300 lines
37 KiB
XML
300 lines
37 KiB
XML
<document id="C06912BAE967CD26C21F492CCA168ACC" ID-DOI="10.11646/zootaxa.3850.1.1" ID-GBIF-Dataset="1bfc2e63-3da5-448a-8ebe-71f9d25c9c5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="286868" ID-ZooBank="BFD1F483-4255-429B-9E17-8D4A9E559C5F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460882059799" checkinUser="plazi" docAuthor="Yin, Zi-Wei & Li, Li-Zhen" docDate="2014" docId="CD490758D857FFACFF0147F846DBF85A" docLanguage="en" docName="zt03850p083.pdf" docOrigin="Zootaxa 3850 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Horniella smetanai Yin and Li, new species" docType="treatment" docVersion="10" lastPageNumber="72" masterDocId="31707F20D810FFEBFF964509443BFFAF" masterDocTitle="Revision of the Oriental genus Horniella Raffray (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="72" updateTime="1698358480273" updateUser="plazi">
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<mods:mods id="68B679B22C9744854EA3F9D6E0A3FA55" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="30C310A565D1BB1A451D9A4699BC7672">
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<mods:title id="2DE3D89799EBB3261AE16A01ED2E4B78">Revision of the Oriental genus Horniella Raffray (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="92D17794DF3061493845845D16719F37" type="personal">
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<mods:role id="BFA36DF2086297F55EAF31E9742ED443">
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<mods:roleTerm id="BF79C13159DB70A5014D69D58C47402E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="78164538888457C89E52452066E4E7DD">Yin, Zi-Wei</mods:namePart>
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</mods:name>
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<mods:name id="5CE550D8B1C1FEB347CBCD9E96227757" type="personal">
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<mods:role id="6C1A96D412801D12947603358B5934AB">
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<mods:roleTerm id="99F810C70311D9A3221929CB4E0D3FD5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="936665BE24832832AC5A72D244160BCB">Li, Li-Zhen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="EE56FA50BB89691E7C07DD6F7D839EFF">text</mods:typeOfResource>
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<mods:relatedItem id="73BBE48DEC4A1CEF9C4FB899E244D40A" type="host">
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<mods:titleInfo id="2BFD5213E9818CACD4668ABFB2AC855F">
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||
<mods:title id="3C2E8B1430E2DE11DD8942C707434924">Zootaxa</mods:title>
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</mods:titleInfo>
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||
<mods:part id="6F79A1369DA628C4AE647E765F8E2491">
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||
<mods:date id="7D7E057A71A2C826FE48629ADAB66880">2014</mods:date>
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<mods:detail id="BBA4B4A8746B537B53BCF97C6836BB05" type="pubDate">
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<mods:number id="EC8365E2EF4F5C5F0B504C7C1A022C3D">2014-08-12</mods:number>
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</mods:detail>
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<mods:detail id="4E002B649C2D51DDC4BC75D5C4CB16A6" type="volume">
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<mods:number id="DBB1418D947FAC9D8DBAB7F1464375B1">3850</mods:number>
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</mods:detail>
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<mods:detail id="F8A45CB6ED0CD6E9F31E3FCE1CA04F65" type="issue">
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<mods:number id="58ABA281F22A199DDA19984C75AFC840">1</mods:number>
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</mods:detail>
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<mods:extent id="0EB89F8AE7773529A4B4FC777F357AAA" unit="page">
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<mods:start id="1AF1F6BEC5723444ADEAA7218D78B5A5">1</mods:start>
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<mods:end id="B08B3BB9AE6A73F99AE8F656A02A58A3">83</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="B09865E8BB99D31A5865B985E9131C8F">journal article</mods:classification>
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<mods:identifier id="843D8E5D5268796ADC1449683B293C81" type="DOI">10.11646/zootaxa.3850.1.1</mods:identifier>
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<mods:identifier id="962236836935ACDBEB9B1908B4039AB0" type="GBIF-Dataset">1bfc2e63-3da5-448a-8ebe-71f9d25c9c5a</mods:identifier>
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<mods:identifier id="27F62A2B021CB9CF3BA5475E84C7B08D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="26602BA5A98D093632E459797B0CD325" type="Zenodo-Dep">286868</mods:identifier>
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<mods:identifier id="EABAFCCAF5BB1DE97E20AD24CD4E972A" type="ZooBank">BFD1F483-4255-429B-9E17-8D4A9E559C5F</mods:identifier>
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</mods:mods>
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<treatment id="CD490758D857FFACFF0147F846DBF85A" ID-DOI="http://doi.org/10.5281/zenodo.6142787" ID-GBIF-Taxon="193985273" ID-Zenodo-Dep="6142787" LSID="urn:lsid:plazi:treatment:CD490758D857FFACFF0147F846DBF85A" httpUri="http://treatment.plazi.org/id/CD490758D857FFACFF0147F846DBF85A" lastPageNumber="72" pageId="71" pageNumber="72">
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<subSubSection id="0DFAE5C5D857FFACFF0147F846E3FCA4" box="[151,728,753,779]" pageId="71" pageNumber="72" type="nomenclature">
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||
<paragraph id="455FB64ED857FFACFF0147F846E3FCA4" blockId="71.[151,728,753,813]" box="[151,728,753,779]" pageId="71" pageNumber="72">
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<heading id="1E170122D857FFACFF0147F846E3FCA4" bold="true" box="[151,728,753,779]" fontSize="11" level="1" pageId="71" pageNumber="72" reason="1">
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<emphasis id="77946A5CD857FFACFF0147F846E3FCA4" bold="true" box="[151,728,753,779]" pageId="71" pageNumber="72">
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26.
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<taxonomicName id="82E0CDCDD857FFACFF5547F84600FCA4" authority="Yin and Li" authorityName="Yin and Li" box="[195,571,753,779]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
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<emphasis id="77946A5CD857FFACFF5547F84597FCA4" bold="true" box="[195,428,753,779]" italics="true" pageId="71" pageNumber="72">Horniella smetanai</emphasis>
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Yin and Li
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</taxonomicName>
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,
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<taxonomicNameLabel id="6CA7D727D857FFACFDDE47F846E3FCA4" box="[584,728,753,779]" pageId="71" pageNumber="72" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0DFAE5C5D857FFACFF01461D4587FC82" box="[151,444,788,813]" pageId="71" pageNumber="72" type="description">
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<paragraph id="455FB64ED857FFACFF01461D4587FC82" blockId="71.[151,728,753,813]" box="[151,444,788,813]" pageId="71" pageNumber="72">
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<figureCitation id="DDDBAACBD857FFACFF01461D44D7FC83" box="[151,236,788,813]" captionStart="FIGURE 43" captionStartId="69.[151,250,1552,1574]" captionTargetBox="[159,1426,201,1525]" captionTargetId="figure@69.[151,1435,193,1531]" captionTargetPageId="69" captionText="FIGURE 43. Male habitus of Horniella species. A. H. pilosa sp. n. B. H. smetanai sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/286911/files/figure.png" pageId="71" pageNumber="72">Figs 43</figureCitation>
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B, 45, 49H;
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<figureCitation id="DDDBAACBD857FFACFEE2461C4587FC82" box="[372,444,788,813]" captionStart="MAP 6" captionStartId="68.[151,209,1635,1656]" captionTargetBox="[151,1435,913,1613]" captionTargetId="figure@68.[151,1435,913,1614]" captionTargetPageId="68" captionText="MAP 6. Distribution of Horniella species in Malaysia." figureDoi="http://doi.org/10.5281/zenodo.6353210" httpUri="https://zenodo.org/record/6353210/files/figure.png" pageId="71" pageNumber="72">Map 6</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0DFAE5C5D857FFACFF0146554767FB87" pageId="71" pageNumber="72" type="materials_examined">
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||
<paragraph id="455FB64ED857FFACFF0146554767FB87" blockId="71.[151,1437,860,2037]" pageId="71" pageNumber="72">
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<emphasis id="77946A5CD857FFACFF0146554573FCDA" bold="true" box="[151,328,860,885]" pageId="71" pageNumber="72">
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<typeStatus id="9A5B08ECD857FFACFF01465544E9FCDA" box="[151,210,860,885]" pageId="71" pageNumber="72">Type</typeStatus>
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material
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</emphasis>
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(4 ♂♂, 1 ♀).
