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<document id="D839B5C95CDCA3953751F2966CC7A3D9" ID-ISSN="1175­5334" approvalRequired="298" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="51" approvalRequired_for_textStreams="166" approvalRequired_for_treatments="80" checkinTime="1627004795516" checkinUser="felipe" docAuthor="Oláh, J. &amp; Johanson, K. A." docDate="2008" docId="BC22C322172FA93E989DFCCC79DFFED6" docLanguage="en" docName="zt01802p248.pdf" docOrigin="Zootaxa 1802" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.2:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="2" docTitle="Hydromanicus truncatus Betten" docType="treatment" docVersion="2" lastPageNumber="31" masterDocId="401BBB5A1731A91E980AFFCB7843FFFC" masterDocTitle="Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)" masterLastPageNumber="248" masterPageNumber="1" pageNumber="31" updateTime="1651296210969" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="1D4C4AEC75092E30DAB6B3E9B90C03C4">Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)</mods:title>
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<mods:namePart id="0C33DB8A9F381CBA5F6756F8D0CB2F98">Oláh, J.</mods:namePart>
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<mods:namePart id="C861D76B90BE3F421386E56C5D35B452">Johanson, K. A.</mods:namePart>
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<mods:date id="949BCF23D3AA14B57E12C2527AC42684">2008</mods:date>
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<treatment id="BC22C322172FA93E989DFCCC79DFFED6" LSID="urn:lsid:plazi:treatment:BC22C322172FA93E989DFCCC79DFFED6" httpUri="http://treatment.plazi.org/id/BC22C322172FA93E989DFCCC79DFFED6" lastPageId="32" lastPageNumber="31" pageId="30" pageNumber="31">
<subSubSection id="7C9121BF172FA900989DFCCC7A5EFCDD" box="[151,541,775,801]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="34347234172FA900989DFCCC7A5EFCDD" blockId="30.[151,541,775,842]" box="[151,541,775,801]" pageId="30" pageNumber="31">
<heading id="6F7CC558172FA900989DFCCC7A5EFCDD" bold="true" box="[151,541,775,801]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="F38B09B7172FA900989DFCCC7A5EFCDD" ID-CoL="3N7KB" ID-ENA="761871" authority="Betten" authorityName="Betten" box="[151,541,775,801]" class="Insecta" family="Hydropsychidae" genus="Hydromanicus" kingdom="Animalia" order="Trichoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="06FFAE26172FA900989DFCCC7A5EFCDD" bold="true" box="[151,541,775,801]" pageId="30" pageNumber="31">
<emphasis id="06FFAE26172FA900989DFCCC7987FCDD" bold="true" box="[151,452,775,801]" italics="true" pageId="30" pageNumber="31">Hydromanicus truncatus</emphasis>
Betten
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF172FA900989DFCFB7956FCB6" box="[151,277,816,842]" pageId="30" pageNumber="31" type="description">
<paragraph id="34347234172FA900989DFCFB7956FCB6" blockId="30.[151,541,775,842]" box="[151,277,816,842]" pageId="30" pageNumber="31">
<figureCitation id="ACB06EB1172FA900989DFCFB7956FCB6" box="[151,277,816,842]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="30" pageNumber="31">Fig. 3841</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF172FA900989DFC4B7A29FC6B" box="[151,618,896,919]" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="34347234172FA900989DFC4B7A29FC6B" blockId="30.[151,618,896,919]" box="[151,618,896,919]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="149B551A172FA900989DFC4B7A29FC6B" box="[151,618,896,919]" pageId="30" pageNumber="31">
<treatmentCitation id="B52A5425172FA900989DFC4B7A25FC6B" author="Betten, C." box="[151,614,896,919]" page="235" pageId="30" pageNumber="31" year="1909">
<taxonomicName id="F38B09B7172FA900989DFC4B7A25FC6B" ID-CoL="3N7KB" authority="Betten, 1909: 235" authorityName="Betten" authorityPageNumber="235" authorityYear="1909" box="[151,614,896,919]" class="Insecta" family="Hydropsychidae" genus="Hydromanicus" kingdom="Animalia" order="Trichoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="06FFAE26172FA900989DFC4B79DDFC6B" box="[151,414,896,919]" italics="true" pageId="30" pageNumber="31">Hydromanicus truncatus</emphasis>
<bibRefCitation id="501A0FC5172FA90099AFFC4B7A25FC6B" author="Betten, C." box="[421,614,896,919]" pageId="30" pageNumber="31" pagination="231 - 242" refId="ref106649" refString="Betten, C. (1909) Notes on the Trichoptera in the collection of the Indian Museum. Records of the Indian Museum, 3, 231 - 242." type="journal article" year="1909">Betten, 1909: 235</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
</subSubSection>
<subSubSection id="7C9121BF172FA93E989DFC0C79DFFED6" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="34347234172FA900989DFC0C79B6FC1E" blockId="30.[151,1436,967,2034]" box="[151,501,967,994]" pageId="30" pageNumber="31">
<materialsCitation id="84E37869172FA900989DFC0C79B2FC1E" box="[151,497,967,994]" country="India" location="India" pageId="30" pageNumber="31" specimenCount="1" typeStatus="holotype">
<emphasis id="06FFAE26172FA900989DFC0C797FFC1D" bold="true" box="[151,316,967,993]" pageId="30" pageNumber="31">
<typeStatus id="EB30CC96172FA900989DFC0C7897FC1D" box="[151,212,967,993]" pageId="30" pageNumber="31">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="4C9C32A4172FA900994EFC0379C3FC1E" box="[324,384,968,994]" name="India" pageId="30" pageNumber="31">India</collectingCountry>
(Bengal)
</materialsCitation>
.
