151,420,295,320]" italics="true" pageId="21" pageNumber="22">Belonuchus mexicanus</emphasis>
<bibRefCitation id="CD694B2F4C3DB35DFE50FEF3F562D424" box="[428,567,295,320]" pageId="21" pageNumber="22" refId="ref73023" refString="Solsky, S. (1864) Description de quelques nouvelles especes de staphylinides. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 37 (2), 433 - 451." type="journal article">Solsky 1864</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFE98F36BD4C8" blockId="21.[151,1437,295,536]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFE98F36BD4C8" infered="true" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFE98F5C7D400" ID-CoL="LC95" authority="Solsky 1864: 440" authorityName="Solsky" authorityPageNumber="440" authorityYear="1864" box="[151,658,332,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="9B8CEACC4C3DB35DFF6BFE98F6F7D400" box="[151,418,332,357]" italics="true" pageId="21" pageNumber="22">Belonuchus mexicanus</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE48FE98F5C7D400" author="" box="[436,658,332,357]" page="440" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE48FE98F51BD401" box="[436,590,332,357]" pageId="21" pageNumber="22" refId="ref73023" refString="Solsky, S. (1864) Description de quelques nouvelles especes de staphylinides. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 37 (2), 433 - 451." type="journal article">Solsky 1864</bibRefCitation>
: 440
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD5AFE99F58BD401" box="[678,734,333,357]" pageId="21" pageNumber="22">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3DB35DFCA2FE98F490D401" box="[862,965,332,357]" name="Mexico" pageId="21" pageNumber="22">Mexique</collectingCountry>
. 
<typeStatus id="7643887C4C3DB35DFC26FE99F347D401" box="[986,1042,333,357]" pageId="21" pageNumber="22">Type</typeStatus>
depository: ZIN? (Solsky) HT. Distribution: 
<collectingRegion id="6B3CF83C4C3DB35DFE9DFEBBF6ECD4EC" box="[353,441,367,392]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">
<collectingCountry id="D1EF764E4C3DB35DFE9DFEBBF6ECD4EC" box="[353,441,367,392]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C3DB35DFE36FEBBF544D4EC" box="[458,529,367,392]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3DB35DFDDEFEBBF5C0D4EC" box="[546,661,367,392]" name="Argentina" pageId="21" pageNumber="22">Argentina</collectingCountry>
. Notes: redescription (
<bibRefCitation id="CD694B2F4C3DB35DFC5EFEBBF31FD4EC" box="[930,1098,367,392]" pageId="21" pageNumber="22" refId="ref72647" refString="Silvestri, F. (1945) Descrizione e biologia del coleottero stafilinide Belonuchus formosus Grav. introdotto in Italia per la lotta contro Ditteri Tripaneidi. Bollettino del R. Laboratorio di Entomologia Agraria di Portici, 5, 312 - 328." type="journal article">Silvestri 1945</bibRefCitation>
); distribution within 
<collectingRegion id="6B3CF83C4C3DB35DFABBFEBBF2C9D4EC" box="[1351,1436,367,392]" country="Mexico" name="Mexico" pageId="21" pageNumber="22">
<collectingCountry id="D1EF764E4C3DB35DFABBFEBBF2C9D4EC" box="[1351,1436,367,392]" name="Mexico" pageId="21" pageNumber="22">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C3DB35DFF30FE40F574D4C8" box="[204,545,404,429]" pageId="21" pageNumber="22" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3DB35DFE59FE41F68BD4C9" box="[421,478,404,429]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2002
</bibRefCitation>
); distribution within 
<collectingCountry id="D1EF764E4C3DB35DFCF7FE40F41AD4C9" box="[779,847,404,429]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C3DB35DFC4AFE41F4BAD4C9" box="[950,1007,404,429]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFE6CF4A1D77C" blockId="21.[151,1437,295,536]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFE6CF4A1D77C" infered="true" pageId="21" pageNumber="22">
= 
<taxonomicName id="6EF84D5D4C3DB35DFF51FE63F449D4B4" ID-CoL="LC7P" authority="Solsky 1864: 440" authorityName="Solsky" authorityPageNumber="440" authorityYear="1864" box="[173,796,439,464]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="variety" species="mexicanus" variety="femoralis">
<emphasis id="9B8CEACC4C3DB35DFF51FE63F6F8D4B4" box="[173,429,439,464]" italics="true" pageId="21" pageNumber="22">Belonuchus mexicanus</emphasis>
var. 
<emphasis id="9B8CEACC4C3DB35DFE18FE63F51BD4B4" box="[484,590,439,464]" italics="true" pageId="21" pageNumber="22">femoralis</emphasis>
<treatmentCitation id="285910CF4C3DB35DFDA8FE63F449D4B4" author="" box="[596,796,439,464]" page="440" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFDA8FE63F5B6D4B4" box="[596,739,439,464]" pageId="21" pageNumber="22" refId="ref73023" refString="Solsky, S. (1864) Description de quelques nouvelles especes de staphylinides. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 37 (2), 433 - 451." type="journal article">Solsky 1864</bibRefCitation>
: 440
</treatmentCitation>
</taxonomicName>
. Type locality: Mexique. Type depository: ZIN? (Solsky) 2 ST. Distribution: Mexico. Notes: not in 
<bibRefCitation id="CD694B2F4C3DB35DFD69FE08F4B1D490" box="[661,996,476,501]" pageId="21" pageNumber="22" refId="ref62964" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus. Vol. 57. W. Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer &amp; Schubert (1914)</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C3DB35DFC13FE08F385D490" box="[1007,1232,476,501]" pageId="21" pageNumber="22" refId="ref63204" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum, 185, 1 - 188." type="journal article">Blackwelder (1944)</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C3DB35DFB20FE09F2C2D491" box="[1244,1431,476,501]" pageId="21" pageNumber="22" refId="ref67417" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article">Herman (2001d)</bibRefCitation>
, but as synonym of 
<taxonomicName id="6EF84D5D4C3DB35DFE66FDD5F4BAD77C" ID-CoL="LC95" authority="in Navarrete-Heredia et al. (2002: 335)" authorityName="in Navarrete-Heredia et al." authorityPageNumber="335" authorityYear="2002" box="[410,1007,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="9B8CEACC4C3DB35DFE66FDD5F57AD77C" box="[410,559,513,536]" italics="true" pageId="21" pageNumber="22">B. mexicanus</emphasis>
in 
<treatmentCitation id="285910CF4C3DB35DFDAFFE2BF4B2D77C" author="" box="[595,999,511,536]" page="335" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFDAFFE2BF4FBD77C" box="[595,942,511,536]" pageId="21" pageNumber="22" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3DB35DFCD6FDD5F436D77C" box="[810,867,511,536]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2002
</bibRefCitation>
: 335
</treatmentCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFD93F516D704" blockId="21.[151,1437,583,717]" box="[151,579,583,608]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFD93F516D704" ID-CoL="LC96" authority="Bernhauer 1921" authorityName="Bernhauer" authorityYear="1921" box="[151,579,583,608]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="9B8CEACC4C3DB35DFF6BFD93F6DDD704" bold="true" box="[151,392,583,608]" italics="true" pageId="21" pageNumber="22">Belonuchus militaris</emphasis>
Bernhauer 1921
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFDB8F64BD7A9" blockId="21.[151,1437,583,717]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFDB8F64BD7A9" infered="true" pageId="21" pageNumber="22">
<materialsCitation id="19903C834C3DB35DFF6BFDB8F377D7E1" ID-GBIF-Occurrence="1915497890" box="[151,1058,620,645]" country="Brazil" location="Sao Paulo" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFDB8F5C3D7E0" ID-CoL="LC96" authority="Bernhauer 1921 c: 170" authorityName="Bernhauer" authorityPageNumber="170" authorityYear="1921" box="[151,662,620,645]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="militaris">
<emphasis id="9B8CEACC4C3DB35DFF6BFDB8F6D1D7E1" box="[151,388,620,645]" italics="true" pageId="21" pageNumber="22">Belonuchus militaris</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE6CFDB8F5C3D7E0" author="" box="[400,662,620,645]" page="170" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE6CFDB8F50FD7E0" box="[400,602,620,645]" pageId="21" pageNumber="22" refId="ref62483" refString="Bernhauer, M. (1921 c) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article">Bernhauer 1921c</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD5AFDB9F58BD7E1" box="[678,734,621,645]" pageId="21" pageNumber="22">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C3DB35DFCAEFDB8F49ED7E1" box="[850,971,620,645]" country="Brazil" name="Sao Paulo" pageId="21" pageNumber="22">Sao Paulo</collectingRegion>
, 
<collectingCountry id="D1EF764E4C3DB35DFC26FDB8F377D7E1" box="[986,1058,620,645]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C3DB35DFBCEFDB9F57BD7CC" ID-GBIF-Occurrence="1915497888" collectionCode="FMNH" country="Brazil" location="ST. Distribution" pageId="21" pageNumber="22" specimenCode="FMNH 1" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="7643887C4C3DB35DFBCEFDB9F33CD7E1" box="[1074,1129,621,645]" pageId="21" pageNumber="22">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3DB35DFAFCFDB9F23FD7E0" box="[1280,1386,620,644]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="21" pageNumber="22">FMNH 1</specimenCode>
<location id="AC2760054C3DB35DFA8BFDB8F604D7CC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3DB35DFA8BFDB8F604D7CC" country="Brazil" name="ST. Distribution" pageId="21" pageNumber="22" stateProvince="Sao Paulo">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3DB35DFEA7FD5BF6CAD7CC" box="[347,415,655,680]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C3DB35DFE53FD44F573D7CC" box="[431,550,655,680]" country="Brazil" name="Sao Paulo" pageId="21" pageNumber="22">São Paulo</collectingRegion>
)
</materialsCitation>
. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C3DB35DFBFCFD45F36DD7CC" box="[1024,1080,655,680]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2013). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFD2FF5E1D671" blockId="21.[151,1437,763,861]" box="[151,692,763,789]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFD2FF6A9D671" authority="Bruch 1915" authorityName="Bruch" authorityYear="1915" box="[151,508,763,789]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="9B8CEACC4C3DB35DFF6BFD2FF63BD670" bold="true" box="[151,366,763,788]" italics="true" pageId="21" pageNumber="22">Belonuchus minor</emphasis>
<bibRefCitation id="CD694B2F4C3DB35DFE89FD28F6A9D671" box="[373,508,764,789]" pageId="21" pageNumber="22" refId="ref63589" refString="Bruch, C. (1915) Catalogo sistematico de los coleopteros de la Republica Argentina. II. Revista del Museo de La Plata, 19, 471 - 526." type="journal article">Bruch 1915</bibRefCitation>
</taxonomicName>
, 
<emphasis id="9B8CEACC4C3DB35DFDFBFD28F5E1D671" bold="true" box="[519,692,764,789]" pageId="21" pageNumber="22">nomen nudum</emphasis>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFCCBF3BDD638" blockId="21.[151,1437,763,861]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFCCBF3BDD638" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFCCBF512D65C" authority="Bruch 1915: 504" authorityName="Bruch" authorityPageNumber="504" authorityYear="1915" box="[151,583,799,824]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="9B8CEACC4C3DB35DFF6BFCCBF63ED65C" box="[151,363,799,824]" italics="true" pageId="21" pageNumber="22">Belonuchus minor</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE85FCCBF512D65C" author="" box="[377,583,799,824]" page="504" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE85FCCBF55DD65C" box="[377,520,799,824]" pageId="21" pageNumber="22" refId="ref63589" refString="Bruch, C. (1915) Catalogo sistematico de los coleopteros de la Republica Argentina. II. Revista del Museo de La Plata, 19, 471 - 526." type="journal article">Bruch 1915</bibRefCitation>
: 504
</treatmentCitation>
</taxonomicName>
, attributed to Bernhauer 
<emphasis id="9B8CEACC4C3DB35DFC86FCF5F4B2D65C" box="[890,999,799,824]" italics="true" pageId="21" pageNumber="22">in litteris</emphasis>
; nomen nudum. Locality: Misiones. Depository: MLPA (MLP 180). Distribution: Argentina. Note: see 
<treatmentCitation id="285910CF4C3DB35DFC4DFC90F389D638" author="" box="[945,1244,836,861]" page="38" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFC4DFC90F3E4D638" box="[945,1201,836,861]" pageId="21" pageNumber="22" refId="ref66201" refString="Fernandez, L. A., Cabrera, N., Chani-Posse, M. R. &amp; Gabellone, C. (2007) Catalog of the types of Staphylinidae Insecta, Coleoptera deposited at Museo de La Plata Argentina. Zootaxa, 1611, 25 - 44." type="journal article">
Fernández 
<emphasis id="9B8CEACC4C3DB35DFBD0FC91F330D639" box="[1068,1125,836,861]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
(2007
</bibRefCitation>
: 38
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFC5FF50AD6C1" blockId="21.[151,1436,907,1005]" box="[151,607,907,933]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFC5FF50AD6C1" ID-CoL="LC99" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,607,907,933]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="miripennis">
<emphasis id="9B8CEACC4C3DB35DFF6BFC5FF6F1D6C0" bold="true" box="[151,420,907,932]" italics="true" pageId="21" pageNumber="22">Belonuchus miripennis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFC7BF275D689" blockId="21.[151,1436,907,1005]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFC7BF275D689" pageId="21" pageNumber="22">
<materialsCitation id="19903C834C3DB35DFF6BFC7BF249D689" ID-GBIF-Occurrence="1915497926" collectionCode="FMNH" collectorName="Umgebung von Cali" country="Colombia" elevation="1700" location="ST. Distribution" pageId="21" pageNumber="22" specimenCode="FMNH 3" specimenCount="2" stateProvince="Tolima" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFC7BF437D6AC" ID-CoL="LC99" authority="Bernhauer 1916 c: 267" authorityName="Bernhauer" authorityPageNumber="267" authorityYear="1916" box="[151,866,943,968]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="miripennis" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C3DB35DFF6BFC7BF51BD6AC" box="[151,590,943,968]" italics="true" pageId="21" pageNumber="22">Belonuchus (Musicoderus) miripennis</emphasis>
<treatmentCitation id="285910CF4C3DB35DFDA7FC7BF437D6AC" author="" box="[603,866,943,968]" page="267" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFDA7FC7BF470D6AC" box="[603,805,943,968]" pageId="21" pageNumber="22" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 267
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFC8EFC64F4FFD6AC" box="[882,938,944,968]" pageId="21" pageNumber="22">Type</typeStatus>
locality: Cañon del Mte. 
<collectingRegion id="6B3CF83C4C3DB35DFB19FC7BF262D6AC" box="[1253,1335,943,968]" country="Colombia" name="Tolima" pageId="21" pageNumber="22">Tolima</collectingRegion>
, 
<elevation id="22D5D1ED4C3DB35DFABBFC64F2CDD6AC" box="[1351,1432,944,968]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="22" unit="m" value="1700.0">
<quantity id="6E009B3B4C3DB35DFABBFC64F2CDD6AC" box="[1351,1432,944,968]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="22" unit="m" value="1700.0">1700m</quantity>
</elevation>
, 
<collectorName id="040D53084C3DB35DFF39FC00F6FDD689" box="[197,424,980,1005]" pageId="21" pageNumber="22">Umgebung von Cali</collectorName>
am 
<collectingRegion id="6B3CF83C4C3DB35DFE24FC00F574D688" box="[472,545,980,1004]" country="Colombia" name="Cauca" pageId="21" pageNumber="22">Cauca</collectingRegion>
, Columbien. 
<typeStatus id="7643887C4C3DB35DFD4BFC01F5BAD689" box="[695,751,981,1005]" pageId="21" pageNumber="22">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3DB35DFC86FC01F4B5D688" box="[890,992,980,1004]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="21" pageNumber="22">FMNH 3</specimenCode>
<location id="AC2760054C3DB35DFC1BFC00F3F5D689" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3DB35DFC1BFC00F3F5D689" box="[999,1184,980,1005]" country="Colombia" name="ST. Distribution" pageId="21" pageNumber="22" stateProvince="Tolima">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3DB35DFB56FC00F249D689" box="[1194,1308,980,1005]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFBCFF573D150" blockId="21.[151,1436,1051,1149]" box="[151,550,1051,1077]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFBCFF573D150" ID-CoL="5WGHX" authority="Fauvel 1891" authorityName="Fauvel" authorityYear="1891" box="[151,550,1051,1077]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="9B8CEACC4C3DB35DFF6BFBCFF6C6D150" bold="true" box="[151,403,1051,1076]" italics="true" pageId="21" pageNumber="22">Belonuchus modestus</emphasis>
<bibRefCitation id="CD694B2F4C3DB35DFE67FBC8F573D150" box="[411,550,1052,1077]" pageId="21" pageNumber="22" refId="ref66096" refString="Fauvel, A. (1891) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 11 e memoire: Staphylinides. Revue d'Entomologie, 10, 87 - 127." type="journal article">Fauvel 1891</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFBEBF3A9D119" blockId="21.[151,1436,1051,1149]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFBEBF3A9D119" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFBEBF53BD13C" ID-CoL="5WGHX" authority="Fauvel 1891: 118" authorityName="Fauvel" authorityPageNumber="118" authorityYear="1891" box="[151,622,1087,1112]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="9B8CEACC4C3DB35DFF6BFBEBF6DBD13C" box="[151,398,1087,1112]" italics="true" pageId="21" pageNumber="22">Belonuchus modestus</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE60FBEBF53BD13C" author="" box="[412,622,1087,1112]" page="118" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE60FBEBF564D13C" box="[412,561,1087,1112]" pageId="21" pageNumber="22" refId="ref66096" refString="Fauvel, A. (1891) Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 11 e memoire: Staphylinides. Revue d'Entomologie, 10, 87 - 127." type="journal article">Fauvel 1891</bibRefCitation>
: 118
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD83FB94F5E2D13C" box="[639,695,1088,1112]" pageId="21" pageNumber="22">Type</typeStatus>
locality: Colonie Tovar [
<collectingCountry id="D1EF764E4C3DB35DFC1AFBEBF336D13C" box="[998,1123,1087,1112]" name="Venezuela" pageId="21" pageNumber="22">Venezuela</collectingCountry>
]. 
<typeStatus id="7643887C4C3DB35DFB86FB94F3E7D13C" box="[1146,1202,1088,1112]" pageId="21" pageNumber="22">Type</typeStatus>
depository: IRSNB (Fauvel) 3 ST [MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C3DB35DFD6EFBB0F45DD119" box="[658,776,1124,1149]" name="Venezuela" pageId="21" pageNumber="22">Venezuela</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3DB35DFCEFFBB0F4D0D119" box="[787,901,1124,1149]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
. Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFB7FF52AD1A1" blockId="21.[151,1436,1195,1293]" box="[151,639,1195,1221]" pageId="21" pageNumber="22">
<emphasis id="9B8CEACC4C3DB35DFF6BFB7FF7F6D1A0" bold="true" box="[151,163,1195,1220]" italics="true" pageId="21" pageNumber="22">*</emphasis>
<taxonomicName id="6EF84D5D4C3DB35DFF5EFB7FF52AD1A1" ID-CoL="LC9B" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[162,639,1195,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="montanellus">
<emphasis id="9B8CEACC4C3DB35DFF5EFB7FF690D1A0" bold="true" box="[162,453,1195,1220]" italics="true" pageId="21" pageNumber="22">Belonuchus montanellus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFB1BF3A8D069" blockId="21.[151,1436,1195,1293]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFB1BF3A8D069" pageId="21" pageNumber="22">
<materialsCitation id="19903C834C3DB35DFF6BFB1BF3ACD069" ID-GBIF-Occurrence="1915497923" collectionCode="FMNH" collectorName="Alto &amp; Umgebung von Cali" country="Colombia" elevation="1700" location="ST. Distribution" pageId="21" pageNumber="22" specimenCode="FMNH 4" specimenCount="2" stateProvince="Tolima" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFB1BF5FCD18C" ID-CoL="LC9B" authority="Bernhauer 1916 c: 272" authorityName="Bernhauer" authorityPageNumber="272" authorityYear="1916" box="[151,681,1231,1256]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="montanellus">
<emphasis id="9B8CEACC4C3DB35DFF6BFB1BF6FCD18C" box="[151,425,1231,1256]" italics="true" pageId="21" pageNumber="22">Belonuchus montanellus</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE53FB1BF5FCD18C" author="" box="[431,681,1231,1256]" page="272" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE53FB1BF526D18C" box="[431,627,1231,1256]" pageId="21" pageNumber="22" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 272
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD4FFB04F5BED18C" box="[691,747,1232,1256]" pageId="21" pageNumber="22">Type</typeStatus>
localities: Cañon del Mte. 
<collectingRegion id="6B3CF83C4C3DB35DFBEBFB1BF33CD18C" box="[1047,1129,1231,1256]" country="Colombia" name="Tolima" pageId="21" pageNumber="22">Tolima</collectingRegion>
, 
<elevation id="22D5D1ED4C3DB35DFB8FFB04F390D18C" box="[1139,1221,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="22" unit="m" value="1700.0">
<quantity id="6E009B3B4C3DB35DFB8FFB04F390D18C" box="[1139,1221,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="22" unit="m" value="1700.0">1700m</quantity>
</elevation>
; 
<collectorName id="040D53084C3DB35DFB33FB1BF254D18C" box="[1231,1281,1231,1256]" pageId="21" pageNumber="22">Alto</collectorName>
de las cruces, 
<elevation id="22D5D1ED4C3DB35DFF39FB20F646D068" box="[197,275,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="m" value="2200.0">
<quantity id="6E009B3B4C3DB35DFF39FB20F646D068" box="[197,275,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="m" value="2200.0">2200m</quantity>
</elevation>
(
<collectorName id="040D53084C3DB35DFEDDFB20F553D069" box="[289,518,1268,1293]" pageId="21" pageNumber="22">Umgebung von Cali</collectorName>
am 
<collectingRegion id="6B3CF83C4C3DB35DFDCAFB20F5D4D068" box="[566,641,1268,1292]" country="Colombia" name="Cauca" pageId="21" pageNumber="22">Cauca</collectingRegion>
). 
<typeStatus id="7643887C4C3DB35DFD68FB21F59ED069" box="[660,715,1269,1293]" pageId="21" pageNumber="22">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3DB35DFCABFB21F4E8D068" box="[855,957,1268,1292]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="21" pageNumber="22">FMNH 4</specimenCode>
<location id="AC2760054C3DB35DFC38FB20F328D069" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3DB35DFC38FB20F328D069" box="[964,1149,1268,1293]" country="Colombia" name="ST. Distribution" pageId="21" pageNumber="22" stateProvince="Tolima">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3DB35DFB7BFB20F3ACD069" box="[1159,1273,1268,1293]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFAEFF5D6D030" blockId="21.[151,1436,1339,1509]" box="[151,643,1339,1365]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFAEFF5D6D030" ID-CoL="LC9C" authority="Blackwelder 1944" authorityName="Blackwelder" authorityYear="1944" box="[151,643,1339,1365]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="montanicus">
<emphasis id="9B8CEACC4C3DB35DFF6BFAEFF6FAD030" bold="true" box="[151,431,1339,1364]" italics="true" pageId="21" pageNumber="22">Belonuchus montanicus</emphasis>
<bibRefCitation id="CD694B2F4C3DB35DFE4AFAE8F5D6D030" box="[438,643,1340,1365]" pageId="21" pageNumber="22" refId="ref63204" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum, 185, 1 - 188." type="journal article">Blackwelder 1944</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFA8BF20FD01C" blockId="21.[151,1436,1339,1509]" box="[151,1370,1375,1400]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFA8BF20FD01C" box="[151,1370,1375,1400]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BFA8BF5FBD01C" ID-CoL="LC9C" authority="Blackwelder 1944: 137" authorityName="Blackwelder" authorityPageNumber="137" authorityYear="1944" box="[151,686,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="montanicus">
<emphasis id="9B8CEACC4C3DB35DFF6BFA8BF6F4D01C" box="[151,417,1375,1400]" italics="true" pageId="21" pageNumber="22">Belonuchus montanicus</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE54FA8BF5FBD01C" author="" box="[424,686,1375,1400]" page="137" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE54FA8BF52CD01C" box="[424,633,1375,1400]" pageId="21" pageNumber="22" refId="ref63204" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum, 185, 1 - 188." type="journal article">Blackwelder 1944</bibRefCitation>
: 137
</treatmentCitation>
</taxonomicName>
(new name for 
<taxonomicName id="6EF84D5D4C3DB35DFCA3FA8BF218D01C" ID-CoL="8XVPK" authority="Bernhauer 1916: 273" authorityName="Bernhauer" authorityPageNumber="273" authorityYear="1916" box="[863,1357,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="9B8CEACC4C3DB35DFCA3FA8BF303D01C" box="[863,1110,1375,1400]" italics="true" pageId="21" pageNumber="22">Belonuchus montanus</emphasis>
Bernhauer 1916: 273
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BFA51F489D081" blockId="21.[151,1436,1339,1509]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BFA51F489D081" infered="true" pageId="21" pageNumber="22">
= 
<taxonomicName id="6EF84D5D4C3DB35DFF4CFA50F5FAD0F8" ID-CoL="8XVPK" authority="Bernhauer 1916 c: 273" authorityName="Bernhauer" authorityPageNumber="273" authorityYear="1916" box="[176,687,1412,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="9B8CEACC4C3DB35DFF4CFA50F6FCD0F8" box="[176,425,1412,1437]" italics="true" pageId="21" pageNumber="22">Belonuchus montanus</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE4EFA50F5FAD0F8" author="" box="[434,687,1412,1437]" page="273" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE4EFA50F52DD0F9" box="[434,632,1412,1437]" pageId="21" pageNumber="22" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 273
</treatmentCitation>
</taxonomicName>
(preoccupied, junior primary homonym of 
<taxonomicName id="6EF84D5D4C3DB35DFB5FFA50F6DED0A4" ID-CoL="8XVPK" authority="Fauvel 1895: 267" authorityName="Fauvel" authorityPageNumber="267" authorityYear="1895" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="9B8CEACC4C3DB35DFB5FFA50F2C9D0F8" box="[1187,1436,1412,1437]" italics="true" pageId="21" pageNumber="22">Belonuchus montanus</emphasis>
<treatmentCitation id="285910CF4C3DB35DFF39FA73F6DED0A4" author="" box="[197,395,1447,1472]" page="267" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFF39FA73F606D0A4" box="[197,339,1447,1472]" pageId="21" pageNumber="22" refId="ref66135" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286." type="journal article">Fauvel 1895</bibRefCitation>
: 267
</treatmentCitation>
</taxonomicName>
). Type localities: West-Columbien: Cañon del Mte. Tolima, 
<quantity id="6E009B3B4C3DB35DFBD5FA7CF32FD0A4" box="[1065,1146,1448,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="22" unit="m" value="1700.0">1700m</quantity>
; Rio Vitaco, 
<quantity id="6E009B3B4C3DB35DFAF6FA7CF209D0A4" box="[1290,1372,1448,1472]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="m" value="2200.0">2200m</quantity>
; Villa Elvira, 
<quantity id="6E009B3B4C3DB35DFEEBFA18F63DD080" box="[279,360,1484,1508]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="22" unit="m" value="1800.0">1800m</quantity>
. Type depository: FMNH 3 ST. Distribution: Colombia.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BF9C7F503D348" blockId="21.[151,1436,1555,1653]" box="[151,598,1555,1581]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BF9C7F503D348" ID-CoL="LC9F" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,598,1555,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="9B8CEACC4C3DB35DFF6BF9C7F6CED348" bold="true" box="[151,411,1555,1580]" italics="true" pageId="21" pageNumber="22">Belonuchus monticola</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BF9E3F4B7D311" blockId="21.[151,1436,1555,1653]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BF9E3F4B7D311" pageId="21" pageNumber="22">
<materialsCitation id="19903C834C3DB35DFF6BF9E3F6D3D311" ID-GBIF-Occurrence="1915497929" collectorName="Umgebung von Cali &amp; West-Columbien" elevation="2200" location="Rio Vitaco" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Cauca" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3DB35DFF6BF9E3F5D1D334" ID-CoL="LC9F" authority="Bernhauer 1917 b: 18" authorityName="Bernhauer" authorityPageNumber="18" authorityYear="1917" box="[151,644,1591,1616]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="9B8CEACC4C3DB35DFF6BF9E3F6C5D334" box="[151,400,1591,1616]" italics="true" pageId="21" pageNumber="22">Belonuchus monticola</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE6AF9E3F5D1D334" author="" box="[406,644,1591,1616]" page="18" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE6AF9E3F509D334" box="[406,604,1591,1616]" pageId="21" pageNumber="22" refId="ref62221" refString="Bernhauer, M. (1917 b) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (17. Beitrag - Fortsetzung). Entomologische Blatter, 13 (1 - 3), 17 - 24." type="journal article">Bernhauer 1917b</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD73F9ECF592D334" box="[655,711,1592,1616]" pageId="21" pageNumber="22">Type</typeStatus>
locality: 
<location id="AC2760054C3DB35DFCD3F9E3F4FFD334" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3DB35DFCD3F9E3F4FFD334" box="[815,938,1591,1616]" name="Rio Vitaco" pageId="21" pageNumber="22" stateProvince="Cauca">Rio Vitaco</location>
, 
<elevation id="22D5D1ED4C3DB35DFC48F9ECF350D334" box="[948,1029,1592,1616]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="m" value="2200.0">
<quantity id="6E009B3B4C3DB35DFC48F9ECF350D334" box="[948,1029,1592,1616]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="21" pageNumber="22" unit="m" value="2200.0">2200m</quantity>
</elevation>
, 
<collectorName id="040D53084C3DB35DFBECF9E3F3A4D334" box="[1040,1265,1591,1616]" pageId="21" pageNumber="22">Umgebung von Cali</collectorName>
am Rio 
<collectingRegion id="6B3CF83C4C3DB35DFAB3F9ECF2CDD334" box="[1359,1432,1592,1616]" country="Colombia" name="Cauca" pageId="21" pageNumber="22">Cauca</collectingRegion>
, 
<collectorName id="040D53084C3DB35DFF39F988F6D3D311" box="[197,390,1628,1653]" pageId="21" pageNumber="22">West-Columbien</collectorName>
</materialsCitation>
. 
<materialsCitation id="19903C834C3DB35DFE6CF989F4B7D311" ID-GBIF-Occurrence="1915497887" box="[400,994,1628,1653]" collectionCode="FMNH" country="Colombia" location="Distribution" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3DB35DFE6CF989F692D311" box="[400,455,1629,1653]" pageId="21" pageNumber="22">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3DB35DFDA8F989F5F0D310" box="[596,677,1629,1652]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="21" pageNumber="22">FMNH</collectionCode>
HT. 
<location id="AC2760054C3DB35DFD21F988F43CD311" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3DB35DFD21F988F43CD311" box="[733,873,1628,1653]" country="Colombia" name="Distribution" pageId="21" pageNumber="22">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3DB35DFC8FF988F4B7D311" box="[883,994,1628,1653]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BF977F516D3D8" blockId="21.[151,1436,1699,1869]" box="[151,579,1699,1725]" pageId="21" pageNumber="22">
<emphasis id="9B8CEACC4C3DB35DFF6BF977F7F6D3D8" bold="true" box="[151,163,1699,1724]" italics="true" pageId="21" pageNumber="22">*</emphasis>
<taxonomicName id="6EF84D5D4C3DB35DFF5EF977F516D3D8" ID-CoL="LC9H" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[162,579,1699,1725]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mordens">
<emphasis id="9B8CEACC4C3DB35DFF5EF977F6CFD3D8" bold="true" box="[162,410,1699,1724]" italics="true" pageId="21" pageNumber="22">Belonuchus mordens</emphasis>
<bibRefCitation id="CD694B2F4C3DB35DFE5DF970F516D3D8" box="[417,579,1700,1725]" pageId="21" pageNumber="22" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BF913F618D228" blockId="21.[151,1436,1699,1869]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BF913F618D228" infered="true" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BF913F525D384" ID-CoL="LC9H" authority="Erichson 1840: 422" authorityName="Erichson" authorityPageNumber="422" authorityYear="1840" box="[151,624,1735,1760]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mordens">
<emphasis id="9B8CEACC4C3DB35DFF6BF913F6D7D384" box="[151,386,1735,1760]" italics="true" pageId="21" pageNumber="22">Belonuchus mordens</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE70F913F525D384" author="" box="[396,624,1735,1760]" page="422" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE70F913F561D384" box="[396,564,1735,1760]" pageId="21" pageNumber="22" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 422
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD80F91CF5E6D384" box="[636,691,1736,1760]" pageId="21" pageNumber="22">Type</typeStatus>
locality: Brasilia. 
<typeStatus id="7643887C4C3DB35DFC76F91CF497D384" box="[906,962,1736,1760]" pageId="21" pageNumber="22">Type</typeStatus>
depository: ZMHB 1 ST 
<specimenCount id="BFFEFD574C3DB35DFB12F91CF256D3BB" box="[1262,1283,1736,1759]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
[MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C3DB35DFEA2F939F6C1D260" box="[350,404,1773,1796]" name="Peru" pageId="21" pageNumber="22">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3DB35DFE5FF938F6BFD261" box="[419,490,1772,1797]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3DB35DFE04F938F53FD261" box="[504,618,1772,1797]" name="Argentina" pageId="21" pageNumber="22">Argentina</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3DB35DFD85F938F59AD261" box="[633,719,1772,1797]" name="Bolivia" pageId="21" pageNumber="22">Bolivia</collectingCountry>
. Notes: frequent guest of 
<taxonomicName id="6EF84D5D4C3DB35DFC02F938F33FD261" ID-CoL="62Z5J" authorityName="Illiger" authorityYear="1806" box="[1022,1130,1772,1797]" class="Insecta" family="Apidae" genus="Melipona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3DB35DFC02F938F33FD261" box="[1022,1130,1772,1797]" italics="true" pageId="21" pageNumber="22">Melipona</emphasis>
</taxonomicName>
nests in 
<collectingCountry id="D1EF764E4C3DB35DFB2AF938F24FD261" box="[1238,1306,1772,1797]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
(
<treatmentCitation id="285910CF4C3DB35DFAD0F938F66CD24C" author="" page="482" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFAD0F938F654D24C" pageId="21" pageNumber="22" refId="ref73862" refString="Wasmann, E. (1904) Contribuicao para o estudo dos hospedes de abelhas brasileiras. Revista Museo Paulista, Sao Paulo, 6, 482 - 487." type="journal article">Wasmann 1904</bibRefCitation>
: 482
</treatmentCitation>
, 
<bibRefCitation id="CD694B2F4C3DB35DFEB8F8DBF68CD24C" box="[324,473,1807,1832]" pageId="21" pageNumber="22" refId="ref68222" refString="Kistner, D. H. (1982) The social insects' bestiary. In: Hermann, H. R. (Ed.), Social Insects. Vol. III. Academic Press, New York, pp. 1 - 244." type="book chapter">Kistner 1982</bibRefCitation>
); distribution, including 
<collectingCountry id="D1EF764E4C3DB35DFD11F8C4F474D243" box="[749,801,1808,1831]" name="Peru" pageId="21" pageNumber="22">Peru</collectingCountry>
and 
<collectingCountry id="D1EF764E4C3DB35DFCABF8DBF4FFD24C" box="[855,938,1807,1832]" name="Bolivia" pageId="21" pageNumber="22">Bolivia</collectingCountry>
(
<bibRefCitation id="CD694B2F4C3DB35DFC44F8DBF3C3D24C" box="[952,1174,1807,1832]" pageId="21" pageNumber="22" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b</bibRefCitation>
), within 
<collectingCountry id="D1EF764E4C3DB35DFB04F8DBF269D24C" box="[1272,1340,1807,1832]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C3DB35DFF39F8E1F7A8D229" box="[197,253,1844,1869]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BF8AFF561D2F1" blockId="21.[151,1437,1915,2013]" box="[151,564,1915,1941]" pageId="21" pageNumber="22">
<taxonomicName id="6EF84D5D4C3DB35DFF6BF8AFF561D2F1" ID-CoL="LC9J" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[151,564,1915,1941]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="moritzi">
<emphasis id="9B8CEACC4C3DB35DFF6BF8AFF62CD2F0" bold="true" box="[151,377,1915,1940]" italics="true" pageId="21" pageNumber="22">Belonuchus moritzi</emphasis>
<bibRefCitation id="CD694B2F4C3DB35DFE7CF8A8F561D2F1" box="[384,564,1916,1941]" pageId="21" pageNumber="22" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3DB35DFF6BF874F5F3D2B9" blockId="21.[151,1437,1915,2013]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="89E811F04C3DB35DFF6BF874F5F3D2B9" pageId="21" pageNumber="22">
<materialsCitation id="19903C834C3DB35DFF6BF874F4CCD2DD" ID-GBIF-Occurrence="1915497907" box="[151,921,1952,1977]" country="Venezuela" location="Venezuela" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3DB35DFF6BF874F53AD2DC" ID-CoL="LC9J" authority="Bernhauer 1906: 331" authorityName="Bernhauer" authorityPageNumber="331" authorityYear="1906" box="[151,623,1952,1977]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="moritzi">
<emphasis id="9B8CEACC4C3DB35DFF6BF874F627D2DC" box="[151,370,1952,1977]" italics="true" pageId="21" pageNumber="22">Belonuchus moritzi</emphasis>
<treatmentCitation id="285910CF4C3DB35DFE87F874F53AD2DC" author="" box="[379,623,1952,1977]" page="331" pageId="21" pageNumber="22">
<bibRefCitation id="CD694B2F4C3DB35DFE87F874F560D2DD" box="[379,565,1952,1977]" pageId="21" pageNumber="22" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 331
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3DB35DFD80F875F5E1D2DD" box="[636,692,1953,1977]" pageId="21" pageNumber="22">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3DB35DFCDEF874F4CCD2DD" box="[802,921,1952,1977]" name="Venezuela" pageId="21" pageNumber="22">Venezuela</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C3DB35DFC59F875F5F3D2B9" ID-GBIF-Occurrence="1915497884" collectionCode="FMNH" country="Venezuela" location="ST. Distribution" pageId="21" pageNumber="22" specimenCode="FMNH 2" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3DB35DFC59F875F488D2DD" box="[933,989,1953,1977]" pageId="21" pageNumber="22">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3DB35DFB91F875F380D2DC" box="[1133,1237,1952,1976]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="21" pageNumber="22">FMNH 2</specimenCode>
<location id="AC2760054C3DB35DFB22F874F2CCD2DD" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3DB35DFB22F874F2CCD2DD" box="[1246,1433,1952,1977]" country="Venezuela" name="ST. Distribution" pageId="21" pageNumber="22">ST. Distribution</location>
: Venezuel. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFF43F5D0D5D4" blockId="22.[151,1437,151,320]" box="[151,645,151,176]" pageId="22" pageNumber="23">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFF43F5D0D5D4" ID-CoL="LC9N" authority="(Coiffait 1981)" authorityName="Coiffait" authorityYear="1981" baseAuthorityName="Coiffait" baseAuthorityYear="1981" box="[151,645,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="multipunctatus">
<emphasis id="9B8CEACC4C3EB35EFF6BFF43F683D5D4" bold="true" box="[151,470,151,176]" italics="true" pageId="22" pageNumber="23">Belonuchus multipunctatus</emphasis>
(
<bibRefCitation id="CD694B2F4C3EB35EFE18FF43F528D5D4" box="[484,637,151,176]" pageId="22" pageNumber="23" refId="ref64928" refString="Coiffait, H. (1981) Contribution a la connaissance des Staphylinides des Iles Galapagos (Coleoptera). Annales de la Societe Entomologique de France, New Series, 17, 287 - 310." type="journal article">Coiffait 1981</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFF68F5EAD424" blockId="22.[151,1437,151,320]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BFF68F5EAD424" infered="true" pageId="22" pageNumber="23">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFF68F5D7D5B0" ID-CoL="39QPR" authority="Coiffait 1981: 297" authorityName="Coiffait" authorityPageNumber="297" authorityYear="1981" box="[151,642,188,213]" class="Insecta" family="Staphylinidae" genus="Endeius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="multipunctatus">
<emphasis id="9B8CEACC4C3EB35EFF6BFF68F6F4D5B1" box="[151,417,188,213]" italics="true" pageId="22" pageNumber="23">Endeius multipunctatus</emphasis>
<treatmentCitation id="285910CF4C3EB35EFE57FF68F5D7D5B0" author="" box="[427,642,188,213]" page="297" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFE57FF68F512D5B0" box="[427,583,188,213]" pageId="22" pageNumber="23" refId="ref64928" refString="Coiffait, H. (1981) Contribution a la connaissance des Staphylinides des Iles Galapagos (Coleoptera). Annales de la Societe Entomologique de France, New Series, 17, 287 - 310." type="journal article">Coiffait 1981</bibRefCitation>
: 297
</treatmentCitation>
</taxonomicName>
. Type locality: Galapagos: I. Santa Cruz versant S.E., basse altitude. Type depository: IRSNB HT ♂; IRSNB, MNHN 258 PT. Distribution: Galapagos Is. (Santa Cruz). Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C3EB35EFE94FED0F410D479" ID-CoL="8XL73" authority="(Chani Posse de Maus 2008: 386)" authorityName="Chani Posse de Maus" authorityPageNumber="386" authorityYear="2008" baseAuthorityName="Chani Posse de Maus" baseAuthorityYear="2008" box="[360,837,260,285]" class="Insecta" family="Staphylinidae" genus="Endeius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3EB35EFE94FED0F687D47F" box="[360,466,260,285]" italics="true" pageId="22" pageNumber="23">Endeius (</emphasis>
Chani Posse de 
<treatmentCitation id="285910CF4C3EB35EFD7FFED1F468D479" author="" box="[643,829,260,285]" page="386" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFD7FFED1F451D478" box="[643,772,260,284]" pageId="22" pageNumber="23" refId="ref64115" refString="Chani Posse de Maus, M. (2008) Systematic revision and cladistic analysis of the South American genus Endeius Coiffait &amp; Saiz (Coleoptera: Staphylinidae). Insect Systematics &amp; Evolution, 39, 381 - 406. https: // doi. org / 10.1163 / 187631208788784255" type="journal article">Maus 2008</bibRefCitation>
: 386
</treatmentCitation>
)
</taxonomicName>
; redescription (
<treatmentCitation id="285910CF4C3EB35EFC0DFED0F252D478" author="" box="[1009,1287,260,285]" page="222" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFC0DFED0F252D478" box="[1009,1287,260,285]" pageId="22" pageNumber="23" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b: 222</bibRefCitation>
</treatmentCitation>
); distribution within Galápagos Is. (
<bibRefCitation id="CD694B2F4C3EB35EFE46FEFCF5E6D424" box="[442,691,295,320]" pageId="22" pageNumber="23" refId="ref70441" refString="Newton, A. F. &amp; Peck, S. B. (2006) Family Staphylinidae, the rove beetles. In: Peck, S. B. (Ed.), The beetles of the Galapagos Islands, Ecuador: evolution, ecology and diversity (Insecta: Coleoptera). NRC Research Press, Ottawa, Ontario, pp. 131 - 158." type="book chapter">Newton &amp; Peck 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFEBBF57AD4EC" blockId="22.[151,1437,367,501]" box="[151,559,367,392]" pageId="22" pageNumber="23">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFEBBF57AD4EC" ID-CoL="LC9P" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[151,559,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mundus">
<emphasis id="9B8CEACC4C3EB35EFF6BFEBBF6D0D4EC" bold="true" box="[151,389,367,392]" italics="true" pageId="22" pageNumber="23">Belonuchus mundus</emphasis>
<bibRefCitation id="CD694B2F4C3EB35EFE70FEBBF57AD4EC" box="[396,559,367,392]" pageId="22" pageNumber="23" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFE40F3D0D490" blockId="22.[151,1437,367,501]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BFE40F3D0D490" infered="true" pageId="22" pageNumber="23">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFE40F529D4C9" ID-CoL="LC9P" authority="Erichson 1840: 425" authorityName="Erichson" authorityPageNumber="425" authorityYear="1840" box="[151,636,404,429]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mundus">
<emphasis id="9B8CEACC4C3EB35EFF6BFE40F62AD4C9" box="[151,383,404,429]" italics="true" pageId="22" pageNumber="23">Belonuchus mundus</emphasis>
<treatmentCitation id="285910CF4C3EB35EFE72FE40F529D4C9" author="" box="[398,636,404,429]" page="425" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFE72FE40F569D4C8" box="[398,572,404,429]" pageId="22" pageNumber="23" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 425
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3EB35EFD72FE41F593D4C9" box="[654,710,405,429]" pageId="22" pageNumber="23">Type</typeStatus>
locality: ins. St. Vincentii. 
