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<mods:title id="E346FC5D72C16846D9689B276FCB5B17">Systematics and biogeography of the ant genus Crematogaster Lund subgenus Orthocrema Santschi in Asia (Hymenoptera: Formicidae)</mods:title>
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<treatment id="1E64E12B3E63FFAD7FB0BF1ADC2EFDE2" ID-DOI="http://doi.org/10.5281/zenodo.7015406" ID-GBIF-Taxon="199534313" ID-Zenodo-Dep="7015406" LSID="urn:lsid:plazi:treatment:1E64E12B3E63FFAD7FB0BF1ADC2EFDE2" httpUri="http://treatment.plazi.org/id/1E64E12B3E63FFAD7FB0BF1ADC2EFDE2" lastPageId="29" lastPageNumber="576" pageId="28" pageNumber="575">
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<paragraph id="9672503D3E63FFAC7FB0BF1AD9D0FAFF" blockId="28.[187,715,1281,1334]" pageId="28" pageNumber="575">
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<taxonomicName id="51CD2BBE3E63FFAC7FB0BF1ADA1FFADF" authorityName="Forel" authorityYear="1913" box="[187,560,1281,1302]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="574" phylum="Arthropoda" rank="species" species="bandarensis">
<emphasis id="A4B98C2F3E63FFAC7FB0BF1ADA1FFADF" bold="true" box="[187,560,1281,1302]" italics="true" pageId="28" pageNumber="575">CREMATOGASTER BANDARENSIS</emphasis>
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<bibRefCitation id="F25C2DCC3E63FFAC7D32BF19DAE4FADF" author="Forel A" box="[569,715,1281,1303]" pageId="28" pageNumber="575" pagination="1 - 148" refId="ref39055" refString="Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien, ausgefuhrt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. 2. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel- Reepen in den Jahren 1911 - 1912. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere 36: 1 - 148." type="journal article" year="1913">FOREL, 1913</bibRefCitation>
<emphasis id="A4B98C2F3E63FFAC7E8CBF3ED9D0FAFF" bold="true" box="[391,511,1313,1334]" pageId="28" pageNumber="575">STAT. NOV.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E63FFAC7E9ABF53D9DAFA97" box="[401,501,1352,1374]" pageId="28" pageNumber="575" type="description">
<paragraph id="9672503D3E63FFAC7E9ABF53D9DAFA97" blockId="28.[401,501,1352,1374]" box="[401,501,1352,1374]" pageId="28" pageNumber="575">
(
<figureCitation id="0EF64CB83E63FFAC7E92BF53D9C2FA97" box="[409,493,1352,1374]" captionStart="Figures 96103" captionStartId="27.[164,254,1716,1735]" captionTargetBox="[164,1444,193,1686]" captionTargetId="figure-0@27.[164,1444,193,1686]" captionTargetPageId="27" captionText="Figures 96103. Body in lateral view. 96, Crematogaster baduvi, worker [Endau Rompin National Park, W. Malaysia]; 97, Crematogaster bandarensis, worker [Tai Rom Yen National Park, Surat Thani Prov., S. Thailand]; 98, Crematogaster binghamii, worker [Dry evergreen forest, Lumchangwat Stn., Khao Ang Reu Nai WS, Chacheongsap Prov., E. Thailand]; 99, Crematogaster biroi, worker [Punjavi Univ. Campus, Patiala, Punjab Prov., India]; 100, Crematogaster biroi, intermediate worker [Punjavi Univ. Campus, Patiala, Punjab Prov., India]; 101, Crematogaster brevispina, worker [Tabuc- Tubig, Oriental Dumaguete, Negros, Philippines]; 102, Crematogaster brunensis, worker [Bukit Sulang, nr. Lamunin, Brunei]; 103, Crematogaster celebensis, worker [Kadokaan, P. Batudaka, Togian Island, Sulawesi, Indonesia]." figureDoi="http://doi.org/10.5281/zenodo.5358220" httpUri="https://zenodo.org/record/5358220/files/figure.png" pageId="28" pageNumber="575">FIG. 97</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E63FFAC7F9BBF74DAD9FA4D" box="[144,758,1390,1412]" pageId="28" pageNumber="575" type="reference_group">
<paragraph id="9672503D3E63FFAC7F9BBF74DAD9FA4D" blockId="28.[144,759,1390,1779]" box="[144,758,1390,1412]" pageId="28" pageNumber="575">
<treatmentCitationGroup id="B6DD77133E63FFAC7F9BBF74DAD9FA4D" box="[144,758,1390,1412]" pageId="28" pageNumber="575">
<treatmentCitation id="176C762C3E63FFAC7F9BBF74DADDFA4D" author="Forel A" box="[144,754,1390,1412]" page="76" pageId="28" pageNumber="575" year="1913">
<taxonomicName id="51CD2BBE3E63FFAC7F9BBF74DADDFA4D" ID-CoL="5GNKP" authority="Forel, 1913: 76" authorityName="Forel" authorityPageNumber="76" authorityYear="1913" box="[144,754,1390,1412]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="575" phylum="Arthropoda" rank="variety" species="biroi" variety="bandarensis">
<emphasis id="A4B98C2F3E63FFAC7F9BBF74D95CFA4A" box="[144,371,1390,1412]" italics="true" pageId="28" pageNumber="575">Crematogaster biroi</emphasis>
var.
