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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e84013" ID-Pensoft-Pub="1864-8312-80-575" ID-Pensoft-UUID="81B59BA932BC5DAF90815FD08F5820A6" ID-ZooBank="4A6669D57DDD4AE0A8B455724B9C594A" ModsDocID="1864-8312-80-575" checkinTime="1668813875820" checkinUser="pensoft" docAuthor="Linsky, Marek, Ciamporova-Zatovicova, Zuzana, Lassova, Kristina &amp; Ciampor Jr, Fedor" docDate="2022" docId="32696ED440875D6E924D11DB484CEEF2" docLanguage="en" docName="ArthropodSystPhyl 80: 575-602" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-11-18" docSource="http://dx.doi.org/10.3897/asp.80.e84013" docTitle="Rumilara Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr 2022, gen. nov." docType="treatment" docUuid="111B1F47-F9E2-4654-B112-17B9ACC6D77A" docUuidSource="ZooBank" docVersion="1" id="81B59BA932BC5DAF90815FD08F5820A6" lastPageNumber="575" masterDocId="81B59BA932BC5DAF90815FD08F5820A6" masterDocTitle="Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)" masterLastPageNumber="602" masterPageNumber="575" pageNumber="575" updateTime="1668813875820" updateUser="pensoft">
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<mods:title>Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Linsky, Marek</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7206-2380</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ciamporova-Zatovicova, Zuzana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0506-6212</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lassova, Kristina</mods:namePart>
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<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ciampor Jr, Fedor</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6269-3592</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
<mods:nameIdentifier type="email">f.ciampor@savba.sk</mods:nameIdentifier>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-11-18</mods:number>
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<mods:number>80</mods:number>
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<mods:start>575</mods:start>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e84013</mods:url>
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<treatment LSID="urn:lsid:zoobank.org:act:111B1F47-F9E2-4654-B112-17B9ACC6D77A" httpUri="http://treatment.plazi.org/id/32696ED440875D6E924D11DB484CEEF2" lastPageNumber="575" pageId="0" pageNumber="575">
<subSubSection pageId="0" pageNumber="575" type="nomenclature">
<paragraph pageId="0" pageNumber="575">
3.2.2. 
<taxonomicName LSID="https://zoobank.org/111B1F47-F9E2-4654-B112-17B9ACC6D77A" authority="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr, 2022" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus" status="gen. nov.">Rumilara</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="575">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="type species">
<paragraph pageId="0" pageNumber="575">Type species.</paragraph>
<paragraph pageId="0" pageNumber="575">
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara obscura" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="diagnosis">
<paragraph pageId="0" pageNumber="575">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="575">
A new genus is recognized by the combination of the following characters: compact structure of antennae (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">5a-e</figureCitation>
); mandibles with teeth fused (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mouthparts of Hexanchorus cordillierae (a) and Rumilara obscura sp. nov. (b - i); a adult mandible; b adult mandible; c adult maxilla; d adult labium (right palp omitted); e larval mandible; f larval antenna; g larval labrum; h larval labium; i larval maxilla. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure4" httpUri="https://binary.pensoft.net/fig/770427" pageId="0" pageNumber="575">4b</figureCitation>
); surface of pronotum microreticulate, with complete transverse impression before middle, outline of discal area elevated, partially or fully interrupted medially by a prescutellar indentation (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">6</figureCitation>
), mesotibiae without pubescence on lateral sides, without tubercle on inner apex in males; fourth tarsomere with apicoventral, long, erect seta; abdomen convex, with apical segment rounded in both sexes (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">8</figureCitation>
).
</paragraph>
<caption doi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 2.</emphasis>
Dorsal habiti of 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
species: 
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
<taxonomicName lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
<taxonomicName lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">e</emphasis>
<taxonomicName lsidName="R. leleupi" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">R. leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
male holotype. Scale: 0.5mm.
</paragraph>
</caption>
<caption doi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 3.</emphasis>
Lateral habiti of 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
species. 
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
<taxonomicName lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
<taxonomicName lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">e</emphasis>
<taxonomicName lsidName="R. leleupi" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">R. leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
male holotype. Scale: 0.5mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">Description.</paragraph>
<paragraph pageId="0" pageNumber="575">
Body elongate (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">2</figureCitation>
- 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">3</figureCitation>
), 2.44-3.28 mm long, subparallel, widest near posterior fifth of elytra (EW), 0.83-1.37 mm wide, dorsum moderately convex. Dorsal surface covered, at least partially, with two types of setae: 1) short, prone, more or less densely pubescent, and 2) numerous moderately long, curved, semi-erect, hair-like. Ventral surface clothed with yellowish, densely set, short to moderately long, recumbent pubescence that is confined partly or fully to following areas: genae, ventral face of head, hypomera, epipleura, prosternum, mesoventrite, metaventrite, abdomen, coxae, trochanters, femora, tibiae and tarsi. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Head</emphasis>
narrower than pronotum, retractile, dorsally moderately convex, laterally rounded, ventrally nearly flat. Surface microreticulate, with fine sparse or very dense punctation partly or fully concealed by pubescence. Eyes well developed, hemispherical; with row of long, hair-like setae along dorsal margin (often peeled off in 
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
and 
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
specimens). Labrum about 3 times wider than long; transverse; with row of long, yellowish, hair-like setae near anterior margin; anterolateral angles rounded; surface glabrous; anterior margin feebly emarginate medially. Clypeus shorter than labrum, transverse, surface microreticulate, fully or not at all concealed by pubescence, anterior margin with a row of moderately long, hair-like setae; frontoclypeal suture straight. Antennae (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">5a-e</figureCitation>
) 11-segmented; clubbed, club ca. twice as long as combined length of scape and pedicel. Scape curved, ca. 1.5 times as long as pedicel; pedicel subglobular. Segment 3 subtriangular, longer than wide; segment 4 about as long as wide; segments 5 to 10 wider than long, transverse; segment 11 about as long as wide, subglobular. All segments densely clothed with short, prone setae; scape and pedicel also with long, stout, hair-like setae; segments 5 to 11 with a few erect, hair-like setae. Mandible (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mouthparts of Hexanchorus cordillierae (a) and Rumilara obscura sp. nov. (b - i); a adult mandible; b adult mandible; c adult maxilla; d adult labium (right palp omitted); e larval mandible; f larval antenna; g larval labrum; h larval labium; i larval maxilla. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure4" httpUri="https://binary.pensoft.net/fig/770427" pageId="0" pageNumber="575">4b</figureCitation>
) short, moderately broad, nearly symmetrical with apical teeth fused; outer lateral margin strongly curved at base with a few long, curved, hair-like setae; ventral condyle bearing row of long, hair-like setae; molar part large, finely grooved. Prostheca hyaline, long, with numerous apical finger-like outgrowths. Maxilla (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mouthparts of Hexanchorus cordillierae (a) and Rumilara obscura sp. nov. (b - i); a adult mandible; b adult mandible; c adult maxilla; d adult labium (right palp omitted); e larval mandible; f larval antenna; g larval labrum; h larval labium; i larval maxilla. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure4" httpUri="https://binary.pensoft.net/fig/770427" pageId="0" pageNumber="575">4c</figureCitation>
) with cardo densely clothed with hair-like setae; stipes subtriangular, with moderately long, densely arranged, hair-like setae; palpifer nearly half as long as stipes, with moderately long, hair-like setae on external face, and with long, hair-like setae near anterior margin. Maxillary palpi 4-segmented; surface of segments 2 and 3 obscured by densely arranged, moderately long, hair-like setae; segment 1 very short, about as long as wide; segment 2 about twice as long as wide, widening apically; segment 3 slightly longer than the previous segment; terminal segment longest. Galea elongate, external outline convex, inner outline concave; apex rounded, with numerous slightly longer, finger-like setae; outer outline densely clothed with long, hair-like setae. Lacinia about twice as wide as galea, longer than wide, subrectangular; clothed with long, densely arranged, hair-like setae and apically with transverse rows of stout, bent, finger-like setae. Labium (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mouthparts of Hexanchorus cordillierae (a) and Rumilara obscura sp. nov. (b - i); a adult mandible; b adult mandible; c adult maxilla; d adult labium (right palp omitted); e larval mandible; f larval antenna; g larval labrum; h larval labium; i larval maxilla. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure4" httpUri="https://binary.pensoft.net/fig/770427" pageId="0" pageNumber="575">4d</figureCitation>
) with palpiger short, clothed with tuft of very long, densely set, hair-like setae. Labial palpi 3-segmented; segments 1 and 2 subequal in length, with tuft of conspicuous, long, hair-like setae; terminal segment largest, suboval, widest apically, apex truncate, with a field of short, finger-like setae. Ligula almost as long as palpi, widest anteriorly; anterolateral angles broadly rounded; anterior and anterolateral margin with densely arranged, moderately long, curved, hair-like setae; medial portion covered with short, stout, finger-like setae. Prementum short, transverse, widest posteriorly; mentum longer then prementum, densely clothed with long, semi-erect, hair-like setae. 
<emphasis bold="true" pageId="0" pageNumber="575">
<emphasis italics="true" pageId="0" pageNumber="575">Thorax</emphasis>
.
</emphasis>
Pronotum (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">6</figureCitation>
) wider than long, subparallel, widest near middle; disc convex, posteriorly to transverse impression distinctly elevated, partially of fully interrupted medially by prescutellar indentation, surface microreticulate; with broad transverse impression in anterior third; sublateral carinae absent; with arcuate impression near posterolateral sides; anterior margin arcuate; anterolateral angles rounded; lateral margins almost straight in anterior third then broadly rounded; posterior margin trisinuate, broadly arcuate on each side and narrowly in front of scutellum; posterior angles suborthogonal. Surface microreticulate (feebly visible in 
<taxonomicName lsidName="R. leleupi" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">R. leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
). Hypomeron widest in anterior third, then distinctly narrowed, almost straight. Prosternum (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">7</figureCitation>
) extremely short in front of procoxae; procoxae strongly transverse in lateral view; prosternal process wide; parallel-sided between procoxae, then straight or concavely tapering towards rounded apex. Mesoventrite short and wide, with deep narrow groove for reception of prosternal process. Metaventrite slightly wider than long, moderately smooth, disc convex with a narrow longitudinal impression on midline. Elytra elongate, parallel or finely subparallel, widest in posterior 4/5, then evenly arched towards rounded apex. Each elytron with 10 rows of shallowly to deeply impressed punctures. Intervals nearly flat, slightly elevated or partly depressed. Scutellum wider than long, rounded. Epipleuron oblique until posterior margin of metaventrite then inflexed horizontally, posteriorly tapering, not reaching elytral apex. Hind wing without radial cross vein; without anal cell; with first and second branches of anal vein 2 fused; anal vein 1 present only apically; cubito-anal cross vein incomplete or complete and joining cubitus to anal vein 1. Legs long, at least partially covered by pubescence; femora widest across middle; tibiae moderately bent, subequal in length with femora. Tarsi 5-segmented, fourth segment with an apicoventral, long, erect seta, fifth segment slightly shorter than remaining segments combined; claws strong, long, without basal or subbasal teeth. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Abdomen</emphasis>
(Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">8</figureCitation>
) with 5 ventrites; first ventrite with intercoxal process subtriangular, about twice as wide as long, feebly incissed medially; ventrites 2-4 gradually shorter and slightly narrower than previous one; posterior margin of ventrite 5 broadly rounded. 
<emphasis bold="true" pageId="0" pageNumber="575">
<emphasis italics="true" pageId="0" pageNumber="575">Male</emphasis>
.
</emphasis>
Sternite VIII (Figs 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Rumilara obscura sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure11" httpUri="https://binary.pensoft.net/fig/770434" pageId="0" pageNumber="575">11e</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Rumilara suppressa sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure12" httpUri="https://binary.pensoft.net/fig/770435" pageId="0" pageNumber="575">12e</figureCitation>
, 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Rumilara riberai sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure13" httpUri="https://binary.pensoft.net/fig/770436" pageId="0" pageNumber="575">13e</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rumilara paterna sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure14" httpUri="https://binary.pensoft.net/fig/770437" pageId="0" pageNumber="575">14d</figureCitation>
, 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rumilara leleupi comb. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure15" httpUri="https://binary.pensoft.net/fig/770438" pageId="0" pageNumber="575">15d</figureCitation>
) with long, rather wide medial apodeme; apically broadly bisinuate, densely set with moderately long, curved, semi-erect, hair-like setae. Aedeagus of trilobate type; long and slender; penis long, tapering apically, corona present, with or without long, dorsal, rod-like sclerite, membranous sack with one smaller and one large, oblong ventral sclerites; parameres feebly to strongly asymmetrical, reaching from below half to 4/5 of penis; phallobase longer than penis. 
<emphasis bold="true" pageId="0" pageNumber="575">
<emphasis italics="true" pageId="0" pageNumber="575">Female</emphasis>
.
</emphasis>
Sternite VIII (Figs 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Rumilara obscura sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure11" httpUri="https://binary.pensoft.net/fig/770434" pageId="0" pageNumber="575">11f</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Rumilara suppressa sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure12" httpUri="https://binary.pensoft.net/fig/770435" pageId="0" pageNumber="575">12f</figureCitation>
, 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Rumilara riberai sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure13" httpUri="https://binary.pensoft.net/fig/770436" pageId="0" pageNumber="575">13f</figureCitation>
) with long, narrow, medial apodeme; apically broadly bi- or quadrisinuate, densely set with moderately long, curved, semi-erect hairs. Ovipositor (Figs 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Rumilara obscura sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure11" httpUri="https://binary.pensoft.net/fig/770434" pageId="0" pageNumber="575">11d</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Rumilara suppressa sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure12" httpUri="https://binary.pensoft.net/fig/770435" pageId="0" pageNumber="575">12d</figureCitation>
, 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Rumilara riberai sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure13" httpUri="https://binary.pensoft.net/fig/770436" pageId="0" pageNumber="575">13d</figureCitation>
) with coxites and styli short; coxites broad at base; struts long and slender.
</paragraph>
<caption doi="10.3897/asp.80.e84013.figure4" httpUri="https://binary.pensoft.net/fig/770427" pageId="0" pageNumber="575" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 4.</emphasis>
Mouthparts of 
<taxonomicName baseAuthorityName="Guerin Meneville" baseAuthorityYear="1843" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus cordillierae" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="species" species="cordillierae">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus cordillierae</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
) and 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara obscura" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
(
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="575">i</emphasis>
); 
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
adult mandible; 
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
adult mandible; 
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
adult maxilla; 
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
adult labium (right palp omitted); 
<emphasis bold="true" pageId="0" pageNumber="575">e</emphasis>
larval mandible; 
<emphasis bold="true" pageId="0" pageNumber="575">f</emphasis>
larval antenna; 
<emphasis bold="true" pageId="0" pageNumber="575">g</emphasis>
larval labrum; 
<emphasis bold="true" pageId="0" pageNumber="575">h</emphasis>
larval labium; 
<emphasis bold="true" pageId="0" pageNumber="575">i</emphasis>
larval maxilla. Scale: 0.1mm.
</paragraph>
</caption>
<caption doi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 5.</emphasis>
Antennae of 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
and 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
: 
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
<taxonomicName lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
<taxonomicName lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="575">e</emphasis>
<taxonomicName lsidName="R. leleupi" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">R. leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="575">f</emphasis>
<taxonomicName lsidName="H. mcdiarmidi" pageId="0" pageNumber="575" rank="species" species="mcdiarmidi">
<emphasis italics="true" pageId="0" pageNumber="575">H. mcdiarmidi</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">g</emphasis>
<taxonomicName lsidName="H. virilis" pageId="0" pageNumber="575" rank="species" species="virilis">
<emphasis italics="true" pageId="0" pageNumber="575">H. virilis</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">h</emphasis>
<taxonomicName lsidName="H. shannoni" pageId="0" pageNumber="575" rank="species" species="shannoni">
<emphasis italics="true" pageId="0" pageNumber="575">H. shannoni</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">i</emphasis>
<taxonomicName lsidName="H. angeli" pageId="0" pageNumber="575" rank="species" species="angeli">
<emphasis italics="true" pageId="0" pageNumber="575">H. angeli</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">j</emphasis>
<taxonomicName lsidName="H. rostratus" pageId="0" pageNumber="575" rank="species" species="rostratus">
<emphasis italics="true" pageId="0" pageNumber="575">H. rostratus</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">k</emphasis>
<taxonomicName lsidName="H. cordillierae" pageId="0" pageNumber="575" rank="species" species="cordillierae">
<emphasis italics="true" pageId="0" pageNumber="575">H. cordillierae</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">l</emphasis>
<taxonomicName lsidName="H. onorei" pageId="0" pageNumber="575" rank="species" species="onorei">
<emphasis italics="true" pageId="0" pageNumber="575">H. onorei</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">m</emphasis>
<taxonomicName lsidName="H. tarsalis" pageId="0" pageNumber="575" rank="species" species="tarsalis">
<emphasis italics="true" pageId="0" pageNumber="575">H. tarsalis</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">n</emphasis>
<taxonomicName lsidName="H. usitatus" pageId="0" pageNumber="575" rank="species" species="usitatus">
<emphasis italics="true" pageId="0" pageNumber="575">H. usitatus</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">o</emphasis>
<taxonomicName lsidName="H. gracilipes" pageId="0" pageNumber="575" rank="species" species="gracilipes">
<emphasis italics="true" pageId="0" pageNumber="575">H. gracilipes</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">p</emphasis>
<taxonomicName lsidName="H. caraibus" pageId="0" pageNumber="575" rank="species" species="caraibus">
<emphasis italics="true" pageId="0" pageNumber="575">H. caraibus</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">q</emphasis>
<taxonomicName lsidName="H. crinitus" pageId="0" pageNumber="575" rank="species" species="crinitus">
<emphasis italics="true" pageId="0" pageNumber="575">H. crinitus</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">r</emphasis>
<taxonomicName lsidName="H. dimorphus" pageId="0" pageNumber="575" rank="species" species="dimorphus">
<emphasis italics="true" pageId="0" pageNumber="575">H. dimorphus</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">s</emphasis>
<taxonomicName lsidName="H. emarginatus" pageId="0" pageNumber="575" rank="species" species="emarginatus">
<emphasis italics="true" pageId="0" pageNumber="575">H. emarginatus</emphasis>
</taxonomicName>
; 
<emphasis bold="true" pageId="0" pageNumber="575">t</emphasis>
<taxonomicName lsidName="H. browni" pageId="0" pageNumber="575" rank="species" species="browni">
<emphasis italics="true" pageId="0" pageNumber="575">H. browni</emphasis>
</taxonomicName>
. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="etymology">
<paragraph pageId="0" pageNumber="575">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="575">
The name of the genus is inspired by the language of the indigenous Quechua people who live in the territory where the new genus originated. In the Quechua language (
<bibRefCitation author="Gomez Rendon, JR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="575" publicationUrl="http://wold.clld.org/vocabulary/37" refId="B14" refString="Gomez Rendon, JR, 2009. Imbabura Quechua vocabulary. In: Haspelmath, Martin &amp; Tadmor, Uri (eds.) World Loanword Database. Leipzig: Max Planck Institute for Evolutionary Anthropology, 1319 entries. il (Available online at http://wold.clld.org/vocabulary/37" title="Imbabura Quechua vocabulary. In: Haspelmath, Martin &amp; Tadmor, Uri (eds.) World Loanword Database. Leipzig: Max Planck Institute for Evolutionary Anthropology, 1319 entries. il (Available online at" url="http://wold.clld.org/vocabulary/37" year="2009">
<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
<normalizedToken originalValue="Rendón">Rendon</normalizedToken>
2009
</bibRefCitation>
), &quot; 
<emphasis italics="true" pageId="0" pageNumber="575">rumi</emphasis>
&quot; means rock, referring to the nature of the type locality and other habitats of the species, i.e., steep rocks with water flowing in thin layer, or fast-flowing streams with bottoms composed largely of larger rocks. The adjective ' 
<emphasis italics="true" pageId="0" pageNumber="575">lara</emphasis>
' refers to type-genus of the subfamily 
<taxonomicName authorityName="LeConte" authorityYear="1861" lsidName="" pageId="0" pageNumber="575" rank="subFamily" subFamily="Larainae">Larainae</taxonomicName>
. Gender feminine.
</paragraph>
<caption doi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 6.</emphasis>
Pronota of 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
species: 
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
<taxonomicName lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
<taxonomicName lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
male holotype; 
<emphasis bold="true" pageId="0" pageNumber="575">e</emphasis>
<taxonomicName lsidName="R. leleupi" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">R. leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
male holotype. Scale: 0.2mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="distribution">
<paragraph pageId="0" pageNumber="575">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="575">
The genus is distributed in central Ecuador, so far known from four provinces (Cotopaxi Province, Napo Province, Morona-Santiago Province, Santo Domingo de los 
<normalizedToken originalValue="Tsáchilas">Tsachilas</normalizedToken>
Province).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="comparisons">
<paragraph pageId="0" pageNumber="575">Comparative notes.</paragraph>
<paragraph pageId="0" pageNumber="575">
The new genus is the closest to 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
, both share a combination of following synapomorphies which separate them from remaining 
<taxonomicName authorityName="LeConte" authorityYear="1861" lsidName="" pageId="0" pageNumber="575" rank="subFamily" subFamily="Larainae">Larainae</taxonomicName>
genera: smaller body size (length never reaches 5 mm); pronotum with distinct transverse impression in anterior half; hind wing venation without anal cell; apex of fourth tarsal segment ventrally with a fine nearly erect seta; and membranous sac of penis with laterally placed fibula. Adults of 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
differs from 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
mainly in: (i) microreticulate pronotum (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">6</figureCitation>
) (less distinct in 
<taxonomicName lsidName="R. leleupi" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">R. leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
); outline of discal area elevated (nearly flat in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
); discal area partially to fully interrupted by prescutellar indentation in middle (medial impression only superficial in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
); transverse impression shifted more to the middle (anterior third in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
); (ii) mesotibiae without pubescence (e.g., Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">3c-e</figureCitation>
) on lateral sides, without tubercle on inner apex (lateral pubescence is present in all 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
species, all males have tubercle on inner apex); (iii) abdomen (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">8</figureCitation>
) with fifth ventrite apically rounded, ventrites convex (in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
fifth ventrite is apically emarginate in both sexes, male has ventrites impressed, female convex); (iv) male genitalia (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Rumilara obscura sp. nov. (a - d): a prosternum and mesosternum; b foreleg (without coxa); c midleg; d hindleg. Schematic illustrations: e midleg of Hexanchorus cordillierae (alp - anterior lateral pubescence; mst - mesotibial tubercle; plp - posterior lateral pubescence; vs - ventral seta); f aedeagus of Rumilara riberai sp. nov. (ap - apophyse; co - corona; df - dorsal fibula; en - endophallus; mi - fibula minor; mj - fibula major; ml - median lobe; pa - paramere; ph - phallobase). Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure10" httpUri="https://binary.pensoft.net/fig/770433" pageId="0" pageNumber="575">10f</figureCitation>
) with fibula major and minor (only major in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
), some species with dorsal fibula (rod-like sclerite unknown in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
); (v) antennae (Figs 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">5a-e</figureCitation>
) more compact, almost club-like (more loosely connected in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
; Figs 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">5f-t</figureCitation>
), and (vi) mandibula (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mouthparts of Hexanchorus cordillierae (a) and Rumilara obscura sp. nov. (b - i); a adult mandible; b adult mandible; c adult maxilla; d adult labium (right palp omitted); e larval mandible; f larval antenna; g larval labrum; h larval labium; i larval maxilla. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure4" httpUri="https://binary.pensoft.net/fig/770427" pageId="0" pageNumber="575">4b</figureCitation>
) with apical teeth fused (as opposed to three apical teeth present in 
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus</emphasis>
</taxonomicName>
; Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mouthparts of Hexanchorus cordillierae (a) and Rumilara obscura sp. nov. (b - i); a adult mandible; b adult mandible; c adult maxilla; d adult labium (right palp omitted); e larval mandible; f larval antenna; g larval labrum; h larval labium; i larval maxilla. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure4" httpUri="https://binary.pensoft.net/fig/770427" pageId="0" pageNumber="575">4a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="remarks">
<paragraph pageId="0" pageNumber="575">Remark.</paragraph>
<paragraph pageId="0" pageNumber="575">
Three 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara" order="Coleoptera" pageId="0" pageNumber="575" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">gen. nov.</emphasis>
species (
<taxonomicName lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
, 
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
, 
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
) differ from 
<taxonomicName lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
and 
<taxonomicName lsidName="R. leleupi" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">R. leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
by femoral pubescence, which is restricted to long, dark, densely set, hair-like setae along dorsal margin, and by pro- and metatibial pubescence that is in a form of fringe at tibial apex.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="included species">
<paragraph pageId="0" pageNumber="575">Included species.</paragraph>
<paragraph pageId="0" pageNumber="575">
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" baseAuthorityName="Deleve" baseAuthorityYear="1968" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara leleupi" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
, 
<taxonomicName lsidName="R. obscura" pageId="0" pageNumber="575" rank="species" species="obscura">
<emphasis italics="true" pageId="0" pageNumber="575">R. obscura</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
, 
<taxonomicName lsidName="R. paterna" pageId="0" pageNumber="575" rank="species" species="paterna">
<emphasis italics="true" pageId="0" pageNumber="575">R. paterna</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
, 
<taxonomicName lsidName="R. riberai" pageId="0" pageNumber="575" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="575">R. riberai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
, 
<taxonomicName lsidName="R. suppressa" pageId="0" pageNumber="575" rank="species" species="suppressa">
<emphasis italics="true" pageId="0" pageNumber="575">R. suppressa</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">sp. nov.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>