<typeStatus id="7643887C4C3EB35EFBE1FE41F300D4C9" box="[1053,1109,405,429]" pageId="22" pageNumber="23">Type</typeStatus>
depository: ZMHB? #?ST. Distribution: Lesser Antilles (
<collectingCountry id="D1EF764E4C3EB35EFDD4FE6CF5C2D4B4" box="[552,663,439,464]" name="Saint Lucia" pageId="22" pageNumber="23">St. Lucia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3EB35EFD55FE6CF47AD4B4" box="[681,815,439,464]" name="Saint Vincent and the Grenadines" pageId="22" pageNumber="23">St. Vincent</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3EB35EFCBDFE63F4F1D4B4" box="[833,932,439,464]" name="Grenada" pageId="22" pageNumber="23">Grenada</collectingCountry>
), 
<collectingCountry id="D1EF764E4C3EB35EFC42FE63F353D4B4" box="[958,1030,439,464]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
. Notes: distribution within Lesser Antilles (
<collectingCountry id="D1EF764E4C3EB35EFED1FE08F6F9D491" box="[301,428,476,501]" name="Saint Vincent and the Grenadines" pageId="22" pageNumber="23">St. Vincent</collectingCountry>
) (
<bibRefCitation id="CD694B2F4C3EB35EFE3DFE08F56BD490" box="[449,574,476,501]" pageId="22" pageNumber="23" refId="ref71023" refString="Peck, S. B. (2010) The beetles of the island of St. Vincent, Lesser Antilles (Insecta: Coleoptera); diversity and distributions. Insecta Mundi, 0 144, 1 - 77." type="journal article">Peck 2010</bibRefCitation>
); within (and including) 
<collectingCountry id="D1EF764E4C3EB35EFCAEFE08F4C3D491" box="[850,918,476,501]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C3EB35EFC01FE09F360D491" box="[1021,1077,476,501]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFDF7F402D758" blockId="22.[151,1436,547,968]" box="[151,855,547,573]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFDF7F7F6D758" bold="true" box="[151,163,547,572]" italics="true" pageId="22" pageNumber="23">*</emphasis>
<taxonomicName id="6EF84D5D4C3EB35EFF5EFDF7F57BD759" ID-CoL="LC9R" authority="(Sharp 1876)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1876" box="[162,558,547,573]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="muticus" status="comb. nov.">
<emphasis id="9B8CEACC4C3EB35EFF5EFDF7F6C7D758" bold="true" box="[162,402,547,572]" italics="true" pageId="22" pageNumber="23">Belonuchus muticus</emphasis>
(Sharp 1876)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C3EB35EFDC6FDF0F5EFD759" box="[570,698,548,573]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFDC6FDF0F5EFD759" bold="true" box="[570,698,548,573]" pageId="22" pageNumber="23">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C3EB35EFD36FDF0F41AD758" box="[714,847,548,573]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="22" pageNumber="23">Figs. 11–12</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFD93F380D7E0" blockId="22.[151,1436,547,968]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BFD93F380D7E0" infered="true" pageId="22" pageNumber="23">
<materialsCitation id="19903C834C3EB35EFF6BFD93F529D7E0" ID-GBIF-Occurrence="1915497927" collectionCode="MCP" country="Brazil" county="Amazonas" location="Ega" municipality="Distribution" pageId="22" pageNumber="23" specimenCode="BMNH 4" specimenCount="2" stateProvince="Sao Paulo" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFD93F569D704" ID-CoL="4GBBK" authority="Sharp 1876 a: 167" authorityName="Sharp" authorityPageNumber="167" authorityYear="1876" box="[151,572,583,608]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="muticus">
<emphasis id="9B8CEACC4C3EB35EFF6BFD93F63BD704" box="[151,366,583,608]" italics="true" pageId="22" pageNumber="23">Philonthus muticus</emphasis>
<treatmentCitation id="285910CF4C3EB35EFE88FD93F569D704" author="" box="[372,572,583,608]" page="167" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFE88FD93F550D704" box="[372,517,583,608]" pageId="22" pageNumber="23" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 167
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3EB35EFDBAFD9CF52BD704" box="[582,638,584,608]" pageId="22" pageNumber="23">Type</typeStatus>
locality: Ega. 
<typeStatus id="7643887C4C3EB35EFCDCFD9CF402D704" box="[800,855,584,608]" pageId="22" pageNumber="23">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3EB35EFC1EFD9CF31ED704" box="[994,1099,584,608]" collectionCode="BMNH" pageId="22" pageNumber="23">BMNH 4</specimenCode>
ST: 3 
<location id="AC2760054C3EB35EFB6FFD9CF394D704" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3EB35EFB6FFD9CF394D704" box="[1171,1217,584,608]" country="Brazil" county="Amazonas" municipality="Distribution" name="Ega" pageId="22" pageNumber="23" stateProvince="Sao Paulo">Ega</location>
, 1 
<collectingRegion id="6B3CF83C4C3EB35EFB1DFD9CF204D704" box="[1249,1361,583,608]" country="Brazil" name="Sao Paulo" pageId="22" pageNumber="23">Sao Paulo</collectingRegion>
[
<collectionCode id="CFE9AE1B4C3EB35EFAA2FD9CF2C9D704" box="[1374,1436,584,608]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="22" pageNumber="23">MCP</collectionCode>
seen]. 
<collectingMunicipality id="4923ACA44C3EB35EFEF2FDB8F6CFD7E1" box="[270,410,620,645]" pageId="22" pageNumber="23">Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C3EB35EFE58FDB8F6BDD7E1" box="[420,488,620,645]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
(
<collectingCounty id="40264E524C3EB35EFE0BFDB8F521D7E0" box="[503,628,620,644]" pageId="22" pageNumber="23">Amazonas</collectingCounty>
)
</materialsCitation>
. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C3EB35EFBB1FDB9F3D0D7E1" box="[1101,1157,620,645]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BFD5BF546D794" pageId="22" pageNumber="23" type="discussion">
<paragraph id="A94736DE4C3EB35EFF6BFD5BF546D794" blockId="22.[151,1436,547,968]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFD5BF65ED7CC" bold="true" box="[151,267,655,680]" pageId="22" pageNumber="23">Remarks.</emphasis>
Sharp explicitly mentioned four specimens of both sexes as part of the 
<typeStatus id="7643887C4C3EB35EFBD0FD44F309D7CC" box="[1068,1116,656,680]" pageId="22" pageNumber="23">type</typeStatus>
series. The specimens listed below agree with the original description by 
<bibRefCitation id="CD694B2F4C3EB35EFD4BFD60F403D7A9" box="[695,854,692,717]" pageId="22" pageNumber="23" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp (1876a)</bibRefCitation>
and is considered to have been 
<typeStatus id="7643887C4C3EB35EFB44FD61F24FD7A9" box="[1208,1306,693,717]" pageId="22" pageNumber="23" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C3EB35EFEE1FD03F6D6D794" box="[285,387,727,752]" pageId="22" pageNumber="23" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BFD28F5D6D6C1" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="A94736DE4C3EB35EFF6BFD28F5D6D6C1" blockId="22.[151,1436,547,968]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFD28F694D671" bold="true" box="[151,449,764,789]" pageId="22" pageNumber="23">
<typeStatus id="7643887C4C3EB35EFF6BFD28F787D671" box="[151,210,764,789]" pageId="22" pageNumber="23">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C3EB35EFE35FD29F56CD671" box="[457,569,765,789]" pageId="22" pageNumber="23" type="lectotype">Lectotype</typeStatus>
<specimenCount id="BFFEFD574C3EB35EFDBDFD29F50DD670" box="[577,600,765,788]" pageId="22" pageNumber="23" type="male">♂</specimenCount>
, here designated (
<collectionCode id="CFE9AE1B4C3EB35EFCD8FD28F428D670" box="[804,893,764,788]" pageId="22" pageNumber="23">BMNH</collectionCode>
, 
<figureCitation id="31C32A5B4C3EB35EFC75FD28F347D670" box="[905,1042,764,789]" captionStart="FIGURES 1–12" captionStartId="9.[151,264,1848,1871]" captionTargetBox="[227,1359,267,1824]" captionTargetId="figure@9.[227,1359,267,1824]" captionTargetPageId="9" captionText="FIGURES 1–12 [BMNH ©]. Type material (habitus and labels): 1, 2, Philonthus abnormalis Sharp, lectotype; 3, 4, Philonthus celatus Sharp, lectotype; 5, 6, Philonthus corticalis Sharp, holotype; 7, 8, Philonthus extremus Sharp, holotype; 9, 10, Philonthus latecinctus Sharp, lectotype; 11, 12, Philonthus muticus Sharp, lectotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454307/files/figure.png" pageId="22" pageNumber="23">Figs. 11–12</figureCitation>
): card-mounted, “ 
<specimenCount id="BFFEFD574C3EB35EFB21FD28F3A6D670" box="[1245,1267,764,788]" pageId="22" pageNumber="23" type="male">♂</specimenCount>
// 
<typeStatus id="7643887C4C3EB35EFAEEFD29F21FD671" box="[1298,1354,765,789]" pageId="22" pageNumber="23">Type</typeStatus>
(round white label with red margin) // Ega (round green label) [in Sharp’s hand]// S America 
<collectingCountry id="D1EF764E4C3EB35EFB47FCCBF3AAD65C" box="[1211,1279,799,824]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
// 
<taxonomicName id="6EF84D5D4C3EB35EFAD8FCCBF64BD639" authorityName="Sharp" authorityYear="1876" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="muticus">Philonthus muticus</taxonomicName>
<typeStatus id="7643887C4C3EB35EFEDBFC91F60AD639" box="[295,351,837,861]" pageId="22" pageNumber="23">Type</typeStatus>
D.S. [in Sharp’s hand] // Sharp Coll. 1905.-313. (white label)”. 
<typeStatus id="7643887C4C3EB35EFBB6FC90F3B8D639" box="[1098,1261,836,861]" pageId="22" pageNumber="23" type="paralectotype">Paralectotypes</typeStatus>
(
<collectionCode id="CFE9AE1B4C3EB35EFB03FC90F20FD638" box="[1279,1370,836,860]" pageId="22" pageNumber="23">BMNH</collectionCode>
): 
<specimenCount id="BFFEFD574C3EB35EFA93FC90F2C9D638" box="[1391,1436,836,860]" pageId="22" pageNumber="23" type="female">2 ♀</specimenCount>
(one without head), card mounted, same locality and collection data as 
<typeStatus id="7643887C4C3EB35EFBDEFCB3F3DCD6E4" box="[1058,1161,871,896]" pageId="22" pageNumber="23" type="lectotype">lectotype</typeStatus>
but with “ 
<typeStatus id="7643887C4C3EB35EFAF3FCBCF2C7D6E4" box="[1295,1426,872,896]" pageId="22" pageNumber="23" type="syntype">SYNTYPE</typeStatus>
” (rounded white label with blue margin).
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BFC7BF33DD6AC" box="[151,1128,943,968]" pageId="22" pageNumber="23" type="discussion">
<paragraph id="A94736DE4C3EB35EFF6BFC7BF33DD6AC" blockId="22.[151,1436,547,968]" box="[151,1128,943,968]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFC7BF64AD6AC" bold="true" box="[151,287,943,968]" pageId="22" pageNumber="23">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C3EB35EFD46FC7BF331D6AC" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,943,968]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3EB35EFD46FC7BF468D6AC" box="[698,829,943,968]" italics="true" pageId="22" pageNumber="23">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C3EB35EFCB7FC7BF309D6AC" box="[843,1116,943,968]" pageId="22" pageNumber="23" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C3EB35EFC1CFC65F34DD6AC" box="[992,1048,943,968]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BFC23F3CAD01C" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="A94736DE4C3EB35EFF6BFC23F525D174" blockId="22.[151,1437,1015,1112]" box="[151,624,1015,1040]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFC23F7F6D174" bold="true" box="[151,163,1015,1040]" italics="true" pageId="22" pageNumber="23">*</emphasis>
<taxonomicName id="6EF84D5D4C3EB35EFF5EFC23F525D174" ID-CoL="LC9T" authority="Bernhauer 1918" authorityName="Bernhauer" authorityYear="1918" box="[162,624,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="9B8CEACC4C3EB35EFF5EFC23F6E0D174" bold="true" box="[162,437,1015,1040]" italics="true" pageId="22" pageNumber="23">Belonuchus nigerrimus</emphasis>
<bibRefCitation id="CD694B2F4C3EB35EFE41FC23F525D174" box="[445,624,1015,1040]" pageId="22" pageNumber="23" refId="ref62367" refString="Bernhauer, M. (1918) Vier neue Belonuchus aus Mexiko [cont.]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 67, 225 - 226." type="journal article">Bernhauer 1918</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFBC8F26CD13C" blockId="22.[151,1437,1015,1112]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BFBC8F26CD13C" pageId="22" pageNumber="23">
<materialsCitation id="19903C834C3EB35EFF6BFBC8F380D150" ID-GBIF-Occurrence="1915497911" box="[151,1237,1052,1077]" country="Mexico" location="Jalappa" municipality="Xalapa" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFBC8F5C7D151" ID-CoL="LC9T" authority="Bernhauer 1918: 225" authorityName="Bernhauer" authorityPageNumber="225" authorityYear="1918" box="[151,658,1052,1077]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="9B8CEACC4C3EB35EFF6BFBC8F6CED150" box="[151,411,1052,1077]" italics="true" pageId="22" pageNumber="23">Belonuchus nigerrimus</emphasis>
<treatmentCitation id="285910CF4C3EB35EFE5EFBC8F5C7D151" author="" box="[418,658,1052,1077]" page="225" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFE5EFBC8F50FD150" box="[418,602,1052,1077]" pageId="22" pageNumber="23" refId="ref62367" refString="Bernhauer, M. (1918) Vier neue Belonuchus aus Mexiko [cont.]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 67, 225 - 226." type="journal article">Bernhauer 1918</bibRefCitation>
: 225
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3EB35EFD61FBC9F580D151" box="[669,725,1053,1077]" pageId="22" pageNumber="23">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3EB35EFCC2FBC8F4CDD151" box="[830,920,1052,1077]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
: 
<location id="AC2760054C3EB35EFC5FFBC8F4A2D151" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3EB35EFC5FFBC8F4A2D151" box="[931,1015,1052,1077]" country="Mexico" municipality="Xalapa" name="Jalappa" pageId="22" pageNumber="23" stateProvince="Veracruz">Jalappa</location>
[
<collectingMunicipality id="4923ACA44C3EB35EFBF8FBC8F30CD151" box="[1028,1113,1052,1077]" pageId="22" pageNumber="23">Xalapa</collectingMunicipality>
, 
<collectingRegion id="6B3CF83C4C3EB35EFB99FBC9F39AD150" box="[1125,1231,1053,1076]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="22" pageNumber="23">Veracruz</collectingRegion>
]
</materialsCitation>
. 
<materialsCitation id="19903C834C3EB35EFB1CFBC9F51BD13C" ID-GBIF-Occurrence="1915497904" collectionCode="FMNH" country="Mexico" location="ST. Distribution" pageId="22" pageNumber="23" specimenCode="FMNH 2" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3EB35EFB1CFBC9F242D151" box="[1248,1303,1053,1077]" pageId="22" pageNumber="23">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C3EB35EFF39FB94F67ED13C" box="[197,299,1088,1112]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="22" pageNumber="23">FMNH 2</specimenCode>
<location id="AC2760054C3EB35EFECEFB94F6BFD13C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3EB35EFECEFB94F6BFD13C" box="[306,490,1087,1112]" country="Mexico" name="ST. Distribution" pageId="22" pageNumber="23">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C3EB35EFE09FBEBF51BD13C" box="[501,590,1087,1112]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C3EB35EFC24FBEBF278D13C" box="[984,1325,1087,1112]" pageId="22" pageNumber="23" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3EB35EFB4DFB95F3BCD13C" box="[1201,1257,1087,1112]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFB53F501D1C4" blockId="22.[151,1437,1159,1256]" box="[151,596,1159,1184]" pageId="22" pageNumber="23">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFB53F501D1C4" ID-CoL="68GHW" authority="Blackwelder, 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,596,1159,1184]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="oakleyi">
<emphasis id="9B8CEACC4C3EB35EFF6BFB53F62FD1C4" bold="true" box="[151,378,1159,1184]" italics="true" pageId="22" pageNumber="23">Belonuchus oakleyi</emphasis>
<bibRefCitation id="CD694B2F4C3EB35EFE7EFB53F501D1C4" box="[386,596,1159,1184]" pageId="22" pageNumber="23" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder, 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFB78F6BDD18C" blockId="22.[151,1437,1159,1256]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BFB78F6BDD18C" pageId="22" pageNumber="23">
<materialsCitation id="19903C834C3EB35EFF6BFB78F313D1A1" ID-GBIF-Occurrence="1915497921" box="[151,1094,1196,1221]" country="Puerto Rico" location="Mayaguez" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Mayaguez" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFB78F5D2D1A0" ID-CoL="68GHW" authority="Blackwelder 1943: 422" authorityName="Blackwelder" authorityPageNumber="422" authorityYear="1943" box="[151,647,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="oakleyi">
<emphasis id="9B8CEACC4C3EB35EFF6BFB78F626D1A1" box="[151,371,1196,1221]" italics="true" pageId="22" pageNumber="23">Belonuchus oakleyi</emphasis>
<treatmentCitation id="285910CF4C3EB35EFE87FB78F5D2D1A0" author="" box="[379,647,1196,1221]" page="422" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFE87FB78F518D1A0" box="[379,589,1196,1221]" pageId="22" pageNumber="23" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 422
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3EB35EFD6FFB79F59FD1A1" box="[659,714,1197,1221]" pageId="22" pageNumber="23">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C3EB35EFCCBFB79F497D1A1" box="[823,962,1196,1221]" name="Puerto Rico" pageId="22" pageNumber="23">Puerto Rico</collectingCountry>
: 
<collectingRegion id="6B3CF83C4C3EB35EFC32FB79F313D1A1" box="[974,1094,1197,1221]" country="Puerto Rico" name="Mayaguez" pageId="22" pageNumber="23">Mayagüez</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C3EB35EFBAEFB79F6B1D18C" ID-GBIF-Occurrence="1915497901" collectionCode="USNM" country="Puerto Rico" location="Distribution" pageId="22" pageNumber="23" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C3EB35EFBAEFB79F3DCD1A1" box="[1106,1161,1197,1221]" pageId="22" pageNumber="23">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3EB35EFAE4FB78F23CD1A0" box="[1304,1385,1196,1220]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23">USNM</collectionCode>
HT. 
<location id="AC2760054C3EB35EFF39FB1BF604D18C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3EB35EFF39FB1BF604D18C" box="[197,337,1231,1256]" country="Puerto Rico" name="Distribution" pageId="22" pageNumber="23">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3EB35EFEA7FB04F6B1D18C" box="[347,484,1231,1256]" name="Puerto Rico" pageId="22" pageNumber="23">Puerto Rico</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFAC3F402D054" blockId="22.[151,1436,1303,1616]" box="[151,855,1303,1328]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFAC3F7F6D054" bold="true" box="[151,163,1303,1328]" italics="true" pageId="22" pageNumber="23">*</emphasis>
<taxonomicName id="6EF84D5D4C3EB35EFF5EFAC3F578D054" ID-CoL="68GHD" authority="(Sharp, 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,557,1303,1328]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="optatus" status="comb. nov.">
<emphasis id="9B8CEACC4C3EB35EFF5EFAC3F6CFD054" bold="true" box="[162,410,1303,1328]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF5EFAC3F6DFD04B" bold="true" box="[162,394,1303,1328]" italics="true" pageId="22" pageNumber="23">Belonuchus optatus</emphasis>
(
</emphasis>
<bibRefCitation id="CD694B2F4C3EB35EFE66FAC3F570D054" box="[410,549,1303,1328]" pageId="22" pageNumber="23" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp, 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C3EB35EFDC5FAC3F5ECD054" box="[569,697,1303,1328]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFDC5FAC3F5ECD054" bold="true" box="[569,697,1303,1328]" pageId="22" pageNumber="23">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C3EB35EFD34FAC3F41AD054" box="[712,847,1303,1328]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="22" pageNumber="23">Figs. 13–14</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BFAE8F3CAD01C" blockId="22.[151,1436,1303,1616]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BFAE8F3CAD01C" pageId="22" pageNumber="23">
<materialsCitation id="19903C834C3EB35EFF6BFAE8F43ED031" ID-GBIF-Occurrence="1915497876" box="[151,875,1340,1365]" country="Mexico" location="Mexico" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3EB35EFF6BFAE8F518D030" ID-CoL="76Z55" authority="Sharp 1885: 427" authorityName="Sharp" authorityPageNumber="427" authorityYear="1885" box="[151,589,1340,1365]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="optatus">
<emphasis id="9B8CEACC4C3EB35EFF6BFAE8F627D030" box="[151,370,1340,1365]" italics="true" pageId="22" pageNumber="23">Philonthus optatus</emphasis>
<treatmentCitation id="285910CF4C3EB35EFE7DFAE8F518D030" author="" box="[385,589,1340,1365]" page="427" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFE7DFAE8F558D031" box="[385,525,1340,1365]" pageId="22" pageNumber="23" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 427
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3EB35EFD9CFAE9F5C2D031" box="[608,663,1341,1365]" pageId="22" pageNumber="23">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C3EB35EFCEDFAE8F43ED031" box="[785,875,1340,1365]" country="Mexico" name="Mexico" pageId="22" pageNumber="23">
<collectingCountry id="D1EF764E4C3EB35EFCEDFAE8F43ED031" box="[785,875,1340,1365]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
</collectingRegion>
</materialsCitation>
: 
<materialsCitation id="19903C834C3EB35EFC82FAE8F356D031" ID-GBIF-Occurrence="1915497928" box="[894,1027,1340,1365]" country="Mexico" location="Guanajuato" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Guanajuato" typeStatus="holotype">
<collectingRegion id="6B3CF83C4C3EB35EFC82FAE8F356D031" box="[894,1027,1340,1365]" country="Mexico" name="Guanajuato" pageId="22" pageNumber="23">Guanajuato</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C3EB35EFBE9FAE9F6E1D01C" ID-GBIF-Occurrence="1915497882" collectionCode="BMNH" country="Mexico" location="Distribution" pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1" stateProvince="Mexico" typeStatus="holotype">
<typeStatus id="7643887C4C3EB35EFBE9FAE9F318D031" box="[1045,1101,1341,1365]" pageId="22" pageNumber="23">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3EB35EFB14FAE9F269D030" box="[1256,1340,1341,1364]" pageId="22" pageNumber="23">BMNH</collectionCode>
HT 
<specimenCount id="BFFEFD574C3EB35EFA7CFAE9F2C3D030" box="[1408,1430,1341,1364]" pageId="22" pageNumber="23" type="female">♀</specimenCount>
. 
<location id="AC2760054C3EB35EFF39FA8BF604D01C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3EB35EFF39FA8BF604D01C" box="[197,337,1375,1400]" country="Mexico" name="Distribution" pageId="22" pageNumber="23" stateProvince="Mexico">Distribution</location>
: 
<collectingRegion id="6B3CF83C4C3EB35EFEA7FA8BF6E1D01C" box="[347,436,1375,1400]" country="Mexico" name="Mexico" pageId="22" pageNumber="23">
<collectingCountry id="D1EF764E4C3EB35EFEA7FA8BF6E1D01C" box="[347,436,1375,1400]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
</collectingRegion>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C3EB35EFCC2FA8BF3C6D01C" box="[830,1171,1375,1400]" pageId="22" pageNumber="23" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3EB35EFBEBFAB5F305D01C" box="[1047,1104,1375,1400]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BFA50F48BD0F9" box="[151,990,1412,1437]" pageId="22" pageNumber="23" type="discussion">
<paragraph id="A94736DE4C3EB35EFF6BFA50F48BD0F9" blockId="22.[151,1436,1303,1616]" box="[151,990,1412,1437]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFA50F65ED0F9" bold="true" box="[151,267,1412,1437]" pageId="22" pageNumber="23">Remarks.</emphasis>
The “unique” specimen in BMNH is confirmed as the 
<typeStatus id="7643887C4C3EB35EFC88FA50F48FD0F9" box="[884,986,1412,1437]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BFA73F452D349" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="A94736DE4C3EB35EFF6BFA73F452D349" blockId="22.[151,1436,1303,1616]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BFA73F69FD0A4" bold="true" box="[151,458,1447,1472]" pageId="22" pageNumber="23">Type material examined.</emphasis>
Holotype ♀ (BMNH, 
<figureCitation id="31C32A5B4C3EB35EFD23FA73F43BD0A4" box="[735,878,1447,1472]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="22" pageNumber="23">Figs. 13–14</figureCitation>
): card-mounted, “ 
<taxonomicName id="6EF84D5D4C3EB35EFBBDFA73F24CD0DB" box="[1089,1305,1447,1472]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="optatus">Philonthus optatus</taxonomicName>
Type D.S. Guanajuato. Mexico. Sallé. [in Sharp’s hand] // Type (round white label with red margin) //Guanajuato (white label) // Mexico. Salle Coll. (white label) // B. C. A. Col. 
<date id="DD46101E4C3EB35EFC86FA24F4FED36C" box="[890,939,1520,1544]" pageId="22" pageNumber="23">I. 2.</date>
<taxonomicName id="6EF84D5D4C3EB35EFC4AFA3BF3B0D36C" authority=", Sharp." authorityName="Sharp." box="[950,1253,1519,1544]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="optatus">Philonthus optatus, Sharp.</taxonomicName>
(white label) // 
<taxonomicName id="6EF84D5D4C3EB35EFF39F9C0F6EDD349" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[197,440,1556,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="formosus">Belonuchus formosus</taxonomicName>
apud Sallé [in Sallé’s hand]”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BF9E3F33DD334" box="[151,1128,1591,1616]" pageId="22" pageNumber="23" type="discussion">
<paragraph id="A94736DE4C3EB35EFF6BF9E3F33DD334" blockId="22.[151,1436,1303,1616]" box="[151,1128,1591,1616]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BF9E3F64AD334" bold="true" box="[151,287,1591,1616]" pageId="22" pageNumber="23">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C3EB35EFD46F9E3F331D334" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1591,1616]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3EB35EFD46F9E3F468D334" box="[698,829,1591,1616]" italics="true" pageId="22" pageNumber="23">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C3EB35EFCB7F9E3F309D334" box="[843,1116,1591,1616]" pageId="22" pageNumber="23" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C3EB35EFC1CF9EDF34DD334" box="[992,1048,1591,1616]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BF9ABF58AD2F1" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="A94736DE4C3EB35EFF6BF9ABF504D3FC" blockId="22.[151,1436,1663,1797]" box="[151,593,1663,1688]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BF9ABF7F6D3FC" bold="true" box="[151,163,1663,1688]" italics="true" pageId="22" pageNumber="23">*</emphasis>
<taxonomicName id="6EF84D5D4C3EB35EFF5EF9ABF504D3FC" ID-CoL="5WGHC" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,593,1663,1688]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="oxyporinus">
<emphasis id="9B8CEACC4C3EB35EFF5EF9ABF6E2D3FC" bold="true" box="[162,439,1663,1688]" italics="true" pageId="22" pageNumber="23">Belonuchus oxyporinus</emphasis>
(
<bibRefCitation id="CD694B2F4C3EB35EFE3AF9ABF51CD3FC" box="[454,585,1663,1688]" pageId="22" pageNumber="23" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BF970F3AAD260" blockId="22.[151,1436,1663,1797]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BF970F3AAD260" infered="true" pageId="22" pageNumber="23">
<taxonomicName id="6EF84D5D4C3EB35EFF6BF970F52DD3D8" ID-CoL="76Z4X" authority="Sharp 1885: 419" authorityName="Sharp" authorityPageNumber="419" authorityYear="1885" box="[151,632,1700,1725]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="oxyporinus">
<emphasis id="9B8CEACC4C3EB35EFF6BF970F6C9D3D8" box="[151,412,1700,1725]" italics="true" pageId="22" pageNumber="23">Philonthus oxyporinus</emphasis>
<treatmentCitation id="285910CF4C3EB35EFE57F970F52DD3D8" author="" box="[427,632,1700,1725]" page="419" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFE57F970F56DD3D9" box="[427,568,1700,1725]" pageId="22" pageNumber="23" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 419
</treatmentCitation>
</taxonomicName>
. Type locality: Mexico: Oaxaca. Type depository: BMNH HT ♀. Distribution: Mexico. Notes: to 
<taxonomicName id="6EF84D5D4C3EB35EFDC7F913F5EBD384" ID-CoL="8VRHV" box="[571,702,1735,1760]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3EB35EFDC7F913F5EBD384" box="[571,702,1735,1760]" italics="true" pageId="22" pageNumber="23">Belonuchus</emphasis>
</taxonomicName>
(after 
<bibRefCitation id="CD694B2F4C3EB35EFCECF91CF4E2D384" box="[784,951,1735,1760]" pageId="22" pageNumber="23" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995</bibRefCitation>
); distribution, biology (Navarrete-Heredia 1995, 
<bibRefCitation id="CD694B2F4C3EB35EFEF7F938F505D260" box="[267,592,1772,1797]" pageId="22" pageNumber="23" refId="ref67947" refString="Jimenez-Sanchez, E., Navarrete-Heredia, J. L. &amp; Padilla-Ramirez, J. R. (2000) Estafilinidos (Coleoptera: Staphylinidae) necrofilos de la Sierra de Nanchititla, Estado de Mexico, Mexico. Folia Entomologica Mexicana, 108, 53 - 78." type="journal article">
Jiménez-Sánchez 
<emphasis id="9B8CEACC4C3EB35EFE28F939F559D261" box="[468,524,1772,1797]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2000
</bibRefCitation>
); distribution within Mexico (
<bibRefCitation id="CD694B2F4C3EB35EFC62F938F3A6D260" box="[926,1267,1772,1797]" pageId="22" pageNumber="23" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C3EB35EFB8BF939F3FAD261" box="[1143,1199,1772,1797]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BF8E7F513D229" blockId="22.[151,1436,1843,1977]" box="[151,582,1843,1869]" pageId="22" pageNumber="23">
<taxonomicName id="6EF84D5D4C3EB35EFF6BF8E7F513D229" ID-CoL="68GHJ" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,582,1843,1869]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="palustris">
<emphasis id="9B8CEACC4C3EB35EFF6BF8E7F6D9D228" bold="true" box="[151,396,1843,1868]" italics="true" pageId="22" pageNumber="23">Belonuchus palustris</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3EB35EFF6BF883F58AD2F1" blockId="22.[151,1436,1843,1977]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="89E811F04C3EB35EFF6BF883F58AD2F1" pageId="22" pageNumber="23">
<materialsCitation id="19903C834C3EB35EFF6BF883F58ED2F1" ID-GBIF-Occurrence="1915497859" collectionCode="FMNH" country="Colombia" elevation="2200" location="Distribution" pageId="22" pageNumber="23" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C3EB35EFF6BF883F41AD214" ID-CoL="68GHJ" authority="Bernhauer 1916 c: 268" authorityName="Bernhauer" authorityPageNumber="268" authorityYear="1916" box="[151,847,1879,1904]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="palustris" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C3EB35EFF6BF883F56CD214" box="[151,569,1879,1904]" italics="true" pageId="22" pageNumber="23">Belonuchus (Musicoderus) palustris</emphasis>
<treatmentCitation id="285910CF4C3EB35EFDBAF883F41AD214" author="" box="[582,847,1879,1904]" page="268" pageId="22" pageNumber="23">
<bibRefCitation id="CD694B2F4C3EB35EFDBAF883F444D214" box="[582,785,1879,1904]" pageId="22" pageNumber="23" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 268
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3EB35EFC9CF88CF4CDD214" box="[864,920,1880,1904]" pageId="22" pageNumber="23">Type</typeStatus>
locality: Alto de las Cruces, 
<quantity id="6E009B3B4C3EB35EFAFEF88CF201D214" box="[1282,1364,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="23" unit="m" value="2200.0">
<elevation id="22D5D1ED4C3EB35EFAFEF88CF201D214" box="[1282,1364,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="23" unit="m" value="2200.0">2200m</elevation>
</quantity>
. 
<typeStatus id="7643887C4C3EB35EFA98F88CF2C9D214" box="[1380,1436,1880,1904]" pageId="22" pageNumber="23">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C3EB35EFEB6F8A9F6CED2F0" box="[330,411,1917,1940]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="22" pageNumber="23">FMNH</collectionCode>
HT. 
<location id="AC2760054C3EB35EFE2FF8A8F50AD2F1" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C3EB35EFE2FF8A8F50AD2F1" box="[467,607,1916,1941]" country="Colombia" name="Distribution" pageId="22" pageNumber="23">Distribution</location>
: 
<collectingCountry id="D1EF764E4C3EB35EFD95F8A8F58ED2F1" box="[617,731,1916,1941]" name="Colombia" pageId="22" pageNumber="23">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C3EB35EFF6BF874F465D2DD" box="[151,816,1952,1977]" pageId="22" pageNumber="23" type="discussion">
<paragraph id="A94736DE4C3EB35EFF6BF874F465D2DD" blockId="22.[151,1436,1843,1977]" box="[151,816,1952,1977]" pageId="22" pageNumber="23">
<emphasis id="9B8CEACC4C3EB35EFF6BF874F641D2DD" bold="true" box="[151,276,1952,1977]" pageId="22" pageNumber="23">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C3EB35EFEB0F874F69AD2DD" box="[332,463,1952,1977]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C3EB35EFEB0F874F69AD2DD" box="[332,463,1952,1977]" italics="true" pageId="22" pageNumber="23">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<caption id="FD8766564C3FB35FFF6BF916F2DDD272" httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,264,1730,1753]" targetBox="[227,1360,195,1709]" targetPageId="23">
<paragraph id="A94736DE4C3FB35FFF6BF916F2DDD272" blockId="23.[151,1436,1730,1814]" pageId="23" pageNumber="24">
<emphasis id="9B8CEACC4C3FB35FFF6BF916F606D3BC" bold="true" box="[151,339,1730,1753]" pageId="23" pageNumber="24">FIGURES 13–24</emphasis>
[BMNH ©]. Type material (habitus and labels): 13, 14
<taxonomicName id="6EF84D5D4C3FB35FFC4AF917F381D3BD" authority="Sharp" authorityName="Sharp" box="[950,1236,1731,1753]" pageId="23" pageNumber="24" rank="subSpecies" subSpecies="optatus">
, 
<emphasis id="9B8CEACC4C3FB35FFC3BF917F3D9D3BD" box="[967,1164,1731,1753]" italics="true" pageId="23" pageNumber="24">Philonthus optatus</emphasis>
Sharp
</taxonomicName>
, holotype; 15, 16
<taxonomicName id="6EF84D5D4C3FB35FFA6BF917F697D393" authority="Sharp" authorityName="Sharp" pageId="23" pageNumber="24" rank="subSpecies" subSpecies="platypterus">
, 
<emphasis id="9B8CEACC4C3FB35FFF6BF935F62BD393" box="[151,382,1761,1783]" italics="true" pageId="23" pageNumber="24">Philonthus platypterus</emphasis>
Sharp
</taxonomicName>
, lectotype; 17, 18
<taxonomicName id="6EF84D5D4C3FB35FFD87F935F4FED393" authority="Sharp" authorityName="Sharp" box="[635,939,1761,1783]" pageId="23" pageNumber="24" rank="subSpecies" subSpecies="rufiventris">
, 
<emphasis id="9B8CEACC4C3FB35FFD74F935F432D393" box="[648,871,1761,1783]" italics="true" pageId="23" pageNumber="24">Philonthus rufiventris</emphasis>
Sharp
</taxonomicName>
, holotype; 19, 20
<taxonomicName id="6EF84D5D4C3FB35FFB9CF935F2CDD393" authority="Sharp" authorityName="Sharp" authorityYear="1885" box="[1120,1432,1761,1783]" class="Insecta" family="Staphylinidae" genus="Bisnius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="platypterus" subSpecies="rufocaudus">
, 
<emphasis id="9B8CEACC4C3FB35FFB90F935F206D393" box="[1132,1363,1761,1783]" italics="true" pageId="23" pageNumber="24">Philonthus rufocaudus</emphasis>
Sharp
</taxonomicName>
, lectotype; 21, 22
<taxonomicName id="6EF84D5D4C3FB35FFEBEF8D4F53CD272" authority="Sharp" authorityName="Sharp" box="[322,617,1792,1814]" pageId="23" pageNumber="24" rank="subSpecies" subSpecies="rufopygus">
, 
<emphasis id="9B8CEACC4C3FB35FFEB1F8D4F573D272" box="[333,550,1792,1814]" italics="true" pageId="23" pageNumber="24">Philonthus rufopygus</emphasis>
Sharp
</taxonomicName>
, lectotype; 23, 24
<taxonomicName id="6EF84D5D4C3FB35FFCDCF8D4F331D272" authority="Sharp" authorityName="Sharp" box="[800,1124,1792,1814]" pageId="23" pageNumber="24" rank="subSpecies" subSpecies="serraticornis">
, 
<emphasis id="9B8CEACC4C3FB35FFCD6F8D4F374D272" box="[810,1057,1792,1814]" italics="true" pageId="23" pageNumber="24">Philonthus serraticornis</emphasis>
Sharp
</taxonomicName>
, holotype. Scale bar= 2 mm.
</paragraph>
</caption>
<subSubSection id="E1E265554C3FB350FF6BF8E9F591D4C8" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="A94736DE4C3FB35FFF6BF8E9F513D232" blockId="23.[151,1436,1853,1986]" box="[151,582,1853,1878]" pageId="23" pageNumber="24">
<emphasis id="9B8CEACC4C3FB35FFF6BF8E9F7F6D232" bold="true" box="[151,163,1853,1878]" italics="true" pageId="23" pageNumber="24">*</emphasis>
<taxonomicName id="6EF84D5D4C3FB35FFF5EF8E9F513D232" ID-CoL="5WGHH" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,582,1853,1878]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="9B8CEACC4C3FB35FFF5EF8E9F6EAD232" bold="true" box="[162,447,1853,1878]" italics="true" pageId="23" pageNumber="24">Belonuchus panamensis</emphasis>
<bibRefCitation id="CD694B2F4C3FB35FFE3AF8E9F513D232" box="[454,582,1853,1878]" pageId="23" pageNumber="24" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C3FB35FFF6BF8B5F6F8D2A5" blockId="23.[151,1436,1853,1986]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="89E811F04C3FB35FFF6BF8B5F6F8D2A5" pageId="23" pageNumber="24">
<taxonomicName id="6EF84D5D4C3FB35FFF6BF8B5F52FD21E" ID-CoL="5WGHH" authority="Sharp 1885: 434" authorityName="Sharp" authorityPageNumber="434" authorityYear="1885" box="[151,634,1889,1914]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="9B8CEACC4C3FB35FFF6BF8B5F6FFD21D" box="[151,426,1889,1914]" italics="true" pageId="23" pageNumber="24">Belonuchus panamensis</emphasis>
<treatmentCitation id="285910CF4C3FB35FFE48F8B5F52FD21E" author="" box="[436,634,1889,1914]" page="434" pageId="23" pageNumber="24">
<bibRefCitation id="CD694B2F4C3FB35FFE48F8B5F568D21E" box="[436,573,1889,1914]" pageId="23" pageNumber="24" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 434
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C3FB35FFD75F8B6F594D21E" box="[649,705,1890,1914]" pageId="23" pageNumber="24">Type</typeStatus>
localities: 
<collectingCountry id="D1EF764E4C3FB35FFCB9F8B5F4EED21E" box="[837,955,1889,1914]" name="Nicaragua" pageId="23" pageNumber="24">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C3FB35FFC36F8B5F36BD21E" box="[970,1086,1889,1914]" country="Nicaragua" name="Chontales" pageId="23" pageNumber="24">Chontales</collectingRegion>
; 
<collectingRegion id="6B3CF83C4C3FB35FFBB2F8B6F3FFD21D" box="[1102,1194,1890,1913]" country="Panama" name="Panama" pageId="23" pageNumber="24">
<collectingCountry id="D1EF764E4C3FB35FFBB2F8B6F3FFD21D" box="[1102,1194,1890,1913]" name="Panama" pageId="23" pageNumber="24">Panama</collectingCountry>
</collectingRegion>
, Volcan de 
<collectingRegion id="6B3CF83C4C3FB35FFAC4F8B5F2CDD21E" box="[1336,1432,1889,1914]" country="Panama" name="Chiriqui" pageId="23" pageNumber="24">Chiriqui</collectingRegion>
, Bugaba, Tolé. 
<typeStatus id="7643887C4C3FB35FFE90F852F6F1D2FA" box="[364,420,1926,1950]" pageId="23" pageNumber="24">Type</typeStatus>
depository: BMNH 11 ST: from all 
<typeStatus id="7643887C4C3FB35FFCB6F852F42FD2FA" box="[842,890,1926,1950]" pageId="23" pageNumber="24">type</typeStatus>
localities [MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C3FB35FFADEF851F2CDD2FA" box="[1314,1432,1925,1950]" name="Nicaragua" pageId="23" pageNumber="24">Nicaragua</collectingCountry>
, 
<collectingCountry id="D1EF764E4C3FB35FFF39F87EF617D2A6" box="[197,322,1961,1986]" name="Costa Rica" pageId="23" pageNumber="24">
<collectingRegion id="6B3CF83C4C3FB35FFF39F87EF650D2A6" box="[197,261,1962,1986]" country="Tanzania" name="Pwani" pageId="23" pageNumber="24">Costa</collectingRegion>
Rica
</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C3FB35FFEB1F87EF6FCD2A5" box="[333,425,1962,1985]" country="Panama" name="Panama" pageId="23" pageNumber="24">
<collectingCountry id="D1EF764E4C3FB35FFEB1F87EF6FCD2A5" box="[333,425,1962,1985]" name="Panama" pageId="23" pageNumber="24">Panama</collectingCountry>
</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFF43F5EED5D4" blockId="24.[151,1436,151,285]" box="[151,699,151,176]" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFF43F5EED5D4" ID-CoL="68GHH" authority="Bernhauer 1921" authorityName="Bernhauer" authorityYear="1921" box="[151,699,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paradoxipunctatus">
<emphasis id="9B8CEACC4C30B350FF6BFF43F555D5D4" bold="true" box="[151,512,151,176]" italics="true" pageId="24" pageNumber="25">Belonuchus paradoxipunctatus</emphasis>
Bernhauer 1921
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFF68F44AD479" blockId="24.[151,1436,151,285]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFF68F44AD479" infered="true" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFF68F453D5B0" ID-CoL="68GHH" authority="Bernhauer 1921 c: 171" authorityName="Bernhauer" authorityPageNumber="171" authorityYear="1921" box="[151,774,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paradoxipunctatus">
<emphasis id="9B8CEACC4C30B350FF6BFF68F6A2D5B1" box="[151,503,188,213]" italics="true" pageId="24" pageNumber="25">Belonuchus paradoxipunctatus</emphasis>
<treatmentCitation id="285910CF4C30B350FDFEFF68F453D5B0" author="" box="[514,774,188,213]" page="171" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FDFEFF68F59FD5B0" box="[514,714,188,213]" pageId="24" pageNumber="25" refId="ref62483" refString="Bernhauer, M. (1921 c) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article">Bernhauer 1921c</bibRefCitation>
: 171
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FCE9FF69F418D5B1" box="[789,845,189,213]" pageId="24" pageNumber="25">Type</typeStatus>
locality: San Roque, 
<collectingRegion id="6B3CF83C4C30B350FBACFF68F390D5B1" box="[1104,1221,188,213]" country="Argentina" name="Corrientes" pageId="24" pageNumber="25">Corrientes</collectingRegion>
[
<collectingCountry id="D1EF764E4C30B350FB2BFF68F205D5B1" box="[1239,1360,188,213]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
]. 
<typeStatus id="7643887C4C30B350FA98FF69F2C9D5B1" box="[1380,1436,189,213]" pageId="24" pageNumber="25">Type</typeStatus>
depository: FMNH 1 ST. Distribution: 
<collectingCountry id="D1EF764E4C30B350FD68FF0BF453D59C" box="[660,774,223,248]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
, 
<collectingCountry id="D1EF764E4C30B350FCEBFF0BF40AD59C" box="[791,863,223,248]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
. Note: distribution within (and including) 
<collectingCountry id="D1EF764E4C30B350FAA4FF0BF2C9D59C" box="[1368,1436,223,248]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C30B350FED8FED1F608D479" box="[292,349,260,285]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2013). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFE9FF509D401" blockId="24.[151,1437,331,464]" box="[151,604,331,357]" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFE9FF509D401" ID-CoL="5WGHG" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[151,604,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="9B8CEACC4C30B350FF6BFE9FF6F4D400" bold="true" box="[151,417,331,356]" italics="true" pageId="24" pageNumber="25">Belonuchus paradoxus</emphasis>
<bibRefCitation id="CD694B2F4C30B350FE55FE98F509D401" box="[425,604,332,357]" pageId="24" pageNumber="25" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFEBBF591D4C8" blockId="24.[151,1437,331,464]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFEBBF591D4C8" pageId="24" pageNumber="25">
<materialsCitation id="19903C834C30B350FF6BFEBBF372D4E3" ID-GBIF-Occurrence="1915497850" box="[151,1063,367,392]" country="Peru" location="Pachitea" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C30B350FF6BFEBBF5E7D4EC" ID-CoL="5WGHG" authority="Bernhauer 1906: 333" authorityName="Bernhauer" authorityPageNumber="333" authorityYear="1906" box="[151,690,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="9B8CEACC4C30B350FF6BFEBBF6F5D4EC" box="[151,416,367,392]" italics="true" pageId="24" pageNumber="25">Belonuchus paradoxus</emphasis>
<treatmentCitation id="285910CF4C30B350FE4CFEBBF5E7D4EC" author="" box="[432,690,367,392]" page="333" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE4CFEBBF524D4EC" box="[432,625,367,392]" pageId="24" pageNumber="25" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 333
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FD3AFEA4F5ABD4EC" box="[710,766,368,392]" pageId="24" pageNumber="25">Type</typeStatus>
locality: 
<location id="AC2760054C30B350FC86FEBBF489D4EC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FC86FEBBF489D4EC" box="[890,988,367,392]" country="Peru" name="Pachitea" pageId="24" pageNumber="25">Pachitea</location>
, 
<collectingCountry id="D1EF764E4C30B350FC0CFEA4F372D4E3" box="[1008,1063,368,391]" name="Peru" pageId="24" pageNumber="25">Peru</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C30B350FBC7FEA4F6C7D4C8" ID-GBIF-Occurrence="1915497840" collectionCode="FMNH" country="Peru" location="Distribution" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C30B350FBC7FEA4F326D4EC" box="[1083,1139,368,392]" pageId="24" pageNumber="25">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C30B350FAEDFEA4F237D4E3" box="[1297,1378,368,391]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="25">FMNH</collectionCode>
HT. 
<location id="AC2760054C30B350FF39FE40F604D4C9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FF39FE40F604D4C9" box="[197,337,404,429]" country="Peru" name="Distribution" pageId="24" pageNumber="25">Distribution</location>
: 
<collectingCountry id="D1EF764E4C30B350FEA7FE41F6C7D4C8" box="[347,402,405,428]" name="Peru" pageId="24" pageNumber="25">Peru</collectingCountry>
</materialsCitation>
. Note: Peru (
<bibRefCitation id="CD694B2F4C30B350FDDFFE40F5ECD4C8" box="[547,697,404,429]" pageId="24" pageNumber="25" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BFE63F465D4B4" box="[151,816,439,464]" pageId="24" pageNumber="25" type="discussion">
<paragraph id="A94736DE4C30B350FF6BFE63F465D4B4" blockId="24.[151,1437,331,464]" box="[151,816,439,464]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BFE63F641D4B4" bold="true" box="[151,276,439,464]" pageId="24" pageNumber="25">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C30B350FEB0FE63F69AD4B4" box="[332,463,439,464]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C30B350FEB0FE63F69AD4B4" box="[332,463,439,464]" italics="true" pageId="24" pageNumber="25">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BFDD4F6EED16B" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="A94736DE4C30B350FF6BFDD4F531D77C" blockId="24.[151,1436,511,608]" box="[151,612,511,536]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BFDD4F7F0D773" box="[151,165,512,535]" italics="true" pageId="24" pageNumber="25">*</emphasis>
<taxonomicName id="6EF84D5D4C30B350FF59FE2BF531D77C" ID-CoL="5WGHN" authority="Bernhauer 1910" authorityName="Bernhauer" authorityYear="1910" box="[165,612,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="9B8CEACC4C30B350FF59FE2BF6FCD77C" bold="true" box="[165,425,511,536]" italics="true" pageId="24" pageNumber="25">Belonuchus pectinipes</emphasis>
<bibRefCitation id="CD694B2F4C30B350FE4CFE2BF531D77C" box="[432,612,511,536]" pageId="24" pageNumber="25" refId="ref61912" refString="Bernhauer, M. (1910) Beitrag zur Kenntnis der Staphyliniden-Fauna von Zentralamerika. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 60, 350 - 393." type="journal article">Bernhauer 1910</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFDF0F3CAD704" blockId="24.[151,1436,511,608]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFDF0F3CAD704" pageId="24" pageNumber="25">
<materialsCitation id="19903C834C30B350FF6BFDF0F37DD758" ID-GBIF-Occurrence="1915497760" box="[151,1064,548,573]" location="Cordoba, Veracruz" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Cordoba, Veracruz" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C30B350FF6BFDF0F5C2D758" ID-CoL="5WGHN" authority="Bernhauer 1910: 379" authorityName="Bernhauer" authorityPageNumber="379" authorityYear="1910" box="[151,663,548,573]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pectinipes">
<emphasis id="9B8CEACC4C30B350FF6BFDF0F6C3D758" box="[151,406,548,573]" italics="true" pageId="24" pageNumber="25">Belonuchus pectinipes</emphasis>
<treatmentCitation id="285910CF4C30B350FE5CFDF0F5C2D758" author="" box="[416,663,548,573]" page="379" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE5CFDF0F509D758" box="[416,604,548,573]" pageId="24" pageNumber="25" refId="ref61912" refString="Bernhauer, M. (1910) Beitrag zur Kenntnis der Staphyliniden-Fauna von Zentralamerika. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 60, 350 - 393." type="journal article">Bernhauer 1910</bibRefCitation>
: 379
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FD5AFDF1F588D759" box="[678,733,549,573]" pageId="24" pageNumber="25">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C30B350FCB3FDF0F37DD758" box="[847,1064,548,573]" isEnumeration="true" pageId="24" pageNumber="25">Cordoba, Veracruz</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C30B350FBCAFDF1F6E1D704" ID-GBIF-Occurrence="1915497762" collectionCode="FMNH" country="Mexico" location="ST. Distribution" pageId="24" pageNumber="25" specimenCode="FMNH 4" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C30B350FBCAFDF1F33BD759" box="[1078,1134,549,573]" pageId="24" pageNumber="25">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C30B350FAFEFDF1F23ED758" box="[1282,1387,548,572]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="25">FMNH 4</specimenCode>
<location id="AC2760054C30B350FA8AFDF0F604D704" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FA8AFDF0F604D704" country="Mexico" name="ST. Distribution" pageId="24" pageNumber="25">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C30B350FEA7FD93F6E1D704" box="[347,436,583,608]" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingCountry>
</materialsCitation>
. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C30B350FCC2FD93F3C6D704" box="[830,1171,583,608]" pageId="24" pageNumber="25" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C30B350FBEBFD9DF305D704" box="[1047,1104,583,608]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFD5BF403D7CC" blockId="24.[151,1437,655,752]" box="[151,854,655,680]" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFD5BF5CBD7CC" authority="Plavilstschikov 1929" authorityName="Plavilstschikov" authorityYear="1929" box="[151,670,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="9B8CEACC4C30B350FF6BFD5BF6FCD7CC" bold="true" box="[151,425,655,680]" italics="true" pageId="24" pageNumber="25">Belonuchus peruvianus</emphasis>
<bibRefCitation id="CD694B2F4C30B350FE4CFD5BF5CBD7CC" box="[432,670,655,680]" pageId="24" pageNumber="25" refId="ref71306" refString="Plavilstschikov, N. N. (1929) Les Staphylinides recoltes par Mr. Nicolas Iconnicov dans le Perou (Coleoptera, Staphylinidae). Entomologisches Nachrichtsblatt, Troppau, 3 (1), 29 - 30." type="journal article">Plavilstschikov 1929</bibRefCitation>
</taxonomicName>
, 
<emphasis id="9B8CEACC4C30B350FD54FD5BF403D7CC" bold="true" box="[680,854,655,680]" pageId="24" pageNumber="25">nomen nudum</emphasis>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFD60F204D794" blockId="24.[151,1437,655,752]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFD60F204D794" infered="true" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFD60F5E9D7A8" authority="Plavilstschikov 1929: 30" authorityName="Plavilstschikov" authorityPageNumber="30" authorityYear="1929" box="[151,700,692,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="9B8CEACC4C30B350FF6BFD60F6CBD7A8" box="[151,414,692,717]" italics="true" pageId="24" pageNumber="25">Belonuchus peruvianus</emphasis>
<treatmentCitation id="285910CF4C30B350FE58FD60F5E9D7A8" author="" box="[420,700,692,717]" page="30" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE58FD60F5C6D7A8" box="[420,659,692,717]" pageId="24" pageNumber="25" refId="ref71306" refString="Plavilstschikov, N. N. (1929) Les Staphylinides recoltes par Mr. Nicolas Iconnicov dans le Perou (Coleoptera, Staphylinidae). Entomologisches Nachrichtsblatt, Troppau, 3 (1), 29 - 30." type="journal article">Plavilstschikov 1929</bibRefCitation>
: 30
</treatmentCitation>
</taxonomicName>
, as &quot;n. sp. (Bernhauer 
<emphasis id="9B8CEACC4C30B350FC47FD61F374D7A9" box="[955,1057,692,717]" italics="true" pageId="24" pageNumber="25">in litteris</emphasis>
)&quot;; nomen nudum, no description. Distribution: 
<collectingCountry id="D1EF764E4C30B350FEA7FD0CF6C7D78B" box="[347,402,728,751]" name="Peru" pageId="24" pageNumber="25">Peru</collectingCountry>
. Note: as valid sp. in 
<bibRefCitation id="CD694B2F4C30B350FD7FFD03F437D794" box="[643,866,727,752]" pageId="24" pageNumber="25" refId="ref63204" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part. 1. Bulletin of the United States National Museum, 185, 1 - 188." type="journal article">Blackwelder (1944)</bibRefCitation>
and 
<bibRefCitation id="CD694B2F4C30B350FC64FD03F369D794" box="[920,1084,727,752]" pageId="24" pageNumber="25" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo (2004)</bibRefCitation>
; not in 
<bibRefCitation id="CD694B2F4C30B350FB6CFD0CF219D794" box="[1168,1356,727,752]" pageId="24" pageNumber="25" refId="ref67417" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article">Herman (2001d)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFCCBF505D65C" blockId="24.[151,1436,799,896]" box="[151,592,799,824]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BFCCBF7F6D65C" bold="true" box="[151,163,799,824]" italics="true" pageId="24" pageNumber="25">*</emphasis>
<taxonomicName id="6EF84D5D4C30B350FF5EFCCBF505D65C" ID-CoL="68GHM" authority="Bernhauer 1935" authorityName="Bernhauer" authorityYear="1935" box="[162,592,799,824]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="piceolus">
<emphasis id="9B8CEACC4C30B350FF5EFCCBF6C0D65C" bold="true" box="[162,405,799,824]" italics="true" pageId="24" pageNumber="25">Belonuchus piceolus</emphasis>
<bibRefCitation id="CD694B2F4C30B350FE60FCCBF505D65C" box="[412,592,799,824]" pageId="24" pageNumber="25" refId="ref62766" refString="Bernhauer, M. (1935) Neue Staphyliniden (Col.) aus Sudamerika. Stylops, a Journal of Taxonomic Entomology, 4, 90 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00565. x" type="journal article">Bernhauer 1935</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFC90F33ED6E4" blockId="24.[151,1436,799,896]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFC90F33ED6E4" pageId="24" pageNumber="25">
<materialsCitation id="19903C834C30B350FF6BFC90F4DAD639" ID-GBIF-Occurrence="1915497798" box="[151,911,836,861]" country="Brazil" location="Brasilien" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C30B350FF6BFC90F524D638" ID-CoL="68GHM" authority="Bernhauer 1935: 90" authorityName="Bernhauer" authorityPageNumber="90" authorityYear="1935" box="[151,625,836,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="piceolus">
<emphasis id="9B8CEACC4C30B350FF6BFC90F6D5D639" box="[151,384,836,861]" italics="true" pageId="24" pageNumber="25">Belonuchus piceolus</emphasis>
<treatmentCitation id="285910CF4C30B350FE76FC90F524D638" author="" box="[394,625,836,861]" page="90" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE76FC90F510D639" box="[394,581,836,861]" pageId="24" pageNumber="25" refId="ref62766" refString="Bernhauer, M. (1935) Neue Staphyliniden (Col.) aus Sudamerika. Stylops, a Journal of Taxonomic Entomology, 4, 90 - 96. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00565. x" type="journal article">Bernhauer 1935</bibRefCitation>
: 90
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FD83FC91F5E3D639" box="[639,694,837,861]" pageId="24" pageNumber="25">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C30B350FCDAFC90F4DAD639" box="[806,911,836,861]" name="Brazil" pageId="24" pageNumber="25">Brasilien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C30B350FC60FC91F6F6D6E4" ID-GBIF-Occurrence="1915497830" collectionCode="BMNH" country="Brazil" location="ST. Distribution" pageId="24" pageNumber="25" specimenCode="FMNH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C30B350FC60FC91F481D639" box="[924,980,837,861]" pageId="24" pageNumber="25">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C30B350FB99FC91F3ECD638" box="[1125,1209,837,860]" pageId="24" pageNumber="25">BMNH</collectionCode>
#?ST; 
<specimenCode id="F95E9EA54C30B350FAF2FC91F226D638" box="[1294,1395,836,860]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="24" pageNumber="25">FMNH 1</specimenCode>
<location id="AC2760054C30B350FA8EFC90F604D6E4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FA8EFC90F604D6E4" country="Brazil" name="ST. Distribution" pageId="24" pageNumber="25">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C30B350FEA7FCB3F6F6D6E4" box="[347,419,871,896]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
</materialsCitation>
. Note: distribution within Brazil unknown (Asenjo 
<emphasis id="9B8CEACC4C30B350FC18FCBDF349D6E4" box="[996,1052,871,896]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFC7BF578D6AC" blockId="24.[151,1436,943,1112]" box="[151,557,943,968]" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFC7BF578D6AC" ID-CoL="68GHL" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[151,557,943,968]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="9B8CEACC4C30B350FF6BFC7BF6F3D6AC" bold="true" box="[151,422,943,968]" italics="true" pageId="24" pageNumber="25">Belonuchus pictipennis</emphasis>
<bibRefCitation id="CD694B2F4C30B350FE51FC7BF578D6AC" box="[429,557,943,968]" pageId="24" pageNumber="25" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFC00F6EED16B" blockId="24.[151,1436,943,1112]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFC00F6EED16B" pageId="24" pageNumber="25">
<materialsCitation id="19903C834C30B350FF6BFC00F4ADD688" ID-GBIF-Occurrence="1915497833" box="[151,1016,980,1005]" country="Panama" location="Bugaba" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C30B350FF6BFC00F523D689" ID-CoL="68GHL" authority="Sharp 1885: 436" authorityName="Sharp" authorityPageNumber="436" authorityYear="1885" box="[151,630,980,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="9B8CEACC4C30B350FF6BFC00F6F4D688" box="[151,417,980,1005]" italics="true" pageId="24" pageNumber="25">Belonuchus pictipennis</emphasis>
<treatmentCitation id="285910CF4C30B350FE52FC00F523D689" author="" box="[430,630,980,1005]" page="436" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE52FC00F56DD689" box="[430,568,980,1005]" pageId="24" pageNumber="25" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 436
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FD7AFC01F5EBD689" box="[646,702,981,1005]" pageId="24" pageNumber="25">Type</typeStatus>
locality: 
<location id="AC2760054C30B350FCCFFC00F4DED689" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FCCFFC00F4DED689" box="[819,907,980,1005]" country="Panama" name="Bugaba" pageId="24" pageNumber="25">Bugaba</location>
, 
<collectingCountry id="D1EF764E4C30B350FC60FC01F4ADD688" box="[924,1016,981,1004]" name="Panama" pageId="24" pageNumber="25">Panama</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C30B350FBF4FC01F6E2D16B" ID-GBIF-Occurrence="1915497804" collectionCode="BMNH" country="Panama" location="Distribution" pageId="24" pageNumber="25" specimenCode="BMNH 2" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C30B350FBF4FC01F36AD689" box="[1032,1087,981,1005]" pageId="24" pageNumber="25">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C30B350FB2BFC01F213D688" box="[1239,1350,980,1004]" collectionCode="BMNH" pageId="24" pageNumber="25">BMNH 2</specimenCode>
ST 
<specimenCount id="BFFEFD574C30B350FA7CFC01F2CDD688" box="[1408,1432,981,1004]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
. 
<location id="AC2760054C30B350FF39FC23F604D174" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FF39FC23F604D174" box="[197,337,1015,1040]" country="Panama" name="Distribution" pageId="24" pageNumber="25">Distribution</location>
: 
<collectingCountry id="D1EF764E4C30B350FEA7FC2CF6E2D16B" box="[347,439,1016,1039]" name="Panama" pageId="24" pageNumber="25">Panama</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BFBC8F512D13C" pageId="24" pageNumber="25" type="discussion">
<paragraph id="A94736DE4C30B350FF6BFBC8F512D13C" blockId="24.[151,1436,943,1112]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BFBC8F641D151" bold="true" box="[151,276,1052,1077]" pageId="24" pageNumber="25">Comment.</emphasis>
It may belong to 
<taxonomicName id="6EF84D5D4C30B350FE11FBC8F5B5D151" authority="Sharp" authorityName="Sharp" authorityYear="1885" box="[493,736,1052,1077]" class="Insecta" family="Staphylinidae" genus="Paederomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C30B350FE11FBC8F5C1D151" box="[493,660,1052,1077]" italics="true" pageId="24" pageNumber="25">Paederomimus</emphasis>
Sharp
</taxonomicName>
(Sharp: 436, “very closely allied to 
<emphasis id="9B8CEACC4C30B350FB6DFBC9F263D150" box="[1169,1334,1053,1076]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FB6DFBC9F278D150" authorityName="Sharp" authorityYear="1885" box="[1169,1325,1053,1076]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mimeticus">B. mimeticus</taxonomicName>
”
</emphasis>
) but see 
<taxonomicName id="6EF84D5D4C30B350FF39FBEBF517D13C" authority="(Sharp)" authorityName="Sharp" box="[197,578,1087,1112]" class="Insecta" family="Staphylinidae" genus="Paederomimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mimeticus">
<emphasis id="9B8CEACC4C30B350FF39FBEBF6B0D13C" box="[197,485,1087,1112]" italics="true" pageId="24" pageNumber="25">Paederomimus mimeticus</emphasis>
(Sharp)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BFB53F58ED18C" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="A94736DE4C30B350FF6BFB53F56AD1C4" blockId="24.[151,1436,1159,1293]" box="[151,575,1159,1184]" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFB53F56AD1C4" ID-CoL="5WGHK" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[151,575,1159,1184]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="planiceps">
<emphasis id="9B8CEACC4C30B350FF6BFB53F6C0D1C4" bold="true" box="[151,405,1159,1184]" italics="true" pageId="24" pageNumber="25">Belonuchus planiceps</emphasis>
<bibRefCitation id="CD694B2F4C30B350FE60FB53F56AD1C4" box="[412,575,1159,1184]" pageId="24" pageNumber="25" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFB78F58ED18C" blockId="24.[151,1436,1159,1293]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFB78F58ED18C" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BFB78F52CD1A1" ID-CoL="5WGHK" authority="Erichson 1840: 425" authorityName="Erichson" authorityPageNumber="425" authorityYear="1840" box="[151,633,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="planiceps">
<emphasis id="9B8CEACC4C30B350FF6BFB78F6DBD1A1" box="[151,398,1196,1221]" italics="true" pageId="24" pageNumber="25">Belonuchus planiceps</emphasis>
<treatmentCitation id="285910CF4C30B350FE6BFB78F52CD1A1" author="" box="[407,633,1196,1221]" page="425" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE6BFB78F56AD1A0" box="[407,575,1196,1221]" pageId="24" pageNumber="25" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 425
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FD7AFB79F5EBD1A1" box="[646,702,1197,1221]" pageId="24" pageNumber="25">Type</typeStatus>
locality: Columbia. 
<typeStatus id="7643887C4C30B350FC56FB79F4B7D1A1" box="[938,994,1197,1221]" pageId="24" pageNumber="25">Type</typeStatus>
depository: ZMHB 4 ST [MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C30B350FEA7FB04F68DD18C" box="[347,472,1231,1256]" name="Costa Rica" pageId="24" pageNumber="25">
<collectingRegion id="6B3CF83C4C30B350FEA7FB04F6CED18C" box="[347,411,1232,1256]" country="Tanzania" name="Pwani" pageId="24" pageNumber="25">Costa</collectingRegion>
Rica
</collectingCountry>
, 
<collectingCountry id="D1EF764E4C30B350FE1FFB1BF501D18C" box="[483,596,1231,1256]" name="Colombia" pageId="24" pageNumber="25">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C30B350FDA3FB1BF583D18C" box="[607,726,1231,1256]" name="Venezuela" pageId="24" pageNumber="25">Venezuela</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BFB20F4D5D069" box="[151,896,1268,1293]" pageId="24" pageNumber="25" type="discussion">
<paragraph id="A94736DE4C30B350FF6BFB20F4D5D069" blockId="24.[151,1436,1159,1293]" box="[151,896,1268,1293]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BFB20F641D069" bold="true" box="[151,276,1268,1293]" pageId="24" pageNumber="25">Comment.</emphasis>
It probably belongs to 
<taxonomicName id="6EF84D5D4C30B350FDE4FB20F429D069" authority="Erichson" authorityName="Erichson" box="[536,892,1268,1293]" class="Insecta" family="Staphylinidae" genus="Holisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C30B350FDE4FB20F53BD069" box="[536,622,1268,1293]" italics="true" pageId="24" pageNumber="25">Holisus</emphasis>
Erichson (Hyptiomina)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BFAEFF52CD0F9" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="A94736DE4C30B350FF6BFAEFF429D031" blockId="24.[151,1437,1339,1760]" box="[151,892,1339,1365]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BFAEFF7F6D030" bold="true" box="[151,163,1339,1364]" italics="true" pageId="24" pageNumber="25">*</emphasis>
<taxonomicName id="6EF84D5D4C30B350FF5EFAEFF507D031" ID-CoL="68GHK" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,594,1339,1365]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platypterus" status="comb. nov.">
<emphasis id="9B8CEACC4C30B350FF5EFAEFF6E0D030" bold="true" box="[162,437,1339,1364]" italics="true" pageId="24" pageNumber="25">Belonuchus platypterus</emphasis>
(
<bibRefCitation id="CD694B2F4C30B350FE38FAE8F51CD031" box="[452,585,1340,1365]" pageId="24" pageNumber="25" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C30B350FDA2FAE8F58BD031" box="[606,734,1340,1365]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FDA2FAE8F58BD031" bold="true" box="[606,734,1340,1365]" pageId="24" pageNumber="25">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C30B350FD11FAE8F421D031" box="[749,884,1340,1365]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="24" pageNumber="25">Figs. 15–16</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C30B350FF6BFA8BF52CD0F9" blockId="24.[151,1437,1339,1760]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BFA8BF52CD0F9" pageId="24" pageNumber="25">
<materialsCitation id="19903C834C30B350FF6BFA8BF302D01C" ID-GBIF-Occurrence="1915497789" box="[151,1111,1375,1400]" country="Guatemala" location="Cerro Zunil" pageId="24" pageNumber="25" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C30B350FF6BFA8BF508D01C" ID-CoL="4GBK2" authority="Sharp 1885: 412" authorityName="Sharp" authorityPageNumber="412" authorityYear="1885" box="[151,605,1375,1400]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platypterus">
<emphasis id="9B8CEACC4C30B350FF6BFA8BF6C3D01C" box="[151,406,1375,1400]" italics="true" pageId="24" pageNumber="25">Philonthus platypterus</emphasis>
<treatmentCitation id="285910CF4C30B350FE62FA8BF508D01C" author="" box="[414,605,1375,1400]" page="412" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE62FA8BF576D01C" box="[414,547,1375,1400]" pageId="24" pageNumber="25" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 412
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FD94FAB4F5CAD01C" box="[616,671,1376,1400]" pageId="24" pageNumber="25">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C30B350FCF7FA8BF4D2D01C" box="[779,903,1375,1400]" name="Guatemala" pageId="24" pageNumber="25">Guatemala</collectingCountry>
, 
<location id="AC2760054C30B350FC6FFAB4F34DD01C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FC6FFAB4F34DD01C" box="[915,1048,1375,1400]" country="Guatemala" name="Cerro Zunil" pageId="24" pageNumber="25">Cerro Zunil</location>
4 000
</materialsCitation>
– 
<materialsCitation id="19903C834C30B350FB9AFA8BF521D0F9" ID-GBIF-Occurrence="1915497791" collectionCode="BMNH" country="Guatemala" elevation="1524" location="ST. Distribution" pageId="24" pageNumber="25" specimenCode="BMNH 3" specimenCount="1" typeStatus="holotype">
<elevation id="22D5D1ED4C30B350FB9AFA8BF386D01C" box="[1126,1235,1375,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="24" pageNumber="25" unit="ft" value="5000.0">
<quantity id="6E009B3B4C30B350FB9AFA8BF386D01C" box="[1126,1235,1375,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.524" pageId="24" pageNumber="25" unit="ft" value="5000.0">5000 feet</quantity>
</elevation>
. 
<typeStatus id="7643887C4C30B350FB22FAB4F243D01C" box="[1246,1302,1376,1400]" pageId="24" pageNumber="25">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C30B350FF39FA51F67BD0F8" box="[197,302,1412,1436]" collectionCode="BMNH" pageId="24" pageNumber="25">BMNH 3</specimenCode>
<location id="AC2760054C30B350FEC9FA50F6B8D0F9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C30B350FEC9FA50F6B8D0F9" box="[309,493,1412,1437]" country="Guatemala" name="ST. Distribution" pageId="24" pageNumber="25">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C30B350FE0BFA50F521D0F9" box="[503,628,1412,1437]" name="Guatemala" pageId="24" pageNumber="25">Guatemala</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BFA73F455D36C" pageId="24" pageNumber="25" type="discussion">
<paragraph id="A94736DE4C30B350FF6BFA73F455D36C" blockId="24.[151,1437,1339,1760]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BFA73F65ED0A4" bold="true" box="[151,267,1447,1472]" pageId="24" pageNumber="25">Remarks.</emphasis>
Sharp explicitly mentioned three specimens (two males and one female) as part of the 
<typeStatus id="7643887C4C30B350FB15FA7CF24CD0A4" box="[1257,1305,1448,1472]" pageId="24" pageNumber="25">type</typeStatus>
series. The specimens listed below agree with the original description by 
<bibRefCitation id="CD694B2F4C30B350FC5BFA18F36AD081" box="[935,1087,1484,1509]" pageId="24" pageNumber="25" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp (1885)</bibRefCitation>
and considered to have been 
<typeStatus id="7643887C4C30B350FF39FA24F672D36C" box="[197,295,1520,1544]" pageId="24" pageNumber="25" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C30B350FDF5FA3BF525D36C" box="[521,624,1519,1544]" pageId="24" pageNumber="25" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BF9C0F48AD3D9" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="A94736DE4C30B350FF6BF9C0F48AD3D9" blockId="24.[151,1437,1339,1760]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BF9C0F698D349" bold="true" box="[151,461,1556,1581]" pageId="24" pageNumber="25">
<typeStatus id="7643887C4C30B350FF6BF9C0F787D349" box="[151,210,1556,1581]" pageId="24" pageNumber="25">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C30B350FE26F9C1F51FD349" box="[474,586,1557,1581]" pageId="24" pageNumber="25" type="lectotype">Lectotype</typeStatus>
<specimenCount id="BFFEFD574C30B350FDA4F9C1F53AD348" box="[600,623,1557,1580]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
, here designated (
<collectionCode id="CFE9AE1B4C30B350FCB2F9C0F4F3D348" box="[846,934,1556,1580]" pageId="24" pageNumber="25">BMNH</collectionCode>
, 
<figureCitation id="31C32A5B4C30B350FC44F9C0F31DD349" box="[952,1096,1556,1581]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="24" pageNumber="25">Figs. 15–16</figureCitation>
): card-mounted, “ 
<specimenCount id="BFFEFD574C30B350FAE2F9C0F261D348" box="[1310,1332,1556,1580]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
(in ink) 
<taxonomicName id="6EF84D5D4C30B350FF39F9E3F694D334" box="[197,449,1591,1616]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platypterus">Philonthus platypterus</taxonomicName>
<typeStatus id="7643887C4C30B350FE3BF9ECF6AAD334" box="[455,511,1592,1616]" pageId="24" pageNumber="25">Type</typeStatus>
D.S. Cerro Zunil. 
<quantity id="6E009B3B4C30B350FD2DF9E3F414D334" box="[721,833,1591,1616]" metricMagnitude="2" metricUnit="m" metricValue="7.6260959999999995" metricValueMax="15.24" metricValueMin="0.012192" pageId="24" pageNumber="25" unit="ft" value="2502.0" valueMax="5000.0" valueMin="4.0">4–5000 ft</quantity>
. [in Sharp’s hand] // 
<typeStatus id="7643887C4C30B350FBD7F9ECF336D334" box="[1067,1123,1592,1616]" pageId="24" pageNumber="25">Type</typeStatus>
(round white label with red margin) // Cerro Zunil. 
<collectingRegion id="6B3CF83C4C30B350FE37F988F51DD311" box="[459,584,1628,1653]" country="Guatemala" name="Guatemala" pageId="24" pageNumber="25">
<collectingCountry id="D1EF764E4C30B350FE37F988F51DD311" box="[459,584,1628,1653]" name="Guatemala" pageId="24" pageNumber="25">Guatemala</collectingCountry>
</collectingRegion>
. Champion. // B. C. A. Col. 
<date id="DD46101E4C30B350FC78F989F4FAD310" box="[900,943,1628,1652]" pageId="24" pageNumber="25">I. 2.</date>
<taxonomicName id="6EF84D5D4C30B350FC4AF988F25DD311" authority=", Sharp." authorityName="Sharp." box="[950,1288,1628,1653]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="platypterus">Philonthus platypterus, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”. 
<typeStatus id="7643887C4C30B350FD6EF9ABF460D3FC" box="[658,821,1663,1688]" pageId="24" pageNumber="25" type="paralectotype">Paralectotypes</typeStatus>
(
<collectionCode id="CFE9AE1B4C30B350FCB9F954F4F5D3FC" box="[837,928,1664,1688]" pageId="24" pageNumber="25">BMNH</collectionCode>
): 
<specimenCount id="BFFEFD574C30B350FC48F954F4B4D3F3" box="[948,993,1664,1688]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
, 
<specimenCount id="BFFEFD574C30B350FC12F954F34CD3F3" box="[1006,1049,1664,1688]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
, same locality and collection data as 
<typeStatus id="7643887C4C30B350FF1FF970F61CD3D9" box="[227,329,1700,1725]" pageId="24" pageNumber="25" type="lectotype">lectotype</typeStatus>
but male with additional printed white label “Sp. figured”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B350FF6BF913F33DD384" box="[151,1128,1735,1760]" pageId="24" pageNumber="25" type="discussion">
<paragraph id="A94736DE4C30B350FF6BF913F33DD384" blockId="24.[151,1437,1339,1760]" box="[151,1128,1735,1760]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BF913F64AD384" bold="true" box="[151,287,1735,1760]" pageId="24" pageNumber="25">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C30B350FD46F913F331D384" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1735,1760]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C30B350FD46F913F468D384" box="[698,829,1735,1760]" italics="true" pageId="24" pageNumber="25">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C30B350FCB7F913F309D384" box="[843,1116,1735,1760]" pageId="24" pageNumber="25" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C30B350FC1CF91DF34DD384" box="[992,1048,1735,1760]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C30B351FF6BF8DBF59ED6AC" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="A94736DE4C30B350FF6BF8DBF545D24C" blockId="24.[151,1436,1807,1977]" box="[151,528,1807,1832]" pageId="24" pageNumber="25">
<emphasis id="9B8CEACC4C30B350FF6BF8DBF7F6D24C" bold="true" box="[151,163,1807,1832]" italics="true" pageId="24" pageNumber="25">*</emphasis>
<taxonomicName id="6EF84D5D4C30B350FF5EF8DBF545D24C" ID-CoL="5WGHR" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,528,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pollens">
<emphasis id="9B8CEACC4C30B350FF5EF8DBF6DDD24C" bold="true" box="[162,392,1807,1832]" italics="true" pageId="24" pageNumber="25">Belonuchus pollens</emphasis>
<bibRefCitation id="CD694B2F4C30B350FE6CF8DBF545D24C" box="[400,528,1807,1832]" pageId="24" pageNumber="25" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C30B350FF6BF8E0F30FD2DC" blockId="24.[151,1436,1807,1977]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="89E811F04C30B350FF6BF8E0F30FD2DC" infered="true" pageId="24" pageNumber="25">
<taxonomicName id="6EF84D5D4C30B350FF6BF8E0F56CD228" ID-CoL="5WGHR" authority="Sharp 1885: 431" authorityName="Sharp" authorityPageNumber="431" authorityYear="1885" box="[151,569,1844,1869]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pollens">
<emphasis id="9B8CEACC4C30B350FF6BF8E0F626D229" box="[151,371,1844,1869]" italics="true" pageId="24" pageNumber="25">Belonuchus pollens</emphasis>
<treatmentCitation id="285910CF4C30B350FE87F8E0F56CD228" author="" box="[379,569,1844,1869]" page="431" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FE87F8E0F555D229" box="[379,512,1844,1869]" pageId="24" pageNumber="25" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 431
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C30B350FDB9F8E1F528D229" box="[581,637,1845,1869]" pageId="24" pageNumber="25">Type</typeStatus>
localities: North America; 
<collectingRegion id="6B3CF83C4C30B350FC48F8E0F35BD229" box="[948,1038,1844,1869]" country="Mexico" name="Mexico" pageId="24" pageNumber="25">
<collectingCountry id="D1EF764E4C30B350FC48F8E0F35BD229" box="[948,1038,1844,1869]" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingCountry>
</collectingRegion>
: 
<collectingRegion id="6B3CF83C4C30B350FBE6F8E0F327D228" box="[1050,1138,1844,1868]" country="Mexico" name="Oaxaca" pageId="24" pageNumber="25">Oaxaca</collectingRegion>
; 
<collectingRegion id="6B3CF83C4C30B350FB82F8E0F3A9D229" box="[1150,1276,1844,1869]" country="Guatemala" name="Guatemala" pageId="24" pageNumber="25">
<collectingCountry id="D1EF764E4C30B350FB82F8E0F3A9D229" box="[1150,1276,1844,1869]" name="Guatemala" pageId="24" pageNumber="25">Guatemala</collectingCountry>
</collectingRegion>
: near the city; 
<collectingCountry id="D1EF764E4C30B350FF39F883F663D214" box="[197,310,1879,1904]" name="Honduras" pageId="24" pageNumber="25">Honduras</collectingCountry>
. 
<typeStatus id="7643887C4C30B350FEC3F88CF622D214" box="[319,375,1880,1904]" pageId="24" pageNumber="25">Type</typeStatus>
depository: BMNH 8 ST: from all 
<typeStatus id="7643887C4C30B350FCFCF88CF465D214" box="[768,816,1880,1904]" pageId="24" pageNumber="25">type</typeStatus>
localities except North America [MCP seen]; FMNH 2 ST. Distribution: 
<collectingRegion id="6B3CF83C4C30B350FE73F8A8F6B2D2F1" box="[399,487,1916,1941]" country="Mexico" name="Mexico" pageId="24" pageNumber="25">
<collectingCountry id="D1EF764E4C30B350FE73F8A8F6B2D2F1" box="[399,487,1916,1941]" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C30B350FE0AF8A8F527D2F1" box="[502,626,1916,1941]" country="Guatemala" name="Guatemala" pageId="24" pageNumber="25">
<collectingCountry id="D1EF764E4C30B350FE0AF8A8F527D2F1" box="[502,626,1916,1941]" name="Guatemala" pageId="24" pageNumber="25">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C30B350FD7DF8A8F5BBD2F1" box="[641,750,1916,1941]" name="Honduras" pageId="24" pageNumber="25">Honduras</collectingCountry>
[&quot;North America&quot; in error]. Notes: not in 
<collectingCountry id="D1EF764E4C30B350FB18F8A8F249D2F0" box="[1252,1308,1916,1940]" name="United States of America" pageId="24" pageNumber="25">USA</collectingCountry>
or 
<collectingCountry id="D1EF764E4C30B350FAB4F8A8F2C9D2F1" box="[1352,1436,1916,1941]" name="Canada" pageId="24" pageNumber="25">Canada</collectingCountry>
(
<treatmentCitation id="285910CF4C30B350FF30F874F6FFD2DC" author="" box="[204,426,1952,1977]" page="733" pageId="24" pageNumber="25">
<bibRefCitation id="CD694B2F4C30B350FF30F874F6FFD2DC" box="[204,426,1952,1977]" pageId="24" pageNumber="25" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 733</bibRefCitation>
</treatmentCitation>
); distribution within 
<collectingRegion id="6B3CF83C4C30B350FD69F874F5BFD2DD" box="[661,746,1952,1977]" country="Mexico" name="Mexico" pageId="24" pageNumber="25">
<collectingCountry id="D1EF764E4C30B350FD69F874F5BFD2DD" box="[661,746,1952,1977]" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C30B350FD05F874F31BD2DC" box="[761,1102,1952,1977]" pageId="24" pageNumber="25" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C30B350FC2EF875F35ED2DD" box="[978,1035,1952,1977]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFF43F592D5D4" blockId="25.[151,1436,151,285]" box="[151,711,151,176]" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFF43F592D5D4" ID-CoL="68GHR" authority="Laporte de Castelnau 1840" authorityName="Laporte de Castelnau" authorityYear="1840" box="[151,711,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="posticalis">
<emphasis id="9B8CEACC4C31B351FF6BFF43F6C6D5D4" bold="true" box="[151,403,151,176]" italics="true" pageId="25" pageNumber="26">Belonuchus posticalis</emphasis>
Laporte de 
<bibRefCitation id="CD694B2F4C31B351FDE6FF43F592D5D4" box="[538,711,151,176]" pageId="25" pageNumber="26" refId="ref68445" refString="Laporte de Castelnau, F. L. (1840) Histoire naturelle des Insectes Coleopteres, vol. 1. In: Histoire naturelle des animaux articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. Vol. 2. P. Dumenil, Paris, 324 pp., 19 pls." type="book">Castelnau 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFF68F532D478" blockId="25.[151,1436,151,285]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BFF68F532D478" infered="true" pageId="25" pageNumber="26">
<materialsCitation id="19903C834C31B351FF6BFF68F369D5B1" ID-GBIF-Occurrence="1915497776" box="[151,1084,188,213]" country="Brazil" location="Bresil" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C31B351FF6BFF68F47AD5B1" ID-CoL="6ZG8B" authority="Laporte de Castelnau 1840: 176" authorityName="Laporte de Castelnau" authorityPageNumber="176" authorityYear="1840" box="[151,815,188,213]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="posticalis">
<emphasis id="9B8CEACC4C31B351FF6BFF68F6C6D5B1" box="[151,403,188,213]" italics="true" pageId="25" pageNumber="26">Staphylinus posticalis</emphasis>
Laporte de 
<treatmentCitation id="285910CF4C31B351FDC8FF68F47AD5B1" author="" box="[564,815,188,213]" page="176" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FDC8FF68F5BBD5B0" box="[564,750,188,213]" pageId="25" pageNumber="26" refId="ref68445" refString="Laporte de Castelnau, F. L. (1840) Histoire naturelle des Insectes Coleopteres, vol. 1. In: Histoire naturelle des animaux articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. Vol. 2. P. Dumenil, Paris, 324 pp., 19 pls." type="book">Castelnau 1840</bibRefCitation>
: 176
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C31B351FCBEFF69F42FD5B1" box="[834,890,189,213]" pageId="25" pageNumber="26">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C31B351FC09FF68F369D5B1" box="[1013,1084,188,213]" name="Brazil" pageId="25" pageNumber="26">Brésil</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C31B351FBB3FF69F6F0D59C" ID-GBIF-Occurrence="1915497774" country="Brazil" location="ST. Distribution" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C31B351FBB3FF69F3D3D5B1" box="[1103,1158,189,213]" pageId="25" pageNumber="26">Type</typeStatus>
depository: ?? #? 
<location id="AC2760054C31B351FA8BFF68F604D59C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C31B351FA8BFF68F604D59C" country="Brazil" name="ST. Distribution" pageId="25" pageNumber="26">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C31B351FEA1FF0BF6F0D59C" box="[349,421,223,248]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
</materialsCitation>
. Note: comb. nov. ex 
<taxonomicName id="6EF84D5D4C31B351FD5DFF0BF3E8D59C" ID-CoL="8VS43" authority="(Bernhauer &amp; Schubert 1914: 371)" authorityName="Bernhauer &amp; Schubert" authorityPageNumber="371" authorityYear="1914" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[673,1213,223,248]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C31B351FD5DFF0BF476D59C" box="[673,803,223,248]" italics="true" pageId="25" pageNumber="26">Staphylinus</emphasis>
(
<treatmentCitation id="285910CF4C31B351FCCFFF0BF3E1D59C" author="" box="[819,1204,223,248]" page="371" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FCCFFF0BF3E1D59C" box="[819,1204,223,248]" pageId="25" pageNumber="26" refId="ref62964" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus. Vol. 57. W. Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer &amp; Schubert 1914: 371</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; distribution within Brazil unknown (Asenjo 
<emphasis id="9B8CEACC4C31B351FE23FED1F542D479" box="[479,535,260,285]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFE9FF5D5D400" blockId="25.[151,1437,331,429]" box="[151,640,331,357]" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFE9FF5D5D400" ID-CoL="5WGHQ" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,640,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="prasinipennis">
<emphasis id="9B8CEACC4C31B351FF6BFE9FF690D400" bold="true" box="[151,453,331,356]" italics="true" pageId="25" pageNumber="26">Belonuchus prasinipennis</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFEBBF4FAD4C9" blockId="25.[151,1437,331,429]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BFEBBF4FAD4C9" pageId="25" pageNumber="26">
<materialsCitation id="19903C834C31B351FF6BFEBBF2CDD4EC" ID-GBIF-Occurrence="1915497788" box="[151,1432,367,392]" country="Bolivia" location="Yuracares" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C31B351FF6BFEBBF5E0D4EC" ID-CoL="5WGHQ" authority="Bernhauer 1917 b: 18" authorityName="Bernhauer" authorityPageNumber="18" authorityYear="1917" box="[151,693,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="prasinipennis">
<emphasis id="9B8CEACC4C31B351FF6BFEBBF6E9D4EC" box="[151,444,367,392]" italics="true" pageId="25" pageNumber="26">Belonuchus prasinipennis</emphasis>
<treatmentCitation id="285910CF4C31B351FE38FEBBF5E0D4EC" author="" box="[452,693,367,392]" page="18" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE38FEBBF5D9D4EC" box="[452,652,367,392]" pageId="25" pageNumber="26" refId="ref62221" refString="Bernhauer, M. (1917 b) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (17. Beitrag - Fortsetzung). Entomologische Blatter, 13 (1 - 3), 17 - 24." type="journal article">Bernhauer 1917b</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
(attributed to Fauvel 
<emphasis id="9B8CEACC4C31B351FC4DFEA5F4A3D4EC" box="[945,1014,367,392]" italics="true" pageId="25" pageNumber="26">in litt.</emphasis>
). 
<typeStatus id="7643887C4C31B351FBECFEA4F312D4EC" box="[1040,1095,368,392]" pageId="25" pageNumber="26">Type</typeStatus>
locality: 
<location id="AC2760054C31B351FB49FEA4F273D4EC" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C31B351FB49FEA4F273D4EC" box="[1205,1318,368,392]" country="Bolivia" name="Yuracares" pageId="25" pageNumber="26">Yuracarès</location>
, 
<collectingCountry id="D1EF764E4C31B351FAC8FEBBF2CDD4EC" box="[1332,1432,367,392]" name="Bolivia" pageId="25" pageNumber="26">Bolivien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C31B351FF39FE41F4FFD4C9" ID-GBIF-Occurrence="1915497844" box="[197,938,404,429]" collectionCode="FMNH, ZMUH" country="Bolivia" location="ST. Distribution" pageId="25" pageNumber="26" specimenCode="FMNH 1, ZMUH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C31B351FF39FE41F7A8D4C9" box="[197,253,405,429]" pageId="25" pageNumber="26">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C31B351FE74FE41F6BBD4C8" box="[392,494,404,428]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26">FMNH 1</specimenCode>
ST; 
<specimenCode id="F95E9EA54C31B351FDDFFE41F5DED4C8" box="[547,651,404,428]" collectionCode="ZMUH" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="25" pageNumber="26">ZMUH 1</specimenCode>
<location id="AC2760054C31B351FD6EFE40F41FD4C9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C31B351FD6EFE40F41FD4C9" box="[658,842,404,429]" country="Bolivia" name="ST. Distribution" pageId="25" pageNumber="26">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C31B351FCA8FE40F4FFD4C9" box="[852,938,404,429]" name="Bolivia" pageId="25" pageNumber="26">Bolivia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFE0FF5D6D490" blockId="25.[151,1436,475,608]" box="[151,643,475,501]" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFE0FF5D6D490" ID-CoL="68GHQ" authority="Bernhauer 1908" authorityName="Bernhauer" authorityYear="1908" box="[151,643,475,501]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pulchripennis">
<emphasis id="9B8CEACC4C31B351FF6BFE0FF69DD490" bold="true" box="[151,456,475,500]" italics="true" pageId="25" pageNumber="26">Belonuchus pulchripennis</emphasis>
<bibRefCitation id="CD694B2F4C31B351FE33FE08F5D6D490" box="[463,643,476,501]" pageId="25" pageNumber="26" refId="ref61850" refString="Bernhauer, M. (1908) Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte, 74 (1), 283 - 372." type="journal article">Bernhauer 1908</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFE2BF64BD704" blockId="25.[151,1436,475,608]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BFE2BF64BD704" pageId="25" pageNumber="26">
<materialsCitation id="19903C834C31B351FF6BFE2BF203D773" ID-GBIF-Occurrence="1915497843" box="[151,1366,511,536]" country="Peru" location="Montana" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Montana" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C31B351FF6BFE2BF594D77C" ID-CoL="68GHQ" authority="Bernhauer 1908: 330" authorityName="Bernhauer" authorityPageNumber="330" authorityYear="1908" box="[151,705,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pulchripennis">
<emphasis id="9B8CEACC4C31B351FF6BFE2BF6EAD77C" box="[151,447,511,536]" italics="true" pageId="25" pageNumber="26">Belonuchus pulchripennis</emphasis>
<treatmentCitation id="285910CF4C31B351FE36FE2BF594D77C" author="" box="[458,705,511,536]" page="330" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE36FE2BF5D0D77C" box="[458,645,511,536]" pageId="25" pageNumber="26" refId="ref61850" refString="Bernhauer, M. (1908) Beitrag zur Staphylinidenfauna von Sudamerika. Archiv fur Naturgeschichte, 74 (1), 283 - 372." type="journal article">Bernhauer 1908</bibRefCitation>
: 330
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C31B351FD2CFDD4F45DD77C" box="[720,776,512,536]" pageId="25" pageNumber="26">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C31B351FC86FDD4F48BD773" box="[890,990,512,535]" country="United States of America" name="Montana" pageId="25" pageNumber="26">Montana</collectingRegion>
de Nancho, 7000', central 
<collectingCountry id="D1EF764E4C31B351FAE3FDD4F203D773" box="[1311,1366,512,535]" name="Peru" pageId="25" pageNumber="26">Peru</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C31B351FA98FDD4F464D758" ID-GBIF-Occurrence="1915497807" collectionCode="FMNH" country="Peru" location="ST. Distribution" pageId="25" pageNumber="26" specimenCode="FMNH 2" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C31B351FA98FDD4F2C9D77C" box="[1380,1436,512,536]" pageId="25" pageNumber="26">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C31B351FEB3FDF1F6EFD758" box="[335,442,548,572]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="25" pageNumber="26">FMNH 2</specimenCode>
ST; ?? 1 
<location id="AC2760054C31B351FDD5FDF0F5BED759" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C31B351FDD5FDF0F5BED759" box="[553,747,548,573]" country="Peru" name="ST. Distribution" pageId="25" pageNumber="26">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C31B351FD06FDF1F464D758" box="[762,817,549,572]" name="Peru" pageId="25" pageNumber="26">Peru</collectingCountry>
</materialsCitation>
. Note: Peru (
<bibRefCitation id="CD694B2F4C31B351FC2FFDF0F338D758" box="[979,1133,548,573]" pageId="25" pageNumber="26" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFD5BF5DDD7CC" blockId="25.[151,1436,655,789]" box="[151,648,655,680]" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFD5BF5DDD7CC" ID-CoL="5WGJ7" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,648,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="purpuripennis">
<emphasis id="9B8CEACC4C31B351FF6BFD5BF698D7CC" bold="true" box="[151,461,655,680]" italics="true" pageId="25" pageNumber="26">Belonuchus purpuripennis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFD60F64BD671" blockId="25.[151,1436,655,789]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BFD60F64BD671" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFD60F596D7A8" ID-CoL="5WGJ7" authority="Bernhauer 1916 c: 278" authorityName="Bernhauer" authorityPageNumber="278" authorityYear="1916" box="[151,707,692,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="purpuripennis">
<emphasis id="9B8CEACC4C31B351FF6BFD60F695D7A8" box="[151,448,692,717]" italics="true" pageId="25" pageNumber="26">Belonuchus purpuripennis</emphasis>
<treatmentCitation id="285910CF4C31B351FE3AFD60F596D7A8" author="" box="[454,707,692,717]" page="278" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE3AFD60F5DED7A9" box="[454,651,692,717]" pageId="25" pageNumber="26" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 278
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C31B351FD32FD61F453D7A9" box="[718,774,693,717]" pageId="25" pageNumber="26">Type</typeStatus>
locality: West-Columbien: Umgebung Cali am Rio 
<collectingRegion id="6B3CF83C4C31B351FAB3FD60F2CDD7A8" box="[1359,1432,692,716]" country="Colombia" name="Cauca" pageId="25" pageNumber="26">Cauca</collectingRegion>
, St. Antonio, 
<quantity id="6E009B3B4C31B351FEA9FD0CF6FBD78B" box="[341,430,728,752]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="m" value="2000.0">2000 m</quantity>
. 
<typeStatus id="7643887C4C31B351FE45FD0CF6A4D794" box="[441,497,728,752]" pageId="25" pageNumber="26">Type</typeStatus>
depository: FMNH 2 ST. Distribution: 
<collectingCountry id="D1EF764E4C31B351FC4CFD03F374D794" box="[944,1057,727,752]" name="Colombia" pageId="25" pageNumber="26">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C31B351FBD1FD03F3D6D794" box="[1069,1155,727,752]" name="Bolivia" pageId="25" pageNumber="26">Bolivia</collectingCountry>
. Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFC97F504D638" blockId="25.[151,1436,835,1005]" box="[151,593,835,861]" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFC97F504D638" ID-CoL="LCB2" authority="(Bernhauer, 1912)" authorityName="Bernhauer" authorityYear="1912" baseAuthorityName="Bernhauer" baseAuthorityYear="1912" box="[151,593,835,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="richteri">
<emphasis id="9B8CEACC4C31B351FF6BFC97F628D638" bold="true" box="[151,381,835,860]" italics="true" pageId="25" pageNumber="26">Belonuchus richteri</emphasis>
(Bernhauer, 1912)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFCB3F59ED6AC" blockId="25.[151,1436,835,1005]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BFCB3F59ED6AC" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFCB3F410D6E4" ID-CoL="4G9MH" authority="Bernhauer 1912 a: 44" authorityName="Bernhauer" authorityPageNumber="44" authorityYear="1912" box="[151,837,871,896]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="richteri" subGenus="Chroaptomus">
<emphasis id="9B8CEACC4C31B351FF6BFCB3F564D6E4" box="[151,561,871,896]" italics="true" pageId="25" pageNumber="26">Philonthus (Chroaptomus) richteri</emphasis>
<treatmentCitation id="285910CF4C31B351FDBFFCB3F410D6E4" author="" box="[579,837,871,896]" page="44" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FDBFFCB3F444D6E4" box="[579,785,871,896]" pageId="25" pageNumber="26" refId="ref61973" refString="Bernhauer, M. (1912 a) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article">Bernhauer 1912a</bibRefCitation>
: 44
</treatmentCitation>
</taxonomicName>
. Type locality: Tucuman prov., Argentina. Type depository: FMNH HT ♀. Distribution: Argentina. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C31B351FBE7FC58F7AED6AC" ID-CoL="8X778" authority="(Chani-Posse 2006: 365)" authorityName="Chani-Posse" authorityPageNumber="365" authorityYear="2006" baseAuthorityName="Chani-Posse" baseAuthorityYear="2006" class="Insecta" family="Staphylinidae" genus="Chroaptomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C31B351FBE7FC58F3E7D6C1" box="[1051,1202,908,933]" italics="true" pageId="25" pageNumber="26">Chroaptomus</emphasis>
(
<treatmentCitation id="285910CF4C31B351FB38FC58F7A6D6AC" author="" page="365" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FB38FC58F7A6D6AC" pageId="25" pageNumber="26" refId="ref64063" refString="Chani-Posse, M. (2006) Systematic revision and cladistic analysis of the Neotropical genus Chroaptomus Sharp (Coleoptera: Staphylinidae), with descriptions of two new species. Insect Systematics &amp; Evolution, 37, 361 - 383. https: // doi. org / 10.1163 / 187631206788831452" type="journal article">Chani-Posse 2006: 365</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; redescription (
<treatmentCitation id="285910CF4C31B351FE5BFC7BF5EBD6AC" author="" box="[423,702,943,968]" page="223" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE5BFC7BF5EBD6AC" box="[423,702,943,968]" pageId="25" pageNumber="26" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b: 223</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C31B351FF6BFC00F465D689" box="[151,816,980,1005]" pageId="25" pageNumber="26" type="discussion">
<paragraph id="A94736DE4C31B351FF6BFC00F465D689" blockId="25.[151,1436,835,1005]" box="[151,816,980,1005]" pageId="25" pageNumber="26">
<emphasis id="9B8CEACC4C31B351FF6BFC00F641D689" bold="true" box="[151,276,980,1005]" pageId="25" pageNumber="26">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C31B351FEB0FC00F69AD689" box="[332,463,980,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C31B351FEB0FC00F69AD689" box="[332,463,980,1005]" italics="true" pageId="25" pageNumber="26">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C31B351FF6BFBC0F65BD207" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="A94736DE4C31B351FF6BFBC0F53AD14A" blockId="25.[151,1437,1044,1718]" box="[151,623,1044,1070]" pageId="25" pageNumber="26">
<emphasis id="9B8CEACC4C31B351FF6BFBC0F7F6D149" bold="true" box="[151,163,1044,1069]" italics="true" pageId="25" pageNumber="26">*</emphasis>
<taxonomicName id="6EF84D5D4C31B351FF5EFBC0F53AD14A" ID-CoL="LCB5" authority="(Fabricius 1801)" authorityName="Fabricius" authorityYear="1801" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[162,623,1044,1070]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufipennis">
<emphasis id="9B8CEACC4C31B351FF5EFBC0F6FBD149" bold="true" box="[162,430,1044,1069]" italics="true" pageId="25" pageNumber="26">Belonuchus rufipennis</emphasis>
(
<bibRefCitation id="CD694B2F4C31B351FE41FBC1F533D14A" box="[445,614,1045,1070]" pageId="25" pageNumber="26" refId="ref65632" refString="Fabricius, J. C. (1801) Systema eleutheratorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Vol. 2. Bibliopolii Academici Novi, Kiliae (Kiel), 687 pp." type="book">Fabricius 1801</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFBEDF5EDD0F2" blockId="25.[151,1437,1044,1718]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BFBEDF5EDD0F2" infered="true" pageId="25" pageNumber="26">
<taxonomicName id="6EF84D5D4C31B351FF6BFBEDF5DCD136" ID-CoL="4ZKC5" authority="Fabricius 1801: 596" authorityName="Fabricius" authorityPageNumber="596" authorityYear="1801" box="[151,649,1081,1106]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufipennis">
<emphasis id="9B8CEACC4C31B351FF6BFBEDF6C0D136" box="[151,405,1081,1106]" italics="true" pageId="25" pageNumber="26">Staphylinus rufipennis</emphasis>
<treatmentCitation id="285910CF4C31B351FE63FBEDF5DCD136" author="" box="[415,649,1081,1106]" page="596" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE63FBEDF518D136" box="[415,589,1081,1106]" pageId="25" pageNumber="26" refId="ref65632" refString="Fabricius, J. C. (1801) Systema eleutheratorum secundum ordines, genera, species, adjectis synonymis, locis, observationibus, descriptionibus. Vol. 2. Bibliopolii Academici Novi, Kiliae (Kiel), 687 pp." type="book">Fabricius 1801</bibRefCitation>
: 596
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C31B351FD6BFBEEF59AD136" box="[663,719,1082,1106]" pageId="25" pageNumber="26">Type</typeStatus>
locality: America meridionali. 
<typeStatus id="7643887C4C31B351FBC0FBEEF321D136" box="[1084,1140,1082,1106]" pageId="25" pageNumber="26">Type</typeStatus>
depository: ZMUK 3 ST (until recently at ZMUC, 
<bibRefCitation id="CD694B2F4C31B351FE0DFB89F5DED112" box="[497,651,1117,1142]" pageId="25" pageNumber="26" refId="ref74168" refString="Zimsen, E. (1964) The type material of I. C. Fabricius. Munksgaard, Copenhagen, 656 pp." type="book">Zimsen 1964</bibRefCitation>
), not seen by 
<bibRefCitation id="CD694B2F4C31B351FCCDFB8AF4BFD112" box="[817,1002,1117,1142]" pageId="25" pageNumber="26" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana (1995)</bibRefCitation>
. Distribution: 
<collectingCountry id="D1EF764E4C31B351FB6DFB89F3BDD112" box="[1169,1256,1117,1142]" name="Canada" pageId="25" pageNumber="26">Canada</collectingCountry>
, 
<collectingCountry id="D1EF764E4C31B351FB0AFB8AF264D112" box="[1270,1329,1118,1142]" name="United States of America" pageId="25" pageNumber="26">USA</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C31B351FABCFB89F2CDD112" box="[1344,1432,1117,1142]" country="Mexico" name="Mexico" pageId="25" pageNumber="26">
<collectingCountry id="D1EF764E4C31B351FABCFB89F2CDD112" box="[1344,1432,1117,1142]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C31B351FF39FB55F614D1FE" box="[197,321,1153,1178]" country="Guatemala" name="Guatemala" pageId="25" pageNumber="26">
<collectingCountry id="D1EF764E4C31B351FF39FB55F614D1FE" box="[197,321,1153,1178]" name="Guatemala" pageId="25" pageNumber="26">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C31B351FEB1FB55F6C2D1FE" box="[333,407,1153,1178]" country="Belize" name="Belize" pageId="25" pageNumber="26">
<collectingCountry id="D1EF764E4C31B351FEB1FB55F6C2D1FE" box="[333,407,1153,1178]" name="Belize" pageId="25" pageNumber="26">Belize</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C31B351FE5FFB55F57CD1FE" box="[419,553,1153,1178]" name="El Salvador" pageId="25" pageNumber="26">El Salvador</collectingCountry>
, 
<collectingCountry id="D1EF764E4C31B351FDC9FB55F5FED1FE" box="[565,683,1153,1178]" name="Nicaragua" pageId="25" pageNumber="26">Nicaragua</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C31B351FD4BFB56F446D1FD" box="[695,787,1154,1177]" country="Panama" name="Panama" pageId="25" pageNumber="26">
<collectingCountry id="D1EF764E4C31B351FD4BFB56F446D1FD" box="[695,787,1154,1177]" name="Panama" pageId="25" pageNumber="26">Panama</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C31B351FCE3FB55F4C5D1FE" box="[799,912,1153,1178]" name="Colombia" pageId="25" pageNumber="26">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C31B351FC60FB55F347D1FE" box="[924,1042,1153,1178]" name="Venezuela" pageId="25" pageNumber="26">Venezuela</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C31B351FBE2FB55F39DD1FE" box="[1054,1224,1153,1178]" country="France" name="Guyane" pageId="25" pageNumber="26">
<collectingCountry id="D1EF764E4C31B351FBE2FB55F39DD1FE" box="[1054,1224,1153,1178]" name="French Guiana" pageId="25" pageNumber="26">French Guiana</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C31B351FB2FFB55F263D1FE" box="[1235,1334,1153,1178]" name="Trinidad and Tobago" pageId="25" pageNumber="26">Trinidad</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C31B351FABEFB55F2CDD1FE" box="[1346,1432,1153,1178]" country="Trinidad and Tobago" name="Tobago" pageId="25" pageNumber="26">
<collectingCountry id="D1EF764E4C31B351FABEFB55F2CDD1FE" box="[1346,1432,1153,1178]" name="Trinidad and Tobago" pageId="25" pageNumber="26">Tobago</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C31B351FF39FB71F677D1DA" box="[197,290,1189,1214]" name="Jamaica" pageId="25" pageNumber="26">Jamaica</collectingCountry>
, 
<collectingCountry id="D1EF764E4C31B351FED2FB72F631D1D9" box="[302,356,1190,1213]" name="Peru" pageId="25" pageNumber="26">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C31B351FE8CFB71F6E2D1DA" box="[368,439,1189,1214]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C31B351FE3FFB71F560D1DA" box="[451,565,1189,1214]" name="Argentina" pageId="25" pageNumber="26">Argentina</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C31B351FDBDFB71F5E2D1DA" box="[577,695,1189,1214]" country="Ecuador" name="Galapagos" pageId="25" pageNumber="26">Galapagos</collectingRegion>
Is. (
<collectingRegion id="6B3CF83C4C31B351FD16FB71F46BD1DA" box="[746,830,1189,1214]" country="Puerto Rico" name="Isabela" pageId="25" pageNumber="26">Isabela</collectingRegion>
, S. Crist., S. Cruz), 
<collectingRegion id="6B3CF83C4C31B351FBDDFB71F320D1DA" box="[1057,1141,1189,1214]" country="United States of America" name="Hawaii" pageId="25" pageNumber="26">Hawaii</collectingRegion>
; introduced for biocontrol in 
<collectingCountry id="D1EF764E4C31B351FF15FB1DF648D186" box="[233,285,1225,1250]" name="Italy" pageId="25" pageNumber="26">Italy</collectingCountry>
but not established. Notes: as synonym of 
<taxonomicName id="6EF84D5D4C31B351FCCBFB1EF384D186" ID-CoL="4ZJZ5" authority="(Erichson 1840: 426)" authorityName="Erichson" authorityPageNumber="426" authorityYear="1840" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[823,1233,1225,1250]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="9B8CEACC4C31B351FCCBFB1EF491D186" box="[823,964,1226,1250]" italics="true" pageId="25" pageNumber="26">B. formosus</emphasis>
(
<treatmentCitation id="285910CF4C31B351FC25FB1DF39FD186" author="" box="[985,1226,1225,1250]" page="426" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FC25FB1DF39FD186" box="[985,1226,1225,1250]" pageId="25" pageNumber="26" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840: 426</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; introduction for biocontrol, biology, immature stages (
<bibRefCitation id="CD694B2F4C31B351FD89FB39F44FD062" box="[629,794,1261,1286]" pageId="25" pageNumber="26" refId="ref72647" refString="Silvestri, F. (1945) Descrizione e biologia del coleottero stafilinide Belonuchus formosus Grav. introdotto in Italia per la lotta contro Ditteri Tripaneidi. Bollettino del R. Laboratorio di Entomologia Agraria di Portici, 5, 312 - 328." type="journal article">Silvestri 1945</bibRefCitation>
); redescription and distribution (
<treatmentCitation id="285910CF4C31B351FB72FB3AF224D062" author="" box="[1166,1393,1261,1286]" page="726" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FB72FB3AF224D062" box="[1166,1393,1261,1286]" pageId="25" pageNumber="26" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 726</bibRefCitation>
</treatmentCitation>
); in 
<taxonomicName id="6EF84D5D4C31B351FF39FAC6F63AD04D" ID-CoL="JMC4" baseAuthorityName="Kistner" baseAuthorityYear="1982" box="[197,367,1298,1321]" class="Insecta" family="Formicidae" genus="Atta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="mexicana">
<emphasis id="9B8CEACC4C31B351FF39FAC6F63AD04D" box="[197,367,1298,1321]" italics="true" pageId="25" pageNumber="26">Atta mexicana</emphasis>
</taxonomicName>
external debris (occasional) (
<bibRefCitation id="CD694B2F4C31B351FD21FAC5F4A9D04E" box="[733,1020,1297,1322]" pageId="25" pageNumber="26" refId="ref69955" refString="Navarrete-Heredia, J. L. (2001) Beetles associated with Atta and Acromyrmex ants (Hymenoptera: Formicidae: Attini). Transactions of the American Entomological Society, 127, 381 - 429." type="journal article">Navarrete-Heredia 2001</bibRefCitation>
); distribution, also within 
<collectingRegion id="6B3CF83C4C31B351FABAFAC5F2C9D04E" box="[1350,1436,1297,1322]" country="Mexico" name="Mexico" pageId="25" pageNumber="26">
<collectingCountry id="D1EF764E4C31B351FABAFAC5F2C9D04E" box="[1350,1436,1297,1322]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C31B351FF30FAE1F579D02A" box="[204,556,1333,1358]" pageId="25" pageNumber="26" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C31B351FE55FAE2F6B0D02A" box="[425,485,1333,1358]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2002
</bibRefCitation>
, 
<bibRefCitation id="CD694B2F4C31B351FDC7FAE2F5B3D02A" box="[571,742,1333,1358]" pageId="25" pageNumber="26" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
); distribution within 
<collectingRegion id="6B3CF83C4C31B351FC20FAE1F307D02A" box="[988,1106,1333,1358]" country="Ecuador" name="Galapagos" pageId="25" pageNumber="26">Galapagos</collectingRegion>
Is. (
<bibRefCitation id="CD694B2F4C31B351FB77FAE2F2C5D02A" box="[1163,1424,1333,1358]" pageId="25" pageNumber="26" refId="ref70441" refString="Newton, A. F. &amp; Peck, S. B. (2006) Family Staphylinidae, the rove beetles. In: Peck, S. B. (Ed.), The beetles of the Galapagos Islands, Ecuador: evolution, ecology and diversity (Insecta: Coleoptera). NRC Research Press, Ottawa, Ontario, pp. 131 - 158." type="book chapter">Newton &amp; Peck 2006</bibRefCitation>
); distribution within 
<collectingCountry id="D1EF764E4C31B351FE5EFA8DF6B3D016" box="[418,486,1369,1394]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C31B351FDA8FA8EF5C5D016" box="[596,656,1369,1394]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2013); in 
<taxonomicName id="6EF84D5D4C31B351FCF1FA8DF429D016" ID-CoL="92B8T" box="[781,892,1369,1394]" class="Insecta" family="Nymphalidae" genus="Heliconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C31B351FCF1FA8DF429D016" box="[781,892,1369,1394]" italics="true" pageId="25" pageNumber="26">Heliconia</emphasis>
</taxonomicName>
bracts (
<bibRefCitation id="CD694B2F4C31B351FC27FA8DF3B1D016" box="[987,1252,1369,1394]" pageId="25" pageNumber="26" refId="ref66551" refString="Frank, J. H. &amp; Barrera, R. (2010) Natural history of Belonuchus Nordmann spp. and allies (Coleoptera: Staphylinidae) in Heliconia L. (Zingiberales: Heliconiaceae) flower bracts. Insecta Mundi, 110, 1 - 12." type="journal article">Frank &amp; Barrera 2010</bibRefCitation>
); distribution in carcasses in 
<collectingCountry id="D1EF764E4C31B351FEADFAA9F694D0F2" box="[337,449,1405,1430]" name="Argentina" pageId="25" pageNumber="26">Argentina</collectingCountry>
(
<bibRefCitation id="CD694B2F4C31B351FE33FAA9F5F9D0F2" box="[463,684,1405,1430]" pageId="25" pageNumber="26" refId="ref61334" refString="Aballay, F., Chani-Posse, M., Ayon, R., Maldonado, M. B. &amp; Centeno, N. (2014) An illustrated key to and diagnoses of the species of Staphylinidae (Coleoptera) associated with decaying carcasses in Argentina. Zootaxa, 3860 (2), 101 - 124. https: // doi. org / 10.11646 / zootaxa. 3860.2.1" type="journal article">
Aballay 
<emphasis id="9B8CEACC4C31B351FDCCFAAAF53DD0F2" box="[560,616,1405,1430]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2014
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BFA76F613D366" blockId="25.[151,1437,1044,1718]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BFA76F613D366" infered="true" pageId="25" pageNumber="26">
= 
<taxonomicName id="6EF84D5D4C31B351FF51FA75F5CCD0DE" ID-CoL="4ZJZ5" authority="Gravenhorst 1806: 72" authorityName="Gravenhorst" authorityPageNumber="72" authorityYear="1806" box="[173,665,1441,1466]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="9B8CEACC4C31B351FF51FA75F6C9D0DE" box="[173,412,1441,1466]" italics="true" pageId="25" pageNumber="26">Staphylinus formosus</emphasis>
<treatmentCitation id="285910CF4C31B351FE5EFA75F5CCD0DE" author="" box="[418,665,1441,1466]" page="72" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE5EFA75F53AD0DE" box="[418,623,1441,1466]" pageId="25" pageNumber="26" refId="ref67002" refString="Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. H. Dieterich, Gottingae (Gottingen), xvi + 248 pp., 1 pl." type="book">Gravenhorst 1806</bibRefCitation>
: 72
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C31B351FD5FFA76F58FD0DE" box="[675,730,1442,1466]" pageId="25" pageNumber="26">Type</typeStatus>
depository: ZMHB LT 
<specimenCount id="BFFEFD574C31B351FC18FA76F4ACD0DD" box="[996,1017,1442,1465]" pageId="25" pageNumber="26" type="female">♀</specimenCount>
, 1 PLT. 
<typeStatus id="7643887C4C31B351FBA9FA76F3D8D0DE" box="[1109,1165,1442,1466]" pageId="25" pageNumber="26">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C31B351FB0AFA75F20FD0DE" box="[1270,1370,1441,1466]" country="Puerto Rico" name="Carolina" pageId="25" pageNumber="26">Carolina</collectingRegion>
, 
<collectingCountry id="D1EF764E4C31B351FA98FA76F2C9D0DE" box="[1380,1436,1442,1466]" name="United States of America" pageId="25" pageNumber="26">USA</collectingCountry>
(&quot; 
<collectingRegion id="6B3CF83C4C31B351FF2AFA11F614D0BA" box="[214,321,1477,1502]" country="Puerto Rico" name="Carolina" pageId="25" pageNumber="26">Carolina</collectingRegion>
&quot;). Note: as synonym of 
<taxonomicName id="6EF84D5D4C31B351FDA4FA12F5BED0BA" ID-CoL="4ZKC5" authorityName="Fabricius" authorityYear="1801" box="[600,747,1478,1502]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufipennis">
<emphasis id="9B8CEACC4C31B351FDA4FA12F5BED0BA" box="[600,747,1478,1502]" italics="true" pageId="25" pageNumber="26">B. rufipennis</emphasis>
</taxonomicName>
(
<bibRefCitation id="CD694B2F4C31B351FD00FA11F48ED0BA" box="[764,987,1477,1502]" pageId="25" pageNumber="26" refId="ref65944" refString="Fauvel, A. (1877) Staphylinides du Bresil et de La Plata, recueillis par Camille van Volxem. Annales de la Societe Entomologique de Belgique, 20 (Compte-Rendu des Seances), xxiii - xxviii." type="journal article">Fauvel 1877: xxvii</bibRefCitation>
). Note: 
<typeStatus id="7643887C4C31B351FBCBFA11F3CBD0BA" box="[1079,1182,1477,1502]" pageId="25" pageNumber="26" type="lectotype">lectotype</typeStatus>
designation (
<treatmentCitation id="285910CF4C31B351FAC6FA12F66FD366" author="" page="728" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FAC6FA12F66FD366" pageId="25" pageNumber="26" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 728</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BF9DAF43ED30A" blockId="25.[151,1437,1044,1718]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BF9DAF43ED30A" infered="true" pageId="25" pageNumber="26">
= 
<taxonomicName id="6EF84D5D4C31B351FF4BF9D9F5EDD342" ID-CoL="5WGHJ" authority="Melsheimer 1844: 35" authorityName="Melsheimer" authorityPageNumber="35" authorityYear="1844" box="[183,696,1549,1574]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="9B8CEACC4C31B351FF4BF9D9F6F6D342" box="[183,419,1549,1574]" italics="true" pageId="25" pageNumber="26">Belonuchus pallipes</emphasis>
<treatmentCitation id="285910CF4C31B351FE4FF9D9F5EDD342" author="" box="[435,696,1549,1574]" page="35" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE4FF9D9F5D0D342" box="[435,645,1549,1574]" pageId="25" pageNumber="26" refId="ref69527" refString="Melsheimer, F. E. (1844) Descriptions of new species of Coleoptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 2 (2 &amp; 5), 26 - 43 &amp; 98 - 118." type="journal article">Melsheimer 1844</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
. Type depository: MCZ? #?ST. Type locality: Pennsylvania. Distribution: USA. Note: as synonym of 
<taxonomicName id="6EF84D5D4C31B351FD6BF9E6F47CD32E" ID-CoL="4ZKC5" authorityName="Fabricius" authorityYear="1801" box="[663,809,1586,1610]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufipennis">
<emphasis id="9B8CEACC4C31B351FD6BF9E6F47CD32E" box="[663,809,1586,1610]" italics="true" pageId="25" pageNumber="26">B. rufipennis</emphasis>
</taxonomicName>
(
<bibRefCitation id="CD694B2F4C31B351FCC4F9E5F341D32E" box="[824,1044,1585,1610]" pageId="25" pageNumber="26" refId="ref65944" refString="Fauvel, A. (1877) Staphylinides du Bresil et de La Plata, recueillis par Camille van Volxem. Annales de la Societe Entomologique de Belgique, 20 (Compte-Rendu des Seances), xxiii - xxviii." type="journal article">Fauvel 1877: xxvii</bibRefCitation>
); no specimens under this name in the Melsheimer collection at the MCZ (
<treatmentCitation id="285910CF4C31B351FD7CF982F40AD30A" author="" box="[640,863,1621,1646]" page="728" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FD7CF982F40AD30A" box="[640,863,1621,1646]" pageId="25" pageNumber="26" refId="ref72912" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946. https: // doi. org / 10.2307 / 3495761" type="journal article">Smetana 1995: 728</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BF9AEF329D3D2" blockId="25.[151,1437,1044,1718]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BF9AEF329D3D2" pageId="25" pageNumber="26">
= 
<taxonomicName id="6EF84D5D4C31B351FF4DF9AEF57DD3F6" ID-CoL="PB7F" authority="Dejean 1833: 63" authorityName="Dejean" authorityPageNumber="63" authorityYear="1833" box="[177,552,1657,1682]" class="Insecta" family="Staphylinidae" genus="Cafius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="9B8CEACC4C31B351FF4DF9AEF608D3F6" box="[177,349,1657,1682]" italics="true" pageId="25" pageNumber="26">Cafius apicalis</emphasis>
<treatmentCitation id="285910CF4C31B351FE9AF9ADF57DD3F6" author="" box="[358,552,1657,1682]" page="63" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE9AF9ADF6A9D3F6" box="[358,508,1657,1682]" pageId="25" pageNumber="26" refId="ref65252" refString="Dejean, P. F. M. A. (1833) Catalogue des Coleopteres de la collection de M. le Comte Dejean. 2 nd Edition. Mequignon-Marvis Pere et Fils, Paris, 176 pp. https: // doi. org / 10.5962 / bhl. title. 8771" type="book">Dejean 1833</bibRefCitation>
: 63
</treatmentCitation>
</taxonomicName>
, unavailable, published in synonymy of unavailable name (
<bibRefCitation id="CD694B2F4C31B351FB25F9AEF7ACD3D2" pageId="25" pageNumber="26" refId="ref67781" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition, adopted by the International Union of Biological Sciences. International Trust for Zoological Nomenclature, London, xxix + 306 pp. [http: // www. nhm. ac. uk / hosted-sites / iczn / code /]" type="book">ICZN 1999: Art. 11.6</bibRefCitation>
). Distribution: 
<collectingCountry id="D1EF764E4C31B351FE5EF949F555D3D2" box="[418,512,1693,1718]" name="Jamaica" pageId="25" pageNumber="26">Jamaica</collectingCountry>
. Note: as synonym of 
<taxonomicName id="6EF84D5D4C31B351FD06F94AF32DD3D2" ID-CoL="4ZJZ5" authority="(Erichson 1840: 426)" authorityName="Erichson" authorityPageNumber="426" authorityYear="1840" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[762,1144,1693,1718]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="9B8CEACC4C31B351FD06F94AF4D5D3D2" box="[762,896,1694,1718]" italics="true" pageId="25" pageNumber="26">B. formosus</emphasis>
(
<treatmentCitation id="285910CF4C31B351FC72F949F325D3D2" author="" box="[910,1136,1693,1718]" page="426" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FC72F949F325D3D2" box="[910,1136,1693,1718]" pageId="25" pageNumber="26" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840: 426</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C31B351FF6BF90AF420D393" blockId="25.[151,1437,1758,2035]" box="[151,885,1758,1783]" pageId="25" pageNumber="26">
<emphasis id="9B8CEACC4C31B351FF6BF90AF7F6D393" bold="true" box="[151,163,1758,1783]" italics="true" pageId="25" pageNumber="26">*</emphasis>
<taxonomicName id="6EF84D5D4C31B351FF5EF90AF519D393" ID-CoL="LCB6" authority="(Sharp 1887)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[162,588,1758,1783]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufiventris" status="comb. nov.">
<emphasis id="9B8CEACC4C31B351FF5EF90AF6FAD393" bold="true" box="[162,431,1758,1783]" italics="true" pageId="25" pageNumber="26">Belonuchus rufiventris</emphasis>
(
<bibRefCitation id="CD694B2F4C31B351FE42F90AF511D393" box="[446,580,1758,1783]" pageId="25" pageNumber="26" refId="ref72534" refString="Sharp, D. (1887) Introduction, Supplement, Index. In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. v - x + 748 - 802 + 803 - 824. [http: // www. sil. si. edu / digitalcollections / bca /]" type="book chapter">Sharp 1887</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C31B351FDABF90AF58DD393" box="[599,728,1758,1783]" pageId="25" pageNumber="26">
<emphasis id="9B8CEACC4C31B351FDABF90AF58DD393" bold="true" box="[599,728,1758,1783]" pageId="25" pageNumber="26">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C31B351FD1BF90AF438D393" box="[743,877,1758,1783]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="25" pageNumber="26">Figs. 17–18</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C31B351FF6BF8D6F65BD207" blockId="25.[151,1437,1758,2035]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="89E811F04C31B351FF6BF8D6F65BD207" pageId="25" pageNumber="26">
<materialsCitation id="19903C834C31B351FF6BF8D6F6BBD25A" ID-GBIF-Occurrence="1915497842" collectionCode="BMNH" country="Mexico" location="Morelos" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Morelos" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C31B351FF6BF8D6F503D27F" ID-CoL="4GBQQ" authority="Sharp 1887: 787" authorityName="Sharp" authorityPageNumber="787" authorityYear="1887" box="[151,598,1794,1819]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufiventris">
<emphasis id="9B8CEACC4C31B351FF6BF8D6F6D8D27F" box="[151,397,1794,1819]" italics="true" pageId="25" pageNumber="26">Philonthus rufiventris</emphasis>
<treatmentCitation id="285910CF4C31B351FE6AF8D6F503D27F" author="" box="[406,598,1794,1819]" page="787" pageId="25" pageNumber="26">
<bibRefCitation id="CD694B2F4C31B351FE6AF8D6F549D27F" box="[406,540,1794,1819]" pageId="25" pageNumber="26" refId="ref72534" refString="Sharp, D. (1887) Introduction, Supplement, Index. In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. v - x + 748 - 802 + 803 - 824. [http: // www. sil. si. edu / digitalcollections / bca /]" type="book chapter">Sharp 1887</bibRefCitation>
: 787
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C31B351FD9FF8D7F5CED27F" box="[611,667,1795,1819]" pageId="25" pageNumber="26">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C31B351FCF4F8D6F437D27F" box="[776,866,1794,1819]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
: &quot;Amecameca in 
<collectingRegion id="6B3CF83C4C31B351FBD7F8D6F3DED27F" box="[1067,1163,1794,1819]" country="Mexico" name="Morelos" pageId="25" pageNumber="26">Morelos</collectingRegion>
&quot; (=Est. 
<collectingCountry id="D1EF764E4C31B351FB15F8D6F210D27F" box="[1257,1349,1794,1819]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
?). 
<typeStatus id="7643887C4C31B351FA98F8D7F2C9D27F" box="[1380,1436,1795,1819]" pageId="25" pageNumber="26">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C31B351FEB0F8F3F6F5D25A" box="[332,416,1831,1854]" pageId="25" pageNumber="26">BMNH</collectionCode>
HT 
<specimenCount id="BFFEFD574C31B351FE24F8F3F6BBD25A" box="[472,494,1831,1854]" pageId="25" pageNumber="26" type="female">♀</specimenCount>
</materialsCitation>
. Distribution: Mexico. Note: distribution within Mexico (
<bibRefCitation id="CD694B2F4C31B351FB7AF8F2F657D207" pageId="25" pageNumber="26" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C31B351FA9DF8F3F2C9D25B" box="[1377,1436,1830,1855]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C31B351FF6BF8BAF34CD2E3" box="[151,1049,1902,1927]" pageId="25" pageNumber="26" type="discussion">
<paragraph id="A94736DE4C31B351FF6BF8BAF34CD2E3" blockId="25.[151,1437,1758,2035]" box="[151,1049,1902,1927]" pageId="25" pageNumber="26">
<emphasis id="9B8CEACC4C31B351FF6BF8BAF65ED2E3" bold="true" box="[151,267,1902,1927]" pageId="25" pageNumber="26">Remarks.</emphasis>
The unique female specimen in BMNH is confirmed as the 
<typeStatus id="7643887C4C31B351FC53F8BAF340D2E3" box="[943,1045,1902,1927]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C31B351FF6BF846F2CCD297" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="A94736DE4C31B351FF6BF846F2CCD297" blockId="25.[151,1437,1758,2035]" pageId="25" pageNumber="26">
<emphasis id="9B8CEACC4C31B351FF6BF846F690D2CF" bold="true" box="[151,453,1938,1963]" pageId="25" pageNumber="26">Type material examined.</emphasis>
Holotype ♀ (BMNH, 
<figureCitation id="31C32A5B4C31B351FD32F846F40FD2CF" box="[718,858,1938,1963]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="25" pageNumber="26">Figs. 17–18</figureCitation>
): card-mounted, “ 
<taxonomicName id="6EF84D5D4C31B351FBDAF846F24BD2CF" box="[1062,1310,1938,1963]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufiventris">Philonthus rufiventris</taxonomicName>
Type D.S. Amecameca. Hoge [in Sharp’s hand] // Type (round white label with red margin) // Amecameca, Morelos. Höge. // B. C. A. Col. 
<date id="DD46101E4C31B351FE3CF80FF6B9D297" box="[448,492,2011,2035]" pageId="25" pageNumber="26">I. 2.</date>
<taxonomicName id="6EF84D5D4C31B351FE0FF80EF46BD297" authority=", Sharp." authorityName="Sharp." box="[499,830,2010,2035]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="rufiventris">Philonthus rufiventris, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFF43F33DD5D4" box="[151,1128,151,176]" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A94736DE4C32B352FF6BFF43F33DD5D4" blockId="26.[151,1128,151,176]" box="[151,1128,151,176]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFF43F64AD5D4" bold="true" box="[151,287,151,176]" pageId="26" pageNumber="27">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C32B352FD46FF43F331D5D4" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C32B352FD46FF43F468D5D4" box="[698,829,151,176]" italics="true" pageId="26" pageNumber="27">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C32B352FCB7FF43F309D5D4" box="[843,1116,151,176]" pageId="26" pageNumber="27" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C32B352FC1CFF4DF34DD5D4" box="[992,1048,151,176]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFF0BF4B5D45B" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="A94736DE4C32B352FF6BFF0BF4D4D59C" blockId="26.[151,1437,223,717]" box="[151,897,223,248]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFF0BF7F6D59C" bold="true" box="[151,163,223,248]" italics="true" pageId="26" pageNumber="27">*</emphasis>
<taxonomicName id="6EF84D5D4C32B352FF5EFF0BF503D59C" ID-CoL="LCB7" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,598,223,248]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufocaudus" status="comb. nov.">
<emphasis id="9B8CEACC4C32B352FF5EFF0BF6EFD59C" bold="true" box="[162,442,223,248]" italics="true" pageId="26" pageNumber="27">Belonuchus rufocaudus</emphasis>
(
<bibRefCitation id="CD694B2F4C32B352FE35FF0BF51BD59C" box="[457,590,223,248]" pageId="26" pageNumber="27" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C32B352FD9EFF0BF5B6D59C" box="[610,739,223,248]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FD9EFF0BF5B6D59C" bold="true" box="[610,739,223,248]" pageId="26" pageNumber="27">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C32B352FD0EFF0BF42DD59C" box="[754,888,223,248]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="26" pageNumber="27">Figs. 19–20</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C32B352FF6BFED0F4B5D45B" blockId="26.[151,1437,223,717]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="89E811F04C32B352FF6BFED0F4B5D45B" pageId="26" pageNumber="27">
<materialsCitation id="19903C834C32B352FF6BFED0F398D479" ID-GBIF-Occurrence="1915497820" box="[151,1229,260,285]" country="Panama" location="Bugaba" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C32B352FF6BFED0F53CD478" ID-CoL="76YRK" authority="Sharp 1885: 412" authorityName="Sharp" authorityPageNumber="412" authorityYear="1885" box="[151,617,260,285]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufocaudus">
<emphasis id="9B8CEACC4C32B352FF6BFED0F6CCD479" box="[151,409,260,285]" italics="true" pageId="26" pageNumber="27">Philonthus rufocaudus</emphasis>
<treatmentCitation id="285910CF4C32B352FE58FED0F53CD478" author="" box="[420,617,260,285]" page="412" pageId="26" pageNumber="27">
<bibRefCitation id="CD694B2F4C32B352FE58FED0F578D479" box="[420,557,260,285]" pageId="26" pageNumber="27" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 412
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C32B352FD8BFED1F5FBD479" box="[631,686,261,285]" pageId="26" pageNumber="27">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C32B352FCDDFED1F428D478" box="[801,893,261,284]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
, 
<location id="AC2760054C32B352FC77FED0F4B1D479" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C32B352FC77FED0F4B1D479" box="[907,996,260,285]" country="Panama" name="Bugaba" pageId="26" pageNumber="27" stateProvince="Chiriqui">Bugaba</location>
, Volcan in 
<collectingRegion id="6B3CF83C4C32B352FB90FED0F398D479" box="[1132,1229,260,285]" country="Panama" name="Chiriqui" pageId="26" pageNumber="27">Chiriqui</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C32B352FB27FED1F48ED45B" ID-GBIF-Occurrence="1915497783" collectionCode="MCP" country="Panama" location="Bugaba" municipality="Distribution" pageId="26" pageNumber="27" specimenCode="BMNH 8" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<typeStatus id="7643887C4C32B352FB27FED1F246D479" box="[1243,1299,261,285]" pageId="26" pageNumber="27">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C32B352FF39FEFCF67BD424" box="[197,302,296,320]" collectionCode="BMNH" pageId="26" pageNumber="27">BMNH 8</specimenCode>
ST: 5 
<location id="AC2760054C32B352FE84FEF3F684D424" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C32B352FE84FEF3F684D424" box="[376,465,295,320]" country="Panama" municipality="Distribution" name="Bugaba" pageId="26" pageNumber="27" stateProvince="Chiriqui">Bugaba</location>
, 3 
<collectingRegion id="6B3CF83C4C32B352FE0CFEF3F518D424" box="[496,589,295,320]" country="Panama" name="Chiriqui" pageId="26" pageNumber="27">Chiriqui</collectingRegion>
[
<collectionCode id="CFE9AE1B4C32B352FDA7FEFCF5CCD424" box="[603,665,296,320]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="26" pageNumber="27">MCP</collectionCode>
seen]. 
<collectingMunicipality id="4923ACA44C32B352FD15FEF3F420D424" box="[745,885,295,320]" pageId="26" pageNumber="27">Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C32B352FC83FEFCF48ED45B" box="[895,987,296,319]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFE98F46ED4B4" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A94736DE4C32B352FF6BFE98F46ED4B4" blockId="26.[151,1437,223,717]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFE98F65ED401" bold="true" box="[151,267,332,357]" pageId="26" pageNumber="27">Remarks.</emphasis>
Sharp did not specify the number of specimens he examined for naming this species, but obviously he based his description on at least one male and one female. Eight specimens were located in the BMNH under this name. The specimens below agree with the original description by 
<bibRefCitation id="CD694B2F4C32B352FC04FE40F3D9D4C9" box="[1016,1164,404,429]" pageId="26" pageNumber="27" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp (1885)</bibRefCitation>
and considered to have been 
<typeStatus id="7643887C4C32B352FEFDFE6CF636D4B4" box="[257,355,440,464]" pageId="26" pageNumber="27" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C32B352FDB9FE63F5F9D4B4" box="[581,684,439,464]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFE08F4BFD7CC" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="A94736DE4C32B352FF6BFE08F4BFD7CC" blockId="26.[151,1437,223,717]" pageId="26" pageNumber="27">
<materialsCitation id="19903C834C32B352FF6BFE08F554D759" ID-GBIF-Occurrence="1915497866" collectionCode="BMNH" collectorName="Type D. S. &amp; Sharp's &amp; Type &amp; Bugaba" country="Panama" location="Panama" pageId="26" pageNumber="27" specimenCount="2" specimenCount-male="2" typeStatus="lectotype">
<emphasis id="9B8CEACC4C32B352FF6BFE08F698D491" bold="true" box="[151,461,476,501]" pageId="26" pageNumber="27">
<typeStatus id="7643887C4C32B352FF6BFE08F787D491" box="[151,210,476,501]" pageId="26" pageNumber="27">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C32B352FE26FE09F51FD491" box="[474,586,477,501]" pageId="26" pageNumber="27" type="lectotype">Lectotype</typeStatus>
<specimenCount id="BFFEFD574C32B352FDA4FE09F53AD490" box="[600,623,477,500]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
, here designated (
<collectionCode id="CFE9AE1B4C32B352FCB2FE08F4F3D490" box="[846,934,476,500]" pageId="26" pageNumber="27">BMNH</collectionCode>
, 
<figureCitation id="31C32A5B4C32B352FC44FE08F31DD490" box="[952,1096,476,501]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="26" pageNumber="27">Figs. 19–20</figureCitation>
): card-mounted, “ 
<specimenCount id="BFFEFD574C32B352FAE2FE08F261D490" box="[1310,1332,476,500]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
(in ink) 
<taxonomicName id="6EF84D5D4C32B352FF39FE2BF691D77C" box="[197,452,511,536]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufocaudus">Philonthus rufocaudus</taxonomicName>
<collectorName id="040D53084C32B352FE31FDD4F514D77C" box="[461,577,512,536]" pageId="26" pageNumber="27">
<typeStatus id="7643887C4C32B352FE31FDD4F550D77C" box="[461,517,512,536]" pageId="26" pageNumber="27">Type</typeStatus>
D.S.
</collectorName>
<collectingCountry id="D1EF764E4C32B352FDB7FDD4F5F2D773" box="[587,679,512,535]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
. Bugaba. Champion [in 
<collectorName id="040D53084C32B352FC38FE2BF34DD77C" box="[964,1048,511,536]" pageId="26" pageNumber="27">Sharp’s</collectorName>
hand] // 
<collectorName id="040D53084C32B352FB7BFDD4F3EBD77C" box="[1159,1214,512,536]" pageId="26" pageNumber="27">
<typeStatus id="7643887C4C32B352FB7BFDD4F3EBD77C" box="[1159,1214,512,536]" pageId="26" pageNumber="27">Type</typeStatus>
</collectorName>
(round white label with red margin) // 
<collectorName id="040D53084C32B352FE55FDF0F554D759" box="[425,513,548,573]" pageId="26" pageNumber="27">Bugaba</collectorName>
</materialsCitation>
, 
<materialsCitation id="19903C834C32B352FDF3FDF1F5C6D7E1" ID-GBIF-Occurrence="1915497795" collectingDate="I. 2." collectionCode="BMNH" collectorName="B. C. A. Col." country="Panama" location="Panama" pageId="26" pageNumber="27" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paralectotype">
<collectingCountry id="D1EF764E4C32B352FDF3FDF1F53ED758" box="[527,619,549,572]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
. Champion. // 
<collectorName id="040D53084C32B352FCEBFDF1F4E7D759" box="[791,946,548,573]" pageId="26" pageNumber="27">B. C. A. Col.</collectorName>
<collectingDate id="CD02E9F64C32B352FC47FDF1F4BFD758" box="[955,1002,548,572]" pageId="26" pageNumber="27">
<date id="DD46101E4C32B352FC47FDF1F4BFD758" box="[955,1002,548,572]" pageId="26" pageNumber="27">I. 2.</date>
</collectingDate>
<taxonomicName id="6EF84D5D4C32B352FC08FDF0F21ED759" authority=", Sharp." authorityName="Sharp." box="[1012,1355,548,573]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufocaudus">Philonthus rufocaudus, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”. 
<typeStatus id="7643887C4C32B352FD1CFD93F4D6D704" box="[736,899,583,608]" pageId="26" pageNumber="27" type="paralectotype">Paralectotypes</typeStatus>
(
<collectionCode id="CFE9AE1B4C32B352FC69FD9CF4A5D704" box="[917,1008,584,608]" pageId="26" pageNumber="27">BMNH</collectionCode>
): 
<specimenCount id="BFFEFD574C32B352FBF8FD9CF367D73B" box="[1028,1074,584,608]" pageId="26" pageNumber="27" type="male">1 ♂</specimenCount>
, 
<specimenCount id="BFFEFD574C32B352FBBCFD9CF339D73B" box="[1088,1132,584,608]" pageId="26" pageNumber="27" type="female">3 ♀</specimenCount>
, card-mounted, with same locality and collection data as 
<typeStatus id="7643887C4C32B352FDD5FDB8F5C6D7E1" box="[553,659,620,645]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
</materialsCitation>
; 3 ♀, card-mounted, “V. de Chriqui 
<quantity id="6E009B3B4C32B352FBBCFDB8F39DD7E1" box="[1088,1224,620,645]" metricMagnitude="2" metricUnit="m" metricValue="6.1341" metricValueMax="12.192" metricValueMin="0.0762" pageId="26" pageNumber="27" unit="ft" value="2012.5" valueMax="4000.0" valueMin="25.0">25–4,000 ft</quantity>
. Champion (white label) // B. C. A. Col. 
<date id="DD46101E4C32B352FE40FD44F6BDD7CC" box="[444,488,656,680]" pageId="26" pageNumber="27">I. 2.</date>
<taxonomicName id="6EF84D5D4C32B352FE13FD5BF417D7CC" authority=", Sharp." authorityName="Sharp." box="[495,834,655,680]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufocaudus">Philonthus rufocaudus, Sharp.</taxonomicName>
(white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFD60F33DD7A8" box="[151,1128,692,717]" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A94736DE4C32B352FF6BFD60F33DD7A8" blockId="26.[151,1437,223,717]" box="[151,1128,692,717]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFD60F64AD7A9" bold="true" box="[151,287,692,717]" pageId="26" pageNumber="27">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C32B352FD46FD60F331D7A8" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,692,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C32B352FD46FD60F468D7A9" box="[698,829,692,717]" italics="true" pageId="26" pageNumber="27">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C32B352FCB7FD60F309D7A8" box="[843,1116,692,717]" pageId="26" pageNumber="27" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C32B352FC1CFD61F34DD7A9" box="[992,1048,692,717]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFD2FF4C2D6E4" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="A94736DE4C32B352FF6BFD2FF424D670" blockId="26.[151,1437,763,1221]" box="[151,881,763,789]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFD2FF7F6D670" bold="true" box="[151,163,763,788]" italics="true" pageId="26" pageNumber="27">*</emphasis>
<taxonomicName id="6EF84D5D4C32B352FF5EFD2FF512D671" ID-CoL="LCB9" authority="(Sharp 1885)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,583,763,789]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufopygus" status="comb. nov.">
<emphasis id="9B8CEACC4C32B352FF5EFD2FF6FFD670" bold="true" box="[162,426,763,788]" italics="true" pageId="26" pageNumber="27">Belonuchus rufopygus</emphasis>
(
<bibRefCitation id="CD694B2F4C32B352FE46FD28F56AD671" box="[442,575,764,789]" pageId="26" pageNumber="27" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C32B352FDAFFD28F586D671" box="[595,723,764,789]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FDAFFD28F586D671" bold="true" box="[595,723,764,789]" pageId="26" pageNumber="27">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C32B352FD1FFD28F43CD670" box="[739,873,764,789]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="26" pageNumber="27">Figs. 21–22</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C32B352FF6BFCCBF4C2D6E4" blockId="26.[151,1437,763,1221]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="89E811F04C32B352FF6BFCCBF4C2D6E4" infered="true" pageId="26" pageNumber="27">
<taxonomicName id="6EF84D5D4C32B352FF6BFCCBF524D65C" ID-CoL="4GBQX" authority="Sharp 1885: 411" authorityName="Sharp" authorityPageNumber="411" authorityYear="1885" box="[151,625,799,824]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufopygus">
<emphasis id="9B8CEACC4C32B352FF6BFCCBF6C4D65C" box="[151,401,799,824]" italics="true" pageId="26" pageNumber="27">Philonthus rufopygus</emphasis>
<treatmentCitation id="285910CF4C32B352FE5EFCCBF524D65C" author="" box="[418,625,799,824]" page="411" pageId="26" pageNumber="27">
<bibRefCitation id="CD694B2F4C32B352FE5EFCCBF565D65C" box="[418,560,799,824]" pageId="26" pageNumber="27" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 411
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C32B352FD7AFCF4F5E8D65C" box="[646,701,800,824]" pageId="26" pageNumber="27">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C32B352FCC0FCF4F4CCD653" box="[828,921,800,823]" country="Panama" name="Panama" pageId="26" pageNumber="27">
<collectingCountry id="D1EF764E4C32B352FCC0FCF4F4CCD653" box="[828,921,800,823]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
</collectingRegion>
: Bugaba. 
<typeStatus id="7643887C4C32B352FBE7FCF4F306D65C" box="[1051,1107,800,824]" pageId="26" pageNumber="27">Type</typeStatus>
depository: BMNH 8 ST. Distribution: 
<collectingRegion id="6B3CF83C4C32B352FEA0FC91F6E0D638" box="[348,437,837,860]" country="Panama" name="Panama" pageId="26" pageNumber="27">
<collectingCountry id="D1EF764E4C32B352FEA0FC91F6E0D638" box="[348,437,837,860]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
</collectingRegion>
[
<collectingRegion id="6B3CF83C4C32B352FE3FFC90F54ED639" box="[451,539,836,861]" country="Mexico" name="Mexico" pageId="26" pageNumber="27">
<collectingCountry id="D1EF764E4C32B352FE3FFC90F54ED639" box="[451,539,836,861]" name="Mexico" pageId="26" pageNumber="27">Mexico</collectingCountry>
</collectingRegion>
in error]. Note: reported from 
<collectingRegion id="6B3CF83C4C32B352FC86FC90F49AD639" box="[890,975,836,861]" country="Mexico" name="Mexico" pageId="26" pageNumber="27">
<collectingCountry id="D1EF764E4C32B352FC86FC90F49AD639" box="[890,975,836,861]" name="Mexico" pageId="26" pageNumber="27">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C32B352FC23FC90F254D639" box="[991,1281,836,861]" pageId="26" pageNumber="27" refId="ref74121" refString="Zaragoza Caballero, S. (1976) Coleopteros de algunas bromelias epifitas y doce nuevos registros de especies para la fauna mexicana. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Zoologia, 45 (1974), 111 - 118." type="journal article">Zaragoza Caballero 1976</bibRefCitation>
, but based on misidentified specimens [J.L. Navarrete-Heredia, unpublished]).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFC58F6EED689" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A94736DE4C32B352FF6BFC58F6EED689" blockId="26.[151,1437,763,1221]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFC58F65ED6C1" bold="true" box="[151,267,908,933]" pageId="26" pageNumber="27">Remarks.</emphasis>
Sharp explicitly mentioned several specimens as part of the 
<typeStatus id="7643887C4C32B352FC2DFC59F354D6C1" box="[977,1025,909,933]" pageId="26" pageNumber="27">type</typeStatus>
series. The specimens listed below agree with the original description by 
<bibRefCitation id="CD694B2F4C32B352FD89FC7BF45CD6AC" box="[629,777,943,968]" pageId="26" pageNumber="27" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp (1885)</bibRefCitation>
and considered to have been 
<typeStatus id="7643887C4C32B352FBA7FC64F3E8D6AC" box="[1115,1213,944,968]" pageId="26" pageNumber="27" type="syntype">syntypes</typeStatus>
prior to the present 
<typeStatus id="7643887C4C32B352FF39FC00F679D689" box="[197,300,980,1005]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
designation.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFC23F424D1C4" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="A94736DE4C32B352FF6BFC23F424D1C4" blockId="26.[151,1437,763,1221]" pageId="26" pageNumber="27">
<materialsCitation id="19903C834C32B352FF6BFC23F694D13C" ID-GBIF-Occurrence="1915497865" collectionCode="BMNH" collectorName="Type D. S." country="Panama" location="Panama" pageId="26" pageNumber="27" specimenCount="2" specimenCount-male="2" typeStatus="lectotype">
<emphasis id="9B8CEACC4C32B352FF6BFC23F698D174" bold="true" box="[151,461,1015,1040]" pageId="26" pageNumber="27">
<typeStatus id="7643887C4C32B352FF6BFC23F787D174" box="[151,210,1015,1040]" pageId="26" pageNumber="27">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C32B352FE26FC2CF51FD174" box="[474,586,1016,1040]" pageId="26" pageNumber="27" type="lectotype">Lectotype</typeStatus>
<specimenCount id="BFFEFD574C32B352FDA4FC2CF53AD16B" box="[600,623,1016,1039]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
, here designated (
<collectionCode id="CFE9AE1B4C32B352FCB2FC2CF4F3D174" box="[846,934,1016,1040]" pageId="26" pageNumber="27">BMNH</collectionCode>
, 
<figureCitation id="31C32A5B4C32B352FC44FC23F31CD174" box="[952,1097,1015,1040]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="26" pageNumber="27">Figs. 21–22</figureCitation>
): card-mounted, “ 
<specimenCount id="BFFEFD574C32B352FAE2FC2CF261D174" box="[1310,1332,1016,1040]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
(in ink) 
<taxonomicName id="6EF84D5D4C32B352FF39FBC8F668D151" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[197,317,1052,1077]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
rufo-pygus 
<collectorName id="040D53084C32B352FE38FBC9F561D150" box="[452,564,1052,1077]" pageId="26" pageNumber="27">
<typeStatus id="7643887C4C32B352FE38FBC9F6A9D151" box="[452,508,1053,1077]" pageId="26" pageNumber="27">Type</typeStatus>
D.S.
</collectorName>
<collectingCountry id="D1EF764E4C32B352FDC6FBC9F5C3D150" box="[570,662,1053,1076]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
. Bugaba. Champion [in Sharp’s hand] // 
<typeStatus id="7643887C4C32B352FBA6FBC9F3C4D151" box="[1114,1169,1053,1077]" pageId="26" pageNumber="27">Type</typeStatus>
(round white label with red margin) // Bugaba
</materialsCitation>
, 
<materialsCitation id="19903C834C32B352FE31FB94F438D1C4" ID-GBIF-Occurrence="1915497831" collectingDate="I. 2." collectionCode="BMNH" collectorName="B. C. A. Col." country="Panama" location="Panama" pageId="26" pageNumber="27" specimenCount="7" specimenCount-female="7" typeStatus="paralectotype">
<collectingCountry id="D1EF764E4C32B352FE31FB94F57CD133" box="[461,553,1088,1111]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
. Champion. // 
<collectorName id="040D53084C32B352FD2CFB94F432D13C" box="[720,871,1087,1112]" pageId="26" pageNumber="27">B. C. A. Col.</collectorName>
<collectingDate id="CD02E9F64C32B352FC93FB94F4C9D13C" box="[879,924,1088,1112]" pageId="26" pageNumber="27">
<date id="DD46101E4C32B352FC93FB94F4C9D13C" box="[879,924,1088,1112]" pageId="26" pageNumber="27">I. 2.</date>
</collectingDate>
<taxonomicName id="6EF84D5D4C32B352FC58FBEBF3BBD13C" authority=", Sharp." authorityName="Sharp." box="[932,1262,1087,1112]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rufopygus">Philonthus rufopygus, Sharp.</taxonomicName>
(white label) // Sharp Coll. 1905.-313. (white label)”. 
<typeStatus id="7643887C4C32B352FD71FBB0F464D119" box="[653,817,1124,1149]" pageId="26" pageNumber="27" type="paralectotype">Paralectotypes</typeStatus>
(
<collectionCode id="CFE9AE1B4C32B352FCB9FBB0F4F5D118" box="[837,928,1124,1148]" pageId="26" pageNumber="27">BMNH</collectionCode>
): 
<specimenCount id="BFFEFD574C32B352FC4BFBB1F4B3D118" box="[951,998,1125,1148]" pageId="26" pageNumber="27" type="female">7 ♀</specimenCount>
(2 of them mounted together), cardmounted, with same locality and collection data as 
<typeStatus id="7643887C4C32B352FCFEFB53F438D1C4" box="[770,877,1159,1184]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFB78F505D069" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A94736DE4C32B352FF6BFB78F33DD1A0" blockId="26.[151,1437,763,1221]" box="[151,1128,1196,1221]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFB78F64AD1A1" bold="true" box="[151,287,1196,1221]" pageId="26" pageNumber="27">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C32B352FD46FB78F331D1A0" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C32B352FD46FB78F468D1A1" box="[698,829,1196,1221]" italics="true" pageId="26" pageNumber="27">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C32B352FCB7FB78F309D1A0" box="[843,1116,1196,1221]" pageId="26" pageNumber="27" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C32B352FC1CFB79F34DD1A1" box="[992,1048,1196,1221]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="A94736DE4C32B352FF6BFB27F505D069" blockId="26.[151,1437,1267,1365]" box="[151,592,1267,1293]" pageId="26" pageNumber="27">
<taxonomicName id="6EF84D5D4C32B352FF6BFB27F505D069" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,592,1267,1293]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="9B8CEACC4C32B352FF6BFB27F6C0D068" bold="true" box="[151,405,1267,1292]" italics="true" pageId="26" pageNumber="27">Belonuchus rugicollis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BFAC3F4FFD3F7" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="A94736DE4C32B352FF6BFAC3F435D031" blockId="26.[151,1437,1267,1365]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="89E811F04C32B352FF6BFAC3F435D031" pageId="26" pageNumber="27">
<materialsCitation id="19903C834C32B352FF6BFAC3F423D04B" ID-GBIF-Occurrence="1915497849" box="[151,886,1303,1328]" country="Peru" location="Peru" pageId="26" pageNumber="27" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C32B352FF6BFAC3F5C4D054" ID-CoL="LCBB" authority="Bernhauer 1916 c: 276" authorityName="Bernhauer" authorityPageNumber="276" authorityYear="1916" box="[151,657,1303,1328]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="9B8CEACC4C32B352FF6BFAC3F6D8D054" box="[151,397,1303,1328]" italics="true" pageId="26" pageNumber="27">Belonuchus rugicollis</emphasis>
<treatmentCitation id="285910CF4C32B352FE68FAC3F5C4D054" author="" box="[404,657,1303,1328]" page="276" pageId="26" pageNumber="27">
<bibRefCitation id="CD694B2F4C32B352FE68FAC3F50CD054" box="[404,601,1303,1328]" pageId="26" pageNumber="27" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 276
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C32B352FD61FACCF580D054" box="[669,725,1304,1328]" pageId="26" pageNumber="27">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C32B352FCC3FACCF423D04B" box="[831,886,1304,1327]" name="Peru" pageId="26" pageNumber="27">Peru</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C32B352FC7DFACCF2CCD04B" ID-GBIF-Occurrence="1915497824" box="[897,1433,1303,1328]" collectionCode="FMNH" collectorName="HT Distribution" country="Peru" location="Peru" pageId="26" pageNumber="27" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C32B352FC7DFACCF4EDD054" box="[897,952,1304,1328]" pageId="26" pageNumber="27">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C32B352FBBAFACCF3C2D04B" box="[1094,1175,1304,1327]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="26" pageNumber="27">FMNH</collectionCode>
<collectorName id="040D53084C32B352FB62FACCF202D054" box="[1182,1367,1303,1328]" pageId="26" pageNumber="27">HT Distribution</collectorName>
: 
<collectingCountry id="D1EF764E4C32B352FA9EFACCF2CCD04B" box="[1378,1433,1304,1327]" name="Peru" pageId="26" pageNumber="27">Peru</collectingCountry>
</materialsCitation>
. Note: Peru (
<bibRefCitation id="CD694B2F4C32B352FEB0FAE8F6B4D030" box="[332,481,1340,1365]" pageId="26" pageNumber="27" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
). Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C32B352FF6BFA57F567D0F8" blockId="26.[151,1437,1411,1544]" box="[151,562,1411,1437]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BFA57F7F6D0F8" bold="true" box="[151,163,1411,1436]" italics="true" pageId="26" pageNumber="27">*</emphasis>
<taxonomicName id="6EF84D5D4C32B352FF5EFA57F567D0F8" ID-CoL="LCBC" authority="Erichson 1840" authorityName="Erichson" authorityYear="1840" box="[162,562,1411,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="satyrus">
<emphasis id="9B8CEACC4C32B352FF5EFA57F6DDD0FF" bold="true" box="[162,392,1411,1436]" italics="true" pageId="26" pageNumber="27">Belonuchus satyrus</emphasis>
<bibRefCitation id="CD694B2F4C32B352FE6CFA50F567D0F8" box="[400,562,1412,1437]" pageId="26" pageNumber="27" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C32B352FF6BFA73F65BD36C" blockId="26.[151,1437,1411,1544]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="89E811F04C32B352FF6BFA73F65BD36C" pageId="26" pageNumber="27">
<taxonomicName id="6EF84D5D4C32B352FF6BFA73F534D0A4" ID-CoL="LCBC" authority="Erichson 1840: 421" authorityName="Erichson" authorityPageNumber="421" authorityYear="1840" box="[151,609,1447,1472]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="satyrus">
<emphasis id="9B8CEACC4C32B352FF6BFA73F621D0A4" box="[151,372,1447,1472]" italics="true" pageId="26" pageNumber="27">Belonuchus satyrus</emphasis>
<treatmentCitation id="285910CF4C32B352FE81FA73F534D0A4" author="" box="[381,609,1447,1472]" page="421" pageId="26" pageNumber="27">
<bibRefCitation id="CD694B2F4C32B352FE81FA73F572D0A4" box="[381,551,1447,1472]" pageId="26" pageNumber="27" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840</bibRefCitation>
: 421
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C32B352FD92FA7CF5F0D0A4" box="[622,677,1448,1472]" pageId="26" pageNumber="27">Type</typeStatus>
locality: Columbia. 
<typeStatus id="7643887C4C32B352FC68FA7CF49ED0A4" box="[916,971,1448,1472]" pageId="26" pageNumber="27">Type</typeStatus>
depository: ZMHB? #?ST. Distribution: 
<collectingRegion id="6B3CF83C4C32B352FF39FA19F674D080" box="[197,289,1485,1508]" country="Panama" name="Panama" pageId="26" pageNumber="27">
<collectingCountry id="D1EF764E4C32B352FF39FA19F674D080" box="[197,289,1485,1508]" name="Panama" pageId="26" pageNumber="27">Panama</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C32B352FED3FA18F6F5D081" box="[303,416,1484,1509]" name="Colombia" pageId="26" pageNumber="27">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C32B352FE53FA18F570D081" box="[431,549,1484,1509]" name="Venezuela" pageId="26" pageNumber="27">Venezuela</collectingCountry>
, 
<collectingCountry id="D1EF764E4C32B352FDCFFA18F5C3D081" box="[563,662,1484,1509]" name="Trinidad and Tobago" pageId="26" pageNumber="27">Trinidad</collectingCountry>
; 
<collectingCountry id="D1EF764E4C32B352FD59FA18F473D081" box="[677,806,1484,1509]" name="Costa Rica" pageId="26" pageNumber="27">
<collectingRegion id="6B3CF83C4C32B352FD59FA18F5B0D080" box="[677,741,1484,1508]" country="Tanzania" name="Pwani" pageId="26" pageNumber="27">Costa</collectingRegion>
Rica
</collectingCountry>
. Notes: biology, in 
<taxonomicName id="6EF84D5D4C32B352FBF0FA18F32FD081" ID-CoL="92B8T" box="[1036,1146,1484,1509]" class="Insecta" family="Nymphalidae" genus="Heliconia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C32B352FBF0FA18F32FD081" box="[1036,1146,1484,1509]" italics="true" pageId="26" pageNumber="27">Heliconia</emphasis>
</taxonomicName>
bracts (
<bibRefCitation id="CD694B2F4C32B352FB26FA18F657D36C" pageId="26" pageNumber="27" refId="ref66551" refString="Frank, J. H. &amp; Barrera, R. (2010) Natural history of Belonuchus Nordmann spp. and allies (Coleoptera: Staphylinidae) in Heliconia L. (Zingiberales: Heliconiaceae) flower bracts. Insecta Mundi, 110, 1 - 12." type="journal article">Frank &amp; Barrera 2010</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C32B352FF6BF9E6F4DAD32F" blockId="26.[151,1436,1586,1863]" box="[151,911,1586,1611]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BF9E6F7F6D32F" bold="true" box="[151,163,1586,1611]" italics="true" pageId="26" pageNumber="27">*</emphasis>
<taxonomicName id="6EF84D5D4C32B352FF5EF9E6F530D32F" ID-CoL="LCBG" authority="(Sharp 1876)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Sharp" baseAuthorityYear="1876" box="[162,613,1586,1611]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="serraticornis" status="comb. nov.">
<emphasis id="9B8CEACC4C32B352FF5EF9E6F69DD32F" bold="true" box="[162,456,1586,1611]" italics="true" pageId="26" pageNumber="27">Belonuchus serraticornis</emphasis>
(Sharp 1876)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C32B352FD8CF9E6F5A4D32F" box="[624,753,1586,1611]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FD8CF9E6F5A4D32F" bold="true" box="[624,753,1586,1611]" pageId="26" pageNumber="27">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C32B352FCFCF9E6F4D2D32F" box="[768,903,1586,1611]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="26" pageNumber="27">Figs. 23–24</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C32B352FF6BF982F4FFD3F7" blockId="26.[151,1436,1586,1863]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="89E811F04C32B352FF6BF982F4FFD3F7" pageId="26" pageNumber="27">
<materialsCitation id="19903C834C32B352FF6BF982F604D3F7" ID-GBIF-Occurrence="1915497787" collectionCode="BMNH" country="Brazil" location="Distribution" pageId="26" pageNumber="27" specimenCount="2" stateProvince="Amazonas" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C32B352FF6BF982F52BD30B" ID-CoL="4GBTH" authority="Sharp 1876 a: 179" authorityName="Sharp" authorityPageNumber="179" authorityYear="1876" box="[151,638,1622,1647]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="serraticornis">
<emphasis id="9B8CEACC4C32B352FF6BF982F6FFD30A" box="[151,426,1622,1647]" italics="true" pageId="26" pageNumber="27">Philonthus serraticornis</emphasis>
<treatmentCitation id="285910CF4C32B352FE4EF982F52BD30B" author="" box="[434,638,1622,1647]" page="179" pageId="26" pageNumber="27">
<bibRefCitation id="CD694B2F4C32B352FE4EF982F511D30B" box="[434,580,1622,1647]" pageId="26" pageNumber="27" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 179
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C32B352FD76F983F594D30B" box="[650,705,1623,1647]" pageId="26" pageNumber="27">Type</typeStatus>
locality: Ega. 
<typeStatus id="7643887C4C32B352FC96F983F4F7D30B" box="[874,930,1623,1647]" pageId="26" pageNumber="27">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C32B352FBCCF983F3D1D30A" box="[1072,1156,1623,1646]" pageId="26" pageNumber="27">BMNH</collectionCode>
HT. 
<location id="AC2760054C32B352FB3CF982F219D30B" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C32B352FB3CF982F219D30B" box="[1216,1356,1622,1647]" country="Brazil" name="Distribution" pageId="26" pageNumber="27" stateProvince="Amazonas">Distribution</location>
: 
<collectingCountry id="D1EF764E4C32B352FAA4F982F2C9D30B" box="[1368,1436,1622,1647]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C32B352FF31F9AFF61FD3F7" box="[205,330,1659,1683]" country="Venezuela" name="Amazonas" pageId="26" pageNumber="27">Amazonas</collectingRegion>
)
</materialsCitation>
. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C32B352FCDFF9A8F40ED3F7" box="[803,859,1658,1683]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BF94AF48BD3D3" box="[151,990,1694,1719]" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A94736DE4C32B352FF6BF94AF48BD3D3" blockId="26.[151,1436,1586,1863]" box="[151,990,1694,1719]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BF94AF65ED3D3" bold="true" box="[151,267,1694,1719]" pageId="26" pageNumber="27">Remarks.</emphasis>
The “unique” specimen in BMNH is confirmed as the 
<typeStatus id="7643887C4C32B352FC88F94AF48FD3D3" box="[884,986,1694,1719]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BF916F381D247" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="A94736DE4C32B352FF6BF916F381D247" blockId="26.[151,1436,1586,1863]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BF916F690D3BF" bold="true" box="[151,453,1730,1755]" pageId="26" pageNumber="27">
<typeStatus id="7643887C4C32B352FF6BF916F787D3BF" box="[151,210,1730,1755]" pageId="26" pageNumber="27">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="7643887C4C32B352FE32F916F563D3BF" box="[462,566,1730,1755]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
<specimenCount id="BFFEFD574C32B352FDBCF917F500D3BE" box="[576,597,1731,1754]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
(incomplete genitalia) (
<collectionCode id="CFE9AE1B4C32B352FC96F917F496D3BF" box="[874,963,1731,1755]" pageId="26" pageNumber="27">BMNH</collectionCode>
, 
<figureCitation id="31C32A5B4C32B352FC2DF916F308D3BF" box="[977,1117,1730,1755]" captionStart="FIGURES 13–24" captionStartId="23.[151,264,1730,1753]" captionTargetBox="[227,1360,195,1709]" captionTargetId="figure@23.[227,1360,195,1709]" captionTargetPageId="23" captionText="FIGURES 13–24 [BMNH ©]. Type material (habitus and labels): 13, 14, Philonthus optatus Sharp, holotype; 15, 16, Philonthus platypterus Sharp, lectotype; 17, 18, Philonthus rufiventris Sharp, holotype; 19, 20, Philonthus rufocaudus Sharp, lectotype; 21, 22, Philonthus rufopygus Sharp, lectotype; 23, 24, Philonthus serraticornis Sharp, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454309/files/figure.png" pageId="26" pageNumber="27">Figs. 23–24</figureCitation>
): card-mounted, “ 
<specimenCount id="BFFEFD574C32B352FAD7F917F214D3BF" box="[1323,1345,1731,1755]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
// 
<typeStatus id="7643887C4C32B352FA98F917F2C9D3BF" box="[1380,1436,1731,1755]" pageId="26" pageNumber="27">Type</typeStatus>
(round white label with red margin) // Ega (round green label) [in Sharp’s hand]// S America 
<collectingCountry id="D1EF764E4C32B352FAC7F932F22AD39B" box="[1339,1407,1766,1791]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
// 
<taxonomicName id="6EF84D5D4C32B352FF39F8DEF69AD247" authorityName="Sharp" authorityYear="1876" box="[197,463,1802,1827]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="serraticornis">Philonthus serraticornis</taxonomicName>
<typeStatus id="7643887C4C32B352FE29F8DFF559D247" box="[469,524,1803,1827]" pageId="26" pageNumber="27">Type</typeStatus>
D.S. [in Sharp’s hand] // Sharp Coll. 1905.-313. (white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B352FF6BF8FAF6A9D2EE" pageId="26" pageNumber="27" type="discussion">
<paragraph id="A94736DE4C32B352FF6BF8FAF33DD223" blockId="26.[151,1436,1586,1863]" box="[151,1128,1838,1863]" pageId="26" pageNumber="27">
<emphasis id="9B8CEACC4C32B352FF6BF8FAF64AD223" bold="true" box="[151,287,1838,1863]" pageId="26" pageNumber="27">Comments.</emphasis>
In accordance with the diagnosis of 
<taxonomicName id="6EF84D5D4C32B352FD46F8FAF331D223" authority="(Chani-Posse et al. 2018)" authorityName="Chani-Posse et al." authorityYear="2018" box="[698,1124,1838,1863]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C32B352FD46F8FAF468D223" box="[698,829,1838,1863]" italics="true" pageId="26" pageNumber="27">Belonuchus</emphasis>
(
<bibRefCitation id="CD694B2F4C32B352FCB7F8FAF309D223" box="[843,1116,1838,1863]" pageId="26" pageNumber="27" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C32B352FC1CF8FBF34DD223" box="[992,1048,1838,1863]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2018
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="A94736DE4C32B352FF6BF8A5F6A9D2EE" blockId="26.[151,1436,1905,2002]" box="[151,508,1905,1930]" pageId="26" pageNumber="27">
<taxonomicName id="6EF84D5D4C32B352FF6BF8A5F6A9D2EE" authority="Sharp 1876" authorityName="Sharp" authorityYear="1876" box="[151,508,1905,1930]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="setiger">
<emphasis id="9B8CEACC4C32B352FF6BF8A5F621D2EE" bold="true" box="[151,372,1905,1930]" italics="true" pageId="26" pageNumber="27">Belonuchus setiger</emphasis>
Sharp 1876
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C32B353FF6BF841F470D6C0" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="A94736DE4C32B352FF6BF841F3D9D2B6" blockId="26.[151,1436,1905,2002]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="89E811F04C32B352FF6BF841F3D9D2B6" pageId="26" pageNumber="27">
<materialsCitation id="19903C834C32B352FF6BF841F566D2B6" ID-GBIF-Occurrence="1915497766" collectionCode="MCP" country="Brazil" location="Distribution" pageId="26" pageNumber="27" specimenCode="BMNH 2" specimenCount="2" specimenCount-male="2" stateProvince="Amazonas" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C32B352FF6BF841F506D2CA" ID-CoL="LCBH" authority="Sharp 1876 a: 164" authorityName="Sharp" authorityPageNumber="164" authorityYear="1876" box="[151,595,1941,1966]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="setiger">
<emphasis id="9B8CEACC4C32B352FF6BF841F627D2C9" box="[151,370,1941,1966]" italics="true" pageId="26" pageNumber="27">Belonuchus setiger</emphasis>
<treatmentCitation id="285910CF4C32B352FE83F841F506D2CA" author="" box="[383,595,1941,1966]" page="164" pageId="26" pageNumber="27">
<bibRefCitation id="CD694B2F4C32B352FE83F841F540D2CA" box="[383,533,1941,1966]" pageId="26" pageNumber="27" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 164
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C32B352FD9FF842F5CED2CA" box="[611,667,1942,1966]" pageId="26" pageNumber="27">Type</typeStatus>
locality: Ega. 
<typeStatus id="7643887C4C32B352FCB3F842F4D3D2CA" box="[847,902,1942,1966]" pageId="26" pageNumber="27">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C32B352FBE2F842F3D9D2CA" box="[1054,1164,1942,1966]" collectionCode="BMNH" pageId="26" pageNumber="27">BMNH 2</specimenCode>
ST: 
<specimenCount id="BFFEFD574C32B352FB31F842F3A9D2C9" box="[1229,1276,1942,1966]" pageId="26" pageNumber="27" type="male">2 ♂</specimenCount>
[
<collectionCode id="CFE9AE1B4C32B352FAF3F842F218D2CA" box="[1295,1357,1942,1966]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="26" pageNumber="27">MCP</collectionCode>
seen]. 
<location id="AC2760054C32B352FF39F86DF604D2B6" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C32B352FF39F86DF604D2B6" box="[197,337,1977,2002]" country="Brazil" name="Distribution" pageId="26" pageNumber="27" stateProvince="Amazonas">Distribution</location>
: 
<collectingCountry id="D1EF764E4C32B352FEA7F86DF6CAD2B6" box="[347,415,1977,2002]" name="Brazil" pageId="26" pageNumber="27">Brazil</collectingCountry>
(
<collectingRegion id="6B3CF83C4C32B352FE52F86EF57ED2B6" box="[430,555,1978,2002]" country="Venezuela" name="Amazonas" pageId="26" pageNumber="27">Amazonas</collectingRegion>
)
</materialsCitation>
. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C32B352FBF8F86EF369D2B6" box="[1028,1084,1977,2002]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFF43F56FD5D4" blockId="27.[151,1436,151,357]" box="[151,570,151,176]" pageId="27" pageNumber="28">
<emphasis id="9B8CEACC4C33B353FF6BFF43F7F6D5D4" bold="true" box="[151,163,151,176]" italics="true" pageId="27" pageNumber="28">*</emphasis>
<taxonomicName id="6EF84D5D4C33B353FF5EFF43F56FD5D4" ID-CoL="LCBJ" authority="Bernhauer 1907" authorityName="Bernhauer" authorityYear="1907" box="[162,570,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sharpi">
<emphasis id="9B8CEACC4C33B353FF5EFF43F62AD5D4" bold="true" box="[162,383,151,176]" italics="true" pageId="27" pageNumber="28">Belonuchus sharpi</emphasis>
<bibRefCitation id="CD694B2F4C33B353FE7AFF43F56FD5D4" box="[390,570,151,176]" pageId="27" pageNumber="28" refId="ref61805" refString="Bernhauer, M. (1907) Neue Staphyliniden aus Sudamerika. 4. Stuck. Wiener Entomologische Zeitung, 26 (7 - 9), 281 - 289. https: // doi. org / 10.5962 / bhl. part. 8892" type="journal article">Bernhauer 1907</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFF68F258D59C" blockId="27.[151,1436,151,357]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFF68F258D59C" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFF68F523D5B1" ID-CoL="LCBJ" authority="Bernhauer 1907: 285" authorityName="Bernhauer" authorityPageNumber="285" authorityYear="1907" box="[151,630,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sharpi">
<emphasis id="9B8CEACC4C33B353FF6BFF68F63BD5B1" box="[151,366,188,213]" italics="true" pageId="27" pageNumber="28">Belonuchus sharpi</emphasis>
<treatmentCitation id="285910CF4C33B353FE80FF68F523D5B1" author="" box="[380,630,188,213]" page="285" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE80FF68F56ED5B0" box="[380,571,188,213]" pageId="27" pageNumber="28" refId="ref61805" refString="Bernhauer, M. (1907) Neue Staphyliniden aus Sudamerika. 4. Stuck. Wiener Entomologische Zeitung, 26 (7 - 9), 281 - 289. https: // doi. org / 10.5962 / bhl. part. 8892" type="journal article">Bernhauer 1907</bibRefCitation>
: 285
</treatmentCitation>
</taxonomicName>
(replacement name for 
<taxonomicName id="6EF84D5D4C33B353FC61FF68F2C5D5B1" ID-CoL="LC87" authority="Sharp 1876 a: 158" authorityName="Sharp" authorityPageNumber="158" authorityYear="1876" box="[925,1424,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="9B8CEACC4C33B353FC61FF68F3FED5B1" box="[925,1195,188,213]" italics="true" pageId="27" pageNumber="28">Belonuchus grandiceps</emphasis>
<treatmentCitation id="285910CF4C33B353FB44FF68F2C5D5B1" author="" box="[1208,1424,188,213]" page="158" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FB44FF68F205D5B1" box="[1208,1360,188,213]" pageId="27" pageNumber="28" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 158
</treatmentCitation>
</taxonomicName>
). Distribution: Brazil (Amazonas, Pará), Peru. Note: distribution within Brazil (Asenjo 
<emphasis id="9B8CEACC4C33B353FB79FF35F3E8D59C" box="[1157,1213,223,248]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFED0F54FD400" blockId="27.[151,1436,151,357]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFED0F54FD400" infered="true" pageId="27" pageNumber="28">
<emphasis id="9B8CEACC4C33B353FF6BFED0F7F3D479" box="[151,166,260,285]" italics="true" pageId="27" pageNumber="28">=</emphasis>
<taxonomicName id="6EF84D5D4C33B353FF5AFED0F529D479" ID-CoL="LC87" authority="Sharp 1876 a: 158" authorityName="Sharp" authorityPageNumber="158" authorityYear="1876" box="[166,636,260,285]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="9B8CEACC4C33B353FF5AFED0F6E5D479" box="[166,432,260,285]" italics="true" pageId="27" pageNumber="28">Belonuchus grandiceps</emphasis>
<treatmentCitation id="285910CF4C33B353FE4BFED0F529D479" author="" box="[439,636,260,285]" page="158" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE4BFED0F51DD479" box="[439,584,260,285]" pageId="27" pageNumber="28" refId="ref72342" refString="Sharp, D. (1876 a) Contributions to an insect fauna of the Amazon Valley. Coleoptera-Staphylinidae. Transactions of the Entomological Society of London, 1876, 27 - 424. https: // doi. org / 10.5962 / bhl. title. 5536" type="journal article">Sharp 1876a</bibRefCitation>
: 158
</treatmentCitation>
</taxonomicName>
(preoccupied, junior secondary homonym in 
<taxonomicName id="6EF84D5D4C33B353FB86FED0F3A8D479" ID-CoL="8VRHV" box="[1146,1277,260,285]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C33B353FB86FED0F3A8D479" box="[1146,1277,260,285]" italics="true" pageId="27" pageNumber="28">Belonuchus</emphasis>
</taxonomicName>
of 
<taxonomicName id="6EF84D5D4C33B353FADFFED0F55ED424" ID-CoL="8Y89V" authority="Kraatz 1859 a: 95" authorityName="Kraatz" authorityPageNumber="95" authorityYear="1859" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="9B8CEACC4C33B353FADFFED0F617D424" italics="true" pageId="27" pageNumber="28">Philonthus grandiceps</emphasis>
<treatmentCitation id="285910CF4C33B353FEB4FEFCF55ED424" author="" box="[328,523,295,320]" page="95" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FEB4FEFCF6B4D424" box="[328,481,295,320]" pageId="27" pageNumber="28" refId="ref68268" refString="Kraatz, G. (1859 a) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan. Archiv fur Naturgeschichte, 25 (1), 1 - 193, pls. 1 - 3." type="journal article">Kraatz 1859a</bibRefCitation>
: 95
</treatmentCitation>
</taxonomicName>
). Type locality: Ega, Tapajos &amp; St. Paulo [Amazonas]. Type depository: BMNH 5 ST. Note: Peru (
<bibRefCitation id="CD694B2F4C33B353FE84FE98F55BD400" box="[376,526,332,357]" pageId="27" pageNumber="28" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFE47F513D4C8" blockId="27.[151,1435,403,501]" box="[151,582,403,429]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFE47F513D4C8" ID-CoL="LCBK" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,582,403,429]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="silvestris">
<emphasis id="9B8CEACC4C33B353FF6BFE47F6D9D4C8" bold="true" box="[151,396,403,428]" italics="true" pageId="27" pageNumber="28">Belonuchus silvestris</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFE63F4FDD491" blockId="27.[151,1435,403,501]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFE63F4FDD491" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BFE63F4F1D491" ID-GBIF-Occurrence="1915497869" collectionCode="FMNH" country="Colombia" elevation="2200" location="Distribution" pageId="27" pageNumber="28" specimenCount="2" stateProvince="Cauca" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BFE63F5D4D4B4" ID-CoL="LCBK" authority="Bernhauer 1917 b: 17" authorityName="Bernhauer" authorityPageNumber="17" authorityYear="1917" box="[151,641,439,464]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="silvestris">
<emphasis id="9B8CEACC4C33B353FF6BFE63F6D3D4B4" box="[151,390,439,464]" italics="true" pageId="27" pageNumber="28">Belonuchus silvestris</emphasis>
<treatmentCitation id="285910CF4C33B353FE72FE63F5D4D4B4" author="" box="[398,641,439,464]" page="17" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE72FE63F500D4B4" box="[398,597,439,464]" pageId="27" pageNumber="28" refId="ref62221" refString="Bernhauer, M. (1917 b) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (17. Beitrag - Fortsetzung). Entomologische Blatter, 13 (1 - 3), 17 - 24." type="journal article">Bernhauer 1917b</bibRefCitation>
: 17
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FD72FE6CF590D4B4" box="[654,709,440,464]" pageId="27" pageNumber="28">Type</typeStatus>
locality: Alto de las Cruces, 
<elevation id="22D5D1ED4C33B353FBE9FE6CF333D4B4" box="[1045,1126,440,464]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="27" pageNumber="28" unit="m" value="2200.0">
<quantity id="6E009B3B4C33B353FBE9FE6CF333D4B4" box="[1045,1126,440,464]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="27" pageNumber="28" unit="m" value="2200.0">2200m</quantity>
</elevation>
, Cali am Rio 
<collectingRegion id="6B3CF83C4C33B353FAFAFE6CF21AD4B4" box="[1286,1359,440,464]" country="Colombia" name="Cauca" pageId="27" pageNumber="28">Cauca</collectingRegion>
, West- Columbien. 
<typeStatus id="7643887C4C33B353FEB3FE09F6D3D491" box="[335,390,477,501]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C33B353FDEFFE09F531D490" box="[531,612,477,500]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="28">FMNH</collectionCode>
HT. 
<location id="AC2760054C33B353FD60FE08F47DD491" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FD60FE08F47DD491" box="[668,808,476,501]" country="Colombia" name="Distribution" pageId="27" pageNumber="28" stateProvince="Cauca">Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FCCEFE08F4F1D491" box="[818,932,476,501]" name="Colombia" pageId="27" pageNumber="28">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFDF7F6AFD759" blockId="27.[151,1437,547,645]" box="[151,506,547,573]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFDF7F6AFD759" ID-CoL="LCBL" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[151,506,547,573]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="9B8CEACC4C33B353FF6BFDF7F626D758" bold="true" box="[151,371,547,572]" italics="true" pageId="27" pageNumber="28">Belonuchus similis</emphasis>
<bibRefCitation id="CD694B2F4C33B353FE86FDF0F6AFD759" box="[378,506,548,573]" pageId="27" pageNumber="28" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFD93F330D7E1" blockId="27.[151,1437,547,645]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFD93F330D7E1" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BFD93F385D704" ID-GBIF-Occurrence="1915497857" box="[151,1232,583,608]" country="Panama" elevation="1295" location="Chiriqui" pageId="27" pageNumber="28" specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BFD93F562D704" ID-CoL="LCBL" authority="Sharp 1885: 436" authorityName="Sharp" authorityPageNumber="436" authorityYear="1885" box="[151,567,583,608]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="9B8CEACC4C33B353FF6BFD93F639D704" box="[151,364,583,608]" italics="true" pageId="27" pageNumber="28">Belonuchus similis</emphasis>
<treatmentCitation id="285910CF4C33B353FE89FD93F562D704" author="" box="[373,567,583,608]" page="436" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE89FD93F6A9D704" box="[373,508,583,608]" pageId="27" pageNumber="28" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 436
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FDB8FD9CF529D704" box="[580,636,584,608]" pageId="27" pageNumber="28">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C33B353FD16FD9CF410D73B" box="[746,837,584,607]" name="Panama" pageId="27" pageNumber="28">Panama</collectingCountry>
, Volcan de 
<collectingRegion id="6B3CF83C4C33B353FC32FD93F37ED704" box="[974,1067,583,608]" country="Panama" name="Chiriqui" pageId="27" pageNumber="28">Chiriqui</collectingRegion>
<elevation id="22D5D1ED4C33B353FBC8FD93F385D704" box="[1076,1232,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.2954" metricValueMax="1.8288" metricValueMin="0.762" pageId="27" pageNumber="28" unit="ft" value="4250.0" valueMax="6000.0" valueMin="2500.0">
<quantity id="6E009B3B4C33B353FBC8FD93F385D704" box="[1076,1232,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.2954" metricValueMax="1.8288" metricValueMin="0.762" pageId="27" pageNumber="28" unit="ft" value="4250.0" valueMax="6000.0" valueMin="2500.0">2500–6000 ft</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C33B353FB21FD9CF330D7E1" ID-GBIF-Occurrence="1915497826" collectionCode="BMNH, FMNH" country="Panama" location="ST. Distribution" pageId="27" pageNumber="28" specimenCode="BMNH 3, FMNH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C33B353FB21FD9CF240D704" box="[1245,1301,584,608]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C33B353FF39FDB9F67BD7E0" box="[197,302,620,644]" collectionCode="BMNH" pageId="27" pageNumber="28">BMNH 3</specimenCode>
ST; 
<specimenCode id="F95E9EA54C33B353FE9FFDB9F69CD7E0" box="[355,457,620,644]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="28">FMNH 1</specimenCode>
<location id="AC2760054C33B353FE2CFDB8F5DDD7E1" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FE2CFDB8F5DDD7E1" box="[464,648,620,645]" country="Panama" name="ST. Distribution" pageId="27" pageNumber="28">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FD6EFDB9F5BBD7E0" box="[658,750,621,644]" name="Panama" pageId="27" pageNumber="28">Panama</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFD67F50ED7A8" blockId="27.[151,1437,691,789]" box="[151,603,691,717]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFD67F50ED7A8" ID-CoL="LCBM" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,603,691,717]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="simillimus">
<emphasis id="9B8CEACC4C33B353FF6BFD67F6F5D7A8" bold="true" box="[151,416,691,716]" italics="true" pageId="27" pageNumber="28">Belonuchus simillimus</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFD03F50FD671" blockId="27.[151,1437,691,789]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFD03F50FD671" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BFD03F503D671" ID-GBIF-Occurrence="1915497895" collectionCode="FMNH" country="Colombia" elevation="1800" location="Distribution" pageId="27" pageNumber="28" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BFD03F5C0D794" ID-CoL="LCBM" authority="Bernhauer 1917 c: 106" authorityName="Bernhauer" authorityPageNumber="106" authorityYear="1917" box="[151,661,727,752]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="simillimus">
<emphasis id="9B8CEACC4C33B353FF6BFD03F6C6D794" box="[151,403,727,752]" italics="true" pageId="27" pageNumber="28">Belonuchus simillimus</emphasis>
<treatmentCitation id="285910CF4C33B353FE66FD03F5C0D794" author="" box="[410,661,727,752]" page="106" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE66FD03F50BD794" box="[410,606,727,752]" pageId="27" pageNumber="28" refId="ref62261" refString="Bernhauer, M. (1917 c) Neue sudamerikanische Staphyliniden (18. Beitrag). Wiener Entomologische Zeitung, 36 (3 - 5), 102 - 116. https: // doi. org / 10.5962 / bhl. part. 12925" type="journal article">Bernhauer 1917c</bibRefCitation>
: 106
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FD63FD0CF583D794" box="[671,726,728,752]" pageId="27" pageNumber="28">Type</typeStatus>
locality: Villa Elvira 
<elevation id="22D5D1ED4C33B353FC34FD0CF34DD794" box="[968,1048,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="1800.0">
<quantity id="6E009B3B4C33B353FC34FD0CF34DD794" box="[968,1048,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="1800.0">1800m</quantity>
</elevation>
, Westkolumbien. 
<typeStatus id="7643887C4C33B353FB1CFD0CF24DD794" box="[1248,1304,728,752]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C33B353FF39FD29F643D670" box="[197,278,765,788]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="28">FMNH</collectionCode>
HT. 
<location id="AC2760054C33B353FEB2FD28F68FD671" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FEB2FD28F68FD671" box="[334,474,764,789]" country="Colombia" name="Distribution" pageId="27" pageNumber="28">Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FE18FD28F503D671" box="[484,598,764,789]" name="Colombia" pageId="27" pageNumber="28">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFC97F51BD639" blockId="27.[151,1437,835,968]" box="[151,590,835,861]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFC97F51BD639" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[151,590,835,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="skalitskyi">
<emphasis id="9B8CEACC4C33B353FF6BFC97F6C6D638" bold="true" box="[151,403,835,860]" italics="true" pageId="27" pageNumber="28">Belonuchus skalitskyi</emphasis>
<bibRefCitation id="CD694B2F4C33B353FE66FC90F51BD639" box="[410,590,836,861]" pageId="27" pageNumber="28" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFCB3F470D6C0" blockId="27.[151,1437,835,968]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFCB3F470D6C0" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFCB3F5CBD6E4" authority="Bernhauer 1906: 334" authorityName="Bernhauer" authorityPageNumber="334" authorityYear="1906" box="[151,670,871,896]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="skalitskyi">
<emphasis id="9B8CEACC4C33B353FF6BFCB3F6C5D6E4" box="[151,400,871,896]" italics="true" pageId="27" pageNumber="28">Belonuchus skalitskyi</emphasis>
<treatmentCitation id="285910CF4C33B353FE63FCB3F5CBD6E4" author="" box="[415,670,871,896]" page="334" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE63FCB3F50BD6E4" box="[415,606,871,896]" pageId="27" pageNumber="28" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 334
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FD4CFCBCF5BDD6E4" box="[688,744,872,896]" pageId="27" pageNumber="28">Type</typeStatus>
locality: Pumananea, 
<collectingCountry id="D1EF764E4C33B353FC0BFCBCF37BD61B" box="[1015,1070,872,895]" name="Peru" pageId="27" pageNumber="28">Peru</collectingCountry>
. 
<typeStatus id="7643887C4C33B353FBBCFCBCF322D6E4" box="[1088,1143,872,896]" pageId="27" pageNumber="28">Type</typeStatus>
depository: FMNH HT. Distribution: 
<collectingCountry id="D1EF764E4C33B353FEA7FC59F6C4D6C0" box="[347,401,909,932]" name="Peru" pageId="27" pageNumber="28">Peru</collectingCountry>
, 
<collectingCountry id="D1EF764E4C33B353FE60FC58F6A7D6C1" box="[412,498,908,933]" name="Bolivia" pageId="27" pageNumber="28">Bolivia</collectingCountry>
. Note: 
<collectingCountry id="D1EF764E4C33B353FDBEFC59F520D6C0" box="[578,629,909,932]" name="Peru" pageId="27" pageNumber="28">Peru</collectingCountry>
(
<bibRefCitation id="CD694B2F4C33B353FD78FC58F44CD6C0" box="[644,793,908,933]" pageId="27" pageNumber="28" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C33B353FF6BFC7BF465D6AC" box="[151,816,943,968]" pageId="27" pageNumber="28" type="discussion">
<paragraph id="A94736DE4C33B353FF6BFC7BF465D6AC" blockId="27.[151,1437,835,968]" box="[151,816,943,968]" pageId="27" pageNumber="28">
<emphasis id="9B8CEACC4C33B353FF6BFC7BF641D6AC" bold="true" box="[151,276,943,968]" pageId="27" pageNumber="28">Comment.</emphasis>
Not 
<taxonomicName id="6EF84D5D4C33B353FEB0FC7BF69AD6AC" box="[332,463,943,968]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C33B353FEB0FC7BF69AD6AC" box="[332,463,943,968]" italics="true" pageId="27" pageNumber="28">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C33B354FF6BFC23F452D4EC" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="A94736DE4C33B353FF6BFC23F520D174" blockId="27.[151,1436,1015,1112]" box="[151,629,1015,1040]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFC23F520D174" ID-CoL="LCBR" authority="(Champion 1913)" authorityName="Champion" authorityYear="1913" baseAuthorityName="Champion" baseAuthorityYear="1913" box="[151,629,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="spinicornis">
<emphasis id="9B8CEACC4C33B353FF6BFC23F6F2D174" bold="true" box="[151,423,1015,1040]" italics="true" pageId="27" pageNumber="28">Belonuchus spinicornis</emphasis>
(
<bibRefCitation id="CD694B2F4C33B353FE4AFC23F538D174" box="[438,621,1015,1040]" pageId="27" pageNumber="28" refId="ref63989" refString="Champion, G. C. (1913) Coleoptera, &amp; c., in bromeliads. Entomologist's Monthly Magazine, 49, 2 - 7." type="journal article">Champion 1913</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFBC8F448D13C" blockId="27.[151,1436,1015,1112]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFBC8F448D13C" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BFBC8F32FD150" ID-GBIF-Occurrence="1915497860" box="[151,1146,1052,1077]" country="Costa Rica" elevation="1400" location="Pitahaya" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BFBC8F5D3D150" ID-CoL="6RR37" authority="Champion 1913: 4" authorityName="Champion" authorityPageNumber="4" authorityYear="1913" box="[151,646,1052,1077]" class="Insecta" family="Staphylinidae" genus="Musicoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="spinicornis">
<emphasis id="9B8CEACC4C33B353FF6BFBC8F6FFD150" box="[151,426,1052,1077]" italics="true" pageId="27" pageNumber="28">Musicoderus spinicornis</emphasis>
<treatmentCitation id="285910CF4C33B353FE4EFBC8F5D3D150" author="" box="[434,646,1052,1077]" page="4" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE4EFBC8F53FD150" box="[434,618,1052,1077]" pageId="27" pageNumber="28" refId="ref63989" refString="Champion, G. C. (1913) Coleoptera, &amp; c., in bromeliads. Entomologist's Monthly Magazine, 49, 2 - 7." type="journal article">Champion 1913</bibRefCitation>
: 4
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FD6CFBC9F59DD151" box="[656,712,1053,1077]" pageId="27" pageNumber="28">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C33B353FCCEFBC8F4FAD151" box="[818,943,1052,1077]" name="Costa Rica" pageId="27" pageNumber="28">Costa Rica</collectingCountry>
, 
<location id="AC2760054C33B353FC46FBC8F348D151" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FC46FBC8F348D151" box="[954,1053,1052,1077]" country="Costa Rica" name="Pitahaya" pageId="27" pageNumber="28">Pitahaya</location>
, 
<elevation id="22D5D1ED4C33B353FBD5FBC8F32FD150" box="[1065,1146,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="28" unit="m" value="1400.0">
<quantity id="6E009B3B4C33B353FBD5FBC8F32FD150" box="[1065,1146,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="27" pageNumber="28" unit="m" value="1400.0">1400m</quantity>
</elevation>
</materialsCitation>
. 
<materialsCitation id="19903C834C33B353FB78FBC9F44CD13C" ID-GBIF-Occurrence="1915497837" collectionCode="MCP" country="Costa Rica" location="Distribution" pageId="27" pageNumber="28" specimenCode="BMNH 2" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C33B353FB78FBC9F3E9D151" box="[1156,1212,1053,1077]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C33B353FAB4FBC9F786D13C" collectionCode="BMNH" pageId="27" pageNumber="28">BMNH 2</specimenCode>
ST: 
<specimenCount id="BFFEFD574C33B353FEF4FB94F661D133" box="[264,308,1088,1112]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
, 
<specimenCount id="BFFEFD574C33B353FEBCFB94F63FD133" box="[320,362,1088,1112]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
[
<collectionCode id="CFE9AE1B4C33B353FE8BFB94F6E0D13C" box="[375,437,1088,1112]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="27" pageNumber="28">MCP</collectionCode>
seen]. 
<location id="AC2760054C33B353FDF9FBEBF5C4D13C" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FDF9FBEBF5C4D13C" box="[517,657,1087,1112]" country="Costa Rica" name="Distribution" pageId="27" pageNumber="28">Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FD60FB94F44CD13C" box="[668,793,1087,1112]" name="Costa Rica" pageId="27" pageNumber="28">Costa Rica</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFB53F56FD1C4" blockId="27.[151,1436,1159,1293]" box="[151,570,1159,1184]" pageId="27" pageNumber="28">
<emphasis id="9B8CEACC4C33B353FF6BFB53F7F6D1C4" bold="true" box="[151,163,1159,1184]" italics="true" pageId="27" pageNumber="28">*</emphasis>
<taxonomicName id="6EF84D5D4C33B353FF5EFB53F56FD1C4" ID-CoL="LCBS" authority="Sharp 1885" authorityName="Sharp" authorityYear="1885" box="[162,570,1159,1184]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="stenoderus">
<emphasis id="9B8CEACC4C33B353FF5EFB53F6E7D1C4" bold="true" box="[162,434,1159,1184]" italics="true" pageId="27" pageNumber="28">Belonuchus stenoderus</emphasis>
<bibRefCitation id="CD694B2F4C33B353FE46FB53F56FD1C4" box="[442,570,1159,1184]" pageId="27" pageNumber="28" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFB78F5A0D068" blockId="27.[151,1436,1159,1293]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFB78F5A0D068" infered="true" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFB78F50BD1A0" ID-CoL="LCBS" authority="Sharp 1885: 434" authorityName="Sharp" authorityPageNumber="434" authorityYear="1885" box="[151,606,1196,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="stenoderus">
<emphasis id="9B8CEACC4C33B353FF6BFB78F6CFD1A1" box="[151,410,1196,1221]" italics="true" pageId="27" pageNumber="28">Belonuchus stenoderus</emphasis>
<treatmentCitation id="285910CF4C33B353FE5DFB78F50BD1A0" author="" box="[417,606,1196,1221]" page="434" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE5DFB78F570D1A1" box="[417,549,1196,1221]" pageId="27" pageNumber="28" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 434
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FD94FB79F5CAD1A1" box="[616,671,1197,1221]" pageId="27" pageNumber="28">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C33B353FCF6FB79F433D1A0" box="[778,870,1197,1220]" country="Panama" name="Panama" pageId="27" pageNumber="28">
<collectingCountry id="D1EF764E4C33B353FCF6FB79F433D1A0" box="[778,870,1197,1220]" name="Panama" pageId="27" pageNumber="28">Panama</collectingCountry>
</collectingRegion>
, Bugaba. 
<typeStatus id="7643887C4C33B353FC28FB79F35ED1A1" box="[980,1035,1197,1221]" pageId="27" pageNumber="28">Type</typeStatus>
depository: BMNH 4 ST; FMNH 1 ST. Distribution: 
<collectingCountry id="D1EF764E4C33B353FE68FB04F543D18C" box="[404,534,1231,1256]" name="Costa Rica" pageId="27" pageNumber="28">
<collectingRegion id="6B3CF83C4C33B353FE68FB04F681D18C" box="[404,468,1232,1256]" country="Tanzania" name="Pwani" pageId="27" pageNumber="28">Costa</collectingRegion>
Rica
</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C33B353FDD4FB04F5D1D183" box="[552,644,1232,1255]" country="Panama" name="Panama" pageId="27" pageNumber="28">
<collectingCountry id="D1EF764E4C33B353FDD4FB04F5D1D183" box="[552,644,1232,1255]" name="Panama" pageId="27" pageNumber="28">Panama</collectingCountry>
</collectingRegion>
. Note: the 
<typeStatus id="7643887C4C33B353FCF3FB04F433D18C" box="[783,870,1232,1256]" pageId="27" pageNumber="28" type="syntype">syntype</typeStatus>
in FMNH does not belong to the ‘ 
<taxonomicName id="6EF84D5D4C33B353FAE5FB1BF62AD069" ID-CoL="LC89" authorityName="" baseAuthorityName="Fabricius" baseAuthorityYear="1801" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
<emphasis id="9B8CEACC4C33B353FAE5FB1BF62AD069" italics="true" pageId="27" pageNumber="28">Belonuchus haemorrhoidalis</emphasis>
</taxonomicName>
clade’ (
<bibRefCitation id="CD694B2F4C33B353FE25FB20F5BCD068" box="[473,745,1268,1293]" pageId="27" pageNumber="28" refId="ref64541" refString="Chani-Posse, M. R., Brunke, A. J., Chatzimanolis, S., Schillhammer, H. &amp; Solodovnikov, A. (2018) Phylogeny of the hyperdiverse rove beetle subtribe Philonthina with implications for classification of the tribe Staphylinini (Coleoptera: Staphylinidae). Cladistics, 34 (1), 1 - 40. https: // doi. org / 10.1111 / cla. 12188" type="journal article">
Chani-Posse 
<emphasis id="9B8CEACC4C33B353FD91FB21F5F0D069" box="[621,677,1268,1293]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2018
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFAEFF5C3D031" blockId="27.[151,1437,1339,1472]" box="[151,662,1339,1365]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFAEFF5C3D031" ID-CoL="LCBT" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,662,1339,1365]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subaeneipennis">
<emphasis id="9B8CEACC4C33B353FF6BFAEFF68ED030" bold="true" box="[151,475,1339,1364]" italics="true" pageId="27" pageNumber="28">Belonuchus subaeneipennis</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFA8BF66ED0A4" blockId="27.[151,1437,1339,1472]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BFA8BF66ED0A4" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BFA8BF662D0A4" ID-GBIF-Occurrence="1915497829" collectionCode="FMNH" collectorName="West-Columbien &amp; Umgebung von Cali &amp; Canon" country="Colombia" elevation="1700" location="ST. Distribution" pageId="27" pageNumber="28" specimenCode="FMNH 1" specimenCount="2" stateProvince="Cauca" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BFA8BF42DD01C" ID-CoL="LCBT" authority="Bernhauer 1916 c: 269" authorityName="Bernhauer" authorityPageNumber="269" authorityYear="1916" box="[151,888,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subaeneipennis" subGenus="Musicoderus">
<emphasis id="9B8CEACC4C33B353FF6BFA8BF522D01C" box="[151,631,1375,1400]" italics="true" pageId="27" pageNumber="28">Belonuchus (Musicoderus) subaeneipennis</emphasis>
<treatmentCitation id="285910CF4C33B353FD81FA8BF42DD01C" author="" box="[637,888,1375,1400]" page="269" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FD81FA8BF414D01C" box="[637,833,1375,1400]" pageId="27" pageNumber="28" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 269
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FC7FFAB4F4EFD01C" box="[899,954,1376,1400]" pageId="27" pageNumber="28">Type</typeStatus>
locality: 
<collectorName id="040D53084C33B353FBD8FA8BF3B3D01C" box="[1060,1254,1375,1400]" pageId="27" pageNumber="28">West-Columbien</collectorName>
: 
<collectorName id="040D53084C33B353FB13FA8BF7A1D0F9" pageId="27" pageNumber="28">Umgebung von Cali</collectorName>
am Rio 
<collectingRegion id="6B3CF83C4C33B353FE9FFA50F6FCD0F8" box="[355,425,1412,1436]" country="Colombia" name="Cauca" pageId="27" pageNumber="28">Cauca</collectingRegion>
(
<collectorName id="040D53084C33B353FE41FA50F55DD0F8" box="[445,520,1412,1436]" pageId="27" pageNumber="28">Cañon</collectorName>
del Mte 
<collectingRegion id="6B3CF83C4C33B353FD80FA50F59BD0F9" box="[636,718,1412,1437]" country="Colombia" name="Tolima" pageId="27" pageNumber="28">Tolima</collectingRegion>
, 
<elevation id="22D5D1ED4C33B353FD22FA50F464D0F8" box="[734,817,1412,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="27" pageNumber="28" unit="m" value="1700.0">
<quantity id="6E009B3B4C33B353FD22FA50F464D0F8" box="[734,817,1412,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="27" pageNumber="28" unit="m" value="1700.0">1700m</quantity>
</elevation>
). 
<typeStatus id="7643887C4C33B353FCB4FA51F4D5D0F9" box="[840,896,1413,1437]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C33B353FBEAFA51F328D0F8" box="[1046,1149,1412,1436]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="28">FMNH 1</specimenCode>
ST;? 1 
<location id="AC2760054C33B353FB2BFA50F2CCD0F9" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FB2BFA50F2CCD0F9" box="[1239,1433,1412,1437]" country="Colombia" name="ST. Distribution" pageId="27" pageNumber="28" stateProvince="Cauca">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FF39FA73F662D0A4" box="[197,311,1447,1472]" name="Colombia" pageId="27" pageNumber="28">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BFA3BF509D36C" blockId="27.[151,1436,1519,1616]" box="[151,604,1519,1544]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BFA3BF509D36C" ID-CoL="68GJ5" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,604,1519,1544]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="9B8CEACC4C33B353FF6BFA3BF6F4D36C" bold="true" box="[151,417,1519,1544]" italics="true" pageId="27" pageNumber="28">Belonuchus subaeneus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BF9C0F4D5D334" blockId="27.[151,1436,1519,1616]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BF9C0F4D5D334" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BF9C0F20FD349" ID-GBIF-Occurrence="1915497816" box="[151,1370,1556,1581]" country="Bolivia" location="Yuracares" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BF9C0F5C7D349" ID-CoL="68GJ5" authority="Bernhauer 1916 c: 268" authorityName="Bernhauer" authorityPageNumber="268" authorityYear="1916" box="[151,658,1556,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subaeneus">
<emphasis id="9B8CEACC4C33B353FF6BF9C0F6C0D349" box="[151,405,1556,1581]" italics="true" pageId="27" pageNumber="28">Belonuchus subaeneus</emphasis>
<treatmentCitation id="285910CF4C33B353FE67F9C0F5C7D349" author="" box="[411,658,1556,1581]" page="268" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE67F9C0F50AD349" box="[411,607,1556,1581]" pageId="27" pageNumber="28" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 268
</treatmentCitation>
</taxonomicName>
(attributed to Fauvel 
<emphasis id="9B8CEACC4C33B353FC7EF9C1F491D349" box="[898,964,1556,1581]" italics="true" pageId="27" pageNumber="28">in litt.</emphasis>
). 
<typeStatus id="7643887C4C33B353FC27F9C1F346D349" box="[987,1043,1557,1581]" pageId="27" pageNumber="28">Type</typeStatus>
locality: 
<location id="AC2760054C33B353FB87F9C0F3B9D348" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FB87F9C0F3B9D348" box="[1147,1260,1556,1580]" country="Bolivia" name="Yuracares" pageId="27" pageNumber="28">Yuracarès</location>
, 
<collectingCountry id="D1EF764E4C33B353FB0AF9C0F20FD349" box="[1270,1370,1556,1581]" name="Bolivia" pageId="27" pageNumber="28">Bolivien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C33B353FA98F9C1F429D334" ID-GBIF-Occurrence="1915497802" collectionCode="FMNH, ZMUH" country="Bolivia" location="Distribution" pageId="27" pageNumber="28" specimenCode="FMNH 1, ZMUH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C33B353FA98F9C1F2C9D349" box="[1380,1436,1557,1581]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C33B353FEB6F9ECF6F9D334" box="[330,428,1592,1616]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="28">FMNH 1</specimenCode>
ST; 
<specimenCode id="F95E9EA54C33B353FE22F9ECF514D334" box="[478,577,1592,1616]" collectionCode="ZMUH" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="27" pageNumber="28">ZMUH 1</specimenCode>
+ST?. 
<location id="AC2760054C33B353FD6CF9E3F449D334" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FD6CF9E3F449D334" box="[656,796,1591,1616]" country="Bolivia" name="Distribution" pageId="27" pageNumber="28">Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FCDAF9E3F429D334" box="[806,892,1591,1616]" name="Bolivia" pageId="27" pageNumber="28">Bolivia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BF9ABF536D3FC" blockId="27.[151,1437,1663,1760]" box="[151,611,1663,1688]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BF9ABF536D3FC" ID-CoL="LCBX" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,611,1663,1688]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subsellatus">
<emphasis id="9B8CEACC4C33B353FF6BF9ABF6F2D3FC" bold="true" box="[151,423,1663,1688]" italics="true" pageId="27" pageNumber="28">Belonuchus subsellatus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BF970F277D384" blockId="27.[151,1437,1663,1760]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BF970F277D384" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BF970F2CCD3D9" ID-GBIF-Occurrence="1915497839" box="[151,1433,1700,1725]" country="Bolivia" location="Yuracares" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BF970F5FED3D9" ID-CoL="LCBX" authority="Bernhauer 1916 c: 276" authorityName="Bernhauer" authorityPageNumber="276" authorityYear="1916" box="[151,683,1700,1725]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subsellatus">
<emphasis id="9B8CEACC4C33B353FF6BF970F6F4D3D9" box="[151,417,1700,1725]" italics="true" pageId="27" pageNumber="28">Belonuchus subsellatus</emphasis>
<treatmentCitation id="285910CF4C33B353FE57F970F5FED3D9" author="" box="[427,683,1700,1725]" page="276" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE57F970F526D3D9" box="[427,627,1700,1725]" pageId="27" pageNumber="28" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 276
</treatmentCitation>
</taxonomicName>
(attributed to Fauvel 
<emphasis id="9B8CEACC4C33B353FC57F971F4A7D3D9" box="[939,1010,1700,1725]" italics="true" pageId="27" pageNumber="28">in litt.</emphasis>
). 
<typeStatus id="7643887C4C33B353FBF0F971F311D3D9" box="[1036,1092,1701,1725]" pageId="27" pageNumber="28">Type</typeStatus>
locality: 
<location id="AC2760054C33B353FB49F970F273D3D8" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FB49F970F273D3D8" box="[1205,1318,1700,1724]" country="Bolivia" name="Yuracares" pageId="27" pageNumber="28">Yuracarès</location>
, 
<collectingCountry id="D1EF764E4C33B353FAC9F970F2CCD3D9" box="[1333,1433,1700,1725]" name="Bolivia" pageId="27" pageNumber="28">Bolivien</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C33B353FF39F91CF277D384" ID-GBIF-Occurrence="1915497801" box="[197,1314,1735,1760]" collectionCode="FMNH, ZMUH" country="Bolivia" location="ST. Distribution" pageId="27" pageNumber="28" specimenCode="FMNH 1, ZMUH 1" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C33B353FF39F91CF7A8D384" box="[197,253,1736,1760]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C33B353FE74F91CF6BFD384" box="[392,490,1736,1760]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="28">FMNH 1</specimenCode>
ST, 
<specimenCode id="F95E9EA54C33B353FDE7F91CF52BD384" box="[539,638,1736,1760]" collectionCode="ZMUH" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="27" pageNumber="28">ZMUH 1</specimenCode>
+ 
<location id="AC2760054C33B353FD73F91CF41ED384" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FD73F91CF41ED384" box="[655,843,1735,1760]" country="Bolivia" name="ST. Distribution" pageId="27" pageNumber="28">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FCA9F913F4FED384" box="[853,939,1735,1760]" name="Bolivia" pageId="27" pageNumber="28">Bolivia</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BF8DBF51ED24C" blockId="27.[151,1437,1807,1941]" box="[151,587,1807,1832]" pageId="27" pageNumber="28">
<taxonomicName id="6EF84D5D4C33B353FF6BF8DBF51ED24C" ID-CoL="LCBY" authority="Bernhauer 1916" authorityName="Bernhauer" authorityYear="1916" box="[151,587,1807,1832]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="superbus">
<emphasis id="9B8CEACC4C33B353FF6BF8DBF6C5D24C" bold="true" box="[151,400,1807,1832]" italics="true" pageId="27" pageNumber="28">Belonuchus superbus</emphasis>
Bernhauer 1916
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C33B353FF6BF8E0F5F8D2F1" blockId="27.[151,1437,1807,1941]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="89E811F04C33B353FF6BF8E0F5F8D2F1" infered="true" pageId="27" pageNumber="28">
<materialsCitation id="19903C834C33B353FF6BF8E0F32AD228" ID-GBIF-Occurrence="1915497823" box="[151,1151,1844,1869]" elevation="2200" location="Alto" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C33B353FF6BF8E0F5DAD228" ID-CoL="LCBY" authority="Bernhauer 1916 c: 273" authorityName="Bernhauer" authorityPageNumber="273" authorityYear="1916" box="[151,655,1844,1869]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="superbus">
<emphasis id="9B8CEACC4C33B353FF6BF8E0F6D2D229" box="[151,391,1844,1869]" italics="true" pageId="27" pageNumber="28">Belonuchus superbus</emphasis>
<treatmentCitation id="285910CF4C33B353FE6CF8E0F5DAD228" author="" box="[400,655,1844,1869]" page="273" pageId="27" pageNumber="28">
<bibRefCitation id="CD694B2F4C33B353FE6CF8E0F503D229" box="[400,598,1844,1869]" pageId="27" pageNumber="28" refId="ref62149" refString="Bernhauer, M. (1916 c) Neue Staphyliniden aus den columbischen Cordilleren und dem ubrigen Sudamerika (16. Beitrag). Entomologische Blatter, 12 (10 - 12), 263 - 279." type="journal article">Bernhauer 1916c</bibRefCitation>
: 273
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C33B353FD60F8E1F586D229" box="[668,723,1845,1869]" pageId="27" pageNumber="28">Type</typeStatus>
localities: 
<location id="AC2760054C33B353FCADF8E0F4D6D229" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FCADF8E0F4D6D229" box="[849,899,1844,1869]" name="Alto" pageId="27" pageNumber="28">Alto</location>
de las cruces, 
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<quantity id="6E009B3B4C33B353FBD2F8E0F32AD228" box="[1070,1151,1844,1868]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="27" pageNumber="28" unit="m" value="2200.0">2200m</quantity>
</elevation>
</materialsCitation>
; 
<materialsCitation id="19903C834C33B353FB70F8E0F232D228" ID-GBIF-Occurrence="1915497878" box="[1164,1383,1844,1869]" elevation="2000" location="Rio Vitaco" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<location id="AC2760054C33B353FB70F8E0F25CD229" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FB70F8E0F25CD229" box="[1164,1289,1844,1869]" name="Rio Vitaco" pageId="27" pageNumber="28">Rio Vitaco</location>
, 
<elevation id="22D5D1ED4C33B353FAE9F8E0F232D228" box="[1301,1383,1844,1868]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="m" value="2000.0">
<quantity id="6E009B3B4C33B353FAE9F8E0F232D228" box="[1301,1383,1844,1868]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="m" value="2000.0">2000m</quantity>
</elevation>
</materialsCitation>
; 
<materialsCitation id="19903C834C33B353FA88F8E0F62BD214" ID-GBIF-Occurrence="1915497813" elevation="1800" location="Rio Aquatal" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
<location id="AC2760054C33B353FA88F8E0F674D214" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FA88F8E0F674D214" name="Rio Aquatal" pageId="27" pageNumber="28">Rio Aquatal</location>
, 
<elevation id="22D5D1ED4C33B353FED1F88CF62BD214" box="[301,382,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="1800.0">
<quantity id="6E009B3B4C33B353FED1F88CF62BD214" box="[301,382,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="1800.0">1800m</quantity>
</elevation>
</materialsCitation>
; S. Antonio, 
<quantity id="6E009B3B4C33B353FDE8F88CF533D214" box="[532,614,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="m" value="2000.0">2000m</quantity>
; 
<materialsCitation id="19903C834C33B353FD8EF883F5F8D2F1" ID-GBIF-Occurrence="1915497875" collectionCode="FMNH" country="Colombia" elevation="1800" location="ST. Distribution" pageId="27" pageNumber="28" specimenCode="FMNH 3" specimenCount="1" typeStatus="holotype">
Villa Elvira, 
<elevation id="22D5D1ED4C33B353FCF8F88CF403D214" box="[772,854,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="1800.0">
<quantity id="6E009B3B4C33B353FCF8F88CF403D214" box="[772,854,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="27" pageNumber="28" unit="m" value="1800.0">1800m</quantity>
</elevation>
. 