<emphasis id="A4B98C2F3E63FFAC7EA4BF75DA15FA4A" box="[431,570,1390,1411]" italics="true" pageId="28" pageNumber="575">bandarensis</emphasis>
<bibRefCitation id="F25C2DCC3E63FFAC7D48BF75DADDFA4D" author="Forel A" box="[579,754,1390,1412]" pageId="28" pageNumber="575" pagination="1 - 148" refId="ref39055" refString="Forel A. 1913. Wissenschaftliche Ergebnisse einer Forschungsreise nach Ostindien, ausgefuhrt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von H. v. Buttel-Reepen. 2. Ameisen aus Sumatra, Java, Malacca und Ceylon. Gesammelt von Herrn Prof. Dr. v. Buttel- Reepen in den Jahren 1911 - 1912. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere 36: 1 - 148." type="journal article" year="1913">Forel, 1913: 76</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E63FFAC7F9BBF96D9EDFA37" pageId="28" pageNumber="575" type="materials_examined">
<paragraph id="9672503D3E63FFAC7F9BBF96D9EDFA37" blockId="28.[144,759,1390,1779]" pageId="28" pageNumber="575">
<materialsCitation id="26A55A603E63FFAC7F9BBF96DA7AFA08" ID-GBIF-Occurrence="3891597470" collectorName="v. Buttel-Reepen" country="Indonesia" location="Bandar Baroe" pageId="28" pageNumber="575" specimenCount="1" specimenCount-worker="1" stateProvince="Sumatra" typeStatus="syntype">
<typeStatus id="4976EE9F3E63FFAC7F9BBF96D8C6FA6B" box="[144,233,1421,1442]" pageId="28" pageNumber="575" type="syntype">syntype</typeStatus>
<specimenCount id="80CB9BB43E63FFAC7FF9BF96D97FFA6A" box="[242,336,1421,1443]" pageId="28" pageNumber="575" type="worker">workers</specimenCount>
,
<location id="931206E63E63FFAC7E54BF96DA2BFA6B" LSID="urn:lsid:plazi:treatment:1E64E12B3E63FFAD7FB0BF1ADC2EFDE2:931206E63E63FFAC7E54BF96DA2BFA6B" box="[351,516,1421,1443]" country="Indonesia" name="Bandar Baroe" pageId="28" pageNumber="575" stateProvince="Sumatra">Bandar Baroe</location>
,
<collectingRegion id="54099EDF3E63FFAC7D19BF96DA56FA6B" box="[530,633,1421,1442]" country="Indonesia" name="Sumatera Utara" pageId="28" pageNumber="575">Sumatra</collectingRegion>
,
<collectingCountry id="EEDA10AD3E63FFAC7D8CBF96DAD8FA6A" box="[647,759,1421,1443]" name="Indonesia" pageId="28" pageNumber="575">Indonesia</collectingCountry>
(
<collectorName id="3B3835EB3E63FFAC7F93BFB7D975FA08" box="[152,346,1451,1473]" pageId="28" pageNumber="575">v. Buttel-Reepen</collectorName>
) (MHNG, examined)
</materialsCitation>
.
<materialsCitation id="26A55A603E63FFAC7D68BFB7D9EDFA37" ID-GBIF-Occurrence="3891597386" collectionCode="MHNG" pageId="28" pageNumber="575" specimenCount="1" specimenCount-worker="1" typeStatus="lectotype">
One syntype
<specimenCount id="80CB9BB43E63FFAC7F9BBFD1D8F2FA29" box="[144,221,1482,1504]" pageId="28" pageNumber="575" type="worker">worker</specimenCount>
(middle specimen of three on one pin) in
<collectionCode id="F0DCC8F83E63FFAC7DAEBFD1DAD9FA16" box="[677,758,1482,1503]" pageId="28" pageNumber="575">MHNG</collectionCode>
here designated
<emphasis id="A4B98C2F3E63FFAC7E5ABFF2D995FA37" box="[337,442,1513,1534]" italics="true" pageId="28" pageNumber="575">
<typeStatus id="4976EE9F3E63FFAC7E5ABFF2D995FA37" box="[337,442,1513,1534]" pageId="28" pageNumber="575" type="lectotype">Lectotype</typeStatus>
</emphasis>
.
</materialsCitation>
</paragraph>
</subSubSection>
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<paragraph id="9672503D3E63FFAC7FA3BC13DAD8F9D5" blockId="28.[144,759,1390,1779]" box="[168,759,1543,1565]" pageId="28" pageNumber="575">
<treatmentCitationGroup id="B6DD77133E63FFAC7FA3BC13DAD8F9D5" box="[168,759,1543,1565]" pageId="28" pageNumber="575">
<treatmentCitation id="176C762C3E63FFAC7FA3BC13DADCF9D5" author="Santschi F" box="[168,755,1543,1565]" page="129" pageId="28" pageNumber="575" year="1928">
<taxonomicName id="51CD2BBE3E63FFAC7FA3BC13DADCF9D5" ID-CoL="5GNKN" authority="Santschi 1928: 129" authorityName="Santschi" authorityPageNumber="129" authorityYear="1928" box="[168,755,1543,1565]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="575" phylum="Arthropoda" rank="variety" species="biroi" variety="andelis">
<emphasis id="A4B98C2F3E63FFAC7FA3BC13D9A8F9D5" box="[168,391,1543,1565]" italics="true" pageId="28" pageNumber="575">Crematogaster biroi</emphasis>
var.
<emphasis id="A4B98C2F3E63FFAC7ECABC1CDA3DF9D5" box="[449,530,1543,1564]" italics="true" pageId="28" pageNumber="575">andelis</emphasis>
<bibRefCitation id="F25C2DCC3E63FFAC7D11BC1CDADCF9D5" author="Santschi F" box="[538,755,1543,1565]" pageId="28" pageNumber="575" pagination="119 - 140" refId="ref40343" refString="Santschi F. 1928. Fourmis de Sumatra, recoltees par Mr. J. B. Corporaal. Tijdschrift Voor Entomologie 71: 119 - 140." type="journal article" year="1928">Santschi 1928: 129</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="9672503D3E63FFAC7F9BBC3DD8D6F9B1" blockId="28.[144,759,1390,1779]" pageId="28" pageNumber="575">
<materialsCitation id="26A55A603E63FFAC7F9BBC3DD8D6F9B1" ID-GBIF-Occurrence="3891597343" collectingDate="1919-09-02" collectionCode="NHMB" country="Indonesia" county="Pulu Berhala" location="Ile Varela" pageId="28" pageNumber="575" specimenCount="1" specimenCount-worker="1" stateProvince="Sumatra" typeStatus="syntype">
One
<typeStatus id="4976EE9F3E63FFAC7FCFBC3DD936F9F2" box="[196,281,1574,1595]" pageId="28" pageNumber="575" type="syntype">syntype</typeStatus>
<specimenCount id="80CB9BB43E63FFAC7E2BBC3DD941F9F5" box="[288,366,1574,1596]" pageId="28" pageNumber="575" type="worker">worker</specimenCount>
,
<location id="931206E63E63FFAC7E73BC3DD9CAF9F5" LSID="urn:lsid:plazi:treatment:1E64E12B3E63FFAD7FB0BF1ADC2EFDE2:931206E63E63FFAC7E73BC3DD9CAF9F5" box="[376,485,1574,1596]" country="Indonesia" county="Pulu Berhala" name="Ile Varela" pageId="28" pageNumber="575" stateProvince="Sumatra">Ile Varela</location>
,
<collectingCounty id="7F1328B13E63FFAC7EFBBC3DDAABF9F5" box="[496,644,1574,1596]" pageId="28" pageNumber="575">Pulu Berhala</collectingCounty>
,
<collectingRegion id="54099EDF3E63FFAC7D9BBC3DDADCF9F2" box="[656,755,1574,1595]" country="Indonesia" name="Sumatera Utara" pageId="28" pageNumber="575">Sumatra</collectingRegion>
,
<collectingCountry id="EEDA10AD3E63FFAC7F9BBC5FD92DF993" box="[144,258,1604,1626]" name="Indonesia" pageId="28" pageNumber="575">Indonesia</collectingCountry>
,
<date id="E27376FD3E63FFAC7E1BBC5FD957F993" box="[272,376,1604,1626]" pageId="28" pageNumber="575" value="1919-09-02">
<collectingDate id="F2378F153E63FFAC7E1BBC5FD957F993" box="[272,376,1604,1626]" pageId="28" pageNumber="575" value="1919-09-02">2.ix.1919</collectingDate>
</date>
(Corporaal) (
<collectionCode id="F0DCC8F83E63FFAC7D1DBC5EDA44F993" box="[534,619,1605,1626]" pageId="28" pageNumber="575">NHMB</collectionCode>
, examined). Syn. nov.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E63FFAC7FA3BC99D9A5F93A" pageId="28" pageNumber="575" type="reference_group">
<paragraph id="9672503D3E63FFAC7FA3BC99DA7FF97F" blockId="28.[144,759,1390,1779]" pageId="28" pageNumber="575">
<treatmentCitationGroup id="B6DD77133E63FFAC7FA3BC99DA7FF97F" pageId="28" pageNumber="575">
<taxonomicName id="51CD2BBE3E63FFAC7FA3BC99DA7EF95E" ID-CoL="5GNKP" authorityName="Forel" authorityYear="1913" box="[168,593,1666,1687]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="575" phylum="Arthropoda" rank="variety" species="biroi" variety="bandarensis">
<emphasis id="A4B98C2F3E63FFAC7FA3BC99D9A4F95E" box="[168,395,1666,1687]" italics="true" pageId="28" pageNumber="575">Crematogaster biroi</emphasis>
var.
<emphasis id="A4B98C2F3E63FFAC7ECDBC99DA7EF95E" box="[454,593,1666,1687]" italics="true" pageId="28" pageNumber="575">bandarensis</emphasis>
</taxonomicName>
;
<treatmentCitation id="176C762C3E63FFAC7D6BBC99D896F97C" author="Emery C" page="132" pageId="28" pageNumber="575" year="1922">
<bibRefCitation id="F25C2DCC3E63FFAC7D6BBC99D896F97C" author="Emery C" pageId="28" pageNumber="575" pagination="95 - 206" refId="ref38677" refString="Emery C. 1922. In Wytsman, P. Genera Insectorum. Hymenoptera, Fam. Formicidae, subfam. Myrmicinae. Fasc. 174 B, 95 - 206. Bruxelles." type="book chapter" year="1922">Emery, 1922: 132</bibRefCitation>
</treatmentCitation>
[Combination in
<emphasis id="A4B98C2F3E63FFAC7E83BCBAD9B5F97F" box="[392,410,1697,1718]" italics="true" pageId="28" pageNumber="575">C</emphasis>
. (
<taxonomicName id="51CD2BBE3E63FFAC7EB8BCBBDA15F97C" authorityName="Santschi" box="[435,570,1696,1717]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="605" phylum="Arthropoda" rank="subGenus" subGenus="Orthocrema">
<emphasis id="A4B98C2F3E63FFAC7EB8BCBBDA15F97C" box="[435,570,1696,1717]" italics="true" pageId="28" pageNumber="575">Orthocrema</emphasis>
</taxonomicName>
)].
</treatmentCitationGroup>
</paragraph>
<paragraph id="9672503D3E63FFAC7FA3BCA4D9A5F93A" blockId="28.[144,759,1390,1779]" pageId="28" pageNumber="575">
<treatmentCitationGroup id="B6DD77133E63FFAC7FA3BCA4D9A5F93A" pageId="28" pageNumber="575">
<taxonomicName id="51CD2BBE3E63FFAC7FA3BCA4DA3AF91D" ID-CoL="5GNKN" authorityName="Santschi" authorityYear="1928" box="[168,533,1727,1748]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="575" phylum="Arthropoda" rank="variety" species="biroi" variety="andelis">
<emphasis id="A4B98C2F3E63FFAC7FA3BCA4D9A6F91D" box="[168,393,1727,1748]" italics="true" pageId="28" pageNumber="575">Crematogaster biroi</emphasis>
var.
<emphasis id="A4B98C2F3E63FFAC7EC8BCA4DA3AF91D" box="[451,533,1727,1748]" italics="true" pageId="28" pageNumber="575">andelis</emphasis>
</taxonomicName>
;
<treatmentCitation id="176C762C3E63FFAC7D28BCA4DAD8F91D" author="Starcke A" box="[547,759,1727,1749]" page="373" pageId="28" pageNumber="575" year="1930">
<bibRefCitation id="F25C2DCC3E63FFAC7D28BCA4DAD8F91D" author="Starcke A" box="[547,759,1727,1749]" pageId="28" pageNumber="575" pagination="371 - 381" refId="ref40411" refString="Starcke A. 1930. Verzeichnis der bis jetzt von der Insel Pulau Berhala bekannt gewordenen Ameisen. Treubia 12: 371 - 381." type="journal article" year="1930">Stärcke, 1930: 373</bibRefCitation>
</treatmentCitation>
[Description of male].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E63FFAD7F9BBD00DA84FE5B" lastPageId="29" lastPageNumber="576" pageId="28" pageNumber="575" type="description">
<paragraph id="9672503D3E63FFAC7F9BBD00DAD8F8A4" blockId="28.[144,759,1819,1902]" pageId="28" pageNumber="575">
<emphasis id="A4B98C2F3E63FFAC7F9BBD00D9A1F8F9" box="[144,398,1819,1840]" italics="true" pageId="28" pageNumber="575">Worker measurements (</emphasis>
n
<emphasis id="A4B98C2F3E63FFAC7EA8BD00D9CFF8F9" box="[419,480,1819,1840]" italics="true" pageId="28" pageNumber="575">= 10):</emphasis>
HW 0.400.47; HL 0.42 0.50; CI 9298; SL 0.340.39; SI 8186; EL 0.09 0.11; PW 0.260.29; WL 0.480.54; PSL 0.050.08; PtL
</paragraph>
<paragraph id="9672503D3E63FFAC7C2DBADEDC06FEDE" blockId="28.[806,1421,197,280]" pageId="28" pageNumber="575">0.140.17; PtW 0.120.15; PtH 0.100.12; PpL 0.09 0.11; PpW 0.130.16; PtHI 6782; PtWI 8088; PpWI 127150; WI 100117.</paragraph>
<paragraph id="9672503D3E63FFAC7C2DBB24DDA1FE9C" blockId="28.[806,1422,319,1901]" box="[806,1422,319,341]" pageId="28" pageNumber="575">
<emphasis id="A4B98C2F3E63FFAC7C2DBB24DC52FE9D" box="[806,1149,319,340]" italics="true" pageId="28" pageNumber="575">General description of worker:</emphasis>
Workers monomorphic.
</paragraph>
<paragraph id="9672503D3E63FFAC7C34BB45DBCEFDE2" blockId="28.[806,1422,319,1901]" pageId="28" pageNumber="575">Head subquadratic in full-face view. Mandibles with four teeth arranged at an equal distance, apical and subapical teeth large, basal two teeth smaller. Anterior clypeal margin convex in medial portion. Compound eyes slightly projecting beyond lateral margins of head in full-face view. Scapes reaching posterolateral corners of head.</paragraph>
<paragraph id="9672503D3E63FFAC7C34B82FDDA1FC11" blockId="28.[806,1422,319,1901]" pageId="28" pageNumber="575">Pronotal collar with weakly concave anterior margin in dorsal view, distinctly lower than pronotum in lateral view. Pronotal dorsum without distinct ridges laterally. Mesonotal dorsum with lateral ridges that irregularly extend posteriad to tips of propodeal spines. Pronotum and mesonotum in lateral view not clearly forming continuous dorsal outline; mesonotal dorsum flat. Metanotal groove in dorsal view transverse, almost straight in median portion, forming deep concavity that is laterally margined by lamellate ridges. Propodeal spiracles elliptical, situated at posterolateral corners of propodeum, touching to metapleural gland bullae. Propodeal spines developed, as long as diameter of propodeal spiracles, in dorsal view directed posteriad.</paragraph>
<paragraph id="9672503D3E63FFAC7C34B9FBDD6EFB23" blockId="28.[806,1422,319,1901]" pageId="28" pageNumber="575">Petiole in dorsal view with subparallel sides, longer than wide. Posterior portion of petiole with short process that is slightly higher than posterior margin of petiole disc in lateral view. Subpetiolar process weakly developed as angulate tubercle. Postpetiole in lateral view with weakly convex dorsum, as high as petiole, in dorsal view as wide as petiole, weakly bilobed posteriorly but without longitudinal sulcus. Subpostpetiolar process undeveloped, but venter of postpetiole convex.</paragraph>
<paragraph id="9672503D3E63FFAC7C34BEEFDC98F95E" blockId="28.[806,1422,319,1901]" pageId="28" pageNumber="575">Integument essentially smooth and shining. Dorsal surface of head smooth and shining. Mandibles with feeble rugulae and smooth interspaces. Clypeus generally smooth and shining, but with one pair of longitudinal rugulae; rugulae not extending to posterior clypeal margin. Dorsal and lateral surfaces of pronotum smooth and shining; anterolateral shoulders of pronotum with rugulae. Mesopleura smooth and shining. Dorsal surface of propodeum smooth and shining, but one pair of rugulae running from metanotal groove to tips of propodeal spines. Dorsal surface pf petiole smooth. Lateral surface of petiole generally smooth, but with one longitudinal rugula. Dorsal and lateral surfaces of postpetiole smooth and shining.</paragraph>
<paragraph id="9672503D3E63FFAD7C34BCBBDA8BFEBD" blockId="28.[806,1422,319,1901]" lastBlockId="29.[164,779,197,402]" lastPageId="29" lastPageNumber="576" pageId="28" pageNumber="575">Standing pilosity sparse. Dorsal face of head with suberect setae sparsely. Clypeus with two pairs of long setae in anterior portion, one directed upward and the other downward. Anterior clypeal margin with one pair of long setae medially and short setae laterally. Scapes with suberect setae. Mesosoma with five pairs of long erect and stout setae (ps1PN, ps2PN, psaMN, pspMN, and ps1PS) that are much longer than other erect setae. Posterolateral tubercles of petiole posteriorly with two pairs of long setae. Postpetiole with three pairs of long setae on disc anteriorly, posteriorly and laterally. Fourth abdominal tergite with erect to suberect setae abundantly, and short decumbent setae sparsely.</paragraph>
<paragraph id="9672503D3E62FFAD7FB7BB67DA84FE5B" blockId="29.[164,779,197,402]" box="[188,683,380,402]" pageId="29" pageNumber="576">Body yellow. All flagellar segments yellow.</paragraph>
</subSubSection>
<subSubSection id="DED703B63E62FFAD7FAFBBA1DAA0FCCB" pageId="29" pageNumber="576" type="discussion">
<paragraph id="9672503D3E62FFAD7FAFBBA1DAA0FCCB" blockId="29.[164,779,442,770]" pageId="29" pageNumber="576">
<emphasis id="A4B98C2F3E62FFAD7FAFBBA1D90DFE06" box="[164,290,442,463]" italics="true" pageId="29" pageNumber="576">Comments:</emphasis>
In the worker this species can be distinguished from all other members of the
<taxonomicName id="51CD2BBE3E62FFAD7D6CBBC2DB25FE24" authorityName="Forel" authorityYear="1911" box="[615,778,472,494]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="576" phylum="Arthropoda" rank="species" species="quadriruga">
<emphasis id="A4B98C2F3E62FFAD7D6CBBC2DA56FE27" box="[615,633,473,494]" italics="true" pageId="29" pageNumber="576">C</emphasis>
.
