340 lines
48 KiB
XML
340 lines
48 KiB
XML
<document id="07F3F67F786A316FCA74E909EF7B85AD" ID-DOI="10.11646/zootaxa.4019.1.21" ID-GBIF-Dataset="845d040b-01f0-4bd4-a545-e7972311d1d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="233296" ID-ZooBank="4638C91A-F573-4DF1-9E8F-C25C9BBABDDF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461266503283" checkinUser="plazi" docAuthor="Paxton, Hannelore & Budaeva, Nataliya" docDate="2015" docId="B31F87A5FF9EBA2BC3A2FDFBFED59D4E" docLanguage="en" docName="zt04019p634.pdf" docOrigin="Zootaxa 4019 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhamphobrachium (Minibrachium) nutrix Paxton & Budaeva, 2015, n. sp." docType="treatment" docVersion="8" lastPageNumber="626" masterDocId="4F26FFDDFF9CBA2EC335FFE7FFC89827" masterDocTitle="Minibrachium, a new subgenus of Rhamphobrachium (Annelida: Onuphidae) from Australia with the description of three new species" masterLastPageNumber="634" masterPageNumber="621" pageNumber="623" updateTime="1698610514063" updateUser="plazi">
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<mods:mods id="36EE1CAF31583C4CACFF9B3B6144661D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BCA1FED901AA7AF42C7F939884A85789">
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<mods:title id="CA349983799F759E852DD1E9C759CDEC">Minibrachium, a new subgenus of Rhamphobrachium (Annelida: Onuphidae) from Australia with the description of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="4E107434A545DF9EF266AC86991F75F0" type="personal">
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<mods:role id="8A84BA490E9DF62A5D4AE50D2D964EAC">
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<mods:roleTerm id="5D93AD1F35A757C08F98A10EC62A6829">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E3DB044753539E9A094843606EEB865D">Paxton, Hannelore</mods:namePart>
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</mods:name>
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<mods:name id="B3BB029D219524F71EBD392412198858" type="personal">
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<mods:role id="A92B4168B008BE1634876BB5A777DBB7">
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<mods:roleTerm id="1DE374E101EBDB20C50B2F150DC974C5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="74C7B7CB4F69876E5B9E3284CE15BBB4">Budaeva, Nataliya</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="12B93DF24965E3DFD7CC2F4FA17C9FEA">text</mods:typeOfResource>
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<mods:titleInfo id="C2B69331C385259FB9219144060C383D">
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<mods:title id="4EC2710B3CDE4DEF66B3289E25ED9FCF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="798A16C7CF87B721A0F15A79A76BE0C7">
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<mods:date id="6CC600C7C8B919388237FE3DB04CEAB0">2015</mods:date>
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<mods:detail id="757FB9EF846BC886C7145CB5ED7AF374" type="volume">
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<mods:number id="786468613621D6F6CA2CFF98BBF810A0">4019</mods:number>
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</mods:detail>
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<mods:detail id="99CE112F2B132AE3A3A6805628374636" type="issue">
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<mods:number id="08F6BF34DC28F98DA825E26BC802262E">1</mods:number>
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<mods:start id="FD3FAF67A64524154397FBF7B90456F2">621</mods:start>
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<mods:end id="B3B4BCDE49DF5693A78AF921EC79A9BD">634</mods:end>
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</mods:extent>
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<mods:classification id="9F024F4396824D95C5910813FC7D16F1">journal article</mods:classification>
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<mods:identifier id="8ABE6F0291544A235E85AAE45E941B12" type="DOI">10.11646/zootaxa.4019.1.21</mods:identifier>
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<mods:identifier id="1B0C27DA73416A4444340FE17D02B645" type="GBIF-Dataset">845d040b-01f0-4bd4-a545-e7972311d1d0</mods:identifier>
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<mods:identifier id="7E60E64907F857E0D76790E0B62ABB2D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="52B48A26AD9223CEAAD1A11D4442700D" type="Zenodo-Dep">233296</mods:identifier>
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<mods:identifier id="2AD82ECB2A904930981E8E3CA4B9CF62" type="ZooBank">4638C91A-F573-4DF1-9E8F-C25C9BBABDDF</mods:identifier>
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</mods:mods>
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<treatment id="B31F87A5FF9EBA2BC3A2FDFBFED59D4E" ID-DOI="http://doi.org/10.5281/zenodo.5662755" ID-GBIF-Taxon="119638084" ID-Zenodo-Dep="5662755" LSID="urn:lsid:plazi:treatment:B31F87A5FF9EBA2BC3A2FDFBFED59D4E" httpUri="http://treatment.plazi.org/id/B31F87A5FF9EBA2BC3A2FDFBFED59D4E" lastPageId="5" lastPageNumber="626" pageId="2" pageNumber="623">
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<subSubSection id="73AC6538FF9EBA2CC3A2FDFBFEC39A7F" pageId="2" pageNumber="623" type="nomenclature">
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<paragraph id="3B0936B3FF9EBA2CC3A2FDFBFD219A10" blockId="2.[151,745,540,600]" box="[151,745,540,567]" pageId="2" pageNumber="623">
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<heading id="604181DFFF9EBA2CC3A2FDFBFD219A10" bold="true" box="[151,745,540,567]" fontSize="11" level="1" pageId="2" pageNumber="623" reason="1">
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<emphasis id="09C2EAA1FF9EBA2CC3A2FDFBFD219A10" bold="true" box="[151,745,540,567]" pageId="2" pageNumber="623">
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<taxonomicName id="FCB64D30FF9EBA2CC3A2FDFBFD689A11" ID-CoL="4S2LK" box="[151,672,540,566]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="2" pageNumber="623" phylum="Annelida" rank="species" species="nutrix" status="sp. nov." subGenus="Minibrachium">
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<emphasis id="09C2EAA1FF9EBA2CC3A2FDFBFD689A11" bold="true" box="[151,672,540,566]" italics="true" pageId="2" pageNumber="623">Rhamphobrachium (Minibrachium) nutrix</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="12F157DAFF9EBA2CC192FDFAFD219A10" box="[679,745,541,567]" pageId="2" pageNumber="623" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="3B0936B3FF9EBA2CC3A2FDD8FEC39A7F" blockId="2.[151,745,540,600]" box="[151,267,575,600]" pageId="2" pageNumber="623">
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(
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<figureCitation id="A38D2A36FF9EBA2CC3AAFDD8FECA9A7F" box="[159,258,575,600]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1435,1457]" captionStartId-1="4.[151,250,1561,1583]" captionStartId-2="5.[151,250,1130,1152]" captionStartId-3="6.[151,250,1273,1295]" captionTargetBox-0="[151,1435,193,1413]" captionTargetBox-1="[170,1416,268,1539]" captionTargetBox-2="[150,1436,193,1108]" captionTargetBox-3="[151,1436,193,1252]" captionTargetId-0="figure@3.[151,1437,193,1414]" captionTargetId-1="figure@4.[170,1417,265,1539]" captionTargetId-2="figure@5.[150,1436,193,1109]" captionTargetId-3="figure@6.[151,1436,193,1252]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Lateral view; B. Anterior part of body, lateral view; C. Prostomium and chaetigers 1 – 2, ventral view; D. Recurved hooks from chaetiger 1; E. Parapodium from chaetiger 1; F. Parapodium from chaetiger 2. Abbreviations: dc = dorsal cirrus, fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, per = peristomium, psl = postchaetal lobe, ul = upper lip, pl = papilliform lobe, vc = ventral cirrus." captionText-1="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." captionText-2="FIGURE 3. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A – B. Distal bidentate parts of the recurved hooks from chaetiger 1; C. Spiny shaft of recurved hook from chaetiger 1; D. Parapodium from chaetiger 3; E. Pectinate chaeta from chaetiger 25; F. Parapodium from chaetiger 5; G. Subacicular hooks from chaetiger 6; H. The same from chaetiger 14; I. The same from chaetiger 22." captionText-3="FIGURE 4. Juvenile of Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Ventral view; B. Anterior part of body, ventral view; C. Parapodia from chaetigers 3 and 4; D. Parapodium from chaetiger 6; E. Subacicular hooks from chaetiger 9; F. The same from chaetiger 12. Abbreviations: fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, ul = upper lip." httpUri-0="https://zenodo.org/record/233297/files/figure.png" httpUri-1="https://zenodo.org/record/233298/files/figure.png" httpUri-2="https://zenodo.org/record/233299/files/figure.png" httpUri-3="https://zenodo.org/record/233300/files/figure.png" pageId="2" pageNumber="623">Figs 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="73AC6538FF9EBA2CC3A2FD66FAFA9B9D" pageId="2" pageNumber="623" type="materials_examined">
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<paragraph id="3B0936B3FF9EBA2CC3A2FD66FA489B21" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
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<emphasis id="09C2EAA1FF9EBA2CC3A2FD66FE8F9ABD" bold="true" box="[151,327,641,666]" pageId="2" pageNumber="623">
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<typeStatus id="E40D8811FF9EBA2CC3A2FD66FF1A9ABD" box="[151,210,641,666]" pageId="2" pageNumber="623">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="E40D8811FF9EBA2CC27AFD66FE749ABD" box="[335,444,641,666]" pageId="2" pageNumber="623" type="holotype">Holotype</typeStatus>
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: AM W.46046, Queensland, Lizard Island, outer Yonge Reef,
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<geoCoordinate id="5E825074FF9EBA2CC7B0FD66FB289ABD" box="[1157,1248,641,666]" direction="south" orientation="latitude" pageId="2" pageNumber="623" precision="925" value="-14.6">14º36’S</geoCoordinate>
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,
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<geoCoordinate id="5E825074FF9EBA2CC7DBFD66FA919ABD" box="[1262,1369,641,666]" direction="east" orientation="longitude" pageId="2" pageNumber="623" precision="925" value="145.46666">145º28’E</geoCoordinate>
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, dead coral,
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<quantity id="FC4E9B56FF9EBA2CC3EBFD41FED29A9A" box="[222,282,678,702]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="623" unit="m" value="10.0">10 m</quantity>
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, coll. P.A. Hutchings & P.B. Weate,
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<date id="4F081073FF9EBA2CC18DFD42FC8C9A99" box="[696,836,677,702]" pageId="2" pageNumber="623" value="1977-01-25">25 Jan 1977</date>
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, Collection Event Code 77 LIZ 56-2.
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<typeStatus id="E40D8811FF9EBA2CC7C6FD41FAAC9A99" box="[1267,1380,678,702]" pageId="2" pageNumber="623" type="paratype">Paratypes</typeStatus>
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: AM W.47873 (3), same data as
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<typeStatus id="E40D8811FF9EBA2CC2E3FD2EFDF49AC5" box="[470,572,713,738]" pageId="2" pageNumber="623" type="holotype">holotype</typeStatus>
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; AM W.46045 (9), Queensland, Lizard Island, outer Yonge Reef,
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<geoCoordinate id="5E825074FF9EBA2CC609FD2EFA5F9AC5" box="[1340,1431,713,738]" direction="south" orientation="latitude" pageId="2" pageNumber="623" precision="925" value="-14.6">14º36’S</geoCoordinate>
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,
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<geoCoordinate id="5E825074FF9EBA2CC3A2FD0AFECA9B21" box="[151,258,749,774]" direction="east" orientation="longitude" pageId="2" pageNumber="623" precision="925" value="145.46666">145º28’E</geoCoordinate>
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, dead coral,
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<quantity id="FC4E9B56FF9EBA2CC2A5FD09FE759B22" box="[400,445,750,774]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="623" unit="m" value="9.0">9 m</quantity>
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, coll. P.A. Hutchings & P.B. Weate,
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<date id="4F081073FF9EBA2CC060FD0AFC169B21" box="[853,990,749,774]" pageId="2" pageNumber="623" value="1977-01-15">15 Jan 1977</date>
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, Collection Event Code 77 LIZ 48-4.
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</paragraph>
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<paragraph id="3B0936B3FF9EBA2CC3F2FCF6FAFA9B9D" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
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<emphasis id="09C2EAA1FF9EBA2CC3F2FCF6FDCE9B0D" bold="true" box="[199,518,785,810]" pageId="2" pageNumber="623">Other material examined.</emphasis>
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AM W.47874 (35, 1 on slide, 1 on SEM stub), same data as
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<typeStatus id="E40D8811FF9EBA2CC7CCFCF6FA979B0D" box="[1273,1375,785,810]" pageId="2" pageNumber="623" type="holotype">holotype</typeStatus>
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; AM W.47875 (28, 1 on slide, 2 and
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<specimenCount id="2DB0FD3AFF9EBA2CC137FCD1FDBA9B69" box="[514,626,821,846]" pageId="2" pageNumber="623" type="juvenile">1 juvenile</specimenCount>
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on SEM stub), Queensland, Lizard Island, outer Yonge Reef,
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<geoCoordinate id="5E825074FF9EBA2CC609FCD2FA5F9B69" box="[1340,1431,821,846]" direction="south" orientation="latitude" pageId="2" pageNumber="623" precision="925" value="-14.6">14º36’S</geoCoordinate>
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,
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<geoCoordinate id="5E825074FF9EBA2CC3A2FCBEFECA9B55" box="[151,258,857,882]" direction="east" orientation="longitude" pageId="2" pageNumber="623" precision="925" value="145.46666">145º28’E</geoCoordinate>
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, dead coral,
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<quantity id="FC4E9B56FF9EBA2CC2A1FCBDFE0B9B56" box="[404,451,858,882]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="623" unit="m" value="9.0">9 m</quantity>
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, coll. P.A. Hutchings & P.B. Weate,
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<date id="4F081073FF9EBA2CC051FCBEFC3A9B55" box="[868,1010,857,882]" pageId="2" pageNumber="623" value="1977-01-15">15 Jan 1977</date>
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, Collection Event Code 77 LIZ 48-4; AM W.46043(1), Queensland, Yonge Reef, outer slope,
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<geoCoordinate id="5E825074FF9EBA2CC07DFC9AFC6B9BB1" box="[840,931,893,918]" direction="south" orientation="latitude" pageId="2" pageNumber="623" precision="925" value="-14.6">14º36’S</geoCoordinate>
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,
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<geoCoordinate id="5E825074FF9EBA2CC08DFC9AFBEB9BB1" box="[952,1059,893,918]" direction="east" orientation="longitude" pageId="2" pageNumber="623" precision="925" value="145.63333">145º38’E</geoCoordinate>
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, reef rock with
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<taxonomicName id="FCB64D30FF9EBA2CC7C0FC9AFAAB9BB2" box="[1269,1379,893,917]" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="GBIF" kingdom="Plantae" order="Bryopsidales" pageId="2" pageNumber="623" phylum="Chlorophyta" rank="genus">
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||
<emphasis id="09C2EAA1FF9EBA2CC7C0FC9AFAAB9BB2" box="[1269,1379,893,917]" italics="true" pageId="2" pageNumber="623">Halimeda</emphasis>
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||
</taxonomicName>
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||
and
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<emphasis id="09C2EAA1FF9EBA2CC3A2FC46FE8B9B9E" box="[151,323,929,953]" italics="true" pageId="2" pageNumber="623">
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<taxonomicName id="FCB64D30FF9EBA2CC3A2FC46FEF79B9E" box="[151,319,929,953]" class="Florideophyceae" family="Corallinaceae" genus="Lithothamnion" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="2" pageNumber="623" phylum="Rhodophyta" rank="genus">Lithothamnion</taxonomicName>
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,
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</emphasis>
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<quantity id="FC4E9B56FF9EBA2CC27FFC46FE4D9B9E" box="[330,389,929,954]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="623" unit="m" value="36.0">36 m</quantity>
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, coll. P.A. Hutchings & P.B. Weate,
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<date id="4F081073FF9EBA2CC028FC45FC519B9D" box="[797,921,929,954]" pageId="2" pageNumber="623" value="1975-01-09">9 Jan 1975</date>
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, Collection Event Code 75 LIZ D-1.
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</paragraph>
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</subSubSection>
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<subSubSection id="73AC6538FF9EBA2CC3F2FC22FE9B9C49" pageId="2" pageNumber="623" type="diagnosis">
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<paragraph id="3B0936B3FF9EBA2CC3F2FC22FE9B9C49" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
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<emphasis id="09C2EAA1FF9EBA2CC3F2FC22FE899BF9" bold="true" box="[199,321,965,990]" pageId="2" pageNumber="623">Diagnosis.</emphasis>
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Very small, up to
|
||
<quantity id="FC4E9B56FF9EBA2CC126FC22FDA19BFA" box="[531,617,965,990]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="623" unit="mm" value="0.5">0.5 mm</quantity>
|
||
wide. Diagnosis and description based on
|
||
<typeStatus id="E40D8811FF9EBA2CC77AFC22FB799BF9" box="[1103,1201,965,990]" pageId="2" pageNumber="623" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="E40D8811FF9EBA2CC7DCFC21FA9C9BF9" box="[1257,1364,966,990]" pageId="2" pageNumber="623" type="paratype">paratypes</typeStatus>
|
||
wider than
|
||
<quantity id="FC4E9B56FF9EBA2CC3E5FC0DFEEF9C26" box="[208,295,1002,1026]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="623" unit="mm" value="0.2">0.2 mm</quantity>
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at chaetiger 10 excluding parapodia. Bidentate distally recurved spiny hooks on chaetigers 1 and 2; two subacicular hooks with spiny hoods from chaetiger 5, hoods becoming gradually less spiny, resembling typical subacicular hooks by about chaetiger 20; pectinate chaetae with 12–14 moderately long teeth. Pygidium with one pair of anal cirri.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF9EBA2CC3F2FB9EFF229FE9" pageId="2" pageNumber="623" type="description">
|
||
<paragraph id="3B0936B3FF9EBA2CC3F2FB9EFD9A9CFE" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
|
||
<emphasis id="09C2EAA1FF9EBA2CC3F2FB9EFE919CB5" bold="true" box="[199,345,1145,1170]" pageId="2" pageNumber="623">Description.</emphasis>
|
||
<typeStatus id="E40D8811FF9EBA2CC251FB9EFE049CB5" box="[356,460,1145,1170]" pageId="2" pageNumber="623" type="holotype">Holotype</typeStatus>
|
||
incomplete, measuring
|
||
<quantity id="FC4E9B56FF9EBA2CC1DDFB9EFCF49CB6" box="[744,828,1145,1170]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="623" unit="mm" value="16.0">16 mm</quantity>
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||
in length for 55 chaetigers,
|
||
<quantity id="FC4E9B56FF9EBA2CC7A1FB9EFB359CB6" box="[1172,1277,1145,1170]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="623" unit="mm" value="0.55">0.55 mm</quantity>
|
||
wide; largest complete specimens 9.4–11.0 mm long for 40–54 chaetigers,
|
||
<quantity id="FC4E9B56FF9EBA2CC07AFB7AFC1F9C92" box="[847,983,1181,1206]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="623" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
|
||
wide. Alcohol stored specimens overall cream-coloured, lacking colour pattern.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF9EBA2CC3F2FB02FD829DA9" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
|
||
Prostomium anteriorly rounded, wider than long, with paired globular frontal lips (
|
||
<figureCitation id="A38D2A36FF9EBA2CC740FB02FB7D9CD9" box="[1141,1205,1253,1278]" captionStart="FIGURE 1" captionStartId="3.[151,250,1435,1457]" captionTargetBox="[151,1435,193,1413]" captionTargetId="figure@3.[151,1437,193,1414]" captionTargetPageId="3" captionText="FIGURE 1. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Lateral view; B. Anterior part of body, lateral view; C. Prostomium and chaetigers 1 – 2, ventral view; D. Recurved hooks from chaetiger 1; E. Parapodium from chaetiger 1; F. Parapodium from chaetiger 2. Abbreviations: dc = dorsal cirrus, fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, per = peristomium, psl = postchaetal lobe, ul = upper lip, pl = papilliform lobe, vc = ventral cirrus." httpUri="https://zenodo.org/record/233297/files/figure.png" pageId="2" pageNumber="623">Fig. 1</figureCitation>
|
||
A–C). Ceratophores of palps and antennae with one proximal ring and one longer distal ring; ceratostyles short and subulate, palps reaching to peristomium, lateral and median antennae reaching to chaetiger 1. Nuchal grooves curved laterally, with small middorsal separation. One pair of small eyespots at bases of lateral antennae. Peristomium shorter than first chaetiger; peristomial cirri absent.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF9EBA2CC3F2FA7EFA4B9E39" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
|
||
First two pairs of parapodia modified, projecting anterolaterally, directed ventrally (
|
||
<figureCitation id="A38D2A36FF9EBA2CC7F4FA7EFAC19D95" box="[1217,1289,1433,1458]" captionStart="FIGURE 1" captionStartId="3.[151,250,1435,1457]" captionTargetBox="[151,1435,193,1413]" captionTargetId="figure@3.[151,1437,193,1414]" captionTargetPageId="3" captionText="FIGURE 1. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Lateral view; B. Anterior part of body, lateral view; C. Prostomium and chaetigers 1 – 2, ventral view; D. Recurved hooks from chaetiger 1; E. Parapodium from chaetiger 1; F. Parapodium from chaetiger 2. Abbreviations: dc = dorsal cirrus, fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, per = peristomium, psl = postchaetal lobe, ul = upper lip, pl = papilliform lobe, vc = ventral cirrus." httpUri="https://zenodo.org/record/233297/files/figure.png" pageId="2" pageNumber="623">Fig. 1</figureCitation>
|
||
B). Each of parapodia 1 and 2 with three short papilliform lobes and one ovate postchaetal lobe (
|
||
<figureCitation id="A38D2A36FF9EBA2CC7B4FA5AFB0C9DF1" box="[1153,1220,1469,1494]" captionStart="FIGURE 1" captionStartId="3.[151,250,1435,1457]" captionTargetBox="[151,1435,193,1413]" captionTargetId="figure@3.[151,1437,193,1414]" captionTargetPageId="3" captionText="FIGURE 1. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Lateral view; B. Anterior part of body, lateral view; C. Prostomium and chaetigers 1 – 2, ventral view; D. Recurved hooks from chaetiger 1; E. Parapodium from chaetiger 1; F. Parapodium from chaetiger 2. Abbreviations: dc = dorsal cirrus, fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, per = peristomium, psl = postchaetal lobe, ul = upper lip, pl = papilliform lobe, vc = ventral cirrus." httpUri="https://zenodo.org/record/233297/files/figure.png" pageId="2" pageNumber="623">Fig. 1</figureCitation>
|
||
E, F). Unmodified parapodia lacking distinct parapodial lobes. Dorsal cirri subulate on chaetiger 1 and 2, then becoming digitate. Ventral cirri subulate on chaetiger 1 and 2, replaced by rounded ventral pads from chaetiger 3. Branchiae absent.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF9EBA2CC3F2F9CEFB469E89" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
|
||
Modified parapodia with three long, pseudocompound recurved hooks (
|
||
<figureCitation id="A38D2A36FF9EBA2CC0C0F9CEFBF29E65" box="[1013,1082,1577,1602]" captionStart="FIGURE 1" captionStartId="3.[151,250,1435,1457]" captionTargetBox="[151,1435,193,1413]" captionTargetId="figure@3.[151,1437,193,1414]" captionTargetPageId="3" captionText="FIGURE 1. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Lateral view; B. Anterior part of body, lateral view; C. Prostomium and chaetigers 1 – 2, ventral view; D. Recurved hooks from chaetiger 1; E. Parapodium from chaetiger 1; F. Parapodium from chaetiger 2. Abbreviations: dc = dorsal cirrus, fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, per = peristomium, psl = postchaetal lobe, ul = upper lip, pl = papilliform lobe, vc = ventral cirrus." httpUri="https://zenodo.org/record/233297/files/figure.png" pageId="2" pageNumber="623">Figs 1</figureCitation>
|
||
D, 2A), each projecting from a papilliform lobe (
|
||
<figureCitation id="A38D2A36FF9EBA2CC26FF9AAFE519E41" box="[346,409,1613,1638]" captionStart="FIGURE 1" captionStartId="3.[151,250,1435,1457]" captionTargetBox="[151,1435,193,1413]" captionTargetId="figure@3.[151,1437,193,1414]" captionTargetPageId="3" captionText="FIGURE 1. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Lateral view; B. Anterior part of body, lateral view; C. Prostomium and chaetigers 1 – 2, ventral view; D. Recurved hooks from chaetiger 1; E. Parapodium from chaetiger 1; F. Parapodium from chaetiger 2. Abbreviations: dc = dorsal cirrus, fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, per = peristomium, psl = postchaetal lobe, ul = upper lip, pl = papilliform lobe, vc = ventral cirrus." httpUri="https://zenodo.org/record/233297/files/figure.png" pageId="2" pageNumber="623">Fig. 1</figureCitation>
|
||
C, E). Shafts of hooks with two rows of long moveable spines below pseudoarticulation and irregularly distributed small spines; slight swelling before distal curvature of hook and small secondary hook, resulting in bifid recurved hook (
|
||
<figureCitation id="A38D2A36FF9EBA2CC13DF972FD879E89" box="[520,591,1685,1710]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="2" pageNumber="623">Figs 2</figureCitation>
|
||
A, 3A–C). Internal chaetal sacs to chaetiger 20–22.
|
||
</paragraph>
|
||
<paragraph id="3B0936B3FF9EBA2CC3F2F95EFF229FE9" blockId="2.[151,1437,641,1998]" pageId="2" pageNumber="623">
|
||
Unmodified parapodia from chaetiger 3, supported by two to three very thick aciculae (
|
||
<figureCitation id="A38D2A36FF9EBA2CC79AF95EFB399EF5" box="[1199,1265,1721,1746]" captionStart="FIGURE 3" captionStartId="5.[151,250,1130,1152]" captionTargetBox="[150,1436,193,1108]" captionTargetId="figure@5.[150,1436,193,1109]" captionTargetPageId="5" captionText="FIGURE 3. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A – B. Distal bidentate parts of the recurved hooks from chaetiger 1; C. Spiny shaft of recurved hook from chaetiger 1; D. Parapodium from chaetiger 3; E. Pectinate chaeta from chaetiger 25; F. Parapodium from chaetiger 5; G. Subacicular hooks from chaetiger 6; H. The same from chaetiger 14; I. The same from chaetiger 22." httpUri="https://zenodo.org/record/233299/files/figure.png" pageId="2" pageNumber="623">Fig. 3</figureCitation>
|
||
D, F). Limbate chaetae from chaetiger 3, decreasing in length from upper to lower position (
|
||
<figureCitation id="A38D2A36FF9EBA2CC713F93AFBA49ED1" box="[1062,1132,1757,1782]" captionStart="FIGURE 3" captionStartId="5.[151,250,1130,1152]" captionTargetBox="[150,1436,193,1108]" captionTargetId="figure@5.[150,1436,193,1109]" captionTargetPageId="5" captionText="FIGURE 3. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A – B. Distal bidentate parts of the recurved hooks from chaetiger 1; C. Spiny shaft of recurved hook from chaetiger 1; D. Parapodium from chaetiger 3; E. Pectinate chaeta from chaetiger 25; F. Parapodium from chaetiger 5; G. Subacicular hooks from chaetiger 6; H. The same from chaetiger 14; I. The same from chaetiger 22." httpUri="https://zenodo.org/record/233299/files/figure.png" pageId="2" pageNumber="623">Fig. 3</figureCitation>
|
||
D). Spinigers absent. Two bidentate subacicular hooks with spiny hoods from chaetiger 5 (
|
||
<figureCitation id="A38D2A36FF9EBA2CC0B1F8E6FC079F3D" box="[900,975,1793,1818]" captionStart="FIGURE 2" captionStartId="4.[151,250,1561,1583]" captionTargetBox="[170,1416,268,1539]" captionTargetId="figure@4.[170,1417,265,1539]" captionTargetPageId="4" captionText="FIGURE 2. Rhamphobrachium (Minibrachium) nutrix n. sp. (A, B), R. (M.) fractum n. sp., (C – G) and R. (M.) talboti n. sp., (H – I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown)." httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="2" pageNumber="623">Figs 2</figureCitation>
|
||
B, 3F); hoods initially almost enclosing distal part of hooks, gradually opening up, becoming smoother and changing to divided hoods, one on either side of distal part of hook by about chaetiger 20, resembling typical subacicular hooks (
|
||
<figureCitation id="A38D2A36FF9EBA2CC761F8AEFB539F45" box="[1108,1179,1865,1890]" captionStart="FIGURE 3" captionStartId="5.[151,250,1130,1152]" captionTargetBox="[150,1436,193,1108]" captionTargetId="figure@5.[150,1436,193,1109]" captionTargetPageId="5" captionText="FIGURE 3. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A – B. Distal bidentate parts of the recurved hooks from chaetiger 1; C. Spiny shaft of recurved hook from chaetiger 1; D. Parapodium from chaetiger 3; E. Pectinate chaeta from chaetiger 25; F. Parapodium from chaetiger 5; G. Subacicular hooks from chaetiger 6; H. The same from chaetiger 14; I. The same from chaetiger 22." httpUri="https://zenodo.org/record/233299/files/figure.png" pageId="2" pageNumber="623">Fig. 3</figureCitation>
|
||
G–I). Slightly oblique pectinate chaetae with 12–14 moderately long teeth from chaetiger 12 (
|
||
<figureCitation id="A38D2A36FF9EBA2CC0FDF88AFBC59FA1" box="[968,1037,1901,1926]" captionStart="FIGURE 3" captionStartId="5.[151,250,1130,1152]" captionTargetBox="[150,1436,193,1108]" captionTargetId="figure@5.[150,1436,193,1109]" captionTargetPageId="5" captionText="FIGURE 3. Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A – B. Distal bidentate parts of the recurved hooks from chaetiger 1; C. Spiny shaft of recurved hook from chaetiger 1; D. Parapodium from chaetiger 3; E. Pectinate chaeta from chaetiger 25; F. Parapodium from chaetiger 5; G. Subacicular hooks from chaetiger 6; H. The same from chaetiger 14; I. The same from chaetiger 22." httpUri="https://zenodo.org/record/233299/files/figure.png" pageId="2" pageNumber="623">Fig. 3</figureCitation>
|
||
E). Pygidium with one pair of anal cirri. Fragile mucous tube without incrustations. Mandibles and maxillae extremely small and fragile, not clearly visible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6FC9663BFF9FBA2DC3A2FA7CFAF09E2A" httpUri="https://zenodo.org/record/233297/files/figure.png" pageId="3" pageNumber="624" targetBox="[151,1435,193,1413]" targetPageId="3">
|
||
<paragraph id="3B0936B3FF9FBA2DC3A2FA7CFAF09E2A" blockId="3.[151,1436,1435,1549]" pageId="3" pageNumber="624">
|
||
<emphasis id="09C2EAA1FF9FBA2DC3A2FA7CFEDB9D96" bold="true" box="[151,275,1435,1457]" pageId="3" pageNumber="624">FIGURE 1.</emphasis>
|
||
<taxonomicName id="FCB64D30FF9FBA2DC22FFA7CFD009D96" box="[282,712,1435,1457]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="624" phylum="Annelida" rank="species" species="nutrix" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF9FBA2DC22FFA7CFE149D96" box="[282,476,1435,1457]" italics="true" pageId="3" pageNumber="624">Rhamphobrachium</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF9FBA2DC2D9FA7CFDB59D96" box="[492,637,1435,1457]" italics="true" pageId="3" pageNumber="624">Minibrachium</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF9FBA2DC1B9FA7BFD009D96" box="[652,712,1436,1457]" italics="true" pageId="3" pageNumber="624">nutrix</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF9FBA2DC1FAFA7BFCC09D96" box="[719,776,1436,1457]" pageId="3" pageNumber="624" rank="species">n. sp.</taxonomicNameLabel>
|
||
, AM W.47875. A. Lateral view; B. Anterior part of body, lateral view; C. Prostomium and chaetigers 1–2,ventral view; D. Recurved hooks from chaetiger 1; E. Parapodium from chaetiger 1; F. Parapodium from chaetiger 2. Abbreviations: dc = dorsal cirrus, fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, per = peristomium, psl = postchaetal lobe, ul = upper lip, pl = papilliform lobe, vc = ventral cirrus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="73AC6538FF9FBA2DC3F2F9B9FB7C9F20" pageId="3" pageNumber="624" type="discussion">
|
||
<paragraph id="3B0936B3FF9FBA2DC3F2F9B9FB7C9F20" blockId="3.[151,1436,1630,2015]" pageId="3" pageNumber="624">
|
||
<emphasis id="09C2EAA1FF9FBA2DC3F2F9B9FEF39E50" bold="true" box="[199,315,1630,1655]" pageId="3" pageNumber="624">Remarks.</emphasis>
|
||
<taxonomicName id="FCB64D30FF9FBA2DC277F9B9FD539E51" box="[322,667,1630,1654]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="624" phylum="Annelida" rank="species" species="nutrix" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF9FBA2DC277F9B9FD539E51" box="[322,667,1630,1654]" italics="true" pageId="3" pageNumber="624">Rhamphobrachium (M.) nutrix</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF9FBA2DC197F9B8FD169E51" box="[674,734,1631,1654]" pageId="3" pageNumber="624" rank="species">n. sp.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="FCB64D30FF9FBA2DC046F9B8FBE99E51" box="[883,1057,1630,1655]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="624" phylum="Annelida" rank="species" species="fractum" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF9FBA2DC046F9B8FC439E51" box="[883,907,1631,1654]" italics="true" pageId="3" pageNumber="624">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF9FBA2DC0AEF9B8FC729E51" box="[923,954,1631,1654]" italics="true" pageId="3" pageNumber="624">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF9FBA2DC0FFF9B9FBE99E51" box="[970,1057,1630,1654]" italics="true" pageId="3" pageNumber="624">fractum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF9FBA2DC71DF9B8FBAC9E51" box="[1064,1124,1631,1654]" pageId="3" pageNumber="624" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="FCB64D30FF9FBA2DC7AEF9B8FAF49E51" box="[1179,1340,1630,1655]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="624" phylum="Annelida" rank="species" species="talboti" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF9FBA2DC7AEF9B8FB7B9E51" box="[1179,1203,1631,1654]" italics="true" pageId="3" pageNumber="624">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF9FBA2DC7F1F9B8FB2A9E51" box="[1220,1250,1631,1654]" italics="true" pageId="3" pageNumber="624">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF9FBA2DC7C6F9B9FAF49E51" box="[1267,1340,1630,1654]" italics="true" pageId="3" pageNumber="624">talboti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF9FBA2DC676F9B8FAB79E51" box="[1347,1407,1631,1654]" pageId="3" pageNumber="624" rank="species">n. sp.</taxonomicNameLabel>
|
||
in being much smaller, having bidentate instead of unidentate distally recurved hooks, having two subacicular hooks per parapodium from chaetiger 5 instead of multiple hooks from chaetiger 3, and having one pair of anal cirri instead of two. Furthermore,
|
||
<taxonomicName id="FCB64D30FF9FBA2DC2EBF92CFDB09EC5" box="[478,632,1738,1762]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="624" phylum="Annelida" rank="species" species="nutrix" subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF9FBA2DC2EBF92CFE3E9EC5" box="[478,502,1739,1762]" italics="true" pageId="3" pageNumber="624">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF9FBA2DC133F92CFDED9EC5" box="[518,549,1739,1762]" italics="true" pageId="3" pageNumber="624">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF9FBA2DC100F92CFDB09EC5" box="[565,632,1739,1762]" italics="true" pageId="3" pageNumber="624">nutrix</emphasis>
|
||
</taxonomicName>
|
||
has pectinate chaetae with 12–14 moderately long teeth while the other two new species have pectinate chaetae with 9–13 very long and 8–10 long teeth respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF9FBA2DC3F2F8F5FE4C9F68" pageId="3" pageNumber="624" type="etymology">
|
||
<paragraph id="3B0936B3FF9FBA2DC3F2F8F5FE4C9F68" blockId="3.[151,1436,1630,2015]" pageId="3" pageNumber="624">
|
||
<emphasis id="09C2EAA1FF9FBA2DC3F2F8F5FE8E9F0C" bold="true" box="[199,326,1810,1835]" pageId="3" pageNumber="624">Etymology</emphasis>
|
||
. The species name
|
||
<taxonomicName id="FCB64D30FF9FBA2DC111F8F4FDAF9F0D" box="[548,615,1811,1834]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="3" pageNumber="624" phylum="Annelida" rank="species" species="nutrix">
|
||
<emphasis id="09C2EAA1FF9FBA2DC111F8F4FDAF9F0D" box="[548,615,1811,1834]" italics="true" pageId="3" pageNumber="624">nutrix</emphasis>
|
||
</taxonomicName>
|
||
(nurse) in Latin refers to the ability of the individuals of the new species to brood their young.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF9FBA2AC3F2F8BDFCE69FE0" lastPageId="4" lastPageNumber="625" pageId="3" pageNumber="624" type="biology_ecology">
|
||
<paragraph id="3B0936B3FF9FBA2AC3F2F8BDFBDF98F3" blockId="3.[151,1436,1630,2015]" lastBlockId="4.[151,1437,151,213]" lastPageId="4" lastPageNumber="625" pageId="3" pageNumber="624">
|
||
<emphasis id="09C2EAA1FF9FBA2DC3F2F8BDFEEF9F54" bold="true" box="[199,295,1882,1907]" pageId="3" pageNumber="624">Biology.</emphasis>
|
||
Despite their small size the specimens of the new species brood their young and thus are considered fully grown individuals. The
|
||
<typeStatus id="E40D8811FF9FBA2DC2D7F899FD8C9FB0" box="[482,580,1918,1943]" pageId="3" pageNumber="624" type="holotype">holotype</typeStatus>
|
||
and several
|
||
<typeStatus id="E40D8811FF9FBA2DC1E1F898FCF79FB0" box="[724,831,1919,1943]" pageId="3" pageNumber="624" type="paratype">paratypes</typeStatus>
|
||
have two to four juveniles inside their tubes. Several similarly sized juveniles are present in the examined samples; presumably they have detached from their parents because of the destruction of the very fragile tubes. None of the specimens has an intact tube and the number of juveniles per parent could not be established with certainty; however, judging from the size of the juveniles, only few of them (possibly not more than 10) could fit with a single brooding parent.
|
||
</paragraph>
|
||
<caption id="6FC9663BFF98BA2AC3A2F9FEFE3E9E8E" httpUri="https://zenodo.org/record/233298/files/figure.png" pageId="4" pageNumber="625" targetBox="[170,1416,268,1539]" targetPageId="4">
|
||
<paragraph id="3B0936B3FF98BA2AC3A2F9FEFE3E9E8E" blockId="4.[151,1436,1561,1705]" pageId="4" pageNumber="625">
|
||
<emphasis id="09C2EAA1FF98BA2AC3A2F9FEFEDC9E08" bold="true" box="[151,276,1561,1583]" pageId="4" pageNumber="625">FIGURE 2.</emphasis>
|
||
<taxonomicName id="FCB64D30FF98BA2AC229F9FEFD029E08" box="[284,714,1561,1583]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="4" pageNumber="625" phylum="Annelida" rank="species" species="nutrix" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF98BA2AC229F9FEFE169E08" box="[284,478,1561,1583]" italics="true" pageId="4" pageNumber="625">Rhamphobrachium</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF98BA2AC2D8F9FEFDB69E08" box="[493,638,1561,1583]" italics="true" pageId="4" pageNumber="625">Minibrachium</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF98BA2AC1BBF9FDFD029E08" box="[654,714,1562,1583]" italics="true" pageId="4" pageNumber="625">nutrix</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF98BA2AC1E6F9FDFCC29E08" box="[723,778,1562,1583]" pageId="4" pageNumber="625" rank="species">n. sp.</taxonomicNameLabel>
|
||
(A, B),
|
||
<taxonomicName id="FCB64D30FF98BA2AC057F9FDFBCA9E08" box="[866,1026,1561,1583]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="4" pageNumber="625" phylum="Annelida" rank="species" species="fractum" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF98BA2AC057F9FDFCB09E08" box="[866,888,1562,1583]" italics="true" pageId="4" pageNumber="625">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF98BA2AC0B2F9FDFC549E08" box="[903,924,1562,1583]" italics="true" pageId="4" pageNumber="625">M</emphasis>
|
||
.)
|
||
<emphasis id="09C2EAA1FF98BA2AC086F9FEFBCA9E08" box="[947,1026,1561,1583]" italics="true" pageId="4" pageNumber="625">fractum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF98BA2AC73CF9FDFB8B9E08" box="[1033,1091,1562,1583]" pageId="4" pageNumber="625" rank="species">n. sp.</taxonomicNameLabel>
|
||
, (C–G) and
|
||
<taxonomicName id="FCB64D30FF98BA2AC7F6F9FDFA9E9E08" box="[1219,1366,1561,1583]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="4" pageNumber="625" phylum="Annelida" rank="species" species="talboti" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF98BA2AC7F6F9FDFB119E08" box="[1219,1241,1562,1583]" italics="true" pageId="4" pageNumber="625">R.</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF98BA2AC7DCF9FDFACD9E08" box="[1257,1285,1562,1583]" italics="true" pageId="4" pageNumber="625">M.</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF98BA2AC621F9FEFA9E9E08" box="[1300,1366,1561,1583]" italics="true" pageId="4" pageNumber="625">talboti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF98BA2AC66BF9FDFA5F9E08" box="[1374,1431,1562,1583]" pageId="4" pageNumber="625" rank="species">n. sp.</taxonomicNameLabel>
|
||
, (H–I). A. Distal part of recurved hook from chaetiger 1; B. Subacicular hook from chaetiger 20; C. Anterior part of body; D. Distal part of recurved hook from chaetiger 1; E. Compound subacicular hook from chaetiger 3; F. Pseudocompound subacicular hook from chaetiger 3; G. Simple subacicular hook from chaetiger 4; H. Mandibles; I. Part of maxillary apparatus (MxI, MxII and MxIII are shown).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3B0936B3FF98BA2AC3F2F91DFCE69FE0" blockId="4.[151,1437,1786,2027]" pageId="4" pageNumber="625">
|
||
Examined juveniles have about 16–20 chaetigers and are about
|
||
<quantity id="FC4E9B56FF98BA2AC0BFF91DFC039F35" box="[906,971,1786,1810]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="625" unit="mm" value="1.0">1 mm</quantity>
|
||
long,
|
||
<quantity id="FC4E9B56FF98BA2AC725F91DFBAE9F35" box="[1040,1126,1786,1810]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="625" unit="mm" value="0.2">0.2 mm</quantity>
|
||
wide, with developed palps and antennae but lacking fully formed frontal, upper and lower lips (
|
||
<figureCitation id="A38D2A36FF98BA2AC0E4F8F9FBD29F10" box="[977,1050,1822,1847]" captionStart="FIGURE 4" captionStartId="6.[151,250,1273,1295]" captionTargetBox="[151,1436,193,1252]" captionTargetId="figure@6.[151,1436,193,1252]" captionTargetPageId="6" captionText="FIGURE 4. Juvenile of Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Ventral view; B. Anterior part of body, ventral view; C. Parapodia from chaetigers 3 and 4; D. Parapodium from chaetiger 6; E. Subacicular hooks from chaetiger 9; F. The same from chaetiger 12. Abbreviations: fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, ul = upper lip." httpUri="https://zenodo.org/record/233300/files/figure.png" pageId="4" pageNumber="625">Fig. 4</figureCitation>
|
||
A, B). The anterior two pairs of parapodia have subulate dorsal and ventral cirri but are not enlarged and lack chaetal lobes and recurved hooks. Limbate chaetae are present from chaetiger 3 (
|
||
<figureCitation id="A38D2A36FF98BA2AC1AAF881FD2B9F58" box="[671,739,1894,1919]" captionStart="FIGURE 4" captionStartId="6.[151,250,1273,1295]" captionTargetBox="[151,1436,193,1252]" captionTargetId="figure@6.[151,1436,193,1252]" captionTargetPageId="6" captionText="FIGURE 4. Juvenile of Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Ventral view; B. Anterior part of body, ventral view; C. Parapodia from chaetigers 3 and 4; D. Parapodium from chaetiger 6; E. Subacicular hooks from chaetiger 9; F. The same from chaetiger 12. Abbreviations: fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, ul = upper lip." httpUri="https://zenodo.org/record/233300/files/figure.png" pageId="4" pageNumber="625">Fig. 4</figureCitation>
|
||
C); pectinate chaetae are absent. Subacicular hooks with spiny hoods number up to three per parapodium from chaetiger 3 (
|
||
<figureCitation id="A38D2A36FF98BA2AC04EF86DFC0D9F85" box="[891,965,1930,1955]" captionStart="FIGURE 4" captionStartId="6.[151,250,1273,1295]" captionTargetBox="[151,1436,193,1252]" captionTargetId="figure@6.[151,1436,193,1252]" captionTargetPageId="6" captionText="FIGURE 4. Juvenile of Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Ventral view; B. Anterior part of body, ventral view; C. Parapodia from chaetigers 3 and 4; D. Parapodium from chaetiger 6; E. Subacicular hooks from chaetiger 9; F. The same from chaetiger 12. Abbreviations: fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, ul = upper lip." httpUri="https://zenodo.org/record/233300/files/figure.png" pageId="4" pageNumber="625">Fig. 4</figureCitation>
|
||
D, E). In posterior chaetigers hoods on subacicular hooks become smaller and less spiny (
|
||
<figureCitation id="A38D2A36FF98BA2AC1FEF849FCD89FE0" box="[715,784,1966,1991]" captionStart="FIGURE 4" captionStartId="6.[151,250,1273,1295]" captionTargetBox="[151,1436,193,1252]" captionTargetId="figure@6.[151,1436,193,1252]" captionTargetPageId="6" captionText="FIGURE 4. Juvenile of Rhamphobrachium (Minibrachium) nutrix n. sp., AM W. 47875. A. Ventral view; B. Anterior part of body, ventral view; C. Parapodia from chaetigers 3 and 4; D. Parapodium from chaetiger 6; E. Subacicular hooks from chaetiger 9; F. The same from chaetiger 12. Abbreviations: fl = frontal lip, la = lateral antenna, ll = lower lip, ma = median antenna, p = palp, ul = upper lip." httpUri="https://zenodo.org/record/233300/files/figure.png" pageId="4" pageNumber="625">Fig. 4</figureCitation>
|
||
F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AC6538FF98BA2AC3F2F835FB349FCC" box="[199,1276,2002,2027]" pageId="4" pageNumber="625" type="materials_examined">
|
||
<paragraph id="3B0936B3FF98BA2AC3F2F835FB349FCC" blockId="4.[151,1437,1786,2027]" box="[199,1276,2002,2027]" pageId="4" pageNumber="625">
|
||
<emphasis id="09C2EAA1FF98BA2AC3F2F835FEAD9FCC" bold="true" box="[199,357,2002,2027]" pageId="4" pageNumber="625">
|
||
<typeStatus id="E40D8811FF98BA2AC3F2F835FECA9FCC" box="[199,258,2002,2027]" pageId="4" pageNumber="625">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Pacific Ocean, Outer Yonge Reef, Lizard Island, Queensland;
|
||
<geoCoordinate id="5E825074FF98BA2AC711F835FBB79FCC" box="[1060,1151,2002,2027]" direction="south" orientation="latitude" pageId="4" pageNumber="625" precision="925" value="-14.6">14º36’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5E825074FF98BA2AC7B9F835FB309FCC" box="[1164,1272,2002,2027]" direction="east" orientation="longitude" pageId="4" pageNumber="625" precision="925" value="145.46666">145º28’E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="6FC9663BFF99BA2BC3A2FB8DFE449CFB" httpUri="https://zenodo.org/record/233299/files/figure.png" pageId="5" pageNumber="626" targetBox="[150,1436,193,1108]" targetPageId="5">
|
||
<paragraph id="3B0936B3FF99BA2BC3A2FB8DFE449CFB" blockId="5.[151,1436,1130,1244]" pageId="5" pageNumber="626">
|
||
<emphasis id="09C2EAA1FF99BA2BC3A2FB8DFEDA9CA7" bold="true" box="[151,274,1130,1152]" pageId="5" pageNumber="626">FIGURE 3.</emphasis>
|
||
<taxonomicName id="FCB64D30FF99BA2BC22CFB8DFD0D9CA7" box="[281,709,1130,1152]" class="Polychaeta" family="Onuphidae" genus="Rhamphobrachium" kingdom="Animalia" order="Eunicida" pageId="5" pageNumber="626" phylum="Annelida" rank="species" species="nutrix" status="sp. nov." subGenus="Minibrachium">
|
||
<emphasis id="09C2EAA1FF99BA2BC22CFB8DFE139CA7" box="[281,475,1130,1152]" italics="true" pageId="5" pageNumber="626">Rhamphobrachium</emphasis>
|
||
(
|
||
<emphasis id="09C2EAA1FF99BA2BC2DFFB8DFDB39CA7" box="[490,635,1130,1152]" italics="true" pageId="5" pageNumber="626">Minibrachium</emphasis>
|
||
)
|
||
<emphasis id="09C2EAA1FF99BA2BC1BCFB8CFD0D9CA7" box="[649,709,1131,1152]" italics="true" pageId="5" pageNumber="626">nutrix</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="12F157DAFF99BA2BC1F9FB8CFCCC9CA7" box="[716,772,1131,1152]" pageId="5" pageNumber="626" rank="species">n. sp.</taxonomicNameLabel>
|
||
, AM W.47875. A–B. Distal bidentate parts of the recurved hooks from chaetiger 1; C. Spiny shaft of recurved hook from chaetiger 1; D. Parapodium from chaetiger 3; E. Pectinate chaeta from chaetiger 25; F. Parapodium from chaetiger 5; G. Subacicular hooks from chaetiger 6; H. The same from chaetiger 14; I. The same from chaetiger 22.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="73AC6538FF99BA2BC3F2FACBFED59D4E" pageId="5" pageNumber="626" type="distribution">
|
||
<paragraph id="3B0936B3FF99BA2BC3F2FACBFED59D4E" blockId="5.[151,1436,1324,1385]" pageId="5" pageNumber="626">
|
||
<emphasis id="09C2EAA1FF99BA2BC3F2FACBFEA89D62" bold="true" box="[199,352,1324,1349]" pageId="5" pageNumber="626">Distribution.</emphasis>
|
||
This species is only known from Lizard Island, Great Barrier Reef, Queensland,
|
||
<collectingCountry id="43A17623FF99BA2BC7DBFACBFA909D62" box="[1262,1368,1324,1349]" name="Australia" pageId="5" pageNumber="626">Australia</collectingCountry>
|
||
; in
|
||
<quantity id="FC4E9B56FF99BA2BC6B5FACAFF189D4F" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="3.6" metricValueMin="0.9" pageId="5" pageNumber="626" unit="m" value="22.5" valueMax="36.0" valueMin="9.0">9– 36 m</quantity>
|
||
depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |