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<document id="4EB277D74A778466272BB74028F96B9B" ID-DOI="10.5281/zenodo.184304" ID-GBIF-Dataset="fe0befe7-de27-4b06-b1ea-0f7f79b6b024" ID-ISSN="1175-5326" ID-Zenodo-Dep="184304" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459967439071" checkinUser="plazi" docAuthor="Feitosa, Rodrigo M. &amp; Brandão, Carlos Roberto F." docDate="2008" docId="03FF87C1FF82D214FF79CBE0FC07ACDA" docLanguage="en" docName="zt01890p049.pdf" docOrigin="Zootaxa 1890" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Lachnomyrmex haskinsi Smith 1944" docType="treatment" docVersion="9" lastPageNumber="19" masterDocId="FFC6FFB9FF92D206FFEECF2FFFEBAF20" masterDocTitle="A taxonomic revision of the Neotropical myrmicine ant genus Lachnomyrmex Wheeler (Hymenoptera: Formicidae)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="17" updateTime="1698231238674" updateUser="plazi">
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<mods:title id="93CC374B63AC8CB9E30377723CAADC97">A taxonomic revision of the Neotropical myrmicine ant genus Lachnomyrmex Wheeler (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="1D56452D19B8F43C23D8C5F486E8385C">Feitosa, Rodrigo M.</mods:namePart>
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<mods:namePart id="258472901987128B20440659ABEE8AFB">Brandão, Carlos Roberto F.</mods:namePart>
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<mods:date id="0A6930234B46C892244D968B633C8A30">2008</mods:date>
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<subSubSection id="C34C655CFF82D216FF79CBE0FEDBAA2C" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BE936D7FF82D216FF79CBE0FDB6ABC9" blockId="16.[151,605,1231,1292]" box="[151,605,1231,1257]" pageId="16" pageNumber="17">
<heading id="D0A181BBFF82D216FF79CBE0FDB6ABC9" bold="true" box="[151,605,1231,1257]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C564D54FF82D216FF79CBE0FDB6ABC9" ID-CoL="3RPV9" authority="Smith, 1944" authorityName="Smith" authorityYear="1944" box="[151,605,1231,1257]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="haskinsi">
<emphasis id="B922EAC5FF82D216FF79CBE0FDB6ABC9" bold="true" box="[151,605,1231,1257]" pageId="16" pageNumber="17">
<emphasis id="B922EAC5FF82D216FF79CBE0FE29ABC9" bold="true" box="[151,450,1231,1257]" italics="true" pageId="16" pageNumber="17">Lachnomyrmex haskinsi</emphasis>
<bibRefCitation id="EFC74B26FF82D216FE27CBE0FDB6ABC9" author="Smith" box="[457,605,1231,1257]" pageId="16" pageNumber="49" refString="Smith, M. R. (1944) The genus Lachnomyrmex, with the description of a second species. Proceedings of the Entomological Society of Washington, 46, 225 - 228." type="journal article" year="1944">Smith, 1944</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE936D7FF82D216FF79CBDDFEDBAA2C" blockId="16.[151,605,1231,1292]" box="[151,304,1266,1292]" pageId="16" pageNumber="17">
<figureCitation id="136D2A52FF82D216FF79CBDDFEEDAA2C" box="[151,262,1266,1292]" captionStart="FIGURE 5" captionStartId="17.[151,255,1793,1817]" captionTargetBox="[304,1283,197,1768]" captionTargetId="figure@17.[304,1283,197,1769]" captionTargetPageId="17" captionText="FIGURE 5. Worker of Lachnomyrmex haskinsi from Heredia, Costa Rica: A, head in full face view; B, lateral view. Image by John Longino, specimen INBIOCRI 001237374." httpUri="https://zenodo.org/record/184309/files/figure.png" pageId="16" pageNumber="17">Figures 5</figureCitation>
,
<figureCitation id="136D2A52FF82D216FEFDCBDDFEDBAA2C" box="[275,304,1266,1292]" captionStart="FIGURE 19" captionStartId="45.[151,255,1922,1946]" captionTargetBox="[301,1287,194,1897]" captionTargetId="figure@45.[300,1288,194,1898]" captionTargetPageId="45" captionText="FIGURE 19. Distribution maps for the Lachnomyrmex species (L. haskinsi, L. mackayi, L. nordestinus, L. plaumanni, L. regularis, and L. scrobiculatus)." httpUri="https://zenodo.org/record/184322/files/figure.png" pageId="16" pageNumber="17">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34C655CFF82D216FF79CA6DFCA1AA59" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BE936D7FF82D216FF79CA6DFCA1AA59" blockId="16.[151,1436,1346,1401]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB4611F9FF82D216FF79CA6DFCA1AA59" pageId="16" pageNumber="17">
<taxonomicName id="4C564D54FF82D216FF79CA6DFDB5AA79" ID-CoL="3RPV9" authority="Smith, 1944: 227" authorityName="Smith" authorityPageNumber="227" authorityYear="1944" box="[151,606,1346,1369]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="haskinsi">
<emphasis id="B922EAC5FF82D216FF79CA6DFE70AA79" box="[151,411,1346,1369]" italics="true" pageId="16" pageNumber="17">Lachnomyrmex haskinsi</emphasis>
<treatmentCitation id="0AF710C6FF82D216FE4FCA6DFDB5AA79" author="Smith" box="[417,606,1346,1369]" page="227" pageId="16" pageNumber="17" year="1944">
<bibRefCitation id="EFC74B26FF82D216FE4FCA6DFDC3AA79" author="Smith" box="[417,552,1346,1369]" pageId="16" pageNumber="49" refString="Smith, M. R. (1944) The genus Lachnomyrmex, with the description of a second species. Proceedings of the Entomological Society of Washington, 46, 225 - 228." type="journal article" year="1944">Smith, 1944</bibRefCitation>
: 227
</treatmentCitation>
</taxonomicName>
, figs. 12.
<typeStatus id="54ED8875FF82D216FD3BCA6DFCD3AA79" box="[725,824,1346,1369]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
worker,
<collectingCountry id="F3417647FF82D216FC77CA6DFC19AA79" box="[921,1010,1346,1369]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
: Canal Zone, Barro Colorado Island, iiiii.1994, J. Zetek col., no. 5121/56906 [USNM] (examined).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C655CFF82D216FF79CA85FB54A934" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE936D7FF82D216FF79CA85FB54A934" blockId="16.[151,1437,1450,1996]" pageId="16" pageNumber="17">
<emphasis id="B922EAC5FF82D216FF79CA85FE97AAE4" bold="true" box="[151,380,1450,1476]" pageId="16" pageNumber="17">Worker diagnosis.</emphasis>
Body covered by continuous vermiculate rugae; petiolar node considerably elevated and subtriangular; dorsum of postpetiole with about six long hairs, and with a raised prominence medially; subpostpetiolar process present and well developed; first tergite of gaster devoid of long hairs.
</paragraph>
</subSubSection>
<subSubSection id="C34C655CFF82D216FF28C90DFD37A944" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BE936D7FF82D216FF28C90DFD37A944" blockId="16.[151,1437,1450,1996]" pageId="16" pageNumber="17">
<emphasis id="B922EAC5FF82D216FF28C90DFE10A91C" bold="true" box="[198,507,1570,1596]" pageId="16" pageNumber="17">
<typeStatus id="54ED8875FF82D216FF28C90DFED3A91C" box="[198,312,1570,1596]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
measurements.
</emphasis>
HL 0.60; HW 0.55; ML 0.22; SL 0.38; EL 0.09; WL 0.68; PSL 0.11; PL 0.24; PPL 0.13; GL 0.79; TL 2.65; CI 92; SI 68; OI 17.
</paragraph>
</subSubSection>
<subSubSection id="C34C655CFF82D214FF28C95DFE7DAE82" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE936D7FF82D216FF28C95DFE8CA9FC" blockId="16.[151,1437,1450,1996]" pageId="16" pageNumber="17">
<emphasis id="B922EAC5FF82D216FF28C95DFE0DA9AC" bold="true" box="[198,486,1650,1676]" pageId="16" pageNumber="17">Worker measurements</emphasis>
(n=2). HL 0.610.65; HW 0.600.63; ML 0.210.24; SL 0.39; EL 0.110.13; WL 0.710.74; PSL 0.160.17; PL 0.250.30; PPL 0.130.14; GL 0.790.80; TL 2.702.94; CI 9798; SI 6266; OI 1721.
</paragraph>
<paragraph id="8BE936D7FF82D216FF28C9C5FAE2A8EC" blockId="16.[151,1437,1450,1996]" pageId="16" pageNumber="17">
<emphasis id="B922EAC5FF82D216FF28C9C5FE23A824" bold="true" box="[198,456,1770,1796]" pageId="16" pageNumber="17">Worker description.</emphasis>
Color red ferruginous to brown, with lighter appendages. Head and mesosoma finely covered by continuous vermiculate rugae, transverse on anterior portion of pronotum, grading to longitudinal on the rest of promesonotum, mesopleura and lateral faces of propodeum; mandibles with few and short longitudinal striae restricted to basal portion; petiole and postpetiole finely and irregularly rugose. Abundant pilosity on head and dorsum of promesonotum; dorsum of petiolar node and postpetiole with about three and six long hairs, respectively; first tergite of gaster entirely devoid of long flexuous hairs.
</paragraph>
<caption id="DF29665FFF83D217FF79C82EFCEFA81B" httpUri="https://zenodo.org/record/184309/files/figure.png" pageId="17" pageNumber="18" targetBox="[304,1283,197,1768]" targetPageId="17">
<paragraph id="8BE936D7FF83D217FF79C82EFCEFA81B" blockId="17.[151,1436,1793,1851]" pageId="17" pageNumber="18">
<emphasis id="B922EAC5FF83D217FF79C82EFEF7A839" bold="true" box="[151,284,1793,1817]" pageId="17" pageNumber="18">FIGURE 5.</emphasis>
Worker of
<taxonomicName id="4C564D54FF83D217FE72C82DFD4AA839" box="[412,673,1794,1817]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="haskinsi">
<emphasis id="B922EAC5FF83D217FE72C82DFD4AA839" box="[412,673,1794,1817]" italics="true" pageId="17" pageNumber="18">Lachnomyrmex haskinsi</emphasis>
</taxonomicName>
from Heredia, Costa Rica: A, head in full face view; B, lateral view. Image by John Longino, specimen INBIOCRI001237374.
</paragraph>
</caption>
<paragraph id="8BE936D7FF83D214FF28C840FEE4AE22" blockId="17.[151,1436,1903,2009]" lastBlockId="18.[151,1437,152,1018]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">Head longer than broad, with vertexal margin convex; frontal lobes slightly rounded laterally; eyes with six to seven facets at greatest diameter. Promesonotum evenly convex in profile; metanotal groove relatively narrow and deeply impressed; propodeal spines straight; teeth of propodeal lobes widen basally, with about half-length of propodeal spines length. Petiolar node strongly elevated and subtriangular in lateral view; dorsum of postpetiole with a raised prominence medially and a well developed anteroventral projection directed anteriorly.</paragraph>
<paragraph id="8BE936D7FF80D214FF28CE20FE7DAE82" blockId="18.[151,1437,152,1018]" pageId="18" pageNumber="19">
<emphasis id="B922EAC5FF80D214FF28CE20FE1DAE09" bold="true" box="[198,502,271,297]" pageId="18" pageNumber="19">Gyne (first description).</emphasis>
Differing from worker by having about 12 facets at eyes maximum diameter; dorsum of petiolar node and postpetiole with about five and 10 long hairs, respectively; first tergite of gaster with around 10 long flexuous hairs on the anterior third, near postpetiolar insertion (see picture in the “Ants of
<collectingCountry id="F3417647FF80D214FF79CEA7FEF1AE82" box="[151,282,392,418]" name="Costa Rica" pageId="18" pageNumber="19">Costa Rica</collectingCountry>
” website).
</paragraph>
</subSubSection>
<subSubSection id="C34C655CFF80D214FF28CE80FC53AED2" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BE936D7FF80D214FF28CE80FC53AED2" blockId="18.[151,1437,152,1018]" pageId="18" pageNumber="19">
<emphasis id="B922EAC5FF80D214FF28CE80FEB8AEE9" bold="true" box="[198,339,431,457]" pageId="18" pageNumber="19">Etymology.</emphasis>
Dr. Marion
<bibRefCitation id="EFC74B26FF80D214FE0ACE9FFD4CAEEA" author="Smith" box="[484,679,432,458]" pageId="18" pageNumber="49" refString="Smith, M. R. (1944) The genus Lachnomyrmex, with the description of a second species. Proceedings of the Entomological Society of Washington, 46, 225 - 228." type="journal article" year="1944">M. Smith (1944)</bibRefCitation>
named this species in honor of Dr. Caryl Parker Haskins (
<date id="FFE81017FF80D214FABFCE9FFF3DAED2" pageId="18" pageNumber="19" value="1908" valueMax="2001">1908 2001</date>
), in recognition for his contribution to the study of ant biology.
</paragraph>
</subSubSection>
<subSubSection id="C34C655CFF80D214FF28CED0FD4EAC12" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BE936D7FF80D214FF28CED0FC5DADB2" blockId="18.[151,1437,152,1018]" pageId="18" pageNumber="19">
<emphasis id="B922EAC5FF80D214FF28CED0FEBFAD39" bold="true" box="[198,340,511,537]" pageId="18" pageNumber="19">Comments.</emphasis>
This species is most similar to
<taxonomicName id="4C564D54FF80D214FD2CCD2FFC0CAD39" box="[706,999,512,537]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="regularis">
<emphasis id="B922EAC5FF80D214FD2CCD2FFC0CAD39" box="[706,999,512,537]" italics="true" pageId="18" pageNumber="19">Lachnomyrmex regularis</emphasis>
</taxonomicName>
, but can be easily distinguished from the latter by the smaller size, more irregular rugulation, and presence of fine short rugae on postpetiole dorsum. Workers of
<taxonomicName id="4C564D54FF80D214FE8FCD7FFE17AD49" box="[353,508,592,617]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="B922EAC5FF80D214FE8FCD7FFE17AD49" box="[353,508,592,617]" italics="true" pageId="18" pageNumber="19">L. fernandezi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C564D54FF80D214FDDBCD7FFD5DAD49" box="[565,694,592,617]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="mackayi">
<emphasis id="B922EAC5FF80D214FDDBCD7FFD5DAD49" box="[565,694,592,617]" italics="true" pageId="18" pageNumber="19">L. mackayi</emphasis>
</taxonomicName>
can also be confounded with
<taxonomicName id="4C564D54FF80D214FBF8CD7FFB7DAD49" box="[1046,1174,592,617]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="haskinsi">
<emphasis id="B922EAC5FF80D214FBF8CD7FFB7DAD49" box="[1046,1174,592,617]" italics="true" pageId="18" pageNumber="19">L. haskinsi</emphasis>
</taxonomicName>
, but they present long flexuous hairs at least on the anterior third of the first gastral tergite.
</paragraph>
<paragraph id="8BE936D7FF80D214FF28CD8FFD4EAC12" blockId="18.[151,1437,152,1018]" pageId="18" pageNumber="19">
<taxonomicName id="4C564D54FF80D214FF28CD8FFE09AD99" box="[198,482,672,697]" class="Insecta" family="Formicidae" genus="Lachnomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="haskinsi">
<emphasis id="B922EAC5FF80D214FF28CD8FFE09AD99" box="[198,482,672,697]" italics="true" pageId="18" pageNumber="19">Lachnomyrmex haskinsi</emphasis>
</taxonomicName>
has been recorded from
<collectingCountry id="F3417647FF80D214FCEBCD8FFC6FAD9A" box="[773,900,672,698]" name="Costa Rica" pageId="18" pageNumber="19">Costa Rica</collectingCountry>
and
<collectingCountry id="F3417647FF80D214FC53CD8FFBF6AD9A" box="[957,1053,672,698]" name="Panama" pageId="18" pageNumber="19">Panama</collectingCountry>
, in
<quantity id="4CAE9B32FF80D214FBA9CD8FFB2FAD9A" box="[1095,1220,672,698]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="18" pageNumber="19" unit="m" value="500.0" valueMax="800.0" valueMin="200.0">200800m</quantity>
elevation wet forests. Workers are usually collected in samples of sifted leaf litter. Dr. John Longino observed a few workers, one with a larva, scattered throughout the debris of a
<taxonomicName id="4C564D54FF80D214FCE4CDDFFC50AC29" box="[778,955,752,777]" class="Insecta" family="Formicidae" genus="Brachymyrmex" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAC5FF80D214FCE4CDDFFC50AC29" box="[778,955,752,777]" italics="true" pageId="18" pageNumber="19">Brachymyrmex</emphasis>
</taxonomicName>
nest he was collecting from beneath rotten wood on the forest floor (
<bibRefCitation id="EFC74B26FF80D214FE02CC37FD72AC12" author="Longino" box="[492,665,792,818]" pageId="18" pageNumber="49" refString="Longino, J. T. (2007) Ants of Costa Rica. Available from: http: // www. evergreen. edu / ants / AntsofCostaRica. html (accessed: February 2008)." type="book chapter" year="2007">Longino 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C655CFF80D214FF28CC10FC07ACDA" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BE936D7FF80D214FF28CC10FC07ACDA" blockId="18.[151,1437,152,1018]" pageId="18" pageNumber="19">
<emphasis id="B922EAC5FF80D214FF28CC10FC91AC79" bold="true" box="[198,890,831,857]" pageId="18" pageNumber="19">
Additional material examined.
<collectingCountry id="F3417647FF80D214FDA7CC10FCE8AC79" box="[585,771,831,857]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
: Alajuela
</emphasis>
: Río San Lorencito, 1990, P. Hammond col. (
<specimenCount id="9D50FD5EFF80D214FA61CC6FFF07ACA2" pageId="18" pageNumber="19" type="worker">1 worker</specimenCount>
) [
<collectionCode id="ED47AE12FF80D214FEEBCC47FE8DACA2" box="[261,358,872,898]" pageId="18" pageNumber="19">BMNH</collectionCode>
];
<emphasis id="B922EAC5FF80D214FE92CC48FE0BACA1" bold="true" box="[380,480,871,897]" pageId="18" pageNumber="19">Heredia</emphasis>
: La Selva Biological Station, CCL,
<geoCoordinate id="EE625010FF80D214FC70CC47FBEBACA2" box="[926,1024,872,898]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="925" value="10.716666">10º43N</geoCoordinate>
<geoCoordinate id="EE625010FF80D214FBE5CC47FB9FACA2" box="[1035,1140,872,898]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="925" value="-84.01667">84º01W</geoCoordinate>
,
<date id="FFE81017FF80D214FB6DCC47FAE7ACA2" box="[1155,1292,872,898]" pageId="18" pageNumber="19" value="1986-07-15">15.vii.1986</date>
, J. Longino col., no. INBIOCRI002280138 (
<specimenCount id="9D50FD5EFF80D214FDD5CCBFFD5AAC8A" box="[571,689,912,938]" pageId="18" pageNumber="19" type="worker">1 worker</specimenCount>
) [
<collectionCode id="ED47AE12FF80D214FD3DCCBFFCC9AC8A" box="[723,802,912,938]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="18" pageNumber="19">INBC</collectionCode>
]; same locality,
<date id="FFE81017FF80D214FBEDCCBFFB91AC8A" box="[1027,1146,912,938]" pageId="18" pageNumber="19" value="1991-10-23">23.x.1991</date>
, J. Longino col., no. INBIOCRI001237374 (1 gyne, examined by images) [
<collectionCode id="ED47AE12FF80D214FCC7CC97FC93ACF2" box="[809,888,952,978]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="18" pageNumber="19">INBC</collectionCode>
];
<emphasis id="B922EAC5FF80D214FC60CC98FC0AACF1" bold="true" box="[910,993,951,977]" pageId="18" pageNumber="19">Limón</emphasis>
: Cerro Cocori, Fca. de E. Rojas,
<date id="FFE81017FF80D214FA91CC97FEFCACDA" pageId="18" pageNumber="19" value="1992-11-09" valueMax="1992-11-30" valueMin="1992-11-09">9 30.xi.1992</date>
, E. Rojas col., no. INBIOCRI000900198 (
<specimenCount id="9D50FD5EFF80D214FCE2CCCFFC9EACDA" box="[780,885,992,1018]" pageId="18" pageNumber="19" type="worker">1 worker</specimenCount>
) [
<collectionCode id="ED47AE12FF80D214FC65CCCFFC0AACDA" box="[907,993,992,1018]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="18" pageNumber="19">MZSP</collectionCode>
].
</paragraph>
</subSubSection>
</treatment>
</document>