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<mods:title id="7B91A48D7E89A4F996BA98CE1CA2D206">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<heading id="720281F7FFD86D69D6DDDCB4BA7803DE" bold="true" box="[151,1151,1410,1437]" fontSize="11" level="1" pageId="49" pageNumber="300" reason="1">
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Key to species of the genus 
<taxonomicName id="EEF54D18FFD86D69D7A6DCB5BCB603DE" ID-CoL="8W25K" authorityName="Steel" authorityYear="1960" box="[492,689,1410,1437]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFD86D69D7A6DCB5BCB603DE" bold="true" box="[492,689,1410,1437]" italics="true" pageId="49" pageNumber="300">Paraphloeostiba</emphasis>
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of New 
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and 
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<paragraph id="294A369BFFD86D69D6DDDCFFBB9B039E" blockId="49.[151,1437,1480,2030]" box="[151,1436,1480,1502]" pageId="49" pageNumber="300">1 Middle portion of pronotum with two distinct longitudinal impressions.......................................... 2</paragraph>
<paragraph id="294A369BFFD86D69D6DDDCD2BB9B03B9" blockId="49.[151,1437,1480,2030]" box="[151,1436,1509,1531]" pageId="49" pageNumber="300">- Middle portion of pronotum without or with indistinct longitudinal impressions................................... 8</paragraph>
<paragraph id="294A369BFFD86D69D6DDDF35BB9B0054" blockId="49.[151,1437,1480,2030]" box="[151,1436,1538,1560]" pageId="49" pageNumber="300">2 Body more than 2.00 mm.............................................................................. 4</paragraph>
<paragraph id="294A369BFFD86D69D6DDDF17BB9B0076" blockId="49.[151,1437,1480,2030]" box="[151,1436,1568,1590]" pageId="49" pageNumber="300">- Body distinctly less than 2.00 mm....................................................................... 7</paragraph>
<paragraph id="294A369BFFD86D69D6DDDF0ABB9B00CE" blockId="49.[151,1437,1480,2030]" pageId="49" pageNumber="300">
4 Antennomeres 8–10 slightly transverse. Elytra slightly more than twice as long as pronotum. Median lobe wide, gradually narrowed toward apex; parameres significantly longer than apex of median lobe, with wide apical portions (
<figureCitation id="B1CE2A1EFFD86D69D34BDF6DBB51002C" box="[1281,1366,1626,1648]" captionStart="FIGURES 148–151" captionStartId="48.[151,264,1862,1886]" captionTargetBox="[168,1427,193,1830]" captionTargetId="figure-22@48.[151,1436,181,1838]" captionTargetPageId="48" captionText="FIGURES 148–151. Paraphloeostiba vitiosa (holotype): 148–aedeagus, parameral view, 149–aedeagus, lateral view, 150–male abdominal tergite VIII, 151–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821293" httpUri="https://zenodo.org/record/10821293/files/figure.png" pageId="49" pageNumber="300">Fig. 148</figureCitation>
). Body yellow-brown. Body length: 
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. 
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(Western New 
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).............................. 
<taxonomicName id="EEF54D18FFD86D69D343DF4FBB6700CE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1289,1376,1656,1677]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="vitiosa" status="sp. n.">
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<paragraph id="294A369BFFD86D69D6DDDFA2BB9B008B" blockId="49.[151,1437,1480,2030]" pageId="49" pageNumber="300">- Antennomeres 8–10 distinctly transverse. Elytra less than twice as long as pronotum. Median lobe different, parameres shorter............................................................................................. 5</paragraph>
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5 Head and pronotum without punctation. Aedeagus as in 
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Fig. 
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in 
<bibRefCitation id="4D644B6AFFD86D69D529DFE7BA0400A5" author="Shavrin, A. V." box="[867,1027,1744,1766]" pageId="49" pageNumber="300" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Body brown. Habitus as in 
<figureCitation id="B1CE2A1EFFD86D69D35CDFE7BB5D00A6" box="[1302,1370,1744,1766]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="49" pageNumber="300">Fig. 39</figureCitation>
and in 
<figureCitation id="B1CE2A1EFFD86D69D6A8DFDABF1D0141" box="[226,282,1773,1795]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="49" pageNumber="300">
Fig. 
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in 
<bibRefCitation id="4D644B6AFFD86D69D773DFDABFDF0140" author="Shavrin, A. V." box="[313,472,1773,1795]" pageId="49" pageNumber="300" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFD86D69D545DFDABD510140" box="[783,854,1773,1795]" captionStart="FIGURES 53–56" captionStartId="21.[151,264,1187,1211]" captionTargetBox="[159,1428,187,1164]" captionTargetId="figure-16@21.[151,1436,181,1164]" captionTargetPageId="21" captionText="FIGURES 53–56. Paraphloeostiba coriacea: 53–aedeagus, parameral view (Western New Guinea), 54–female abdominal tergite VIII, 55–female abdominal sternite VIII, 56–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821243" httpUri="https://zenodo.org/record/10821243/files/figure.png" pageId="49" pageNumber="300">Fig. 56</figureCitation>
. Body length: 
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<taxonomicName id="EEF54D18FFD86D69D2CEDFD9BB9B0140" authority="(Cameron, 1952)" baseAuthorityName="Cameron" baseAuthorityYear="1952" box="[1156,1436,1773,1795]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="coriacea">
<emphasis id="1B81EA89FFD86D69D2CEDFD9BA940140" box="[1156,1171,1774,1795]" italics="true" pageId="49" pageNumber="300">P</emphasis>
. 
<emphasis id="1B81EA89FFD86D69D2D4DFD9BAF50140" box="[1182,1266,1774,1795]" italics="true" pageId="49" pageNumber="300">coriacea</emphasis>
(
<bibRefCitation id="4D644B6AFFD86D69D2B5DFDABB920140" author="Cameron, M." box="[1279,1429,1773,1795]" pageId="49" pageNumber="300" pagination="241 - 256" refId="ref35440" refString="Cameron, M. (1952) Results of the Archibold Expeditions. New species of Staphylinidae (Col.) from New Guinea and Misool. Treubia, 21 (2), 241 - 256." type="journal article" year="1952">Cameron, 1952</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFD86D69D6DDDE3CBB9B0163" blockId="49.[151,1437,1480,2030]" box="[151,1436,1802,1824]" pageId="49" pageNumber="300">- Head and pronotum with distinct punctation................................................................ 6</paragraph>
<paragraph id="294A369BFFD86D69D6DDDE1FBB9B01D6" blockId="49.[151,1437,1480,2030]" pageId="49" pageNumber="300">
6 Pronotum widest slightly above middle, strongly narrowed posteriad, with slightly concave laterobasal margins. Apical part of median lobe moderately narrow; parameres slightly not reaching apex of median lobe, with distinctly broadened apical portions (
<figureCitation id="B1CE2A1EFFD86D69D6A3DE55BF35013B" box="[233,306,1890,1912]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="49" pageNumber="300">Fig. 57</figureCitation>
). Body yellow-brown to reddish-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFD86D69D50CDE55BD880134" box="[838,911,1890,1912]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="49" pageNumber="300">Fig. 42</figureCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFD86D69D286DE55BB13013B" box="[1228,1300,1890,1912]" captionStart="FIGURES 57–63" captionStartId="23.[151,264,1646,1670]" captionTargetBox="[159,1428,189,1614]" captionTargetId="figure-16@23.[151,1436,181,1622]" captionTargetPageId="23" captionText="FIGURES 57–63. Paraphloeostiba guadalcanalensis (paratypes): 57–aedeagus, parameral view, 58–aedeagus, lateral view, 59–male abdominal tergite VIII, 60–male abdominal sternite VIII, 61–female abdominal tergite VIII, 62–female abdominal sternite VIII, 63–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821245" httpUri="https://zenodo.org/record/10821245/files/figure.png" pageId="49" pageNumber="300">Fig. 63</figureCitation>
. Body length: 
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. 
<collectingCountry id="51E2760BFFD86D69D733DEB7BC1D01D5" box="[377,538,1920,1942]" name="Solomon Islands" pageId="49" pageNumber="300">Solomon Islands</collectingCountry>
..................................................... 
<taxonomicName id="EEF54D18FFD86D69D2EBDEB7BB6701D6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1185,1376,1920,1941]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="300" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFD86D69D2EBDEB7BB6701D6" box="[1185,1376,1920,1941]" italics="true" pageId="49" pageNumber="300">P. guadalcanalensis</emphasis>
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<emphasis id="1B81EA89FFD86D69D32CDEB7BB9B01D6" bold="true" box="[1382,1436,1920,1941]" pageId="49" pageNumber="300">
<taxonomicNameLabel id="00B257F2FFD86D69D32CDEB7BB9B01D6" box="[1382,1436,1920,1941]" pageId="49" pageNumber="300" rank="species">sp. n.</taxonomicNameLabel>
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</paragraph>
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- Pronotum widest in middle, relatively gradually narowed toward hind angles, without distinctly concave laterobasal margins. Apical part of median lobe moderately broad; parameres reaching or slightly exceeding apex of median lobe, with slightly broadened apical portions (
<figureCitation id="B1CE2A1EFFD86D69D7ADDEEFBC2B01AE" box="[487,556,2007,2030]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="49" pageNumber="300">
Fig. 
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</figureCitation>
in Shavrin (2020)). Body brown, usually with paler middle portions of elytra. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD6A8D9A0BF1206EE" box="[226,277,151,173]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">
Fig 
<quantity id="EE0D9B7EFFDB6D6AD743D9AFBF1206EE" box="[265,277,152,173]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="50" pageNumber="301" unit="in" value="4.0">4</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD77FD9A0BFD206EE" author="Shavrin, A. V." box="[309,469,151,173]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDB6D6AD55AD9A0BD5206EE" box="[784,853,151,173]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="50" pageNumber="301">
Fig. 
<quantity id="EE0D9B7EFFDB6D6AD577D9AFBD5206EE" box="[829,853,152,173]" metricMagnitude="0" metricUnit="m" metricValue="2.5146" pageId="50" pageNumber="301" unit="in" value="99.0">99</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD53FD9A0BDE806EE" author="Steel, W. O." box="[885,1007,151,173]" pageId="50" pageNumber="301" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
. Body length: 
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. Cosmopolitan species................................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD21DD982BB9B068A" authority="(MacLeay, 1873)" baseAuthorityName="MacLeay" baseAuthorityYear="1873" box="[1111,1436,180,202]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="gayndahensis">
<emphasis id="1B81EA89FFDB6D6AD21DD982BAF5068A" box="[1111,1266,180,202]" italics="true" pageId="50" pageNumber="301">P. gayndahensis</emphasis>
(
<bibRefCitation id="4D644B6AFFDB6D6AD2B4D983BB92068A" author="MacLeay, W. J." box="[1278,1429,180,201]" pageId="50" pageNumber="301" pagination="79 - 205" refId="ref35618" refString="MacLeay, W. J. (1873) Notes on a collection of insects from Gayndah. The Transactions of the Entomological Society of New South Wales, 2, 79 - 205." type="journal article" year="1873">MacLeay, 1873</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD9E5BB9B0763" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
7 Elytra about twice as long as pronotum. Apical part of median lobe moderately wide; parameres slightly exceeding apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD72FD9D9BFAA0747" box="[357,429,238,260]" captionStart="FIGURES 64–67" captionStartId="25.[151,264,1719,1743]" captionTargetBox="[159,1426,190,1687]" captionTargetId="figure-16@25.[151,1436,181,1695]" captionTargetPageId="25" captionText="FIGURES 64–67. Paraphloeostiba iriana (holotype): 64–aedeagus, parameral view, 65–aedeagus, lateral view, 66–male abdominal tergite VIII, 67–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821247" httpUri="https://zenodo.org/record/10821247/files/figure.png" pageId="50" pageNumber="301">Fig. 64</figureCitation>
). Body brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD48ED9D9BD0B0740" box="[708,780,238,260]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="50" pageNumber="301">Fig. 40</figureCitation>
. Body length: 
<quantity id="EE0D9B7EFFDB6D6AD5DDD9D9BA220740" box="[919,1061,238,260]" metricMagnitude="-3" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.96" metricValueMin="1.72" pageId="50" pageNumber="301" unit="mm" value="1.8399999999999999" valueMax="1.96" valueMin="1.72">1.72–1.96 mm</quantity>
. 
<collectingCountry id="51E2760BFFDB6D6AD265D9D9BA8B0747" box="[1071,1164,238,260]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New 
<collectingCountry id="51E2760BFFDB6D6AD36BD9D9BB6F0747" box="[1313,1384,238,260]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
)........................................................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD344D83CBB660763" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1294,1377,267,288]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD344D83CBB660763" box="[1294,1377,267,288]" italics="true" pageId="50" pageNumber="301">P. iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CD83CBB9B0763" bold="true" box="[1382,1436,267,288]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CD83CBB9B0763" box="[1382,1436,267,288]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD81FBB9B0735" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Elytra distinctly less than twice as long as pronotum. Apex of median lobe narrow; parameres distinctly not exceeding apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD72FD873BFAA0719" box="[357,429,324,346]" captionStart="FIGURES 85–91" captionStartId="31.[151,264,1937,1961]" captionTargetBox="[164,1428,677,1906]" captionTargetId="figure-229@31.[151,1436,669,1914]" captionTargetPageId="31" captionText="FIGURES 85–91. Paraphloeostiba okapensis (paratypes): 85–aedeagus, parameral view, 86–aedeagus, lateral view, 87–male abdominal tergite VIII, 88–male abdominal sternite VIII, 89–female abdominal tergite VIII, 90–female abdominal sternite VIII, 91–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821259" httpUri="https://zenodo.org/record/10821259/files/figure.png" pageId="50" pageNumber="301">Fig. 85</figureCitation>
). Body reddish-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD559D873BD5D0719" box="[787,858,324,346]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 75</figureCitation>
. Body length: 
<quantity id="EE0D9B7EFFDB6D6AD5ACD873BA730719" box="[998,1140,324,346]" metricMagnitude="-3" metricUnit="m" metricValue="1.745" metricValueMax="1.86" metricValueMin="1.63" pageId="50" pageNumber="301" unit="mm" value="1.745" valueMax="1.86" valueMin="1.63">1.63–1.86 mm</quantity>
. 
<collectingCountry id="51E2760BFFDB6D6AD237D872BB310719" box="[1149,1334,324,346]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
............................................................................................ 
<taxonomicName id="EEF54D18FFDB6D6AD2A0D855BB660735" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1258,1377,353,375]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="okapensis" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2A0D855BB660735" box="[1258,1377,353,375]" italics="true" pageId="50" pageNumber="301">P. okapensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CD856BB9B0735" bold="true" box="[1382,1436,353,374]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CD856BB9B0735" box="[1382,1436,353,374]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD849BB9B07D0" blockId="50.[151,1437,151,2017]" box="[151,1436,382,404]" pageId="50" pageNumber="301">8 Antennomeres 6–10 strongly transverse, preapical antennomere three times as broad as long......................... 9</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD8ACBB9B07F3" blockId="50.[151,1437,151,2017]" box="[151,1436,411,433]" pageId="50" pageNumber="301">- Antennomeres 6–10 less transverse, preapical antennomere about twice as broad as long........................... 10</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDD88FBB9B0460" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
9 Elytra about twice as long as pronotum. Parameres significantly longer than apex of median lobe, with wide apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD6A3D8E3BF3B07AA" box="[233,316,468,490]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="50" pageNumber="301">Fig. 111</figureCitation>
). Head and elytra reddish or darker; pronotum and abdomen yellow-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD2EBD8E3BAEF07AA" box="[1185,1256,468,490]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 79</figureCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDB6D6AD72AD8C6BFB50445" box="[352,434,497,519]" captionStart="FIGURES 111–118" captionStartId="39.[151,264,1464,1488]" captionTargetBox="[159,1428,162,1442]" captionTargetId="figure-16@39.[151,1436,151,1447]" captionTargetPageId="39" captionText="FIGURES 111–118. Paraphloeostiba rhopalocera: 111–aedeagus, parameral view (holotype), 112–aedeagus, lateral view (holotype), 113–male abdominal tergite VIII, 114–male abdominal sternite VIII, 115–female abdominal tergite VIII, 116–female abdominal sternite VIII, 117–female accessory sclerite, 118–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821271" httpUri="https://zenodo.org/record/10821271/files/figure.png" pageId="50" pageNumber="301">Fig. 117</figureCitation>
. Body smaller: 
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. 
<collectingCountry id="51E2760BFFDB6D6AD4AAD8C6BD3A0444" box="[736,829,497,519]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(
<collectingRegion id="EB31F879FFDB6D6AD500D8C6BD980445" box="[842,927,497,518]" country="Indonesia" name="Sumatera Utara" pageId="50" pageNumber="301">Sumatra</collectingRegion>
), 
<collectingCountry id="51E2760BFFDB6D6AD5E5D8C5BA6F0444" box="[943,1128,497,519]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
................................................................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD22EDB38BB9B0460" authority="(Cameron, 1928)" baseAuthorityName="Cameron" baseAuthorityYear="1928" box="[1124,1436,526,548]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="rhopalocera">
<emphasis id="1B81EA89FFDB6D6AD22EDB38BAF50460" box="[1124,1266,526,548]" italics="true" pageId="50" pageNumber="301">P. rhopalocera</emphasis>
(
<bibRefCitation id="4D644B6AFFDB6D6AD2B5DB39BB920460" author="Cameron, M." box="[1279,1429,526,547]" pageId="50" pageNumber="301" pagination="90 - 110" refId="ref35375" refString="Cameron, M. (1928) Fauna sumatrensis. Staphylinidae (Col.). Entomologische Mitteilungen, 17 (2), 90 - 110." type="journal article" year="1928">Cameron, 1928</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDB1CBB9B0439" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Elytra distinctly less than twice as long as pronotum. Parameres distinctly exceeding apex of median lobe, with very wide apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD777DB70BF86041E" box="[317,385,583,605]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="50" pageNumber="301">
Fig. 
<quantity id="EE0D9B7EFFDB6D6AD723DB7FBF86041E" box="[361,385,584,605]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="50" pageNumber="301" unit="in" value="32.0">32</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD7D5DB70BC0F041E" author="Steel, W. O." box="[415,520,583,605]" pageId="50" pageNumber="301" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel 1960</bibRefCitation>
). Body reddish-brown. Forebody as in 
<figureCitation id="B1CE2A1EFFDB6D6AD536DB70BDB4041E" box="[892,947,583,605]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="50" pageNumber="301">
Fig. 
<quantity id="EE0D9B7EFFDB6D6AD5EDDB7FBDB4041E" box="[935,947,584,605]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="50" pageNumber="301" unit="in" value="9.0">9</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD59BDB70BA4E041E" author="Steel, W. O." box="[977,1097,583,605]" pageId="50" pageNumber="301" pagination="141 - 172" refId="ref36092" refString="Steel, W. O. (1960) Three new omaliine genera from Asia and Australasia previously confused with Phloeonomus Thomson (Coleoptera: Staphylinidae). The Transactions of the Royal Entomological Society of London, 112, 141 - 172. https: // doi. org / 10.1111 / j. 1365 - 2311.1960. tb 00497. x" type="journal article" year="1960">Steel (1960)</bibRefCitation>
. Body larger: 2.00 mm. 
<collectingCountry id="51E2760BFFDB6D6AD365DB7FBF200439" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
(
<collectingRegion id="EB31F879FFDB6D6AD77EDB53BFAB0439" box="[308,428,612,634]" country="Papua New Guinea" name="West New Britain" pageId="50" pageNumber="301">New Britain</collectingRegion>
)......................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD227DB52BB9B0439" authority="(Bernhauer, 1915)" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[1133,1436,612,634]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="specularis">
<emphasis id="1B81EA89FFDB6D6AD227DB52BAEF043A" box="[1133,1256,612,634]" italics="true" pageId="50" pageNumber="301">P. specularis</emphasis>
(
<bibRefCitation id="4D644B6AFFDB6D6AD2BEDB53BB920439" author="Bernhauer, M" box="[1268,1429,612,634]" pageId="50" pageNumber="301" pagination="179 - 202" refId="ref35314" refString="Bernhauer, M (1915) Beitrag zur Staphylinidenfauna von Neu-Guinea. Deutsche Entomologische Zeitschrift, 1915, 179 - 202. https: // doi. org / 10.1002 / mmnd. 48019150207" type="journal article" year="1915">Bernhauer, 1915</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDBB6BB9B04D5" blockId="50.[151,1437,151,2017]" box="[151,1436,641,663]" pageId="50" pageNumber="301">10 Laterobasal margins of pronotum distinctly concave in front of hind angles...................................... 11</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDBA9BB9B04F0" blockId="50.[151,1437,151,2017]" box="[151,1436,670,692]" pageId="50" pageNumber="301">- Laterobasal margins of pronotum gradually narrowed toward hind angles....................................... 13</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDB8CBB9B054A" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
11 Pronotum twice as broad as long, with widely rounded apical angles slighlty protruded apicad. Parameres very narrow, significantly shorter than apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD4E2DBE0BCFB04AE" box="[680,764,727,749]" captionStart="FIGURES 144–147" captionStartId="47.[151,264,1614,1638]" captionTargetBox="[171,1427,190,1582]" captionTargetId="figure-16@47.[151,1436,181,1591]" captionTargetPageId="47" captionText="FIGURES 144–147. Paraphloeostiba telnovi (holotype): 144–aedeagus, parameral view, 145–aedeagus, lateral view, 146–male abdominal tergite VIII, 147–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821291" httpUri="https://zenodo.org/record/10821291/files/figure.png" pageId="50" pageNumber="301">Fig. 144</figureCitation>
). Body reddish-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD228DBE0BAB204AE" box="[1122,1205,727,749]" captionStart="FIGURES 141–143" captionStartId="46.[151,264,1009,1033]" captionTargetBox="[212,1369,193,974]" captionTargetId="figure-22@46.[206,1381,181,986]" captionTargetPageId="46" captionText="FIGURES 141–143. Habitus of Omaliini: 141–Paraphloeostiba telnovi (holotype), 142–Xanthonomus guineensis (holotype), 143–X. thayerae (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821285" httpUri="https://zenodo.org/record/10821285/files/figure.png" pageId="50" pageNumber="301">Fig. 141</figureCitation>
. Body length: 
<quantity id="EE0D9B7EFFDB6D6AD30BDBEFBB9E04AE" box="[1345,1433,728,749]" metricMagnitude="-3" metricUnit="m" metricValue="1.82" pageId="50" pageNumber="301" unit="mm" value="1.82">1.82 mm</quantity>
. 
<collectingCountry id="51E2760BFFDB6D6AD6A8DBC2BF9C0549" box="[226,411,756,778]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
........................................................................ 
<taxonomicName id="EEF54D18FFDB6D6AD34DDBC2BB67054A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1287,1376,756,778]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="telnovi" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD34DDBC2BB67054A" box="[1287,1376,756,778]" italics="true" pageId="50" pageNumber="301">P. telnovi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDBC3BB9B054A" box="[1382,1436,756,777]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDA25BB9B0500" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">- Pronotum distinctly less than twice as broad as long, with narrower apical angles not protruded apicad. Parameres wider and longer............................................................................................. 12</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDA7CBB9B05DA" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
12 Head without or with indistinct fine punctation. Median lobe narrow; parameres narrow, reaching apex of median lobe, with narrow apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD78DDA50BC09053E" box="[455,526,871,893]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="50" pageNumber="301">Fig. 20</figureCitation>
). Body yellowish-brown to reddish brown, with paler elytra. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD296DA50BB11053E" box="[1244,1302,871,893]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">Fig. 4</figureCitation>
. Body length: 1.58–2.00 mm. 
<collectingCountry id="51E2760BFFDB6D6AD733DAB2BC3505D9" box="[377,562,900,922]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
......................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD2A4DAB2BB6605D9" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1262,1377,901,922]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="attenuata" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2A4DAB2BB6605D9" box="[1262,1377,901,922]" italics="true" pageId="50" pageNumber="301">P. attenuata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDAB3BB9B05DA" bold="true" box="[1382,1436,900,921]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDAB3BB9B05DA" box="[1382,1436,900,921]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDA95BB9B024E" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Head without punctation. Median lobe moderately wide; parameres slightly exceeding apex of median lobe, with distinctly broadened apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD7A0DA89BC230590" box="[490,548,958,980]" captionStart="FIGURES 7–10" captionStartId="4.[151,264,1664,1688]" captionTargetBox="[171,1427,192,1632]" captionTargetId="figure-22@4.[151,1435,181,1641]" captionTargetPageId="4" captionText="FIGURES 7–10. Nipponophloeostiba hornabrooki (holotype): 7–aedeagus, parameral view, 8–aedeagus, lateral view, 9–male abdominal tergite VIII, 10–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821218" httpUri="https://zenodo.org/record/10821218/files/figure.png" pageId="50" pageNumber="301">
Fig. 
<quantity id="EE0D9B7EFFDB6D6AD452DA89BC230590" box="[536,548,958,979]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="50" pageNumber="301" unit="in" value="7.0">7</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD40EDA89BCEA0597" author="Shavrin, A. V." box="[580,749,958,980]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b))</bibRefCitation>
. Body reddish-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD21BDA89BA8D0590" box="[1105,1162,958,980]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">
Fig. 
<quantity id="EE0D9B7EFFDB6D6AD234DA89BA8D0590" box="[1150,1162,958,979]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="50" pageNumber="301" unit="in" value="1.0">1</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD2E1DA89BB4C0597" author="Shavrin, A. V." box="[1195,1355,958,980]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDB6D6AD789DAECBC0D05B3" box="[451,522,987,1009]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="50" pageNumber="301">Fig. 38</figureCitation>
. Body length: 
<quantity id="EE0D9B7EFFDB6D6AD4DCDAECBD2305B3" box="[662,804,987,1009]" metricMagnitude="-3" metricUnit="m" metricValue="2.065" metricValueMax="2.35" metricValueMin="1.78" pageId="50" pageNumber="301" unit="mm" value="2.065" valueMax="2.35" valueMin="1.78">1.78–2.35 mm</quantity>
. 
<collectingCountry id="51E2760BFFDB6D6AD567DAECBD8D05B2" box="[813,906,987,1009]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New 
<collectingCountry id="51E2760BFFDB6D6AD26ADAECBA6E05B2" box="[1056,1129,987,1009]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
), 
<collectingCountry id="51E2760BFFDB6D6AD233DAECBB3505B2" box="[1145,1330,987,1009]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
...................................................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD2DCDACFBB9B024E" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1174,1436,1015,1037]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFDB6D6AD2DCDACFBAA2024E" box="[1174,1189,1016,1037]" italics="true" pageId="50" pageNumber="301">P</emphasis>
. 
<emphasis id="1B81EA89FFDB6D6AD2FBDACFBB17024E" box="[1201,1296,1016,1037]" italics="true" pageId="50" pageNumber="301">conjuncta</emphasis>
Shavrin, 2023
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDD23BB9B026A" blockId="50.[151,1437,151,2017]" box="[151,1436,1044,1066]" pageId="50" pageNumber="301">13 Head and pronotum without punctation.................................................................. 14</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDD05BB9B0204" blockId="50.[151,1437,151,2017]" box="[151,1436,1073,1095]" pageId="50" pageNumber="301">- Head without or with distinct punctation, but pronotum always with punctures................................... 15</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDD79BB9B02FA" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
14 Pronotum transverse, slightly more than twice as broad as long; surface of elytra without elevations between punctures. Parameres slightly exceeding apex of median lobe, with wide apical portions, with several short apical and preapical setae (
<figureCitation id="B1CE2A1EFFDB6D6AD33FDD5CBEFA02DE" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="50" pageNumber="301">Fig. 97</figureCitation>
). Body yellow-brown, with reddish-brown apical portions of elytra. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD254DDB0BA6102DE" box="[1054,1126,1159,1181]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 76</figureCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDB6D6AD6A8DD93BF3202FA" box="[226,309,1188,1210]" captionStart="FIGURES 97–103" captionStartId="35.[151,264,1467,1491]" captionTargetBox="[160,1427,159,1442]" captionTargetId="figure-16@35.[151,1436,151,1450]" captionTargetPageId="35" captionText="FIGURES 97–103. Paraphloeostiba penelopeae (paratypes): 97–aedeagus, parameral view, 98–aedeagus, lateral view, 99– male abdominal tergite VIII, 100–male abdominal sternite VIII, 101–female abdominal tergite VIII, 102–female abdominal sternite VIII, 103–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821267" httpUri="https://zenodo.org/record/10821267/files/figure.png" pageId="50" pageNumber="301">Fig. 103</figureCitation>
. Body length: 1.40–2.00 mm. 
<collectingCountry id="51E2760BFFDB6D6AD412DD93BCFE02F9" box="[600,761,1188,1210]" name="Solomon Islands" pageId="50" pageNumber="301">Solomon Islands</collectingCountry>
....................................... 
<taxonomicName id="EEF54D18FFDB6D6AD291DD92BB6702FA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1243,1376,1188,1210]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="penelopeae" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD291DD92BB6702FA" box="[1243,1376,1188,1210]" italics="true" pageId="50" pageNumber="301">P. penelopeae</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDD93BB9B02FA" bold="true" box="[1382,1436,1188,1209]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDD93BB9B02FA" box="[1382,1436,1188,1209]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDDF5BB9B036E" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Pronotum less transverse, about one and a half times as broad as long; surface of elytra with distinct elevations between punctures. Parameres significantly exceeding apes of median lobe, with numerous very long apical and preapical setae (
<figureCitation id="B1CE2A1EFFDB6D6AD33FDDE9BF0E0353" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="50" pageNumber="301">Fig. 104</figureCitation>
). Body dark-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD41BDDCCBC900353" box="[593,663,1275,1297]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="50" pageNumber="301">Fig. 77</figureCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDB6D6AD586DDCCBA190353" box="[972,1054,1275,1297]" captionStart="FIGURES 104–110" captionStartId="37.[151,264,1945,1969]" captionTargetBox="[159,1428,605,1914]" captionTargetId="figure-180@37.[151,1436,597,1922]" captionTargetPageId="37" captionText="FIGURES 104–110. Paraphloeostiba pilosa: 104–aedeagus, parameral view (holotype), 105–aedeagus, lateral view (holotype), 106–male abdominal tergite VIII, 107–male abdominal sternite VIII, 108–female abdominal tergite VIII, 109–female abdominal sternite VIII, 110–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821269" httpUri="https://zenodo.org/record/10821269/files/figure.png" pageId="50" pageNumber="301">Fig. 110</figureCitation>
. Body length: 
<quantity id="EE0D9B7EFFDB6D6AD2E2DDCCBB310353" box="[1192,1334,1275,1297]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.65" metricValueMin="1.95" pageId="50" pageNumber="301" unit="mm" value="2.3" valueMax="2.65" valueMin="1.95">1.95–2.65 mm</quantity>
. 
<collectingCountry id="51E2760BFFDB6D6AD375DDCCBB9B0352" box="[1343,1436,1275,1297]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New 
<collectingCountry id="51E2760BFFDB6D6AD73BDC20BFBF036E" box="[369,440,1303,1325]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
)..................................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD344DC2FBB66036F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1294,1377,1303,1325]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="pilosa" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD344DC2FBB66036F" box="[1294,1377,1303,1325]" italics="true" pageId="50" pageNumber="301">P. pilosa</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDC2FBB9B036E" bold="true" box="[1382,1436,1304,1325]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDC2FBB9B036E" box="[1382,1436,1304,1325]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDC03BB9B03C0" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
15 Antennomeres 6–7 about as broad as long. Aedeagus long, with moderately wide and elongate median lobe with rounded apex (
<figureCitation id="B1CE2A1EFFDB6D6AD6A3DC66BF360325" box="[233,305,1361,1383]" captionStart="FIGURES 81–84" captionStartId="29.[151,264,1649,1673]" captionTargetBox="[168,1428,710,1617]" captionTargetId="figure-145@29.[151,1436,704,1625]" captionTargetPageId="29" captionText="FIGURES 81–84. Paraphloeostiba margineguttata (holotype): 81–aedeagus, parameral view, 82–aedeagus, lateral view, 83– male abdominal tergite VIII, 84–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821255" httpUri="https://zenodo.org/record/10821255/files/figure.png" pageId="50" pageNumber="301">Fig. 81</figureCitation>
). Body reddish-brown, with paler elytra and abdomen. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD59ADC66BA1F0325" box="[976,1048,1361,1383]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="50" pageNumber="301">Fig. 43</figureCitation>
. Body length: 
<quantity id="EE0D9B7EFFDB6D6AD2ECDC66BB320324" box="[1190,1333,1361,1383]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="2.77" metricValueMin="2.57" pageId="50" pageNumber="301" unit="mm" value="2.67" valueMax="2.77" valueMin="2.57">2.57–2.77 mm</quantity>
. 
<collectingCountry id="51E2760BFFDB6D6AD375DC66BB9B0324" box="[1343,1436,1361,1383]" name="Indonesia" pageId="50" pageNumber="301">Indonesia</collectingCountry>
(Western New 
<collectingCountry id="51E2760BFFDB6D6AD73BDC59BFBF03C7" box="[369,440,1390,1412]" name="Guinea-Bissau" pageId="50" pageNumber="301">Guinea</collectingCountry>
)............................................................. 
<taxonomicName id="EEF54D18FFDB6D6AD2FCDC58BB6703C7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1206,1376,1391,1412]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="margineguttata" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2FCDC58BB6703C7" box="[1206,1376,1391,1412]" italics="true" pageId="50" pageNumber="301">P. margineguttata</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDC59BB9B03C0" bold="true" box="[1382,1436,1390,1411]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDC59BB9B03C0" box="[1382,1436,1390,1411]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDCBCBB9B03E2" blockId="50.[151,1437,151,2017]" box="[151,1436,1419,1441]" pageId="50" pageNumber="301">- Antennomeres 6–10 distinctly transverse. Aedeagus and coloration of body different.............................. 16</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDC90BB9B03FE" blockId="50.[151,1437,151,2017]" box="[151,1436,1447,1469]" pageId="50" pageNumber="301">16 Head without punctation.............................................................................. 17</paragraph>
<paragraph id="294A369BFFDB6D6AD6D7DCF2BB9B039A" blockId="50.[151,1437,151,2017]" box="[157,1436,1476,1498]" pageId="50" pageNumber="301">- Head with distinct punctation.......................................................................... 22</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDCD6BB9B0073" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
17 Elytra with large and deep punctation and with distinct elevations between punctures. Apical portion of median lobe very narrow; parameres with very wide apical portions (
<figureCitation id="B1CE2A1EFFDB6D6AD49BDCC9BD210050" box="[721,806,1534,1556]" captionStart="FIGURES 133–140" captionStartId="44.[151,264,1943,1967]" captionTargetBox="[185,1403,678,1912]" captionTargetId="figure-200@44.[177,1411,669,1920]" captionTargetPageId="44" captionText="FIGURES 133–140. Paraphloeostiba steeli: 133–aedeagus, parameral view (holotype), 134–aedeagus, lateral view (holotype), 135–male abdominal tergite VIII, 136–male abdominal sternite VIII, 137–female abdominal tergite VIII, 138–female abdominal sternite VIII, 139–female accessory sclerite, 140–spermatheca. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821281" httpUri="https://zenodo.org/record/10821281/files/figure.png" pageId="50" pageNumber="301">Fig. 133</figureCitation>
). Body reddish-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD2DBDCC9BADF0050" box="[1169,1240,1534,1556]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="50" pageNumber="301">Fig. 44</figureCitation>
. Body length: 
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. 
<collectingCountry id="51E2760BFFDB6D6AD709DF2CBFFB0072" box="[323,508,1563,1585]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
................................................................ 
<taxonomicName id="EEF54D18FFDB6D6AD35CDF2CBB660073" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1302,1377,1563,1584]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD35CDF2CBB660073" box="[1302,1377,1563,1584]" italics="true" pageId="50" pageNumber="301">P. steeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDF2CBB9B0073" bold="true" box="[1382,1436,1563,1584]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDF2CBB9B0073" box="[1382,1436,1563,1584]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDF0FBB9B002A" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">- Elytra with distinctly finer punctation, without elevations between punctures.Apical portion of median lobe narrower; parameres different. Coloration of body different................................................................... 18</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDF46BB9B0083" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
18 Elytra longer, twice as long as pronotum. Aedeagus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD546DF46BD5300C4" box="[780,852,1649,1671]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="50" pageNumber="301">Fig. 46</figureCitation>
. Body yellow-brown to brown, with slightly paler elytra and darkened apical portions. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDB6D6AD542DFB9BD4800E7" box="[776,847,1678,1700]" captionStart="FIGURES 46–52" captionStartId="19.[151,264,1524,1548]" captionTargetBox="[172,1415,189,1492]" captionTargetId="figure-16@19.[164,1423,181,1500]" captionTargetPageId="19" captionText="FIGURES 46–52. Paraphloeostiba coprophila (paratypes): 46–aedeagus, parameral view, 47–aedeagus, lateral view, 48–male abdominal tergite VIII, 49–male abdominal sternite VIII, 50–female abdominal tergite VIII, 51–female abdominal sternite VIII, 52–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821239" httpUri="https://zenodo.org/record/10821239/files/figure.png" pageId="50" pageNumber="301">Fig. 52</figureCitation>
. Body length: 
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. 
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............................................................................................ 
<taxonomicName id="EEF54D18FFDB6D6AD2AADF9CBB670083" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1248,1376,1707,1728]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2AADF9CBB670083" box="[1248,1376,1707,1728]" italics="true" pageId="50" pageNumber="301">P. coprophila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDF9CBB9B0083" bold="true" box="[1382,1436,1707,1728]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDF9CBB9B0083" box="[1382,1436,1707,1728]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDFFFBB9B009E" blockId="50.[151,1437,151,2017]" box="[151,1436,1735,1757]" pageId="50" pageNumber="301">- Elytra shorter, distinctly less than twice as long as pronotum. Coloration of body different.......................... 19</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDFD3BB9B00BA" blockId="50.[151,1437,151,2017]" box="[151,1436,1764,1786]" pageId="50" pageNumber="301">19 Body yellow-brown. Median lobe and apical portions of parameres wide........................................ 20</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDE35BB9B0155" blockId="50.[151,1437,151,2017]" box="[151,1436,1793,1815]" pageId="50" pageNumber="301">- Body darker. Median lobe and apical portions of parameres narrow............................................ 21</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDE29BB9B012E" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
20 Lateral sides of pronotum gradually narrowed both anteriad and posteriad. Body yellow-brown, with reddish-brown apical portions of elytra. Median lobe slightly broader; parameres significantly longer than apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD295DE0CBB200113" box="[1247,1319,1851,1873]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="50" pageNumber="301">Fig. 12</figureCitation>
). Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD6B1DE60BF28012E" box="[251,303,1879,1901]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">Fig 2</figureCitation>
. Body smaller: 
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. 
<collectingCountry id="51E2760BFFDB6D6AD46DDE6FBCE7012E" box="[551,736,1879,1901]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
........................................... 
<taxonomicName id="EEF54D18FFDB6D6AD2B0DE6FBB66012F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1274,1377,1879,1901]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="assimile" status="sp. n.">
<emphasis id="1B81EA89FFDB6D6AD2B0DE6FBB66012F" box="[1274,1377,1879,1901]" italics="true" pageId="50" pageNumber="301">P. assimile</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDB6D6AD32CDE6FBB9B012E" bold="true" box="[1382,1436,1880,1901]" pageId="50" pageNumber="301">
<taxonomicNameLabel id="00B257F2FFDB6D6AD32CDE6FBB9B012E" box="[1382,1436,1880,1901]" pageId="50" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDB6D6AD6DDDE42BB9B01A3" blockId="50.[151,1437,151,2017]" pageId="50" pageNumber="301">
- Lateral sides of pronotum slightly more narrowed posteriad than anteriad. Body yellow-brown. Median lobe relatively narrower; parameres slightly exceeding apex of median lobe (
<figureCitation id="B1CE2A1EFFDB6D6AD480DEA6BD0901E4" box="[714,782,1937,1959]" captionStart="FIGURES 12–15" captionStartId="7.[151,264,1981,2005]" captionTargetBox="[160,1427,418,1948]" captionTargetId="figure-91@7.[151,1436,417,1957]" captionTargetPageId="7" captionText="FIGURES 12–15. Paraphloeostiba assimile (holotype): 12–aedeagus, parameral view, 13–aedeagus, lateral view, 14–male abdominal tergite VIII, 15–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821223" httpUri="https://zenodo.org/record/10821223/files/figure.png" pageId="50" pageNumber="301">
Fig. 
<quantity id="EE0D9B7EFFDB6D6AD4BCDEA6BD0901E4" box="[758,782,1937,1959]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="50" pageNumber="301" unit="in" value="15.0">15</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD561DEA6BDD501E4" author="Shavrin, A. V." box="[811,978,1937,1959]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b))</bibRefCitation>
. Habitus as in 
<figureCitation id="B1CE2A1EFFDB6D6AD211DEA6BA9501E5" box="[1115,1170,1937,1959]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="50" pageNumber="301">
Fig. 
<quantity id="EE0D9B7EFFDB6D6AD2CCDEA6BA9501E5" box="[1158,1170,1937,1958]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="50" pageNumber="301" unit="in" value="3.0">3</quantity>
</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD2E5DEA6BB4A01E4" author="Shavrin, A. V." box="[1199,1357,1937,1959]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite asi n 
<figureCitation id="B1CE2A1EFFDB6D6AD789DE99BC000180" box="[451,519,1966,1988]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="50" pageNumber="301">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDB6D6AD46CDE99BCC20187" author="Shavrin, A. V." box="[550,709,1966,1988]" pageId="50" pageNumber="301" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Body larger: 2.00– 
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. 
<collectingCountry id="51E2760BFFDB6D6AD5AEDE99BA9A0187" box="[996,1181,1966,1988]" name="Papua New Guinea" pageId="50" pageNumber="301">Papua New Guinea</collectingCountry>
................................................................................................... 
<taxonomicName id="EEF54D18FFDB6D6AD2E2DEFBBB9B01A3" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1192,1436,1995,2017]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="301" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="1B81EA89FFDB6D6AD2E2DEFBBAB001A2" box="[1192,1207,1996,2017]" italics="true" pageId="50" pageNumber="301">P</emphasis>
. 
<emphasis id="1B81EA89FFDB6D6AD288DEFCBB0801A3" box="[1218,1295,1995,2016]" italics="true" pageId="50" pageNumber="301">flaveola</emphasis>
Shavrin, 2023
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD9AFBB9B0689" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
21 Disc of pronotum without longitudinal impressions. Apical parts of median lobe and parameres narrow (
<figureCitation id="B1CE2A1EFFDA6D6BD28DD9A0BB0906EE" box="[1223,1294,151,173]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="51" pageNumber="302">Fig. 32</figureCitation>
). Body brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD72FD982BF980688" box="[357,415,181,203]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">Fig. 6</figureCitation>
. Body smaller: 
<quantity id="EE0D9B7EFFDA6D6BD47DD982BC880689" box="[567,655,181,202]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="51" pageNumber="302" unit="mm" value="1.42">1.42 mm</quantity>
. 
<collectingCountry id="51E2760BFFDA6D6BD4D2D982BD560688" box="[664,849,181,203]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
................................. 
<taxonomicName id="EEF54D18FFDA6D6BD2A3D982BB670689" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1257,1376,181,202]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD2A3D982BB670689" box="[1257,1376,181,202]" italics="true" pageId="51" pageNumber="302">P. betlephila</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD982BB9B0689" bold="true" box="[1382,1436,181,202]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD982BB9B0689" box="[1382,1436,181,202]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD9E4BB9B0761" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
- Disc of pronotum with indistinct longitudinal impressions. Apical parts of median lobe and parameres distinctly broader (
<figureCitation id="B1CE2A1EFFDA6D6BD33FD9E5BF0E0746" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="51" pageNumber="302">Fig. 119</figureCitation>
). Body yellowish brown to reddish-brown, with slightly paler elytra. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD26FD9D8BA6B0746" box="[1061,1132,239,261]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="51" pageNumber="302">Fig. 78</figureCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDA6D6BD6A8D83ABF320760" box="[226,309,269,291]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="51" pageNumber="302">Fig. 125</figureCitation>
. Body larger: 
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. 
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(Western New 
<collectingCountry id="51E2760BFFDA6D6BD50CD83ABD8A0760" box="[838,909,269,291]" name="Guinea-Bissau" pageId="51" pageNumber="302">Guinea</collectingCountry>
)............................. 
<taxonomicName id="EEF54D18FFDA6D6BD340D83ABB660761" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1290,1377,269,290]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD340D83ABB660761" box="[1290,1377,269,290]" italics="true" pageId="51" pageNumber="302">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD83ABB9B0761" bold="true" box="[1382,1436,269,290]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD83ABB9B0761" box="[1382,1436,269,290]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD81DBB9B071E" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
22 Body dark brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD46DD81DBC64077C" box="[551,611,298,320]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">Fig. 3</figureCitation>
. Body distinctly larger: 
<quantity id="EE0D9B7EFFDA6D6BD51ED81DBDA90703" box="[852,942,298,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="51" pageNumber="302" unit="mm" value="2.75">2.75 mm</quantity>
. Aedeagus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD21CD81DBAA70703" box="[1110,1184,298,320]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="51" pageNumber="302">Fig. 16</figureCitation>
. 
<collectingCountry id="51E2760BFFDA6D6BD2E1D81DBB0F0703" box="[1195,1288,298,320]" name="Indonesia" pageId="51" pageNumber="302">Indonesia</collectingCountry>
(Western New 
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)........................................................................... 
<taxonomicName id="EEF54D18FFDA6D6BD280D87FBB66071E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1226,1377,328,349]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD280D87FBB66071E" box="[1226,1377,328,349]" italics="true" pageId="51" pageNumber="302">P. atramentaria</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD87FBB9B071E" bold="true" box="[1382,1436,328,349]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD87FBB9B071E" box="[1382,1436,328,349]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD852BB9B0739" blockId="51.[151,1437,151,877]" box="[151,1436,357,379]" pageId="51" pageNumber="302">- Body paler and smaller............................................................................... 23</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD8B5BB9B07F6" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
23 Median lobe wide, with widely rounded apex (
<figureCitation id="B1CE2A1EFFDA6D6BD4EDD8B5BCF707DB" box="[679,752,386,408]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="51" pageNumber="302">Fig. 68</figureCitation>
). Body reddish brown, with paler elytra. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD34ED8B5BB4B07D4" box="[1284,1356,386,408]" captionStart="FIGURES 39–44" captionStartId="17.[151,264,1881,1905]" captionTargetBox="[177,1411,194,1844]" captionTargetId="figure-16@17.[164,1424,181,1857]" captionTargetPageId="17" captionText="FIGURES 39–44. Habitus of Paraphloeostiba: 39–P. coriacea (Western New Guinea), 40–P. iriana (paratype), 41–P. latissima (paratype), 42–P. guadalcanalensis (paratype), 43–P. margineguttata (holotype), 44–P. steeli (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821233" httpUri="https://zenodo.org/record/10821233/files/figure.png" pageId="51" pageNumber="302">Fig. 41</figureCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDA6D6BD789D8A8BC0D07F6" box="[451,522,415,437]" captionStart="FIGURES 68–74" captionStartId="27.[151,264,1913,1937]" captionTargetBox="[172,1428,642,1882]" captionTargetId="figure-207@27.[151,1436,633,1890]" captionTargetPageId="27" captionText="FIGURES 68–74. Paraphloeostiba latissima (paratypes): 68–aedeagus, parameral view, 69–aedeagus, lateral view, 70–male abdominal tergite VIII, 71–male abdominal sternite VIII, 72–female abdominal tergite VIII, 73–female abdominal sternite VIII, 74–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821251" httpUri="https://zenodo.org/record/10821251/files/figure.png" pageId="51" pageNumber="302">Fig. 74</figureCitation>
. Body length: 
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. 
<collectingCountry id="51E2760BFFDA6D6BD567D897BDE107F6" box="[813,998,415,437]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
.................... 
<taxonomicName id="EEF54D18FFDA6D6BD2B8D897BB6607F7" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1266,1377,415,437]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="latissima" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD2B8D897BB6607F7" box="[1266,1377,415,437]" italics="true" pageId="51" pageNumber="302">P. latissima</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFDA6D6BD32CD897BB9B07F6" bold="true" box="[1382,1436,416,437]" pageId="51" pageNumber="302">
<taxonomicNameLabel id="00B257F2FFDA6D6BD32CD897BB9B07F6" box="[1382,1436,416,437]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD88ABB9B0791" blockId="51.[151,1437,151,877]" box="[151,1436,445,467]" pageId="51" pageNumber="302">- Median lobe distinctly narrower, with narrower apex. Coloration different....................................... 24</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD8EDBB9B07B3" blockId="51.[151,1437,151,877]" box="[151,1436,474,496]" pageId="51" pageNumber="302">24 Median lobe wide; parameres with distinctly broadened apical portions......................................... 25</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDD8CFBB9B044E" blockId="51.[151,1437,151,877]" box="[151,1436,503,525]" pageId="51" pageNumber="302">- Median lobe and apical portion of parameres narrower...................................................... 26</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDDB22BB9B0426" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
25 Pronotum strongly transverse. Apical portions of parameres very wide (
<figureCitation id="B1CE2A1EFFDA6D6BD5D0DB22BDE40469" box="[922,995,533,555]" captionStart="FIGURES 92–95" captionStartId="33.[151,264,1780,1804]" captionTargetBox="[160,1427,189,1747]" captionTargetId="figure-16@33.[151,1436,181,1756]" captionTargetPageId="33" captionText="FIGURES 92–95. Paraphloeostiba papuana (Western New Guinea): 92–aedeagus, parameral view, 93–aedeagus, lateral view, 94–male abdominal tergite VIII, 95–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821261" httpUri="https://zenodo.org/record/10821261/files/figure.png" pageId="51" pageNumber="302">Fig. 92</figureCitation>
). Body yellow-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD303DB22BB850468" box="[1353,1410,533,555]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDA6D6BD6A8DB05BF86040B" author="Shavrin, A. V." box="[226,385,562,584]" pageId="51" pageNumber="302" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDA6D6BD4FCDB05BCFD0404" box="[694,762,562,584]" captionStart="FIGURES 20–25" captionStartId="11.[151,264,1928,1952]" captionTargetBox="[159,1428,711,1895]" captionTargetId="figure-258@11.[151,1436,705,1905]" captionTargetPageId="11" captionText="FIGURES 20–25. Paraphloeostiba attenuata (paratypes): 20–aedeagus, parameral view, 21–aedeagus, lateral view, 22–male abdominal tergite VIII, 23–male abdominal sternite VIII, 24–female abdominal tergite VIII, 25–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821227" httpUri="https://zenodo.org/record/10821227/files/figure.png" pageId="51" pageNumber="302">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="4D644B6AFFDA6D6BD552DB05BDB0040B" author="Shavrin, A. V." box="[792,951,562,584]" pageId="51" pageNumber="302" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Body smaller: 
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. 
<collectingCountry id="51E2760BFFDA6D6BD2A9DB05BB47040B" box="[1251,1344,562,584]" name="Indonesia" pageId="51" pageNumber="302">Indonesia</collectingCountry>
(Western New 
<collectingCountry id="51E2760BFFDA6D6BD75FDB78BF590426" box="[277,350,591,613]" name="Guinea-Bissau" pageId="51" pageNumber="302">Guinea</collectingCountry>
), 
<collectingCountry id="51E2760BFFDA6D6BD724DB67BC200426" box="[366,551,591,613]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
................................................. 
<taxonomicName id="EEF54D18FFDA6D6BD2CEDB67BB9B0426" authority="(Cameron, 1931)" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[1156,1436,592,613]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="papuana">
<emphasis id="1B81EA89FFDA6D6BD2CEDB67BA940426" box="[1156,1171,592,613]" italics="true" pageId="51" pageNumber="302">P</emphasis>
. 
<emphasis id="1B81EA89FFDA6D6BD2D4DB67BAF50426" box="[1182,1266,592,613]" italics="true" pageId="51" pageNumber="302">papuana</emphasis>
(
<bibRefCitation id="4D644B6AFFDA6D6BD2B5DB67BB920426" author="Cameron, M." box="[1279,1429,592,613]" pageId="51" pageNumber="302" pagination="355 - 364" refId="ref35403" refString="Cameron, M. (1931) Staphylinidae (Coleoptera) from New Guinea, in the South Australian Museum. Records of the South Australian Museum, 4 (3), 355 - 364." type="journal article" year="1931">Cameron, 1931</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDDB5ABB9B04FE" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
- Pronotum slightly transverse. Apical portions of parameres distinctly narrower (
<figureCitation id="B1CE2A1EFFDA6D6BD597DB5ABA3604C0" box="[989,1073,621,643]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="51" pageNumber="302">Fig. 126</figureCitation>
). Body reddish-brown, with distinctly paler elytra. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD794DBBDBC2204DC" box="[478,549,650,672]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="51" pageNumber="302">Fig. 80</figureCitation>
. Female accessory sclerite as in 
<figureCitation id="B1CE2A1EFFDA6D6BD516DBBDBDA804DC" box="[860,943,650,672]" captionStart="FIGURES 126–132" captionStartId="42.[151,264,1947,1971]" captionTargetBox="[159,1428,660,1914]" captionTargetId="figure-230@42.[151,1436,653,1924]" captionTargetPageId="42" captionText="FIGURES 126–132. Paraphloeostiba rufula: 126–aedeagus, parameral view (holotype), 127–aedeagus, lateral view (holotype), 128–male abdominal tergite VIII, 129–male abdominal sternite VIII, 130–female abdominal tergite VIII, 131–female abdominal sternite VIII, 132–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821277" httpUri="https://zenodo.org/record/10821277/files/figure.png" pageId="51" pageNumber="302">Fig. 132</figureCitation>
. Body larger: 2.00– 
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. 
<collectingCountry id="51E2760BFFDA6D6BD287DBBCBB8104E3" box="[1229,1414,650,672]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
......................................................................................... 
<taxonomicName id="EEF54D18FFDA6D6BD344DB9FBB6604FF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1294,1377,679,701]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="rufula" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD344DB9FBB6604FF" box="[1294,1377,679,701]" italics="true" pageId="51" pageNumber="302">P. rufula</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFDA6D6BD32CDB9FBB9B04FE" box="[1382,1436,680,701]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDDBF2BB9B0556" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
26 Basal portion of aedeagus narrow; median lobe moderately wide and long; parameres narrow, distinctly not reaching apex of median lobe (
<figureCitation id="B1CE2A1EFFDA6D6BD729DBD5BFAF04BB" box="[355,424,738,760]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="51" pageNumber="302">
Fig. 
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in 
<bibRefCitation id="4D644B6AFFDA6D6BD78FDBD5BC6B04BB" author="Shavrin, A. V." box="[453,620,738,760]" pageId="51" pageNumber="302" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b))</bibRefCitation>
. Body reddish-brown, with paler elytra. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD227DBD5BAA304BB" box="[1133,1188,738,760]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">
Fig. 
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in 
<bibRefCitation id="4D644B6AFFDA6D6BD28BDBD5BB6704BB" author="Shavrin, A. V." box="[1217,1376,738,760]" pageId="51" pageNumber="302" pagination="486 - 500" refId="ref35934" refString="Shavrin, A. V. (2023 b) New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines. Zootaxa, 5301 (4), 486 - 500. https: // doi. org / 10.11646 / zootaxa. 5301.4.6" type="journal article" year="2023">Shavrin (2023b)</bibRefCitation>
. Body length: 
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. 
<collectingCountry id="51E2760BFFDA6D6BD788DA37BC7C0556" box="[450,635,767,789]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
.............................................. 
<taxonomicName id="EEF54D18FFDA6D6BD2E4DA37BB9B0556" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1198,1436,767,789]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="ullrichi">
<emphasis id="1B81EA89FFDA6D6BD2E4DA37BABA0556" box="[1198,1213,768,789]" italics="true" pageId="51" pageNumber="302">P</emphasis>
. 
<emphasis id="1B81EA89FFDA6D6BD283DBC8BB170557" box="[1225,1296,767,788]" italics="true" pageId="51" pageNumber="302">ullrichi</emphasis>
Shavrin, 2023
</taxonomicName>
</paragraph>
<paragraph id="294A369BFFDA6D6BD6DDDA2ABB9B052E" blockId="51.[151,1437,151,877]" pageId="51" pageNumber="302">
- Basal portion of aedeagus wide; median lobe very narrow, moderately short; parameres distinctly exceeding apex of median lobe, with broadened apical portions (
<figureCitation id="B1CE2A1EFFDA6D6BD414DA0DBCAD0513" box="[606,682,826,848]" captionStart="FIGURES 26–31" captionStartId="13.[151,264,1583,1607]" captionTargetBox="[187,1428,185,1557]" captionTargetId="figure-16@13.[151,1436,181,1560]" captionTargetPageId="13" captionText="FIGURES 26–31. Paraphloeostiba barclayi (holotype): 26–aedeagus, parameral view, 27–aedeagus, lateral view, 28–male abdominal tergite VIII, 29–male abdominal sternite VIII, 30–female abdominal tergite VIII, 31–female abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821229" httpUri="https://zenodo.org/record/10821229/files/figure.png" pageId="51" pageNumber="302">Fig. 26</figureCitation>
). Body yellow-brown to reddish-brown. Habitus as in 
<figureCitation id="B1CE2A1EFFDA6D6BD29BDA0DBB080513" box="[1233,1295,826,848]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="51" pageNumber="302">Fig. 5</figureCitation>
. Body length: 
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. 
<collectingCountry id="51E2760BFFDA6D6BD733DA6FBC35052E" box="[377,562,855,877]" name="Papua New Guinea" pageId="51" pageNumber="302">Papua New Guinea</collectingCountry>
.......................................................... 
<taxonomicName id="EEF54D18FFDA6D6BD2B0DA6FBB67052F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1274,1376,855,877]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="302" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
<emphasis id="1B81EA89FFDA6D6BD2B0DA6FBB67052F" box="[1274,1376,855,877]" italics="true" pageId="51" pageNumber="302">P. barclayi</emphasis>
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<taxonomicNameLabel id="00B257F2FFDA6D6BD32CDA6FBB9B052E" box="[1382,1436,856,877]" pageId="51" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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