</paragraph>
<paragraph id="34347234172FA90098CCFC24793CFBA6" blockId="30.[151,1436,967,2034]" pageId="30" pageNumber="31">
<emphasis id="06FFAE26172FA90098CCFC24792FFBF5" bold="true" box="[198,364,1007,1033]" pageId="30" pageNumber="31">New Records</emphasis>
.
<materialsCitation id="84E37869172FA9009976FC2478B9FBCE" collectionCode="NHML" collectorName="Nat. Coll. McLachlan &amp; Coll. McLachlan &amp; Coll. B. M." country="India" location="Khasia Hills" pageId="30" pageNumber="31" specimenCount="2" specimenCount-male="2" stateProvince="Meghalaya">
<emphasis id="06FFAE26172FA9009976FC247991FBF5" bold="true" box="[380,466,1007,1033]" pageId="30" pageNumber="31">
<collectingCountry id="4C9C32A4172FA9009976FC247991FBF5" box="[380,466,1007,1033]" name="India" pageId="30" pageNumber="31">INDIA</collectingCountry>
</emphasis>
:
<collectingRegion id="F64FBCD6172FA90099E8FC3B7A24FBF6" box="[482,615,1008,1034]" country="India" name="Meghalaya" pageId="30" pageNumber="31">Meghalaya</collectingRegion>
,
<location id="315424EF172FA9009A7EFC3B7B45FBF6" LSID="urn:lsid:plazi:treatment:BC22C322172FA93E989DFCCC79DFFED6:315424EF172FA9009A7EFC3B7B45FBF6" box="[628,774,1008,1034]" country="India" name="Khasia Hills" pageId="30" pageNumber="31" stateProvince="Meghalaya">Khasia Hills</location>
<emphasis id="06FFAE26172FA9009B04FC247B5BFBF5" bold="true" box="[782,792,1007,1033]" pageId="30" pageNumber="31">-</emphasis>
<specimenCount id="228DB9BD172FA9009B2AFC3B7B38FBF6" box="[800,891,1008,1034]" pageId="30" pageNumber="31" type="male">2 males</specimenCount>
(
<collectorName id="997E17E2172FA9009B86FC3B7CD2FBF6" box="[908,1169,1008,1034]" pageId="30" pageNumber="31">Nat. Coll. McLachlan</collectorName>
,
<collectorName id="997E17E2172FA9009C94FC3B7D5FFBF6" box="[1182,1308,1008,1034]" pageId="30" pageNumber="31">Coll. B.M.</collectorName>
1938-674,
<collectionCode id="529AEAF1172FA900989DFBD378B0FBCE" box="[151,243,1048,1074]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="30" pageNumber="31">NHML</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E37869172FA900990CFBD37938FBA6" collectingDate="1961-03-11" collectionCode="ROM" collectorName="F. Schmid" country="India" location="Amatulla" municipality="Kameng" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Assam">
<collectingRegion id="F64FBCD6172FA900990CFBD3791AFBCE" box="[262,345,1048,1074]" country="India" name="Assam" pageId="30" pageNumber="31">Assam</collectingRegion>
,
<collectingMunicipality id="D450E84E172FA900996EFBD37989FBCE" box="[356,458,1048,1074]" pageId="30" pageNumber="31">Kameng</collectingMunicipality>
,
<location id="315424EF172FA90099DFFBD37A06FBCE" LSID="urn:lsid:plazi:treatment:BC22C322172FA93E989DFCCC79DFFED6:315424EF172FA90099DFFBD37A06FBCE" box="[469,581,1048,1074]" country="India" municipality="Kameng" name="Amatulla" pageId="30" pageNumber="31" stateProvince="Assam">Amatulla</location>
,
<date id="403554F4172FA9009A5BFBD37A8CFBCE" box="[593,719,1048,1074]" pageId="30" pageNumber="31" value="1961-03-11">
<collectingDate id="5071AD1C172FA9009A5BFBD37A8CFBCE" box="[593,719,1048,1074]" pageId="30" pageNumber="31" value="1961-03-11">11.iii.1961</collectingDate>
</date>
[
<collectorName id="997E17E2172FA9009AD7FBD37B1BFBCE" box="[733,856,1048,1074]" pageId="30" pageNumber="31">F. Schmid</collectorName>
] -
<specimenCount id="228DB9BD172FA9009B7CFBD37B87FBCE" box="[886,964,1048,1074]" pageId="30" pageNumber="31" type="male">1 male</specimenCount>
(
<collectionCode id="529AEAF1172FA9009BDEFBD37C58FBCE" box="[980,1051,1048,1074]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="30" pageNumber="31" type="Museum">ROM</collectionCode>
, pinned with cleared abdomen in plastic genital vial)
</materialsCitation>
.
</paragraph>
<paragraph id="34347234172FA90098CCFBA37B04FAB6" blockId="30.[151,1436,967,2034]" pageId="30" pageNumber="31">
Male. Body and wings brown. Forewing membrane medium brown with maculation pattern, pale stripe present on anterior margin of forewing median cell. Sternum
<collectionCode id="529AEAF1172FA9009B7BFB5B7BC5FB56" box="[881,902,1168,1194]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="31" type="Museum">V</collectionCode>
without visible lobe or protuberance. Maxillary palp formula I-(
<collectionCode id="529AEAF1172FA9009983FB7379DCFB2E" box="[393,415,1208,1234]" pageId="30" pageNumber="31">II</collectionCode>
,
<collectionCode id="529AEAF1172FA90099A1FB737988FB2E" box="[427,459,1208,1234]" pageId="30" pageNumber="31">III</collectionCode>
,
<collectionCode id="529AEAF1172FA90099D2FB7379BBFB2E" box="[472,504,1208,1234]" pageId="30" pageNumber="31">IV</collectionCode>
)-
<collectionCode id="529AEAF1172FA9009A03FB737A5CFB2E" box="[521,543,1208,1234]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="31" type="Museum">V</collectionCode>
, segment
<collectionCode id="529AEAF1172FA9009A9FFB737AE9FB2E" box="[661,682,1208,1234]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="31" type="Museum">V</collectionCode>
as long as length of segments I
<collectionCode id="529AEAF1172FA9009C24FB737C0EFB2E" box="[1070,1101,1208,1234]" pageId="30" pageNumber="31">IV</collectionCode>
together. Head dorsum dark brown, with 7 dark brown setal warts. Swollen setal wart absent on proepisternum. Small setal wart present on precoxale (proepisternum). Pretarsal claw asymmetrical, laterally flanked by setal bundle. Spur formula 244. Protibia with posteroapical spur equal to anteroapical spur.
</paragraph>
<paragraph id="34347234172FA90098CFFA937CFFFA3E" blockId="30.[151,1436,967,2034]" pageId="30" pageNumber="31">
Wings. Venation typical for the genus. Forewing crossveins m-cu and cu separate. Sc and
<collectionCode id="529AEAF1172FA9009D17FA937D73FA8E" box="[1309,1328,1368,1394]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="30" pageNumber="31">R</collectionCode>
separate before costa. Cu2 and A fusing before wing margin. Hind wing Sc and
<collectionCode id="529AEAF1172FA9009BEBFA4B7BB7FA66" box="[993,1012,1408,1434]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="30" pageNumber="31">R</collectionCode>
meeting before costa at crossvein r. Stem of M and Cu1 separate. Median cell open. Fork 1 present. Forewing length 11.2 mm.
</paragraph>
<paragraph id="34347234172FA93E98CFFA1B79DFFED6" blockId="30.[151,1436,967,2034]" lastBlockId="32.[151,1436,152,298]" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
Male genitalia. Abdominal segment IX fused annularly. Slightly two-partite, longitudinal suture visible at dorsal end of broad apical lobe (
<figureCitation id="ACB06EB1172FA9009A19FA337A28F9EE" box="[531,619,1528,1554]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="30" pageNumber="31">Fig. 38</figureCitation>
); almost dividing segment IX into large ventral and small dorsal part. Median keel narrow, dorsally and laterally granulose apex open in dorsal view (
<figureCitation id="ACB06EB1172FA9009C36F9EB7CD7F9C6" box="[1084,1172,1568,1594]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="30" pageNumber="31">Fig. 39</figureCitation>
); anterior margin arciform, dorsum with keel shifted posterad. Antecosta forming single line narrowing dorsally before margin, with external groove of antecostal suture. Apical lobe on posterolateral margin very broad, rounded. Posterior spine row continuous on broad apical lobe and on two-partite dorsal lobe to median keel. Depression between segments IX and X shallow, stepwise. Segment X long, narrow, with capitate apex in lateral view (
<figureCitation id="ACB06EB1172FA9009D12F90B7D33F926" box="[1304,1392,1728,1754]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="30" pageNumber="31">Fig. 38</figureCitation>
); in dorsal view (
<figureCitation id="ACB06EB1172FA9009939F92379C9F8FE" box="[307,394,1768,1794]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="30" pageNumber="31">Fig. 39</figureCitation>
) broad, with pronounced triangular, lateral protuberances. Preanal appendages forming long processes being broadest at mid-length, covered by setae nested in elevated alveoli. Apicoventral setose lobes forming pair of digitiform and capitate processes; slightly down-curving in lateral view; straight in dorsal view; bearing setae on dorsal surfaces. Dorsal interlobular gap narrow, deep (
<figureCitation id="ACB06EB1172FA9009C47F8AB7CE6F886" box="[1101,1189,1888,1914]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="30" pageNumber="31">Fig. 39</figureCitation>
); demarcated by apicoventral digitiform processes. Apicodorsal setose lobes reduced into pair of small, setose, longitudinal surfaces between triangular lateral protuberances (
<figureCitation id="ACB06EB1172FA9009AEFF87B7B7EF836" box="[741,829,1968,1994]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="30" pageNumber="31">Fig. 39</figureCitation>
); smooth cavity on segment X invisible. Longitudinal sutures running along dorsal part of segment X, visible in lateral and dorsal view. Coxopodite long, slen- der, straight, without significant apically broadening. Each harpagones with two-lobes, median lobes larger than lateral lobes (
<figureCitation id="ACB06EB1172EA9019967FF0B7986FF26" box="[365,453,192,218]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="31" pageNumber="32">Fig. 40</figureCitation>
). Phallic apparatus (
<figureCitation id="ACB06EB1172EA9019ABBFF0B7B46FF26" box="[689,773,192,218]" captionStart="FIGURES 3841" captionStartId="31.[151,269,1911,1935]" captionTargetBox="[171,1385,472,1845]" captionTargetId="figure-142@31.[151,1436,439,1885]" captionTargetPageId="31" captionText="FIGURES 3841. Hydromanicus truncatus Betten, 1909, non-type, male. 38 — genitalia, lateral; 39 — genitalia, dorsal; 40 — gonocoxite, ventral; 41 — phallus, lateral." pageId="31" pageNumber="32">Fig. 41</figureCitation>
) basal part curving ventrad, right angled; anterior part of horizontal phallotheca very broad, slightly broader than basal part; narrowing gradually before apical ventral keel. Phallotheca ending distally in regularly circular, sclerous, endothecal process. Median rim thickened, incurving; pair of small, irregular, phallotremal sclerites located in middle position beneath sclerous, endothecal process.
<taxonomicName id="F38B09B7172EA9019927FEAB79FCFE86" authorityName="M. Zang &amp; R.H. Petersen" authorityYear="1989" box="[301,447,352,378]" class="Agaricomycetes" family="Phallaceae" genus="Endophallus" kingdom="Fungi" order="Phallales" pageId="31" pageNumber="9" phylum="Basidiomycota" rank="genus">Endophallus</taxonomicName>
with shape of phallotheca ending anteriorly in narrow, ventrad curving tube with gonopore; broadly rounded, half-ovoid, sub-apical ventral keel dominating apex of phallotheca in lateral view. Remarks: As the
<typeStatus id="EB30CC961711A93E999FFF5379B8FF4E" box="[405,507,152,178]" pageId="32" pageNumber="33">holotype</typeStatus>
of this species is not available,
<specimenCount id="228DB9BD1711A93E9B76FF537C4DFF4E" box="[892,1038,152,178]" pageId="32" pageNumber="33" type="generic">3 specimens</specimenCount>
collected in different regions but not far from the type locality, permit us to re-describe the species. The specimens from the 2 new localities, Khasia Hills in
<collectingRegion id="F64FBCD61711A93E995EFF237995FEFE" box="[340,470,232,258]" country="India" name="Meghalaya" pageId="32" pageNumber="33">Meghalaya</collectingRegion>
and Kameng in
<collectingRegion id="F64FBCD61711A93E9A95FF237AB1FEFE" box="[671,754,232,258]" country="India" name="Assam" pageId="32" pageNumber="33">Assam</collectingRegion>
, have slightly different length ratio between the lobes on the harpagonal apices.
</paragraph>
</subSubSection>
</treatment>
</document>