<typeStatus id="7643887C4C33B353FC9DF88CF4CDD214" box="[865,920,1880,1904]" pageId="27" pageNumber="28">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C33B353FBDBF88CF3D8D214" box="[1063,1165,1880,1904]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="28">FMNH 3</specimenCode>
ST;? 2 
<location id="AC2760054C33B353FB23F88CF2CCD214" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C33B353FB23F88CF2CCD214" box="[1247,1433,1879,1904]" country="Colombia" name="ST. Distribution" pageId="27" pageNumber="28">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C33B353FF39F8A8F662D2F1" box="[197,311,1916,1941]" name="Colombia" pageId="27" pageNumber="28">Colombia</collectingCountry>
. Note: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFF43F42AD5D4" blockId="28.[151,1437,151,717]" box="[151,895,151,176]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BFF43F7F6D5D4" bold="true" box="[151,163,151,176]" italics="true" pageId="28" pageNumber="29">*</emphasis>
<taxonomicName id="6EF84D5D4C34B354FF5EFF43F500D5D4" ID-CoL="LCBZ" authority="(Herman 2001)" authorityName="Chani-Posse &amp; Newton &amp; Hansen &amp; Solodovnikov" authorityYear="2018" baseAuthorityName="Herman" baseAuthorityYear="2001" box="[162,597,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="supernus" status="comb. nov.">
<emphasis id="9B8CEACC4C34B354FF5EFF43F6E5D5D4" bold="true" box="[162,432,151,176]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF5EFF43F6F5D5CB" bold="true" box="[162,416,151,176]" italics="true" pageId="28" pageNumber="29">Belonuchus supernus</emphasis>
(
</emphasis>
Herman 2001)
</taxonomicName>
, 
<taxonomicNameLabel id="80BF57B74C34B354FD9DFF43F5B4D5D4" box="[609,737,151,176]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FD9DFF43F5B4D5D4" bold="true" box="[609,737,151,176]" pageId="28" pageNumber="29">comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="31C32A5B4C34B354FD0CFF43F422D5D4" box="[752,887,151,176]" captionStart="FIGURES 45–52" captionStartId="82.[151,264,1313,1336]" captionTargetBox="[234,1352,195,1292]" captionTargetId="figure@82.[234,1352,195,1292]" captionTargetPageId="82" captionText="FIGURES 45–52 [FMNH ©]. Type and non-type material (habitus and labels): 45, 46, Belonuchus penetrans Silvestri, nontype; 47, 48, Philonthus excelsus Bernhauer, syntype; 49, 50 Philonthus peruvianus Bernhauer, syntype; 51, 52, Philonthus subaeneipennis Bernhauer, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454315/files/figure.png" pageId="28" pageNumber="29">Figs. 47–48</figureCitation>
)
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFF68F655D593" blockId="28.[151,1437,151,717]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BFF68F655D593" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BFF68F508D5B0" ID-CoL="4GBWZ" authority="Herman 2001 a: 48" authorityName="Herman" authorityPageNumber="48" authorityYear="2001" box="[151,605,188,213]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="supernus">
<emphasis id="9B8CEACC4C34B354FF6BFF68F62AD5B0" box="[151,383,188,213]" italics="true" pageId="28" pageNumber="29">Philonthus supernus</emphasis>
<treatmentCitation id="285910CF4C34B354FE74FF69F508D5B0" author="" box="[392,605,188,212]" page="48" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE74FF69F560D5B0" box="[392,565,188,212]" pageId="28" pageNumber="29" refId="ref67257" refString="Herman, L. H. (2001 a) Nomenclatural changes in the Staphylinidae (Insecta: Coleoptera). Bulletin of the American Museum of Natural History, 264, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 (2001) 264 % 3 C 0003: NCITSI % 3 E 2.0. CO; 2" type="journal article">Herman 2001a</bibRefCitation>
: 48
</treatmentCitation>
</taxonomicName>
(new name for 
<taxonomicName id="6EF84D5D4C34B354FCE4FF68F3A2D5B0" ID-CoL="6V9NZ" authority="Bernhauer 1941: 287" authorityName="Bernhauer" authorityPageNumber="287" authorityYear="1941" box="[792,1271,188,213]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="excelsus">
<emphasis id="9B8CEACC4C34B354FCE4FF68F4ADD5B1" box="[792,1016,188,213]" italics="true" pageId="28" pageNumber="29">Philonthus excelsus</emphasis>
<treatmentCitation id="285910CF4C34B354FBFEFF68F3A2D5B0" author="" box="[1026,1271,188,213]" page="287" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FBFEFF68F3EED5B0" box="[1026,1211,188,213]" pageId="28" pageNumber="29" refId="ref62815" refString="Bernhauer, M. (1941) Staphylinidae (Col.). In: Titschack, E. (Ed.), Beitrage zur Fauna Perus. Vol. 1. Conrad Behre, Hamburg, pp. 277 - 293. [most copies destroyed; reprinted in 1951 with different pagination by G. Fischer, Jena]" type="journal article">Bernhauer 1941</bibRefCitation>
: 287
</treatmentCitation>
</taxonomicName>
). Distribution: 
<collectingCountry id="D1EF764E4C34B354FF39FF34F7A9D593" box="[197,252,224,247]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFED1F452D4EC" blockId="28.[151,1437,151,717]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BFED1F452D4EC" infered="true" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BFED1F7FFD478" box="[151,170,261,284]" italics="true" pageId="28" pageNumber="29">=</emphasis>
<taxonomicName id="6EF84D5D4C34B354FF45FED0F5E5D478" ID-CoL="6V9NZ" authority="Bernhauer 1941: 287" authorityName="Bernhauer" authorityPageNumber="287" authorityYear="1941" box="[185,688,260,285]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="excelsus">
<emphasis id="9B8CEACC4C34B354FF45FED0F6CAD479" box="[185,415,260,285]" italics="true" pageId="28" pageNumber="29">Philonthus excelsus</emphasis>
<treatmentCitation id="285910CF4C34B354FE53FED0F5E5D478" author="" box="[431,688,260,285]" page="287" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE53FED0F53AD478" box="[431,623,260,285]" pageId="28" pageNumber="29" refId="ref62815" refString="Bernhauer, M. (1941) Staphylinidae (Col.). In: Titschack, E. (Ed.), Beitrage zur Fauna Perus. Vol. 1. Conrad Behre, Hamburg, pp. 277 - 293. [most copies destroyed; reprinted in 1951 with different pagination by G. Fischer, Jena]" type="journal article">Bernhauer 1941</bibRefCitation>
: 287
</treatmentCitation>
</taxonomicName>
, also 1951: 277 (preoccupied, junior secondary homonym in 
<taxonomicName id="6EF84D5D4C34B354FF39FEF3F66BD424" ID-CoL="8VRYR" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[197,318,295,320]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C34B354FF39FEF3F66BD424" box="[197,318,295,320]" italics="true" pageId="28" pageNumber="29">Philonthus</emphasis>
</taxonomicName>
of 
<taxonomicName id="6EF84D5D4C34B354FE96FEFDF479D424" ID-CoL="6M37X" authority="Cameron 1931: 361" authorityName="Cameron" authorityPageNumber="361" authorityYear="1931" box="[362,812,295,320]" class="Insecta" family="Staphylinidae" genus="Hesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="excelsus">
<emphasis id="9B8CEACC4C34B354FE96FEFDF56ED424" box="[362,571,295,320]" italics="true" pageId="28" pageNumber="29">Hesperus excelsus</emphasis>
<treatmentCitation id="285910CF4C34B354FDB9FEFCF479D424" author="" box="[581,812,295,320]" page="361" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FDB9FEFCF5A7D424" box="[581,754,296,320]" pageId="28" pageNumber="29" refId="ref63849" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article">Cameron 1931</bibRefCitation>
: 361
</treatmentCitation>
</taxonomicName>
, and junior primary homonym of 
<taxonomicName id="6EF84D5D4C34B354FB47FEF3F6F2D400" ID-CoL="6V9NZ" authority="Cameron 1932: 147" authorityName="Cameron" authorityPageNumber="147" authorityYear="1932" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="excelsus">
<emphasis id="9B8CEACC4C34B354FB47FEF3F2CED424" box="[1211,1435,295,320]" italics="true" pageId="28" pageNumber="29">Philonthus excelsus</emphasis>
<treatmentCitation id="285910CF4C34B354FF39FE98F6F2D400" author="" box="[197,423,332,356]" page="147" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FF39FE98F63BD400" box="[197,366,332,356]" pageId="28" pageNumber="29" refId="ref63886" refString="Cameron, M. (1932) Coleoptera, Staphylinidae. Vol. III. In: Stephenson, J. (Ed.), Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, pp. i - xiii + 1 - 443, pls. 1 - 2." type="book chapter">Cameron 1932</bibRefCitation>
: 147
</treatmentCitation>
</taxonomicName>
). 
<typeStatus id="7643887C4C34B354FE46FE99F6A4D401" box="[442,497,333,357]" pageId="28" pageNumber="29">Type</typeStatus>
locality: Ocaña, 
<quantity id="6E009B3B4C34B354FD4CFE98F454D400" box="[688,769,332,356]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="28" pageNumber="29" unit="m" value="3100.0">3100m</quantity>
, etc., s 
<collectingCountry id="D1EF764E4C34B354FCAFFE99F4DCD400" box="[851,905,333,356]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
, Oberh. Paleo 
<quantity id="6E009B3B4C34B354FBD1FE98F32BD400" box="[1069,1150,332,356]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="28" pageNumber="29" unit="m" value="3240.0">3240m</quantity>
, S 
<collectingCountry id="D1EF764E4C34B354FB63FE99F383D400" box="[1183,1238,333,356]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
. 
<typeStatus id="7643887C4C34B354FB1CFE99F242D401" box="[1248,1303,333,357]" pageId="28" pageNumber="29">Type</typeStatus>
depository: FMNH 21 ST: 1 Ocaña. Note: 
<collectingCountry id="D1EF764E4C34B354FDD9FEA4F50DD4E3" box="[549,600,368,391]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
(
<bibRefCitation id="CD694B2F4C34B354FD9AFEBBF5A9D4EC" box="[614,764,367,392]" pageId="28" pageNumber="29" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C34B354FF6BFE40F5AAD77C" pageId="28" pageNumber="29" type="discussion">
<paragraph id="A94736DE4C34B354FF6BFE40F5AAD77C" blockId="28.[151,1437,151,717]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BFE40F65ED4C9" bold="true" box="[151,267,404,429]" pageId="28" pageNumber="29">Remarks.</emphasis>
Although references about three different 
<typeStatus id="7643887C4C34B354FD01FE41F478D4C9" box="[765,813,405,429]" pageId="28" pageNumber="29">type</typeStatus>
localities on different dates and a male character are given in the original description, there is no indication about how many specimens were examined. Two specimens were located in the FMNH under this name. The specimen below agrees with the original description and considered to have been a 
<typeStatus id="7643887C4C34B354FD5CFDD4F5AED77C" box="[672,763,512,536]" pageId="28" pageNumber="29" type="syntype">syntype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C34B354FF6BFDF0F52BD7A9" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="A94736DE4C34B354FF6BFDF0F52BD7A9" blockId="28.[151,1437,151,717]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BFDF0F69FD759" bold="true" box="[151,458,548,573]" pageId="28" pageNumber="29">Type material examined.</emphasis>
Syntype ♂ (FMNH, 
<figureCitation id="31C32A5B4C34B354FD2CFDF0F40AD758" box="[720,863,548,573]" captionStart="FIGURES 45–52" captionStartId="82.[151,264,1313,1336]" captionTargetBox="[234,1352,195,1292]" captionTargetId="figure@82.[234,1352,195,1292]" captionTargetPageId="82" captionText="FIGURES 45–52 [FMNH ©]. Type and non-type material (habitus and labels): 45, 46, Belonuchus penetrans Silvestri, nontype; 47, 48, Philonthus excelsus Bernhauer, syntype; 49, 50 Philonthus peruvianus Bernhauer, syntype; 51, 52, Philonthus subaeneipennis Bernhauer, holotype. Scale bar= 2 mm." httpUri="https://zenodo.org/record/1454315/files/figure.png" pageId="28" pageNumber="29">Figs. 47–48</figureCitation>
): card mounted with cutouts at head-thorax and abdomen // Süd – Peru Oberh. Paleo, 
<quantity id="6E009B3B4C34B354FD87FD9CF59CD704" box="[635,713,584,608]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="28" pageNumber="29" unit="m" value="3240.0">3240m</quantity>
<date id="DD46101E4C34B354FD2EFD93F417D704" box="[722,834,583,608]" pageId="28" pageNumber="29" value="1936-10-04">10.4.1936</date>
[in Bernhauer’s hand] (green label) // Hbg. – Süd – Peru Sammelreise 1936. Eing. Nr. 1, 1937 (green label) // 219 (white label) // 
<taxonomicName id="6EF84D5D4C34B354FBD8FDB8F2C9D7E0" authority="Brh. Cotypus" authorityName="Brh. Cotypus" box="[1060,1436,620,645]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="excelsus">Philonthus excelsus Brh. Cotypus</taxonomicName>
(white label) // 
<taxonomicName id="6EF84D5D4C34B354FE76FD5BF56ED7CC" authority="Benh." authorityName="Bernhauer" authorityYear="1941" box="[394,571,655,680]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="excelsus">excelsus Benh.</taxonomicName>
Typus 
<taxonomicName id="6EF84D5D4C34B354FD61FD5BF440D7CC" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[669,789,655,680]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="95" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
[in Bernhauer’s hand](red label) // Chicago NHMus M.Bernhauer Collection (white label)”.
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C34B354FF6BFD2FF45AD202" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="A94736DE4C34B354FF6BFD2FF53DD670" blockId="28.[151,1436,763,861]" box="[151,616,763,789]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BFD2FF7F6D670" bold="true" box="[151,163,763,788]" italics="true" pageId="28" pageNumber="29">*</emphasis>
<taxonomicName id="6EF84D5D4C34B354FF5EFD2FF53DD670" ID-CoL="LCC2" authority="(Solsky 1872)" authorityName="Solsky" authorityYear="1872" baseAuthorityName="Solsky" baseAuthorityYear="1872" box="[162,616,763,789]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="taczanovskii">
<emphasis id="9B8CEACC4C34B354FF5EFD2FF696D670" bold="true" box="[162,451,763,788]" italics="true" pageId="28" pageNumber="29">Belonuchus taczanovskii</emphasis>
(
<bibRefCitation id="CD694B2F4C34B354FE2EFD28F535D670" box="[466,608,764,789]" pageId="28" pageNumber="29" refId="ref73125" refString="Solsky, S. (1872) Enumeration et description des Coleopteres de la famille des Staphylinides recueillis par Mrs. C. Ielsky et le Baron de Nolken pendant leurs voyages dans l'Amerique du Sud en 1870 et 1871. Premiere partie. Horae Societatis Entomologicae Rossicae, 8, 289 - 314." type="journal article">Solsky 1872</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFCCBF420D638" blockId="28.[151,1436,763,861]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BFCCBF420D638" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BFCCBF5D4D65C" authority="Solsky 1872: 303" authorityName="Solsky" authorityPageNumber="303" authorityYear="1872" box="[151,641,799,824]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="taczanovskii">
<emphasis id="9B8CEACC4C34B354FF6BFCCBF6F2D65C" box="[151,423,799,824]" italics="true" pageId="28" pageNumber="29">Philonthus taczanovskii</emphasis>
<treatmentCitation id="285910CF4C34B354FE4DFCCBF5D4D65C" author="" box="[433,641,799,824]" page="303" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE4DFCCBF510D65C" box="[433,581,799,824]" pageId="28" pageNumber="29" refId="ref73125" refString="Solsky, S. (1872) Enumeration et description des Coleopteres de la famille des Staphylinides recueillis par Mrs. C. Ielsky et le Baron de Nolken pendant leurs voyages dans l'Amerique du Sud en 1870 et 1871. Premiere partie. Horae Societatis Entomologicae Rossicae, 8, 289 - 314." type="journal article">Solsky 1872</bibRefCitation>
: 303
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C34B354FD6DFCF4F59DD65C" box="[657,712,800,824]" pageId="28" pageNumber="29">Type</typeStatus>
locality: Monte Rico, 
<collectingCountry id="D1EF764E4C34B354FC2BFCF4F349D653" box="[983,1052,800,823]" name="Peru" pageId="28" pageNumber="29">Perou</collectingCountry>
. 
<typeStatus id="7643887C4C34B354FBD6FCF4F334D65C" box="[1066,1121,800,824]" pageId="28" pageNumber="29">Type</typeStatus>
depository: ZIN? (Solsky) #?ST. Distribution: 
<collectingCountry id="D1EF764E4C34B354FE5EFC91F68DD638" box="[418,472,837,860]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C34B354FE1FFC90F516D639" box="[483,579,836,861]" country="Democratic Republic of the Congo" name="Equateur" pageId="28" pageNumber="29">
<collectingCountry id="D1EF764E4C34B354FE1FFC90F516D639" box="[483,579,836,861]" name="Ecuador" pageId="28" pageNumber="29">Ecuador</collectingCountry>
</collectingRegion>
. Note: 
<collectingCountry id="D1EF764E4C34B354FD6EFC91F590D638" box="[658,709,837,860]" name="Peru" pageId="28" pageNumber="29">Peru</collectingCountry>
(
<bibRefCitation id="CD694B2F4C34B354FD28FC90F43CD638" box="[724,873,836,861]" pageId="28" pageNumber="29" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFC5FF5B1D6C0" blockId="28.[151,1436,907,1077]" box="[151,740,907,933]" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BFC5FF5B1D6C0" ID-CoL="LCC4" authority="(Laporte de Castelnau 1840)" authorityName="Laporte de Castelnau" authorityYear="1840" baseAuthorityName="Laporte de Castelnau" baseAuthorityYear="1840" box="[151,740,907,933]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="terminalis">
<emphasis id="9B8CEACC4C34B354FF6BFC5FF6C9D6C0" bold="true" box="[151,412,907,932]" italics="true" pageId="28" pageNumber="29">Belonuchus terminalis</emphasis>
(Laporte de 
<bibRefCitation id="CD694B2F4C34B354FDD1FC58F58ED6C0" box="[557,731,908,933]" pageId="28" pageNumber="29" refId="ref68445" refString="Laporte de Castelnau, F. L. (1840) Histoire naturelle des Insectes Coleopteres, vol. 1. In: Histoire naturelle des animaux articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. Vol. 2. P. Dumenil, Paris, 324 pp., 19 pls." type="book">Castelnau 1840</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFC7BF46BD150" blockId="28.[151,1436,907,1077]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BFC7BF46BD150" infered="true" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BFC7BF46DD6AC" ID-CoL="4ZKF6" authority="Laporte de Castelnau 1840: 176" authorityName="Laporte de Castelnau" authorityPageNumber="176" authorityYear="1840" box="[151,824,943,968]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="terminalis">
<emphasis id="9B8CEACC4C34B354FF6BFC7BF6CED6AC" box="[151,411,943,968]" italics="true" pageId="28" pageNumber="29">Staphylinus terminalis</emphasis>
Laporte de 
<treatmentCitation id="285910CF4C34B354FDC0FC7BF46DD6AC" author="" box="[572,824,943,968]" page="176" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FDC0FC7BF5A2D6AC" box="[572,759,943,968]" pageId="28" pageNumber="29" refId="ref68445" refString="Laporte de Castelnau, F. L. (1840) Histoire naturelle des Insectes Coleopteres, vol. 1. In: Histoire naturelle des animaux articules, Annelides, Crustaces, Arachnides, Myriapodes et Insectes. Vol. 2. P. Dumenil, Paris, 324 pp., 19 pls." type="book">Castelnau 1840</bibRefCitation>
: 176
</treatmentCitation>
</taxonomicName>
. Type locality: Brésil. Type depository:? #? ST. Distribution: Brazil. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C34B354FD5FFC00F3E3D688" ID-CoL="8VS43" authority="(Bernhauer &amp; Schubert 1914: 371)" authorityName="Bernhauer &amp; Schubert" authorityPageNumber="371" authorityYear="1914" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[675,1206,980,1005]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C34B354FD5FFC00F470D689" box="[675,805,980,1005]" italics="true" pageId="28" pageNumber="29">Staphylinus</emphasis>
(
<treatmentCitation id="285910CF4C34B354FCCFFC00F3FBD688" author="" box="[819,1198,980,1005]" page="371" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FCCFFC00F3FBD688" box="[819,1198,980,1005]" pageId="28" pageNumber="29" refId="ref62964" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed), Coleopterorum Catalogus. Vol. 57. W. Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer &amp; Schubert 1914: 371</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; preoccupied, junior primary homonym of 
<taxonomicName id="6EF84D5D4C34B354FE3EFC23F4FED174" ID-CoL="4ZKF6" authority="Erichson 1839: 396" authorityName="Erichson" authorityPageNumber="396" authorityYear="1839" box="[450,939,1015,1040]" class="Insecta" family="Staphylinidae" genus="Staphylinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="terminalis">
<emphasis id="9B8CEACC4C34B354FE3EFC23F5EBD174" box="[450,702,1015,1040]" italics="true" pageId="28" pageNumber="29">Staphylinus terminalis</emphasis>
<treatmentCitation id="285910CF4C34B354FD34FC23F4FED174" author="" box="[712,939,1015,1040]" page="396" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FD34FC23F425D174" box="[712,880,1015,1040]" pageId="28" pageNumber="29" refId="ref65468" refString="Erichson, W. F. (1839) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, viii + 400 pp." type="book">Erichson 1839</bibRefCitation>
: 396
</treatmentCitation>
</taxonomicName>
; conserved by Opinion 2053 (
<bibRefCitation id="CD694B2F4C34B354FAFBFC2CF2DAD174" box="[1287,1423,1016,1040]" pageId="28" pageNumber="29" refId="ref67836" refString="ICZN (2003) Opinion 2053 (Case 3207). Staphylinidae Latreille, 1804 (Insecta, Coleoptera): 65 specific names conserved. Bulletin of Zoological Nomenclature, 60 (3), 237 - 246." type="journal article">ICZN 2003</bibRefCitation>
); distribution within Brazil unknown (Asenjo 
<emphasis id="9B8CEACC4C34B354FD4AFBC9F5BBD151" box="[694,750,1052,1077]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFBB7F521D118" blockId="28.[151,1437,1123,1221]" box="[151,628,1123,1149]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BFBB7F7F6D118" bold="true" box="[151,163,1123,1148]" italics="true" pageId="28" pageNumber="29">*</emphasis>
<taxonomicName id="6EF84D5D4C34B354FF5EFBB7F521D118" ID-CoL="LCC7" authority="Bernhauer 1912" authorityName="Bernhauer" authorityYear="1912" box="[162,628,1123,1149]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tremolerasi">
<emphasis id="9B8CEACC4C34B354FF5EFBB7F6ECD118" bold="true" box="[162,441,1123,1148]" italics="true" pageId="28" pageNumber="29">Belonuchus tremolerasi</emphasis>
Bernhauer 1912
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFB53F54AD1A1" blockId="28.[151,1437,1123,1221]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BFB53F54AD1A1" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BFB53F5C0D1C4" ID-CoL="LCC7" authority="Bernhauer 1912 a: 40" authorityName="Bernhauer" authorityPageNumber="40" authorityYear="1912" box="[151,661,1159,1184]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tremolerasi">
<emphasis id="9B8CEACC4C34B354FF6BFB53F6F5D1C4" box="[151,416,1159,1184]" italics="true" pageId="28" pageNumber="29">Belonuchus tremolerasi</emphasis>
<treatmentCitation id="285910CF4C34B354FE5BFB53F5C0D1C4" author="" box="[423,661,1159,1184]" page="40" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE5BFB53F539D1C4" box="[423,620,1159,1184]" pageId="28" pageNumber="29" refId="ref61973" refString="Bernhauer, M. (1912 a) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article">Bernhauer 1912a</bibRefCitation>
: 40
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C34B354FD5CFB5CF582D1C4" box="[672,727,1160,1184]" pageId="28" pageNumber="29">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C34B354FCBEFB5CF4F3D1C4" box="[834,934,1160,1184]" name="Uruguay" pageId="28" pageNumber="29">Uruguay</collectingCountry>
. 
<typeStatus id="7643887C4C34B354FC4CFB5CF4BDD1C4" box="[944,1000,1160,1184]" pageId="28" pageNumber="29">Type</typeStatus>
depository: FMNH 1 ST. Distribution: 
<collectingCountry id="D1EF764E4C34B354FF39FB79F67DD1A1" box="[197,296,1197,1221]" name="Uruguay" pageId="28" pageNumber="29">Uruguay</collectingCountry>
, 
<collectingCountry id="D1EF764E4C34B354FECFFB78F6F0D1A1" box="[307,421,1196,1221]" name="Argentina" pageId="28" pageNumber="29">Argentina</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C34B354FE4CFB79F54FD1A1" box="[432,538,1197,1221]" country="Paraguay" name="Asuncion" pageId="28" pageNumber="29">
<collectingCountry id="D1EF764E4C34B354FE4CFB79F54FD1A1" box="[432,538,1197,1221]" name="Paraguay" pageId="28" pageNumber="29">Paraguay</collectingCountry>
</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFB27F508D068" blockId="28.[151,1437,1267,1400]" box="[151,605,1267,1293]" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BFB27F508D068" ID-CoL="LCC8" authority="Blackwelder 1943" authorityName="Blackwelder" authorityYear="1943" box="[151,605,1267,1293]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trinitatis">
<emphasis id="9B8CEACC4C34B354FF6BFB27F6DFD068" bold="true" box="[151,394,1267,1292]" italics="true" pageId="28" pageNumber="29">Belonuchus trinitatis</emphasis>
<bibRefCitation id="CD694B2F4C34B354FE6DFB20F508D068" box="[401,605,1268,1293]" pageId="28" pageNumber="29" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFAC3F679D01C" blockId="28.[151,1437,1267,1400]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BFAC3F679D01C" pageId="28" pageNumber="29">
<materialsCitation id="19903C834C34B354FF6BFAC3F2CDD054" ID-GBIF-Occurrence="1915497821" box="[151,1432,1303,1328]" country="Trinidad and Tobago" location="Department of Agriculture Stock Farm" municipality="St" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C34B354FF6BFAC3F5C5D054" ID-CoL="LCC8" authority="Blackwelder 1943: 423" authorityName="Blackwelder" authorityPageNumber="423" authorityYear="1943" box="[151,656,1303,1328]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trinitatis">
<emphasis id="9B8CEACC4C34B354FF6BFAC3F6D5D054" box="[151,384,1303,1328]" italics="true" pageId="28" pageNumber="29">Belonuchus trinitatis</emphasis>
<treatmentCitation id="285910CF4C34B354FE74FAC3F5C5D054" author="" box="[392,656,1303,1328]" page="423" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE74FAC3F50DD054" box="[392,600,1303,1328]" pageId="28" pageNumber="29" refId="ref63152" refString="Blackwelder, R. E. (1943) Monograph of the West Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, i - viii, 1 - 658. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book chapter">Blackwelder 1943</bibRefCitation>
: 423
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C34B354FD67FACCF587D054" box="[667,722,1304,1328]" pageId="28" pageNumber="29">Type</typeStatus>
locality: 
<collectingCountry id="D1EF764E4C34B354FCC0FAC3F4F5D054" box="[828,928,1303,1328]" name="Trinidad and Tobago" pageId="28" pageNumber="29">Trinidad</collectingCountry>
: 
<location id="AC2760054C34B354FC56FACCF209D04B" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C34B354FC56FACCF209D04B" box="[938,1372,1303,1328]" country="Trinidad and Tobago" municipality="St" name="Department of Agriculture Stock Farm" pageId="28" pageNumber="29">Department of Agriculture Stock Farm</location>
at 
<collectingMunicipality id="4923ACA44C34B354FA82FACCF2CDD054" box="[1406,1432,1304,1328]" pageId="28" pageNumber="29">St</collectingMunicipality>
</materialsCitation>
. 
<materialsCitation id="19903C834C34B354FF39FAE8F5D6D031" ID-GBIF-Occurrence="1915497852" box="[197,643,1340,1365]" country="Trinidad and Tobago" location="Port of Spain" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Port of Spain" typeStatus="holotype">
Joseph, 
<quantity id="6E009B3B4C34B354FED8FAE8F62ED031" box="[292,379,1340,1365]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="28" pageNumber="29" unit="mi" value="6.0">6 miles</quantity>
east of 
<collectingRegion id="6B3CF83C4C34B354FE1EFAE9F5D6D031" box="[482,643,1340,1365]" country="Trinidad and Tobago" name="Port of Spain" pageId="28" pageNumber="29">Port of Spain</collectingRegion>
</materialsCitation>
. 
<materialsCitation id="19903C834C34B354FD6FFAE9F67DD01C" ID-GBIF-Occurrence="1915497847" collectionCode="USNM" country="Trinidad and Tobago" location="Distribution" pageId="28" pageNumber="29" specimenCode="FMNH 1" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="7643887C4C34B354FD6FFAE9F59ED031" box="[659,715,1341,1365]" pageId="28" pageNumber="29">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C34B354FC9DFAE8F4E7D030" box="[865,946,1340,1364]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29">USNM</collectionCode>
HT 
<specimenCount id="BFFEFD574C34B354FC13FAE9F353D030" box="[1007,1030,1341,1364]" pageId="28" pageNumber="29" type="male">♂</specimenCount>
, 4 PT; 
<specimenCode id="F95E9EA54C34B354FB98FAE9F39AD030" box="[1124,1231,1340,1364]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="28" pageNumber="29">FMNH 1</specimenCode>
PT. 
<location id="AC2760054C34B354FAF1FAE8F2CCD031" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C34B354FAF1FAE8F2CCD031" box="[1293,1433,1340,1365]" country="Trinidad and Tobago" name="Distribution" pageId="28" pageNumber="29">Distribution</location>
: 
<collectingCountry id="D1EF764E4C34B354FF39FA8BF67DD01C" box="[197,296,1375,1400]" name="Trinidad and Tobago" pageId="28" pageNumber="29">Trinidad</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFA73F52ED0A4" blockId="28.[151,1437,1447,1581]" box="[151,635,1447,1472]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BFA73F7F6D0A4" bold="true" box="[151,163,1447,1472]" italics="true" pageId="28" pageNumber="29">*</emphasis>
<taxonomicName id="6EF84D5D4C34B354FF5EFA73F52ED0A4" ID-CoL="5WGJ9" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,635,1447,1472]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trochanterinus">
<emphasis id="9B8CEACC4C34B354FF5EFA73F6B4D0A4" bold="true" box="[162,481,1447,1472]" italics="true" pageId="28" pageNumber="29">Belonuchus trochanterinus</emphasis>
(
<bibRefCitation id="CD694B2F4C34B354FE0CFA73F526D0A4" box="[496,627,1447,1472]" pageId="28" pageNumber="29" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BFA18F6B0D349" blockId="28.[151,1437,1447,1581]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BFA18F6B0D349" infered="true" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BFA18F5D6D080" ID-CoL="4GC27" authority="Sharp 1885: 417" authorityName="Sharp" authorityPageNumber="417" authorityYear="1885" box="[151,643,1484,1509]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trochanterinus">
<emphasis id="9B8CEACC4C34B354FF6BFA18F6E9D081" box="[151,444,1484,1509]" italics="true" pageId="28" pageNumber="29">Philonthus trochanterinus</emphasis>
<treatmentCitation id="285910CF4C34B354FE38FA18F5D6D080" author="" box="[452,643,1484,1509]" page="417" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE38FA18F51FD081" box="[452,586,1484,1509]" pageId="28" pageNumber="29" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 417
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C34B354FD73FA19F592D081" box="[655,711,1485,1509]" pageId="28" pageNumber="29">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C34B354FCCFFA18F4FAD081" box="[819,943,1484,1509]" country="Guatemala" name="Guatemala" pageId="28" pageNumber="29">
<collectingCountry id="D1EF764E4C34B354FCCFFA18F4FAD081" box="[819,943,1484,1509]" name="Guatemala" pageId="28" pageNumber="29">Guatemala</collectingCountry>
</collectingRegion>
, Capetillo. 
<typeStatus id="7643887C4C34B354FBCDFA19F33CD081" box="[1073,1129,1485,1509]" pageId="28" pageNumber="29">Type</typeStatus>
depository: BMNH #? ST. Distribution: 
<collectingRegion id="6B3CF83C4C34B354FEA3FA3BF6E2D36C" box="[351,439,1519,1544]" country="Mexico" name="Mexico" pageId="28" pageNumber="29">
<collectingCountry id="D1EF764E4C34B354FEA3FA3BF6E2D36C" box="[351,439,1519,1544]" name="Mexico" pageId="28" pageNumber="29">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C34B354FE3BFA3BF511D36C" box="[455,580,1519,1544]" name="Guatemala" pageId="28" pageNumber="29">
<collectingRegion id="6B3CF83C4C34B354FE3BFA3BF511D36C" box="[455,580,1519,1544]" country="Guatemala" name="Guatemala" pageId="28" pageNumber="29">Guatemala</collectingRegion>
</collectingCountry>
. Note: comb. nov. ex 
<emphasis id="9B8CEACC4C34B354FCB2FA3BF49AD36C" box="[846,975,1519,1544]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FCB2FA3BF49FD36C" ID-CoL="8VRYR" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[846,970,1519,1544]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
,
</emphasis>
distribution within 
<collectingRegion id="6B3CF83C4C34B354FB46FA3BF25AD36C" box="[1210,1295,1519,1544]" country="Mexico" name="Mexico" pageId="28" pageNumber="29">
<collectingCountry id="D1EF764E4C34B354FB46FA3BF25AD36C" box="[1210,1295,1519,1544]" name="Mexico" pageId="28" pageNumber="29">Mexico</collectingCountry>
</collectingRegion>
(Navarrete- 
<collectingRegion id="6B3CF83C4C34B354FF39F9C0F64BD349" box="[197,286,1556,1581]" country="Costa Rica" name="Heredia" pageId="28" pageNumber="29">Heredia</collectingRegion>
<emphasis id="9B8CEACC4C34B354FED8F9C1F608D349" box="[292,349,1556,1581]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
2002: 335).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BF982F5D6D30B" blockId="28.[151,1437,1622,1719]" box="[151,643,1622,1647]" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BF982F5D6D30B" ID-CoL="LCCC" authority="Bernhauer 1934" authorityName="Bernhauer" authorityYear="1934" box="[151,643,1622,1647]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="9B8CEACC4C34B354FF6BF982F69DD30B" bold="true" box="[151,456,1622,1647]" italics="true" pageId="28" pageNumber="29">Belonuchus tucumanensis</emphasis>
Bernhauer 1934
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BF9AEF687D3D3" blockId="28.[151,1437,1622,1719]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BF9AEF687D3D3" pageId="28" pageNumber="29">
<materialsCitation id="19903C834C34B354FF6BF9AEF69BD3D3" ID-GBIF-Occurrence="1915497827" collectionCode="FMNH" country="Argentina" location="Distribution" pageId="28" pageNumber="29" specimenCode="FMNH 1" specimenCount="2" stateProvince="Tucuman" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C34B354FF6BF9AEF58BD3F7" ID-CoL="LCCC" authority="Bernhauer 1934 b: 213" authorityName="Bernhauer" authorityPageNumber="213" authorityYear="1934" box="[151,734,1658,1683]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="9B8CEACC4C34B354FF6BF9AEF694D3F7" box="[151,449,1658,1683]" italics="true" pageId="28" pageNumber="29">Belonuchus tucumanensis</emphasis>
<treatmentCitation id="285910CF4C34B354FE2CF9AEF58BD3F7" author="" box="[464,734,1658,1683]" page="213" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE2CF9AEF5CBD3F7" box="[464,670,1658,1683]" pageId="28" pageNumber="29" refId="ref62723" refString="Bernhauer, M. (1934 b) Neue Staphyliniden aus Argentinien (IV) (Col.) (34. Beitrag zur Kenntnis der sudamerikanischen Fauna). Revista de Entomologia, 4 (2), 212 - 221." type="journal article">Bernhauer 1934b</bibRefCitation>
: 213
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C34B354FD0DF9AFF47DD3F7" box="[753,808,1659,1683]" pageId="28" pageNumber="29">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C34B354FC5EF9AFF35AD3F7" box="[930,1039,1659,1683]" country="Argentina" name="Tucuman" pageId="28" pageNumber="29">Tucumán</collectingRegion>
. 
<typeStatus id="7643887C4C34B354FBDDF9AFF30DD3F7" box="[1057,1112,1659,1683]" pageId="28" pageNumber="29">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C34B354FB09F9AFF236D3F7" box="[1269,1379,1659,1683]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="28" pageNumber="29">FMNH 1</specimenCode>
HT. 
<location id="AC2760054C34B354FF39F94AF604D3D3" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C34B354FF39F94AF604D3D3" box="[197,337,1694,1719]" country="Argentina" name="Distribution" pageId="28" pageNumber="29" stateProvince="Tucuman">Distribution</location>
: 
<collectingCountry id="D1EF764E4C34B354FEA7F94AF69BD3D3" box="[347,462,1694,1719]" name="Argentina" pageId="28" pageNumber="29">Argentina</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BF935F56AD39E" blockId="28.[151,1436,1761,1894]" box="[151,575,1761,1786]" pageId="28" pageNumber="29">
<emphasis id="9B8CEACC4C34B354FF6BF935F7F6D39E" bold="true" box="[151,163,1761,1786]" italics="true" pageId="28" pageNumber="29">*</emphasis>
<taxonomicName id="6EF84D5D4C34B354FF5EF935F56AD39E" ID-CoL="68GJ9" authority="Solsky 1870" authorityName="Solsky" authorityYear="1870" box="[162,575,1761,1786]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variolosus">
<emphasis id="9B8CEACC4C34B354FF5EF935F6F9D39E" bold="true" box="[162,428,1761,1786]" italics="true" pageId="28" pageNumber="29">Belonuchus variolosus</emphasis>
<bibRefCitation id="CD694B2F4C34B354FE48F935F56AD39E" box="[436,575,1761,1786]" pageId="28" pageNumber="29" refId="ref73086" refString="Solsky, S. (1870) Staphylins de l'Amerique meridionale et du Mexique. II. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 42 (4), 257 - 267. [1869]" type="journal article">Solsky 1870</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BF8D1F3F6D226" blockId="28.[151,1436,1761,1894]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BF8D1F3F6D226" pageId="28" pageNumber="29">
<materialsCitation id="19903C834C34B354FF6BF8D1F3D4D27A" ID-GBIF-Occurrence="1915497812" box="[151,1153,1797,1822]" country="French Guiana" location="Cayenne" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C34B354FF6BF8D1F52ED27A" ID-CoL="68GJ9" authority="Solsky 1870: 263" authorityName="Solsky" authorityPageNumber="263" authorityYear="1870" box="[151,635,1797,1822]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variolosus">
<emphasis id="9B8CEACC4C34B354FF6BF8D1F6CED27A" box="[151,411,1797,1822]" italics="true" pageId="28" pageNumber="29">Belonuchus variolosus</emphasis>
<treatmentCitation id="285910CF4C34B354FE54F8D1F52ED27A" author="" box="[424,635,1797,1822]" page="263" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE54F8D1F568D27A" box="[424,573,1797,1822]" pageId="28" pageNumber="29" refId="ref73086" refString="Solsky, S. (1870) Staphylins de l'Amerique meridionale et du Mexique. II. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 42 (4), 257 - 267. [1869]" type="journal article">Solsky 1870</bibRefCitation>
: 263
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C34B354FD70F8D2F591D27A" box="[652,708,1798,1822]" pageId="28" pageNumber="29">Type</typeStatus>
locality: &quot; 
<location id="AC2760054C34B354FCBEF8D2F4FFD27A" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C34B354FCBEF8D2F4FFD27A" box="[834,938,1798,1822]" country="French Guiana" name="Cayenne" pageId="28" pageNumber="29">Cayenne</location>
&quot; [
<collectingCountry id="D1EF764E4C34B354FC39F8D1F32FD27A" box="[965,1146,1797,1822]" name="French Guiana" pageId="28" pageNumber="29">French Guiana</collectingCountry>
]
</materialsCitation>
. 
<materialsCitation id="19903C834C34B354FB6DF8D2F3F6D226" ID-GBIF-Occurrence="1915497796" collectionCode="ZIN" country="French Guiana" county="Note" location="Solsky" municipality="Cayenne" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C34B354FB6DF8D2F39CD27A" box="[1169,1225,1798,1822]" pageId="28" pageNumber="29">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C34B354FA9DF8D2F2C7D27A" box="[1377,1426,1798,1822]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="28" pageNumber="29">ZIN</collectionCode>
? (
<location id="AC2760054C34B354FF31F8FDF649D226" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C34B354FF31F8FDF649D226" box="[205,284,1833,1858]" country="French Guiana" county="Note" municipality="Cayenne" name="Solsky" pageId="28" pageNumber="29">Solsky</location>
) #? 
<location id="AC2760054C34B354FEB9F8FEF6ABD226" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C34B354FEB9F8FEF6ABD226" box="[325,510,1833,1858]" country="French Guiana" county="Note" municipality="Cayenne" name="ST. Distribution" pageId="28" pageNumber="29">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C34B354FDF4F8FDF5FBD226" box="[520,686,1833,1858]" name="French Guiana" pageId="28" pageNumber="29">French Guiana</collectingCountry>
(
<collectingMunicipality id="4923ACA44C34B354FD40F8FEF471D226" box="[700,804,1834,1858]" pageId="28" pageNumber="29">Cayenne</collectingMunicipality>
). 
<collectingCounty id="40264E524C34B354FCCBF8FEF427D225" box="[823,882,1834,1857]" pageId="28" pageNumber="29">Note</collectingCounty>
: generic affiliation unclear.
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C34B354FF6BF89AF45AD202" blockId="28.[151,1436,1761,1894]" box="[151,783,1869,1894]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BF89AF45AD202" box="[151,783,1869,1894]" pageId="28" pageNumber="29">
= 
<taxonomicName id="6EF84D5D4C34B354FF51F899F6F0D202" ID-CoL="8XVPM" box="[173,421,1869,1894]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="variolaris">
<emphasis id="9B8CEACC4C34B354FF51F899F6F0D202" box="[173,421,1869,1894]" italics="true" pageId="28" pageNumber="29">Belonuchus variolaris</emphasis>
</taxonomicName>
; 
<treatmentCitation id="285910CF4C34B354FE48F899F52ED202" author="" box="[436,635,1869,1894]" page="305" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE48F899F516D202" box="[436,579,1869,1894]" pageId="28" pageNumber="29" refId="ref73125" refString="Solsky, S. (1872) Enumeration et description des Coleopteres de la famille des Staphylinides recueillis par Mrs. C. Ielsky et le Baron de Nolken pendant leurs voyages dans l'Amerique du Sud en 1870 et 1871. Premiere partie. Horae Societatis Entomologicae Rossicae, 8, 289 - 314." type="journal article">Solsky 1872</bibRefCitation>
: 305
</treatmentCitation>
, misspelling.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C34B354FF6BF840F525D2CA" box="[151,624,1940,1966]" pageId="28" pageNumber="29" type="description">
<paragraph id="A94736DE4C34B354FF6BF840F525D2CA" blockId="28.[151,1436,1940,2038]" box="[151,624,1940,1966]" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BF840F525D2CA" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,624,1940,1966]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="venezolanus">
<emphasis id="9B8CEACC4C34B354FF6BF840F6E0D2C9" bold="true" box="[151,437,1940,1965]" italics="true" pageId="28" pageNumber="29">Belonuchus venezolanus</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E1E265554C34B355FF6BF86DF4ACD3FC" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="A94736DE4C34B354FF6BF86DF20ED292" blockId="28.[151,1436,1940,2038]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="89E811F04C34B354FF6BF86DF20ED292" pageId="28" pageNumber="29">
<taxonomicName id="6EF84D5D4C34B354FF6BF86DF5FED2B6" ID-CoL="LCCF" authority="Bernhauer 1917 c: 104" authorityName="Bernhauer" authorityPageNumber="104" authorityYear="1917" box="[151,683,1977,2002]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="venezolanus">
<emphasis id="9B8CEACC4C34B354FF6BF86DF6FCD2B6" box="[151,425,1977,2002]" italics="true" pageId="28" pageNumber="29">Belonuchus venezolanus</emphasis>
<treatmentCitation id="285910CF4C34B354FE4CF86DF5FED2B6" author="" box="[432,683,1977,2002]" page="104" pageId="28" pageNumber="29">
<bibRefCitation id="CD694B2F4C34B354FE4CF86DF521D2B6" box="[432,628,1977,2002]" pageId="28" pageNumber="29" refId="ref62261" refString="Bernhauer, M. (1917 c) Neue sudamerikanische Staphyliniden (18. Beitrag). Wiener Entomologische Zeitung, 36 (3 - 5), 102 - 116. https: // doi. org / 10.5962 / bhl. part. 12925" type="journal article">Bernhauer 1917c</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C34B354FD4AF86EF5B8D2B6" box="[694,749,1978,2002]" pageId="28" pageNumber="29">Type</typeStatus>
localities: 
<collectingCountry id="D1EF764E4C34B354FC94F86DF48ED2B6" box="[872,987,1977,2002]" name="Venezuela" pageId="28" pageNumber="29">Venezuela</collectingCountry>
&amp; Columbien. 
<typeStatus id="7643887C4C34B354FB74F86EF395D2B6" box="[1160,1216,1978,2002]" pageId="28" pageNumber="29">Type</typeStatus>
depository: FMNH 3ST; NMW 1+ST?; ZMUH 1+ST?. Distribution: 
<collectingCountry id="D1EF764E4C34B354FD0DF809F432D292" box="[753,871,2013,2038]" name="Venezuela" pageId="28" pageNumber="29">Venezuela</collectingCountry>
, 
<collectingCountry id="D1EF764E4C34B354FC8EF809F4B1D292" box="[882,996,2013,2038]" name="Colombia" pageId="28" pageNumber="29">Colombia</collectingCountry>
. Note: generic affiliation unclear.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFF43F5C9D5D4" blockId="29.[151,1436,151,464]" box="[151,668,151,176]" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFF43F5C9D5D4" ID-CoL="LCCH" authority="Baudi di Selve 1848" authorityName="Baudi di Selve" authorityYear="1848" box="[151,668,151,176]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis id="9B8CEACC4C35B355FF6BFF43F6E4D5D4" bold="true" box="[151,433,151,176]" italics="true" pageId="29" pageNumber="30">Belonuchus viridipennis</emphasis>
Baudi di 
<bibRefCitation id="CD694B2F4C35B355FDE3FF43F5C9D5D4" box="[543,668,151,176]" pageId="29" pageNumber="30" refId="ref61718" refString="Baudi di Selve, F. (1848) Alcuni specie nuove di Stafilini. In: Baudi di Selve, F. &amp; Truqui, E. (Eds.), Studi Entomologici. Tomo Primo. Fascicolo II. Stamperia Sociale degli Artisti Tipogafi, Torino, pp. 113 - 148." type="book chapter">Selve 1848</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFF68F407D424" blockId="29.[151,1436,151,464]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFF68F407D424" infered="true" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFF68F586D5B1" ID-CoL="LCCH" authority="Baudi di Selve 1848: 125" authorityName="Baudi di Selve" authorityPageNumber="125" authorityYear="1848" box="[151,723,188,213]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis id="9B8CEACC4C35B355FF6BFF68F6FDD5B1" box="[151,424,188,213]" italics="true" pageId="29" pageNumber="30">Belonuchus viridipennis</emphasis>
Baudi di 
<treatmentCitation id="285910CF4C35B355FDE4FF68F586D5B1" author="" box="[536,723,188,213]" page="125" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FDE4FF68F5CFD5B0" box="[536,666,188,213]" pageId="29" pageNumber="30" refId="ref61718" refString="Baudi di Selve, F. (1848) Alcuni specie nuove di Stafilini. In: Baudi di Selve, F. &amp; Truqui, E. (Eds.), Studi Entomologici. Tomo Primo. Fascicolo II. Stamperia Sociale degli Artisti Tipogafi, Torino, pp. 113 - 148." type="book chapter">Selve 1848</bibRefCitation>
: 125
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C35B355FD23FF69F443D5B1" box="[735,790,189,213]" pageId="29" pageNumber="30">Type</typeStatus>
locality: Novae Valenciae in Columbia [
<collectingCountry id="D1EF764E4C35B355FB18FF68F208D5B1" box="[1252,1373,188,213]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
, teste 
<treatmentCitation id="285910CF4C35B355FF39FF34F694D59C" author="" box="[197,449,223,248]" page="1594" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FF39FF34F626D59C" box="[197,371,224,248]" pageId="29" pageNumber="30" refId="ref67370" refString="Herman, L. H. (2001 c) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part III. Bulletin of the American Museum of Natural History, 265, 1067 - 1806." type="journal article">Herman 2001c</bibRefCitation>
: 1594
</treatmentCitation>
]. 
<typeStatus id="7643887C4C35B355FE2AFF34F55BD59C" box="[470,526,224,248]" pageId="29" pageNumber="30">Type</typeStatus>
depository: MRSN/MCSN? #?ST. Distribution: 
<collectingRegion id="6B3CF83C4C35B355FBB4FF0BF3F5D59C" box="[1096,1184,223,248]" country="Mexico" name="Mexico" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FBB4FF0BF3F5D59C" box="[1096,1184,223,248]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C35B355FB4CFF0BF279D59C" box="[1200,1324,223,248]" country="Guatemala" name="Guatemala" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FB4CFF0BF279D59C" box="[1200,1324,223,248]" name="Guatemala" pageId="29" pageNumber="30">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C35B355FAC0FF34F2CDD593" box="[1340,1432,224,247]" country="Panama" name="Panama" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FAC0FF34F2CDD593" box="[1340,1432,224,247]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
</collectingRegion>
, 
<collectingCountry id="D1EF764E4C35B355FF39FED0F663D479" box="[197,310,260,285]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
, 
<collectingCountry id="D1EF764E4C35B355FEB6FED0F694D479" box="[330,449,260,285]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
. Note: distribution within 
<collectingRegion id="6B3CF83C4C35B355FCF4FED0F408D479" box="[776,861,260,285]" country="Mexico" name="Mexico" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FCF4FED0F408D479" box="[776,861,260,285]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C35B355FC88FED0F3B4D478" box="[884,1249,260,285]" pageId="29" pageNumber="30" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C35B355FBA9FED1F3C0D479" box="[1109,1173,260,285]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2002
</bibRefCitation>
). Note: generic affiliation unclear (based on non-type material in FMNH).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFE99F59AD4B4" blockId="29.[151,1436,151,464]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFE99F59AD4B4" infered="true" pageId="29" pageNumber="30">
= 
<taxonomicName id="6EF84D5D4C35B355FF51FE98F501D400" ID-CoL="LC76" authority="Sharp 1885: 433" authorityName="Sharp" authorityPageNumber="433" authorityYear="1885" box="[173,596,332,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="dejectus">
<emphasis id="9B8CEACC4C35B355FF51FE98F6C7D401" box="[173,402,332,357]" italics="true" pageId="29" pageNumber="30">Belonuchus dejectus</emphasis>
<treatmentCitation id="285910CF4C35B355FE65FE98F501D400" author="" box="[409,596,332,357]" page="433" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE65FE98F548D401" box="[409,541,332,357]" pageId="29" pageNumber="30" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 433
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C35B355FDA2FE99F5C3D401" box="[606,662,333,357]" pageId="29" pageNumber="30">Type</typeStatus>
localities: 
<collectingRegion id="6B3CF83C4C35B355FCECFE98F43FD401" box="[784,874,332,357]" country="Mexico" name="Mexico" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FCECFE98F43FD401" box="[784,874,332,357]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
</collectingRegion>
: 
<collectingRegion id="6B3CF83C4C35B355FC88FE98F482D401" box="[884,983,332,357]" country="Argentina" name="Cordoba" pageId="29" pageNumber="30">Cordova</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C35B355FC1EFE98F37ED401" box="[994,1067,332,357]" country="Guatemala" name="Jalapa" pageId="29" pageNumber="30">Jalapa</collectingRegion>
, Las Vigas, Mirador; 
<collectingRegion id="6B3CF83C4C35B355FAE0FE98F2CDD401" box="[1308,1432,332,357]" country="Guatemala" name="Guatemala" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FAE0FE98F2CDD401" box="[1308,1432,332,357]" name="Guatemala" pageId="29" pageNumber="30">Guatemala</collectingCountry>
</collectingRegion>
, Capetillo, Cerro Zunil, Balheu, Tamahu, San Joaquin, Purula, El Tumbador, San Gerónimo; 
<collectingRegion id="6B3CF83C4C35B355FB3BFEA4F271D4E3" box="[1223,1316,368,391]" country="Panama" name="Panama" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FB3BFEA4F271D4E3" box="[1223,1316,368,391]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
</collectingRegion>
: Volcan de 
<collectingRegion id="6B3CF83C4C35B355FF39FE40F677D4C9" box="[197,290,404,429]" country="Panama" name="Chiriqui" pageId="29" pageNumber="30">Chiriqui</collectingRegion>
<quantity id="6E009B3B4C35B355FED7FE40F693D4C9" box="[299,454,404,429]" metricMagnitude="2" metricUnit="m" metricValue="9.906" metricValueMax="12.192" metricValueMin="7.62" pageId="29" pageNumber="30" unit="ft" value="3250.0" valueMax="4000.0" valueMin="2500.0">2500–4000 ft</quantity>
. 
<typeStatus id="7643887C4C35B355FE2EFE41F55CD4C9" box="[466,521,405,429]" pageId="29" pageNumber="30">Type</typeStatus>
depository: BMNH 14+ST. Distribution: 
<collectingRegion id="6B3CF83C4C35B355FC1BFE40F36AD4C9" box="[999,1087,404,429]" country="Mexico" name="Mexico" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FC1BFE40F36AD4C9" box="[999,1087,404,429]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C35B355FBB1FE40F39CD4C9" box="[1101,1225,404,429]" country="Guatemala" name="Guatemala" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FBB1FE40F39CD4C9" box="[1101,1225,404,429]" name="Guatemala" pageId="29" pageNumber="30">Guatemala</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C35B355FB2AFE41F267D4C8" box="[1238,1330,405,428]" country="Panama" name="Panama" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FB2AFE41F267D4C8" box="[1238,1330,405,428]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
</collectingRegion>
. Note: as synonym of 
<taxonomicName id="6EF84D5D4C35B355FEADFE6DF59ED4B4" ID-CoL="LCCH" authority="(Sharp 1887: 789)" authorityName="Sharp" authorityPageNumber="789" authorityYear="1887" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[337,715,439,464]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="viridipennis">
<emphasis id="9B8CEACC4C35B355FEADFE6DF6A3D4B4" box="[337,502,439,464]" italics="true" pageId="29" pageNumber="30">B. viridipennis</emphasis>
(
<treatmentCitation id="285910CF4C35B355FDF8FE63F596D4B4" author="" box="[516,707,439,464]" page="789" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FDF8FE63F596D4B4" box="[516,707,439,464]" pageId="29" pageNumber="30" refId="ref72534" refString="Sharp, D. (1887) Introduction, Supplement, Index. In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2). Taylor &amp; Francis, London, pp. v - x + 748 - 802 + 803 - 824. [http: // www. sil. si. edu / digitalcollections / bca /]" type="book chapter">Sharp 1887: 789</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFE2BF50BD77C" blockId="29.[151,1437,511,608]" box="[151,606,511,536]" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFE2BF50BD77C" ID-CoL="LCCJ" authority="Bernhauer 1906" authorityName="Bernhauer" authorityYear="1906" box="[151,606,511,536]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vulneratus">
<emphasis id="9B8CEACC4C35B355FF6BFE2BF6F6D77C" bold="true" box="[151,419,511,536]" italics="true" pageId="29" pageNumber="30">Belonuchus vulneratus</emphasis>
<bibRefCitation id="CD694B2F4C35B355FE56FE2BF50BD77C" box="[426,606,511,536]" pageId="29" pageNumber="30" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFDF0F24AD704" blockId="29.[151,1437,511,608]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFDF0F24AD704" pageId="29" pageNumber="30">
<materialsCitation id="19903C834C35B355FF6BFDF0F372D758" ID-GBIF-Occurrence="1915497871" box="[151,1063,548,573]" country="Peru" location="Pachitea" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C35B355FF6BFDF0F5E7D759" ID-CoL="LCCJ" authority="Bernhauer 1906: 335" authorityName="Bernhauer" authorityPageNumber="335" authorityYear="1906" box="[151,690,548,573]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="vulneratus">
<emphasis id="9B8CEACC4C35B355FF6BFDF0F6F5D759" box="[151,416,548,573]" italics="true" pageId="29" pageNumber="30">Belonuchus vulneratus</emphasis>
<treatmentCitation id="285910CF4C35B355FE4CFDF0F5E7D759" author="" box="[432,690,548,573]" page="335" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE4CFDF0F524D759" box="[432,625,548,573]" pageId="29" pageNumber="30" refId="ref61774" refString="Bernhauer, M. (1906) Neue Staphyliniden aus Sudamerika. 3. Stuck. Verhandlungen der Kaiserlich-Koniglichen Zoologisch- Botanischen Gesellschaft in Wien, 56, 322 - 339." type="journal article">Bernhauer 1906</bibRefCitation>
: 335
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C35B355FD3AFDF1F5ABD759" box="[710,766,549,573]" pageId="29" pageNumber="30">Type</typeStatus>
locality: 
<location id="AC2760054C35B355FC86FDF0F489D759" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C35B355FC86FDF0F489D759" box="[890,988,548,573]" country="Peru" name="Pachitea" pageId="29" pageNumber="30">Pachitea</location>
, 
<collectingCountry id="D1EF764E4C35B355FC0CFDF1F372D758" box="[1008,1063,549,572]" name="Peru" pageId="29" pageNumber="30">Peru</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="19903C834C35B355FBC7FDF1F6C7D73B" ID-GBIF-Occurrence="1915497874" collectionCode="FMNH" country="Peru" location="Distribution" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C35B355FBC7FDF1F326D759" box="[1083,1139,549,573]" pageId="29" pageNumber="30">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C35B355FAEDFDF1F237D758" box="[1297,1378,549,572]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="29" pageNumber="30">FMNH</collectionCode>
HT. 
<location id="AC2760054C35B355FF39FD93F604D704" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C35B355FF39FD93F604D704" box="[197,337,583,608]" country="Peru" name="Distribution" pageId="29" pageNumber="30">Distribution</location>
: 
<collectingCountry id="D1EF764E4C35B355FEA7FD9CF6C7D73B" box="[347,402,584,607]" name="Peru" pageId="29" pageNumber="30">Peru</collectingCountry>
</materialsCitation>
. Note: Peru (
<bibRefCitation id="CD694B2F4C35B355FDDFFD93F5ECD704" box="[547,697,583,608]" pageId="29" pageNumber="30" refId="ref61407" refString="Asenjo, A. (2004) Lista preliminar de las especies de Staphylinidae (Coleoptera) registradas para Peru. Revista Peruana de Entomologia, 44, 55 - 64." type="journal article">Asenjo 2004</bibRefCitation>
). Note: not 
<taxonomicName id="6EF84D5D4C35B355FCC7FD93F4EBD704" ID-CoL="8VRHV" box="[827,958,583,608]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C35B355FCC7FD93F4EBD704" box="[827,958,583,608]" italics="true" pageId="29" pageNumber="30">Belonuchus</emphasis>
</taxonomicName>
, to be placed in a further study.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFD5BF52BD7CC" blockId="29.[151,1436,655,789]" box="[151,638,655,680]" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFD5BF52BD7CC" ID-CoL="LCCK" authority="(Bernhauer 1921)" authorityName="Bernhauer" authorityYear="1921" baseAuthorityName="Bernhauer" baseAuthorityYear="1921" box="[151,638,655,680]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="weiserianus">
<emphasis id="9B8CEACC4C35B355FF6BFD5BF6E4D7CC" bold="true" box="[151,433,655,680]" italics="true" pageId="29" pageNumber="30">Belonuchus weiserianus</emphasis>
(Bernhauer 1921)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFD60F4C7D670" blockId="29.[151,1436,655,789]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFD60F4C7D670" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFD60F59ED7A9" ID-CoL="4GC59" authority="Bernhauer 1921 c: 106" authorityName="Bernhauer" authorityPageNumber="106" authorityYear="1921" box="[151,715,692,717]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="weiserianus">
<emphasis id="9B8CEACC4C35B355FF6BFD60F6FDD7A8" box="[151,424,692,717]" italics="true" pageId="29" pageNumber="30">Philonthus weiserianus</emphasis>
<treatmentCitation id="285910CF4C35B355FE46FD60F59ED7A9" author="" box="[442,715,692,717]" page="106" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE46FD60F5DCD7A8" box="[442,649,692,717]" pageId="29" pageNumber="30" refId="ref62483" refString="Bernhauer, M. (1921 c) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article">Bernhauer 1921c</bibRefCitation>
: 106
</treatmentCitation>
</taxonomicName>
. Type locality: Hornadita, 
<quantity id="6E009B3B4C35B355FBDEFD60F327D7A8" box="[1058,1138,692,716]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="29" pageNumber="30" unit="m" value="3400.0">3400m</quantity>
, Jujuy, Argentina. Type depository: FMNH LT ♀. Distribution: Argentina. Note: comb. nov. ex 
<taxonomicName id="6EF84D5D4C35B355FC04FD03F324D794" ID-CoL="8VRYR" baseAuthorityName="Bernhauer &amp; Schubert" baseAuthorityYear="1914" box="[1016,1137,727,752]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C35B355FC04FD03F324D794" box="[1016,1137,727,752]" italics="true" pageId="29" pageNumber="30">Philonthus</emphasis>
</taxonomicName>
and lectotype designation (
<treatmentCitation id="285910CF4C35B355FF30FD28F6E2D670" author="" box="[204,439,764,789]" page="8" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FF30FD28F6E2D670" box="[204,439,764,789]" pageId="29" pageNumber="30" refId="ref64208" refString="Chani-Posse, M. (2010) Revision of the southern South American species of Philonthus Stephens Coleoptera: Staphylinidae. Zootaxa, 2595, 1 - 70." type="journal article">Chani-Posse 2010: 8</bibRefCitation>
</treatmentCitation>
); redescription (
<treatmentCitation id="285910CF4C35B355FD93FD28F4D3D670" author="" box="[623,902,764,789]" page="224" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FD93FD28F4D3D670" box="[623,902,764,789]" pageId="29" pageNumber="30" refId="ref64276" refString="Chani-Posse, M. (2011 b) Redescription of some South American species of Belonuchus Coleoptera: Staphylinidae, with a checklist of the southern South American species of the genus. Revista de la Sociedad Entomologica Argentina, 70 (3 - 4), 213 - 229." type="journal article">Chani-Posse 2011b: 224</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFC97F5DBD638" blockId="29.[151,1436,835,933]" box="[151,654,835,861]" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFC97F5DBD638" ID-CoL="LCCL" authority="Bernhauer 1917" authorityName="Bernhauer" authorityYear="1917" box="[151,654,835,861]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xantholinoides">
<emphasis id="9B8CEACC4C35B355FF6BFC97F686D638" bold="true" box="[151,467,835,860]" italics="true" pageId="29" pageNumber="30">Belonuchus xantholinoides</emphasis>
Bernhauer 1917
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFCB3F4B9D6C1" blockId="29.[151,1436,835,933]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFCB3F4B9D6C1" pageId="29" pageNumber="30">
<materialsCitation id="19903C834C35B355FF6BFCB3F4B2D6C1" ID-GBIF-Occurrence="1915497855" collectionCode="FMNH" collectorName="Villa Elvira &amp; S. Antonio" country="Colombia" elevation="1800" location="ST. Distribution" pageId="29" pageNumber="30" specimenCode="FMNH 2" specimenCount="2" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C35B355FF6BFCB3F58BD6E4" ID-CoL="LCCL" authority="Bernhauer 1917 c: 106" authorityName="Bernhauer" authorityPageNumber="106" authorityYear="1917" box="[151,734,871,896]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xantholinoides">
<emphasis id="9B8CEACC4C35B355FF6BFCB3F69ED6E4" box="[151,459,871,896]" italics="true" pageId="29" pageNumber="30">Belonuchus xantholinoides</emphasis>
<treatmentCitation id="285910CF4C35B355FE2BFCB3F58BD6E4" author="" box="[471,734,871,896]" page="106" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE2BFCB3F5F4D6E4" box="[471,673,871,896]" pageId="29" pageNumber="30" refId="ref62261" refString="Bernhauer, M. (1917 c) Neue sudamerikanische Staphyliniden (18. Beitrag). Wiener Entomologische Zeitung, 36 (3 - 5), 102 - 116. https: // doi. org / 10.5962 / bhl. part. 12925" type="journal article">Bernhauer 1917c</bibRefCitation>
: 106
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C35B355FD12FCBCF473D6E4" box="[750,806,872,896]" pageId="29" pageNumber="30">Type</typeStatus>
localities: 
<collectorName id="040D53084C35B355FC50FCB3F366D6E4" box="[940,1075,871,896]" pageId="29" pageNumber="30">Villa Elvira</collectorName>
<elevation id="22D5D1ED4C35B355FBBCFCBCF3DBD6E4" box="[1088,1166,872,896]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">
<quantity id="6E009B3B4C35B355FBBCFCBCF3DBD6E4" box="[1088,1166,872,896]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">1800m</quantity>
</elevation>
&amp; 
<collectorName id="040D53084C35B355FB40FCBCF26ED6E4" box="[1212,1339,871,896]" pageId="29" pageNumber="30">S. Antonio</collectorName>
<elevation id="22D5D1ED4C35B355FABBFCBCF2CDD6E4" box="[1351,1432,872,896]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">
<quantity id="6E009B3B4C35B355FABBFCBCF2CDD6E4" box="[1351,1432,872,896]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">2200m</quantity>
</elevation>
, Westkolumbien. 
<typeStatus id="7643887C4C35B355FE7EFC59F6ECD6C1" box="[386,441,909,933]" pageId="29" pageNumber="30">Type</typeStatus>
depository: 
<specimenCode id="F95E9EA54C35B355FDBAFC59F5FED6C0" box="[582,683,908,932]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="29" pageNumber="30">FMNH 2</specimenCode>
<location id="AC2760054C35B355FD4EFC58F43ED6C1" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C35B355FD4EFC58F43ED6C1" box="[690,875,908,933]" country="Colombia" name="ST. Distribution" pageId="29" pageNumber="30">ST. Distribution</location>
: 
<collectingCountry id="D1EF764E4C35B355FC89FC58F4B2D6C1" box="[885,999,908,933]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
</materialsCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFC07F509D689" blockId="29.[151,1436,979,1149]" box="[151,604,979,1005]" pageId="29" pageNumber="30">
<emphasis id="9B8CEACC4C35B355FF6BFC07F7F6D688" bold="true" box="[151,163,979,1004]" italics="true" pageId="29" pageNumber="30">*</emphasis>
<taxonomicName id="6EF84D5D4C35B355FF5EFC07F509D689" ID-CoL="LCCM" authority="Solsky 1868" authorityName="Solsky" authorityYear="1868" box="[162,604,979,1005]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xanthomelas">
<emphasis id="9B8CEACC4C35B355FF5EFC07F69FD688" bold="true" box="[162,458,979,1004]" italics="true" pageId="29" pageNumber="30">Belonuchus xanthomelas</emphasis>
<bibRefCitation id="CD694B2F4C35B355FE2DFC00F509D689" box="[465,604,980,1005]" pageId="29" pageNumber="30" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFC23F65BD119" blockId="29.[151,1436,979,1149]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFC23F65BD119" infered="true" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFC23F5DFD174" ID-CoL="LCCM" authority="Solsky 1868: 139" authorityName="Solsky" authorityPageNumber="139" authorityYear="1868" box="[151,650,1015,1040]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xanthomelas">
<emphasis id="9B8CEACC4C35B355FF6BFC23F6E4D174" box="[151,433,1015,1040]" italics="true" pageId="29" pageNumber="30">Belonuchus xanthomelas</emphasis>
<treatmentCitation id="285910CF4C35B355FE40FC23F5DFD174" author="" box="[444,650,1015,1040]" page="139" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE40FC23F51BD174" box="[444,590,1015,1040]" pageId="29" pageNumber="30" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
: 139
</treatmentCitation>
</taxonomicName>
. Type locality: [Mexique et probablement de l'Etat d'Oaxaca]. Type depository: ZIN? (Solsky) #?ST. Distribution: USA (TX), Mexico. Note: generic placement (Navarrete- 
<bibRefCitation id="CD694B2F4C35B355FF39FBEBF635D13C" box="[197,352,1087,1112]" pageId="29" pageNumber="30" refId="ref69886" refString="Navarrete-Heredia, J. L. (1995) Aspectos biologicos de Philonthus apiciventris y P. oxyporinus (Co 1 eoptera: Staphylinidae: Staphylininae) en una zona de Morelos, Mexico, con una lista de las especies mexicanas de Philonthus. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Serie Zoologia, 66 (1), 81 - 106." type="journal article">Heredia 1995</bibRefCitation>
); distribution, also within Mexico (
<bibRefCitation id="CD694B2F4C35B355FD14FBEBF36FD13C" box="[744,1082,1087,1112]" pageId="29" pageNumber="30" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C35B355FC3DFB95F4ACD13C" box="[961,1017,1087,1112]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2002
</bibRefCitation>
; Márquez-Luna 2001, 
<bibRefCitation id="CD694B2F4C35B355FAC4FB94F657D119" pageId="29" pageNumber="30" refId="ref69151" refString="Marquez, J. (2006) Primeros registros estatales y datos de distribucion geografica de especies mexicanas de Staphylinidae (Coleoptera). Boletin de la Sociedad Entomologica Aragonesa, 38, 181 - 198." type="journal article">Márquez 2006</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFB7FF5CAD1A0" blockId="29.[151,1437,1195,1328]" box="[151,671,1195,1221]" pageId="29" pageNumber="30">
<emphasis id="9B8CEACC4C35B355FF6BFB7FF7F6D1A0" bold="true" box="[151,163,1195,1220]" italics="true" pageId="29" pageNumber="30">*</emphasis>
<taxonomicName id="6EF84D5D4C35B355FF5EFB7FF5CAD1A0" ID-CoL="LCCN" authority="(Nordmann 1837)" authorityName="Nordmann" authorityYear="1837" baseAuthorityName="Nordmann" baseAuthorityYear="1837" box="[162,671,1195,1221]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xanthopterus">
<emphasis id="9B8CEACC4C35B355FF5EFB7FF69AD1A0" bold="true" box="[162,463,1195,1220]" italics="true" pageId="29" pageNumber="30">Belonuchus xanthopterus</emphasis>
(
<bibRefCitation id="CD694B2F4C35B355FE21FB78F5C2D1A0" box="[477,663,1196,1221]" pageId="29" pageNumber="30" refId="ref70688" refString="Nordmann, A. (1837) Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum, 4, 1 - 167. https: // doi. org / 10.5962 / bhl. title. 144129" type="journal article">Nordmann 1837</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFB1BF3A1D054" blockId="29.[151,1437,1195,1328]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFB1BF3A1D054" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BFB1BF5F9D18C" ID-CoL="4GC5L" authority="Nordmann 1837: 99" authorityName="Nordmann" authorityPageNumber="99" authorityYear="1837" box="[151,684,1231,1256]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xanthopterus">
<emphasis id="9B8CEACC4C35B355FF6BFB1BF6FAD18C" box="[151,431,1231,1256]" italics="true" pageId="29" pageNumber="30">Philonthus xanthopterus</emphasis>
<treatmentCitation id="285910CF4C35B355FE40FB1BF5F9D18C" author="" box="[444,684,1231,1256]" page="99" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE40FB1BF528D18C" box="[444,637,1231,1256]" pageId="29" pageNumber="30" refId="ref70688" refString="Nordmann, A. (1837) Symbolae ad monographiam staphylinorum. Ex Academiae Caesareae Scientiarum, 4, 1 - 167. https: // doi. org / 10.5962 / bhl. title. 144129" type="journal article">Nordmann 1837</bibRefCitation>
: 99
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C35B355FD41FB04F5A0D18C" box="[701,757,1232,1256]" pageId="29" pageNumber="30">Type</typeStatus>
locality: Parae in America meridionali. 
<typeStatus id="7643887C4C35B355FB26FB04F244D18C" box="[1242,1297,1232,1256]" pageId="29" pageNumber="30">Type</typeStatus>
depository: ZMHB 1 ST 
<specimenCount id="BFFEFD574C35B355FEA3FB21F621D068" box="[351,372,1269,1292]" pageId="29" pageNumber="30" type="male">♂</specimenCount>
[MCP seen]. Distribution: 
<collectingCountry id="D1EF764E4C35B355FD50FB20F5A6D069" box="[684,755,1268,1293]" name="Brazil" pageId="29" pageNumber="30">Brazil</collectingCountry>
, 
<collectingCountry id="D1EF764E4C35B355FCFDFB21F462D068" box="[769,823,1269,1292]" name="Peru" pageId="29" pageNumber="30">Peru</collectingCountry>
, 
<collectingRegion id="6B3CF83C4C35B355FCB8FB20F4F7D068" box="[836,930,1268,1292]" country="Panama" name="Panama" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FCB8FB20F4F7D068" box="[836,930,1268,1292]" name="Panama" pageId="29" pageNumber="30">Panama</collectingCountry>
</collectingRegion>
?, 
<collectingRegion id="6B3CF83C4C35B355FC45FB20F349D069" box="[953,1052,1268,1293]" country="Democratic Republic of the Congo" name="Equateur" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FC45FB20F349D069" box="[953,1052,1268,1293]" name="Ecuador" pageId="29" pageNumber="30">Ecuador</collectingCountry>
</collectingRegion>
?. Note: comb. nov. ex 
<taxonomicName id="6EF84D5D4C35B355FADFFB20F6E3D054" ID-CoL="8VRYR" authority="(Erichson 1840: 421)" authorityName="Erichson" authorityPageNumber="421" authorityYear="1840" baseAuthorityName="Erichson" baseAuthorityYear="1840" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C35B355FADFFB20F2C9D069" box="[1315,1436,1268,1293]" italics="true" pageId="29" pageNumber="30">Philonthus</emphasis>
(
<treatmentCitation id="285910CF4C35B355FF30FAC3F6F8D054" author="" box="[204,429,1303,1328]" page="421" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FF30FAC3F6F8D054" box="[204,429,1303,1328]" pageId="29" pageNumber="30" refId="ref65498" refString="Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 554 pp. [pp. 401 - 954] https: // doi. org / 10.5962 / bhl. title. 59644" type="book chapter">Erichson 1840: 421</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
; 
<collectingCountry id="D1EF764E4C35B355FE3EFACCF6A0D04B" box="[450,501,1304,1327]" name="Peru" pageId="29" pageNumber="30">Peru</collectingCountry>
(
<treatmentCitation id="285910CF4C35B355FDFFFAC3F583D054" author="" box="[515,726,1303,1328]" page="177" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FDFFFAC3F583D054" box="[515,726,1303,1328]" pageId="29" pageNumber="30" refId="ref73180" refString="Soukup, J. (1942) Apuntes para la zoogeografia entomologica (Entomografia peruana). Boletin del Museo de Historia Natural &quot; Javier Prado &quot;, Lima, 6 (21), 172 - 192." type="journal article">Soukup 1942: 177</bibRefCitation>
</treatmentCitation>
); distribution within 
<collectingCountry id="D1EF764E4C35B355FC3DFAC3F350D054" box="[961,1029,1303,1328]" name="Brazil" pageId="29" pageNumber="30">Brazil</collectingCountry>
(Asenjo 
<emphasis id="9B8CEACC4C35B355FB90FACDF3F0D054" box="[1132,1189,1303,1328]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2013).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFA8BF517D01C" blockId="29.[151,1436,1375,1509]" box="[151,578,1375,1400]" pageId="29" pageNumber="30">
<emphasis id="9B8CEACC4C35B355FF6BFA8BF7F6D01C" bold="true" box="[151,163,1375,1400]" italics="true" pageId="29" pageNumber="30">*</emphasis>
<taxonomicName id="6EF84D5D4C35B355FF5EFA8BF517D01C" ID-CoL="LCCP" authority="Solsky 1868" authorityName="Solsky" authorityYear="1868" box="[162,578,1375,1400]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xanthopus">
<emphasis id="9B8CEACC4C35B355FF5EFA8BF6FAD01C" bold="true" box="[162,431,1375,1400]" italics="true" pageId="29" pageNumber="30">Belonuchus xanthopus</emphasis>
<bibRefCitation id="CD694B2F4C35B355FE4AFA8BF517D01C" box="[438,578,1375,1400]" pageId="29" pageNumber="30" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BFA50F434D080" blockId="29.[151,1436,1375,1509]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BFA50F434D080" infered="true" pageId="29" pageNumber="30">
<materialsCitation id="19903C834C35B355FF6BFA50F201D0F9" ID-GBIF-Occurrence="1915497818" box="[151,1364,1412,1437]" collectorName="de l'Etat d'Oaxaca" country="Mexico" location="Mexique" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<taxonomicName id="6EF84D5D4C35B355FF6BFA50F522D0F8" ID-CoL="LCCP" authority="Solsky 1868: 139" authorityName="Solsky" authorityPageNumber="139" authorityYear="1868" box="[151,631,1412,1437]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="xanthopus">
<emphasis id="9B8CEACC4C35B355FF6BFA50F6CCD0F9" box="[151,409,1412,1437]" italics="true" pageId="29" pageNumber="30">Belonuchus xanthopus</emphasis>
<treatmentCitation id="285910CF4C35B355FE59FA50F522D0F8" author="" box="[421,631,1412,1437]" page="139" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE59FA50F56FD0F9" box="[421,570,1412,1437]" pageId="29" pageNumber="30" refId="ref73057" refString="Solsky, S. (1868) Etudes sur les Staphylinides du Mexique. Horae Societatis Entomologicae Rossicae, 5, 119 - 144, pl. 4." type="journal article">Solsky 1868</bibRefCitation>
: 139
</treatmentCitation>
</taxonomicName>
. 
<typeStatus id="7643887C4C35B355FD7BFA51F5EAD0F9" box="[647,703,1413,1437]" pageId="29" pageNumber="30">Type</typeStatus>
locality: [
<collectingCountry id="D1EF764E4C35B355FCC6FA50F4F4D0F9" box="[826,929,1412,1437]" name="Mexico" pageId="29" pageNumber="30">Mexique</collectingCountry>
et probablement 
<collectorName id="040D53084C35B355FB88FA50F21AD0F9" box="[1140,1359,1412,1437]" pageId="29" pageNumber="30">de l'Etat d'Oaxaca</collectorName>
]
</materialsCitation>
. 
<materialsCitation id="19903C834C35B355FA98FA51F463D0A4" ID-GBIF-Occurrence="1915497757" collectionCode="ZIN" country="Mexico" location="Solsky" municipality="ST. Distribution" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<typeStatus id="7643887C4C35B355FA98FA51F2C9D0F9" box="[1380,1436,1413,1437]" pageId="29" pageNumber="30">Type</typeStatus>
depository: 
<collectionCode id="CFE9AE1B4C35B355FEB0FA7CF628D0A4" box="[332,381,1448,1472]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="29" pageNumber="30">ZIN</collectionCode>
? (
<location id="AC2760054C35B355FE65FA73F6BDD0A4" LSID="urn:lsid:plazi:treatment:215187C84C20B355FF6BFC00F4ACD3FC:AC2760054C35B355FE65FA73F6BDD0A4" box="[409,488,1447,1472]" country="Mexico" municipality="ST. Distribution" name="Solsky" pageId="29" pageNumber="30">Solsky</location>
) #? 
<collectingMunicipality id="4923ACA44C35B355FDE9FA7CF585D0A4" box="[533,720,1447,1472]" pageId="29" pageNumber="30">ST. Distribution</collectingMunicipality>
: 
<collectingCountry id="D1EF764E4C35B355FD21FA73F463D0A4" box="[733,822,1447,1472]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
</materialsCitation>
. Notes: generic placement (Navarrete-Heredia 1995); distribution within Mexico (
<bibRefCitation id="CD694B2F4C35B355FE03FA18F400D080" box="[511,853,1484,1509]" pageId="29" pageNumber="30" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C35B355FD25FA19F444D081" box="[729,785,1484,1509]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2002
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BF9C7F516D349" blockId="29.[151,1437,1555,1688]" box="[151,579,1555,1581]" pageId="29" pageNumber="30">
<emphasis id="9B8CEACC4C35B355FF6BF9C7F7F6D348" bold="true" box="[151,163,1555,1580]" italics="true" pageId="29" pageNumber="30">*</emphasis>
<taxonomicName id="6EF84D5D4C35B355FF5EF9C7F516D349" ID-CoL="LCCQ" authority="(Sharp 1885)" authorityName="Sharp" authorityYear="1885" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[162,579,1555,1581]" class="Insecta" family="Staphylinidae" genus="Belonuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="9B8CEACC4C35B355FF5EF9C7F6FCD348" bold="true" box="[162,425,1555,1580]" italics="true" pageId="29" pageNumber="30">Belonuchus zunilensis</emphasis>
(
<bibRefCitation id="CD694B2F4C35B355FE44F9C0F56ED349" box="[440,571,1556,1581]" pageId="29" pageNumber="30" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="A94736DE4C35B355FF6BF9E3F4ACD3FC" blockId="29.[151,1437,1555,1688]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="89E811F04C35B355FF6BF9E3F4ACD3FC" infered="true" pageId="29" pageNumber="30">
<taxonomicName id="6EF84D5D4C35B355FF6BF9E3F50CD334" ID-CoL="4GC64" authority="Sharp 1885: 418" authorityName="Sharp" authorityPageNumber="418" authorityYear="1885" box="[151,601,1591,1616]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="9B8CEACC4C35B355FF6BF9E3F6DCD334" box="[151,393,1591,1616]" italics="true" pageId="29" pageNumber="30">Philonthus zunilensis</emphasis>
<treatmentCitation id="285910CF4C35B355FE68F9E3F50CD334" author="" box="[404,601,1591,1616]" page="418" pageId="29" pageNumber="30">
<bibRefCitation id="CD694B2F4C35B355FE68F9E3F548D334" box="[404,541,1591,1616]" pageId="29" pageNumber="30" refId="ref72492" refString="Sharp, D. (1885) Staphylinidae. Biologia Centrali-Americana. Insecta. Coleoptera, 1 (2), 393 - 536. [http: // www. sil. si. edu / digitalcollections / bca /]" type="journal article">Sharp 1885</bibRefCitation>
: 418
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</taxonomicName>
. 
<typeStatus id="7643887C4C35B355FD94F9ECF5CAD334" box="[616,671,1592,1616]" pageId="29" pageNumber="30">Type</typeStatus>
locality: 
<collectingRegion id="6B3CF83C4C35B355FCEDF9E3F4DAD334" box="[785,911,1591,1616]" country="Guatemala" name="Guatemala" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FCEDF9E3F4DAD334" box="[785,911,1591,1616]" name="Guatemala" pageId="29" pageNumber="30">Guatemala</collectingCountry>
</collectingRegion>
: Cerro Zunil 
<quantity id="6E009B3B4C35B355FBCCF9E3F399D334" box="[1072,1228,1591,1616]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" metricValueMax="1.524" metricValueMin="1.2192" pageId="29" pageNumber="30" unit="ft" value="4500.0" valueMax="5000.0" valueMin="4000.0">4000–5000 ft</quantity>
. 
<typeStatus id="7643887C4C35B355FB27F9ECF246D334" box="[1243,1299,1592,1616]" pageId="29" pageNumber="30">Type</typeStatus>
depository: BMNH 4 ST. Distribution: 
<collectingRegion id="6B3CF83C4C35B355FDFBF988F50AD311" box="[519,607,1628,1653]" country="Mexico" name="Mexico" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FDFBF988F50AD311" box="[519,607,1628,1653]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
</collectingRegion>
, 
<collectingRegion id="6B3CF83C4C35B355FD93F988F5B9D311" box="[623,748,1628,1653]" country="Guatemala" name="Guatemala" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FD93F988F5B9D311" box="[623,748,1628,1653]" name="Guatemala" pageId="29" pageNumber="30">Guatemala</collectingCountry>
</collectingRegion>
. Notes: comb. nov. ex 
<taxonomicName id="6EF84D5D4C35B355FC02F988F322D311" ID-CoL="8VRYR" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[1022,1143,1628,1653]" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="9B8CEACC4C35B355FC02F988F322D311" box="[1022,1143,1628,1653]" italics="true" pageId="29" pageNumber="30">Philonthus</emphasis>
</taxonomicName>
, distribution, also within 
<collectingRegion id="6B3CF83C4C35B355FF39F9ABF64FD3FC" box="[197,282,1663,1688]" country="Mexico" name="Mexico" pageId="29" pageNumber="30">
<collectingCountry id="D1EF764E4C35B355FF39F9ABF64FD3FC" box="[197,282,1663,1688]" name="Mexico" pageId="29" pageNumber="30">Mexico</collectingCountry>
</collectingRegion>
(
<bibRefCitation id="CD694B2F4C35B355FED5F9ABF52BD3FC" box="[297,638,1663,1688]" pageId="29" pageNumber="30" refId="ref70158" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Illustrated guide to the genera of Staphylinidae (Coleoptera) of Mexico. Universidad de Guadalajara y Conabio, Mexico, 401 pp." type="book">
Navarrete-Heredia 
<emphasis id="9B8CEACC4C35B355FDFEF955F56FD3FC" box="[514,570,1663,1688]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2002
</bibRefCitation>
); distribution (
<bibRefCitation id="CD694B2F4C35B355FCDDF9ABF4BBD3FC" box="[801,1006,1663,1688]" pageId="29" pageNumber="30" refId="ref61503" refString="Asiain J., Marquez J. &amp; Bueno-Villegas, J. (2011) The Staphylinidae (Coleoptera) Fauna of Los Marmoles National Park, Hidalgo, Mexico. Coleopterists Bulletin, 65 (4), 393 - 402." type="journal article">
Asiain 
<emphasis id="9B8CEACC4C35B355FC8FF955F4FED3FC" box="[883,939,1663,1688]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2011
</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>