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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.537.6090" ID-GBIF-Dataset="ae220ec2-27f7-43de-b96d-3b5e0dd853b6" ID-PMC="PMC4714052" ID-Pensoft-Pub="1313-2970-537-131" ID-PubMed="26798242" ID-ZBK="EA901A84CB134889A77C07E89EDA172E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-537-131" ModsDocOrigin="ZooKeys 537" ModsDocTitle="Three new species of Hagnagora Druce, 1885 (Lepidoptera, Geometridae, Larentiinae) from Ecuador and Costa Rica and a concise revision of the genus" checkinTime="1451243721882" checkinUser="pensoft" docAuthor="Brehm, Gunnar" docDate="2015" docId="BCEB2CEF5BBF5E09E3608365D82BFC9A" docLanguage="en" docName="ZooKeys 537: 131-156" docOrigin="ZooKeys 537" docSource="http://dx.doi.org/10.3897/zookeys.537.6090" docTitle="Hagnagora richardi Brehm, sp. n." docType="treatment" docUuid="406E12C4-4231-49F2-BE51-61E504E395F7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="139" masterDocId="FFCE8A69FFE9FFA2FFE1FFD8EE5D462F" masterDocTitle="Three new species of Hagnagora Druce, 1885 (Lepidoptera, Geometridae, Larentiinae) from Ecuador and Costa Rica and a concise revision of the genus" masterLastPageNumber="156" masterPageNumber="131" pageNumber="138" updateTime="1668160957273" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of Hagnagora Druce, 1885 (Lepidoptera, Geometridae, Larentiinae) from Ecuador and Costa Rica and a concise revision of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brehm, Gunnar</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>537</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>131</mods:start>
<mods:end>156</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.537.6090</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.537.6090</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-537-131</mods:identifier>
<mods:identifier type="ZBK">EA901A84CB134889A77C07E89EDA172E</mods:identifier>
<mods:identifier type="ZooBank">EA901A84CB134889A77C07E89EDA172E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127897772" LSID="urn:lsid:zoobank.org:act:406E12C4-4231-49F2-BE51-61E504E395F7" httpUri="http://treatment.plazi.org/id/BCEB2CEF5BBF5E09E3608365D82BFC9A" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138">
<subSubSection pageId="7" pageNumber="138" type="multiple">
<paragraph pageId="7" pageNumber="138">Taxon classification Animalia Lepidoptera Geometridae</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="138" type="nomenclature">
<paragraph pageId="7" pageNumber="138">
<taxonomicName LSID="http://zoobank.org/406E12C4-4231-49F2-BE51-61E504E395F7" authority="Brehm" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi Brehm</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="138">sp. n.</taxonomicNameLabel>
Figs 9, 13
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138" type="type material">
<paragraph pageId="7" pageNumber="138">Type material.</paragraph>
<paragraph pageId="7" pageNumber="138">
Holotype: male (Fig. 9): Ecuador, Loja province, Parque Nacional Podocarpus, Cajanuma,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-4.1141667">04°06.85'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-79.1745">79°10.47'W</geoCoordinate>
, 2916 m, 20 November 2008, G. Brehm leg. (ID 18080, genitalia preparation, barcode sequence 658 bp) (PMJ).
</paragraph>
<paragraph pageId="8" pageNumber="139">
<pageBreakToken pageId="8" pageNumber="139" start="start">Paratypes</pageBreakToken>
: (deposited in CISEC, PMJ, RCGB, ZSM) 4 males, 2 females. 1 female: same as holotype but
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-4.114333">04°06.86'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-79.17433">79°10.46'W</geoCoordinate>
, 2897 m, F. Bodner leg. (ID 15855, barcode sequence 658 bp); 1 female (Fig. 13): Ecuador, Zamora Chinchipe, Reserva
<normalizedToken originalValue="Biológica">Biologica</normalizedToken>
San Francisco,
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-3.9786668">03°58.72'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-79.074">79°04.44'W</geoCoordinate>
, 2180 m, 16 November 2008, F. Bodner leg. (ID 16285, barcode sequence 658 bp); 1 male same as previous but 28 October 1999, D.
<normalizedToken originalValue="Süßenbach">Suessenbach</normalizedToken>
leg. (BC ZSM Lep 04774, barcode sequence 529 bp); 1 male: same as previous but
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-3.9941666">03°59.65'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-79.06834">79°04.10'W</geoCoordinate>
, 2670 m, G. Brehm leg. (ID 17328, barcode sequence 658 bp); 1 male as previous but
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-3.9946666">03°59.68'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-79.06834">79°04.10'W</geoCoordinate>
, 2677 m, 18 November 2008 (ID 16119, barcode sequence 658 bp); 1 male as previous but 25 November 2008 (ID 17863, barcode sequence 621 bp).
</paragraph>
<caption pageId="8" pageNumber="139">
<paragraph pageId="8" pageNumber="139">
Figures 12-14. 12
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
(F&amp;R), female from Ecuador as reference specimen with Barcode Index Number (BIN) a dorsal view b ventral view c genitalia 13
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
sp. n., female, paratype a dorsal view b ventral view c genitalia 14
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora hedwigae" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="hedwigae">Hagnagora hedwigae</taxonomicName>
sp. n., female, paratype a dorsal view b ventral view c genitalia.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="8" pageNumber="139" type="description">
<paragraph pageId="8" pageNumber="139">Description.</paragraph>
<paragraph pageId="8" pageNumber="139">As illustrated in Figs 9, 13. The wing length of the holotype (male) is 19 mm. The wing length of a female paratype (Fig. 13) is 21 mm.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="139" type="distribution">
<paragraph pageId="8" pageNumber="139">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="139">Only known from a small region around Podocarpus National Park, provinces Zamora-Chinchipe and Loja, Ecuador, with an observed elevational range of 2180-3021 m a.s.l. Apart from the</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="139" type="type locality">
<paragraph pageId="8" pageNumber="139">Type locality</paragraph>
<paragraph pageId="8" pageNumber="139">
and nearby sites, specimens were collected at elevations at ca. 3000 m at Cerro Toledo in the same National Park (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-4.383333">04°23'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-79.11667">79°07'W</geoCoordinate>
). However, this record is not fully reliable because genitalia preparation or barcoding was not conducted for these specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="139" type="diagnosis">
<paragraph pageId="8" pageNumber="139">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="139">
Closely resembles other species of the
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
clade. On average significantly larger than
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
, but the female has about the same size as
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora hedwigae" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="hedwigae">Hagnagora hedwigae</taxonomicName>
. The uncus of the male is larger and broader than in
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
. The signum of the bursa copulatrix is less complex than in
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
and
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora hedwigae" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="hedwigae">Hagnagora hedwigae</taxonomicName>
. Easily distinguishable from
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora marionae" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="marionae">Hagnagora marionae</taxonomicName>
by the cream-white colour of the blotches on the forewing. COI barcode: The minimum observed distance to the presumably most closely related species (
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora marionae" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="marionae">Hagnagora marionae</taxonomicName>
) is 6.6%.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="139" type="etymology">
<paragraph pageId="8" pageNumber="139">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="139">
<taxonomicName class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
is named in honour of Richard Philipp from Jena, Germany, in recognition of his and his
<normalizedToken originalValue="parents">parents'</normalizedToken>
support for the taxonomy of Neotropical geometrid moths.
</paragraph>
</subSubSection>
</treatment>
</document>