807 lines
107 KiB
XML
807 lines
107 KiB
XML
<document id="896FF4AE7B7BCAEBB465459D298BE108" ID-DOI="10.1080/00222933.2021.1936679" ID-ISSN="1464-5262" ID-Zenodo-Dep="5466018" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630920019635" checkinUser="plazi" docAuthor="Ma, Zhen-Xing, Lei, Zhi-Ming, Li, Wen-Juan & Zhou, Chang-Fa" docDate="2021" docId="650487C7FF94FF9EFE66FF5CFDECFB21" docLanguage="en" docName="JNATHIST.55.17-18.1131-1159.pdf" docOrigin="Journal of Natural History 55 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2021.1936679" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Epeorus (Proepeorus) falcatus Ma & Lei & Li & Zhou 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1152" masterDocId="993DFFBFFF99FF8BFFE7FFD2FFF1FFB4" masterDocTitle="A revision of Epeorus (Proepeorus) in China, with descriptions of two new species (Ephemeroptera, Heptageniidae)" masterLastPageNumber="1159" masterPageNumber="1131" pageNumber="1144" updateTime="1699282404856" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="00FC1003809BCD06BCBA050A3123D160">A revision of Epeorus (Proepeorus) in China, with descriptions of two new species (Ephemeroptera, Heptageniidae)</mods:title>
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<mods:namePart id="8EE573DC20C40534D77747A5D819CD59">Ma, Zhen-Xing</mods:namePart>
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<mods:namePart id="32691B264CF323157E7315DBC99C7009">Lei, Zhi-Ming</mods:namePart>
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<mods:namePart id="0AA8CD63A67A9A19473524B978F66262">Li, Wen-Juan</mods:namePart>
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<mods:namePart id="9ABDC978FE3C3E85985AAA574F4A6AB9">Zhou, Chang-Fa</mods:namePart>
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<mods:title id="3778CD46ED45679EF171051EFB464793">Journal of Natural History</mods:title>
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<mods:part id="88A28E25FAD59D3E53178C84DF305640">
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<mods:date id="87EA1680A83B5314B38B91A435547F84">2021</mods:date>
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<mods:number id="7E2AF24B1E576ECECBDF87F781D5B769">2021-07-21</mods:number>
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<mods:number id="545FDF32C8830BD694C3EB8F8FCF2D77">17 - 18</mods:number>
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<heading id="B65A81BDFF94FF86FE66FF5CFC52FF1C" box="[385,931,142,168]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="1144" reason="2">
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<taxonomicName id="2AAD4D52FF94FF86FE66FF5CFD1EFF1C" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[385,751,142,168]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="1144" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
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<emphasis id="DFD9EAC3FF94FF86FE66FF5CFE12FF1C" bold="true" box="[385,483,142,168]" italics="true" pageId="13" pageNumber="1144">Epeorus</emphasis>
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(
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<emphasis id="DFD9EAC3FF94FF86FE14FF5CFD8FFF1C" bold="true" box="[499,638,142,168]" italics="true" pageId="13" pageNumber="1144">Proepeorus</emphasis>
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)
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<emphasis id="DFD9EAC3FF94FF86FD69FF5CFD1EFF1C" bold="true" box="[654,751,142,168]" italics="true" pageId="13" pageNumber="1144">falcatus</emphasis>
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<taxonomicNameLabel id="C4EA57B8FF94FF86FD10FF5CFCA8FF1C" box="[759,857,142,168]" pageId="13" pageNumber="1144" rank="species">sp. nov.</taxonomicNameLabel>
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Zhou
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</emphasis>
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</paragraph>
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<subSubSection id="A5B7655AFF94FF86FED3FF63FC1EFF7F" box="[308,1007,177,203]" pageId="13" pageNumber="1144" type="description">
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<paragraph id="ED1236D1FF94FF86FED3FF63FC1EFF7F" blockId="13.[308,1007,142,203]" box="[308,1007,177,203]" pageId="13" pageNumber="1144">
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<figureCitation id="75962A54FF94FF86FED3FF63FE64FF7F" box="[308,405,177,203]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStart-2="Figure 9" captionStartId-0="13.[160,225,1645,1668]" captionStartId-1="14.[160,225,999,1022]" captionStartId-2="15.[160,225,913,936]" captionTargetBox-0="[163,1153,809,1614]" captionTargetBox-1="[173,1147,142,968]" captionTargetBox-2="[164,1151,143,864]" captionTargetId-0="figure-157@13.[163,1153,809,1614]" captionTargetId-1="figure-212@14.[163,1153,142,968]" captionTargetId-2="figure-239@15.[163,1153,142,881]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." captionText-1="Figure 8. Mouthparts of Epeorus (Proepeorus) falcatus sp. nov. (digital photos): (a) Labrum; (b) Left mandible; (c) Right mandible; (d) Hypopharynx; (e) Maxillae; (f) Labium." captionText-2="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi-0="http://doi.org/10.5281/zenodo.5466041" figureDoi-1="http://doi.org/10.5281/zenodo.5466043" figureDoi-2="http://doi.org/10.5281/zenodo.5466047" httpUri-0="https://zenodo.org/record/5466041/files/figure.png" httpUri-1="https://zenodo.org/record/5466043/files/figure.png" httpUri-2="https://zenodo.org/record/5466047/files/figure.png" pageId="13" pageNumber="1144">Figs 7–9</figureCitation>
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,
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<figureCitation id="75962A54FF94FF86FE44FF63FDC2FF7F" box="[419,563,177,203]" captionStart="Figure 10" captionStartId="16.[160,225,1118,1141]" captionTargetBox="[163,1153,142,1086]" captionTargetId="figure-136@16.[163,1153,142,1086]" captionTargetPageId="16" captionText="Figure 10. Male adults of two new species: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) General view, (b) Wings; (c, d) Epeorus (Proepeorus) falcatus sp. nov., (c) General view, (d) Wings; (e, f) Thorax of Epeorus (Proepeorus) gibbus sp. nov., (e) Dorsal view, (f) Ventral view; (g, h) Thorax of Epeorus (Proepeorus) falcatus sp. nov., (g) Dorsal view, (h) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5466049" httpUri="https://zenodo.org/record/5466049/files/figure.png" pageId="13" pageNumber="1144">10C–D, G–H</figureCitation>
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,
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<figureCitation id="75962A54FF94FF86FDD9FF63FD7AFF7F" box="[574,651,177,203]" captionStart="Figure 11" captionStartId="17.[160,225,668,691]" captionTargetBox="[163,1153,142,637]" captionTargetId="figure-303@17.[163,1153,142,637]" captionTargetPageId="17" captionText="Figure 11. Abdominal terga of imagoes of two new species: (a–d) Epeorus (Proepeorus) gibbus sp. nov., (a) Dorsal view of male, (b) Ventral view of male, (c) Dorsal view of female, (d) Ventral view of female; (e–h) Epeorus (Proepeorus) falcatus sp. nov., (e) Dorsal view of male, (f) Ventral view of male, (g) Dorsal view of female, (h) Ventral view of female; Abbreviations: ma, medio-anterior myo-sigillum; mp, medio-posterior myo-sigillum." figureDoi="http://doi.org/10.5281/zenodo.5466051" httpUri="https://zenodo.org/record/5466051/files/figure.png" pageId="13" pageNumber="1144">11E–H</figureCitation>
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,
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<figureCitation id="75962A54FF94FF86FD71FF63FD33FF7F" box="[662,706,177,203]" captionStart="Figure 12" captionStartId="18.[160,225,1161,1184]" captionTargetBox="[163,1153,142,1130]" captionTargetId="figure-131@18.[163,1153,142,1130]" captionTargetPageId="18" captionText="Figure 12. Genitalia of male: (a) Epeorus (Proepeorus) gibbus sp. nov. (ventral view); (b) Epeorus (Proepeorus) falcatus sp. nov. (ventral view); (c) Epeorus (Proepeorus) nipponicus (ventral view); (d) Epeorus (Proepeorus) aculeatus (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5466053" httpUri="https://zenodo.org/record/5466053/files/figure.png" pageId="13" pageNumber="1144">12B</figureCitation>
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,
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<figureCitation id="75962A54FF94FF86FD37FF63FCEFFF7F" box="[720,798,177,203]" captionStart="Figure 13" captionStartId="19.[160,225,918,941]" captionTargetBox="[166,1150,143,884]" captionTargetId="figure-194@19.[163,1153,142,887]" captionTargetPageId="19" captionText="Figure 13. Genitalia of male: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) Ventral view, (b) Dorsal view; (c) Epeorus (Proepeorus) nipponicus (ventral view); (d, e) Epeorus (Proepeorus) falcatus sp. nov., (d) Ventral view, (e) Dorsal view; (f) Epeorus (Proepeorus) aculeatus (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5466055" httpUri="https://zenodo.org/record/5466055/files/figure.png" pageId="13" pageNumber="1144">13D–E</figureCitation>
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,
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<figureCitation id="75962A54FF94FF86FCCEFF63FC89FF7F" box="[809,888,177,203]" captionStart="Figure 14" captionStartId="20.[160,225,1164,1187]" captionTargetBox="[163,1153,142,1122]" captionTargetId="figure-177@20.[163,1153,142,1132]" captionTargetPageId="20" captionText="Figure 14. Subgenital plate and sternum IX of female: (a, b) Epeorus (Proepeorus) gibbus sp. nov.; (c, d) Epeorus (Proepeorus) falcatus sp. nov.; (e, f) Epeorus (Proepeorus) nipponicus." figureDoi="http://doi.org/10.5281/zenodo.5466059" httpUri="https://zenodo.org/record/5466059/files/figure.png" pageId="13" pageNumber="1144">14C–D</figureCitation>
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,
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<figureCitation id="75962A54FF94FF86FC63FF63FC42FF7F" box="[900,947,177,203]" captionStart="Figure 15" captionStartId="21.[160,225,946,969]" captionTargetBox="[163,1153,187,914]" captionTargetId="figure-223@21.[163,1153,142,914]" captionTargetPageId="21" captionText="Figure 15. Mouthparts of Epeorus (Proepeorus) spp.: (a–f) labrum, (a) E. (P.) nipponicus; (b) E. (P.) unispinosus; (c) E. (P.) bifurcatus; (d) E. (P.) aculeatus; (e) E. (P.) rhithralis; (f) E. (P.) bispinosus; (g–i) base of apical tooth complex of maxilla, (g) E. (P.) gibbus sp. nov.; (h) E. (P.) falcatus sp. nov.; (i) E. (P.) nipponicus." figureDoi="http://doi.org/10.5281/zenodo.5466061" httpUri="https://zenodo.org/record/5466061/files/figure.png" pageId="13" pageNumber="1144">15H</figureCitation>
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,
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<figureCitation id="75962A54FF94FF86FC26FF63FC1EFF7F" box="[961,1007,177,203]" captionStart="Figure 16" captionStartId="22.[160,225,917,940]" captionTargetBox="[166,1153,142,885]" captionTargetId="figure-264@22.[163,1153,142,885]" captionTargetPageId="22" captionText="Figure 16. Incisors of left mandible in ventral view: (a) Epeorus (Epeorus) melli; (b) Epeorus nguyeni; (c) Epeorus (Proepeorus) nipponicus; (d) Epeorus (Belovius) pellucidus; (e) Epeorus (Proepeorus) unispinosus; (f) Epeorus (Proepeorus) gibbus sp. nov.; (g) Epeorus (Iron) alexantri; (h) Epeorus (Proepeorus) falcatus sp. nov.; (i) Epeorus (Proepeorus) aculeatus; (j) Epeorus (Proepeorus) bifurcatus; (k) Epeorus (Proepeorus) bispinosus; (l) Epeorus (Proepeorus) rhithralis." figureDoi="http://doi.org/10.5281/zenodo.5466065" httpUri="https://zenodo.org/record/5466065/files/figure.png" pageId="13" pageNumber="1144">16H</figureCitation>
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<emphasis id="DFD9EAC3FF94FF86FF47FEDFFE7BFE93" bold="true" box="[160,394,269,295]" italics="true" pageId="13" pageNumber="1144">Materials examined</emphasis>
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, collected by
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<collectorName id="40585307FF94FF86FDD1FE81FD16FED8" box="[566,743,339,364]" pageId="13" pageNumber="1144">Zhen-Xing MA</collectorName>
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, Xu-Hong-Yi
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<collectingDate id="8957E9F9FF94FF86FC0BFE81FB71FED8" box="[1004,1152,339,364]" pageId="13" pageNumber="1144" value="2019-08-01" valueMax="2019-08-05" valueMin="2019-08-01">1–5-VIII-2019</collectingDate>
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:
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imagoes
|
||
<specimenCount id="FBABFD58FF94FF86FE3FFEA4FDCAFE3B" box="[472,571,374,399]" pageId="13" pageNumber="1144" type="female">2 female</specimenCount>
|
||
imagoes 10 nymphs, same as
|
||
<typeStatus id="32168873FF94FF86FC7CFEA4FBF3FE3B" box="[923,1026,374,399]" pageId="13" pageNumber="1144" type="holotype">holotype</typeStatus>
|
||
; 5 nymphs, Huaping Nature Reserve,
|
||
<collectingRegion id="2F69F833FF94FF86FE27FE4BFD79FE06" box="[448,648,409,434]" country="China" name="Guangxi" pageId="13" pageNumber="1144">Guangxi Province</collectingRegion>
|
||
, collected by Zhen-Xing MA,
|
||
<date id="99131011FF94FF86FC31FE4BFB71FE06" box="[982,1152,409,434]" pageId="13" pageNumber="1144" value="2020-08-20" valueMax="2020-08-21" valueMin="2020-08-20">20-21-VIII-2020</date>
|
||
; 30 nymphs, Jintong Mountain, Shaoyang City,
|
||
<collectingRegion id="2F69F833FF94FF86FD50FE69FC9FFE60" box="[695,878,443,468]" country="China" name="Hunan" pageId="13" pageNumber="1144">Hunan Province</collectingRegion>
|
||
, collected by Zhen-Xing MA,
|
||
<date id="99131011FF94FF86FF34FE0CFEABFE43" box="[211,346,478,503]" pageId="13" pageNumber="1144" value="2020-08-23">23-VIII-2020</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF94FF86FF47FDFAFEC5FD3C" blockId="13.[160,1156,552,752]" pageId="13" pageNumber="1144">
|
||
<emphasis id="DFD9EAC3FF94FF86FF47FDFAFE7CFDF6" bold="true" box="[160,397,552,578]" italics="true" pageId="13" pageNumber="1144">Nymph (in ethanol).</emphasis>
|
||
Body length 10.0–12.0 mm (female),
|
||
<quantity id="2A559B34FF94FF86FCA8FDFBFC26FDF6" box="[847,983,553,578]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="13" pageNumber="1144" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.5–9.5 mm</quantity>
|
||
(male), caudal filaments 14.0–15.0 mm (female), 11.0 mm (male). Body generally pale to brown (
|
||
<figureCitation id="75962A54FF94FF86FF4FFDBDFEF8FD3C" box="[168,265,623,648]" captionStart="Figure 7" captionStartId="13.[160,225,1645,1668]" captionTargetBox="[163,1153,809,1614]" captionTargetId="figure-157@13.[163,1153,809,1614]" captionTargetPageId="13" captionText="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="13" pageNumber="1144">Figure 7</figureCitation>
|
||
(a)).
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF94FF84FF47FD42FE2EFB1C" blockId="13.[160,1156,552,752]" lastBlockId="15.[160,1156,1098,1712]" lastPageId="15" lastPageNumber="1146" pageId="13" pageNumber="1144">
|
||
<emphasis id="DFD9EAC3FF94FF86FF47FD42FF13FD1E" box="[160,226,656,682]" italics="true" pageId="13" pageNumber="1144">Head.</emphasis>
|
||
ellipsoid, head width/length ratio 1.39, anterior and lateral margins evenly convex, posterior margin slightly convex (nearly straight); anterior margin with a row of dense setae directed anteriorly, posterior margin with two tufts of setae. Head generally pale, with median brown or dark grey irregular markings (
|
||
<figureCitation id="75962A54FF97FF85FC53FB88FBEEFBC7" box="[948,1055,1114,1139]" captionStart="Figure 7" captionStartId="13.[160,225,1645,1668]" captionTargetBox="[163,1153,809,1614]" captionTargetId="figure-157@13.[163,1153,809,1614]" captionTargetPageId="13" captionText="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="14" pageNumber="1145">Figures 7</figureCitation>
|
||
(c) and 9 (a)).
|
||
<emphasis id="DFD9EAC3FF97FF85FF30FBAFFEC4FB22" bold="true" box="[215,309,1149,1174]" pageId="14" pageNumber="1145">Labrum</emphasis>
|
||
: lateral margins straight and widened anteriorly, anterior margin with a distinct median notch, dorsal surface with 6 bristles antero-medially and 2 bristles near each side of anterolateral margin. Ventral surface with a row of bristles along lateral margin extend to anterior margin and brush of fine setae medially (
|
||
<figureCitation id="75962A54FF97FF85FBFFFB37FB75FB4A" box="[1048,1156,1253,1278]" captionStart="Figure 8" captionStartId="14.[160,225,999,1022]" captionTargetBox="[173,1147,142,968]" captionTargetId="figure-212@14.[163,1153,142,968]" captionTargetPageId="14" captionText="Figure 8. Mouthparts of Epeorus (Proepeorus) falcatus sp. nov. (digital photos): (a) Labrum; (b) Left mandible; (c) Right mandible; (d) Hypopharynx; (e) Maxillae; (f) Labium." figureDoi="http://doi.org/10.5281/zenodo.5466043" httpUri="https://zenodo.org/record/5466043/files/figure.png" pageId="14" pageNumber="1145">Figures 8</figureCitation>
|
||
(a) and 9(b)).
|
||
<emphasis id="DFD9EAC3FF97FF85FEA0FAD5FE37FA94" bold="true" box="[327,454,1287,1312]" pageId="14" pageNumber="1145">Mandibles</emphasis>
|
||
: outer incisor with 2 (left mandible) or 3 (right mandible) apical lobes and serrated margins, inner incisor with 2 (right mandible) or 3 (left mandible) apical lobes and serrated outer margins. Prostheca represented by a tuft of plumose setae (
|
||
<figureCitation id="75962A54FF97FF85FE9FFABDFE12FA3C" box="[376,483,1391,1416]" captionStart="Figure 8" captionStartId="14.[160,225,999,1022]" captionTargetBox="[173,1147,142,968]" captionTargetId="figure-212@14.[163,1153,142,968]" captionTargetPageId="14" captionText="Figure 8. Mouthparts of Epeorus (Proepeorus) falcatus sp. nov. (digital photos): (a) Labrum; (b) Left mandible; (c) Right mandible; (d) Hypopharynx; (e) Maxillae; (f) Labium." figureDoi="http://doi.org/10.5281/zenodo.5466043" httpUri="https://zenodo.org/record/5466043/files/figure.png" pageId="14" pageNumber="1145">Figures 8</figureCitation>
|
||
(b,c) and 9(c,d)).
|
||
<emphasis id="DFD9EAC3FF97FF85FD42FABDFCFAFA3C" bold="true" box="[677,779,1391,1416]" pageId="14" pageNumber="1145">Maxillae</emphasis>
|
||
: with 1 fimbriated setae at base of apex of maxilla (
|
||
<figureCitation id="75962A54FF97FF85FE6EFA40FE0AFA1F" box="[393,507,1426,1451]" captionStart="Figure 15" captionStartId="21.[160,225,946,969]" captionTargetBox="[163,1153,187,914]" captionTargetId="figure-223@21.[163,1153,142,914]" captionTargetPageId="21" captionText="Figure 15. Mouthparts of Epeorus (Proepeorus) spp.: (a–f) labrum, (a) E. (P.) nipponicus; (b) E. (P.) unispinosus; (c) E. (P.) bifurcatus; (d) E. (P.) aculeatus; (e) E. (P.) rhithralis; (f) E. (P.) bispinosus; (g–i) base of apical tooth complex of maxilla, (g) E. (P.) gibbus sp. nov.; (h) E. (P.) falcatus sp. nov.; (i) E. (P.) nipponicus." figureDoi="http://doi.org/10.5281/zenodo.5466061" httpUri="https://zenodo.org/record/5466061/files/figure.png" pageId="14" pageNumber="1145">Figure 15</figureCitation>
|
||
(h)); basal segment of maxillary palp slightly expanded proximally and with a tuft of (4–5) distal bristles on outer margin; apical segment 1.35× length of basal segment, with slender basal half and extremely expanded distal half (
|
||
<figureCitation id="75962A54FF97FF85FF01FA28FEA0F9A7" box="[230,337,1530,1555]" captionStart="Figure 8" captionStartId="14.[160,225,999,1022]" captionTargetBox="[173,1147,142,968]" captionTargetId="figure-212@14.[163,1153,142,968]" captionTargetPageId="14" captionText="Figure 8. Mouthparts of Epeorus (Proepeorus) falcatus sp. nov. (digital photos): (a) Labrum; (b) Left mandible; (c) Right mandible; (d) Hypopharynx; (e) Maxillae; (f) Labium." figureDoi="http://doi.org/10.5281/zenodo.5466043" httpUri="https://zenodo.org/record/5466043/files/figure.png" pageId="14" pageNumber="1145">Figure 8</figureCitation>
|
||
(e)).
|
||
<emphasis id="DFD9EAC3FF97FF85FE68FA28FDC4F9A7" bold="true" box="[399,565,1530,1555]" pageId="14" pageNumber="1145">Hypopharynx</emphasis>
|
||
: lingua with anterolateral lobes (
|
||
<figureCitation id="75962A54FF97FF85FC0AFA28FBA9F9A7" box="[1005,1112,1530,1555]" captionStart="Figure 8" captionStartId="14.[160,225,999,1022]" captionTargetBox="[173,1147,142,968]" captionTargetId="figure-212@14.[163,1153,142,968]" captionTargetPageId="14" captionText="Figure 8. Mouthparts of Epeorus (Proepeorus) falcatus sp. nov. (digital photos): (a) Labrum; (b) Left mandible; (c) Right mandible; (d) Hypopharynx; (e) Maxillae; (f) Labium." figureDoi="http://doi.org/10.5281/zenodo.5466043" httpUri="https://zenodo.org/record/5466043/files/figure.png" pageId="14" pageNumber="1145">Figure 8</figureCitation>
|
||
(d)).
|
||
<emphasis id="DFD9EAC3FF97FF85FF47F9CEFF0AF981" bold="true" box="[160,251,1564,1589]" pageId="14" pageNumber="1145">Labium</emphasis>
|
||
: with narrow U-shaped separation between glossae; glossa triangular, with a shallow subapical projection, dorsum with a row of longitudinal dense setae, both inner margin and outer margin of glossa with a row of bristles; apical segment of labial palpi slightly shorter than basal segment, apical/basal segment length ratio 0.86 (
|
||
<figureCitation id="75962A54FF96FF84FF05FB5DFEBEFB1C" box="[226,335,1167,1192]" captionStart="Figure 8" captionStartId="14.[160,225,999,1022]" captionTargetBox="[173,1147,142,968]" captionTargetId="figure-212@14.[163,1153,142,968]" captionTargetPageId="14" captionText="Figure 8. Mouthparts of Epeorus (Proepeorus) falcatus sp. nov. (digital photos): (a) Labrum; (b) Left mandible; (c) Right mandible; (d) Hypopharynx; (e) Maxillae; (f) Labium." figureDoi="http://doi.org/10.5281/zenodo.5466043" httpUri="https://zenodo.org/record/5466043/files/figure.png" pageId="15" pageNumber="1146">Figures 8</figureCitation>
|
||
(f) and 9(e)).
|
||
</paragraph>
|
||
<caption id="B9D26659FF94FF86FF47F9BFFD44F90B" ID-DOI="http://doi.org/10.5281/zenodo.5466041" ID-Zenodo-Dep="5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="13" pageNumber="1144" startId="13.[160,225,1645,1668]" targetBox="[163,1153,809,1614]" targetPageId="13">
|
||
<paragraph id="ED1236D1FF94FF86FF47F9BFFD44F90B" blockId="13.[160,1155,1645,1727]" pageId="13" pageNumber="1144">
|
||
<emphasis id="DFD9EAC3FF94FF86FF47F9BFFF0FF930" bold="true" box="[160,254,1645,1668]" pageId="13" pageNumber="1144">Figure 7.</emphasis>
|
||
Nymphal habitus and structures of
|
||
<taxonomicName id="2AAD4D52FF94FF86FD91F9BFFC69F930" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[630,920,1645,1668]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="1144" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF94FF86FD91F9BFFD31F930" box="[630,704,1645,1668]" italics="true" pageId="13" pageNumber="1144">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF94FF86FD35F9BFFCCDF930" box="[722,828,1645,1668]" italics="true" pageId="13" pageNumber="1144">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF94FF86FCAAF9BFFC69F930" box="[845,920,1645,1668]" italics="true" pageId="13" pageNumber="1144">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF94FF86FC45F9BFFC09F930" box="[930,1016,1645,1668]" pageId="13" pageNumber="1144" rank="species">
|
||
<emphasis id="DFD9EAC3FF94FF86FC45F9BFFC02F930" bold="true" box="[930,1011,1645,1668]" pageId="13" pageNumber="1144">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9D26659FF97FF85FF47FC35FC96FBA8" ID-DOI="http://doi.org/10.5281/zenodo.5466043" ID-Zenodo-Dep="5466043" httpUri="https://zenodo.org/record/5466043/files/figure.png" pageId="14" pageNumber="1145" startId="14.[160,225,999,1022]" targetBox="[173,1147,142,968]" targetPageId="14">
|
||
<paragraph id="ED1236D1FF97FF85FF47FC35FC96FBA8" blockId="14.[160,1155,999,1052]" pageId="14" pageNumber="1145">
|
||
<emphasis id="DFD9EAC3FF97FF85FF47FC35FF0AFC4A" bold="true" box="[160,251,999,1022]" pageId="14" pageNumber="1145">Figure 8.</emphasis>
|
||
Mouthparts of
|
||
<taxonomicName id="2AAD4D52FF97FF85FE7DFC35FD46FC4A" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[410,695,999,1022]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="1145" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF97FF85FE7DFC35FE15FC4A" box="[410,484,999,1022]" italics="true" pageId="14" pageNumber="1145">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF97FF85FE13FC35FDAFFC4A" box="[500,606,999,1022]" italics="true" pageId="14" pageNumber="1145">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF97FF85FD8BFC35FD46FC4A" box="[620,695,999,1022]" italics="true" pageId="14" pageNumber="1145">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF97FF85FD58FC35FCE2FC4A" box="[703,787,999,1022]" pageId="14" pageNumber="1145" rank="species">
|
||
<emphasis id="DFD9EAC3FF97FF85FD58FC35FCFFFC4A" bold="true" box="[703,782,999,1022]" pageId="14" pageNumber="1145">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(digital photos): (a) Labrum; (b) Left mandible; (c) Right mandible; (d) Hypopharynx; (e) Maxillae; (f) Labium.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B9D26659FF96FF84FF47FC43FD4BFBB4" ID-DOI="http://doi.org/10.5281/zenodo.5466047" ID-Zenodo-Dep="5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="15" pageNumber="1146" startId="15.[160,225,913,936]" targetBox="[164,1151,143,864]" targetPageId="15">
|
||
<paragraph id="ED1236D1FF96FF84FF47FC43FD4BFBB4" blockId="15.[160,1155,913,1024]" pageId="15" pageNumber="1146">
|
||
<emphasis id="DFD9EAC3FF96FF84FF47FC43FF08FC1C" bold="true" box="[160,249,913,936]" pageId="15" pageNumber="1146">Figure 9.</emphasis>
|
||
Nymph of
|
||
<taxonomicName id="2AAD4D52FF96FF84FE8FFC43FD70FC1C" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[360,641,913,936]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="1146" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF96FF84FE8FFC43FE43FC1C" box="[360,434,913,936]" italics="true" pageId="15" pageNumber="1146">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF96FF84FE58FC43FDD8FC1C" box="[447,553,913,936]" italics="true" pageId="15" pageNumber="1146">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF96FF84FDD1FC43FD70FC1C" box="[566,641,913,936]" italics="true" pageId="15" pageNumber="1146">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF96FF84FD60FC43FD29FC1C" box="[647,728,913,936]" pageId="15" pageNumber="1146" rank="species">
|
||
<emphasis id="DFD9EAC3FF96FF84FD60FC43FD22FC1C" bold="true" box="[647,723,913,936]" pageId="15" pageNumber="1146">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED1236D1FF96FF84FF47FB63FDAAFA2F" blockId="15.[160,1156,1098,1712]" pageId="15" pageNumber="1146">
|
||
<emphasis id="DFD9EAC3FF96FF84FF47FB63FF04FB7F" box="[160,245,1201,1227]" italics="true" pageId="15" pageNumber="1146">Thorax.</emphasis>
|
||
pale to brown with irregular markings. Femora pale, with a medial dark spot (indistinct in some individuals) and irregular brown markings, outer margin with a row of long setae and sparse short setae (
|
||
<figureCitation id="75962A54FF96FF84FDC6FB25FD8FFAA4" box="[545,638,1271,1296]" captionStart="Figure 7" captionStartId="13.[160,225,1645,1668]" captionTargetBox="[163,1153,809,1614]" captionTargetId="figure-157@13.[163,1153,809,1614]" captionTargetPageId="13" captionText="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="15" pageNumber="1146">Figure 7</figureCitation>
|
||
(f)), dorsum with scattered blunt spatulate and arrow-like setae (
|
||
<figureCitation id="75962A54FF96FF84FEBAFAC8FE48FA87" box="[349,441,1306,1331]" captionStart="Figure 9" captionStartId="15.[160,225,913,936]" captionTargetBox="[164,1151,143,864]" captionTargetId="figure-239@15.[163,1153,142,881]" captionTargetPageId="15" captionText="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="15" pageNumber="1146">Figure 9</figureCitation>
|
||
(f)). Proximal and medial portion of tibiae slightly darkened, base and apex of tarsi darkened (
|
||
<figureCitation id="75962A54FF96FF84FE15FAEEFDA5FAE1" box="[498,596,1340,1365]" captionStart="Figure 7" captionStartId="13.[160,225,1645,1668]" captionTargetBox="[163,1153,809,1614]" captionTargetId="figure-157@13.[163,1153,809,1614]" captionTargetPageId="13" captionText="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="15" pageNumber="1146">Figure 7</figureCitation>
|
||
(f)). Forefemora/foretibiae length ratio 0.96, mid femora/mid tibiae length ratio 1.08, hind femora/hind tibiae length ratio 1.22. Claws with 3 subapical denticles (
|
||
<figureCitation id="75962A54FF96FF84FE34FA50FDC1FA2F" box="[467,560,1410,1435]" captionStart="Figure 9" captionStartId="15.[160,225,913,936]" captionTargetBox="[164,1151,143,864]" captionTargetId="figure-239@15.[163,1153,142,881]" captionTargetPageId="15" captionText="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="15" pageNumber="1146">Figure 9</figureCitation>
|
||
(g)).
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF96FF9BFF47FA76FEE7FA33" blockId="15.[160,1156,1098,1712]" lastBlockId="16.[160,1156,1321,1415]" lastPageId="16" lastPageNumber="1147" pageId="15" pageNumber="1146">
|
||
<emphasis id="DFD9EAC3FF96FF84FF47FA76FEE4FA0A" box="[160,277,1444,1470]" italics="true" pageId="15" pageNumber="1146">Abdomen.</emphasis>
|
||
Pale with brown markings. Terga segments II–IX with paired small submedian spines and a median longitudinal dark stripe. Terga II–IV with paired submedian irregular pale spots. Each tergum with transverse brown band along anterior margin (
|
||
<figureCitation id="75962A54FF96FF84FEE6F9DEFE93F991" box="[257,354,1548,1573]" captionStart="Figure 7" captionStartId="13.[160,225,1645,1668]" captionTargetBox="[163,1153,809,1614]" captionTargetId="figure-157@13.[163,1153,809,1614]" captionTargetPageId="13" captionText="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="15" pageNumber="1146">Figure 7</figureCitation>
|
||
(a,d)). Posterior margins of terga with blunt denticles and mixed with smaller round pointed denticles (
|
||
<figureCitation id="75962A54FF96FF84FDD2F9FDFD6BF9FC" box="[565,666,1583,1608]" captionStart="Figure 9" captionStartId="15.[160,225,913,936]" captionTargetBox="[164,1151,143,864]" captionTargetId="figure-239@15.[163,1153,142,881]" captionTargetPageId="15" captionText="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="15" pageNumber="1146">Figure 9</figureCitation>
|
||
(h)). Sterna colourless (sterna VII–IX with paired brown stripes in some individuals) (
|
||
<figureCitation id="75962A54FF96FF84FD2FF980FCC0F9DF" box="[712,817,1618,1643]" captionStart="Figure 7" captionStartId="13.[160,225,1645,1668]" captionTargetBox="[163,1153,809,1614]" captionTargetId="figure-157@13.[163,1153,809,1614]" captionTargetPageId="13" captionText="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="15" pageNumber="1146">Figure 7</figureCitation>
|
||
(e)). Gills: gill I moderately expanded anteriorly and narrowed apically, outer margin with a row of fine setae (
|
||
<figureCitation id="75962A54FF96FF84FF4FF945FEFDF904" box="[168,268,1687,1712]" captionStart="Figure 9" captionStartId="15.[160,225,913,936]" captionTargetBox="[164,1151,143,864]" captionTargetId="figure-239@15.[163,1153,142,881]" captionTargetPageId="15" captionText="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="15" pageNumber="1146">Figure 9</figureCitation>
|
||
(i)); gills II–
|
||
<collectionCode id="8BBCAE14FF96FF84FE6DF945FE52F904" box="[394,419,1687,1712]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="1146">VI</collectionCode>
|
||
similar, distal half of costal margin with a row of long setae (
|
||
<figureCitation id="75962A54FF89FF9BFF4FFAFBFEF8FAF6" box="[168,265,1321,1346]" captionStart="Figure 9" captionStartId="15.[160,225,913,936]" captionTargetBox="[164,1151,143,864]" captionTargetId="figure-239@15.[163,1153,142,881]" captionTargetPageId="15" captionText="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="16" pageNumber="1147">Figure 9</figureCitation>
|
||
(j)); gill VII oval, with long anal rib, slightly curved and without distinct fold (
|
||
<figureCitation id="75962A54FF89FF9BFF4FFA9EFEE7FAD1" box="[168,278,1356,1381]" captionStart="Figure 7" captionStartId="13.[160,225,1645,1668]" captionTargetBox="[163,1153,809,1614]" captionTargetId="figure-157@13.[163,1153,809,1614]" captionTargetPageId="13" captionText="Figure 7. Nymphal habitus and structures of Epeorus (Proepeorus) falcatus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466041" httpUri="https://zenodo.org/record/5466041/files/figure.png" pageId="16" pageNumber="1147">Figures 7</figureCitation>
|
||
(e) and 9(k)). Cerci pale to brown, with a row of dorsal setae along basal segments.
|
||
</paragraph>
|
||
<caption id="B9D26659FF89FF9BFF47FB8CFCE4FB79" ID-DOI="http://doi.org/10.5281/zenodo.5466049" ID-Zenodo-Dep="5466049" httpUri="https://zenodo.org/record/5466049/files/figure.png" pageId="16" pageNumber="1147" startId="16.[160,225,1118,1141]" targetBox="[163,1153,142,1086]" targetPageId="16">
|
||
<paragraph id="ED1236D1FF89FF9BFF47FB8CFCE4FB79" blockId="16.[160,1156,1118,1229]" pageId="16" pageNumber="1147">
|
||
<emphasis id="DFD9EAC3FF89FF9BFF47FB8CFEF8FBC1" bold="true" box="[160,265,1118,1141]" pageId="16" pageNumber="1147">Figure 10.</emphasis>
|
||
Male adults of two new species: (a, b)
|
||
<taxonomicName id="2AAD4D52FF89FF9BFD7DFB8CFC5DFBC1" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[666,940,1118,1141]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="1147" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF89FF9BFD7DFB8CFD15FBC1" box="[666,740,1118,1141]" italics="true" pageId="16" pageNumber="1147">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF89FF9BFD13FB8CFCAFFBC1" box="[756,862,1118,1141]" italics="true" pageId="16" pageNumber="1147">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF89FF9BFC8AFB8CFC5DFBC1" box="[877,940,1118,1141]" italics="true" pageId="16" pageNumber="1147">gibbus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF89FF9BFC52FB8CFBF9FBC1" box="[949,1032,1118,1141]" pageId="16" pageNumber="1147" rank="species">
|
||
<emphasis id="DFD9EAC3FF89FF9BFC52FB8CFBF2FBC1" bold="true" box="[949,1027,1118,1141]" pageId="16" pageNumber="1147">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, (a) General view, (b) Wings; (c, d)
|
||
<taxonomicName id="2AAD4D52FF89FF9BFE93FBA9FD7AFB26" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[372,651,1147,1170]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="1147" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF89FF9BFE93FBA9FE4FFB26" box="[372,446,1147,1170]" italics="true" pageId="16" pageNumber="1147">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF89FF9BFE2DFBA9FDC5FB26" box="[458,564,1147,1170]" italics="true" pageId="16" pageNumber="1147">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF89FF9BFDA7FBA9FD7AFB26" box="[576,651,1147,1170]" italics="true" pageId="16" pageNumber="1147">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF89FF9BFD77FBA9FD11FB26" box="[656,736,1147,1170]" pageId="16" pageNumber="1147" rank="species">
|
||
<emphasis id="DFD9EAC3FF89FF9BFD77FBA9FD2AFB26" bold="true" box="[656,731,1147,1170]" pageId="16" pageNumber="1147">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, (c) General view, (d) Wings; (e, f) Thorax of
|
||
<taxonomicName id="2AAD4D52FF89FF9BFF47FB4AFE49FB1B" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[160,440,1176,1199]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="1147" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF89FF9BFF47FB4AFF1BFB1B" box="[160,234,1176,1199]" italics="true" pageId="16" pageNumber="1147">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF89FF9BFF1AFB4AFE96FB1B" box="[253,359,1176,1199]" italics="true" pageId="16" pageNumber="1147">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF89FF9BFE9EFB4AFE49FB1B" box="[377,440,1176,1199]" italics="true" pageId="16" pageNumber="1147">gibbus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF89FF9BFE23FB4AFDEBFB1B" box="[452,538,1176,1199]" pageId="16" pageNumber="1147" rank="species">
|
||
<emphasis id="DFD9EAC3FF89FF9BFE23FB4AFDE4FB1B" bold="true" box="[452,533,1176,1199]" pageId="16" pageNumber="1147">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, (e) Dorsal view, (f) Ventral view; (g, h) Thorax of
|
||
<taxonomicName id="2AAD4D52FF89FF9BFBDEFB4AFE9BFB79" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="1147" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF89FF9BFBDEFB4AFB72FB1B" box="[1081,1155,1176,1199]" italics="true" pageId="16" pageNumber="1147">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF89FF9BFF40FB64FEE0FB79" box="[167,273,1206,1229]" italics="true" pageId="16" pageNumber="1147">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF89FF9BFEF8FB64FE9BFB79" box="[287,362,1206,1229]" italics="true" pageId="16" pageNumber="1147">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DFD9EAC3FF89FF9BFE97FB64FE35FB79" bold="true" box="[368,452,1206,1229]" pageId="16" pageNumber="1147">
|
||
<taxonomicNameLabel id="C4EA57B8FF89FF9BFE97FB64FE35FB79" box="[368,452,1206,1229]" pageId="16" pageNumber="1147" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, (g) Dorsal view, (h) Ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED1236D1FF89FF9BFF47FA66FC21FA44" blockId="16.[160,1156,1460,1728]" pageId="16" pageNumber="1147">
|
||
<emphasis id="DFD9EAC3FF89FF9BFF47FA66FE38FA7A" bold="true" box="[160,457,1460,1486]" italics="true" pageId="16" pageNumber="1147">Male imago (in ethanol).</emphasis>
|
||
Body length 9.0–10.0 mm, forewing 9.5–10.0 mm, hindwing 3.0–
|
||
<quantity id="2A559B34FF89FF9BFF2AFA05FED7FA44" box="[205,294,1495,1520]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="1147" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, cerci 28.0–34.0 mm. Body generally brownish (
|
||
<figureCitation id="75962A54FF89FF9BFCD9FA05FC59FA44" box="[830,936,1495,1520]" captionStart="Figure 10" captionStartId="16.[160,225,1118,1141]" captionTargetBox="[163,1153,142,1086]" captionTargetId="figure-136@16.[163,1153,142,1086]" captionTargetPageId="16" captionText="Figure 10. Male adults of two new species: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) General view, (b) Wings; (c, d) Epeorus (Proepeorus) falcatus sp. nov., (c) General view, (d) Wings; (e, f) Thorax of Epeorus (Proepeorus) gibbus sp. nov., (e) Dorsal view, (f) Ventral view; (g, h) Thorax of Epeorus (Proepeorus) falcatus sp. nov., (g) Dorsal view, (h) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5466049" httpUri="https://zenodo.org/record/5466049/files/figure.png" pageId="16" pageNumber="1147">Figure 10</figureCitation>
|
||
(c)).
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF89FF9BFF47FA2BFC10F982" blockId="16.[160,1156,1460,1728]" pageId="16" pageNumber="1147">
|
||
<emphasis id="DFD9EAC3FF89FF9BFF47FA2BFF13F9A7" box="[160,226,1529,1555]" italics="true" pageId="16" pageNumber="1147">Head.</emphasis>
|
||
Face fold dark brown, antennae brown, median area brown. Compound eyes pale (dark brown when alive), contiguous, with darker basal half (
|
||
<figureCitation id="75962A54FF89FF9BFCACF9CFFC49F982" box="[843,952,1565,1590]" captionStart="Figure 10" captionStartId="16.[160,225,1118,1141]" captionTargetBox="[163,1153,142,1086]" captionTargetId="figure-136@16.[163,1153,142,1086]" captionTargetPageId="16" captionText="Figure 10. Male adults of two new species: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) General view, (b) Wings; (c, d) Epeorus (Proepeorus) falcatus sp. nov., (c) General view, (d) Wings; (e, f) Thorax of Epeorus (Proepeorus) gibbus sp. nov., (e) Dorsal view, (f) Ventral view; (g, h) Thorax of Epeorus (Proepeorus) falcatus sp. nov., (g) Dorsal view, (h) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5466049" httpUri="https://zenodo.org/record/5466049/files/figure.png" pageId="16" pageNumber="1147">Figure 10</figureCitation>
|
||
(g)).
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF89FF9AFF47F9ECFEB6FC63" blockId="16.[160,1156,1460,1728]" lastBlockId="17.[160,1156,889,1711]" lastPageId="17" lastPageNumber="1148" pageId="16" pageNumber="1147">
|
||
<emphasis id="DFD9EAC3FF89FF9BFF47F9ECFF04F9EC" box="[160,245,1598,1624]" italics="true" pageId="16" pageNumber="1147">Thorax.</emphasis>
|
||
Dark brown. Prosternum dark brown, median depression of mesothoracic furcasternum convergent anteriorly (
|
||
<figureCitation id="75962A54FF89FF9BFDBBF9B0FD20F9CF" box="[604,721,1634,1659]" captionStart="Figure 10" captionStartId="16.[160,225,1118,1141]" captionTargetBox="[163,1153,142,1086]" captionTargetId="figure-136@16.[163,1153,142,1086]" captionTargetPageId="16" captionText="Figure 10. Male adults of two new species: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) General view, (b) Wings; (c, d) Epeorus (Proepeorus) falcatus sp. nov., (c) General view, (d) Wings; (e, f) Thorax of Epeorus (Proepeorus) gibbus sp. nov., (e) Dorsal view, (f) Ventral view; (g, h) Thorax of Epeorus (Proepeorus) falcatus sp. nov., (g) Dorsal view, (h) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5466049" httpUri="https://zenodo.org/record/5466049/files/figure.png" pageId="16" pageNumber="1147">Figure 10</figureCitation>
|
||
(h)). Most area of wings transparent except semi-hyaline stigma area. Forewings with brown veins and dark brown costal brace, stigma region with 9–10 cross veins between Sc and C; MA forked 2/3 of distance from base to margin,
|
||
<collectionCode id="8BBCAE14FF88FF9AFD60FCABFD5AFC26" box="[647,683,889,914]" country="0" name="Mohonk Preserve, Inc." pageId="17" pageNumber="1148">MP</collectionCode>
|
||
forked 1/3 of distance from base to margin; hindwings transparent,
|
||
<collectionCode id="8BBCAE14FF88FF9AFDCCFC4EFDBEFC01" box="[555,591,924,949]" country="0" name="Mohonk Preserve, Inc." pageId="17" pageNumber="1148">MP</collectionCode>
|
||
forked 1/2 of distance from base to margin (
|
||
<figureCitation id="75962A54FF88FF9AFF4FFC6CFEEDFC63" box="[168,284,958,983]" captionStart="Figure 10" captionStartId="16.[160,225,1118,1141]" captionTargetBox="[163,1153,142,1086]" captionTargetId="figure-136@16.[163,1153,142,1086]" captionTargetPageId="16" captionText="Figure 10. Male adults of two new species: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) General view, (b) Wings; (c, d) Epeorus (Proepeorus) falcatus sp. nov., (c) General view, (d) Wings; (e, f) Thorax of Epeorus (Proepeorus) gibbus sp. nov., (e) Dorsal view, (f) Ventral view; (g, h) Thorax of Epeorus (Proepeorus) falcatus sp. nov., (g) Dorsal view, (h) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.5466049" httpUri="https://zenodo.org/record/5466049/files/figure.png" pageId="17" pageNumber="1148">Figure 10</figureCitation>
|
||
(d)).
|
||
</paragraph>
|
||
<caption id="B9D26659FF88FF9AFF47FD4EFE09FC9D" ID-DOI="http://doi.org/10.5281/zenodo.5466051" ID-Zenodo-Dep="5466051" httpUri="https://zenodo.org/record/5466051/files/figure.png" pageId="17" pageNumber="1148" startId="17.[160,225,668,691]" targetBox="[163,1153,142,637]" targetPageId="17">
|
||
<paragraph id="ED1236D1FF88FF9AFF47FD4EFE09FC9D" blockId="17.[160,1157,668,809]" pageId="17" pageNumber="1148">
|
||
<emphasis id="DFD9EAC3FF88FF9AFF47FD4EFEF8FD07" bold="true" box="[160,265,668,691]" pageId="17" pageNumber="1148">Figure 11.</emphasis>
|
||
Abdominal terga of imagoes of two new species: (a–d)
|
||
<taxonomicName id="2AAD4D52FF88FF9AFCAEFD4EFBADFD07" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[841,1116,668,691]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="1148" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF88FF9AFCAEFD4EFC62FD07" box="[841,915,668,691]" italics="true" pageId="17" pageNumber="1148">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF88FF9AFC44FD4EFBFCFD07" box="[931,1037,668,691]" italics="true" pageId="17" pageNumber="1148">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF88FF9AFBFAFD4EFBADFD07" box="[1053,1116,668,691]" italics="true" pageId="17" pageNumber="1148">gibbus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF88FF9AFB82FD4EFF3CFD65" pageId="17" pageNumber="1148" rank="species">
|
||
<emphasis id="DFD9EAC3FF88FF9AFB82FD4EFF39FD65" bold="true" pageId="17" pageNumber="1148">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, (a) Dorsal view of male, (b) Ventral view of male, (c) Dorsal view of female, (d) Ventral view of female; (e–h)
|
||
<taxonomicName id="2AAD4D52FF88FF9AFECFFD05FDB2FD5A" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[296,579,727,750]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="1148" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF88FF9AFECFFD05FE83FD5A" box="[296,370,727,750]" italics="true" pageId="17" pageNumber="1148">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF88FF9AFE66FD05FE1AFD5A" box="[385,491,727,750]" italics="true" pageId="17" pageNumber="1148">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF88FF9AFE1FFD05FDB2FD5A" box="[504,579,727,750]" italics="true" pageId="17" pageNumber="1148">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF88FF9AFDAEFD05FD6AFD5A" box="[585,667,727,750]" pageId="17" pageNumber="1148" rank="species">
|
||
<emphasis id="DFD9EAC3FF88FF9AFDAEFD05FD66FD5A" bold="true" box="[585,663,727,750]" pageId="17" pageNumber="1148">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, (e) Dorsal view of male, (f) Ventral view of male, (g) Dorsal view of female, (h) Ventral view of female; Abbreviations: ma, medio-anterior myo-sigillum; mp, medio-posterior myo-sigillum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED1236D1FF88FF9AFF47FC32FDEEFB13" blockId="17.[160,1156,889,1711]" pageId="17" pageNumber="1148">
|
||
<emphasis id="DFD9EAC3FF88FF9AFF47FC32FF28FC4E" box="[160,217,992,1018]" italics="true" pageId="17" pageNumber="1148">Legs.</emphasis>
|
||
Forelegs brown, with dark brown tibiae and distal half of femora, paler basal half of femora and tarsi, ratios of femora: tibiae: tarsi = 2.5: 3.2: 5.5, order of tarsal segments arranged in descending order: 1, 2, 3, 4, 5. Middle legs light brown, ratios of femora: tibiae: tarsi = 2.2: 2.1: 1.3, order of tarsal segments arranged in descending length 1, 2, 3, 5, 4 (hind legs missing). Femora of all legs with a median dark brown macula. Claws of each leg dissimilar, one blunt, one hooked.
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF88FF9AFF47FB62FEDCF91B" blockId="17.[160,1156,889,1711]" pageId="17" pageNumber="1148">
|
||
<emphasis id="DFD9EAC3FF88FF9AFF47FB62FEE4FB7E" box="[160,277,1200,1226]" italics="true" pageId="17" pageNumber="1148">Abdomen.</emphasis>
|
||
Light brown, with darker maculae. Terga II–IX with similar colour pattern (
|
||
<figureCitation id="75962A54FF88FF9AFF4FFB01FEE7FB58" box="[168,278,1235,1260]" captionStart="Figure 11" captionStartId="17.[160,225,668,691]" captionTargetBox="[163,1153,142,637]" captionTargetId="figure-303@17.[163,1153,142,637]" captionTargetPageId="17" captionText="Figure 11. Abdominal terga of imagoes of two new species: (a–d) Epeorus (Proepeorus) gibbus sp. nov., (a) Dorsal view of male, (b) Ventral view of male, (c) Dorsal view of female, (d) Ventral view of female; (e–h) Epeorus (Proepeorus) falcatus sp. nov., (e) Dorsal view of male, (f) Ventral view of male, (g) Dorsal view of female, (h) Ventral view of female; Abbreviations: ma, medio-anterior myo-sigillum; mp, medio-posterior myo-sigillum." figureDoi="http://doi.org/10.5281/zenodo.5466051" httpUri="https://zenodo.org/record/5466051/files/figure.png" pageId="17" pageNumber="1148">Figure 11</figureCitation>
|
||
(e)): indistinct brown transverse band along anterior margin; median area brown, with a longitudinal median dark stripe and a pair of paler irregular maculae (like the fusion of two dots); posterior half of each tergum darker and with a dark brown transverse band along posterior margin extended to lateral margin. Tergum X pale, with a median dark stripe. Colouration of sterna II–VIII similar (
|
||
<figureCitation id="75962A54FF88FF9AFC35FA8CFBCEFAC3" box="[978,1087,1374,1399]" captionStart="Figure 11" captionStartId="17.[160,225,668,691]" captionTargetBox="[163,1153,142,637]" captionTargetId="figure-303@17.[163,1153,142,637]" captionTargetPageId="17" captionText="Figure 11. Abdominal terga of imagoes of two new species: (a–d) Epeorus (Proepeorus) gibbus sp. nov., (a) Dorsal view of male, (b) Ventral view of male, (c) Dorsal view of female, (d) Ventral view of female; (e–h) Epeorus (Proepeorus) falcatus sp. nov., (e) Dorsal view of male, (f) Ventral view of male, (g) Dorsal view of female, (h) Ventral view of female; Abbreviations: ma, medio-anterior myo-sigillum; mp, medio-posterior myo-sigillum." figureDoi="http://doi.org/10.5281/zenodo.5466051" httpUri="https://zenodo.org/record/5466051/files/figure.png" pageId="17" pageNumber="1148">Figure 11</figureCitation>
|
||
(f)): 1) dark brown median band; 2) a pair of distinct brown oblique stripes near anterior margin (medio-anterior myo-sigilla, progressively fading posteriorly); 3) a pair of small round blanks inside the median band (medio-posterior myo-sigilla). Sternum IX brown, with longitudinal dark band laterally; styliger light brown, posterior margin slightly convex medially. Forceps segments I–II brown, III light brown and IV pale, combined length of segments III–IV subequal to half of segment II; penis lobes widely divergent, with sharp lateral hooked tips directed dorsally and medio-apical emargination. Submedian titillators distinct (
|
||
<figureCitation id="75962A54FF88FF9AFD51F9A6FCC5F939" box="[694,820,1652,1677]" captionStart="Figure 12" captionStartId="18.[160,225,1161,1184]" captionTargetBox="[163,1153,142,1130]" captionTargetId="figure-131@18.[163,1153,142,1130]" captionTargetPageId="18" captionText="Figure 12. Genitalia of male: (a) Epeorus (Proepeorus) gibbus sp. nov. (ventral view); (b) Epeorus (Proepeorus) falcatus sp. nov. (ventral view); (c) Epeorus (Proepeorus) nipponicus (ventral view); (d) Epeorus (Proepeorus) aculeatus (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5466053" httpUri="https://zenodo.org/record/5466053/files/figure.png" pageId="17" pageNumber="1148">Figures 12</figureCitation>
|
||
(b), 13(d,e)). Cerci with dark annulations.
|
||
</paragraph>
|
||
<caption id="B9D26659FF8BFF99FF47FB5BFD91FB6F" ID-DOI="http://doi.org/10.5281/zenodo.5466053" ID-Zenodo-Dep="5466053" httpUri="https://zenodo.org/record/5466053/files/figure.png" pageId="18" pageNumber="1149" startId="18.[160,225,1161,1184]" targetBox="[163,1153,142,1130]" targetPageId="18">
|
||
<paragraph id="ED1236D1FF8BFF99FF47FB5BFD91FB6F" blockId="18.[160,1156,1161,1243]" pageId="18" pageNumber="1149">
|
||
<emphasis id="DFD9EAC3FF8BFF99FF47FB5BFEFAFB14" bold="true" box="[160,267,1161,1184]" pageId="18" pageNumber="1149">Figure 12.</emphasis>
|
||
Genitalia of male: (a)
|
||
<taxonomicName id="2AAD4D52FF8BFF99FE1FFB5BFCFFFB14" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[504,782,1161,1184]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="1149" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8BFF99FE1FFB5BFDB3FB14" box="[504,578,1161,1184]" italics="true" pageId="18" pageNumber="1149">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8BFF99FDB3FB5BFD4FFB14" box="[596,702,1161,1184]" italics="true" pageId="18" pageNumber="1149">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8BFF99FD28FB5BFCFFFB14" box="[719,782,1161,1184]" italics="true" pageId="18" pageNumber="1149">gibbus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8BFF99FCFEFB5BFC9FFB14" box="[793,878,1161,1184]" pageId="18" pageNumber="1149" rank="species">
|
||
<emphasis id="DFD9EAC3FF8BFF99FCFEFB5BFC9BFB14" bold="true" box="[793,874,1161,1184]" pageId="18" pageNumber="1149">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(ventral view); (b)
|
||
<taxonomicName id="2AAD4D52FF8BFF99FBDEFB5BFE9DFB0A" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="1149" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8BFF99FBDEFB5BFB72FB14" box="[1081,1155,1161,1184]" italics="true" pageId="18" pageNumber="1149">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8BFF99FF40FB75FEE0FB0A" box="[167,273,1191,1214]" italics="true" pageId="18" pageNumber="1149">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8BFF99FEC6FB75FE9DFB0A" box="[289,364,1191,1214]" italics="true" pageId="18" pageNumber="1149">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8BFF99FE92FB75FE38FB0A" box="[373,457,1191,1214]" pageId="18" pageNumber="1149" rank="species">
|
||
<emphasis id="DFD9EAC3FF8BFF99FE92FB75FE34FB0A" bold="true" box="[373,453,1191,1214]" pageId="18" pageNumber="1149">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(ventral view); (c)
|
||
<taxonomicName id="2AAD4D52FF8BFF99FD6BFB75FC36FB0A" authorityName="Ueno" authorityYear="1931" baseAuthorityName="Ueno" baseAuthorityYear="1931" box="[652,967,1191,1214]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="1149" phylum="Arthropoda" rank="species" species="nipponicus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8BFF99FD6BFB75FD27FB0A" box="[652,726,1191,1214]" italics="true" pageId="18" pageNumber="1149">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8BFF99FD01FB75FCA1FB0A" box="[742,848,1191,1214]" italics="true" pageId="18" pageNumber="1149">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8BFF99FC87FB75FC36FB0A" box="[864,967,1191,1214]" italics="true" pageId="18" pageNumber="1149">nipponicus</emphasis>
|
||
</taxonomicName>
|
||
(ventral view); (d)
|
||
<taxonomicName id="2AAD4D52FF8BFF99FF47FB16FE3AFB6F" authorityName=": Nguyen and Bae" authorityYear="2004" box="[160,459,1220,1243]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="1149" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8BFF99FF47FB16FF1BFB6F" box="[160,234,1220,1243]" italics="true" pageId="18" pageNumber="1149">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8BFF99FF1EFB16FE92FB6F" box="[249,355,1220,1243]" italics="true" pageId="18" pageNumber="1149">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8BFF99FE97FB16FE3AFB6F" box="[368,459,1220,1243]" italics="true" pageId="18" pageNumber="1149">aculeatus</emphasis>
|
||
</taxonomicName>
|
||
(ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED1236D1FF8BFF99FF47FAC9FDF2F9D8" blockId="18.[160,1157,1307,1644]" pageId="18" pageNumber="1149">
|
||
<emphasis id="DFD9EAC3FF8BFF99FF47FAC9FEA1FA81" bold="true" box="[160,336,1307,1333]" italics="true" pageId="18" pageNumber="1149">Female imago.</emphasis>
|
||
Body length
|
||
<quantity id="2A559B34FF8BFF99FE19FAC9FD9BFA80" box="[510,618,1307,1332]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="1149" unit="mm" value="10.5">10.5 mm</quantity>
|
||
, forewing
|
||
<quantity id="2A559B34FF8BFF99FD0BFAC9FCA9FA80" box="[748,856,1307,1332]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="1149" unit="mm" value="10.5">10.5 mm</quantity>
|
||
, hindwing
|
||
<quantity id="2A559B34FF8BFF99FC06FAC9FBCEFA80" box="[993,1087,1307,1332]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="1149" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, cerci
|
||
<quantity id="2A559B34FF8BFF99FF47FAECFEE3FAE3" box="[160,274,1342,1367]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="18" pageNumber="1149" unit="mm" value="20.5">20.5 mm</quantity>
|
||
. General reddish-brown, colouration of body similar to male imago. Thorax paler than in male, wings same as male. Length ratios of femora: tibiae: tarsi of fore legs = 2.6: 2.6: 2.1; length ratios of femora: tibiae: tarsi of middle legs = 2.7: 2.6: 1.3; length ratios of femora: tibiae: tarsi of hind legs = 3.0: 2.5: 1.1. Colour pattern of abdominal terga same as male imago but darker (
|
||
<figureCitation id="75962A54FF8BFF99FC03FA1BFBA4FA56" box="[996,1109,1481,1506]" captionStart="Figure 11" captionStartId="17.[160,225,668,691]" captionTargetBox="[163,1153,142,637]" captionTargetId="figure-303@17.[163,1153,142,637]" captionTargetPageId="17" captionText="Figure 11. Abdominal terga of imagoes of two new species: (a–d) Epeorus (Proepeorus) gibbus sp. nov., (a) Dorsal view of male, (b) Ventral view of male, (c) Dorsal view of female, (d) Ventral view of female; (e–h) Epeorus (Proepeorus) falcatus sp. nov., (e) Dorsal view of male, (f) Ventral view of male, (g) Dorsal view of female, (h) Ventral view of female; Abbreviations: ma, medio-anterior myo-sigillum; mp, medio-posterior myo-sigillum." figureDoi="http://doi.org/10.5281/zenodo.5466051" httpUri="https://zenodo.org/record/5466051/files/figure.png" pageId="18" pageNumber="1149">Figure 11</figureCitation>
|
||
(g)). Abdominal sterna somewhat different to male imago (
|
||
<figureCitation id="75962A54FF8BFF99FCA7FA39FC42F9B0" box="[832,947,1515,1540]" captionStart="Figure 11" captionStartId="17.[160,225,668,691]" captionTargetBox="[163,1153,142,637]" captionTargetId="figure-303@17.[163,1153,142,637]" captionTargetPageId="17" captionText="Figure 11. Abdominal terga of imagoes of two new species: (a–d) Epeorus (Proepeorus) gibbus sp. nov., (a) Dorsal view of male, (b) Ventral view of male, (c) Dorsal view of female, (d) Ventral view of female; (e–h) Epeorus (Proepeorus) falcatus sp. nov., (e) Dorsal view of male, (f) Ventral view of male, (g) Dorsal view of female, (h) Ventral view of female; Abbreviations: ma, medio-anterior myo-sigillum; mp, medio-posterior myo-sigillum." figureDoi="http://doi.org/10.5281/zenodo.5466051" httpUri="https://zenodo.org/record/5466051/files/figure.png" pageId="18" pageNumber="1149">Figure 11</figureCitation>
|
||
(h)): sterna VII–IX darker than other segments. Subgenital plate extended to posterior margin of sternum VIII and with rounded posterior margin, sternum IX with shallow median emargination (
|
||
<figureCitation id="75962A54FF8BFF99FEB7F981FE34F9D8" box="[336,453,1619,1644]" captionStart="Figure 14" captionStartId="20.[160,225,1164,1187]" captionTargetBox="[163,1153,142,1122]" captionTargetId="figure-177@20.[163,1153,142,1132]" captionTargetPageId="20" captionText="Figure 14. Subgenital plate and sternum IX of female: (a, b) Epeorus (Proepeorus) gibbus sp. nov.; (c, d) Epeorus (Proepeorus) falcatus sp. nov.; (e, f) Epeorus (Proepeorus) nipponicus." figureDoi="http://doi.org/10.5281/zenodo.5466059" httpUri="https://zenodo.org/record/5466059/files/figure.png" pageId="18" pageNumber="1149">Figure 14</figureCitation>
|
||
(c,d)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B9D26659FF8AFF98FF47FC44FC35FC5C" ID-DOI="http://doi.org/10.5281/zenodo.5466055" ID-Zenodo-Dep="5466055" httpUri="https://zenodo.org/record/5466055/files/figure.png" pageId="19" pageNumber="1150" startId="19.[160,225,918,941]" targetBox="[166,1150,143,884]" targetPageId="19">
|
||
<paragraph id="ED1236D1FF8AFF98FF47FC44FC35FC5C" blockId="19.[160,1155,918,1000]" pageId="19" pageNumber="1150">
|
||
<emphasis id="DFD9EAC3FF8AFF98FF47FC44FEF9FC19" bold="true" box="[160,264,918,941]" pageId="19" pageNumber="1150">Figure 13.</emphasis>
|
||
Genitalia of male: (a, b)
|
||
<taxonomicName id="2AAD4D52FF8AFF98FDE6FC44FCE0FC19" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[513,785,918,941]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="1150" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8AFF98FDE6FC44FDBAFC19" box="[513,587,918,941]" italics="true" pageId="19" pageNumber="1150">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8AFF98FDBDFC44FD35FC19" box="[602,708,918,941]" italics="true" pageId="19" pageNumber="1150">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8AFF98FD35FC44FCE0FC19" box="[722,785,918,941]" italics="true" pageId="19" pageNumber="1150">gibbus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8AFF98FCFEFC44FC9DFC19" box="[793,876,918,941]" pageId="19" pageNumber="1150" rank="species">
|
||
<emphasis id="DFD9EAC3FF8AFF98FCFEFC44FC99FC19" bold="true" box="[793,872,918,941]" pageId="19" pageNumber="1150">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, (a) Ventral view, (b) Dorsal view; (c)
|
||
<taxonomicName id="2AAD4D52FF8AFF98FF1DFC66FDC3FC7F" authorityName="Ueno" authorityYear="1931" baseAuthorityName="Ueno" baseAuthorityYear="1931" box="[250,562,948,971]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="1150" phylum="Arthropoda" rank="species" species="nipponicus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8AFF98FF1DFC66FEB5FC7F" box="[250,324,948,971]" italics="true" pageId="19" pageNumber="1150">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8AFF98FEB4FC66FE4CFC7F" box="[339,445,948,971]" italics="true" pageId="19" pageNumber="1150">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8AFF98FE2CFC66FDC3FC7F" box="[459,562,948,971]" italics="true" pageId="19" pageNumber="1150">nipponicus</emphasis>
|
||
</taxonomicName>
|
||
(ventral view); (d, e)
|
||
<taxonomicName id="2AAD4D52FF8AFF98FCEFFC66FBD5FC7F" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[776,1060,948,971]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="1150" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8AFF98FCEFFC66FCA3FC7F" box="[776,850,948,971]" italics="true" pageId="19" pageNumber="1150">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8AFF98FC86FC66FC3AFC7F" box="[865,971,948,971]" italics="true" pageId="19" pageNumber="1150">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8AFF98FC3EFC66FBD5FC7F" box="[985,1060,948,971]" italics="true" pageId="19" pageNumber="1150">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8AFF98FBCCFC66FB8FFC7F" box="[1067,1150,948,971]" pageId="19" pageNumber="1150" rank="species">
|
||
<emphasis id="DFD9EAC3FF8AFF98FBCCFC66FB88FC7F" bold="true" box="[1067,1145,948,971]" pageId="19" pageNumber="1150">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, (d) Ventral view, (e) Dorsal view; (f)
|
||
<taxonomicName id="2AAD4D52FF8AFF98FDE2FC03FCC1FC5C" authorityName=": Nguyen and Bae" authorityYear="2004" box="[517,816,977,1000]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="1150" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8AFF98FDE2FC03FDBEFC5C" box="[517,591,977,1000]" italics="true" pageId="19" pageNumber="1150">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8AFF98FDBAFC03FD36FC5C" box="[605,711,977,1000]" italics="true" pageId="19" pageNumber="1150">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8AFF98FD32FC03FCC1FC5C" box="[725,816,977,1000]" italics="true" pageId="19" pageNumber="1150">aculeatus</emphasis>
|
||
</taxonomicName>
|
||
(ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A5B7655AFF8AFF98FF47FBCAFE18FBC3" pageId="19" pageNumber="1150" type="etymology">
|
||
<paragraph id="ED1236D1FF8AFF98FF47FBCAFEECFB86" blockId="19.[160,1155,1048,1143]" box="[160,285,1048,1074]" pageId="19" pageNumber="1150">
|
||
<emphasis id="DFD9EAC3FF8AFF98FF47FBCAFEECFB86" bold="true" box="[160,285,1048,1074]" italics="true" pageId="19" pageNumber="1150">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF8AFF98FF47FBE9FE18FBC3" blockId="19.[160,1155,1048,1143]" pageId="19" pageNumber="1150">
|
||
The Latin specific name
|
||
<taxonomicName id="2AAD4D52FF8AFF98FE5BFBE9FDE0FBE0" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[444,529,1083,1108]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="1150" phylum="Arthropoda" rank="species" species="falcatus">
|
||
<emphasis id="DFD9EAC3FF8AFF98FE5BFBE9FDE0FBE0" box="[444,529,1083,1108]" italics="true" pageId="19" pageNumber="1150">falcatus</emphasis>
|
||
</taxonomicName>
|
||
means sickle-shaped, curved or hooked, refers to the sickle-shaped gill I of nymph.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5B7655AFF8AFF98FF47FB60FCD3FAE2" pageId="19" pageNumber="1150" type="biology_ecology">
|
||
<paragraph id="ED1236D1FF8AFF98FF47FB60FF09FB78" blockId="19.[160,1155,1202,1366]" box="[160,248,1202,1228]" pageId="19" pageNumber="1150">
|
||
<emphasis id="DFD9EAC3FF8AFF98FF47FB60FF09FB78" bold="true" box="[160,248,1202,1228]" italics="true" pageId="19" pageNumber="1150">Biology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF8AFF98FF47FB07FCD3FAE2" blockId="19.[160,1155,1202,1366]" pageId="19" pageNumber="1150">
|
||
Nymphs of this species occur in the small-sized mountain streams (1.0–3.0 m wide,
|
||
<quantity id="2A559B34FF8AFF98FBB2FB07FF2FFAA5" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="19" pageNumber="1150" unit="m" value="0.3" valueMax="0.5" valueMin="0.1">0.1– 0.5 m</quantity>
|
||
deep, ca.
|
||
<quantity id="2A559B34FF8AFF98FEA8FB2AFE16FAA5" box="[335,487,1272,1297]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="19" pageNumber="1150" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">1000–1300 m</quantity>
|
||
a.s.l.), and they are found underneath stones in moderately flowing sections where the substrate is mostly stony. Moulting of the subimago was observed in 7:30 PM, and the process lasts ca. 4 minutes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5B7655AFF8AFF9FFF47FA43FEB7F9CD" lastPageId="20" lastPageNumber="1151" pageId="19" pageNumber="1150" type="diagnosis">
|
||
<paragraph id="ED1236D1FF8AFF98FF47FA43FE40FA1F" blockId="19.[160,1156,1425,1728]" box="[160,433,1425,1451]" pageId="19" pageNumber="1150">
|
||
<emphasis id="DFD9EAC3FF8AFF98FF47FA43FE40FA1F" bold="true" box="[160,433,1425,1451]" italics="true" pageId="19" pageNumber="1150">Diagnosis and remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF8AFF98FF47FA67FDE1F9CF" blockId="19.[160,1156,1425,1728]" pageId="19" pageNumber="1150">
|
||
This species can be recognised by the following diagnosis characters. In the adult: 1) penis lobes with lateral hooked projections and distinct median titillators (
|
||
<figureCitation id="75962A54FF8AFF98FC2EFA05FBC8FA44" box="[969,1081,1495,1520]" captionStart="Figure 13" captionStartId="19.[160,225,918,941]" captionTargetBox="[166,1150,143,884]" captionTargetId="figure-194@19.[163,1153,142,887]" captionTargetPageId="19" captionText="Figure 13. Genitalia of male: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) Ventral view, (b) Dorsal view; (c) Epeorus (Proepeorus) nipponicus (ventral view); (d, e) Epeorus (Proepeorus) falcatus sp. nov., (d) Ventral view, (e) Dorsal view; (f) Epeorus (Proepeorus) aculeatus (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5466055" httpUri="https://zenodo.org/record/5466055/files/figure.png" pageId="19" pageNumber="1150">Figure 13</figureCitation>
|
||
(d)); 2) subgenital plate of female oval, extended to posterior margin of sternum VIII (
|
||
<figureCitation id="75962A54FF8AFF98FBFEFA28FB72F9A7" box="[1049,1155,1530,1555]" captionStart="Figure 14" captionStartId="20.[160,225,1164,1187]" captionTargetBox="[163,1153,142,1122]" captionTargetId="figure-177@20.[163,1153,142,1132]" captionTargetPageId="20" captionText="Figure 14. Subgenital plate and sternum IX of female: (a, b) Epeorus (Proepeorus) gibbus sp. nov.; (c, d) Epeorus (Proepeorus) falcatus sp. nov.; (e, f) Epeorus (Proepeorus) nipponicus." figureDoi="http://doi.org/10.5281/zenodo.5466059" httpUri="https://zenodo.org/record/5466059/files/figure.png" pageId="19" pageNumber="1150">Figure 14</figureCitation>
|
||
(c,d)). In the nymph: 1) lamellae of gill I moderately expanded forward and narrowed apically (
|
||
<figureCitation id="75962A54FF8AFF98FEE2F9EDFE95F9EC" box="[261,356,1599,1624]" captionStart="Figure 9" captionStartId="15.[160,225,913,936]" captionTargetBox="[164,1151,143,864]" captionTargetId="figure-239@15.[163,1153,142,881]" captionTargetPageId="15" captionText="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="19" pageNumber="1150">Figure 9</figureCitation>
|
||
(i)); 2) abdominal terga with paired small submedian spines and blunt denticle posteriorly (
|
||
<figureCitation id="75962A54FF8AFF98FE6FF9B0FE14F9CF" box="[392,485,1634,1659]" captionStart="Figure 9" captionStartId="15.[160,225,913,936]" captionTargetBox="[164,1151,143,864]" captionTargetId="figure-239@15.[163,1153,142,881]" captionTargetPageId="15" captionText="Figure 9. Nymph of Epeorus (Proepeorus) falcatus sp. nov.: (a) Head capsule; (b) Labrum (ventral view on left, dorsal view on right); (c) Right mandible; (d) Left mandible; (e) Glossae of labium (dorsal view on left, ventral view on right); (f) Femoral setae; (g) Claw; (h) Posterior margin of tergum VII (vertical line is midline of tergum); (i) Gill I; (j) Gill II; (k) Gill VII." figureDoi="http://doi.org/10.5281/zenodo.5466047" httpUri="https://zenodo.org/record/5466047/files/figure.png" pageId="19" pageNumber="1150">Figure 9</figureCitation>
|
||
(h)).
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF8AFF9FFF27F957FD9EFA5B" blockId="19.[160,1156,1425,1728]" lastBlockId="20.[160,1156,1286,1657]" lastPageId="20" lastPageNumber="1151" pageId="19" pageNumber="1150">
|
||
The nymph of
|
||
<taxonomicName id="2AAD4D52FF8AFF98FE94F957FDE7F92A" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[371,534,1669,1694]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="1155" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8AFF98FE94F957FE71F92A" box="[371,384,1669,1694]" italics="true" pageId="19" pageNumber="1150">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8AFF98FE7EF957FE56F92A" box="[409,423,1669,1694]" italics="true" pageId="19" pageNumber="1150">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8AFF98FE58F957FDE7F92A" box="[447,534,1669,1694]" italics="true" pageId="19" pageNumber="1150">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8AFF98FDC5F957FD75F92A" box="[546,644,1669,1694]" pageId="19" pageNumber="1150" rank="species">
|
||
<emphasis id="DFD9EAC3FF8AFF98FDC5F957FD8FF92A" bold="true" box="[546,638,1669,1694]" pageId="19" pageNumber="1150">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is most similar to that of
|
||
<emphasis id="DFD9EAC3FF8AFF98FC23F957FC20F92A" box="[964,977,1669,1694]" italics="true" pageId="19" pageNumber="1150">E</emphasis>
|
||
.
|
||
<taxonomicName id="2AAD4D52FF8AFF98FC06F957FB8CF92A" authorityName=": Nguyen and Bae" authorityYear="2004" box="[993,1149,1669,1694]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="1150" phylum="Arthropoda" rank="species" species="bifurcatus" subGenus="Proepeorus">
|
||
(
|
||
<emphasis id="DFD9EAC3FF8AFF98FC0EF957FC06F92A" box="[1001,1015,1669,1694]" italics="true" pageId="19" pageNumber="1150">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8AFF98FBE8F957FB8CF92A" box="[1039,1149,1669,1694]" italics="true" pageId="19" pageNumber="1150">bifurcatus</emphasis>
|
||
</taxonomicName>
|
||
. They both have paired small submedian spines and blunt denticles along posterior margin of abdominal terga. But
|
||
<emphasis id="DFD9EAC3FF8DFF9FFDC6FAD4FDDFFAAB" box="[545,558,1286,1311]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
.
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFDD9FAD4FD2BFAAB" authorityName=": Nguyen and Bae" authorityYear="2004" box="[574,730,1286,1311]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1151" phylum="Arthropoda" rank="species" species="bifurcatus" subGenus="Proepeorus">
|
||
(
|
||
<emphasis id="DFD9EAC3FF8DFF9FFDA1FAD4FDA5FAAB" box="[582,596,1286,1311]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD8BFAD4FD2BFAAB" box="[620,730,1286,1311]" italics="true" pageId="20" pageNumber="1151">bifurcatus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished by the somewhat expanded gills I (
|
||
<figureCitation id="75962A54FF8DFF9FFE4AFAFAFDFFFAF5" box="[429,526,1320,1345]" captionStart="Figure 6" captionStartId="7.[160,225,827,850]" captionTargetBox="[163,1142,142,796]" captionTargetId="figure-232@7.[163,1153,142,796]" captionTargetPageId="7" captionText="Figure 6. Nymphal structures of Epeorus spp.: (a–c) Labrum, (a) Epeorus (Epeorus) melli, (b) Epeorus (Propeorus) nipponicus, (c) E. (P.) unispinosus; (d, e) Gill I, (d) E. (P.) unispinosus, (e) E. (P.) bifurcatus; (f–h) Posterior margin of tergum VII, (f) E. (P.) bispinosus, (g) E. (P.) rhithralis; (h) E. (P.) bifurcatus." figureDoi="http://doi.org/10.5281/zenodo.5466039" httpUri="https://zenodo.org/record/5466039/files/figure.png" pageId="20" pageNumber="1151">Figure 6</figureCitation>
|
||
(e)), which are moderately expanded in
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFC03FAFAFB75FAF5" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[996,1156,1320,1345]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1155" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC03FAFAFC00FAF5" box="[996,1009,1320,1345]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8DFF9FFBE0FAFAFBE4FAF5" box="[1031,1045,1320,1345]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFBCAFAFAFB75FAF5" box="[1069,1156,1320,1345]" italics="true" pageId="20" pageNumber="1151">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8DFF9FFF47FA99FEF0FAD0" box="[160,257,1355,1380]" pageId="20" pageNumber="1151" rank="species">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFF47FA99FF0AFAD0" bold="true" box="[160,251,1355,1380]" pageId="20" pageNumber="1151">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
In addition, the imaginal characters of
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFD32FA99FC78FAD0" authorityName=": Nguyen and Bae" authorityYear="2004" box="[725,905,1355,1380]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1155" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD32FA99FD13FAD0" box="[725,738,1355,1380]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD1EFA99FCF6FAD0" box="[761,775,1355,1380]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFCF8FA99FC78FAD0" box="[799,905,1355,1380]" italics="true" pageId="20" pageNumber="1151">aculeatus</emphasis>
|
||
</taxonomicName>
|
||
are very similar to
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFB96FA99FED4FA33" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1155" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFB96FA99FB8FFAD0" box="[1137,1150,1355,1380]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8DFF9FFF4FFABCFF47FA33" box="[168,182,1390,1415]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFF29FABCFED4FA33" box="[206,293,1390,1415]" italics="true" pageId="20" pageNumber="1151">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8DFF9FFED7FABCFE60FA33" box="[304,401,1390,1415]" pageId="20" pageNumber="1151" rank="species">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFED7FABCFE7AFA33" bold="true" box="[304,395,1390,1415]" pageId="20" pageNumber="1151">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, especially the colouration pattern of abdomen (see
|
||
<bibRefCitation id="893C4B20FF8DFF9FFBEDFABCFE9CFA1D" author="Webb JM & McCafferty WP" pageId="20" pageNumber="1151" pagination="65 - 68" refId="ref13791" refString="Webb JM, McCafferty WP. 2006 a. First description of the adult male of Epeorus aculeatus Braasch (Ephemeroptera: Heptageniidae). Zootaxa. 1277: 65 - 68. doi: 10.11646 / zootaxa. 1277.1.5." type="journal article" year="2006">Webb and McCafferty 2006a</bibRefCitation>
|
||
:
|
||
<figureCitation id="75962A54FF8DFF9FFE67FA42FE1FFA1D" box="[384,494,1424,1449]" captionStart="Figure 1" captionStartId="3.[160,225,582,605]" captionTargetBox="[163,1153,142,550]" captionTargetId="figure-40@3.[163,1153,142,551]" captionTargetPageId="3" captionText="Figure 1. Nymph of Epeorus spp.: (a–c) Epeorus (Proepeorus) nipponicus: (a) Nymphal habitus, (b) Head capsule, (c) Abdominal terga VII–X; D–F: Epeorus (Proepeorus) unispinosus: (d) Nymphal habitus, (e) Head capsule, (f) Abdominal terga VII–X." figureDoi="http://doi.org/10.5281/zenodo.5466022" httpUri="https://zenodo.org/record/5466022/files/figure.png" pageId="20" pageNumber="1151">Figures 1</figureCitation>
|
||
and
|
||
<figureCitation id="75962A54FF8DFF9FFDD3FA42FDB3FA1D" box="[564,578,1424,1449]" captionStart="Figure 2" captionStartId="3.[160,225,1504,1527]" captionTargetBox="[163,1153,723,1473]" captionTargetId="figure-86@3.[163,1153,723,1473]" captionTargetPageId="3" captionText="Figure 2. Nymphal habitus and structures of Epeorus (Proepeorus) gibbus sp. nov.: (a) Nymphal habitus (dorsal view); (b) Nymphal habitus (ventral view); (c) Head capsule; (d) Abdominal tergum (segment VII); (e) Gill VII (ventral view); (f) Middle leg." figureDoi="http://doi.org/10.5281/zenodo.5466026" httpUri="https://zenodo.org/record/5466026/files/figure.png" pageId="20" pageNumber="1151">2</figureCitation>
|
||
). However, the penis of
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFC91FA42FBC1FA1D" authorityName=": Nguyen and Bae" authorityYear="2004" box="[886,1072,1424,1449]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1155" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC91FA42FC72FA1D" box="[886,899,1424,1449]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC7AFA42FC5AFA1D" box="[925,939,1424,1449]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC21FA42FBC1FA1D" box="[966,1072,1424,1449]" italics="true" pageId="20" pageNumber="1151">aculeatus</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFED0FA61FE26FA78" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[311,471,1459,1484]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1155" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFED0FA61FEB5FA78" box="[311,324,1459,1484]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8DFF9FFEBCFA61FE98FA78" box="[347,361,1459,1484]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFE67FA61FE26FA78" box="[384,471,1459,1484]" italics="true" pageId="20" pageNumber="1151">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8DFF9FFE07FA61FDB1FA78" box="[480,576,1459,1484]" pageId="20" pageNumber="1151" rank="species">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFE07FA61FDCBFA78" bold="true" box="[480,570,1459,1484]" pageId="20" pageNumber="1151">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by the absence of the lateral hooked projections and the smaller titillators (
|
||
<figureCitation id="75962A54FF8DFF9FFE3DFA04FDBAFA5B" box="[474,587,1494,1519]" captionStart="Figure 13" captionStartId="19.[160,225,918,941]" captionTargetBox="[166,1150,143,884]" captionTargetId="figure-194@19.[163,1153,142,887]" captionTargetPageId="19" captionText="Figure 13. Genitalia of male: (a, b) Epeorus (Proepeorus) gibbus sp. nov., (a) Ventral view, (b) Dorsal view; (c) Epeorus (Proepeorus) nipponicus (ventral view); (d, e) Epeorus (Proepeorus) falcatus sp. nov., (d) Ventral view, (e) Dorsal view; (f) Epeorus (Proepeorus) aculeatus (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5466055" httpUri="https://zenodo.org/record/5466055/files/figure.png" pageId="20" pageNumber="1151">Figure 13</figureCitation>
|
||
(f)).
|
||
</paragraph>
|
||
<caption id="B9D26659FF8DFF9FFF47FB5EFC63FB74" ID-DOI="http://doi.org/10.5281/zenodo.5466059" ID-Zenodo-Dep="5466059" httpUri="https://zenodo.org/record/5466059/files/figure.png" pageId="20" pageNumber="1151" startId="20.[160,225,1164,1187]" targetBox="[163,1153,142,1122]" targetPageId="20">
|
||
<paragraph id="ED1236D1FF8DFF9FFF47FB5EFC63FB74" blockId="20.[160,1155,1164,1216]" pageId="20" pageNumber="1151">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFF47FB5EFEF4FB17" bold="true" box="[160,261,1164,1187]" pageId="20" pageNumber="1151">Figure 14.</emphasis>
|
||
Subgenital plate and sternum IX of female: (a, b)
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFD02FB5EFC00FB17" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[741,1009,1164,1187]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1151" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD02FB5EFCDEFB17" box="[741,815,1164,1187]" italics="true" pageId="20" pageNumber="1151">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8DFF9FFCDBFB5EFC57FB17" box="[828,934,1164,1187]" italics="true" pageId="20" pageNumber="1151">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC55FB5EFC00FB17" box="[946,1009,1164,1187]" italics="true" pageId="20" pageNumber="1151">gibbus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC11FB5EFBBFFB17" bold="true" box="[1014,1102,1164,1187]" pageId="20" pageNumber="1151">
|
||
<taxonomicNameLabel id="C4EA57B8FF8DFF9FFC11FB5EFBBBFB17" box="[1014,1098,1164,1187]" pageId="20" pageNumber="1151" rank="species">sp. nov.</taxonomicNameLabel>
|
||
;
|
||
</emphasis>
|
||
(c, d)
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFF47FB7BFE4AFB74" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[160,443,1193,1216]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1151" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFF47FB7BFF1BFB74" box="[160,234,1193,1216]" italics="true" pageId="20" pageNumber="1151">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8DFF9FFF1EFB7BFE92FB74" box="[249,355,1193,1216]" italics="true" pageId="20" pageNumber="1151">Proepeorus</emphasis>
|
||
)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFE97FB7BFE4AFB74" box="[368,443,1193,1216]" italics="true" pageId="20" pageNumber="1151">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8DFF9FFE25FB7BFDE5FB74" box="[450,532,1193,1216]" pageId="20" pageNumber="1151" rank="species">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFE25FB7BFDE1FB74" bold="true" box="[450,528,1193,1216]" pageId="20" pageNumber="1151">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
; (e, f)
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFDB2FB7BFC7FFB74" authorityName="Ueno" authorityYear="1931" baseAuthorityName="Ueno" baseAuthorityYear="1931" box="[597,910,1193,1216]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1153" phylum="Arthropoda" rank="species" species="nipponicus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFDB2FB7BFD6EFB74" box="[597,671,1193,1216]" italics="true" pageId="20" pageNumber="1151">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD4AFB7BFCE6FB74" box="[685,791,1193,1216]" italics="true" pageId="20" pageNumber="1151">Proepeorus</emphasis>
|
||
) nipponicus
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED1236D1FF8DFF9FFF27FA2AFEB7F9CD" blockId="20.[160,1156,1286,1657]" pageId="20" pageNumber="1151">
|
||
As a new species belonging to the ‘paired-spines’ species in
|
||
<collectingCountry id="95BA7641FF8DFF9FFC96FA2AFC45F9A5" box="[881,948,1528,1553]" name="China" pageId="20" pageNumber="1151">China</collectingCountry>
|
||
,
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFC58FA2AFBA9F9A5" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[959,1112,1528,1553]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1155" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC58FA2AFC3DF9A5" box="[959,972,1528,1553]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8DFF9FFC06FA2AFC1EF9A5" box="[993,1007,1528,1553]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFBE4FA2AFBA9F9A5" box="[1027,1112,1528,1553]" italics="true" pageId="20" pageNumber="1151">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8DFF9FFB86FA2AFF22F980" pageId="20" pageNumber="1151" rank="species">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFB86FA2AFF3CF980" bold="true" pageId="20" pageNumber="1151">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
possess similar imaginal characters as
|
||
<taxonomicName id="2AAD4D52FF8DFF9FFD62F9C9FCDEF980" authorityName=": Nguyen and Bae" authorityYear="2004" box="[645,815,1563,1588]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="1155" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD62F9C9FD63F980" box="[645,658,1563,1588]" italics="true" pageId="20" pageNumber="1151">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD41F9C9FD45F980" box="[678,692,1563,1588]" italics="true" pageId="20" pageNumber="1151">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8DFF9FFD20F9C9FCDEF980" box="[711,815,1563,1588]" italics="true" pageId="20" pageNumber="1151">aculeatus</emphasis>
|
||
</taxonomicName>
|
||
and its titillators on penes are distinct. Therefore, it is possible that all the male imagoes of ‘paired-spines’ species may have titillators.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B9D26659FF8CFF9EFF47FC60FEFCFB95" ID-DOI="http://doi.org/10.5281/zenodo.5466061" ID-Zenodo-Dep="5466061" httpUri="https://zenodo.org/record/5466061/files/figure.png" pageId="21" pageNumber="1152" startId="21.[160,225,946,969]" targetBox="[163,1153,187,914]" targetPageId="21">
|
||
<paragraph id="ED1236D1FF8CFF9EFF47FC60FEFCFB95" blockId="21.[160,1155,946,1057]" pageId="21" pageNumber="1152">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFF47FC60FEFBFC7D" bold="true" box="[160,266,946,969]" pageId="21" pageNumber="1152">Figure 15.</emphasis>
|
||
Mouthparts of
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFE49FC60FD8BFC7D" authority="" box="[430,634,946,969]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1152" phylum="Arthropoda" rank="subGenus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFE49FC60FE09FC7D" box="[430,504,946,969]" italics="true" pageId="21" pageNumber="1152">Epeorus</emphasis>
|
||
(
|
||
<emphasis id="DFD9EAC3FF8CFF9EFDEDFC60FD85FC7D" box="[522,628,946,969]" italics="true" pageId="21" pageNumber="1152">Proepeorus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.: (a–f) labrum, (a)
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFC8CFC60FBE7FC7D" authorityName="Ueno" authorityYear="1931" baseAuthorityName="Ueno" baseAuthorityYear="1931" box="[875,1046,946,969]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1155" phylum="Arthropoda" rank="species" species="nipponicus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC8CFC60FC87FC7D" box="[875,886,946,969]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC6BFC60FC69FC7D" box="[908,920,946,969]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC48FC60FBE7FC7D" box="[943,1046,946,969]" italics="true" pageId="21" pageNumber="1152">nipponicus</emphasis>
|
||
</taxonomicName>
|
||
; (b)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFBADFC60FBA4FC7D" box="[1098,1109,946,969]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
.
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFB83FC60FEFEFC52" authorityName="Braasch" authorityYear="1980" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1152" phylum="Arthropoda" rank="species" species="unispinosus" subGenus="Proepeorus">
|
||
(
|
||
<emphasis id="DFD9EAC3FF8CFF9EFB8CFC60FB86FC7D" box="[1131,1143,946,969]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFF47FC1DFEFEFC52" box="[160,271,975,998]" italics="true" pageId="21" pageNumber="1152">unispinosus</emphasis>
|
||
</taxonomicName>
|
||
; (c)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFEDEFC1DFEB5FC52" box="[313,324,975,998]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
.
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFEA8FC1DFE25FC52" authorityName=": Nguyen and Bae" authorityYear="2004" box="[335,468,975,998]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1152" phylum="Arthropoda" rank="species" species="bifurcatus" subGenus="Proepeorus">
|
||
(
|
||
<emphasis id="DFD9EAC3FF8CFF9EFEB1FC1DFE93FC52" box="[342,354,975,998]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFE92FC1DFE25FC52" box="[373,468,975,998]" italics="true" pageId="21" pageNumber="1152">bifurcatus</emphasis>
|
||
</taxonomicName>
|
||
; (d)
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFDE7FC1DFD66FC52" authorityName=": Nguyen and Bae" authorityYear="2004" box="[512,663,975,998]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1155" phylum="Arthropoda" rank="species" species="aculeatus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFDE7FC1DFDFAFC52" box="[512,523,975,998]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8CFF9EFDFAFC1DFDD8FC52" box="[541,553,975,998]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFDDBFC1DFD66FC52" box="[572,663,975,998]" italics="true" pageId="21" pageNumber="1152">aculeatus</emphasis>
|
||
</taxonomicName>
|
||
; (e)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFD25FC1DFD3CFC52" box="[706,717,975,998]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
.
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFD3FFC1DFCA2FC52" authorityName="Braasch" authorityYear="1980" box="[728,851,975,998]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1152" phylum="Arthropoda" rank="species" species="rhithralis" subGenus="Proepeorus">
|
||
(
|
||
<emphasis id="DFD9EAC3FF8CFF9EFD38FC1DFD1AFC52" box="[735,747,975,998]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFD19FC1DFCA2FC52" box="[766,851,975,998]" italics="true" pageId="21" pageNumber="1152">rhithralis</emphasis>
|
||
</taxonomicName>
|
||
; (f)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC9EFC1DFC75FC52" box="[889,900,975,998]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
.
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFC68FC1DFBE6FC52" authorityName=": Kluge" authorityYear="1997" box="[911,1047,975,998]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1152" phylum="Arthropoda" rank="species" species="bispinosus" subGenus="Proepeorus">
|
||
(
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC71FC1DFC53FC52" box="[918,930,975,998]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC52FC1DFBE6FC52" box="[949,1047,975,998]" italics="true" pageId="21" pageNumber="1152">bispinosus</emphasis>
|
||
</taxonomicName>
|
||
; (g–i) base of apical tooth complex of maxilla, (g)
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFDCBFC3EFD5AFBB7" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[556,683,1004,1027]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1155" phylum="Arthropoda" rank="species" species="gibbus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFDCBFC3EFDC6FBB7" box="[556,567,1004,1027]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8CFF9EFDABFC3EFDA9FBB7" box="[588,600,1004,1027]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFD8BFC3EFD5AFBB7" box="[620,683,1004,1027]" italics="true" pageId="21" pageNumber="1152">gibbus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8CFF9EFD53FC3EFCF9FBB7" box="[692,776,1004,1027]" pageId="21" pageNumber="1152" rank="species">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFD53FC3EFCF2FBB7" bold="true" box="[692,771,1004,1027]" pageId="21" pageNumber="1152">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
; (h)
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFCDDFC3EFC34FBB7" authority="Ma & Lei & Li & Zhou, 2021" authorityName="Ma & Lei & Li & Zhou" authorityYear="2021" box="[826,965,1004,1027]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1155" phylum="Arthropoda" rank="species" species="falcatus" status="sp. nov." subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFCDDFC3EFCB4FBB7" box="[826,837,1004,1027]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8CFF9EFCBEFC3EFC94FBB7" box="[857,869,1004,1027]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.)
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC9DFC3EFC34FBB7" box="[890,965,1004,1027]" italics="true" pageId="21" pageNumber="1152">falcatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4EA57B8FF8CFF9EFC2AFC3EFBD0FBB7" box="[973,1057,1004,1027]" pageId="21" pageNumber="1152" rank="species">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFC2AFC3EFBEDFBB7" bold="true" box="[973,1052,1004,1027]" pageId="21" pageNumber="1152">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
; (i)
|
||
<taxonomicName id="2AAD4D52FF8CFF9EFBACFC3EFEF8FB95" authorityName="Ueno" authorityYear="1931" baseAuthorityName="Ueno" baseAuthorityYear="1931" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="1155" phylum="Arthropoda" rank="species" species="nipponicus" subGenus="Proepeorus">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFBACFC3EFBA7FBB7" box="[1099,1110,1004,1027]" italics="true" pageId="21" pageNumber="1152">E</emphasis>
|
||
. (
|
||
<emphasis id="DFD9EAC3FF8CFF9EFB8CFC3EFB86FBB7" box="[1131,1143,1004,1027]" italics="true" pageId="21" pageNumber="1152">P</emphasis>
|
||
.) nipponicus
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="A5B7655AFF8CFF9EFF47FB8BFDECFB21" box="[160,541,1113,1173]" pageId="21" pageNumber="1152" type="distribution">
|
||
<paragraph id="ED1236D1FF8CFF9EFF47FB8BFEDDFBC7" blockId="21.[160,541,1113,1173]" box="[160,300,1113,1139]" pageId="21" pageNumber="1152">
|
||
<emphasis id="DFD9EAC3FF8CFF9EFF47FB8BFEDDFBC7" bold="true" box="[160,300,1113,1139]" italics="true" pageId="21" pageNumber="1152">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="ED1236D1FF8CFF9EFF47FBAEFDECFB21" blockId="21.[160,541,1113,1173]" box="[160,541,1148,1173]" pageId="21" pageNumber="1152">
|
||
<materialsCitation id="5DC53C8CFF8CFF9EFF47FBAEFDE9FB21" box="[160,536,1148,1173]" country="China" location="Guizhou" pageId="21" pageNumber="1152" specimenCount="1" stateProvince="Guizhou">
|
||
<collectingCountry id="95BA7641FF8CFF9EFF47FBAEFF11FB21" box="[160,224,1148,1173]" name="China" pageId="21" pageNumber="1152">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="2F69F833FF8CFF9EFF09FBAEFEA1FB21" box="[238,336,1148,1173]" country="China" name="Guizhou" pageId="21" pageNumber="1152">Guizhou</collectingRegion>
|
||
,
|
||
<collectingRegion id="2F69F833FF8CFF9EFEBDFBAEFE4BFB21" box="[346,442,1148,1173]" country="China" name="Guangxi" pageId="21" pageNumber="1152">Guangxi</collectingRegion>
|
||
,
|
||
<collectingRegion id="2F69F833FF8CFF9EFE23FBAEFDE3FB21" box="[452,530,1148,1173]" country="China" name="Hunan" pageId="21" pageNumber="1152">Hunan</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |