269 lines
44 KiB
XML
269 lines
44 KiB
XML
<document id="8AB09A3C131A5613BB242DCA6D6E85BE" ID-CLB-Dataset="27794" ID-DOI="10.11646/zootaxa.4531.1.5" ID-GBIF-Dataset="fab96b68-c280-4ced-b0c7-6bb3c325bb50" ID-ISSN="1175-5326" ID-Zenodo-Dep="2614412" ID-ZooBank="237FD9CC-77C2-432F-B912-FBB2AC4F03FB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553786530950" checkinUser="plazi" docAuthor="Komai, Tomoyuki & Fujita, Yoshihisa" docDate="2018" docId="03B2878DDC6FFF82FF5FF8E52DEFF825" docLanguage="en" docName="zootaxa.4531.1.5.pdf" docOrigin="Zootaxa 4531 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mirograpsus Komai & Fujita 2018, n. gen." docType="treatment" docUuid="A491CA9B-98D6-4013-834A-6E258D78A822" docUuidSource="ZooBank" docVersion="9" lastPageNumber="110" masterDocId="FF8BFFF5DC6FFF83FFC8FF8B2911FFD6" masterDocTitle="A new genus and new species assigned to Macrophthalmidae (Decapoda: Brachyura: Thoracotremata) from the Ryukyu Islands, Japan" masterLastPageNumber="116" masterPageNumber="109" pageNumber="109" updateTime="1698714789343" updateUser="ExternalLinkService">
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<mods:titleInfo id="5C53F57ADA997765CD74DBEC78EFD8CB">
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<mods:title id="78EAB9853123432F65C4DA9C8FD3CC11">A new genus and new species assigned to Macrophthalmidae (Decapoda: Brachyura: Thoracotremata) from the Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="82F5ACD640330346BBA5DA1A4480C53F">Komai, Tomoyuki</mods:namePart>
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<mods:roleTerm id="A45C416399484BF27D8AD0F9A9F1E843">Author</mods:roleTerm>
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<mods:namePart id="2D3934B41ABBDABFF9A942BDD2DA9A57">Fujita, Yoshihisa</mods:namePart>
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<mods:title id="625E8D0FFEE51CAEB62060136DBB56DC">Zootaxa</mods:title>
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<mods:part id="6A6270F4F9A082047C5808E29F814B82">
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<mods:date id="98C437F13800B9EFBAC9F551A8CFB81A">2018</mods:date>
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<mods:number id="C49198B72EECA5D9749273A6297131ED">2018-12-11</mods:number>
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<mods:number id="6F1D1D99EEDE076C904AA8BDFC41648D">4531</mods:number>
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<mods:classification id="5E796DC116398EA25B47C09ECBFBFE1D">journal article</mods:classification>
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<mods:identifier id="45F11416C5D838803BBC0B2E99527AD2" type="DOI">10.11646/zootaxa.4531.1.5</mods:identifier>
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<treatment id="03B2878DDC6FFF82FF5FF8E52DEFF825" ID-DOI="http://doi.org/10.5281/zenodo.5951358" ID-GBIF-Taxon="154520751" ID-Zenodo-Dep="5951358" LSID="urn:lsid:zoobank.org:act:A491CA9B-98D6-4013-834A-6E258D78A822" httpUri="http://treatment.plazi.org/id/03B2878DDC6FFF82FF5FF8E52DEFF825" lastPageId="1" lastPageNumber="110" pageId="0" pageNumber="109">
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<subSubSection id="C3016510DC6FFF83FF5FF8E52B8FF87D" pageId="0" pageNumber="109" type="nomenclature">
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<paragraph id="8BA4369BDC6FFF83FF5FF8E5289BF85F" blockId="0.[151,670,1902,1963]" box="[151,394,1902,1929]" pageId="0" pageNumber="109">
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<heading id="D0EC81F7DC6FFF83FF5FF8E5289BF85F" bold="true" box="[151,394,1902,1929]" fontSize="11" level="1" pageId="0" pageNumber="109" reason="1">
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<emphasis id="B96FEA89DC6FFF83FF5FF8E5289BF85F" bold="true" box="[151,394,1902,1929]" pageId="0" pageNumber="109">
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<taxonomicName id="4C1B4D18DC6FFF83FF5FF8E52823F85E" ID-CoL="847XC" authority="Komai & Fujita, 2018" authorityName="Komai & Fujita" authorityYear="2018" box="[151,306,1902,1928]" class="Malacostraca" family="Macrophthalmidae" genus="Mirograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B96FEA89DC6FFF83FF5FF8E52823F85E" bold="true" box="[151,306,1902,1928]" italics="true" pageId="0" pageNumber="109">Mirograpsus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25C57F2DC6FFF83FEF0F8E4289BF85F" box="[312,394,1903,1929]" pageId="0" pageNumber="109" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA4369BDC6FFF83FF5FF8182B8FF87D" blockId="0.[151,670,1902,1963]" box="[151,670,1938,1963]" pageId="0" pageNumber="109">[New Japanese name: Kimen-doro-gani-zoku]</paragraph>
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</subSubSection>
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<subSubSection id="C3016510DC6FFF83FF5FF8502DA1F825" box="[151,1200,2010,2035]" pageId="0" pageNumber="109" type="materials_examined">
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<paragraph id="8BA4369BDC6FFF83FF5FF8502DA1F825" blockId="0.[151,1200,2010,2035]" box="[151,1200,2010,2035]" pageId="0" pageNumber="109">
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<typeStatus id="54A08839DC6FFF83FF5FF85029DEF825" box="[151,207,2011,2035]" pageId="0" pageNumber="109">Type</typeStatus>
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species.
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<taxonomicName id="4C1B4D18DC6FFF83FEFCF8502B18F824" authority="Komai & Fujita, 2018" authorityName="Komai & Fujita" authorityYear="2018" box="[308,521,2011,2034]" class="Malacostraca" family="Macrophthalmidae" genus="Mirograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="asper" status="sp. nov.">
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<emphasis id="B96FEA89DC6FFF83FEFCF8502B18F824" box="[308,521,2011,2034]" italics="true" pageId="0" pageNumber="109">Mirograpsus asper</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25C57F2DC6FFF83FDD8F8502B5DF824" box="[528,588,2011,2034]" pageId="0" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
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Present designation by monotypy. Gender: masculine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3016510DC6EFF82FF5FFF1C2AEAFCE2" pageId="1" pageNumber="110" type="diagnosis">
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<paragraph id="8BA4369BDC6EFF82FF5FFF1C2AEAFCE2" blockId="1.[151,1437,151,2035]" pageId="1" pageNumber="110">
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<emphasis id="B96FEA89DC6EFF82FF5FFF1C2800FF66" bold="true" box="[151,273,151,176]" pageId="1" pageNumber="110">Diagnosis.</emphasis>
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Carapace longer than wide; regions on dorsal surface well defined with gastric, cardiac, intestinal and branchial regions markedly elevated; cardiac region with paired rounded prominences; branchial regions each with 1 conspicuous tubercle on posterior part; external orbital tooth directed anteriorly, subacutely pointed, 2 blunt lateral teeth posterior to external orbital tooth; no lateral constrictions. Front broad, markedly bilobed, not constricted at base. Postfrontal ridges absent. Orbit large; upper orbital margin nearly transverse, markedly sinuous; lower orbital margin bordered with minute granules, inner orbital tooth small, blunt; 1 small but conspicuous subconical tubercle just below inner orbital tooth. Ocular peduncle constricted; cornea large, just reaching external orbital tooth. Antennules folded slightly obliquely; basal article large, but not particularly inflated; inter-antennular septum triangular, narrow in upper part. Antennae in orbital hiatus; flagellum greatly reduced, with long apical filamentous seta. Central region of epistome convex, supported by short median ridge in buccal cavity. Maxillipeds 3 fully closing buccal cavity; lateral margin of merus somewhat expanded, noticeably convex; carpus inserted medially on distal margin of merus; exopodal flagellum well developed. Male cheliped moderately stout; merus slightly narrowing distally, without subdistal spine on upper margin; carpus without prominent tooth or spine on inner surface; occlusal margins of fingers each with row of small teeth; tips of fingers pointed, crossing distally. Ambulatory legs moderately slender; meri each without subdistal spine on anterior margin; propodi unarmed at flexor distal angles. Male pleon gradually narrowing toward telson; somites 4–6 functionally fused though sutures demarcating somites 4/5 and 5/6 on external surface still discernible (while no suture delimited on internal surface); somite 6 with low convexity on lateral margins proximally. Male gonopod 1 relatively slender, not recurved, gradually tapering distally. Female unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3016510DC6EFF82FF0FFCB52C2BF851" pageId="1" pageNumber="110" type="discussion">
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<paragraph id="8BA4369BDC6EFF82FF0FFCB52A02F9BF" blockId="1.[151,1437,151,2035]" pageId="1" pageNumber="110">
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<emphasis id="B96FEA89DC6EFF82FF0FFCB5282AFC81" bold="true" box="[199,315,830,855]" pageId="1" pageNumber="110">Remarks.</emphasis>
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As mentioned above, the new taxon superficially resembles species of the camptrandriid genera
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<taxonomicName id="4C1B4D18DC6EFF82FF5FFCE829FFFCAC" authorityName="Tan & Ng" authorityYear="1999" box="[151,238,867,890]" class="Malacostraca" family="Camptandriidae" genus="Moguai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
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<emphasis id="B96FEA89DC6EFF82FF5FFCE829FFFCAC" box="[151,238,867,890]" italics="true" pageId="1" pageNumber="110">Moguai</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1B4D18DC6EFF82FEEBFCE828BEFCAC" authorityName="Ahyong" authorityYear="2014" box="[291,431,867,890]" class="Malacostraca" family="Camptandriidae" genus="Paramoguai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
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<emphasis id="B96FEA89DC6EFF82FEEBFCE828BEFCAC" box="[291,431,867,890]" italics="true" pageId="1" pageNumber="110">Paramoguai</emphasis>
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</taxonomicName>
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, but the nearly straight, non-recurved gonopod 1 clearly excludes it from
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<taxonomicName id="4C1B4D18DC6EFF82FB2AFCE92C88FCAD" authorityName="Stimpson" authorityYear="1858" box="[1250,1433,866,891]" class="Malacostraca" family="Camptandriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="family">Camptandriidae</taxonomicName>
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.
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<taxonomicName id="4C1B4D18DC6EFF82FF5FFC0C2865FC48" authority="Komai & Fujita, 2018" authorityName="Komai & Fujita" authorityYear="2018" box="[151,372,903,926]" class="Malacostraca" family="Macrophthalmidae" genus="Mirograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="asper" status="n. gen., n. sp.">
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<emphasis id="B96FEA89DC6EFF82FF5FFC0C2865FC48" box="[151,372,903,926]" italics="true" pageId="1" pageNumber="110">Mirograpsus asper</emphasis>
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</taxonomicName>
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<emphasis id="B96FEA89DC6EFF82FE4CFC0D2B2BFC49" bold="true" box="[388,570,902,927]" pageId="1" pageNumber="110">
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<taxonomicNameLabel id="A25C57F2DC6EFF82FE4CFC0D2B2BFC49" box="[388,570,902,927]" pageId="1" pageNumber="110" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</emphasis>
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most closely resembles
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<taxonomicName id="4C1B4D18DC6EFF82FCA7FC0C2D43FC48" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[879,1106,903,926]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
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<emphasis id="B96FEA89DC6EFF82FCA7FC0C2D43FC48" box="[879,1106,903,926]" italics="true" pageId="1" pageNumber="110">Apograpsus paantu</emphasis>
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</taxonomicName>
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, the sole representative of
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<taxonomicName id="4C1B4D18DC6EFF82FF5FFC21280CFC17" authorityName="Komai & Wada" authorityYear="2008" box="[151,285,938,961]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
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<emphasis id="B96FEA89DC6EFF82FF5FFC21280CFC17" box="[151,285,938,961]" italics="true" pageId="1" pageNumber="110">Apograpsus</emphasis>
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</taxonomicName>
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, in the general structure of the carapace, which is longer than wide with markedly elevated gastric, cardiac and intestinal regions, with the cardiac region possessing paired rounded prominences. The following major differences between the two genera, however, warrant the establishment of the new genus (cf.
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<bibRefCitation id="EF8A4B6ADC6EFF82FAF8FC7A2807FBF8" author="Komai, T. & Wada, K." pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
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for character states of
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<taxonomicName id="4C1B4D18DC6EFF82FDD5FB9D2BB2FBFB" authorityName="Komai & Wada" authorityYear="2008" box="[541,675,1046,1069]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
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<emphasis id="B96FEA89DC6EFF82FDD5FB9D2BB2FBFB" box="[541,675,1046,1069]" italics="true" pageId="1" pageNumber="110">Apograpsus</emphasis>
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</taxonomicName>
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): (1) the carapace is devoid of postfrontal ridges (
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<figureCitation id="13202A1EDC6EFF82FB29FB9E2C2AFBF8" box="[1249,1339,1045,1070]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[223,1350,237,1831]" captionTargetId="figure@3.[196,1392,193,1867]" captionTargetPageId="3" captionText="FIGURE 2. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607). A, carapace and cephalic appendages, dorsal view; B, posterior part of carapace dorsal surface, posterior view; C, frontal part of carapace, left eye, antennules, left antenna and epistome, anterior view; D, lower orbital margin and suborbital region, ventral view (setae omitted); E, left maxilliped 3, outer view; F, thoracic sternum and gonopods 1, ventral view; G, pleon, outer view; H, left gonopod 1 and penis, ventral view; I, left gonopod 1, dorsal view; J, right gonopod 2." httpUri="https://zenodo.org/record/2614418/files/figure.png" pageId="1" pageNumber="110">Fig. 2A</figureCitation>
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) (versus longitudinal postfrontal ridges are present in
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<taxonomicName id="4C1B4D18DC6EFF82FD58FBB22BECFB87" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[656,765,1081,1105]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
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<emphasis id="B96FEA89DC6EFF82FD58FBB22BECFB87" box="[656,765,1081,1105]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
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</taxonomicName>
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; cf.
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<bibRefCitation id="EF8A4B6ADC6EFF82FCF8FBB22D32FB87" author="Komai, T. & Wada, K." box="[816,1059,1081,1106]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
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: fig. 17A); (2) the lateral margins of the carapace are devoid of constriction posterior to the external orbital teeth (
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<figureCitation id="13202A1EDC6EFF82FBABFBD72DD0FBA3" box="[1123,1217,1116,1141]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[223,1350,237,1831]" captionTargetId="figure@3.[196,1392,193,1867]" captionTargetPageId="3" captionText="FIGURE 2. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607). A, carapace and cephalic appendages, dorsal view; B, posterior part of carapace dorsal surface, posterior view; C, frontal part of carapace, left eye, antennules, left antenna and epistome, anterior view; D, lower orbital margin and suborbital region, ventral view (setae omitted); E, left maxilliped 3, outer view; F, thoracic sternum and gonopods 1, ventral view; G, pleon, outer view; H, left gonopod 1 and penis, ventral view; I, left gonopod 1, dorsal view; J, right gonopod 2." httpUri="https://zenodo.org/record/2614418/files/figure.png" pageId="1" pageNumber="110">Fig. 2A</figureCitation>
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) (versus distinctly constricted posterior to the external orbital teeth in
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<taxonomicName id="4C1B4D18DC6EFF82FD1FFB0A2A55FB4E" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[727,836,1153,1176]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
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<emphasis id="B96FEA89DC6EFF82FD1FFB0A2A55FB4E" box="[727,836,1153,1176]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
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</taxonomicName>
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; cf.
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<bibRefCitation id="EF8A4B6ADC6EFF82FCBFFB0B2D7BFB4F" author="Komai, T. & Wada, K." box="[887,1130,1152,1177]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
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: fig. 17A); (3) the antennal flagellum is greatly reduced, bearing a long filamentous terminal seta (
|
||
<figureCitation id="13202A1EDC6EFF82FC66FB2F2D15FB6B" box="[942,1028,1188,1213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[223,1350,237,1831]" captionTargetId="figure@3.[196,1392,193,1867]" captionTargetPageId="3" captionText="FIGURE 2. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607). A, carapace and cephalic appendages, dorsal view; B, posterior part of carapace dorsal surface, posterior view; C, frontal part of carapace, left eye, antennules, left antenna and epistome, anterior view; D, lower orbital margin and suborbital region, ventral view (setae omitted); E, left maxilliped 3, outer view; F, thoracic sternum and gonopods 1, ventral view; G, pleon, outer view; H, left gonopod 1 and penis, ventral view; I, left gonopod 1, dorsal view; J, right gonopod 2." httpUri="https://zenodo.org/record/2614418/files/figure.png" pageId="1" pageNumber="110">Fig. 2C</figureCitation>
|
||
) (versus multi-articulated, without a prominent terminal seta in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FE0FFB432B24FB36" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[455,565,1224,1248]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FE0FFB432B24FB36" box="[455,565,1224,1248]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FDA1FB4C2A4CFB36" author="Komai, T. & Wada, K." box="[617,861,1223,1248]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 17A, B); (4) the male chela is large and stout, with the fingers pointed and crossing when closed (
|
||
<figureCitation id="13202A1EDC6EFF82FD1EFB602A5CFAD2" box="[726,845,1259,1284]" captionStart="FIGURE 4" captionStartId="5.[151,250,1951,1973]" captionTargetBox="[211,1388,277,1836]" captionTargetId="figure@5.[151,1436,240,1894]" captionTargetPageId="5" captionText="FIGURE 4. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607), left pereopods. A, cheliped, dorsal view; B, same, chela, outer view; C, pereopod 2, dorsal view; D, pereopod 3, dorsal view; E, pereopod 4, dorsal view; F, pereopod 5, dorsal view." httpUri="https://zenodo.org/record/2614424/files/figure.png" pageId="1" pageNumber="110">Fig. 4A, B</figureCitation>
|
||
) (versus small and slender, tips of fingers are spoonshaped, being similar to the female chela in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FD64FA9B2A0FFAF1" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[684,798,1296,1319]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FD64FA9B2A0FFAF1" box="[684,798,1296,1319]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FC93FA842D4DFAFE" author="Komai, T. & Wada, K." box="[859,1116,1295,1320]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 17G); (5) the meri of pereopods 2–5 are unarmed on the anterior margins (
|
||
<figureCitation id="13202A1EDC6EFF82FD2EFAB82A39FA9D" box="[742,808,1331,1356]" captionStart="FIGURE 4" captionStartId="5.[151,250,1951,1973]" captionTargetBox="[211,1388,277,1836]" captionTargetId="figure@5.[151,1436,240,1894]" captionTargetPageId="5" captionText="FIGURE 4. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607), left pereopods. A, cheliped, dorsal view; B, same, chela, outer view; C, pereopod 2, dorsal view; D, pereopod 3, dorsal view; E, pereopod 4, dorsal view; F, pereopod 5, dorsal view." httpUri="https://zenodo.org/record/2614424/files/figure.png" pageId="1" pageNumber="110">Fig. 4</figureCitation>
|
||
C–F) (versus each armed with a small subterminal spine in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FF7BFADC280EFAB9" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[179,287,1367,1391]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FF7BFADC280EFAB9" box="[179,287,1367,1391]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FE98FADD2B50FAB9" author="Komai, T. & Wada, K." box="[336,577,1366,1391]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 17H, I); and (6) the male pleomeres 4–6 are functionally fused (
|
||
<figureCitation id="13202A1EDC6EFF82FAF4FADD2C85FAB9" box="[1340,1428,1366,1391]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[223,1350,237,1831]" captionTargetId="figure@3.[196,1392,193,1867]" captionTargetPageId="3" captionText="FIGURE 2. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607). A, carapace and cephalic appendages, dorsal view; B, posterior part of carapace dorsal surface, posterior view; C, frontal part of carapace, left eye, antennules, left antenna and epistome, anterior view; D, lower orbital margin and suborbital region, ventral view (setae omitted); E, left maxilliped 3, outer view; F, thoracic sternum and gonopods 1, ventral view; G, pleon, outer view; H, left gonopod 1 and penis, ventral view; I, left gonopod 1, dorsal view; J, right gonopod 2." httpUri="https://zenodo.org/record/2614418/files/figure.png" pageId="1" pageNumber="110">Fig. 2G</figureCitation>
|
||
) (versus all freely articulated in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FE33FAF02B79FA44" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[507,616,1403,1426]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FE33FAF02B79FA44" box="[507,616,1403,1426]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FD56FAF12A85FA45" author="Komai, T. & Wada, K." box="[670,916,1402,1427]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 17D). Other notable differences between
|
||
<taxonomicName id="4C1B4D18DC6EFF82FF5FFA1429EDFA60" authorityName="Komai & Fujita" authorityYear="2018" box="[151,252,1439,1462]" class="Malacostraca" family="Macrophthalmidae" genus="Mirograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="asper">
|
||
<emphasis id="B96FEA89DC6EFF82FF5FFA1429EDFA60" box="[151,252,1439,1462]" italics="true" pageId="1" pageNumber="110">M. asper</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96FEA89DC6EFF82FECDFA1528B1FA61" bold="true" box="[261,416,1438,1463]" pageId="1" pageNumber="110">n. gen, n. sp.</emphasis>
|
||
and
|
||
<taxonomicName id="4C1B4D18DC6EFF82FE12FA142B58FA60" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[474,585,1439,1462]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FE12FA142B58FA60" box="[474,585,1439,1462]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
are: the front lacks long setae on each anterolateral part (
|
||
<figureCitation id="13202A1EDC6EFF82FB28FA152C2BFA61" box="[1248,1338,1438,1463]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[223,1350,237,1831]" captionTargetId="figure@3.[196,1392,193,1867]" captionTargetPageId="3" captionText="FIGURE 2. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607). A, carapace and cephalic appendages, dorsal view; B, posterior part of carapace dorsal surface, posterior view; C, frontal part of carapace, left eye, antennules, left antenna and epistome, anterior view; D, lower orbital margin and suborbital region, ventral view (setae omitted); E, left maxilliped 3, outer view; F, thoracic sternum and gonopods 1, ventral view; G, pleon, outer view; H, left gonopod 1 and penis, ventral view; I, left gonopod 1, dorsal view; J, right gonopod 2." httpUri="https://zenodo.org/record/2614418/files/figure.png" pageId="1" pageNumber="110">Fig. 2A</figureCitation>
|
||
) (versus with a few long setae are present at each anterolateral part of the front in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FC1CFA492D53FA0C" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[980,1090,1474,1498]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FC1CFA492D53FA0C" box="[980,1090,1474,1498]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FBBEFA492C7BFA0C" author="Komai, T. & Wada, K." box="[1142,1386,1474,1499]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 17A, B); the branchial region of the carapace has one conspicuous tubercle in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FBD3FA6C2D91FA28" authority="Komai & Fujita, 2018" authorityName="Komai & Fujita" authorityYear="2018" box="[1051,1152,1511,1534]" class="Malacostraca" family="Macrophthalmidae" genus="Mirograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="asper" status="n. gen., n. sp.">
|
||
<emphasis id="B96FEA89DC6EFF82FBD3FA6C2D91FA28" box="[1051,1152,1511,1534]" italics="true" pageId="1" pageNumber="110">M. asper</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96FEA89DC6EFF82FB41FA6D2C3BFA29" bold="true" box="[1161,1322,1510,1535]" pageId="1" pageNumber="110">
|
||
<taxonomicNameLabel id="A25C57F2DC6EFF82FB41FA6D2C3BFA29" box="[1161,1322,1510,1535]" pageId="1" pageNumber="110" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13202A1EDC6EFF82FAF4FA6D2C85FA28" box="[1340,1428,1510,1535]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[223,1350,237,1831]" captionTargetId="figure@3.[196,1392,193,1867]" captionTargetPageId="3" captionText="FIGURE 2. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607). A, carapace and cephalic appendages, dorsal view; B, posterior part of carapace dorsal surface, posterior view; C, frontal part of carapace, left eye, antennules, left antenna and epistome, anterior view; D, lower orbital margin and suborbital region, ventral view (setae omitted); E, left maxilliped 3, outer view; F, thoracic sternum and gonopods 1, ventral view; G, pleon, outer view; H, left gonopod 1 and penis, ventral view; I, left gonopod 1, dorsal view; J, right gonopod 2." httpUri="https://zenodo.org/record/2614418/files/figure.png" pageId="1" pageNumber="110">Fig. 2B</figureCitation>
|
||
) (versus tubercle absent in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FE0EF9812B27F9F7" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[454,566,1546,1569]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FE0EF9812B27F9F7" box="[454,566,1546,1569]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FDA5F9822A76F9F4" author="Komai, T. & Wada, K." box="[621,871,1545,1570]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 17A); and the male pleomere 6 bears a low convexity on the proximal part of each lateral margin (
|
||
<figureCitation id="13202A1EDC6EFF82FCDEF9A62A61F990" box="[790,880,1581,1606]" captionStart="FIGURE 2" captionStartId="3.[151,250,1882,1904]" captionTargetBox="[223,1350,237,1831]" captionTargetId="figure@3.[196,1392,193,1867]" captionTargetPageId="3" captionText="FIGURE 2. Mirograpsus asper n. gen, n. sp., holotype, male (5.8×5.1 mm; CBM-ZC 14607). A, carapace and cephalic appendages, dorsal view; B, posterior part of carapace dorsal surface, posterior view; C, frontal part of carapace, left eye, antennules, left antenna and epistome, anterior view; D, lower orbital margin and suborbital region, ventral view (setae omitted); E, left maxilliped 3, outer view; F, thoracic sternum and gonopods 1, ventral view; G, pleon, outer view; H, left gonopod 1 and penis, ventral view; I, left gonopod 1, dorsal view; J, right gonopod 2." httpUri="https://zenodo.org/record/2614418/files/figure.png" pageId="1" pageNumber="110">Fig. 2G</figureCitation>
|
||
) (versus no convexity is present on each lateral margin in
|
||
<taxonomicName id="4C1B4D18DC6EFF82FEC1F9DA2867F9BF" baseAuthorityName="Naruse & Kishino" baseAuthorityYear="2006" box="[265,374,1617,1641]" class="Malacostraca" family="Macrophthalmidae" genus="Apograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="paantu">
|
||
<emphasis id="B96FEA89DC6EFF82FEC1F9DA2867F9BF" box="[265,374,1617,1641]" italics="true" pageId="1" pageNumber="110">A. paantu</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FE60F9DA2B8AF9BF" author="Komai, T. & Wada, K." box="[424,667,1617,1642]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 17D).
|
||
</paragraph>
|
||
<paragraph id="8BA4369BDC6EFF82FF0FF9FE29D2F92E" blockId="1.[151,1437,151,2035]" pageId="1" pageNumber="110">
|
||
<taxonomicName id="4C1B4D18DC6EFF82FF0FF9FE285AF95B" authorityName="Barnard" authorityYear="1955" box="[199,331,1653,1677]" class="Malacostraca" family="Macrophthalmidae" genus="Ilyograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96FEA89DC6EFF82FF0FF9FE285AF95B" box="[199,331,1653,1677]" italics="true" pageId="1" pageNumber="110">Ilyograpsus</emphasis>
|
||
</taxonomicName>
|
||
, another representative of
|
||
<taxonomicName id="4C1B4D18DC6EFF82FD5FF9FF2A23F95B" authorityName="Stevcic" authorityYear="2005" box="[663,818,1652,1677]" class="Malacostraca" family="Macrophthalmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="subFamily" subFamily="Ilyograpsinae">Ilyograpsinae</taxonomicName>
|
||
, is readily distinguished from
|
||
<taxonomicName id="4C1B4D18DC6EFF82FB67F9FE2C2EF95A" authorityName="Komai & Fujita" authorityYear="2018" box="[1199,1343,1653,1676]" class="Malacostraca" family="Macrophthalmidae" genus="Mirograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96FEA89DC6EFF82FB67F9FE2C2EF95A" box="[1199,1343,1653,1676]" italics="true" pageId="1" pageNumber="110">Mirograpsus</emphasis>
|
||
</taxonomicName>
|
||
by the carapace being wider than long, the carapace regions being less defined, meri of the ambulatory legs each armed with a subdistal spine on the anterior margin, and the feely articulated pleomeres 4–6 (cf.
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FBB3F9372C7DF903" author="Komai, T. & Wada, K." box="[1147,1388,1724,1749]" pageId="1" pageNumber="110" pagination="357 - 384" refId="ref3775" refString="Komai, T. & Wada, K. (2008) A revision of the estuarine crab genus Ilyograpsus (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletin of Zoology, 56, 357 - 384." type="journal article" year="2008">Komai & Wada 2008</bibRefCitation>
|
||
: fig. 10).
|
||
</paragraph>
|
||
<paragraph id="8BA4369BDC6EFF82FF0FF8882C2BF851" blockId="1.[151,1437,151,2035]" pageId="1" pageNumber="110">
|
||
The family
|
||
<taxonomicName id="4C1B4D18DC6EFF82FEACF8882B29F8CA" authorityName="Dana" authorityYear="1851" box="[356,568,1795,1820]" class="Malacostraca" family="Macrophthalmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="family">Macrophthalmidae</taxonomicName>
|
||
is now represented by 10 genera classified into three subfamilies,
|
||
<taxonomicName id="4C1B4D18DC6EFF82FF5FF8AC287FF896" authorityName="Dana" authorityYear="1851" box="[151,366,1831,1856]" class="Malacostraca" family="Macrophthalmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="subFamily" subFamily="Macrophthalminae">Macrophthalminae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1B4D18DC6EFF82FEB1F8AC2B00F896" authorityName="Stevcic" authorityYear="2005" box="[377,529,1831,1856]" class="Malacostraca" family="Macrophthalmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="subFamily" subFamily="Ilyograpsinae">Ilyograpsinae</taxonomicName>
|
||
and Tritodynaminae (
|
||
<bibRefCitation id="EF8A4B6ADC6EFF82FCC3F8A32AA3F896" author="Ng, P. K. L. & Guinot, D. & Davie, P. J. F." box="[779,946,1831,1856]" pageId="1" pageNumber="110" pagination="1 - 286" refId="ref3939" refString="Ng, P. K. L., Guinot, D. & Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17 (Supplement), 1 - 286." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis id="B96FEA89DC6EFF82FCFEF8A32A79F8E9" box="[822,872,1831,1855]" italics="true" pageId="1" pageNumber="110">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
;
|
||
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Teng
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<emphasis id="B96FEA89DC6EFF82FDC4F8C72B51F8B5" box="[524,576,1867,1891]" italics="true" pageId="1" pageNumber="110">et al</emphasis>
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. 2016
|
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</bibRefCitation>
|
||
), but in all of the known genera, the male pleon consists of freely articulated somites 1–6 + telson. In this regard, consequently,
|
||
<taxonomicName id="4C1B4D18DC6EFF82FC84F8FB2ACAF851" authority="Komai & Fujita, 2018" authorityName="Komai & Fujita" authorityYear="2018" box="[844,987,1904,1927]" class="Malacostraca" family="Macrophthalmidae" genus="Mirograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B96FEA89DC6EFF82FC84F8FB2ACAF851" box="[844,987,1904,1927]" italics="true" pageId="1" pageNumber="110">Mirograpsus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96FEA89DC6EFF82FC2BF8E52D23F851" bold="true" box="[995,1074,1902,1927]" pageId="1" pageNumber="110">
|
||
<taxonomicNameLabel id="A25C57F2DC6EFF82FC2BF8E52D23F851" box="[995,1074,1902,1927]" pageId="1" pageNumber="110" rank="genus">n. gen.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unique in the family.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3016510DC6EFF82FF0FF8192DEFF825" pageId="1" pageNumber="110" type="etymology">
|
||
<paragraph id="8BA4369BDC6EFF82FF0FF8192DEFF825" blockId="1.[151,1437,151,2035]" pageId="1" pageNumber="110">
|
||
<emphasis id="B96FEA89DC6EFF82FF0FF819285FF87D" bold="true" box="[199,334,1938,1963]" pageId="1" pageNumber="110">Etymology.</emphasis>
|
||
The genus name is an arbitrary combination of the Latin “
|
||
<emphasis id="B96FEA89DC6EFF82FC14F8182D0DF87C" box="[988,1052,1939,1962]" italics="true" pageId="1" pageNumber="110">mirus</emphasis>
|
||
”, meaning amazing or marvelous, and the generic name “
|
||
<emphasis id="B96FEA89DC6EFF82FE6AF83C2B10F818" box="[418,513,1975,1998]" italics="true" pageId="1" pageNumber="110">Grapsus</emphasis>
|
||
”, which is a common suffix for the genera of the subfamily
|
||
<taxonomicName id="4C1B4D18DC6EFF82FB09F83D2C4CF819" authorityName="Stevcic" authorityYear="2005" box="[1217,1373,1974,1999]" class="Malacostraca" family="Macrophthalmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="110" phylum="Arthropoda" rank="subFamily" subFamily="Ilyograpsinae">Ilyograpsinae</taxonomicName>
|
||
. This name alludes to the unusual appearance of this crab among the macrophthalmids. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |