411 lines
45 KiB
XML
411 lines
45 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1153.99537" ID-Pensoft-Pub="1313-2970-1153-73" ID-Pensoft-UUID="9A8EFF777E2258F090F13B331AFDFAE2" ID-ZooBank="D9BFB5475F3F45F1A5563F05839B4413" ModsDocID="1313-2970-1153-73" checkinTime="1678945459464" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Feng, Zegang & Zhang, Feng" docDate="2023" docId="EA6A9783425E5DD59F949D1620536FEF" docLanguage="en" docName="ZooKeys 1153: 73-95" docOrigin="ZooKeys 1153" docPubDate="2023-03-15" docSource="http://dx.doi.org/10.3897/zookeys.1153.99537" docTitle="Tyrannochthonius calvatus Hou & Feng & Zhang 2023, sp. nov." docType="treatment" docUuid="789C8A08-776E-47A8-BCC3-B73E4940FB3D" docUuidSource="ZooBank" docVersion="1" id="9A8EFF777E2258F090F13B331AFDFAE2" lastPageNumber="73" masterDocId="9A8EFF777E2258F090F13B331AFDFAE2" masterDocTitle="Three new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from eastern Yunnan, China" masterLastPageNumber="95" masterPageNumber="73" pageNumber="73" updateTime="1678945459464" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Three new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from eastern Yunnan, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hou, Yanmeng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0059-3419</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Feng, Zegang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5809-5984</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China & Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Feng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier type="email">dudu06042001@163.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-15</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1153</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>73</mods:start>
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<mods:end>95</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1153.99537</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1153.99537</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1153-73</mods:identifier>
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<mods:identifier type="ZooBank">D9BFB5475F3F45F1A5563F05839B4413</mods:identifier>
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<mods:identifier type="Pensoft-UUID">9A8EFF777E2258F090F13B331AFDFAE2</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:789C8A08-776E-47A8-BCC3-B73E4940FB3D" httpUri="http://treatment.plazi.org/id/EA6A9783425E5DD59F949D1620536FEF" lastPageNumber="73" pageId="0" pageNumber="73">
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<subSubSection pageId="0" pageNumber="73" type="nomenclature">
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<paragraph pageId="0" pageNumber="73">
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<taxonomicName LSID="https://zoobank.org/789C8A08-776E-47A8-BCC3-B73E4940FB3D" authority="Hou & Feng & Zhang, 2023" authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius calvatus" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="calvatus" status="sp. nov.">Tyrannochthonius calvatus</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="73">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="description">
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<paragraph pageId="0" pageNumber="73">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Unnamed cave (type locality) and Dongtianfu Cave, habitats of Tyrannochthonius calvatus sp. nov. A unnamed cave B Dongtianfu Cave C live female of T. calvatus sp. nov. in its natural environment from Dongtianfu Cave." figureDoi="10.3897/zookeys.1153.99537.figure2" httpUri="https://binary.pensoft.net/fig/825475" pageId="0" pageNumber="73">Figs 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius calvatus sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1153.99537.figure3" httpUri="https://binary.pensoft.net/fig/825476" pageId="0" pageNumber="73">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">
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, 5 Chinese name:
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<normalizedToken originalValue="秃头暴伪蝎">秃头暴伪蝎</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="type material">
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<paragraph pageId="0" pageNumber="73">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
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: China • ♂; Yunnan Province, Fuyuan County, Mohong Town, Puchong Village, unnamed cave;
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<geoCoordinate degrees="25" direction="north" minutes="22.301" orientation="latitude" precision="1" value="25.371683">25°22.301'N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="6.380" orientation="longitude" precision="1" value="104.10633">104°6.380'E</geoCoordinate>
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; 2060 m a.s.l.; 07 Oct. 2021; Zegang Feng, Yanmeng Hou, Lu Zhang and Liu Fu leg.; under a stone in the dark zone; Ps.-MHBU-HBUARA#2021-429-01 (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area, general cave locations, and type locality for each species, Yunnan Province, China. Each color represents an administrative region (green: Yunnan Province; red: Fuyuan County) A Daidai Cave (Lagynochthonius daidaiensis sp. nov.) B Xianren Cave (Tyrannochthonius capito sp. nov.) C, D unnamed cave and Dongtianfu Cave (T. calvatus sp. nov.)." figureDoi="10.3897/zookeys.1153.99537.figure1" httpUri="https://binary.pensoft.net/fig/825474" pageId="0" pageNumber="73">1C</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Unnamed cave (type locality) and Dongtianfu Cave, habitats of Tyrannochthonius calvatus sp. nov. A unnamed cave B Dongtianfu Cave C live female of T. calvatus sp. nov. in its natural environment from Dongtianfu Cave." figureDoi="10.3897/zookeys.1153.99537.figure2" httpUri="https://binary.pensoft.net/fig/825475" pageId="0" pageNumber="73">2A</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Paratypes</emphasis>
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: • 4 ♂; the same data as the holotype; Ps.-MHBU-HBUARA#2021-429-02-HBUARA#2021-429-05 • 1 ♂, 2 ♀; the same collection date and collectors as the holotype; Puchong Village, Dongtianfu Cave;
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<geoCoordinate degrees="25" direction="north" minutes="22.105" orientation="latitude" precision="1" value="25.368416">25°22.105'N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="6.447" orientation="longitude" precision="1" value="104.10745">104°6.447'E</geoCoordinate>
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; 2035 m a.s.l.; under stones and clods in the dark zone; Ps.-MSWU-HBUARA#2021-428-01-HBUARA#2021-428-03 (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area, general cave locations, and type locality for each species, Yunnan Province, China. Each color represents an administrative region (green: Yunnan Province; red: Fuyuan County) A Daidai Cave (Lagynochthonius daidaiensis sp. nov.) B Xianren Cave (Tyrannochthonius capito sp. nov.) C, D unnamed cave and Dongtianfu Cave (T. calvatus sp. nov.)." figureDoi="10.3897/zookeys.1153.99537.figure1" httpUri="https://binary.pensoft.net/fig/825474" pageId="0" pageNumber="73">1D</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Unnamed cave (type locality) and Dongtianfu Cave, habitats of Tyrannochthonius calvatus sp. nov. A unnamed cave B Dongtianfu Cave C live female of T. calvatus sp. nov. in its natural environment from Dongtianfu Cave." figureDoi="10.3897/zookeys.1153.99537.figure2" httpUri="https://binary.pensoft.net/fig/825475" pageId="0" pageNumber="73">2B</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1153.99537.figure1" httpUri="https://binary.pensoft.net/fig/825474" pageId="0" pageNumber="73" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="73">
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<emphasis bold="true" pageId="0" pageNumber="73">Figure 1.</emphasis>
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Study area, general cave locations, and type locality for each species, Yunnan Province, China. Each color represents an administrative region (green: Yunnan Province; red: Fuyuan County)
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<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
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Daidai Cave (
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<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
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<emphasis italics="true" pageId="0" pageNumber="73">Lagynochthonius daidaiensis</emphasis>
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</taxonomicName>
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sp. nov.)
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<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
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Xianren Cave (
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<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius capito" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="capito">
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<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius capito</emphasis>
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</taxonomicName>
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sp. nov.)
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<emphasis bold="true" pageId="0" pageNumber="73">C, D</emphasis>
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unnamed cave and Dongtianfu Cave (
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<taxonomicName lsidName="T. calvatus" pageId="0" pageNumber="73" rank="species" species="calvatus">
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<emphasis italics="true" pageId="0" pageNumber="73">T. calvatus</emphasis>
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</taxonomicName>
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sp. nov.).
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1153.99537.figure2" httpUri="https://binary.pensoft.net/fig/825475" pageId="0" pageNumber="73" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="73">
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<emphasis bold="true" pageId="0" pageNumber="73">Figure 2.</emphasis>
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Unnamed cave (type locality) and Dongtianfu Cave, habitats of
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<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius calvatus" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="calvatus">
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<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius calvatus</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
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unnamed cave
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<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
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Dongtianfu Cave
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<emphasis bold="true" pageId="0" pageNumber="73">C</emphasis>
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live female of
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<taxonomicName lsidName="T. calvatus" pageId="0" pageNumber="73" rank="species" species="calvatus">
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<emphasis italics="true" pageId="0" pageNumber="73">T. calvatus</emphasis>
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</taxonomicName>
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sp. nov. in its natural environment from Dongtianfu Cave.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="diagnosis">
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<paragraph pageId="0" pageNumber="73">Diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="73">
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<emphasis bold="true" pageId="0" pageNumber="73">(♂♀).</emphasis>
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Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, with four setae (including preocular setae) only, without epistome and flanking basal setae; posterior margin of carapace with two setae; tergites I-III each with two setae. Pedipalps slender, femur 8.06-8.44 (♂), 7.94-8.38 (♀)
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than broad; chela 8.05-8.33 (♂), 8.05-8.71 (♀)
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than deep only movable chelal finger with intercalary teeth; movable chelal finger teeth markedly smaller than fixed chelal finger teeth and strongly retrorse and contiguous.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="etymology">
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<paragraph pageId="0" pageNumber="73">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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The specific name is derived from the Latin adjective
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<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius calvatus" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="calvatus">
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<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius calvatus</emphasis>
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</taxonomicName>
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(bald) and refers to the absence of two antero-median setae on the carapace.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="description">
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<paragraph pageId="0" pageNumber="73">Description.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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<emphasis bold="true" pageId="0" pageNumber="73">Adult males</emphasis>
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(Figs
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius calvatus sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1153.99537.figure3" httpUri="https://binary.pensoft.net/fig/825476" pageId="0" pageNumber="73">3A</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4A-F</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Color</emphasis>
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: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Cephalothorax</emphasis>
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(Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4C</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5A</figureCitation>
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): carapace 0.98
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome absent, without two setae flanking base; with 14-15 setae arranged s2s:4:4:0-1:2 (with 14-17 setae arranged s2s:4:4:0-3:2 in females), most setae heavy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, the first two pairs situated middle and lateral to the setae of ocular row respectively, the third situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3-4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 12 or 13 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ca. half their length (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5C</figureCitation>
|
||
); intercoxal tubercle absent; without sub-oral seta.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Chelicera</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5B, D</figureCitation>
|
||
): large, ca. as long as carapace, 2.43-2.48
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 14-16 teeth, distal one largest; movable finger with 13-15 retrorse contiguous small teeth; galea absent (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5B</figureCitation>
|
||
). Serrula exterior with 25-28 blades and serrula interior with 17 or 18 blades. Rallum with eight blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5D</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4A, B, E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5E-G</figureCitation>
|
||
): long and slender, trochanter 1.74-1.88, femur 8.06-8.44, patella 2.37-2.61, chela 8.05-8.33, hand 3.00-3.10
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep femur 2.87
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than patella; movable chelal finger 1.64-1.66
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than hand and 0.61-0.62
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5E</figureCitation>
|
||
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
|
||
<emphasis italics="true" pageId="0" pageNumber="73">ib</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="73">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis italics="true" pageId="0" pageNumber="73">eb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="73">esb</emphasis>
|
||
, and
|
||
<emphasis italics="true" pageId="0" pageNumber="73">ist</emphasis>
|
||
forming a nearly straight oblique row at base of fixed chelal finger;
|
||
<emphasis italics="true" pageId="0" pageNumber="73">it</emphasis>
|
||
slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="73">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis italics="true" pageId="0" pageNumber="73">et</emphasis>
|
||
slightly near to tip of fixed chelal finger, very close to chelal teeth;
|
||
<emphasis italics="true" pageId="0" pageNumber="73">dx</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="73">et</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="73">sb</emphasis>
|
||
situated midway between
|
||
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="73">st</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="73">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis italics="true" pageId="0" pageNumber="73">t</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
|
||
and proximal to
|
||
<emphasis italics="true" pageId="0" pageNumber="73">est</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5F</figureCitation>
|
||
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="73">ist</emphasis>
|
||
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 32-36 macrodenticles, long and pointed, without intercalary teeth; movable chelal finger with 37-41 macrodenticles (smaller than teeth on fixed chelal finger), retrorse and contiguous, plus 10-12 intercalary microdenticles (roughly extending to
|
||
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
|
||
), 47-53 in total (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5F</figureCitation>
|
||
). Chelal fingers straight in dorsal view; microsetae (
|
||
<emphasis italics="true" pageId="0" pageNumber="73">sc</emphasis>
|
||
) present on dorsum of chelal hand (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5G</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Opisthosoma</emphasis>
|
||
: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2:2:2:2:4:4:4:4:4-5:4:T2-3T:0; tergite IX with an unpaired median seta. Sternal chaetotaxy III-XII: 12-14:11-12:8:9:9:9-10:9:7-9:0: 2. Anterior genital operculum with ten setae, genital opening slit-like, with 10-12 marginal setae on each side, 31-33 in total (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4F</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5H</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tyrannochthonius calvatus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73">5I</figureCitation>
|
||
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.88
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than patella and with one lyrifissure at the base of femur; tarsus 2.45
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than tibia. Femoropatella of leg IV 4.83-5.04
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep; tibia 7.60
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep; with basal tactile setae on both tarsal segments: basitarsus 4.00-4.25
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.24-0.28), telotarsus 15.33-15.50
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.71-2.91
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.27-0.28). Arolium slightly shorter than the claws, not divided; claws simple.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1153.99537.figure3" httpUri="https://binary.pensoft.net/fig/825476" pageId="0" pageNumber="73" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<emphasis bold="true" pageId="0" pageNumber="73">Figure 3.</emphasis>
|
||
<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius calvatus" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="calvatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius calvatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
|
||
holotype male, habitus (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
|
||
paratype female, habitus (dorsal view). Scale bars: 0.50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<emphasis bold="true" pageId="0" pageNumber="73">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius calvatus" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="calvatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius calvatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis bold="true" pageId="0" pageNumber="73">A-F</emphasis>
|
||
), paratype female (
|
||
<emphasis bold="true" pageId="0" pageNumber="73">G</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
|
||
left chela (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
|
||
left chela (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">C</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">D</emphasis>
|
||
left chelicera (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">E</emphasis>
|
||
left pedipalp (minus chela, dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">F</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">G</emphasis>
|
||
female genital area (ventral view). Scale bars: 0.25 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="73">E</emphasis>
|
||
); 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="73">A, B</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="73">C, D, F, G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1153.99537.figure5" httpUri="https://binary.pensoft.net/fig/825478" pageId="0" pageNumber="73" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<emphasis bold="true" pageId="0" pageNumber="73">Figure 5.</emphasis>
|
||
<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius calvatus" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="calvatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius calvatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
|
||
left chelicera (dorsal view) with details of dentation
|
||
<emphasis bold="true" pageId="0" pageNumber="73">C</emphasis>
|
||
coxal spines on coxae II (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">D</emphasis>
|
||
rallum
|
||
<emphasis bold="true" pageId="0" pageNumber="73">E</emphasis>
|
||
left pedipalp (minus chela, dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">F</emphasis>
|
||
left chela (lateral view) with details of dentation and trichobothrial pattern
|
||
<emphasis bold="true" pageId="0" pageNumber="73">G</emphasis>
|
||
left chela (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">H</emphasis>
|
||
leg I (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="73">I</emphasis>
|
||
leg IV (lateral view). Scale bars: 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="73">A, B, E-I</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="73">C, D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<emphasis bold="true" pageId="0" pageNumber="73">Adult females</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Unnamed cave (type locality) and Dongtianfu Cave, habitats of Tyrannochthonius calvatus sp. nov. A unnamed cave B Dongtianfu Cave C live female of T. calvatus sp. nov. in its natural environment from Dongtianfu Cave." figureDoi="10.3897/zookeys.1153.99537.figure2" httpUri="https://binary.pensoft.net/fig/825475" pageId="0" pageNumber="73">2C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius calvatus sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1153.99537.figure3" httpUri="https://binary.pensoft.net/fig/825476" pageId="0" pageNumber="73">3B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tyrannochthonius calvatus sp. nov., holotype male (A-F), paratype female (G) A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E left pedipalp (minus chela, dorsal view) F male genital area (ventral view) G female genital area (ventral view). Scale bars: 0.25 mm (E); 0.20 mm (A, B); 0.10 mm (C, D, F, G)." figureDoi="10.3897/zookeys.1153.99537.figure4" httpUri="https://binary.pensoft.net/fig/825477" pageId="0" pageNumber="73">4G</figureCitation>
|
||
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; tergal chaetotaxy I-XII: 2:2:2:2-3:4:4:3-4:4:4-5:4:T2T:0; sternal chaetotaxy IV-XII: 10-12:9-10:7-10:9:9:9:6-8:0:2; anterior genital operculum with 10-13 setae, posterior margin with 10-14 marginal setae, 22-24 in total; leg IV with a long tactile seta on both tarsal segments: basitarsus 3.56-5.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.23-0.25), telotarsus 14.00-16.17
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.77-2.88
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.27-0.29).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Dimensions</emphasis>
|
||
(length/breadth or, in the case of the legs, also for chela and hand, length/depth in mm). Males (females in parentheses): body length 2.16-2.21 (2.13-2.26). Pedipalps: trochanter 0.32-0.33/0.17-0.19 (0.32-0.34/0.18-0.20), femur 1.29-1.35/0.16 (1.34-1.37/0.16-0.17), patella 0.45-0.47/0.18-0.19 (0.49-0.52/0.18-0.20), chela 1.75-1.77/0.21-0.22 (1.77-1.84/0.21-0.22), hand 0.65-0.66/0.21-0.22 (0.65-0.68/0.21-0.22), movable chelal finger length 1.08 (1.11-1.16). Chelicera 0.72-0.73/0.29-0.30 (0.73-0.76/0.29-0.32), movable finger length 0.40 (0.40-0.41). Carapace 0.59/0.59-0.60 (0.59-0.65/0.62-0.66). Leg I: trochanter 0.21/0.13-0.14 (0.20-0.22/0.12-0.16), femur 0.75-0.77/0.09 (0.77-0.82/0.08-0.10), patella 0.40-0.41/0.08 (0.41-0.44/0.07-0.09), tibia 0.33/0.07 (0.34-0.37/0.06-0.07), tarsus 0.81/0.06 (0.79-0.82/0.06-0.07). Leg IV: trochanter 0.29-0.32/0.18 (0.28-0.30/0.15-0.18), femoropatella 1.11-1.16/0.23 (1.11-1.15/0.20-0.24), tibia 0.76/0.10 (0.77-0.81/0.10), basitarsus 0.32-0.34/0.08 (0.32-0.35/0.07-0.09), telotarsus 0.92-0.93/0.06 (0.92-0.98/0.06-0.07).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="remarks">
|
||
<paragraph pageId="0" pageNumber="73">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">
|
||
<taxonomicName authorityName="Hou & Feng & Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius calvatus" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="calvatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius calvatus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be easily distinguished from other Chinese cave-dwelling
|
||
<taxonomicName authorityName="Chamberlin" authorityYear="1929" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="73">Tyrannochthonius</emphasis>
|
||
</taxonomicName>
|
||
species by lacking two carapaceal antero-median setae and the presence of intercalary teeth on the movable chelal finger only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="73" type="distribution">
|
||
<paragraph pageId="0" pageNumber="73">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="73">Known only from the unnamed cave (type locality) and Dongtianfu Cave.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |