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<document ID-DOI="10.5252/zoosystema2023v45a3" ID-GBIF-Dataset="bc090a4d-4bce-42e0-bf6b-fb4bd45ca811" ID-ISSN="1638-9387" ID-Zenodo-Dep="7681295" ID-ZooBank="urn:lsid:zoobank.org:pub:62C2AC78-741C-4675-8C3A-67E359EB369E" checkinTime="1677503147187" checkinUser="felipe" docAuthor="Marchán, Daniel Fernández, Domínguez, Jorge, Hedde, Mickaël &amp; Decaëns, Thibaud" docDate="2023" docId="F03D0809FFC27454FEB6C4A9A1EEFD89" docLanguage="en" docName="Zoosystema.45.3.107-128.pdf" docOrigin="Zoosystema 45 (3)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.3:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="3" docTitle="Scherotheca altarocca Marchan, Decaens &amp; Dominguez 2023, n. sp." docType="treatment" docVersion="7" lastPageNumber="114" masterDocId="0C047071FFC4745DFFE4C278A344FF98" masterDocTitle="The cradle of giants: insights into the origin of Scherotheca Bouché, 1972 (Lumbricidae, Crassiclitellata) with the descriptions of eight new species from Corsica, France" masterLastPageNumber="128" masterPageNumber="107" pageNumber="111" updateTime="1677510542406" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The cradle of giants: insights into the origin of Scherotheca Bouché, 1972 (Lumbricidae, Crassiclitellata) with the descriptions of eight new species from Corsica, France</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marchán, Daniel Fernández</mods:namePart>
<mods:affiliation>CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) danifermch @ gmail. com</mods:affiliation>
<mods:nameIdentifier type="email">danifermch@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Domínguez, Jorge</mods:namePart>
<mods:affiliation>Universidade de Vigo, Grupo de Ecoloxía Animal (GEA), E- 36310 Vigo (Spain)</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hedde, Mickaël</mods:namePart>
<mods:affiliation>Eco &amp; Sols, INRAE, IRD, CIRAD, SupAgro Montpellier, Campus SupAgro, Bâtiment 12, 2 place Viala, F- 34060 Montpellier cedex 2 (France) mickael. hedde @ inrae. fr</mods:affiliation>
<mods:nameIdentifier type="email">hedde@inrae.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Decaëns, Thibaud</mods:namePart>
<mods:affiliation>CEFE, Université de Montpellier, CNRS, EPHE, IRD, 1919, route de Mende, F- 34293 Montpellier cedex 5 (France) thibaud. decaens @ cefe. cnrs. fr</mods:affiliation>
<mods:nameIdentifier type="email">decaens@cefe.cnrs.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoosystema</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>45</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>107</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/zoosystema2023v45a3</mods:identifier>
<mods:identifier type="GBIF-Dataset">bc090a4d-4bce-42e0-bf6b-fb4bd45ca811</mods:identifier>
<mods:identifier type="ISSN">1638-9387</mods:identifier>
<mods:identifier type="Zenodo-Dep">7681295</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:62C2AC78-741C-4675-8C3A-67E359EB369E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7681299" ID-Zenodo-Dep="7681299" LSID="urn:lsid:plazi:treatment:F03D0809FFC27454FEB6C4A9A1EEFD89" httpUri="http://treatment.plazi.org/id/F03D0809FFC27454FEB6C4A9A1EEFD89" lastPageId="9" lastPageNumber="114" pageId="6" pageNumber="111">
<subSubSection pageId="6" pageNumber="111" type="nomenclature">
<paragraph blockId="6.[234,673,1745,1836]" pageId="6" pageNumber="111">
<heading centered="true" fontSize="11" level="2" pageId="6" pageNumber="111" reason="3">
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="111" phylum="Annelida" rank="species" species="altarocca" status="n. sp.">
<emphasis bold="true" box="[338,569,1745,1772]" italics="true" pageId="6" pageNumber="111">Scherotheca altarocca</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[614,673,1778,1804]" pageId="6" pageNumber="111" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="111" type="description">
<paragraph blockId="6.[234,673,1745,1836]" box="[405,502,1810,1836]" pageId="6" pageNumber="111">
(
<figureCitation box="[413,494,1810,1836]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens &amp; Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens &amp; Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens &amp; Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens &amp; Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens &amp; Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens &amp; Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens &amp; Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens &amp; Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="6" pageNumber="111">Fig. 3A</figureCitation>
)
</paragraph>
<paragraph blockId="6.[162,745,1870,1888]" box="[162,745,1870,1888]" pageId="6" pageNumber="111">
<uri box="[162,745,1870,1888]" pageId="6" pageNumber="111">
urn:lsid:zoobank.org:act:
<uuid box="[361,745,1870,1888]" pageId="6" pageNumber="111">89EFE242-73AF-435E-86C8-67B17B93D253</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="111" type="materials_examined">
<paragraph blockId="6.[131,776,1921,2024]" pageId="6" pageNumber="111">
<materialsCitation collectingDate="2021-04-03" collectionCode="MNHN" collectorName="T. Decaens &amp; D. Fernandez Marchan" country="France" county="Adult" elevation="1422" latitude="41.872776" location="Castellu d'Ornucciu" longLatPrecision="19" longitude="9.154722" municipality="Zicavo" pageId="6" pageNumber="111" specimenCode="DFM-0446" specimenCount="1" stateProvince="Corse du Sud" typeStatus="holotype">
TYPE MATERIAL. —
<emphasis bold="true" box="[326,421,1921,1945]" pageId="6" pageNumber="111">
<typeStatus box="[326,421,1921,1945]" pageId="6" pageNumber="111">Holotype</typeStatus>
</emphasis>
.
<emphasis bold="true" box="[433,508,1922,1946]" pageId="6" pageNumber="111">
<collectingCountry box="[433,508,1922,1946]" name="France" pageId="6" pageNumber="111">Corsica</collectingCountry>
</emphasis>
•
<specimenCount box="[531,589,1921,1945]" pageId="6" pageNumber="111" type="adult">Adult</specimenCount>
;
<collectingRegion box="[599,734,1921,1945]" pageId="6" pageNumber="111">Corse du Sud</collectingRegion>
,
<collectingMunicipality pageId="6" pageNumber="111">Zicavo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC27454FEB6C4A9A1EEFD89:7D4BEFC4FFC2745BFF59C5E3A2CCF82B" box="[189,392,1947,1971]" country="France" county="Adult" latitude="41.872776" longLatPrecision="19" longitude="9.154722" municipality="Zicavo" name="Castellu d'Ornucciu" pageId="6" pageNumber="111" stateProvince="Corse du Sud">Castellu d’Ornucciu</location>
;
<geoCoordinate box="[403,525,1947,1971]" degrees="41" direction="north" minutes="52" orientation="latitude" pageId="6" pageNumber="111" precision="15" seconds="22" value="41.872776">41°52’22”N</geoCoordinate>
,
<geoCoordinate box="[536,625,1948,1972]" degrees="9" direction="east" minutes="9" orientation="longitude" pageId="6" pageNumber="111" precision="15" seconds="17" value="9.154722">9°9’17”E</geoCoordinate>
;
<elevation box="[639,770,1947,1972]" metricMagnitude="3" metricUnit="m" metricValue="1.422" pageId="6" pageNumber="111" unit="m" value="1422.0">
<quantity box="[639,717,1948,1972]" metricMagnitude="3" metricUnit="m" metricValue="1.422" pageId="6" pageNumber="111" unit="m" value="1422.0">1422 m</quantity>
a.s.l.
</elevation>
;
<date box="[133,234,1974,1998]" pageId="6" pageNumber="111" value="2021-04-03">
<collectingDate box="[133,234,1974,1998]" pageId="6" pageNumber="111" value="2021-04-03">3.IV.2021</collectingDate>
</date>
;
<collectorName box="[242,350,1974,1997]" pageId="6" pageNumber="111">T. Decaëns</collectorName>
,
<collectorName box="[358,582,1974,1998]" pageId="6" pageNumber="111">D. Fernández Marchán</collectorName>
leg.; BOLD Sample ID:
<specimenCode box="[172,289,2000,2024]" pageId="6" pageNumber="111">DFM-0446</specimenCode>
;
<collectionCode box="[300,386,2001,2024]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="111">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph blockId="6.[811,1457,1369,1895]" pageId="6" pageNumber="111">
<materialsCitation collectingDate="2021-04-06" collectionCode="T, BOLD, MNHN" country="France" elevation="1051" latitude="41.845833" location="Olivese" longLatPrecision="19" longitude="9.081944" municipality="Sud" pageId="6" pageNumber="111" specimenCount="3" specimenCount-adult="3" stateProvince="Corse" typeStatus="paratype">
<emphasis bold="true" box="[813,1004,1369,1394]" pageId="6" pageNumber="111">
<typeStatus box="[813,915,1369,1393]" pageId="6" pageNumber="111">Paratypes</typeStatus>
.
<collectingCountry box="[926,1004,1370,1394]" name="France" pageId="6" pageNumber="111">Corsica</collectingCountry>
</emphasis>
•
<specimenCount box="[1030,1102,1369,1393]" pageId="6" pageNumber="111" type="adult">3 adult</specimenCount>
specimens;
<collectingRegion box="[1227,1286,1369,1393]" country="France" name="Corse" pageId="6" pageNumber="111">Corse</collectingRegion>
du
<collectingMunicipality box="[1327,1369,1369,1393]" pageId="6" pageNumber="111">Sud</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC27454FEB6C4A9A1EEFD89:7D4BEFC4FFC2745BFA81C721A6EBFAE9" box="[1381,1455,1369,1393]" country="France" latitude="41.845833" longLatPrecision="19" longitude="9.081944" municipality="Sud" name="Olivese" pageId="6" pageNumber="111" stateProvince="Corse">Olivese</location>
(D26 road);
<geoCoordinate box="[947,1074,1395,1419]" degrees="41" direction="north" minutes="50" orientation="latitude" pageId="6" pageNumber="111" precision="15" seconds="45" value="41.845833">41°50’45”N</geoCoordinate>
,
<geoCoordinate box="[1087,1179,1395,1419]" degrees="9" direction="east" minutes="4" orientation="longitude" pageId="6" pageNumber="111" precision="15" seconds="55" value="9.081944">9°4’55”E</geoCoordinate>
;
<elevation box="[1195,1333,1395,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.051" pageId="6" pageNumber="111" unit="m" value="1051.0">
<quantity box="[1195,1276,1395,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.051" pageId="6" pageNumber="111" unit="m" value="1051.0">1051 m</quantity>
a.s.l.
</elevation>
;
<date box="[1347,1452,1395,1419]" pageId="6" pageNumber="111" value="2021-04-06">
<collectingDate box="[1347,1452,1395,1419]" pageId="6" pageNumber="111" value="2021-04-06">6.IV.2021</collectingDate>
</date>
;
<collectionCode box="[814,832,1422,1445]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="111">T</collectionCode>
. Decaëns, D. Fernández Marchán leg.;
<collectionCode box="[1254,1327,1422,1446]" pageId="6" pageNumber="111">BOLD</collectionCode>
Sample ID: DFM-0512, DFM-0513, DFM-0514;
<collectionCode box="[1212,1298,1449,1472]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="111">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-06" collectionCode="T, BOLD" country="France" elevation="1178" latitude="41.836945" location="Olivese" longLatPrecision="19" longitude="9.095833" municipality="Sud" pageId="6" pageNumber="111" specimenCount="3" specimenCount-adult="3" stateProvince="Corse" typeStatus="paratype">
<specimenCount box="[1321,1392,1448,1472]" pageId="6" pageNumber="111" type="adult">3 adult</specimenCount>
specimens;
<collectingRegion box="[877,935,1475,1499]" country="France" name="Corse" pageId="6" pageNumber="111">Corse</collectingRegion>
du
<collectingMunicipality box="[971,1012,1475,1499]" pageId="6" pageNumber="111">Sud</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC27454FEB6C4A9A1EEFD89:7D4BEFC4FFC2745BFC19C7BBA702FA43" box="[1021,1094,1475,1499]" country="France" latitude="41.836945" longLatPrecision="19" longitude="9.095833" municipality="Sud" name="Olivese" pageId="6" pageNumber="111" stateProvince="Corse">Olivese</location>
(D69 road);
<geoCoordinate box="[1223,1348,1474,1498]" degrees="41" direction="north" minutes="50" orientation="latitude" pageId="6" pageNumber="111" precision="15" seconds="13" value="41.836945">41°50’13”N</geoCoordinate>
,
<geoCoordinate box="[1357,1448,1474,1498]" degrees="9" direction="east" minutes="5" orientation="longitude" pageId="6" pageNumber="111" precision="15" seconds="45" value="9.095833">9°5’45”E</geoCoordinate>
;
<elevation box="[813,951,1501,1525]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="6" pageNumber="111" unit="m" value="1178.0">
<quantity box="[813,894,1501,1525]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="6" pageNumber="111" unit="m" value="1178.0">1178 m</quantity>
a.s.l.
</elevation>
;
<date box="[965,1071,1501,1525]" pageId="6" pageNumber="111" value="2021-04-06">
<collectingDate box="[965,1071,1501,1525]" pageId="6" pageNumber="111" value="2021-04-06">6.IV.2021</collectingDate>
</date>
;
<collectionCode box="[1083,1100,1502,1525]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="111">T</collectionCode>
. Decaëns, D. Fernández Marchán leg.; Eco&amp;Sols;
<collectionCode box="[983,1055,1528,1552]" pageId="6" pageNumber="111">BOLD</collectionCode>
Sample ID: DFM-0266, DFM-0267, DFM-0268
</materialsCitation>
) •
<materialsCitation collectingDate="2021-04-06" collectionCode="BOLD, MNHN" country="France" elevation="1178" latitude="41.836945" location="Olivese" longLatPrecision="19" longitude="9.095833" municipality="Sud" pageId="6" pageNumber="111" specimenCount="14" specimenCount-adult="2" specimenCount-juvenile="12" stateProvince="Corse" typeStatus="paratype">
<specimenCount box="[976,1053,1554,1578]" pageId="6" pageNumber="111" type="adult">2 adult</specimenCount>
specimens,
<specimenCount box="[1187,1310,1554,1578]" pageId="6" pageNumber="111" type="juvenile">12 juvenile</specimenCount>
; same data as for preceding;
<collectionCode box="[971,1044,1581,1605]" pageId="6" pageNumber="111">BOLD</collectionCode>
Sample ID: DFM-0269, DFM-0270, DFM-0271, DFM-0272, DFM-0274, DFM-0275, DFM-0276, DFM-0277, DFM-0278, DFM-0279, DFM-0280, DFM-0281, DFM-0282, DFM-0283;
<collectionCode box="[1081,1167,1660,1683]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="111">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-03" collectionCode="T, BOLD, MNHN" country="France" county="Sud" elevation="1422" latitude="41.872776" location="Castellu d'Ornucciu" longLatPrecision="19" longitude="9.154722" municipality="Zicavo" pageId="6" pageNumber="111" specimenCount="6" specimenCount-adult="6" stateProvince="Corse" typeStatus="paratype">
<specimenCount box="[1194,1267,1660,1684]" pageId="6" pageNumber="111" type="adult">6 adult</specimenCount>
specimens;
<collectingRegion box="[1394,1452,1660,1684]" country="France" name="Corse" pageId="6" pageNumber="111">Corse</collectingRegion>
du
<collectingCounty box="[850,892,1686,1710]" pageId="6" pageNumber="111">Sud</collectingCounty>
,
<collectingMunicipality box="[905,977,1686,1710]" pageId="6" pageNumber="111">Zicavo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC27454FEB6C4A9A1EEFD89:7D4BEFC4FFC2745BFC39C4EEA7F7F936" box="[989,1203,1686,1710]" country="France" county="Sud" latitude="41.872776" longLatPrecision="19" longitude="9.154722" municipality="Zicavo" name="Castellu d'Ornucciu" pageId="6" pageNumber="111" stateProvince="Corse">Castellu d’Ornucciu</location>
;
<geoCoordinate box="[1216,1344,1686,1710]" degrees="41" direction="north" minutes="52" orientation="latitude" pageId="6" pageNumber="111" precision="15" seconds="22" value="41.872776">41°52’22”N</geoCoordinate>
,
<geoCoordinate box="[1357,1451,1686,1710]" degrees="9" direction="east" minutes="9" orientation="longitude" pageId="6" pageNumber="111" precision="15" seconds="17" value="9.154722">9°9’17”E</geoCoordinate>
;
<elevation box="[813,951,1713,1737]" metricMagnitude="3" metricUnit="m" metricValue="1.422" pageId="6" pageNumber="111" unit="m" value="1422.0">
<quantity box="[813,894,1713,1737]" metricMagnitude="3" metricUnit="m" metricValue="1.422" pageId="6" pageNumber="111" unit="m" value="1422.0">1422 m</quantity>
a.s.l.
</elevation>
;
<date box="[965,1071,1713,1737]" pageId="6" pageNumber="111" value="2021-04-03">
<collectingDate box="[965,1071,1713,1737]" pageId="6" pageNumber="111" value="2021-04-03">3.IV.2021</collectingDate>
</date>
;
<collectionCode box="[1083,1100,1713,1736]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="111">T</collectionCode>
. Decaëns, D. Fernández Marchán leg.;
<collectionCode box="[863,934,1739,1763]" pageId="6" pageNumber="111">BOLD</collectionCode>
Sample ID: DFM-0447, DFM-0448, DFM-0449, DFM-0450, DFM-0451, DFM-0452;
<collectionCode box="[1213,1299,1766,1789]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="111">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation collectingDate="2021-04-03" collectionCode="T, BOLD, MNHN" country="France" county="Sud" elevation="1248" latitude="41.87583" location="Ponte di Valpine" longLatPrecision="19" longitude="9.132778" municipality="Zicavo" pageId="6" pageNumber="111" specimenCount="3" specimenCount-adult="3" stateProvince="Corse" typeStatus="paratype">
<specimenCount box="[1324,1396,1766,1790]" pageId="6" pageNumber="111" type="adult">3 adult</specimenCount>
specimens;
<collectingRegion box="[889,948,1792,1816]" country="France" name="Corse" pageId="6" pageNumber="111">Corse</collectingRegion>
du
<collectingCounty box="[993,1035,1792,1816]" pageId="6" pageNumber="111">Sud</collectingCounty>
,
<collectingMunicipality box="[1048,1120,1792,1816]" pageId="6" pageNumber="111">Zicavo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:F03D0809FFC27454FEB6C4A9A1EEFD89:7D4BEFC4FFC2745BFB89C579A65AF880" box="[1133,1310,1792,1816]" country="France" county="Sud" latitude="41.87583" longLatPrecision="19" longitude="9.132778" municipality="Zicavo" name="Ponte di Valpine" pageId="6" pageNumber="111" stateProvince="Corse">Ponte di Valpine</location>
;
<geoCoordinate box="[1323,1450,1792,1816]" degrees="41" direction="north" minutes="52" orientation="latitude" pageId="6" pageNumber="111" precision="15" seconds="33" value="41.87583">41°52’33”N</geoCoordinate>
,
<geoCoordinate box="[813,905,1818,1842]" degrees="9" direction="east" minutes="7" orientation="longitude" pageId="6" pageNumber="111" precision="15" seconds="58" value="9.132778">9°7’58”E</geoCoordinate>
;
<elevation box="[919,1052,1818,1843]" metricMagnitude="3" metricUnit="m" metricValue="1.248" pageId="6" pageNumber="111" unit="m" value="1248.0">
<quantity box="[919,998,1819,1843]" metricMagnitude="3" metricUnit="m" metricValue="1.248" pageId="6" pageNumber="111" unit="m" value="1248.0">1248 m</quantity>
a.s.l.
</elevation>
;
<date box="[1064,1182,1819,1843]" pageId="6" pageNumber="111" value="2021-04-03">
<collectingDate box="[1064,1182,1819,1843]" pageId="6" pageNumber="111" value="2021-04-03">03.IV.2021</collectingDate>
</date>
;
<collectionCode box="[1192,1209,1819,1842]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="111">T</collectionCode>
. Decaëns, D. Fernández Marchán leg.;
<collectionCode box="[968,1041,1845,1869]" pageId="6" pageNumber="111">BOLD</collectionCode>
Sample ID:, DFM-0443, DFM-0444, DFM-0445;
<collectionCode box="[948,1037,1872,1895]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="111">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="111" type="etymology">
<paragraph blockId="6.[811,1455,1949,2025]" pageId="6" pageNumber="111">
ETYMOLOGY. — The species name is derived from the natural region of Altarocca where the
<typeStatus box="[1039,1081,1976,2000]" pageId="6" pageNumber="111">type</typeStatus>
specimens have been collected (name in apposition).
</paragraph>
</subSubSection>
<caption ID-Table-UUID="2CEBE997FFC3745AFF60C2A3A7D6FE82" httpUri="http://table.plazi.org/id/2CEBE997FFC3745AFF60C2A3A7D6FE82" pageId="7" pageNumber="112" startId="7.[132,143,219,237]" targetBox="[134,1453,340,1072]" targetIsTable="true" targetPageId="7">
<paragraph blockId="7.[132,1457,219,282]" pageId="7" pageNumber="112">
TABLE 2. — Reproduction-related morphological characters of the newly described species of Corsican
<taxonomicName authority="Bouche, 1972" authorityName="Bouche" authorityYear="1972" box="[1002,1230,219,237]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="genus">
<emphasis box="[1002,1107,219,237]" italics="true" pageId="7" pageNumber="112">Scherotheca</emphasis>
<bibRefCitation author="BOUCHE M. B." box="[1113,1230,219,237]" pageId="7" pageNumber="112" refId="ref15653" refString="BOUCHE M. B. 1972. - Lombriciens de France: ecologie et systematique (Vol. 72, No. 2). Paris: Institut National de la Recherche Agronomique." type="book" year="1972">Bouché, 1972</bibRefCitation>
</taxonomicName>
and their closest relatives. Abbreviations:
<emphasis bold="true" box="[257,376,241,260]" pageId="7" pageNumber="112">Sperm. pores</emphasis>
, position of spermathecal pores;
<emphasis bold="true" box="[657,694,241,260]" pageId="7" pageNumber="112">S.V.,</emphasis>
position of seminal vesicles;
<emphasis bold="true" box="[939,969,242,260]" pageId="7" pageNumber="112">T.P.</emphasis>
, position of
<emphasis box="[1072,1238,242,260]" italics="true" pageId="7" pageNumber="112">tubercula pubertatis</emphasis>
. Symbols:
<emphasis bold="true" box="[1332,1340,242,260]" pageId="7" pageNumber="112">*</emphasis>
, according to
<bibRefCitation author="POP V. V." box="[132,225,264,282]" pageId="7" pageNumber="112" pagination="18" refId="ref17161" refString="POP V. V. 1947. - Lombriciens de la Corse. Archives de Zoologie experimentale et generale 85: 18." type="journal article" year="1947">Pop (1947)</bibRefCitation>
: composite between the different populations included in the original description;
<emphasis bold="true" box="[907,922,264,282]" pageId="7" pageNumber="112">**</emphasis>
, according to
<bibRefCitation author="BOUCHE M. B." box="[1043,1167,264,282]" pageId="7" pageNumber="112" refId="ref15653" refString="BOUCHE M. B. 1972. - Lombriciens de France: ecologie et systematique (Vol. 72, No. 2). Paris: Institut National de la Recherche Agronomique." type="book" year="1972">Bouché (1972)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph pageId="7" pageNumber="112">
<table box="[134,1453,340,1072]" gridcols="6" gridrows="30" pageId="7" pageNumber="112">
<tr box="[134,1453,340,360]" gridrow="0" pageId="7" pageNumber="112">
<th box="[134,449,340,360]" gridcol="0" gridrow="0" pageId="7" pageNumber="112">
<emphasis bold="true" box="[134,215,340,360]" pageId="7" pageNumber="112">Species</emphasis>
</th>
<th box="[464,720,340,360]" gridcol="1" gridrow="0" pageId="7" pageNumber="112">
<emphasis bold="true" box="[465,601,340,360]" pageId="7" pageNumber="112">Sperm. pores</emphasis>
</th>
<th box="[738,909,340,360]" gridcol="2" gridrow="0" pageId="7" pageNumber="112">
<emphasis bold="true" box="[738,822,340,360]" pageId="7" pageNumber="112">Clitelum</emphasis>
</th>
<th box="[920,1119,340,360]" gridcol="3" gridrow="0" pageId="7" pageNumber="112">
<emphasis bold="true" box="[920,953,340,360]" pageId="7" pageNumber="112">T.P.</emphasis>
</th>
<th box="[1129,1231,340,360]" gridcol="4" gridrow="0" pageId="7" pageNumber="112">
<emphasis bold="true" box="[1130,1166,340,360]" pageId="7" pageNumber="112">S.V.</emphasis>
</th>
<th box="[1251,1453,340,360]" gridcol="5" gridrow="0" pageId="7" pageNumber="112">
<emphasis bold="true" box="[1252,1400,340,360]" pageId="7" pageNumber="112">Spermathecae</emphasis>
</th>
</tr>
<tr box="[134,1453,375,395]" gridrow="1" pageId="7" pageNumber="112">
<th box="[134,449,375,395]" gridcol="0" gridrow="1" pageId="7" pageNumber="112">
<taxonomicName authority="Bouche, 1972" authorityName="Bouche" authorityYear="1972" box="[134,436,375,395]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="albomaculata">
<emphasis box="[134,298,375,395]" italics="true" pageId="7" pageNumber="112">Sc. albomaculata</emphasis>
<bibRefCitation author="BOUCHE M. B." box="[304,436,375,395]" pageId="7" pageNumber="112" refId="ref15653" refString="BOUCHE M. B. 1972. - Lombriciens de France: ecologie et systematique (Vol. 72, No. 2). Paris: Institut National de la Recherche Agronomique." type="book" year="1972">Bouché, 1972</bibRefCitation>
</taxonomicName>
</th>
<td box="[464,720,375,395]" gridcol="1" gridrow="1" pageId="7" pageNumber="112">11/12-13/14 multiple</td>
<td box="[738,909,375,395]" gridcol="2" gridrow="1" pageId="7" pageNumber="112">(26)27-37(38)</td>
<td box="[920,1119,375,395]" gridcol="3" gridrow="1" pageId="7" pageNumber="112">30-38</td>
<td box="[1129,1231,375,395]" gridcol="4" gridrow="1" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,375,395]" gridcol="5" gridrow="1" pageId="7" pageNumber="112">Multiple 11,12,13</td>
</tr>
<tr box="[134,1453,399,420]" gridrow="2" pageId="7" pageNumber="112">
<th box="[134,449,399,420]" gridcol="0" gridrow="2" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,350,399,420]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="altarocca">
<emphasis box="[134,258,399,420]" italics="true" pageId="7" pageNumber="112">Sc. altarocca</emphasis>
Marchán
</taxonomicName>
,
</th>
<td box="[464,720,399,420]" gridcol="1" gridrow="2" pageId="7" pageNumber="112">(11/12)12/13, 13/14 simple</td>
<td box="[738,909,399,420]" gridcol="2" gridrow="2" pageId="7" pageNumber="112">(29)30-36(37)</td>
<td box="[920,1119,399,420]" gridcol="3" gridrow="2" pageId="7" pageNumber="112">(½ 32)33-36(37)</td>
<td box="[1129,1231,399,420]" gridcol="4" gridrow="2" pageId="7" pageNumber="112">11,12</td>
<td box="[1251,1453,399,420]" gridcol="5" gridrow="2" pageId="7" pageNumber="112">Simple (11),12,13</td>
</tr>
<tr box="[134,1453,423,444]" gridrow="3" pageId="7" pageNumber="112" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th box="[134,449,423,444]" gridcol="0" gridrow="3" pageId="7" pageNumber="112">
Decäens &amp; Domínguez,
<taxonomicNameLabel box="[377,431,424,444]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,448,468]" gridrow="4" pageId="7" pageNumber="112">
<th box="[134,449,448,468]" gridcol="0" gridrow="4" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,392,448,468]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="boccaverghju">
<emphasis box="[134,299,448,468]" italics="true" pageId="7" pageNumber="112">Sc. boccaverghju</emphasis>
Marchán
</taxonomicName>
,
</th>
<td box="[464,720,448,468]" gridcol="1" gridrow="4" pageId="7" pageNumber="112">12/13, 13/14 multiple</td>
<td box="[738,909,448,468]" gridcol="2" gridrow="4" pageId="7" pageNumber="112">27-36</td>
<td box="[920,1119,448,468]" gridcol="3" gridrow="4" pageId="7" pageNumber="112">31-34(½ 35)</td>
<td box="[1129,1231,448,468]" gridcol="4" gridrow="4" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,448,468]" gridcol="5" gridrow="4" pageId="7" pageNumber="112">Multiple 12,13</td>
</tr>
<tr box="[134,1453,472,492]" gridrow="5" pageId="7" pageNumber="112">
<th box="[134,1453,472,492]" colspan="6" colspanRight="5" gridcol="0" gridrow="5" pageId="7" pageNumber="112">
Decäens &amp; Domínguez,
<taxonomicNameLabel box="[377,431,472,492]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,496,516]" gridrow="6" pageId="7" pageNumber="112">
<th box="[134,449,496,516]" gridcol="0" gridrow="6" pageId="7" pageNumber="112">
<taxonomicName authority="Bouche, 1972" authorityName="Bouche" authorityYear="1972" box="[134,397,496,516]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="brevisella">
<emphasis box="[134,259,496,516]" italics="true" pageId="7" pageNumber="112">Sc. brevisella</emphasis>
<bibRefCitation author="BOUCHE M. B." box="[265,397,496,516]" pageId="7" pageNumber="112" refId="ref15653" refString="BOUCHE M. B. 1972. - Lombriciens de France: ecologie et systematique (Vol. 72, No. 2). Paris: Institut National de la Recherche Agronomique." type="book" year="1972">Bouché, 1972</bibRefCitation>
</taxonomicName>
</th>
<td box="[464,720,496,516]" gridcol="1" gridrow="6" pageId="7" pageNumber="112">9/10-13/14 simple</td>
<td box="[738,909,496,516]" gridcol="2" gridrow="6" pageId="7" pageNumber="112">½ 27-37</td>
<td box="[920,1119,496,516]" gridcol="3" gridrow="6" pageId="7" pageNumber="112">30-37</td>
<td box="[1129,1231,496,516]" gridcol="4" gridrow="6" pageId="7" pageNumber="112">(10)11,12</td>
<td box="[1251,1453,496,516]" gridcol="5" gridrow="6" pageId="7" pageNumber="112">Simple 9,10,11,12,13</td>
</tr>
<tr box="[134,1453,520,540]" gridrow="7" pageId="7" pageNumber="112">
<th box="[134,449,520,540]" gridcol="0" gridrow="7" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,374,520,540]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="capcorsana">
<emphasis box="[134,281,520,540]" italics="true" pageId="7" pageNumber="112">Sc. capcorsana</emphasis>
Marchán
</taxonomicName>
,
</th>
<td box="[464,720,520,540]" gridcol="1" gridrow="7" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,520,540]" gridcol="2" gridrow="7" pageId="7" pageNumber="112">(27)28-35(36)</td>
<td box="[920,1119,520,540]" gridcol="3" gridrow="7" pageId="7" pageNumber="112">½ 30-34</td>
<td box="[1129,1231,520,540]" gridcol="4" gridrow="7" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,520,540]" gridcol="5" gridrow="7" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,544,564]" gridrow="8" pageId="7" pageNumber="112">
<th box="[134,1453,544,564]" colspan="6" colspanRight="5" gridcol="0" gridrow="8" pageId="7" pageNumber="112">
Decäens &amp; Domínguez,
<taxonomicNameLabel box="[377,431,544,564]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,568,588]" gridrow="9" pageId="7" pageNumber="112">
<th box="[134,449,568,588]" gridcol="0" gridrow="9" pageId="7" pageNumber="112">
<taxonomicName authority="* (Pop, 1947)" authorityName="* (Pop" authorityYear="1947" baseAuthorityName="Pop" baseAuthorityYear="1947" box="[134,385,568,588]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="corsicana">
<emphasis box="[134,270,568,588]" italics="true" pageId="7" pageNumber="112">Sc. corsicana*</emphasis>
(
<bibRefCitation author="POP V. V." box="[281,380,568,588]" pageId="7" pageNumber="112" pagination="18" refId="ref17161" refString="POP V. V. 1947. - Lombriciens de la Corse. Archives de Zoologie experimentale et generale 85: 18." type="journal article" year="1947">Pop, 1947</bibRefCitation>
)
</taxonomicName>
</th>
<td box="[464,720,568,588]" gridcol="1" gridrow="9" pageId="7" pageNumber="112">12/13,13/14 simple/</td>
<td box="[738,909,568,588]" gridcol="2" gridrow="9" pageId="7" pageNumber="112">27-35,36</td>
<td box="[920,1119,568,588]" gridcol="3" gridrow="9" pageId="7" pageNumber="112">31-34</td>
<td box="[1129,1231,568,588]" gridcol="4" gridrow="9" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,568,588]" gridcol="5" gridrow="9" pageId="7" pageNumber="112">Simple/double/</td>
</tr>
<tr box="[134,1453,592,612]" gridrow="10" pageId="7" pageNumber="112" rowspan-0="1" rowspan-4="1">
<td box="[464,720,592,612]" gridcol="1" gridrow="10" pageId="7" pageNumber="112">double/triple</td>
<td box="[738,909,592,612]" gridcol="2" gridrow="10" pageId="7" pageNumber="112">26-34</td>
<td box="[920,1119,592,612]" gridcol="3" gridrow="10" pageId="7" pageNumber="112">30,31-34,35 29-32</td>
<td box="[1251,1453,592,612]" gridcol="5" gridrow="10" pageId="7" pageNumber="112">triple 12,13</td>
</tr>
<tr box="[134,1453,618,638]" gridrow="11" pageId="7" pageNumber="112">
<th box="[134,449,618,638]" gridcol="0" gridrow="11" pageId="7" pageNumber="112">
<emphasis box="[134,277,618,638]" italics="true" pageId="7" pageNumber="112">
<taxonomicName authorityName="* (Pop" authorityYear="1947" box="[134,262,618,638]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="corsicana">Sc. corsicana</taxonomicName>
**
</emphasis>
</th>
<td box="[464,720,618,638]" gridcol="1" gridrow="11" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,618,638]" gridcol="2" gridrow="11" pageId="7" pageNumber="112">(1/2 26)1/2 27-1/2</td>
<td box="[920,1119,618,638]" gridcol="3" gridrow="11" pageId="7" pageNumber="112">(1/2 30)31-1/2 34(34)</td>
<td box="[1129,1231,618,638]" gridcol="4" gridrow="11" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,618,638]" gridcol="5" gridrow="11" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,642,662]" gridrow="12" pageId="7" pageNumber="112">
<th box="[134,1453,642,662]" colspan="6" colspanRight="5" gridcol="0" gridrow="12" pageId="7" pageNumber="112">36(36)</th>
</tr>
<tr box="[134,1453,666,686]" gridrow="13" pageId="7" pageNumber="112">
<th box="[134,449,666,686]" gridcol="0" gridrow="13" pageId="7" pageNumber="112">
<taxonomicName authority="Bouche, 1972" authorityName="Bouche" authorityYear="1972" box="[134,449,666,686]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="subSpecies" species="corsicana" subSpecies="popi">
<emphasis box="[134,310,666,686]" italics="true" pageId="7" pageNumber="112">Sc. corsicana popi</emphasis>
<bibRefCitation author="BOUCHE M. B." box="[316,449,666,686]" pageId="7" pageNumber="112" refId="ref15653" refString="BOUCHE M. B. 1972. - Lombriciens de France: ecologie et systematique (Vol. 72, No. 2). Paris: Institut National de la Recherche Agronomique." type="book" year="1972">Bouché, 1972</bibRefCitation>
</taxonomicName>
</th>
<td box="[464,720,666,686]" gridcol="1" gridrow="13" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,666,686]" gridcol="2" gridrow="13" pageId="7" pageNumber="112">30-1/2 37</td>
<td box="[920,1119,666,686]" gridcol="3" gridrow="13" pageId="7" pageNumber="112">(33)34-1/2 37</td>
<td box="[1129,1231,666,686]" gridcol="4" gridrow="13" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,666,686]" gridcol="5" gridrow="13" pageId="7" pageNumber="112">Simple 13, 14</td>
</tr>
<tr box="[134,1453,690,710]" gridrow="14" pageId="7" pageNumber="112">
<th box="[134,449,690,710]" gridcol="0" gridrow="14" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan, Decaens" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,407,690,710]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="darioi">
<emphasis box="[134,224,690,710]" italics="true" pageId="7" pageNumber="112">Sc. darioi</emphasis>
Marchán, Decäens
</taxonomicName>
&amp;
</th>
<td box="[464,720,690,710]" gridcol="1" gridrow="14" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,690,710]" gridcol="2" gridrow="14" pageId="7" pageNumber="112">26-35</td>
<td box="[920,1119,690,710]" gridcol="3" gridrow="14" pageId="7" pageNumber="112">29-33</td>
<td box="[1129,1231,690,710]" gridcol="4" gridrow="14" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,690,710]" gridcol="5" gridrow="14" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,714,734]" gridrow="15" pageId="7" pageNumber="112">
<th box="[134,1453,714,734]" colspan="6" colspanRight="5" gridcol="0" gridrow="15" pageId="7" pageNumber="112">
Domínguez,
<taxonomicNameLabel box="[268,323,714,734]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,738,758]" gridrow="16" pageId="7" pageNumber="112">
<th box="[134,449,738,758]" gridcol="0" gridrow="16" pageId="7" pageNumber="112">
<taxonomicName authority="(Qiu &amp; Bouche, 1998)" baseAuthorityName="Qiu &amp; Bouche" baseAuthorityYear="1998" box="[134,445,738,758]" class="Clitellata" family="Lumbricidae" genus="Eumenescolex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="emiliae">
<emphasis box="[134,237,738,758]" italics="true" pageId="7" pageNumber="112">Sc. emiliae</emphasis>
(Qiu &amp; Bouché, 1998)
</taxonomicName>
</th>
<td box="[464,720,738,758]" gridcol="1" gridrow="16" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,738,758]" gridcol="2" gridrow="16" pageId="7" pageNumber="112">26-35</td>
<td box="[920,1119,738,758]" gridcol="3" gridrow="16" pageId="7" pageNumber="112">28-32(33)</td>
<td box="[1129,1231,738,758]" gridcol="4" gridrow="16" pageId="7" pageNumber="112">11,12</td>
<td box="[1251,1453,738,758]" gridcol="5" gridrow="16" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,762,782]" gridrow="17" pageId="7" pageNumber="112">
<th box="[134,1453,762,782]" colspan="6" colspanRight="5" gridcol="0" gridrow="17" pageId="7" pageNumber="112">n. comb.</th>
</tr>
<tr box="[134,1453,786,807]" gridrow="18" pageId="7" pageNumber="112">
<th box="[134,449,786,807]" gridcol="0" gridrow="18" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan, Decaens" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,421,786,807]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="litoralis">
<emphasis box="[134,237,786,807]" italics="true" pageId="7" pageNumber="112">Sc. litoralis</emphasis>
Marchán, Decäens
</taxonomicName>
&amp;
</th>
<td box="[464,720,786,807]" gridcol="1" gridrow="18" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,786,807]" gridcol="2" gridrow="18" pageId="7" pageNumber="112">27-36</td>
<td box="[920,1119,786,807]" gridcol="3" gridrow="18" pageId="7" pageNumber="112">30-34(35)</td>
<td box="[1129,1231,786,807]" gridcol="4" gridrow="18" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,786,807]" gridcol="5" gridrow="18" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,810,831]" gridrow="19" pageId="7" pageNumber="112" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th box="[134,449,810,831]" gridcol="0" gridrow="19" pageId="7" pageNumber="112">
Domínguez,
<taxonomicNameLabel box="[268,323,811,831]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,835,855]" gridrow="20" pageId="7" pageNumber="112">
<th box="[134,449,835,855]" gridcol="0" gridrow="20" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan, Decaens" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,423,835,855]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="marceli">
<emphasis box="[134,239,835,855]" italics="true" pageId="7" pageNumber="112">Sc. marceli</emphasis>
Marchán, Decäens
</taxonomicName>
&amp;
</th>
<td box="[464,720,835,855]" gridcol="1" gridrow="20" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,835,855]" gridcol="2" gridrow="20" pageId="7" pageNumber="112">26-35</td>
<td box="[920,1119,835,855]" gridcol="3" gridrow="20" pageId="7" pageNumber="112">(29)30-33</td>
<td box="[1129,1231,835,855]" gridcol="4" gridrow="20" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,835,855]" gridcol="5" gridrow="20" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,859,879]" gridrow="21" pageId="7" pageNumber="112">
<th box="[134,1453,859,879]" colspan="6" colspanRight="5" gridcol="0" gridrow="21" pageId="7" pageNumber="112">
Domínguez,
<taxonomicNameLabel box="[269,323,859,879]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,883,903]" gridrow="22" pageId="7" pageNumber="112">
<th box="[134,449,883,903]" gridcol="0" gridrow="22" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,378,883,903]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="mausoleana">
<emphasis box="[134,285,883,903]" italics="true" pageId="7" pageNumber="112">Sc. mausoleana</emphasis>
Marchán
</taxonomicName>
,
</th>
<td box="[464,720,883,903]" gridcol="1" gridrow="22" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,883,903]" gridcol="2" gridrow="22" pageId="7" pageNumber="112">(26)27-36(37)</td>
<td box="[920,1119,883,903]" gridcol="3" gridrow="22" pageId="7" pageNumber="112">30-34(35)</td>
<td box="[1129,1231,883,903]" gridcol="4" gridrow="22" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,883,903]" gridcol="5" gridrow="22" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,907,927]" gridrow="23" pageId="7" pageNumber="112">
<th box="[134,1453,907,927]" colspan="6" colspanRight="5" gridcol="0" gridrow="23" pageId="7" pageNumber="112">
Decäens &amp; Domínguez,
<taxonomicNameLabel box="[377,431,907,927]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,931,951]" gridrow="24" pageId="7" pageNumber="112">
<th box="[134,449,931,951]" gridcol="0" gridrow="24" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan" authorityName="Marchan &amp; Decaens" authorityYear="2020" box="[134,373,931,951]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="portcrosana">
<emphasis box="[134,284,931,951]" italics="true" pageId="7" pageNumber="112">Sc. portcrosana</emphasis>
Marchán
</taxonomicName>
&amp;
</th>
<td box="[464,720,931,951]" gridcol="1" gridrow="24" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,931,951]" gridcol="2" gridrow="24" pageId="7" pageNumber="112">26-35(½ 36)</td>
<td box="[920,1119,931,951]" gridcol="3" gridrow="24" pageId="7" pageNumber="112">30-33</td>
<td box="[1129,1231,931,951]" gridcol="4" gridrow="24" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,931,951]" gridcol="5" gridrow="24" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,955,975]" gridrow="25" pageId="7" pageNumber="112">
<th box="[134,1453,955,975]" colspan="6" colspanRight="5" gridcol="0" gridrow="25" pageId="7" pageNumber="112">Decäens, 2020</th>
</tr>
<tr box="[134,1453,979,999]" gridrow="26" pageId="7" pageNumber="112">
<th box="[134,720,979,999]" colspan="2" colspanRight="1" gridcol="0" gridrow="26" pageId="7" pageNumber="112">
<taxonomicName authority="Qiu &amp; Bouche, 1998" authorityName="Qiu &amp; Bouche" authorityYear="1998" box="[134,461,979,999]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="portonana">
<emphasis box="[134,267,979,999]" italics="true" pageId="7" pageNumber="112">Sc. portonana</emphasis>
Qiu &amp; Bouché, 1998
</taxonomicName>
12/13, 13/14 simple
</th>
<td box="[738,909,979,999]" gridcol="2" gridrow="26" pageId="7" pageNumber="112">(25)26-35</td>
<td box="[920,1119,979,999]" gridcol="3" gridrow="26" pageId="7" pageNumber="112">(28)29-32(33)</td>
<td box="[1129,1231,979,999]" gridcol="4" gridrow="26" pageId="7" pageNumber="112">9,10,11,12</td>
<td box="[1251,1453,979,999]" gridcol="5" gridrow="26" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,1003,1024]" gridrow="27" pageId="7" pageNumber="112">
<th box="[134,449,1003,1024]" gridcol="0" gridrow="27" pageId="7" pageNumber="112">
<taxonomicName authority="Marchan, Decaens" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[134,389,1003,1024]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="qiui">
<emphasis box="[134,205,1003,1024]" italics="true" pageId="7" pageNumber="112">Sc. qiui</emphasis>
Marchán, Decäens
</taxonomicName>
&amp;
</th>
<td box="[464,720,1003,1024]" gridcol="1" gridrow="27" pageId="7" pageNumber="112">12/13, 13/14 simple</td>
<td box="[738,909,1003,1024]" gridcol="2" gridrow="27" pageId="7" pageNumber="112">(26)27-35(36)</td>
<td box="[920,1119,1003,1024]" gridcol="3" gridrow="27" pageId="7" pageNumber="112">30-33(1/n 34)</td>
<td box="[1129,1231,1003,1024]" gridcol="4" gridrow="27" pageId="7" pageNumber="112">*11,12</td>
<td box="[1251,1453,1003,1024]" gridcol="5" gridrow="27" pageId="7" pageNumber="112">Simple 12,13</td>
</tr>
<tr box="[134,1453,1027,1048]" gridrow="28" pageId="7" pageNumber="112" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th box="[134,449,1027,1048]" gridcol="0" gridrow="28" pageId="7" pageNumber="112">
Domínguez,
<taxonomicNameLabel box="[269,323,1028,1048]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
</th>
</tr>
<tr box="[134,1453,1052,1072]" gridrow="29" pageId="7" pageNumber="112">
<th box="[134,449,1052,1072]" gridcol="0" gridrow="29" pageId="7" pageNumber="112">
<taxonomicName authority="(Baldasseroni, 1906)" baseAuthorityName="Baldasseroni" baseAuthorityYear="1906" box="[134,445,1052,1072]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="targionii">
<emphasis box="[134,246,1052,1072]" italics="true" pageId="7" pageNumber="112">Sc. targionii</emphasis>
(Baldasseroni, 1906)
</taxonomicName>
</th>
<td box="[464,720,1052,1072]" gridcol="1" gridrow="29" pageId="7" pageNumber="112">11/12,12/13 simple/double</td>
<td box="[738,909,1052,1072]" gridcol="2" gridrow="29" pageId="7" pageNumber="112">(26)27-40</td>
<td box="[920,1119,1052,1072]" gridcol="3" gridrow="29" pageId="7" pageNumber="112">27-40</td>
<td box="[1129,1231,1052,1072]" gridcol="4" gridrow="29" pageId="7" pageNumber="112">11,12</td>
<td box="[1251,1453,1052,1072]" gridcol="5" gridrow="29" pageId="7" pageNumber="112">Simple/double 11,12</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="7" pageNumber="112" type="diagnosis">
<paragraph blockId="7.[130,776,1177,1624]" pageId="7" pageNumber="112">
DIAGNOSIS. —
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[272,772,1177,1201]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="altarocca" status="n. sp.">
<emphasis box="[272,456,1177,1201]" italics="true" pageId="7" pageNumber="112">Scherotheca altarocca</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[130,181,1204,1227]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
can be distinguished from all of the other species of
<taxonomicName authorityName="Bouche" authorityYear="1972" box="[666,772,1203,1227]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="genus">
<emphasis box="[666,772,1203,1227]" italics="true" pageId="7" pageNumber="112">Scherotheca</emphasis>
</taxonomicName>
(except
<taxonomicName authority="Qiu &amp; Bouche, 1998" authorityName="Qiu &amp; Bouche" authorityYear="1998" box="[208,589,1230,1254]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="subSpecies" species="corsicana" subSpecies="popi">
<emphasis box="[208,372,1230,1253]" italics="true" pageId="7" pageNumber="112">Sc. corsicana popi</emphasis>
Qiu &amp; Bouché, 1998
</taxonomicName>
) by the position of the clitellum in segments (29)30-36(37) and
<emphasis box="[586,775,1256,1280]" italics="true" pageId="7" pageNumber="112">tubercula pubertatis</emphasis>
in segments (½ 32)33-36(37) (
<tableCitation box="[437,521,1283,1307]" captionStart="TABLE" captionStartId="5.[131,142,219,237]" captionTargetPageId="5" captionText="TABLE 1. — Morphological characters of the newly described species of Corsican Scherotheca Bouché, 1972 and their closest relatives. Length in millimetres; weight in grams; 1ST D.P., position of first dorsal pore. Symbols: *, according to Pop (1947): composite between the different populations included in the original description; **, according to Bouché (1972)." httpUri="http://table.plazi.org/id/2CEBE997FFC17458FF67C2A3A2B6FE82" pageId="7" pageNumber="112" tableUuid="2CEBE997FFC17458FF67C2A3A2B6FE82">Tables 1</tableCitation>
;
<tableCitation box="[534,548,1283,1307]" captionStart="TABLE" captionStartId="7.[132,143,219,237]" captionTargetPageId="7" captionText="TABLE 2. — Reproduction-related morphological characters of the newly described species of Corsican Scherotheca Bouché, 1972 and their closest relatives. Abbreviations: Sperm. pores, position of spermathecal pores; S.V., position of seminal vesicles; T.P., position of tubercula pubertatis. Symbols: *, according to Pop (1947): composite between the different populations included in the original description; **, according to Bouché (1972)." httpUri="http://table.plazi.org/id/2CEBE997FFC3745AFF60C2A3A7D6FE82" pageId="7" pageNumber="112" tableUuid="2CEBE997FFC3745AFF60C2A3A7D6FE82">2</tableCitation>
;
<figureCitation box="[560,620,1283,1307]" captionStart="FIG" captionStartId="8.[132,143,1935,1953]" captionTargetBox="[204,1366,211,1864]" captionTargetId="figure-101@8.[165,1136,207,1872]" captionTargetPageId="8" captionText="FIG. 2. — External and internal morphological differences between the newly described species of Corsican Scherotheca Bouché,1972 and their closest relatives. The three different shades represent absent, slight and moderate pigmentation. Open circles represent structures that only appear in some individuals. Structures shown in light and dark shades (genital papillae, clitellum and tubercula pubertatis) represent the maximum and minimum extension of these characters, respectively. Phylogenetic relationships are the same as those shown in Figure 1." figureDoi="http://doi.org/10.5281/zenodo.7681301" httpUri="https://zenodo.org/record/7681301/files/figure.png" pageId="7" pageNumber="112">Figs 2</figureCitation>
;
<figureCitation box="[632,664,1283,1307]" captionStart="FIG" captionStartId="10.[131,142,1916,1934]" captionTargetBox="[132,1455,214,1870]" captionTargetId="figure-138@10.[699,1268,817,1382]" captionTargetPageId="10" captionText="FIG. 3. — External morphology of the newly described species of Corsican Scherotheca Bouché,1972:A, Scherotheca altarocca Marchán,Decäens &amp; Domínguez, n. sp.; B, Scherotheca marceli Marchán, Decäens &amp; Domínguez, n. sp.; C, Scherotheca darioi Marchán, Decäens &amp; Domínguez, n. sp.; D, Scherotheca boccaverghju Marchán, Decäens &amp; Domínguez, n. sp.; E, Scherotheca qiui Marchán, Decäens &amp; Domínguez, n. sp.; F, Scherotheca litoralis Marchán, Decäens &amp; Domínguez, n. sp.; G, Scherotheca capcorsana Marchán, Decäens &amp; Domínguez, n. sp.; H, Scherotheca mausoleana Marchán, Decäens &amp; Domínguez, n. sp. Left side, lateral view; right side, ventral view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7681305" httpUri="https://zenodo.org/record/7681305/files/figure.png" pageId="7" pageNumber="112">3A</figureCitation>
). Although
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[133,604,1309,1333]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="altarocca" status="n. sp.">
<emphasis box="[133,254,1309,1333]" italics="true" pageId="7" pageNumber="112">Sc. altarocca</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[616,672,1310,1333]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName authority="Qiu &amp; Bouche, 1998" authorityName="Qiu &amp; Bouche" authorityYear="1998" box="[134,597,1336,1360]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="subSpecies" species="corsicana" subSpecies="popi">
<emphasis box="[134,380,1336,1360]" italics="true" pageId="7" pageNumber="112">Scherotheca corsicana popi</emphasis>
Qiu &amp; Bouché,
<quantity box="[546,597,1336,1360]" metricMagnitude="1" metricUnit="m" metricValue="5.07492" pageId="7" pageNumber="112" unit="in" value="1998.0">1998</quantity>
</taxonomicName>
in characters such as the clitellum and
<emphasis box="[331,518,1362,1386]" italics="true" pageId="7" pageNumber="112">tubercula pubertatis</emphasis>
, it does not exactly fit the description made by
<bibRefCitation author="QIU J. P. &amp; BOUCHE M." box="[332,559,1388,1413]" pageId="7" pageNumber="112" pagination="117 - 139" refId="ref17184" refString="QIU J. P. &amp; BOUCHE M. 1998 a. - Revision morphologique, chorologique et taxonomique du genre Scherotheca Bouche, 1972 (Oligochaeta: Lumbricidae). Documents pedozoologiques et integrologiques 4 (12): 117 - 139." type="journal article" year="1998">Qiu &amp; Bouché (1998a)</bibRefCitation>
, differing significantly in internal characters such as position of spermathecae and number of seminal vesicles (spermathecae in (11)12, 13 vs 13, 14 and 2 pairs of seminal vesicles in
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" box="[329,772,1468,1492]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="altarocca" status="n. sp.">
<emphasis box="[329,442,1468,1492]" italics="true" pageId="7" pageNumber="112">Sc. altarocca</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[131,184,1495,1518]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
vs 4 pairs in
<taxonomicName authorityName="Qiu &amp; Bouche" authorityYear="1998" box="[313,477,1495,1518]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="subSpecies" species="corsicana" subSpecies="popi">
<emphasis box="[313,477,1495,1518]" italics="true" pageId="7" pageNumber="112">Sc. corsicana popi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="7.[130,776,1177,1624]" pageId="7" pageNumber="112">
COI uncorrected average pairwise distances and topology of multilocus molecular phylogenetic trees supports the status of
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="altarocca" status="n. sp.">
<emphasis box="[663,774,1547,1571]" italics="true" pageId="7" pageNumber="112">Sc. altarocca</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[467,518,1574,1597]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
as independent from other morphologically related species for which sequences are available.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="112" type="distribution">
<paragraph blockId="7.[132,776,1648,1778]" pageId="7" pageNumber="112">
DISTRIBUTION AND ECOLOGY.
<emphasis box="[438,463,1648,1671]" italics="true" pageId="7" pageNumber="112">—</emphasis>
<taxonomicName authority="Marchan, Decaens &amp; Dominguez" authorityName="Marchan, Decaens &amp; Dominguez" authorityYear="2023" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="112" phylum="Annelida" rank="species" species="altarocca" status="n. sp.">
<emphasis box="[471,672,1648,1672]" italics="true" pageId="7" pageNumber="112">Scherotheca altarocca</emphasis>
Marchán, Decäens &amp; Domínguez
</taxonomicName>
,
<taxonomicNameLabel box="[376,430,1675,1698]" pageId="7" pageNumber="112" rank="species">n. sp.</taxonomicNameLabel>
has been found in the municipality of Olivese and Zicavo, in the natural region of Altarocca,
<collectingCountry box="[687,744,1701,1725]" name="France" pageId="7" pageNumber="112">Corse</collectingCountry>
du Sud (
<figureCitation box="[184,260,1727,1751]" captionStart="FIG" captionStartId="14.[132,143,1613,1631]" captionTargetBox="[220,1354,216,1568]" captionTargetId="figure-223@14.[220,1354,216,1568]" captionTargetPageId="14" captionText="FIG. 4. — Distribution of the genus Scherotheca Bouché, 1972 in Corsica: A, localities documented in this work of the newly described species and other Scherotheca species sampled during this work; B, distribution of Corsican species of Scherotheca and Eumenescolex Qiu &amp; Bouché, 1998 according to Bouché (1972), Qiu &amp; Bouché (1998) and Szederjesi et al. (2021)." figureDoi="http://doi.org/10.5281/zenodo.7681307" httpUri="https://zenodo.org/record/7681307/files/figure.png" pageId="7" pageNumber="112">Fig. 4A</figureCitation>
), in
<taxonomicName box="[304,442,1727,1751]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="112" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis box="[304,442,1727,1751]" italics="true" pageId="7" pageNumber="112">Fagus sylvatica</emphasis>
</taxonomicName>
L. forests and alpine grasslands at moderately high elevations (
<quantity box="[407,571,1753,1778]" metricMagnitude="3" metricUnit="m" metricValue="1.1545" metricValueMax="1.258" metricValueMin="1.051" pageId="7" pageNumber="112" unit="m" value="1154.5" valueMax="1258.0" valueMin="1051.0">1051 to 1258 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="112" type="description">
<paragraph blockId="7.[130,777,1810,2028]" box="[132,277,1810,1836]" pageId="7" pageNumber="112">DESCRIPTION</paragraph>
<paragraph blockId="7.[130,777,1810,2028]" box="[132,346,1841,1867]" pageId="7" pageNumber="112">
<emphasis box="[132,346,1841,1867]" italics="true" pageId="7" pageNumber="112">External morphology</emphasis>
</paragraph>
<paragraph blockId="7.[130,777,1810,2028]" lastBlockId="7.[810,1456,1172,1517]" pageId="7" pageNumber="112">
Body pigmentation absent. White-beige in fixed specimens (
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). Average length,
<quantity box="[445,531,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="112" unit="mm" value="80.0">80 mm</quantity>
(
<quantity box="[549,679,1905,1932]" metricMagnitude="-2" metricUnit="m" metricValue="8.05" metricValueMax="9.5" metricValueMin="6.6" pageId="7" pageNumber="112" unit="mm" value="80.5" valueMax="95.0" valueMin="66.0">66-95 mm</quantity>
,
<emphasis box="[695,735,1906,1931]" italics="true" pageId="7" pageNumber="112">n =</emphasis>
<specimenCount pageId="7" pageNumber="112" type="adult">12 adults</specimenCount>
); body cylindrical in cross-section; average number of segments 149 (
<quantity box="[297,448,1969,1996]" metricMagnitude="-1" metricUnit="m" metricValue="1.455" metricValueMax="1.56" metricValueMin="1.35" pageId="7" pageNumber="112" unit="mm" value="145.5" valueMax="156.0" valueMin="135.0">135-156 mm</quantity>
,
<emphasis box="[460,498,1970,1995]" italics="true" pageId="7" pageNumber="112">n =</emphasis>
<specimenCount box="[506,611,1969,1995]" pageId="7" pageNumber="112" type="adult">11 adults</specimenCount>
; 135 segments in the
<typeStatus box="[205,307,2001,2028]" pageId="7" pageNumber="112">holotype</typeStatus>
). Average weight (fixed specimens),
<quantity box="[704,775,2001,2028]" metricMagnitude="-3" metricUnit="kg" metricValue="1.26" pageId="7" pageNumber="112" unit="g" value="1.26">1.26 g</quantity>
(
<quantity box="[821,951,1172,1198]" metricMagnitude="-3" metricUnit="kg" metricValue="1.295" metricValueMax="1.78" metricValueMin="0.81" pageId="7" pageNumber="112" unit="g" value="1.295" valueMax="1.78" valueMin="0.81">0.81-1.78 g</quantity>
,
<emphasis box="[961,996,1173,1198]" italics="true" pageId="7" pageNumber="112">n =</emphasis>
<specimenCount box="[1002,1105,1172,1199]" pageId="7" pageNumber="112" type="adult">14 adults</specimenCount>
). Prostomium epilobous, closed. Longitudinal furrows in segments 1 and 2. First dorsal pore at intersegmental furrow 10/11. Nephridial pores “en solfège” (irregularly distributed). Spermathecal pores at intersegmental furrows (11/12) 12/13 and 13/
<quantity box="[1147,1204,1299,1326]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="7" pageNumber="112" unit="in" value="14.0">14 in</quantity>
setal line
<emphasis box="[1311,1321,1300,1325]" italics="true" pageId="7" pageNumber="112">c</emphasis>
. Male pores in segment 15, surrounded by a well-developed porophore. Female pores in segment 14. Clitellum saddle-shaped in segments (29)30-36(37).
<emphasis box="[1097,1315,1395,1421]" italics="true" pageId="7" pageNumber="112">Tubercula pubertatis</emphasis>
in segments (½ 32)33-36(37).
<emphasis box="[1013,1098,1427,1453]" italics="true" pageId="7" pageNumber="112">Chaetae</emphasis>
small and closely paired. Genital papillae in segments (10,11)12-16, (29,30)31,32,37,38(3 9,40,41,42).
</paragraph>
<paragraph blockId="7.[811,1457,1554,2028]" box="[813,992,1554,1580]" pageId="7" pageNumber="112">
<emphasis box="[813,992,1554,1580]" italics="true" pageId="7" pageNumber="112">Internal anatomy</emphasis>
</paragraph>
<paragraph blockId="7.[811,1457,1554,2028]" pageId="7" pageNumber="112">
<collectingRegion box="[813,871,1587,1613]" country="Spain" name="Ceuta" pageId="7" pageNumber="112">Septa</collectingRegion>
5/6-10/11 thickened and muscular. Hearts in segments 6-11, oesophageal. Calciferous glands in segments 10-14, with diverticula in segment 10. Crop in segments 15-16, gizzard in segments 17-19. Typhlosole pinnate.Male sexual system holandric, testes and funnels (not enclosed in testes sacs, but with sperm present) located ventrally in segments 10 and 11. Two pairs of reniform seminal vesicles in segments 11 and 12; in one individual very inconspicuous, vestigial seminal vesicles present in segments 9 and 10. Ovaries and female funnels in segment 13, ovarian receptacles (ovisacs) in segment 14. Two or three pairs of small globular spermathecae in segments (11), 12 and 13 (intersegments 11/12, 12/13, 13/14). Nephridial bladders U-shaped, reclinate in segment 30.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7681301" ID-Zenodo-Dep="7681301" httpUri="https://zenodo.org/record/7681301/files/figure.png" pageId="8" pageNumber="113" startId="8.[132,143,1935,1953]" targetBox="[204,1366,211,1864]" targetPageId="8">
<paragraph blockId="8.[132,1457,1935,2021]" pageId="8" pageNumber="113">
FIG. 2. — External and internal morphological differences between the newly described species of Corsican
<taxonomicName authority="Bouche, 1972" authorityName="Bouche" authorityYear="1972" box="[1013,1238,1935,1953]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis box="[1013,1118,1935,1953]" italics="true" pageId="8" pageNumber="113">Scherotheca</emphasis>
<bibRefCitation author="BOUCHE M. B." box="[1122,1238,1935,1953]" pageId="8" pageNumber="113" refId="ref15653" refString="BOUCHE M. B. 1972. - Lombriciens de France: ecologie et systematique (Vol. 72, No. 2). Paris: Institut National de la Recherche Agronomique." type="book" year="1972">Bouché,1972</bibRefCitation>
</taxonomicName>
and their closest relatives. The three different shades represent absent, slight and moderate pigmentation. Open circles represent structures that only appear in some individuals. Structures shown in light and dark shades (genital papillae, clitellum and
<emphasis box="[706,871,1980,1998]" italics="true" pageId="8" pageNumber="113">tubercula pubertatis</emphasis>
) represent the maximum and minimum extension of these characters, respectively. Phylogenetic relationships are the same as those shown in Figure 1.
</paragraph>
</caption>
<subSubSection pageId="9" pageNumber="114" type="discussion">
<paragraph blockId="9.[131,777,216,529]" box="[132,232,216,242]" pageId="9" pageNumber="114">REMARKS</paragraph>
<paragraph blockId="9.[131,777,216,529]" pageId="9" pageNumber="114">
This species displays some degree of morphological variability, with different combinations of extension of clitellum and
<emphasis box="[183,401,311,337]" italics="true" pageId="9" pageNumber="114">tubercula pubertatis</emphasis>
, position of genital papillae and spermathecae. These different morphotypes do not show any clear association with genetic lineages, and therefore they do not appear to correspond to differentiated taxa within the species described here.
</paragraph>
<paragraph blockId="9.[131,777,216,529]" pageId="9" pageNumber="114">
This species corresponds to
<taxonomicName authorityName="* (Pop" authorityYear="1947" box="[446,575,471,497]" class="Clitellata" family="Lumbricidae" genus="Scherotheca" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="114" phylum="Annelida" rank="species" species="corsicana">
<emphasis box="[446,575,471,497]" italics="true" pageId="9" pageNumber="114">Sc. corsicana</emphasis>
</taxonomicName>
L
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the checklist of Corsican earthworms of
<bibRefCitation author="MARCHAN D. F. &amp; GERARD S. &amp; HEDDE M. &amp; ROUGERIE R. &amp; DECAENS T." box="[427,676,502,529]" pageId="9" pageNumber="114" pagination="439 - 461" refId="ref16506" refString="MARCHAN D. F., GERARD S., HEDDE M., ROUGERIE R. &amp; DECAENS T. 2022 a. - An updated checklist and a DNA barcode library for the earthworms (Crassiclitellata, Oligochaeta) of Corsica, France. Zoosystema 44 (17): 439 - 461. https: // doi. org / 10.5252 / zoosystema 2022 v 44 a 17. http: // zoosystema. com / 44 / 17" type="journal article" year="2022">
Marchán
<emphasis box="[531,583,502,528]" italics="true" pageId="9" pageNumber="114">et al.</emphasis>
(2022a)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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