170 lines
16 KiB
XML
170 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.554.6065" ID-GBIF-Dataset="9c0b4691-c23f-4429-9d2f-2bd8334f7401" ID-PMC="PMC4740830" ID-Pensoft-Pub="1313-2970-554-69" ID-PubMed="26877680" ID-ZBK="04FDA81C59CB4172B3FBFD57BFCD62E3" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-554-69" ModsDocOrigin="ZooKeys 554" ModsDocTitle="A new, widely distributed species of the Exocelinaekari-group from West Papua (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1453145179725" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Panjaitan, Rawati & Balke, Michael" docDate="2016" docId="E9ADC7E0079C3D141839E5BEC974419D" docLanguage="en" docName="ZooKeys 554: 69-85" docOrigin="ZooKeys 554" docSource="http://dx.doi.org/10.3897/zookeys.554.6065" docTitle="Exocelina manokwariensis subsp. nokensis Shaverdo, Panjaitan & Balke, 2016, ssp. n." docType="treatment" docUuid="9F65FAC4-58E8-4667-9C12-236CB7F46016" docUuidSource="ZooBank" docVersion="6" lastPageNumber="82" masterDocId="7E21AA01FFBFFF99FFD1F417ED1D9818" masterDocTitle="A new, widely distributed species of the Exocelinaekari-group from West Papua (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="85" masterPageNumber="69" pageNumber="79" updateTime="1668162486955" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>A new, widely distributed species of the Exocelinaekari-group from West Papua (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Shaverdo, Helena</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Panjaitan, Rawati</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Balke, Michael</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2016</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>554</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>69</mods:start>
|
|
<mods:end>85</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.554.6065</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.554.6065</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-554-69</mods:identifier>
|
|
<mods:identifier type="ZBK">04FDA81C59CB4172B3FBFD57BFCD62E3</mods:identifier>
|
|
<mods:identifier type="ZooBank">04FDA81C59CB4172B3FBFD57BFCD62E3</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="127874692" LSID="urn:lsid:zoobank.org:act:9F65FAC4-58E8-4667-9C12-236CB7F46016" httpUri="http://treatment.plazi.org/id/E9ADC7E0079C3D141839E5BEC974419D" lastPageId="13" lastPageNumber="82" pageId="10" pageNumber="79">
|
|
<subSubSection pageId="10" pageNumber="79" type="multiple">
|
|
<paragraph pageId="10" pageNumber="79">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="10" pageNumber="79" type="nomenclature">
|
|
<paragraph pageId="10" pageNumber="79">
|
|
<taxonomicName LSID="http://zoobank.org/9F65FAC4-58E8-4667-9C12-236CB7F46016" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina manokwariensis" order="Coleoptera" pageId="10" pageNumber="79" phylum="Arthropoda" rank="subSpecies" species="manokwariensis" subSpecies="nokensis">Exocelina manokwariensis nokensis</taxonomicName>
|
|
<taxonomicNameLabel pageId="10" pageNumber="79">ssp. n.</taxonomicNameLabel>
|
|
Figs 7, 8
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="80" type="reference_group">
|
|
<paragraph pageId="11" pageNumber="80">
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="11" pageNumber="80" phylum="Arthropoda" rank="genus">
|
|
<pageBreakToken pageId="11" pageNumber="80" start="start">Exocelina</pageBreakToken>
|
|
</taxonomicName>
|
|
undescribed sp. MB1275:
|
|
<bibRefCitation author="Toussaint, EFA" journalOrPublisher="Nature Communications" pageId="15" pageNumber="84" pagination="1 - 10" title="The towering orogeny of New Guinea as a trigger for arthropod megadiversity." url="10.1038/ncomms5001" volume="1" year="2014">Toussaint et al. 2014</bibRefCitation>
|
|
: Supplementary figs 1-4, tab. 2.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="80" type="type locality">
|
|
<paragraph pageId="11" pageNumber="80">Type locality.</paragraph>
|
|
<paragraph pageId="11" pageNumber="80">
|
|
Indonesia: West Papua Province: Raja Ampat Regency, Waigeo Island, Waifoi, Mountain Nok,
|
|
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-0.08466666">00°05.08'S</geoCoordinate>
|
|
;
|
|
<geoCoordinate direction="east" orientation="longitude" precision="9" value="130.74316">130°44.59'E</geoCoordinate>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="80" type="type material">
|
|
<paragraph pageId="11" pageNumber="80">Type material.</paragraph>
|
|
<paragraph pageId="11" pageNumber="80">Holotype: male "Indonesia: Papua, Waigeo, Waifoi, Mt. Nok, 500m, 00.05.076S 130.44.586E (BH 11)" (MZB). Paratypes: 25 males, 39 females with the same labels as the holotype, one male additionally with a green label "M.Balke 1275" (MZB, NHMW, ZSM). 1 male "Indonesia: Papua, Waigeo, Waifoi, <50m, 00.06.088S 130.42.855E, Balke (BH 10)", "M.Balke 1274" [green] (ZSM). 23 males "N.DUTCH NEW GUINEA: Waigeu. Camp Nok. 2,500 ft. iv.1938. L.E.Cheesman. B.M.1938-593.", one of them additionally with labels "collection 28", "measured J.Parkin 78" (BMNH). 5 males "N.DUTCH NEW GUINEA: Waigeu.Camp 1.Mt.Nok. 2,500 ft. v.1938. L.E.Cheesman. B.M.1938-593." (BMNH).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="80" type="materials_examined">
|
|
<paragraph pageId="11" pageNumber="80">Additional material.</paragraph>
|
|
<paragraph pageId="11" pageNumber="80">
|
|
27 females "N.DUTCH NEW GUINEA: Waigeu. Camp Nok. 2,500 ft. iv.1938. L.E.Cheesman. B.M.1938-593.", one of them additionally with labels "collection 27", "measured J.Parkin 76" (BMNH). 6 females "N.DUTCH NEW GUINEA: Waigeu.Camp 1.Mt.Nok. 2,500 ft. v.1938. L.E.Cheesman. B.M.1938-593." (BMNH). These females might belong to two species:
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina manokwariensis" order="Coleoptera" pageId="11" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="manokwariensis" subSpecies="nokensis">Exocelina manokwariensis nokensis</taxonomicName>
|
|
ssp. n. and
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina waigeoensis" order="Coleoptera" pageId="11" pageNumber="80" phylum="Arthropoda" rank="species" species="waigeoensis">Exocelina waigeoensis</taxonomicName>
|
|
Shaverdo, Hendrich & Balke, 2012.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="80" type="description">
|
|
<paragraph pageId="11" pageNumber="80">Description.</paragraph>
|
|
<paragraph pageId="11" pageNumber="80">As nominative subspecies, except for the following characters.</paragraph>
|
|
<paragraph pageId="11" pageNumber="80">Size: TL-H 3.1-3.6 mm, TL 3.45-4.0 mm, MW 1.65-1.9 mm; holotype: TL-H 3.4 mm, TL 3.75 mm, MW 1.8 mm (Fig. 7).</paragraph>
|
|
<caption pageId="11" pageNumber="80">
|
|
<paragraph pageId="11" pageNumber="80">
|
|
Figure 7. Habitus and coloration of
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina manokwariensis" order="Coleoptera" pageId="11" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="manokwariensis" subSpecies="nokensis">Exocelina manokwariensis nokensis</taxonomicName>
|
|
ssp. n.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="11" pageNumber="80">Surface sculpture: Dorsal microreticulation, especially on head and pronotum, slightly weaker.</paragraph>
|
|
<paragraph pageId="11" pageNumber="80">Structures: Base of prosternum and neck of prosternal process with distinct ridge, less rounded, smooth or with few transverse lines anteriorly.</paragraph>
|
|
<paragraph pageId="11" pageNumber="80">Male: Protarsomere 5 ventrally with anterior row of 9 and posterior row 4 short setae (Fig. 8B). Abdominal ventrite 6 with 3-6 lateral striae on each side. Median lobe slightly larger and somehow more slender. Its apex with relatively short tip and truncate margin almost straight in lateral view and asymmetric in ventral view (Fig. 8C, D). Subdistal part of paramere with less numerous, thick flattened setae (Fig. 8E).</paragraph>
|
|
<caption pageId="11" pageNumber="80">
|
|
<paragraph pageId="11" pageNumber="80">
|
|
Figure 8.
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina manokwariensis" order="Coleoptera" pageId="11" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="manokwariensis" subSpecies="nokensis">Exocelina manokwariensis nokensis</taxonomicName>
|
|
ssp. n. A male antenna B male protarsomeres 4-5 in ventral view C median lobe in ventral view D median lobe in lateral view E paramere in external view.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="11" pageNumber="80">Female: Antennae simple, abdominal ventrite 6 broadly rounded apically, without striae.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="13" lastPageNumber="82" pageId="12" pageNumber="81" type="comparation with the other subspecies and co-inhabiting species">
|
|
<paragraph pageId="12" pageNumber="81">
|
|
<pageBreakToken pageId="12" pageNumber="81" start="start">Comparation</pageBreakToken>
|
|
with the other subspecies and co-inhabiting species.
|
|
</paragraph>
|
|
<paragraph lastPageId="13" lastPageNumber="82" pageId="12" pageNumber="81">
|
|
Although widely separated geographically,
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina manokwariensis" order="Coleoptera" pageId="12" pageNumber="81" phylum="Arthropoda" rank="subSpecies" species="manokwariensis" subSpecies="nokensis">Exocelina manokwariensis nokensis</taxonomicName>
|
|
ssp. n. is very similar to the nominative subspecies, from which can be separated by the larger and somehow more slender median lobe, less numerous flattened setae on the paramere, and its finer dorsal
|
|
<pageBreakToken pageId="13" pageNumber="82" start="start">microreticulation</pageBreakToken>
|
|
. This subspecies was collected together with two other species of the
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ekari" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="ekari">Exocelina ekari</taxonomicName>
|
|
-group:
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina evelyncheesmanae" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="evelyncheesmanae">Exocelina evelyncheesmanae</taxonomicName>
|
|
Shaverdo, Hendrich & Balke, 2012 and
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina waigeoensis" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="waigeoensis">Exocelina waigeoensis</taxonomicName>
|
|
Shaverdo, Hendrich & Balke, 2012; see descriptions and illustrations in
|
|
<bibRefCitation author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="15" pageNumber="84" pagination="1 - 76" title="Introduction of the Exocelinaekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al. 2012</bibRefCitation>
|
|
. From the former (both males and females), it can be easily separated by its smaller size and distinctly modified male antennae (TL-H: 3.75-4.1 mm, MW: 1.9-2.2 mm and slightly modified male antennae: antennomeres 3-7 very slightly enlarged, antennomere 3 slightly more triangular than other antennomeres, in
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina evelyncheesmanae" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="evelyncheesmanae">Exocelina evelyncheesmanae</taxonomicName>
|
|
). From the latter, males can be distinguished by their distinctly modified antennae and the shapes of the median lobes (slightly modified antennae and apex of the median lobe elongate in lateral view in
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina waigeoensis" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="waigeoensis">Exocelina waigeoensis</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="13" pageNumber="82" type="distribution">
|
|
<paragraph pageId="13" pageNumber="82">Distribution.</paragraph>
|
|
<paragraph pageId="13" pageNumber="82">Indonesia: West Papua Province: Raja Ampat Regency, Waigeo Island (Fig. 9).</paragraph>
|
|
<caption pageId="13" pageNumber="82">
|
|
<paragraph pageId="13" pageNumber="82">
|
|
Figure 9. Map of the western part of New Guinea showing distribution of subspecies of
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina manokwariensis" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="manokwariensis">Exocelina manokwariensis</taxonomicName>
|
|
sp. n. and co-occurring species.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="13" pageNumber="82" type="etymology">
|
|
<paragraph pageId="13" pageNumber="82">Etymology.</paragraph>
|
|
<paragraph pageId="13" pageNumber="82">The species is named after Nok Mountain, the type locality of the species. The name is an adjective in the nominative singular.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="13" pageNumber="82" type="ecology">
|
|
<paragraph pageId="13" pageNumber="82">Ecology.</paragraph>
|
|
<paragraph pageId="13" pageNumber="82">
|
|
The species was collected in two localities on Nok Mountain together with
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina evelyncheesmanae" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="evelyncheesmanae">Exocelina evelyncheesmanae</taxonomicName>
|
|
and
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina waigeoensis" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="waigeoensis">Exocelina waigeoensis</taxonomicName>
|
|
and in one locality together with
|
|
<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina evelyncheesmanae" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" species="evelyncheesmanae">Exocelina evelyncheesmanae</taxonomicName>
|
|
in ration ca. 1:1.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |