treatments-xml/data/03/B2/87/03B287E9FFA5FFBFFF0D67DBBA89FDAB.xml

<document id="245E56B2772F8A062088634737071F7A" ID-DOI="10.5281/zenodo.5377199" ID-ISSN="1638-9395" ID-Zenodo-Dep="5377199" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630282681036" checkinUser="marcus" docAuthor="Popov, Vasil V." docDate="2004" docId="03B287E9FFA5FFBFFF0D67DBBA89FDAB" docLanguage="en" docName="Geodiversitas.2004.26.3.403-491.pdf" docOrigin="Geodiversitas 26 (3)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Rhagapodemus hautimagnensis Mein &amp; Michaux 1970" docType="treatment" docVersion="3" lastPageNumber="456" masterDocId="FF8BFF91FF97FF8AFFC5650BB968FFA1" masterDocTitle="Pliocene small mammals (Mammalia, Lipotyphla, Chiroptera, Lagomorpha, Rodentia) from Muselievo (North Bulgaria)" masterLastPageNumber="491" masterPageNumber="403" pageNumber="453" updateTime="1699212936602" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="86658E7B07AAD69509BC27018CF65C76">Pliocene small mammals (Mammalia, Lipotyphla, Chiroptera, Lagomorpha, Rodentia) from Muselievo (North Bulgaria)</mods:title>
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<mods:namePart id="A8C69443BD07B2E66A47C0A2ABAF3BA9">Popov, Vasil V.</mods:namePart>
<mods:affiliation id="430D604C791DBC49AD19A821BA3749DF">Institute of Zoology, Bulgarian Academy of Sciences, Boul. Tsar Osvoboditel 1, 1000 Sofia (Republic of Bulgaria) popov @ zoology. bas. bg</mods:affiliation>
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<mods:title id="F370D2C68DE14DC84799CD1EECD8B0A3">Geodiversitas</mods:title>
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<mods:date id="C241DA0931BFA740173F76CC4590F22A">2004</mods:date>
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<paragraph id="8BA436FFFFA5FFB8FF0D67DBB8B7FC8B" blockId="50.[200,570,720,810]" pageId="50" pageNumber="453">
<heading id="D0EC8193FFA5FFB8FF0D67DBB8B7FC8B" centered="true" fontSize="11" level="2" pageId="50" pageNumber="453" reason="2">
<taxonomicName id="4C1B4D7CFFA5FFB8FF0D67DBBB6CFCAB" authority="Mein &amp; Michaux, 1970" authorityName="Mein &amp; Michaux" authorityYear="1970" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="50" pageNumber="453" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA5FFB8FF0D67DBB802FD4B" bold="true" box="[200,362,720,746]" italics="true" pageId="50" pageNumber="453">Rhagapodemus</emphasis>
cf. 
<emphasis id="B96FEAEDFFA5FFB8FE5767DBBB52FD4B" bold="true" box="[402,570,720,746]" italics="true" pageId="50" pageNumber="453">hautimagnensis</emphasis>
<bibRefCitation id="EF8A4B0EFFA5FFB8FF3B67FBBB6CFCAB" author="MEIN P. &amp; MICHAUX J." box="[254,516,752,778]" pageId="50" pageNumber="453" pagination="2780 - 2783" refId="ref45227" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sud-occidentale. Comptes rendus de l'Academie des Sciences Paris 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux, 1970</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13202A7AFFA5FFB8FEEF661BB815FC8B" box="[298,381,784,810]" captionStart="FIG" captionStartId="51.[118,129,1646,1663]" captionTargetBox="[165,1165,281,1640]" captionTargetId="figure-20@51.[165,1165,281,1643]" captionTargetPageId="51" captionText="FIG. 26. — Rhagapodemus cf. hautimagnensis Mein &amp; Michaux, 1970, upper molars in occlusal view; A, right M1, Ms243; B, right M1, Ms244; C, right M1, Ms246; D, left M2, Ms253; E, right M2, Ms254; F, right M2, Ms255; G, right M3, Ms259; H, right M3, Ms260; I, right M3, Ms242. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5377243" httpUri="https://zenodo.org/record/5377243/files/figure.png" pageId="50" pageNumber="453">Figs 26</figureCitation>
; 
<figureCitation id="13202A7AFFA5FFB8FE4C661BB8CCFC8B" box="[393,420,784,810]" captionStart="FIG" captionStartId="52.[119,130,1646,1663]" captionTargetBox="[161,1170,225,1614]" captionTargetId="figure-18@52.[161,1170,225,1614]" captionTargetPageId="52" captionText="FIG. 27. — A-H, Rhagapodemus cf. hautimagnensis Mein &amp; Michaux, 1970, lower molars in occlusal view; A, right m1, Ms233; B, right m1, Ms234; C, right m1, Ms229; D, left m2, Ms237; E, right m2, Ms238; F, right m2, Ms235; G, left m3, Ms261; H, left m3, Ms262; I, J, Rhagapodemus sp., M1 in occlusal view; I, left, Ms302; J, right, Ms303. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5377245" httpUri="https://zenodo.org/record/5377245/files/figure.png" pageId="50" pageNumber="453">27</figureCitation>
A-H)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016574FFA5FFB8FFB26642B988FC35" pageId="50" pageNumber="453" type="reference_group">
<paragraph id="8BA436FFFFA5FFB8FFB26642B988FC35" blockId="50.[119,652,841,1073]" pageId="50" pageNumber="453">
<treatmentCitationGroup id="AB0B11D1FFA5FFB8FFB26642B988FC35" pageId="50" pageNumber="453">
<treatmentCitation id="0ABA10EEFFA5FFB8FFB26642B985FCDB" author="MEIN P. &amp; MICHAUX J." page="2782" pageId="50" pageNumber="453" year="1970">
<taxonomicName id="4C1B4D7CFFA5FFB8FFB26642B985FCDB" authority="Mein &amp; Michaux, 1970: 2782" authorityName="Mein &amp; Michaux" authorityPageNumber="2782" authorityYear="1970" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="50" pageNumber="453" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA5FFB8FFB26642B8D3FCC0" box="[119,443,841,865]" italics="true" pageId="50" pageNumber="453">Rhagapodemus hautimagnensis</emphasis>
<bibRefCitation id="EF8A4B0EFFA5FFB8FE016642B985FCDB" author="MEIN P. &amp; MICHAUX J." pageId="50" pageNumber="453" pagination="2780 - 2783" refId="ref45227" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sud-occidentale. Comptes rendus de l'Academie des Sciences Paris 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux, 1970: 2782</bibRefCitation>
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. — 
<treatmentCitation id="0ABA10EEFFA5FFB8FEDD666ABB70FCDB" author="VAN DE WEERD A." box="[280,536,865,890]" page="140" pageId="50" pageNumber="453" year="1979">
<bibRefCitation id="EF8A4B0EFFA5FFB8FEDD666ABB70FCDB" author="VAN DE WEERD A." box="[280,536,865,890]" pageId="50" pageNumber="453" pagination="127 - 168" refId="ref48078" refString="VAN DE WEERD A. 1979. - Early Ruscinian rodents and lagomorphs (Mammalia) from the lignites near Ptolemais (Macedonia, Greece). Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B, 82 (2): 127 - 168." type="journal article" year="1979">
<collectingRegion id="49DFF81DFFA5FFB8FEDD666AB829FCD9" box="[280,321,865,888]" country="Turkey" name="Van" pageId="50" pageNumber="453">Van</collectingRegion>
de Weerd 1979: 140
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, 141, pl. 2, figs 10-14.
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</paragraph>
</subSubSection>
<subSubSection id="C3016574FFA5FFB8FFB266A8BB6EFBA9" pageId="50" pageNumber="453" type="materials_examined">
<paragraph id="8BA436FFFFA5FFB8FFB266A8BB6EFBA9" blockId="50.[119,652,841,1073]" pageId="50" pageNumber="453">
<collectionCode id="ED0AAE3AFFA5FFB8FFB266A8B82CFC18" box="[119,324,931,954]" pageId="50" pageNumber="453">MATERIAL EXAMINED</collectionCode>
. — 
<quantity id="4CE39B1AFFA5FFB8FEB566AFB8CFFC1D" box="[368,423,932,956]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="50" pageNumber="453" unit="m" value="23.0">23 m</quantity>
1 (Ms223-235), 1 mandibular fragment (Ms236), 
<quantity id="4CE39B1AFFA5FFB8FE4666B6B8DFFC74" box="[387,439,957,981]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="453" unit="m" value="10.0">10 m</quantity>
2 (Ms235-241), 
<quantity id="4CE39B1AFFA5FFB8FD9D66B6BBE9FC74" box="[600,641,957,981]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="453" unit="m" value="5.0">5 m</quantity>
3 (Ms261-263), 18 M1 (Ms243-251, 264), 7 M2 (Ms252-258), 3 M3 (Ms242, 259, 260).
</paragraph>
</subSubSection>
<subSubSection id="C3016574FFA5FFB8FFB26113BA69FA8B" pageId="50" pageNumber="453" type="description">
<paragraph id="8BA436FFFFA5FFB8FFB26113B8D1FB90" blockId="50.[119,652,841,1073]" box="[119,441,1048,1073]" pageId="50" pageNumber="453">
MEASUREMENTS. — See 
<tableCitation id="C6990344FFA5FFB8FEAD6112B8D1FB90" box="[360,441,1049,1073]" captionStart="TABLE" captionStartId="50.[119,130,206,223]" captionText="TABLE 9. — Molar measurements (mm) of Rhagapodemus cf. hautimagnensis Mein &amp; Michaux, 1970 from Muselievo. For abbreviations see Material and methods." pageId="50" pageNumber="453">Table 9.</tableCitation>
</paragraph>
<paragraph id="8BA436FFFFA5FFB8FFB26164B86FFB26" blockId="50.[119,652,1135,1706]" box="[119,263,1135,1161]" pageId="50" pageNumber="453">DESCRIPTION</paragraph>
<paragraph id="8BA436FFFFA5FFB8FFB2619BB841F908" blockId="50.[119,652,1135,1706]" pageId="50" pageNumber="453">m1: the anterocentral tubercle is fairly well developed and isolated on unworn teeth. In the latter stages of wear, it fuses with the anterior pair of cusps. In unworn teeth, the anterior and middle pairs of cusps are separated by a narrow valley. In some worn specimens, they are connected by a sagital pass. The posterior lamina is always isolated. The talonid is of medium size, oval and slightly elongated transversally. The labial accessory cusps are variable in size and number. The posterior one is always well developed, rounded and quickly becomes confluent with the hypoconid with wear. The other accessory cusps may lack, but more often, they are two, adjacent to the anterior and the middle pairs of cusps, respectively. In some specimens additional weak buds may occur.</paragraph>
<paragraph id="8BA436FFFFA5FFB8FD6F677BBA68FD6B" blockId="50.[682,1216,623,1322]" pageId="50" pageNumber="453">m2: the accessory labial cusps, beside the anterior one, are poor (near the hypoconid) or lacking.</paragraph>
<paragraph id="8BA436FFFFA5FFB8FD6F67DBBD2CFD48" blockId="50.[682,1216,623,1322]" box="[682,1092,719,746]" pageId="50" pageNumber="453">m3: there is a weak anterior add cusp.</paragraph>
<paragraph id="8BA436FFFFA5FFB8FD6F67FBBA39FBAB" blockId="50.[682,1216,623,1322]" pageId="50" pageNumber="453">
M1: these molars possess three roots. The 
<emphasis id="B96FEAEDFFA5FFB8FB4B67E5BDCDFCA9" box="[1166,1189,750,776]" italics="true" pageId="50" pageNumber="453">t1</emphasis>
is considerably pushed to the back and is separated from the 
<emphasis id="B96FEAEDFFA5FFB8FCD66625BA41FCE9" box="[787,809,814,840]" italics="true" pageId="50" pageNumber="453">t2</emphasis>
. In some specimens 
<emphasis id="B96FEAEDFFA5FFB8FBD76625BD40FCE9" box="[1042,1064,814,840]" italics="true" pageId="50" pageNumber="453">t3</emphasis>
has a labially directed spur reaching the base of the 
<emphasis id="B96FEAEDFFA5FFB8FB9B665BBD1CFCCB" box="[1118,1140,848,874]" italics="true" pageId="50" pageNumber="453">t6</emphasis>
. Most often, the 
<emphasis id="B96FEAEDFFA5FFB8FCDB6665BA5CFC29" box="[798,820,878,904]" italics="true" pageId="50" pageNumber="453">t4</emphasis>
and 
<emphasis id="B96FEAEDFFA5FFB8FCB56664BAEEFC28" box="[880,902,879,905]" italics="true" pageId="50" pageNumber="453">t7</emphasis>
are not connected until the late stages of wear. Sometimes the 
<emphasis id="B96FEAEDFFA5FFB8FBEA6685BD2CFC09" box="[1071,1092,910,936]" italics="true" pageId="50" pageNumber="453">t8</emphasis>
and 
<emphasis id="B96FEAEDFFA5FFB8FB456685BDFDFC09" box="[1152,1173,910,936]" italics="true" pageId="50" pageNumber="453">t9</emphasis>
are also separated. The 
<emphasis id="B96FEAEDFFA5FFB8FC4466A4BAFFFC68" box="[897,919,943,969]" italics="true" pageId="50" pageNumber="453">t7</emphasis>
is elongated and looks like a broad ridge. The posterior cingulum (
<emphasis id="B96FEAEDFFA5FFB8FBB366C5BDF3FC49" box="[1142,1179,974,1000]" italics="true" pageId="50" pageNumber="453">t12</emphasis>
) is well developed.
</paragraph>
<paragraph id="8BA436FFFFA5FFB8FD6F611BBA26FB6B" blockId="50.[682,1216,623,1322]" pageId="50" pageNumber="453">
M2: the majority of teeth have three roots, but some specimens show 4 or 5 roots. The 
<emphasis id="B96FEAEDFFA5FFB8FB496125BDCBFBE9" box="[1164,1187,1070,1096]" italics="true" pageId="50" pageNumber="453">t1</emphasis>
is larger than the 
<emphasis id="B96FEAEDFFA5FFB8FC896145BA0AFBC9" box="[844,866,1102,1128]" italics="true" pageId="50" pageNumber="453">t3</emphasis>
and they are set apart. The pairs of adjacent cusps, 
<emphasis id="B96FEAEDFFA5FFB8FCBF6165BAC7FB29" box="[890,943,1134,1160]" italics="true" pageId="50" pageNumber="453">t4/t7</emphasis>
and 
<emphasis id="B96FEAEDFFA5FFB8FC2F6165BD76FB29" box="[1002,1054,1134,1160]" italics="true" pageId="50" pageNumber="453">t8/t9</emphasis>
, are separated during the early and middle wearing stages. The 
<emphasis id="B96FEAEDFFA5FFB8FD6F61A4BBA8FB68" box="[682,704,1199,1225]" italics="true" pageId="50" pageNumber="453">t7</emphasis>
is elongated.
</paragraph>
<paragraph id="8BA436FFFFA5FFB8FD6F61DBBA69FA8B" blockId="50.[682,1216,623,1322]" pageId="50" pageNumber="453">
M3: the tooth is three rooted. The 
<emphasis id="B96FEAEDFFA5FFB8FBF661C5BD21FB49" box="[1075,1097,1230,1256]" italics="true" pageId="50" pageNumber="453">t3</emphasis>
is lacking. The other cusps are connected in various combinations.
</paragraph>
</subSubSection>
<subSubSection id="C3016574FFA5FFBFFD6F6044BA89FDAB" lastPageId="53" lastPageNumber="456" pageId="50" pageNumber="453" type="discussion">
<paragraph id="8BA436FFFFA5FFB8FD6F6044BA66FAC6" blockId="50.[682,1215,1359,1706]" box="[682,782,1359,1385]" pageId="50" pageNumber="453">REMARKS</paragraph>
<paragraph id="8BA436FFFFA5FFBFFD6F607BBB1EFC2B" blockId="50.[682,1215,1359,1706]" lastBlockId="53.[118,652,208,1706]" lastPageId="53" lastPageNumber="456" pageId="50" pageNumber="453">
Some species of large murids, showing pronounced hypsodonty and presence of 
<emphasis id="B96FEAEDFFA5FFB8FB976084BD00FA08" box="[1106,1128,1423,1449]" italics="true" pageId="50" pageNumber="453">t7</emphasis>
on M1 and M2, are known from Europe. The first described species is 
<taxonomicName id="4C1B4D7CFFA5FFB8FC6060DBBA29F9AB" authority="Kretzoi, 1959" authorityName="Kretzoi" authorityYear="1959" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="50" pageNumber="453" phylum="Chordata" rank="species" species="frequens">
<emphasis id="B96FEAEDFFA5FFB8FC6060DBBDD6FA49" box="[933,1214,1486,1514]" italics="true" pageId="50" pageNumber="453">Rhagapodemus frequens</emphasis>
<bibRefCitation id="EF8A4B0EFFA5FFB8FD6F60FBBA29F9AB" author="KRETZOI M." box="[682,833,1520,1546]" pageId="50" pageNumber="453" pagination="237 - 246" refId="ref44793" refString="KRETZOI M. 1959. - Insectivoren, Nagetiere und Lagomorphen der jungstpliozanen Fauna von Csarnota im Villayer Gebirge (Sudungarn). Vertebrata Hungarica 1 (2): 237 - 246." type="journal article" year="1959">Kretzoi, 1959</bibRefCitation>
</taxonomicName>
, from the late Pliocene of Csarnóta (Carpathian basin, 
<collectingCountry id="F30C766FFFA5FFB8FCBE6304BA88F988" box="[891,992,1551,1577]" name="Hungary" pageId="50" pageNumber="453">Hungary</collectingCountry>
). This species is also recorded in 
<collectingCountry id="F30C766FFFA5FFB8FCFD633BBAE1F9EB" box="[824,905,1584,1610]" name="Poland" pageId="50" pageNumber="453">Poland</collectingCountry>
(Weze 1, 
<bibRefCitation id="EF8A4B0EFFA5FFB8FBC1633BBDDAF9EB" author="SULIMSKI A." box="[1028,1202,1584,1610]" pageId="50" pageNumber="453" pagination="149 - 261" refId="ref46924" refString="SULIMSKI A. 1964. - Pliocene Lagomorpha and Rodentia from Weze I. Acta Paleontologica Polonica 9 (2): 149 - 261." type="journal article" year="1964">Sulimski 1964</bibRefCitation>
). Subsequently, a more evolved form, 
<taxonomicName id="4C1B4D7CFFA5FFB8FBF8635BBDE4F92B" authority="De Bruijn &amp; Van der Meulen, 1975" authorityName="De Bruijn &amp; Van der Meulen" authorityYear="1975" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="50" pageNumber="453" phylum="Chordata" rank="subSpecies" species="frequens" subSpecies="athensis">
<emphasis id="B96FEAEDFFA5FFB8FBF8635BBB91F92B" italics="true" pageId="50" pageNumber="453">Rh. frequens athensis</emphasis>
<bibRefCitation id="EF8A4B0EFFA5FFB8FCC56364BDE4F92B" author="DE BRUIJN H. &amp; VAN DER MEULEN A. J." box="[768,1164,1647,1674]" pageId="50" pageNumber="453" pagination="314 - 338" refId="ref43216" refString="DE BRUIJN H. &amp; VAN DER MEULEN A. J. 1975. - The early Pleistocene Rodents from Tourkoubounia- 1 (Athens, Greece). I. Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B, 78 (4): 314 - 338." type="journal article" year="1975">De Bruijn &amp; Van der Meulen, 1975</bibRefCitation>
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, was reported from the south of 
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(the early Pleistocene locality of Tourkobounia). This form is considered as a separate species by 
<bibRefCitation id="EF8A4B0EFFA2FFBFFDFC65FBB8F8FE8B" author="MARTIN SUAREZ E. &amp; MEIN P." pageId="53" pageNumber="456" pagination="87 - 97" refId="ref45020" refString="MARTIN SUAREZ E. &amp; MEIN P. 1998. - Revision of the genera Parapodemus, Apodemus, Rhagamys and Rhagapodemus (Rodentia, Mammalia). Geobios 31 (1): 87 - 97." type="journal article" year="1998">Martin Suarez &amp; Mein (1998)</bibRefCitation>
. Two closely related species differing in size were described from 
<collectingCountry id="F30C766FFFA2FFBFFFB36444B9AEFEC8" box="[118,198,335,361]" name="France" pageId="53" pageNumber="456">France</collectingCountry>
: 
<taxonomicName id="4C1B4D7CFFA2FFBFFF11645BB877FE2B" authority="Mein &amp; Michaux, 1970" authorityName="Mein &amp; Michaux" authorityYear="1970" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA2FFBFFF11645BBB4DFECB" box="[212,549,336,362]" italics="true" pageId="53" pageNumber="456">Rhagapodemus hautimagnensis</emphasis>
<bibRefCitation id="EF8A4B0EFFA2FFBFFDEB645BB877FE2B" author="MEIN P. &amp; MICHAUX J." pageId="53" pageNumber="456" pagination="2780 - 2783" refId="ref45227" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sud-occidentale. Comptes rendus de l'Academie des Sciences Paris 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux, 1970</bibRefCitation>
</taxonomicName>
(large), and 
<taxonomicName id="4C1B4D7CFFA2FFBFFE6A647BB84AFE0B" authority="Mein &amp; Michaux, 1970" authorityName="Mein &amp; Michaux" authorityYear="1970" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="ballesoi">
<emphasis id="B96FEAEDFFA2FFBFFE6A647BBB40FE2B" box="[431,552,368,394]" italics="true" pageId="53" pageNumber="456">Rh. ballesoi</emphasis>
<bibRefCitation id="EF8A4B0EFFA2FFBFFDF4647BB84AFE0B" author="MEIN P. &amp; MICHAUX J." pageId="53" pageNumber="456" pagination="2780 - 2783" refId="ref45227" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sud-occidentale. Comptes rendus de l'Academie des Sciences Paris 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux, 1970</bibRefCitation>
</taxonomicName>
(small). The first species is also known from the early Ruscinian of 
<collectingCountry id="F30C766FFFA2FFBFFDC764BBBB25FE6B" box="[514,589,432,458]" name="Greece" pageId="53" pageNumber="456">Greece</collectingCountry>
(
<bibRefCitation id="EF8A4B0EFFA2FFBFFD9864BBB847FE4B" author="VAN DE WEERD A." pageId="53" pageNumber="456" pagination="127 - 168" refId="ref48078" refString="VAN DE WEERD A. 1979. - Early Ruscinian rodents and lagomorphs (Mammalia) from the lignites near Ptolemais (Macedonia, Greece). Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B, 82 (2): 127 - 168." type="journal article" year="1979">Van de Weerd 1979</bibRefCitation>
). Another species, 
<taxonomicName id="4C1B4D7CFFA2FFBFFDCE64DBBB30FDAB" authority="De Bruijn &amp; Van der Meulen, 1975" authorityName="De Bruijn &amp; Van der Meulen" authorityYear="1975" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="wandeweerdi">
<emphasis id="B96FEAEDFFA2FFBFFDCE64DBB9D2FDAB" italics="true" pageId="53" pageNumber="456">Rh. wandeweerdi</emphasis>
<bibRefCitation id="EF8A4B0EFFA2FFBFFF0664E4BB30FDAB" author="DE BRUIJN H. &amp; VAN DER MEULEN A. J." box="[195,600,495,522]" pageId="53" pageNumber="456" pagination="314 - 338" refId="ref43216" refString="DE BRUIJN H. &amp; VAN DER MEULEN A. J. 1975. - The early Pleistocene Rodents from Tourkoubounia- 1 (Athens, Greece). I. Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B, 78 (4): 314 - 338." type="journal article" year="1975">De Bruijn &amp; Van der Meulen, 1975</bibRefCitation>
</taxonomicName>
, was reported from the isle of Rhodes (the locality Maritsa). Initially this material was described under the name 
<taxonomicName id="4C1B4D7CFFA2FFBFFEEA675BB8F1FDCB" authorityName="Kaup" authorityYear="1829" box="[303,409,592,618]" class="Mammalia" family="Muridae" genus="Apodemus" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFA2FFBFFEEA675BB8F1FDCB" box="[303,409,592,618]" italics="true" pageId="53" pageNumber="456">Apodemus</emphasis>
</taxonomicName>
aff. 
<emphasis id="B96FEAEDFFA2FFBFFE0B6744BB73FDC9" box="[462,539,591,616]" italics="true" pageId="53" pageNumber="456">jeanteti</emphasis>
Michaux, 1967 (
<bibRefCitation id="EF8A4B0EFFA2FFBFFF79677BB8CDFD2B" author="DE BRUIJN H. &amp; DAWSON M. R. &amp; MEIN P." box="[188,421,623,650]" pageId="53" pageNumber="456" pagination="535 - 584" refId="ref43163" refString="DE BRUIJN H., DAWSON M. R. &amp; MEIN P. 1970. - Upper Pliocene Rodentia, Lagomorpha and Insectivora (Mammalia) from the isle of Rhodes (Greece). Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B 73 (5): 535 - 584." type="journal article" year="1970">
De Bruijn 
<emphasis id="B96FEAEDFFA2FFBFFEF56764B80BFD2B" box="[304,355,623,650]" italics="true" pageId="53" pageNumber="456">et al.</emphasis>
1970
</bibRefCitation>
). This short review of the Pliocene findings of the genus shows that it evolved in south and southeastern Europe. The main evolutionary trend is the increase of size and hypsodonty. The latest form, 
<taxonomicName id="4C1B4D7CFFA2FFBFFE7767FBBBEDFCAB" authorityName="De Bruijn &amp; Van der Meulen" authorityYear="1975" box="[434,645,750,778]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="subSpecies" species="frequens" subSpecies="athensis">
<emphasis id="B96FEAEDFFA2FFBFFE7767FBBBEDFCAB" box="[434,645,750,778]" italics="true" pageId="53" pageNumber="456">Rh. frequens athensis</emphasis>
</taxonomicName>
, is the largest one and has higher crowned molars. Therefore, it can be supposed that the forms of this phyletic lineage have stratigraphical significance for the Pliocene and the early Pleistocene.
</paragraph>
<caption id="DF646677FFA4FFB9FFB36365B8C0F90B" ID-DOI="http://doi.org/10.5281/zenodo.5377243" ID-Zenodo-Dep="5377243" httpUri="https://zenodo.org/record/5377243/files/figure.png" pageId="51" pageNumber="454" startId="51.[118,129,1646,1663]" targetBox="[165,1165,281,1640]" targetPageId="51">
<paragraph id="8BA436FFFFA4FFB9FFB36365B8C0F90B" blockId="51.[118,1216,1646,1706]" pageId="51" pageNumber="454">
FIG. 26. — 
<taxonomicName id="4C1B4D7CFFA4FFB9FF106365BBA9F9DE" authority="Mein &amp; Michaux, 1970" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[213,705,1646,1663]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="51" pageNumber="454" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA4FFB9FF106365B830F9DE" box="[213,344,1646,1663]" italics="true" pageId="51" pageNumber="454">Rhagapodemus</emphasis>
cf. 
<emphasis id="B96FEAEDFFA4FFB9FEBC6365B893F9DE" box="[377,507,1646,1663]" italics="true" pageId="51" pageNumber="454">hautimagnensis</emphasis>
<bibRefCitation id="EF8A4B0EFFA4FFB9FDC46365BBA9F9DE" author="MEIN P. &amp; MICHAUX J." box="[513,705,1646,1663]" pageId="51" pageNumber="454" pagination="2780 - 2783" refId="ref45227" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sud-occidentale. Comptes rendus de l'Academie des Sciences Paris 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux, 1970</bibRefCitation>
</taxonomicName>
, upper molars in occlusal view; 
<emphasis id="B96FEAEDFFA4FFB9FC176365BAB7F9DE" bold="true" box="[978,991,1646,1663]" pageId="51" pageNumber="454">A</emphasis>
, right M1, Ms243; 
<emphasis id="B96FEAEDFFA4FFB9FB456365BDE5F9DE" bold="true" box="[1152,1165,1646,1663]" pageId="51" pageNumber="454">B</emphasis>
, right M1, Ms244; 
<emphasis id="B96FEAEDFFA4FFB9FF1B638FB984F934" bold="true" box="[222,236,1668,1685]" pageId="51" pageNumber="454">C</emphasis>
, right M1, Ms246; 
<emphasis id="B96FEAEDFFA4FFB9FE42638FB8FDF934" bold="true" box="[391,405,1668,1685]" pageId="51" pageNumber="454">D</emphasis>
, left M2, Ms253; 
<emphasis id="B96FEAEDFFA4FFB9FDE1638FBB58F934" bold="true" box="[548,560,1668,1685]" pageId="51" pageNumber="454">E</emphasis>
, right M2, Ms254; 
<emphasis id="B96FEAEDFFA4FFB9FD0E638FBBBEF934" bold="true" box="[715,726,1668,1685]" pageId="51" pageNumber="454">F</emphasis>
, right M2, Ms255; 
<emphasis id="B96FEAEDFFA4FFB9FCB4638FBA17F934" bold="true" box="[881,895,1668,1685]" pageId="51" pageNumber="454">G</emphasis>
, right M3, Ms259; 
<emphasis id="B96FEAEDFFA4FFB9FBDF638FBD40F934" bold="true" box="[1050,1064,1668,1685]" pageId="51" pageNumber="454">H</emphasis>
, right M3, Ms260; 
<emphasis id="B96FEAEDFFA4FFB9FFB36392B914F90B" bold="true" box="[118,124,1689,1706]" pageId="51" pageNumber="454">I</emphasis>
, right M3, Ms242. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF646677FFA3FFBEFFB26365BAAEF90B" ID-DOI="http://doi.org/10.5281/zenodo.5377245" ID-Zenodo-Dep="5377245" httpUri="https://zenodo.org/record/5377245/files/figure.png" pageId="52" pageNumber="455" startId="52.[119,130,1646,1663]" targetBox="[161,1170,225,1614]" targetPageId="52">
<paragraph id="8BA436FFFFA3FFBEFFB26365BAAEF90B" blockId="52.[119,1216,1646,1706]" pageId="52" pageNumber="455">
FIG. 27. — 
<emphasis id="B96FEAEDFFA3FFBEFF126365B992F9DE" bold="true" box="[215,250,1646,1663]" pageId="52" pageNumber="455">A -H</emphasis>
, 
<taxonomicName id="4C1B4D7CFFA3FFBEFEC36365BB96F9DE" authority="Mein &amp; Michaux, 1970" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[262,766,1646,1663]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="52" pageNumber="455" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA3FFBEFEC36365B8E4F9DE" box="[262,396,1646,1663]" italics="true" pageId="52" pageNumber="455">Rhagapodemus</emphasis>
cf. 
<emphasis id="B96FEAEDFFA3FFBEFE6A6365BB5BF9DE" box="[431,563,1646,1663]" italics="true" pageId="52" pageNumber="455">hautimagnensis</emphasis>
<bibRefCitation id="EF8A4B0EFFA3FFBEFDFF6365BB96F9DE" author="MEIN P. &amp; MICHAUX J." box="[570,766,1646,1663]" pageId="52" pageNumber="455" pagination="2780 - 2783" refId="ref45227" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sud-occidentale. Comptes rendus de l'Academie des Sciences Paris 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux, 1970</bibRefCitation>
</taxonomicName>
, lower molars in occlusal view; 
<emphasis id="B96FEAEDFFA3FFBEFBD66365BD48F9DE" bold="true" box="[1043,1056,1646,1663]" pageId="52" pageNumber="455">A</emphasis>
, right m1, Ms233; 
<emphasis id="B96FEAEDFFA3FFBEFFB2638FB9ECF934" bold="true" box="[119,132,1668,1685]" pageId="52" pageNumber="455">B</emphasis>
, right m1, Ms234; 
<emphasis id="B96FEAEDFFA3FFBEFEE7638FB858F934" bold="true" box="[290,304,1668,1685]" pageId="52" pageNumber="455">C</emphasis>
, right m1, Ms229; 
<emphasis id="B96FEAEDFFA3FFBEFE0A638FB8B5F934" bold="true" box="[463,477,1668,1685]" pageId="52" pageNumber="455">D</emphasis>
, left m2, Ms237; 
<emphasis id="B96FEAEDFFA3FFBEFDB5638FBB14F934" bold="true" box="[624,636,1668,1685]" pageId="52" pageNumber="455">E</emphasis>
, right m2, Ms238; 
<emphasis id="B96FEAEDFFA3FFBEFCDE638FBA4EF934" bold="true" box="[795,806,1668,1685]" pageId="52" pageNumber="455">F</emphasis>
, right m2, Ms235; 
<emphasis id="B96FEAEDFFA3FFBEFC03638FBABCF934" bold="true" box="[966,980,1668,1685]" pageId="52" pageNumber="455">G</emphasis>
, left m3, Ms261; 
<emphasis id="B96FEAEDFFA3FFBEFBA2638FBD1DF934" bold="true" box="[1127,1141,1668,1685]" pageId="52" pageNumber="455">H</emphasis>
, left m3, Ms262; 
<emphasis id="B96FEAEDFFA3FFBEFF7F6392B9A8F90B" bold="true" box="[186,192,1689,1706]" pageId="52" pageNumber="455">I</emphasis>
, 
<emphasis id="B96FEAEDFFA3FFBEFF0F6392B9BCF90B" bold="true" box="[202,212,1689,1706]" pageId="52" pageNumber="455">J</emphasis>
, 
<taxonomicName id="4C1B4D7CFFA3FFBEFF1B6392B8ECF90B" box="[222,388,1689,1706]" class="Mammalia" family="Muridae" genus="Rhagapodemus" kingdom="Animalia" order="Rodentia" pageId="52" pageNumber="455" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B96FEAEDFFA3FFBEFF1B6392B809F90B" box="[222,353,1689,1706]" italics="true" pageId="52" pageNumber="455">Rhagapodemus</emphasis>
sp.
</taxonomicName>
, M1 in occlusal view; 
<emphasis id="B96FEAEDFFA3FFBEFDFD6392BB56F90B" bold="true" box="[568,574,1689,1706]" pageId="52" pageNumber="455">I</emphasis>
, left, Ms302; 
<emphasis id="B96FEAEDFFA3FFBEFD6A6392BBD1F90B" bold="true" box="[687,697,1689,1706]" pageId="52" pageNumber="455">J</emphasis>
, right, Ms303. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BA436FFFFA2FFBFFFB3669BB874FB48" blockId="53.[118,652,208,1706]" pageId="53" pageNumber="456">
The teeth from Muselievo are larger than these of the Greek population of 
<taxonomicName id="4C1B4D7CFFA2FFBFFE5E66BBB988FC49" authorityName="Mein &amp; Michaux" authorityYear="1970" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA2FFBFFE5E66BBB988FC49" italics="true" pageId="53" pageNumber="456">Rhagapodemus hautimagnensis</emphasis>
</taxonomicName>
. In this respect, they are similar with 
<taxonomicName id="4C1B4D7CFFA2FFBFFFB366FBB828FBAB" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[118,320,1008,1034]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA2FFBFFFB366FBB828FBAB" box="[118,320,1008,1034]" italics="true" pageId="53" pageNumber="456">Rh. hautimagnensis</emphasis>
</taxonomicName>
from its 
<typeStatus id="54A0885DFFA2FFBFFE6766E4B8B8FBA8" box="[418,464,1007,1033]" pageId="53" pageNumber="456">type</typeStatus>
locality (
<bibRefCitation id="EF8A4B0EFFA2FFBFFDF766FBB870FB8B" author="MEIN P. &amp; MICHAUX J." pageId="53" pageNumber="456" pagination="2780 - 2783" refId="ref45227" refString="MEIN P. &amp; MICHAUX J. 1970. - Un nouveau stade dans l'evolution des rongeurs pliocenes de l'Europe sud-occidentale. Comptes rendus de l'Academie des Sciences Paris 270: 2780 - 2783." type="journal article" year="1970">Mein &amp; Michaux 1970</bibRefCitation>
) as well as with 
<taxonomicName id="4C1B4D7CFFA2FFBFFE14611BBBECFB8B" authorityName="De Bruijn &amp; Van der Meulen" authorityYear="1975" box="[465,644,1040,1066]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="wandeweerdi">
<emphasis id="B96FEAEDFFA2FFBFFE14611BBBECFB8B" box="[465,644,1040,1066]" italics="true" pageId="53" pageNumber="456">Rh. wandeweerdi</emphasis>
</taxonomicName>
. On the other hand, the molars available are smaller than the molars of the latter forms (
<taxonomicName id="4C1B4D7CFFA2FFBFFDA2615BB9A3FB29" authorityName="Kretzoi" authorityYear="1959" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="frequens">
<emphasis id="B96FEAEDFFA2FFBFFDA2615BB9A3FB29" italics="true" pageId="53" pageNumber="456">Rh. frequens</emphasis>
</taxonomicName>
[
<bibRefCitation id="EF8A4B0EFFA2FFBFFF1F617BB816FB2B" author="SULIMSKI A." box="[218,382,1136,1162]" pageId="53" pageNumber="456" pagination="149 - 261" refId="ref46924" refString="SULIMSKI A. 1964. - Pliocene Lagomorpha and Rodentia from Weze I. Acta Paleontologica Polonica 9 (2): 149 - 261." type="journal article" year="1964">Sulimski 1964</bibRefCitation>
] and 
<taxonomicName id="4C1B4D7CFFA2FFBFFE7B617BB9E4FB09" authorityName="De Bruijn &amp; Van der Meulen" authorityYear="1975" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="subSpecies" species="frequens" subSpecies="athensis">
<emphasis id="B96FEAEDFFA2FFBFFE7B617BB9E4FB09" italics="true" pageId="53" pageNumber="456">Rh. frequens athensis</emphasis>
</taxonomicName>
). It appears that the 
<taxonomicName id="4C1B4D7CFFA2FFBFFEB6619BBB64FB0B" authorityName="Kretzoi" authorityYear="1959" box="[371,524,1168,1194]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFA2FFBFFEB6619BBB64FB0B" box="[371,524,1168,1194]" italics="true" pageId="53" pageNumber="456">Rhagapodemus</emphasis>
</taxonomicName>
-assemblage under consideration shows an intermediate evolutionary stage.
</paragraph>
<paragraph id="8BA436FFFFA2FFBFFFB361FBBB5EF90B" blockId="53.[118,652,208,1706]" pageId="53" pageNumber="456">
Although similar in size, the molars from Muselievo differs from 
<taxonomicName id="4C1B4D7CFFA2FFBFFEA9601BBBE3FA8B" authorityName="De Bruijn &amp; Van der Meulen" authorityYear="1975" box="[364,651,1296,1322]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="wandeweerdi">
<emphasis id="B96FEAEDFFA2FFBFFEA9601BBBE3FA8B" box="[364,651,1296,1322]" italics="true" pageId="53" pageNumber="456">Rhagapodemus wandeweerdi</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EF8A4B0EFFA2FFBFFF6D6024BB3DFAEB" author="DE BRUIJN H. &amp; VAN DER MEULEN A. J." box="[168,597,1327,1354]" pageId="53" pageNumber="456" pagination="314 - 338" refId="ref43216" refString="DE BRUIJN H. &amp; VAN DER MEULEN A. J. 1975. - The early Pleistocene Rodents from Tourkoubounia- 1 (Athens, Greece). I. Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B, 78 (4): 314 - 338." type="journal article" year="1975">
De Bruijn &amp; 
<collectingRegion id="49DFF81DFFA2FFBFFE8E6024B813FAE8" box="[331,379,1327,1353]" country="Turkey" name="Van" pageId="53" pageNumber="456">Van</collectingRegion>
der Meulen 1975
</bibRefCitation>
) by: 1) having an oval posterior labial accessory cusp (c1) on m1, which fuses quickly with the hypoconid; 2) a poor or lacking posterior accessory cusp on m2; and 3) a large distance between the 
<emphasis id="B96FEAEDFFA2FFBFFDB360A5BBE4FA69" box="[630,652,1454,1480]" italics="true" pageId="53" pageNumber="456">t1</emphasis>
and 
<emphasis id="B96FEAEDFFA2FFBFFF6160C5B9D2FA49" box="[164,186,1486,1512]" italics="true" pageId="53" pageNumber="456">t3</emphasis>
on M2. In these features the material is similar with 
<taxonomicName id="4C1B4D7CFFA2FFBFFF2760FBB8D4F9AB" authorityName="Mein &amp; Michaux" authorityYear="1970" box="[226,444,1520,1546]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="hautimagnensis">
<emphasis id="B96FEAEDFFA2FFBFFF2760FBB8D4F9AB" box="[226,444,1520,1546]" italics="true" pageId="53" pageNumber="456">Rh. hautimagnensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1B4D7CFFA2FFBFFE3C60FBBBECF9A9" authorityName="Kretzoi" authorityYear="1959" box="[505,644,1518,1546]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="species" species="frequens">
<emphasis id="B96FEAEDFFA2FFBFFE3C60FBBBECF9A9" box="[505,644,1518,1546]" italics="true" pageId="53" pageNumber="456">Rh. frequens</emphasis>
</taxonomicName>
. Therefore, if one considers these species as evolutionary stages of one phyletic lineage, the population from Muselievo provides a link between them. The material is tentatively referred to the first species on the basis of the similarity in size.
</paragraph>
<paragraph id="8BA436FFFFA2FFBFFD6F65DBBA89FDAB" blockId="53.[682,1215,207,522]" pageId="53" pageNumber="456">
Although some small molars of 
<taxonomicName id="4C1B4D7CFFA2FFBFFC3A65DBBDF0FF4B" authorityName="Kretzoi" authorityYear="1959" box="[1023,1176,208,234]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFA2FFBFFC3A65DBBDF0FF4B" box="[1023,1176,208,234]" italics="true" pageId="53" pageNumber="456">Rhagapodemus</emphasis>
</taxonomicName>
are reminiscent at first glance of that of 
<taxonomicName id="4C1B4D7CFFA2FFBFFB9565FBBDD6FEAB" authorityName="Ognev" authorityYear="1923" box="[1104,1214,240,266]" class="Mammalia" family="Muridae" genus="Sylvaemus" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFA2FFBFFB9565FBBDD6FEAB" box="[1104,1214,240,266]" italics="true" pageId="53" pageNumber="456">Sylvaemus</emphasis>
</taxonomicName>
they are readily distinguishable by their hypsodonty and occlusal pattern. In 
<taxonomicName id="4C1B4D7CFFA2FFBFFBD9643BBDDFFEEB" authorityName="Kretzoi" authorityYear="1959" box="[1052,1207,304,330]" class="Mammalia" family="Muridae" genus="Rhagapodemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFA2FFBFFBD9643BBDDFFEEB" box="[1052,1207,304,330]" italics="true" pageId="53" pageNumber="456">Rhagapodemus</emphasis>
</taxonomicName>
, the crowns of molars are high and the wear surfaces of the cusps and the ridges all lie in about the same (horizontal) plane. In contrast, in 
<taxonomicName id="4C1B4D7CFFA2FFBFFD6F64BBBA7CFE6B" authorityName="Ognev" authorityYear="1923" box="[682,788,432,458]" class="Mammalia" family="Muridae" genus="Sylvaemus" kingdom="Animalia" order="Rodentia" pageId="53" pageNumber="456" phylum="Chordata" rank="genus">
<emphasis id="B96FEAEDFFA2FFBFFD6F64BBBA7CFE6B" box="[682,788,432,458]" italics="true" pageId="53" pageNumber="456">Sylvaemus</emphasis>
</taxonomicName>
, these surfaces are oblique to the horizontal plane and the ridges between cusps are lower (
<bibRefCitation id="EF8A4B0EFFA2FFBFFD3664FBBABDFDAB" author="VAN DE WEERD A." box="[755,981,496,522]" pageId="53" pageNumber="456" pagination="127 - 168" refId="ref48078" refString="VAN DE WEERD A. 1979. - Early Ruscinian rodents and lagomorphs (Mammalia) from the lignites near Ptolemais (Macedonia, Greece). Koninklijke Nederlandse Akademie van Wetenschappen, Proceedings B, 82 (2): 127 - 168." type="journal article" year="1979">
<collectingRegion id="49DFF81DFFA2FFBFFD3664FBBA49FDAB" box="[755,801,496,522]" country="Turkey" name="Van" pageId="53" pageNumber="456">Van</collectingRegion>
de Weerd 1979
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>