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<document id="F02BBC0F102AFC373FEF5EA17ABB91B9" ID="10.11646/zootaxa.4173.1.9" ID-CLB-Dataset="38079" ID-DOI="10.11646/zootaxa.4173.1.9" ID-GBIF-Dataset="197debd5-a36b-4984-ba7f-7f6a0e2a7d14" ID-ISSN="1175-5326" ID-Zenodo-Dep="159237" ID-ZooBank="A903E6AF-EA63-419A-9871-FBA44B25033C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1475479562258" checkinUser="plazi" docAuthor="Pedraza, Manuel, Tavares, Marcos &amp; Magalhães, Célio" docDate="2016" docId="03FF87E7F946FFCB029AFE9BFD6B51FD" docLanguage="en" docName="zootaxa.4173.1.9.pdf" docOrigin="Zootaxa 4173 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Melothelphusa apiaka Pedraza, Tavares &amp; Magalhães, 2016, n. sp." docType="treatment" docVersion="9" lastPageNumber="99" masterDocId="FFC6FF9FF944FFCD020DFA1DFFB45078" masterDocTitle="A new genus of freshwater crab of the tribe Kingsleyini Bott, 1970 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) with description of a new species from Mato Grosso, Brazil" masterLastPageNumber="100" masterPageNumber="94" pageNumber="96" updateTime="1698428851158" updateUser="ExternalLinkService">
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<mods:title id="6B9F2C6335A4921199D3256689795863">A new genus of freshwater crab of the tribe Kingsleyini Bott, 1970 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) with description of a new species from Mato Grosso, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="41662FF9EF91B9487BE7D20DAB1A07B2" type="personal">
<mods:role id="8F2AC26979C81DF28F217E5DC1B7E6F2">
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<mods:namePart id="FC38024A8477D447D986E42055F54209">Pedraza, Manuel</mods:namePart>
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<mods:name id="578EB93E3445AC510825F40A739EC871" type="personal">
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<mods:namePart id="C333CCD149AACEDC9C586918C24B7365">Tavares, Marcos</mods:namePart>
</mods:name>
<mods:name id="15D5AE968577190ABC902D654C08F870" type="personal">
<mods:role id="8A00593670E6EC4C5C4F5D9F3FC2420E">
<mods:roleTerm id="D8EEDA679790801CF511EBAAF3A749E5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="721F495D08C7BE1B2BDBEB4C528D9E33">Magalhães, Célio</mods:namePart>
</mods:name>
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<mods:title id="FD7C512FB736664ED06FE893854DA1C8">Zootaxa</mods:title>
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<mods:part id="317E0DBDB2A9075D6DBD382AF56F19B7">
<mods:date id="92697B5C92E8500A1EA65C88D483B89A">2016</mods:date>
<mods:detail id="E29883A41D3510F03F0EDB4752FF6E5B" type="volume">
<mods:number id="52B7F0D623CD2581ACDA0234E9C0B6F8">4173</mods:number>
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<mods:identifier id="53FC9E187A188B7154E1433967D86F24" type="DOI">10.11646/zootaxa.4173.1.9</mods:identifier>
<mods:identifier id="D3D781E2A18F8BAA0AAFD9D76394A41B" type="GBIF-Dataset">197debd5-a36b-4984-ba7f-7f6a0e2a7d14</mods:identifier>
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<treatment id="03FF87E7F946FFCB029AFE9BFD6B51FD" ID-DOI="http://doi.org/10.5281/zenodo.6087893" ID-GBIF-Taxon="124556917" ID-Zenodo-Dep="6087893" LSID="urn:lsid:plazi:treatment:03FF87E7F946FFCB029AFE9BFD6B51FD" httpUri="http://treatment.plazi.org/id/03FF87E7F946FFCB029AFE9BFD6B51FD" lastPageId="6" lastPageNumber="99" pageId="2" pageNumber="96">
<subSubSection id="C34C657AF946FFCF029AFE9BFE7A54BA" pageId="2" pageNumber="96" type="nomenclature">
<paragraph id="8BE936F1F946FFCF029AFE9BFE5A54D8" blockId="2.[151,494,1158,1218]" box="[151,494,1158,1184]" pageId="2" pageNumber="96">
<heading id="D0A1819DF946FFCF029AFE9BFE5A54D8" bold="true" box="[151,494,1158,1184]" fontSize="11" level="1" pageId="2" pageNumber="96" reason="1">
<emphasis id="B922EAE3F946FFCF029AFE9BFE5A54D8" bold="true" box="[151,494,1158,1184]" pageId="2" pageNumber="96">
<taxonomicName id="4C564D72F946FFCF029AFE9BFE1054D8" ID-CoL="84FZL" box="[151,420,1158,1184]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="apiaka" status="sp. nov.">
<emphasis id="B922EAE3F946FFCF029AFE9BFE1054D8" bold="true" box="[151,420,1158,1184]" italics="true" pageId="2" pageNumber="96">Melothelphusa apiaka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F946FFCF03A1FE9BFE5A54D8" box="[428,494,1158,1184]" pageId="2" pageNumber="96" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE936F1F946FFCF029AFEB4FE7A54BA" blockId="2.[151,494,1158,1218]" box="[151,462,1193,1218]" pageId="2" pageNumber="96">
(
<figureCitation id="136D2A74F946FFCF0292FEB4FE9654BA" box="[159,290,1193,1218]" captionStart="FIGURE 1 A E" captionStartId="3.[151,250,1081,1103]" captionTargetBox="[162,1425,206,1045]" captionTargetId="figure@3.[150,1437,195,1056]" captionTargetPageId="3" captionText="FIGURE 1 A E. Melothelphusa apiaka n. sp., male holotype, cl 18.8 mm, cw 27.8 mm (MZUSP 32914). A, habitus, dorsal view. B, cephalothorax, ventral view. C, cephalothorax, frontal view. D, E, lateral view of right and left chelipeds, respectively. Abbreviations: CxP 4, coxa of pereopod 4; G 1, gonopod 1; G 2, gonopod 2; St 4 8, thoracic sternites 4 to 8. Scale bars A E, 10 mm." httpUri="https://zenodo.org/record/159238/files/figure.png" pageId="2" pageNumber="96">Figs. 1AE</figureCitation>
; 2AC; 3AD)
</paragraph>
</subSubSection>
<subSubSection id="C34C657AF946FFCF029AFEECFBF3577A" pageId="2" pageNumber="96" type="materials_examined">
<paragraph id="8BE936F1F946FFCF029AFEECFE035636" blockId="2.[151,1437,1265,1974]" pageId="2" pageNumber="96">
<materialsCitation id="3B3E3CACF946FFCF029AFEECFC745556" ID-GBIF-Occurrence="1315974754" collectingDate="1997-02-10" collectingDateMax="1997-02-15" collectingDateMin="1997-02-10" collectionCode="MZUSP" collectorName="G. Skuk &amp; V. Xavier" country="Brazil" elevation="220" latitude="-9.566667" location="Apiacas" longLatPrecision="1299" longitude="-57.383335" pageId="2" pageNumber="96" specimenCode="MZUSP 32914" specimenCount="2" stateProvince="Mato Grosso" typeStatus="holotype">
<emphasis id="B922EAE3F946FFCF029AFEECFEF05572" bold="true" box="[151,324,1265,1290]" pageId="2" pageNumber="96">
<typeStatus id="54ED8853F946FFCF029AFEECFF665572" box="[151,210,1265,1290]" pageId="2" pageNumber="96">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54ED8853F946FFCF0346FEECFE0C5572" box="[331,440,1265,1290]" pageId="2" pageNumber="96" type="holotype">Holotype</typeStatus>
: male (cl 18.8 x cw 27.8),
<collectingCountry id="F3417661F946FFCF00EDFEECFC935572" box="[736,807,1265,1290]" name="Brazil" pageId="2" pageNumber="96">Brazil</collectingCountry>
,
<collectingRegion id="4992F813F946FFCF013FFEEFFC725572" box="[818,966,1266,1290]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="96">Mato Grosso</collectingRegion>
,
<location id="8E89602AF946FFCF01DCFEECFB9A5572" LSID="urn:lsid:plazi:treatment:03FF87E7F946FFCB029AFE9BFD6B51FD:8E89602AF946FFCF01DCFEECFB9A5572" box="[977,1070,1265,1290]" country="Brazil" latitude="-9.566667" longLatPrecision="1299" longitude="-57.383335" name="Apiacas" pageId="2" pageNumber="96" stateProvince="Mato Grosso">Apiacás</location>
, [
<geoCoordinate id="EE625036F946FFCF064CFEEFFB245572" box="[1089,1168,1266,1290]" direction="south" orientation="latitude" pageId="2" pageNumber="96" precision="925" value="-9.566667">9º34S</geoCoordinate>
,
<geoCoordinate id="EE625036F946FFCF0691FEECFAB15572" box="[1180,1285,1265,1290]" direction="west" orientation="longitude" pageId="2" pageNumber="96" precision="925" value="-57.383335">57º23W</geoCoordinate>
], altitude
<elevation id="007BD1C2F946FFCF077FFEEFFF1B5555" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="96" unit="m" value="220.0">
<quantity id="4CAE9B14F946FFCF077FFEEFFF1B5555" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="96" unit="m" value="220.0">220 m</quantity>
</elevation>
,
<collectorName id="26A35327F946FFCF02B0FF0BFEAD5556" box="[189,281,1301,1326]" pageId="2" pageNumber="96">G. Skuk</collectorName>
&amp;
<collectorName id="26A35327F946FFCF034FFF0BFE045556" box="[322,432,1301,1326]" pageId="2" pageNumber="96">V. Xavier</collectorName>
coll.,
<collectingDate id="EFACE9D9F946FFCF03F1FF08FD465556" box="[508,754,1301,1326]" pageId="2" pageNumber="96" value="1997-02-10" valueMax="1997-02-15" valueMin="1997-02-10">
<date id="FFE81031F946FFCF03F1FF08FD465556" box="[508,754,1301,1326]" pageId="2" pageNumber="96" value="1997-02-10" valueMax="1997-02-15" valueMin="1997-02-10">1015 February 1997</date>
</collectingDate>
(
<specimenCode id="DBF09E8AF946FFCF0109FF0BFC0C5556" box="[772,952,1302,1326]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="96">
<collectionCode id="ED47AE34F946FFCF0109FF0BFCD75556" box="[772,867,1302,1326]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="96">MZUSP</collectionCode>
32914
</specimenCode>
)
</materialsCitation>
. Paratypes (same date as holotype unless otherwise stated): 1 male (cl 17.2 x cw 26.3), 3 females (cl 20.6 x cw 32.8; cl 18.5 x cw 28.3; cl 20.3 x cw 32.2) (MZUSP 32915); 1 male (cl 16.1 x cw 24.3), 2 females (cl 14.1 x cw 21.2; cl 17.2 x cw 25.8) (INPA 2183); 1 male (cl 19.2 x cw 28.9); 3 females (cl 21.6 x cw 32.8; cl 19.9 x cw 30.9; cl 16.4 x cw 25.4), Ana Maria Ramos-Costa coll.,
<date id="FFE81031F946FFCF02D8FFB8FE7255C6" box="[213,454,1445,1470]" pageId="2" pageNumber="96" value="1997-02-15" valueMax="1997-02-17" valueMin="1997-02-15">1517 February 1997</date>
(MZUSP 32916): 1 male (cl 18.8 x cw 27.5); 2 females (cl 19.5 x cw 29.5; cl 22.5 x cw 35.0), Ana Maria Ramos-Costa coll.,
<date id="FFE81031F946FFCF004CFFD4FC81559A" box="[577,821,1481,1506]" pageId="2" pageNumber="96" value="1997-02-15" valueMax="1997-02-17" valueMin="1997-02-15">1517 February 1997</date>
(INPA 2184):
<materialsCitation id="3B3E3CACF946FFCF01D3FFD7FDA65652" ID-GBIF-Occurrence="1315974752" collectingDate="1997-06-07" collectingDateMax="1997-06-18" collectingDateMin="1997-06-07" collectionCode="INPA" collectorName="G. Skuk" country="Brazil" county="Juruena" latitude="-10.366667" location="Juruena" longLatPrecision="1298" longitude="-58.333332" municipality="Juruena River" pageId="2" pageNumber="96" specimenCode="INPA 2185" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso" typeStatus="holotype">
<specimenCount id="9D50FD78F946FFCF01D3FFD7FB9E559A" box="[990,1066,1481,1506]" pageId="2" pageNumber="96" type="male">1 male</specimenCount>
(cl 18.9), dry, carapace partially damaged,
<collectingCountry id="F3417661F946FFCF0303FFF0FEE1567E" box="[270,341,1517,1542]" name="Brazil" pageId="2" pageNumber="96">Brazil</collectingCountry>
,
<collectingRegion id="4992F813F946FFCF0368FFF3FE4A567E" box="[357,510,1518,1542]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="96">Mato Grosso</collectingRegion>
,
<collectingCounty id="62884E7DF946FFCF0003FFF3FDD1567D" box="[526,613,1518,1541]" pageId="2" pageNumber="96">Juruena</collectingCounty>
[probably
<collectingMunicipality id="6B8DAC8BF946FFCF00E4FFF3FC3F567E" box="[745,907,1517,1542]" pageId="2" pageNumber="96">Juruena River</collectingMunicipality>
, near
<location id="8E89602AF946FFCF01DBFFF3FB84567D" LSID="urn:lsid:plazi:treatment:03FF87E7F946FFCB029AFE9BFD6B51FD:8E89602AF946FFCF01DBFFF3FB84567D" box="[982,1072,1518,1541]" country="Brazil" county="Juruena" latitude="-10.366667" longLatPrecision="1298" longitude="-58.333332" municipality="Juruena River" name="Juruena" pageId="2" pageNumber="96" stateProvince="Mato Grosso">Juruena</location>
,
<geoCoordinate id="EE625036F946FFCF064DFFF3FB2D567E" box="[1088,1177,1518,1542]" direction="south" orientation="latitude" pageId="2" pageNumber="96" precision="925" value="-10.366667">10º22S</geoCoordinate>
<geoCoordinate id="EE625036F946FFCF06A8FFF0FABA567E" box="[1189,1294,1517,1542]" direction="west" orientation="longitude" pageId="2" pageNumber="96" precision="925" value="-58.333332">58º20W</geoCoordinate>
],
<collectingDate id="EFACE9D9F946FFCF0728FFF3FF665652" pageId="2" pageNumber="96" value="1997-06-07" valueMax="1997-06-18" valueMin="1997-06-07">
<date id="FFE81031F946FFCF0728FFF3FF665652" pageId="2" pageNumber="96" value="1997-06-07" valueMax="1997-06-18" valueMin="1997-06-07">718 June 1997</date>
</collectingDate>
,
<collectorName id="26A35327F946FFCF02D2FC0FFE8E5652" box="[223,314,1553,1578]" pageId="2" pageNumber="96">G. Skuk</collectorName>
coll. (
<specimenCode id="DBF09E8AF946FFCF038EFC0FFDBD5652" box="[387,521,1553,1578]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="96">
<collectionCode id="ED47AE34F946FFCF038EFC0FFE705652" box="[387,452,1554,1578]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="96">INPA</collectionCode>
2185
</specimenCode>
)
</materialsCitation>
; 1 male (cl 14.5 x cw 20.5), Brazil, Mato Grosso, Juruena, G. Skuk coll.,
<date id="FFE81031F946FFCF0769FC0FFEBD5636" pageId="2" pageNumber="96" value="1997-06-07" valueMax="1997-06-18" valueMin="1997-06-07">718 June 1997</date>
(MPEG 1742).
</paragraph>
<paragraph id="8BE936F1F946FFCF02CAFC44FF5456A6" blockId="2.[151,1437,1265,1974]" pageId="2" pageNumber="96">
<materialsCitation id="3B3E3CACF946FFCF02CAFC44FB8E56EE" ID-GBIF-Occurrence="1315974755" collectingDate="1973-07-20" collectingDateMax="1973-07-28" collectingDateMin="1973-07-20" collectionCode="MZUSP" collectorName="Expedicao Permanente da Amazonia" country="Brazil" county="Comparative" location="Tapajos river" municipality="Monte Cristo" pageId="2" pageNumber="96" specimenCode="MZUSP 6550" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<emphasis id="B922EAE3F946FFCF02CAFC44FE68560A" bold="true" box="[199,476,1625,1650]" pageId="2" pageNumber="96">
<collectingCounty id="62884E7DF946FFCF02CAFC44FED0560A" box="[199,356,1625,1650]" pageId="2" pageNumber="96">Comparative</collectingCounty>
material.
</emphasis>
<taxonomicName id="4C564D72F946FFCF03EAFC44FC875609" box="[487,819,1625,1649]" class="Malacostraca" family="Pseudothelphusidae" genus="Brasiliothelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="tapajoense">
<emphasis id="B922EAE3F946FFCF03EAFC44FC875609" box="[487,819,1625,1649]" italics="true" pageId="2" pageNumber="96">Brasiliothelphusa tapajoense</emphasis>
</taxonomicName>
: male
<typeStatus id="54ED8853F946FFCF0184FC44FC5A560A" box="[905,1006,1625,1650]" pageId="2" pageNumber="96" type="holotype">holotype</typeStatus>
,
<collectingCountry id="F3417661F946FFCF01F3FC44FBF1560A" box="[1022,1093,1625,1650]" name="Brazil" pageId="2" pageNumber="96">Brazil</collectingCountry>
,
<collectingRegion id="4992F813F946FFCF0658FC47FB3E560A" box="[1109,1162,1626,1650]" country="Brazil" name="Para" pageId="2" pageNumber="96">Pará</collectingRegion>
,
<collectingMunicipality id="6B8DAC8BF946FFCF0694FC47FA81560A" box="[1177,1333,1625,1650]" pageId="2" pageNumber="96">Monte Cristo</collectingMunicipality>
,
<location id="8E89602AF946FFCF0748FC44FF7A56EE" LSID="urn:lsid:plazi:treatment:03FF87E7F946FFCB029AFE9BFD6B51FD:8E89602AF946FFCF0748FC44FF7A56EE" country="Brazil" county="Comparative" municipality="Monte Cristo" name="Tapajos river" pageId="2" pageNumber="96" stateProvince="Para">Tapajós river</location>
,
<collectorName id="26A35327F946FFCF02D6FC60FD3456EE" box="[219,640,1661,1686]" pageId="2" pageNumber="96">Expedição Permanente da Amazônia</collectorName>
leg.
<collectingDate id="EFACE9D9F946FFCF00B1FC63FCC856EE" box="[700,892,1661,1686]" pageId="2" pageNumber="96" value="1973-07-20" valueMax="1973-07-28" valueMin="1973-07-20">
<date id="FFE81031F946FFCF00B1FC63FCC856EE" box="[700,892,1661,1686]" pageId="2" pageNumber="96" value="1973-07-20" valueMax="1973-07-28" valueMin="1973-07-20">2028 July 1973</date>
</collectingDate>
(
<specimenCode id="DBF09E8AF946FFCF0180FC63FB8756EE" box="[909,1075,1661,1686]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="96">
<collectionCode id="ED47AE34F946FFCF0180FC63FC5856EE" box="[909,1004,1662,1686]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="96">MZUSP</collectionCode>
6550
</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B3E3CACF946FFCF0645FC63FF6F56A6" ID-GBIF-Occurrence="1315974753" collectingDate="1973-07-20" collectingDateMax="1973-07-28" collectingDateMin="1973-07-20" collectionCode="MZUSP" collectorName="Expedicao Permanente da Amazonia" country="Brazil" location="Tapajos river" municipality="Monte Cristo" pageId="2" pageNumber="96" specimenCode="MZUSP 6378" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Para" typeStatus="paratype">
<specimenCount id="9D50FD78F946FFCF0645FC63FB1056EE" box="[1096,1188,1661,1686]" pageId="2" pageNumber="96" type="male">2 males</specimenCount>
,
<specimenCount id="9D50FD78F946FFCF06BCFC63FA9456EE" box="[1201,1312,1661,1686]" pageId="2" pageNumber="96" type="female">2 females</specimenCount>
<typeStatus id="54ED8853F946FFCF0727FC63FA2C56EE" box="[1322,1432,1662,1686]" pageId="2" pageNumber="96" type="paratype">paratypes</typeStatus>
,
<collectingCountry id="F3417661F946FFCF029AFCBCFF6A56C2" box="[151,222,1697,1722]" name="Brazil" pageId="2" pageNumber="96">Brazil</collectingCountry>
,
<collectingRegion id="4992F813F946FFCF02E0FCBFFE9656C2" box="[237,290,1698,1722]" country="Brazil" name="Para" pageId="2" pageNumber="96">Pará</collectingRegion>
,
<collectingMunicipality id="6B8DAC8BF946FFCF033DFCBFFE7856C2" box="[304,460,1697,1722]" pageId="2" pageNumber="96">Monte Cristo</collectingMunicipality>
,
<location id="8E89602AF946FFCF03D6FCBCFDC756C2" LSID="urn:lsid:plazi:treatment:03FF87E7F946FFCB029AFE9BFD6B51FD:8E89602AF946FFCF03D6FCBCFDC756C2" box="[475,627,1697,1722]" country="Brazil" municipality="Monte Cristo" name="Tapajos river" pageId="2" pageNumber="96" stateProvince="Para">Tapajós river</location>
,
<collectorName id="26A35327F946FFCF008FFCBCFB9D56C2" box="[642,1065,1697,1722]" pageId="2" pageNumber="96">Expedição Permanente da Amazônia</collectorName>
leg.
<collectingDate id="EFACE9D9F946FFCF0665FCBFFA9F56C2" box="[1128,1323,1697,1722]" pageId="2" pageNumber="96" value="1973-07-20" valueMax="1973-07-28" valueMin="1973-07-20">
<date id="FFE81031F946FFCF0665FCBFFA9F56C2" box="[1128,1323,1697,1722]" pageId="2" pageNumber="96" value="1973-07-20" valueMax="1973-07-28" valueMin="1973-07-20">2028 July 1973</date>
</collectingDate>
(
<specimenCode id="DBF09E8AF946FFCF0730FCBFFF6056A6" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="96">
<collectionCode id="ED47AE34F946FFCF0730FCBFFA2856C2" box="[1341,1436,1698,1722]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="96">MZUSP</collectionCode>
6378
</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE936F1F946FFCF02CAFCF4FBF3577A" blockId="2.[151,1437,1265,1974]" box="[199,1095,1769,1794]" pageId="2" pageNumber="96">
<emphasis id="B922EAE3F946FFCF02CAFCF4FED1577A" bold="true" box="[199,357,1769,1794]" pageId="2" pageNumber="96">Type locality.</emphasis>
Apiacás [9º34S, 57º23W], altitude
<quantity id="4CAE9B14F946FFCF010AFCF7FCE45779" box="[775,848,1770,1794]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="96" unit="m" value="220.0">220 m</quantity>
, Mato Grosso, Brazil.
</paragraph>
</subSubSection>
<subSubSection id="C34C657AF946FFCF02CAFD10FDB75732" pageId="2" pageNumber="96" type="distribution">
<paragraph id="8BE936F1F946FFCF02CAFD10FDB75732" blockId="2.[151,1437,1265,1974]" pageId="2" pageNumber="96">
<emphasis id="B922EAE3F946FFCF02CAFD10FED4575E" bold="true" box="[199,352,1805,1830]" pageId="2" pageNumber="96">Distribution.</emphasis>
Presently known only from northern
<collectingRegion id="4992F813F946FFCF011EFD13FC1D575E" box="[787,937,1806,1830]" country="Brazil" name="Mato Grosso" pageId="2" pageNumber="96">Mato Grosso</collectingRegion>
,
<collectingCountry id="F3417661F946FFCF01BAFD10FC4A575E" box="[951,1022,1805,1830]" name="Brazil" pageId="2" pageNumber="96">Brazil</collectingCountry>
, in the interfluvial area between the Juruena and São Manuel Rivers.
</paragraph>
</subSubSection>
<subSubSection id="C34C657AF946FFCF02CAFD48FF4E57CE" pageId="2" pageNumber="96" type="diagnosis">
<paragraph id="8BE936F1F946FFCF02CAFD48FF4E57CE" blockId="2.[151,1437,1265,1974]" pageId="2" pageNumber="96">
<emphasis id="B922EAE3F946FFCF02CAFD48FEF55716" bold="true" box="[199,321,1877,1902]" pageId="2" pageNumber="96">Diagnosis.</emphasis>
Mesial process of G1 large, subquadrate; mesial side of apical plate smooth, devoid of spines; subapical bulge noticeable in sternal view; subapical bulge and crest of sternal lobe both covered by abundant spinules.
</paragraph>
</subSubSection>
<caption id="DF296679F947FFCE029AFE24FF7154D2" httpUri="https://zenodo.org/record/159238/files/figure.png" pageId="3" pageNumber="97" targetBox="[162,1425,206,1045]" targetPageId="3">
<paragraph id="8BE936F1F947FFCE029AFE24FF7154D2" blockId="3.[151,1437,1081,1194]" pageId="3" pageNumber="97">
<emphasis id="B922EAE3F947FFCE029AFE24FEFF5436" bold="true" box="[151,331,1081,1103]" pageId="3" pageNumber="97">FIGURE 1 AE.</emphasis>
<taxonomicName id="4C564D72F947FFCE035FFE24FD815436" box="[338,565,1081,1102]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="97" phylum="Arthropoda" rank="species" species="apiaka" status="sp. nov.">
<emphasis id="B922EAE3F947FFCE035FFE24FD815436" box="[338,565,1081,1102]" italics="true" pageId="3" pageNumber="97">Melothelphusa apiaka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F947FFCE0031FE24FDC25437" box="[572,630,1081,1103]" pageId="3" pageNumber="97" rank="species">
<emphasis id="B922EAE3F947FFCE0031FE24FDFE5436" bold="true" box="[572,586,1081,1102]" pageId="3" pageNumber="97">n</emphasis>
.
<emphasis id="B922EAE3F947FFCE0055FE24FDC45436" bold="true" box="[600,624,1081,1102]" pageId="3" pageNumber="97">sp</emphasis>
.
</taxonomicNameLabel>
, male holotype, cl 18.8 mm, cw 27.8 mm (MZUSP 32914). A, habitus, dorsal view. B, cephalothorax, ventral view. C, cephalothorax, frontal view. D, E, lateral view of right and left chelipeds, respectively. Abbreviations: CxP4, coxa of pereopod 4; G1, gonopod 1; G2, gonopod 2; St48, thoracic sternites 4 to 8. Scale bars AE, 10 mm.
</paragraph>
</caption>
<subSubSection id="C34C657AF947FFC802CAFECFFA8A56A9" lastPageId="5" lastPageNumber="99" pageId="3" pageNumber="97" type="description">
<paragraph id="8BE936F1F947FFCE02CAFECFFD97560F" blockId="3.[151,1437,1234,2015]" pageId="3" pageNumber="97">
<emphasis id="B922EAE3F947FFCE02CAFECFFE5B5493" bold="true" box="[199,495,1234,1259]" pageId="3" pageNumber="97">
Description of
<typeStatus id="54ED8853F947FFCE038CFECFFE5E5493" box="[385,490,1234,1259]" pageId="3" pageNumber="97" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace cervical groove nearly straight, faint, ending just before anterolateral margin. Post-frontal lobules obsolete; median groove indistinct; fronto-orbital region width less than half maximum width of carapace (18.84/27.83). Anterolateral margin fringed with minute anteriorly-directed tubercles almost evenly spaced. Posterolateral margin, smooth, poorly defined. Frontal margin carinate, straight in frontal view. Supraorbital margin lined with small granules laterally. Infraorbital margin slightly crenulated, moderately marginated, straight mesially. Exorbital angle moderately prominent, obtuse. Inner orbital tooth poorly developed, narrow, subtriangular, lobe-like. Mxp3 dactylus rather flattened ventrally, mesial surface markedly setate; propodus and carpus subclinical, nearly equal in size. Mxp3 ischium subquadrate, mesial edge setate, dentate, teeth squarish, corneous; ischium lateral margin slightly convex. Mxp3 exopod noticeable, reaching to about 0,5 times length of lateral margin of ischium (left exopod lost in
<typeStatus id="54ED8853F947FFCE0092FC0BFCB25657" box="[671,774,1558,1583]" pageId="3" pageNumber="97" type="holotype">holotype</typeStatus>
). Epistome upper margin straight, strongly carinate; lower margin deeply sinuous to receive folded mxp3 palp, large triangular epistomial tooth in center filling gap between joining palps; scarcely pubescence.
</paragraph>
<paragraph id="8BE936F1F947FFCE02CAFC9EFD5257EF" blockId="3.[151,1437,1234,2015]" pageId="3" pageNumber="97">Merus of major cheliped subtriangular in cross-section, upper, mesial margins distinctly crenulated. Carpus upper surface smooth; mesial margin crenulated, strong acute tooth medially. Propodus massive, mesial, lateral surfaces smooth; dorsal, ventral surfaces with line of small tubercles. Movable finger slightly curved downwards, lateral, mesial, dorsal surfaces with punctations arranged in longitudinal subparallel lines. Cutting edges of fingers with small, massive teeth; fingers fully closing. P2P5 meri, carpi, propodi massive, dactyli slightly laterally compressed, with 5 longitudinal rows of strong acute spines: 3 dorsal, 2 ventral; propodi with minute scattered spines dorsally, ventrally, smooth mesially; carpi minutely spinous dorsally, punctate laterally, smooth mesially, ventrally; meri crenulate dorsally, otherwise smooth.</paragraph>
<paragraph id="8BE936F1F947FFC902CAFDBFFCC550AD" blockId="3.[151,1437,1234,2015]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="98" pageId="3" pageNumber="97">Thoracic sternal sutures 4/5 to 6/7 well marked, ending just before reaching median line of thoracic sternum; median line prolonging between sternites 7 and 8, deeper at interception with sternal suture 7/8; episternites 4-6 triangular posteriorly, episternite 7 posteriorly truncate. Sterno-abdominal cavity strongly pubescent; tubercle of abdominal holding system minute, placed near sternal suture 5/6.</paragraph>
<caption id="DF296679F940FFC9029AFD61FC325795" httpUri="https://zenodo.org/record/159239/files/figure.png" pageId="4" pageNumber="98" targetBox="[251,1316,295,1863]" targetPageId="4">
<paragraph id="8BE936F1F940FFC9029AFD61FC325795" blockId="4.[151,1437,1916,2029]" pageId="4" pageNumber="98">
<emphasis id="B922EAE3F940FFC9029AFD61FEFF57EA" bold="true" box="[151,331,1916,1938]" pageId="4" pageNumber="98">FIGURE 2 AC.</emphasis>
<taxonomicName id="4C564D72F940FFC9035CFD61FD8757E9" box="[337,563,1916,1937]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="98" phylum="Arthropoda" rank="species" species="apiaka" status="sp. nov.">
<emphasis id="B922EAE3F940FFC9035CFD61FD8757E9" box="[337,563,1916,1937]" italics="true" pageId="4" pageNumber="98">Melothelphusa apiaka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F940FFC90037FD61FDC757EA" box="[570,627,1916,1938]" pageId="4" pageNumber="98" rank="species">
<emphasis id="B922EAE3F940FFC90037FD61FDFC57E9" bold="true" box="[570,584,1916,1937]" pageId="4" pageNumber="98">n</emphasis>
.
<emphasis id="B922EAE3F940FFC90058FD61FDD857E9" bold="true" box="[597,620,1916,1937]" pageId="4" pageNumber="98">sp</emphasis>
.
</taxonomicNameLabel>
, male holotype, cl 18.8 mm, cw 27.8 mm (MZUSP 32914). Left first gonopod: A, abdominal view; B, lateral view; C, sternal view. Abbreviations: ap, apical plate; apa, abdominal process of apical plate; al, abdominal lobe; fas, field of apical spines; mp, marginal process; mep, mesial process; ml, marginal lobe; ms, marginal suture; oc, opening of spermatic channel; sap, subapical process. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF296679F941FFC8029AFE92FC5B54BC" httpUri="https://zenodo.org/record/159240/files/figure.png" pageId="5" pageNumber="99" targetBox="[184,1409,184,1141]" targetPageId="5">
<paragraph id="8BE936F1F941FFC8029AFE92FC5B54BC" blockId="5.[151,1437,1167,1220]" pageId="5" pageNumber="99">
<emphasis id="B922EAE3F941FFC8029AFE92FEFF54DC" bold="true" box="[151,331,1167,1189]" pageId="5" pageNumber="99">FIGURE 3 AD.</emphasis>
<taxonomicName id="4C564D72F941FFC8035CFE8DFD8754DD" box="[337,563,1168,1189]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="apiaka" status="sp. nov.">
<emphasis id="B922EAE3F941FFC8035CFE8DFD8754DD" box="[337,563,1168,1189]" italics="true" pageId="5" pageNumber="99">Melothelphusa apiaka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F941FFC80037FE8DFDC754DD" box="[570,627,1168,1189]" pageId="5" pageNumber="99" rank="species">
<emphasis id="B922EAE3F941FFC80037FE8DFDFC54DD" bold="true" box="[570,584,1168,1189]" pageId="5" pageNumber="99">n</emphasis>
.
<emphasis id="B922EAE3F941FFC80058FE8DFDD854DD" bold="true" box="[597,620,1168,1189]" pageId="5" pageNumber="99">sp</emphasis>
.
</taxonomicNameLabel>
, male holotype, cl 18.8 mm, cw 27.8 mm (MZUSP 32914). Left first gonopod: A, abdominal view. B, lateral view. C, sternal view. D, mesial view. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8BE936F1F941FFC802CAFEF6FEA75533" blockId="5.[151,1437,1259,2029]" pageId="5" pageNumber="99">Male abdomen triangular in shape, tapering progressively from third somite onward, margins scarcely pubescent, sutures between somites well distinct, complete. Telson broadly triangular, lateral margins distinctly crenulated.</paragraph>
<paragraph id="8BE936F1F941FFC802CAFF4BFEFD56D6" blockId="5.[151,1437,1259,2029]" pageId="5" pageNumber="99">G1 robust, proximal half slightly convex in lateral view. Marginal suture straight. Lateral suture deep, extending 2/3 of gonopod length. Marginal process short, strongly curved to spermatic channel, apex truncated. Field of apical spines open, almost as wide as long. Opening of spermatic channel delimited by crest of abdominal lobe and marginal process. Crest of sternal lobe strongly curved, covered with abundant small spinules. Apical plate short, subquadrate in abdominal view, mesial side smooth, devoid of spines, bearing two distinct lobes distally (abdominal, sternal); sternal lobe small, uniformly rounded; abdominal lobe narrow; apical plate provided with prominent transversal keel in abdominal view, partially covering marginal process in abdominal view. Subapical bulge noticeable in sternal view, rounded, covered with small spinules. Mesial process large, subquadrate, arising from sternal surface, continuous with apical plate, spine-like distally, slightly oriented to abdominal side.</paragraph>
<paragraph id="8BE936F1F941FFC802CAFCA4FA8A56A9" blockId="5.[151,1437,1259,2029]" box="[199,1342,1720,1745]" pageId="5" pageNumber="99">G2 distinctly slender, distal 3/4 tapered; tip flattened, with numerous short spinules on sternal surface.</paragraph>
</subSubSection>
<subSubSection id="C34C657AF941FFC802CAFCC1FEA75760" pageId="5" pageNumber="99" type="etymology">
<paragraph id="8BE936F1F941FFC802CAFCC1FEA75760" blockId="5.[151,1437,1259,2029]" pageId="5" pageNumber="99">
<emphasis id="B922EAE3F941FFC802CAFCC1FEFA568D" bold="true" box="[199,334,1756,1781]" pageId="5" pageNumber="99">Etymology.</emphasis>
The species is named after the Indian tribe
<taxonomicName id="4C564D72F941FFC80156FCC1FC1B568D" box="[859,943,1756,1781]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="apiaka" status="sp. nov.">Apiaká</taxonomicName>
. The specific epithet is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34C657AF941FFCB02CAFD3EFD6B51FD" lastPageId="6" lastPageNumber="100" pageId="5" pageNumber="99" type="discussion">
<paragraph id="8BE936F1F941FFCB02CAFD3EFD6B51FD" blockId="5.[151,1437,1259,2029]" lastBlockId="6.[151,1436,151,389]" lastPageId="6" lastPageNumber="100" pageId="5" pageNumber="99">
<emphasis id="B922EAE3F941FFC802CAFD3EFE8F5744" bold="true" box="[199,315,1827,1852]" pageId="5" pageNumber="99">Remarks.</emphasis>
<taxonomicName id="4C564D72F941FFC8034CFD3EFD8E5743" box="[321,570,1827,1851]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="apiaka" status="sp. nov.">
<emphasis id="B922EAE3F941FFC8034CFD3EFD8E5743" box="[321,570,1827,1851]" italics="true" pageId="5" pageNumber="99">Melothelphusa apiaka</emphasis>
</taxonomicName>
<emphasis id="B922EAE3F941FFC8004CFD3EFDCB5744" bold="true" box="[577,639,1827,1852]" pageId="5" pageNumber="99">
<taxonomicNameLabel id="A2115798F941FFC8004CFD3EFDCB5744" box="[577,639,1827,1852]" pageId="5" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C564D72F941FFC800B8FD39FC3B5743" box="[693,911,1827,1851]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="dardanelosensis" status="comb. nov.">
<emphasis id="B922EAE3F941FFC800B8FD39FC3B5743" box="[693,911,1827,1851]" italics="true" pageId="5" pageNumber="99">M. dardanelosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F941FFC80198FD3EFC4F5744" box="[917,1019,1827,1852]" pageId="5" pageNumber="99">
<emphasis id="B922EAE3F941FFC80198FD3EFC4F5744" bold="true" box="[917,1019,1827,1852]" pageId="5" pageNumber="99">n. comb.</emphasis>
</taxonomicNameLabel>
are unique in the tribe
<taxonomicName id="4C564D72F941FFC806F1FD3EFACB5744" box="[1276,1407,1827,1852]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="99" phylum="Arthropoda" rank="tribe" tribe="Kingsleyini">Kingsleyini</taxonomicName>
as follows: they both have a G1 with a short marginal process that is strongly curved toward the spermatic channel which has a truncated apex (fig 2B); field of apical spines of G1 semicircular in shape, open (figs. 2B; 3B); the opening of spermatic channel of G1 delimited by both the crest of abdominal lobe and marginal process (figs. 2B, C; 3B); and the apical plate provided with strong, keel-like process, running transversally on the abdominal surface of G1 (figs. 2A, B; 3A, B, D). However,
<taxonomicName id="4C564D72F941FFC8006AFDC8FD6E5794" box="[615,730,2004,2028]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="apiaka" status="sp. nov.">
<emphasis id="B922EAE3F941FFC8006AFDC8FD6E5794" box="[615,730,2004,2028]" italics="true" pageId="5" pageNumber="99">M. apiaka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F941FFC800ECFDC9FC955795" box="[737,801,2004,2029]" pageId="5" pageNumber="99" rank="species">
<emphasis id="B922EAE3F941FFC800ECFDC9FC955795" bold="true" box="[737,801,2004,2029]" pageId="5" pageNumber="99">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C564D72F941FFC801BAFDC8FB265794" box="[951,1170,2004,2028]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="99" phylum="Arthropoda" rank="species" species="dardanelosensis" status="comb. nov.">
<emphasis id="B922EAE3F941FFC801BAFDC8FB265794" box="[951,1170,2004,2028]" italics="true" pageId="5" pageNumber="99">M. dardanelosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F941FFC80697FDC9FAB55795" box="[1178,1281,2004,2029]" pageId="5" pageNumber="99">
<emphasis id="B922EAE3F941FFC80697FDC9FAB55795" bold="true" box="[1178,1281,2004,2029]" pageId="5" pageNumber="99">n. comb.</emphasis>
</taxonomicNameLabel>
in having: (i) the mesial process of G1 distinctly subquadrate (figs. 2C; 3C) (while the mesial process is subtriangular in
<taxonomicName id="4C564D72F942FFCB0770FA84FEF850AB" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="dardanelosensis" status="comb. nov.">
<emphasis id="B922EAE3F942FFCB0770FA84FEF850AB" italics="true" pageId="6" pageNumber="100">M. dardanelosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F942FFCB0358FAA6FE0A50AC" box="[341,446,187,212]" pageId="6" pageNumber="100">
<emphasis id="B922EAE3F942FFCB0358FAA6FE0A50AC" bold="true" box="[341,446,187,212]" pageId="6" pageNumber="100">n. comb.</emphasis>
</taxonomicNameLabel>
); (ii) the mesial side of apical plate of G1 devoid of spines (figs. 2A; 3A) (while the mesial side of apical plate is furnished with a distinct triangular spine in
<taxonomicName id="4C564D72F942FFCB01D8FAFDFB05508F" box="[981,1201,223,247]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="dardanelosensis" status="comb. nov.">
<emphasis id="B922EAE3F942FFCB01D8FAFDFB05508F" box="[981,1201,223,247]" italics="true" pageId="6" pageNumber="100">M. dardanelosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F942FFCB06B4FAC3FA94508F" box="[1209,1312,222,247]" pageId="6" pageNumber="100">
<emphasis id="B922EAE3F942FFCB06B4FAC3FA94508F" bold="true" box="[1209,1312,222,247]" pageId="6" pageNumber="100">n. comb.</emphasis>
</taxonomicNameLabel>
); (iii) well developed subapical bulge in sternal view of G1 (figs. 2B, C; 3BD) (while the subapical bulge is absent and subapical region is smooth in sternal view in
<taxonomicName id="4C564D72F942FFCB0080FB3AFCD35146" box="[653,871,294,318]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="dardanelosensis" status="comb. nov.">
<emphasis id="B922EAE3F942FFCB0080FB3AFCD35146" box="[653,871,294,318]" italics="true" pageId="6" pageNumber="100">M. dardanelosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F942FFCB0160FB38FC675146" box="[877,979,293,318]" pageId="6" pageNumber="100">
<emphasis id="B922EAE3F942FFCB0160FB38FC675146" bold="true" box="[877,979,293,318]" pageId="6" pageNumber="100">n. comb.</emphasis>
</taxonomicNameLabel>
); (iv) the crest of the sternal lobe and the subapical bulge of G1 abundantly covered with spinules (figs. 2 AC; 3AD) (while all regions of the G1 are devoid of spinules in
<taxonomicName id="4C564D72F942FFCB0385FB70FDD651FD" box="[392,610,365,389]" class="Malacostraca" family="Pseudothelphusidae" genus="Melothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="100" phylum="Arthropoda" rank="species" species="dardanelosensis" status="comb. nov.">
<emphasis id="B922EAE3F942FFCB0385FB70FDD651FD" box="[392,610,365,389]" italics="true" pageId="6" pageNumber="100">M. dardanelosensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2115798F942FFCB0064FB71FD7B51FD" box="[617,719,364,389]" pageId="6" pageNumber="100">
<emphasis id="B922EAE3F942FFCB0064FB71FD7B51FD" bold="true" box="[617,719,364,389]" pageId="6" pageNumber="100">n. comb.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
</treatment>
</document>