<emphasis id="A4B98C2F3E62FFAD7D83BBC3DB25FE24" box="[648,778,472,493]" italics="true" pageId="29" pageNumber="576">quadriruga</emphasis>
</taxonomicName>
group by the distinct compound eyes, V-shaped metanotal groove in lateral view, large propodeal spiracles touching metapleural gland bulla, and short propodeal spines. This species is similar to
<taxonomicName id="51CD2BBE3E62FFAD7DFAB848D96AFD4F" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="576" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="A4B98C2F3E62FFAD7DFAB848DB2CFDA1" box="[753,771,595,616]" italics="true" pageId="29" pageNumber="576">C</emphasis>
.
<emphasis id="A4B98C2F3E62FFAD7FAFB86AD96AFD4F" box="[164,325,625,646]" italics="true" pageId="29" pageNumber="576">philippinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="51CD2BBE3E62FFAD7E8BB869DA7DFD4F" authorityName="Hosoishi &amp; Ogata" authorityYear="2016" box="[384,594,625,647]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="576" phylum="Arthropoda" rank="species" species="sundalandensis">
<emphasis id="A4B98C2F3E62FFAD7E8BB869D9BDFD4E" box="[384,402,626,647]" italics="true" pageId="29" pageNumber="576">C</emphasis>
.
<emphasis id="A4B98C2F3E62FFAD7EABB86ADA7DFD4F" box="[416,594,625,646]" italics="true" pageId="29" pageNumber="576">sundalandensis</emphasis>
</taxonomicName>
, but can be distinguished from them by the short and stout propodeal spines (PSL 0.050.08 vs. 0.080.11 and
<quantity id="5135FDD83E62FFAD7D74B8B4DB24FD0D" box="[639,779,686,708]" metricMagnitude="-3" metricUnit="m" metricValue="2.413" metricValueMax="2.7939999999999996" metricValueMin="2.032" pageId="29" pageNumber="576" unit="in" value="0.095" valueMax="0.11" valueMin="0.08">0.080.11 in</quantity>
the latters). This species corresponds to sp. 52 of SKY (
<bibRefCitation id="F25C2DCC3E62FFAD7FA7B8F7D978FCC8" author="Ito F &amp; Yamane S &amp; Eguchi K &amp; Noerdjito WA &amp; Kahono S &amp; Tsuji K &amp; Ohkawara K &amp; Yamauchi K &amp; Nishida T &amp; Nakamura K" box="[172,343,748,769]" pageId="29" pageNumber="576" pagination="379 - 404" refId="ref39608" refString="Ito F, Yamane S, Eguchi K, Noerdjito WA, Kahono S, Tsuji K, Ohkawara K, Yamauchi K, Nishida T, Nakamura K. 2001. Ant species diversity in the Bogor Botanic Garden, West Java, Indonesia, with descriptions of two new species of the genus Leptanilla (Hymenoptera, Formicidae). Tropics 10: 379 - 404." type="journal article" year="2001">
Ito
<emphasis id="A4B98C2F3E62FFAD7FDDB8F7D929FCC8" box="[214,262,748,769]" italics="true" pageId="29" pageNumber="576">et al</emphasis>
., 2001
</bibRefCitation>
;
<bibRefCitation id="F25C2DCC3E62FFAD7E6EB8F7DAABFCCB" author="Eguchi K &amp; Yamane SK" box="[357,644,748,770]" pageId="29" pageNumber="576" pagination="49 - 59" refId="ref38555" refString="Eguchi K, Yamane SK. 2003. Species diversity of ants (Hymenoptera, Formicidae) in a lowland rainforest, northwestern Borneo. New Entomologist 52: 49 - 59." type="journal article" year="2003">Eguchi &amp; Yamane, 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E62FFAD7FAFB932D8CCFC70" pageId="29" pageNumber="576" type="distribution">
<paragraph id="9672503D3E62FFAD7FAFB932D8CCFC70" blockId="29.[164,779,809,953]" pageId="29" pageNumber="576">
<emphasis id="A4B98C2F3E62FFAD7FAFB932D9EAFCF7" box="[164,453,809,830]" italics="true" pageId="29" pageNumber="576">Distribution and biology:</emphasis>
This species is known from S.
<collectingCountry id="EEDA10AD3E62FFAD7FCAB95CD907FC94" box="[193,296,839,861]" name="Thailand" pageId="29" pageNumber="576">Thailand</collectingCountry>
,
<collectingCountry id="EEDA10AD3E62FFAD7E3FB95CD9B5FC94" box="[308,410,839,861]" name="Malaysia" pageId="29" pageNumber="576">Malaysia</collectingCountry>
(Peninsula and Borneo),
<collectingCountry id="EEDA10AD3E62FFAD7DBCB95CDB29FC94" box="[695,774,839,861]" name="Brunei" pageId="29" pageNumber="576">Brunei</collectingCountry>
,
<collectingCountry id="EEDA10AD3E62FFAD7FAFB97DD93BFCB5" box="[164,276,870,892]" name="Indonesia" pageId="29" pageNumber="576">Indonesia</collectingCountry>
(
<collectingRegion id="54099EDF3E62FFAD7E2DB97DD978FCB5" box="[294,343,870,892]" country="Indonesia" name="Bali" pageId="29" pageNumber="576">Bali</collectingRegion>
,
<collectingRegion id="54099EDF3E62FFAD7E6CB97DD9E1FCB2" box="[359,462,870,891]" country="Indonesia" name="Sumatera Utara" pageId="29" pageNumber="576">Sumatra</collectingRegion>
, Krakatau) (
<figureCitation id="0EF64CB83E62FFAD7D6EB97DDA97FCB5" box="[613,696,870,892]" captionStart="Figure 52" captionStartId="18.[144,223,1277,1296]" captionTargetBox="[147,1422,198,1246]" captionTargetId="figure-211@18.[144,1424,195,1248]" captionTargetPageId="18" captionText="Figure 52. Geographical distribution of the species of the C. quadriruga group." figureDoi="http://doi.org/10.5281/zenodo.5358206" httpUri="https://zenodo.org/record/5358206/files/figure.png" pageId="29" pageNumber="576">Fig. 52</figureCitation>
). It inhabits developed forests, and nests in soil or in leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="DED703B63E62FFAD7FAFB9FBDC2EFDE2" pageId="29" pageNumber="576" type="materials_examined">
<paragraph id="9672503D3E62FFAD7FAFB9FBDC2EFDE2" blockId="29.[164,779,992,1902]" lastBlockId="29.[826,1441,197,555]" pageId="29" pageNumber="576">
<emphasis id="A4B98C2F3E62FFAD7FAFB9FBD9BAFC3C" box="[164,405,992,1013]" italics="true" pageId="29" pageNumber="576">Material examined:</emphasis>
<materialsCitation id="26A55A603E62FFAD7EA8B9FADAF5FBDC" ID-GBIF-Occurrence="3891597378" pageId="29" pageNumber="576">
<collectingCountry id="EEDA10AD3E62FFAD7EA8B9FADA3BFC3F" box="[419,532,993,1014]" name="Brunei" pageId="29" pageNumber="576">BRUNEI</collectingCountry>
:
<specimenCount id="80CB9BB43E62FFAD7D2FB9FADA9DFC3F" box="[548,690,992,1014]" pageId="29" pageNumber="576" type="worker">one worker</specimenCount>
, Tasek Merimbun,
<date id="E27376FD3E62FFAD7E2DB9E4D9BDFBDC" box="[294,402,1023,1045]" pageId="29" pageNumber="576" value="1999-02-16">16.ii.1999</date>
, (Eg99-BOR-126) (K. Eguchi)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7DEDB9E4D9C9FB9B" ID-GBIF-Occurrence="3891597328" pageId="29" pageNumber="576">
INDONESIA:
<specimenCount id="80CB9BB43E62FFAD7E2FBE05D9B1FBFD" box="[292,414,1054,1076]" pageId="29" pageNumber="576" type="worker">one worker</specimenCount>
, Sembung,
<quantity id="5135FDD83E62FFAD7D16BE06DA4EFBFD" box="[541,609,1053,1076]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="576" unit="km" value="25.0">25 km</quantity>
N of Denpasar, Bali,
<date id="E27376FD3E62FFAD7FEABE27D975FB9B" box="[225,346,1084,1106]" pageId="29" pageNumber="576" value="1987-03-19">19.iii.1987</date>
, (Imai
<emphasis id="A4B98C2F3E62FFAD7EA0BE26D9F3FB98" box="[427,476,1084,1106]" italics="true" pageId="29" pageNumber="576">et al</emphasis>
)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7EF8BE27D9CAFB46" ID-GBIF-Occurrence="3891597352" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7EF8BE27DABBFB9B" box="[499,660,1084,1106]" pageId="29" pageNumber="576" type="worker">three workers</specimenCount>
, Tuapejat, Pulau Sipora, Mentawai Is., W. Sumatra,
<date id="E27376FD3E62FFAD7D90BE40DB25FBB8" box="[667,778,1115,1137]" pageId="29" pageNumber="576" value="2007-02-24">24.ii.2007</date>
(
<collectionCode id="F0DCC8F83E62FFAD7FA0BE62D8E3FB46" box="[171,204,1145,1167]" country="USA" name="Stanford University" pageId="29" pageNumber="576">SU</collectionCode>
07-SKY-97) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7EFBBE61DB29FB05" ID-GBIF-Occurrence="3891597325" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7EFBBE61DA43FB46" box="[496,620,1145,1167]" pageId="29" pageNumber="576" type="worker">one worker</specimenCount>
, Bodong Jaya, Sumberjaya, Lampung Barat (secondary forest), S. Sumatra,
<date id="E27376FD3E62FFAD7E1DBEADD9A4FB05" box="[278,395,1206,1228]" pageId="29" pageNumber="576" value="2007-09-16">16.ix.2007</date>
(leaf litter / s. soil) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7FAFBECED926FAE1" ID-GBIF-Occurrence="3891597389" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7FAFBECED90DFB22" box="[164,290,1237,1259]" pageId="29" pageNumber="576" type="worker">six workers</specimenCount>
, Bodong Java, Sumberjaya, Lampung Barat, S. Sumatra,
<date id="E27376FD3E62FFAD7E3BBEEFD98FFAC3" box="[304,416,1268,1290]" pageId="29" pageNumber="576" value="2007-09-16">16.ix.2007</date>
(leaf litter) (
<collectionCode id="F0DCC8F83E62FFAD7D27BEEFDA61FAC3" box="[556,590,1268,1290]" country="USA" name="Stanford University" pageId="29" pageNumber="576">SU</collectionCode>
07-SKY-156) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7E1EBF09DB28FA8F" ID-GBIF-Occurrence="3891597339" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7E1EBF09D987FAE1" box="[277,424,1298,1320]" pageId="29" pageNumber="576" type="worker">four workers</specimenCount>
, Pulau Sebesi, Lampung Prov., Sunda Strait,
<date id="E27376FD3E62FFAD7E4BBF2AD992FA8E" box="[320,445,1329,1351]" pageId="29" pageNumber="576" value="2005-08-12">12.viii.2005</date>
(RK05-SKY-24) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7FAFBF4BDAFDFA4D" ID-GBIF-Occurrence="3891597310" pageId="29" pageNumber="576">
<collectingCountry id="EEDA10AD3E62FFAD7FAFBF4BD902FAAC" box="[164,301,1360,1381]" name="Malaysia" pageId="29" pageNumber="576">MALAYSIA</collectingCountry>
:
<specimenCount id="80CB9BB43E62FFAD7E30BF54D9FCFAAC" box="[315,467,1359,1381]" pageId="29" pageNumber="576" type="worker">four workers</specimenCount>
,
<quantity id="5135FDD83E62FFAD7EE9BF4BDA67FAAC" box="[482,584,1359,1381]" metricMagnitude="4" metricUnit="m" metricValue="3.0577536" pageId="29" pageNumber="576" unit="mi" value="19.0">19 miles</quantity>
, Cameron Highlands,
<date id="E27376FD3E62FFAD7FFCBF75D95FFA4D" box="[247,368,1390,1412]" pageId="29" pageNumber="576" value="2010-03-11">11.iii.2010</date>
(SH10-Mal-48) (S. Hosoishi)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7DE9BF74D948FA08" ID-GBIF-Occurrence="3891597376" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7DE9BF74D8DCFA6A" pageId="29" pageNumber="576" type="worker">one worker</specimenCount>
, Ulu Gombak,
<collectingRegion id="54099EDF3E62FFAD7E90BF96D9D1FA6A" box="[411,510,1421,1443]" country="Malaysia" name="Selangor" pageId="29" pageNumber="576">Selangor</collectingRegion>
,
<date id="E27376FD3E62FFAD7D02BF96DA55FA6A" box="[521,634,1421,1443]" pageId="29" pageNumber="576" value="2005-11-27">27.xi.2005</date>
(
<collectionCode id="F0DCC8F83E62FFAD7D83BF96DA80FA6A" box="[648,687,1421,1443]" country="0" name="Mohonk Preserve, Inc." pageId="29" pageNumber="576">MP</collectionCode>
05-SKY- 08) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7E7FBFB0D976FA37" ID-GBIF-Occurrence="3891597396" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7E7FBFB0DA29FA08" box="[372,518,1451,1473]" pageId="29" pageNumber="576" type="worker">four workers</specimenCount>
, UM plantations, Kota Tinggi,
<collectingRegion id="54099EDF3E62FFAD7FF7BFD1D912FA29" box="[252,317,1482,1504]" country="Malaysia" name="Johor" pageId="29" pageNumber="576">Johor</collectingRegion>
,
<date id="E27376FD3E62FFAD7E42BFD1D9E9FA29" box="[329,454,1482,1504]" pageId="29" pageNumber="576" value="2013-12-17">17.xii.2013</date>
(sugar, TD-85) (Sk. Yamane &amp; Y. Miyaguni)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7E6CBFF3D973F9F5" ID-GBIF-Occurrence="3891597432" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7E6CBFF3D9D2FA37" box="[359,509,1512,1534]" pageId="29" pageNumber="576" type="worker">four workers</specimenCount>
, UM plantations, Kota Tinggi,
<collectingRegion id="54099EDF3E62FFAD7FF0BC1CD913F9D4" box="[251,316,1543,1565]" country="Malaysia" name="Johor" pageId="29" pageNumber="576">Johor</collectingRegion>
,
<date id="E27376FD3E62FFAD7E43BC1CD9EBF9D4" box="[328,452,1543,1565]" pageId="29" pageNumber="576" value="2013-12-17">17.xii.2013</date>
(sugar, GD-64) (Sk. Yamane &amp; Y. Miyaguni)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7E60BC3DDA5CF993" ID-GBIF-Occurrence="3891597312" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7E60BC3DD9C0F9F5" box="[363,495,1574,1596]" pageId="29" pageNumber="576" type="worker">one worker</specimenCount>
, Kubah N. P., Kuching,
<collectingRegion id="54099EDF3E62FFAD7FAFBC5FD928F993" box="[164,263,1604,1626]" country="Malaysia" name="Sarawak" pageId="29" pageNumber="576">Sarawak</collectingRegion>
, Borneo,
<date id="E27376FD3E62FFAD7E66BC5FD9CBF993" box="[365,484,1604,1626]" pageId="29" pageNumber="576" value="1997-12-28">28.xii.1997</date>
, (F. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7D75BC5EDAF9F95E" ID-GBIF-Occurrence="3891597393" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7D75BC5EDB28F993" box="[638,775,1604,1626]" pageId="29" pageNumber="576" type="worker">two workers</specimenCount>
, 8 ha plot, Tower Reg., Lambir, Miri,
<collectingRegion id="54099EDF3E62FFAD7D55BC78DAECF9B0" box="[606,707,1635,1657]" country="Malaysia" name="Sarawak" pageId="29" pageNumber="576">Sarawak</collectingRegion>
#117, E Malaysia,
<date id="E27376FD3E62FFAD7E3DBC9AD992F95E" box="[310,445,1665,1687]" pageId="29" pageNumber="576" value="1994-08-27">27.viii.1994</date>
, (
<collectionCode id="F0DCC8F83E62FFAD7ED9BC99D9CBF95E" box="[466,484,1666,1687]" name="Tavera, Department of Geology and Geophysics" pageId="29" pageNumber="576">T</collectionCode>
. Itioka &amp;
<collectionCode id="F0DCC8F83E62FFAD7D52BC99DA44F95E" box="[601,619,1666,1687]" name="Tavera, Department of Geology and Geophysics" pageId="29" pageNumber="576">T</collectionCode>
. Yumoto)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7DE8BC99DA77F91C" ID-GBIF-Occurrence="3891597375" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7DE8BC99D8DAF97F" pageId="29" pageNumber="576" type="worker">one worker</specimenCount>
, Old Tower R., Lambir N. P., Miri,
<collectingRegion id="54099EDF3E62FFAD7D94BCBBDB28F97F" box="[671,775,1696,1718]" country="Malaysia" name="Sarawak" pageId="29" pageNumber="576">Sarawak</collectingRegion>
,
<date id="E27376FD3E62FFAD7FAFBCA4D924F91C" box="[164,267,1727,1749]" pageId="29" pageNumber="576" value="1993-01-22">22.i.1993</date>
, (Sk. Yamane) (Canopy Ecol.)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7D6FBCA4DA27F8D8" ID-GBIF-Occurrence="3891597315" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7D6FBCA4DB29F91C" box="[612,774,1727,1749]" pageId="29" pageNumber="576" type="worker">seven workers</specimenCount>
, Lambir N. P., Miri,
<collectingRegion id="54099EDF3E62FFAD7E9DBCC6D9D1F93A" box="[406,510,1757,1779]" country="Malaysia" name="Sarawak" pageId="29" pageNumber="576">Sarawak</collectingRegion>
,
<date id="E27376FD3E62FFAD7D04BCC6DA56F93B" box="[527,633,1757,1778]" pageId="29" pageNumber="576" value="2011-05-14">14.v.2011</date>
(SR11-SKY- 11) (nest in soil) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7D1CBCE7D8FCF886" ID-GBIF-Occurrence="3891597388" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7D1CBCE7DA88F8DB" box="[535,679,1788,1810]" pageId="29" pageNumber="576" type="worker">two workers</specimenCount>
, Danum Valley,
<collectingRegion id="54099EDF3E62FFAD7FF9BD01D915F8F9" box="[242,314,1818,1840]" country="Malaysia" name="Sabah" pageId="29" pageNumber="576">Sabah</collectingRegion>
, Borneo,
<date id="E27376FD3E62FFAD7EAABD01DA0DF8F9" box="[417,546,1818,1840]" pageId="29" pageNumber="576" value="1995-08-29">29.viii.1995</date>
, (Sk. Yamane) (prim. for.)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7FEBBD22DA0FF8A7" ID-GBIF-Occurrence="3891597397" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7FEBBD22D95FF886" box="[224,368,1849,1871]" pageId="29" pageNumber="576" type="worker">two workers</specimenCount>
, Tawau Hills N. P.,
<collectingRegion id="54099EDF3E62FFAD7D57BD22DA89F886" box="[604,678,1849,1871]" country="Malaysia" name="Sabah" pageId="29" pageNumber="576">Sabah</collectingRegion>
, Borneo, E. Malaysia,
<date id="E27376FD3E62FFAD7E3ABD43D9B5F8A7" box="[305,410,1880,1902]" pageId="29" pageNumber="576" value="1996-07-08">8.vii.1996</date>
, (K. Eguchi)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7D20BD43DCA5FF30" ID-GBIF-Occurrence="3891597377" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7D20BD43DAECF8A7" box="[555,707,1880,1902]" pageId="29" pageNumber="576" type="worker">three workers</specimenCount>
, Inoue Trail, Lambir N. P.,
<collectingRegion id="54099EDF3E62FFAD7B2FBADEDCA3FF12" box="[1060,1164,197,219]" country="Malaysia" name="Sarawak" pageId="29" pageNumber="576">Sarawak</collectingRegion>
, E. Malaysia,
<date id="E27376FD3E62FFAD7A39BADEDD8EFF12" box="[1330,1441,197,219]" pageId="29" pageNumber="576" value="2004-07-02">2.vii.2004</date>
(SR04-SKY-78) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7B92BAFFDCE2FE9D" ID-GBIF-Occurrence="3891597461" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7B92BAFFDD6EFF30" box="[1177,1345,227,249]" pageId="29" pageNumber="576" type="worker">seven workers</specimenCount>
, Maliau Basin (riparian forest),
<collectingRegion id="54099EDF3E62FFAD7B57BB19DC88FED1" box="[1116,1191,258,280]" country="Malaysia" name="Sabah" pageId="29" pageNumber="576">Sabah</collectingRegion>
, Borneo (
<quantity id="5135FDD83E62FFAD7A16BB19DD4AFEDE" box="[1309,1381,258,279]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="576" unit="m" value="300.0">300 m</quantity>
alt.)
<date id="E27376FD3E62FFAD7C31BB3BDB81FEFF" box="[826,942,288,310]" pageId="29" pageNumber="576" value="2011-11-10">10.xi.2011</date>
(ex carton cover on plant stem with scale insects) (SB11-SKY-38) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7BDCBB24DBF9FE79" ID-GBIF-Occurrence="3891597463" pageId="29" pageNumber="576">
<collectingCountry id="EEDA10AD3E62FFAD7BDCBB24DD4FFE9D" box="[1239,1376,319,340]" name="Thailand" pageId="29" pageNumber="576">THAILAND</collectingCountry>
:
<specimenCount id="80CB9BB43E62FFAD7A61BB24DBB7FEBD" pageId="29" pageNumber="576" type="worker">eight workers</specimenCount>
, Khlong Saeng WS, Ratchaprapha D.,
<collectingRegion id="54099EDF3E62FFAD7A6BBB45DBEDFE5B" country="Thailand" name="Surat Thani" pageId="29" pageNumber="576">Surat Thani Prov.</collectingRegion>
,
<date id="E27376FD3E62FFAD7CC6BB67DC17FE5B" box="[973,1080,380,402]" pageId="29" pageNumber="576" value="2011-10-15">15.x.2011</date>
(TH11-SKY-134) (rotting wood) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7CEEBB80DC13FE24" ID-GBIF-Occurrence="3891597340" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7CEEBB80DC59FE78" box="[997,1142,411,433]" pageId="29" pageNumber="576" type="worker">two workers</specimenCount>
, Tai Rom Yen NP (
<quantity id="5135FDD83E62FFAD7A51BB80DD8EFE79" box="[1370,1441,411,432]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="576" unit="m" value="400.0">400 m</quantity>
alt.),
<collectingRegion id="54099EDF3E62FFAD7C79BBA1DC18FE06" box="[882,1079,441,463]" country="Thailand" name="Surat Thani" pageId="29" pageNumber="576">Surat Thani Prov.</collectingRegion>
,
<date id="E27376FD3E62FFAD7B4ABBA1DC8AFE06" box="[1089,1189,442,463]" pageId="29" pageNumber="576" value="2011-10-11">11.x.2011</date>
(decayed wood) (TH11- SKY-016) (Sk. Yamane)
</materialsCitation>
;
<materialsCitation id="26A55A603E62FFAD7B4CBBC3DC2EFDE2" ID-GBIF-Occurrence="3891597467" pageId="29" pageNumber="576">
<specimenCount id="80CB9BB43E62FFAD7B4CBBC3DCCCFE27" box="[1095,1251,472,494]" pageId="29" pageNumber="576" type="worker">three workers</specimenCount>
, Ton Nga Chang, nr. Hat Yai,
<collectingRegion id="54099EDF3E62FFAD7CDCBBECDCBFFDC5" box="[983,1168,503,525]" country="Thailand" name="Songkhla" pageId="29" pageNumber="576">Songkhla Prov.</collectingRegion>
,
<date id="E27376FD3E62FFAD7BABBBECDD37FDC4" box="[1184,1304,503,525]" pageId="29" pageNumber="576" value="2001-09-23">23.ix.2001</date>
(Eg01-TH- 609) (K. Eguchi).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>