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<materialsCitation id="F588BC13D857FFACFD8B4655457BFC13" ID-GBIF-Occurrence="3704617310" collectingDate="1987-06-19" collectionCode="MHNG" collectorName="Burckhardt- & Lobl" determinerName="Yin & Li" elevation="1200" location="Kota Kinabalu" municipality="Crocker Ra." pageId="71" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="SABAH" typeStatus="holotype">
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<typeStatus id="9A5B08ECD857FFACFD8B465546B3FCDA" box="[541,648,860,885]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="53E67DC7D857FFACFD084654468EFCDB" box="[670,693,861,884]" pageId="71" pageNumber="72" type="male">♂</specimenCount>
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, labeled ‘
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<collectingRegion id="872478ACD857FFACFCA5465547A0FCDB" box="[819,923,860,884]" pageId="71" pageNumber="72">SABAH</collectingRegion>
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:
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<collectingMunicipality id="A53B2C34D857FFACFC264655407FFCDB" box="[944,1092,860,885]" pageId="71" pageNumber="72">Crocker Ra.</collectingMunicipality>
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,
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<quantity id="82181BABD857FFACFBCD46554087FCDB" box="[1115,1212,860,884]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="71" pageNumber="72" unit="m" value="1200.0">
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||
<elevation id="CECD517DD857FFACFBCD46554087FCDB" box="[1115,1212,860,884]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="71" pageNumber="72" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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, km 63 rte
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<location id="403FE095D857FFACFAF0465444C5FC37" LSID="urn:lsid:plazi:treatment:CD490758D857FFACFF0147F846DBF85A:403FE095D857FFACFAF0465444C5FC37" municipality="Crocker Ra." name="Kota Kinabalu" pageId="71" pageNumber="72" stateProvince="SABAH">Kota Kinabalu</location>
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–
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<location id="403FE095D857FFACFE9A467645BEFC37" LSID="urn:lsid:plazi:treatment:CD490758D857FFACFF0147F846DBF85A:403FE095D857FFACFE9A467645BEFC37" box="[268,389,895,920]" municipality="Crocker Ra." name="Tambunan" pageId="71" pageNumber="72" stateProvince="SABAH">Tambunan</location>
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||
,
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||
<date id="315E908ED857FFACFE0646764631FC37" box="[400,522,895,920]" pageId="71" pageNumber="72" value="1987-06-19">
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||
<collectingDate id="211A6966D857FFACFE0646764631FC37" box="[400,522,895,920]" pageId="71" pageNumber="72" value="1987-06-19">19.vi.1987</collectingDate>
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||
</date>
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,
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<collectorName id="E815D398D857FFACFD81467646E0FC37" box="[535,731,895,920]" pageId="71" pageNumber="72">Burckhardt- Löbl</collectorName>
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/ Holotype [red], ♂,
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||
<taxonomicName id="82E0CDCDD857FFACFC5A4689409DFC38" box="[972,1190,896,920]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="smetanai" status="sp. nov.">
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||
<emphasis id="77946A5CD857FFACFC5A4689409DFC38" box="[972,1190,896,920]" italics="true" pageId="71" pageNumber="72">Horniella smetanai</emphasis>
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||
</taxonomicName>
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||
<emphasis id="77946A5CD857FFACFB38467640D5FC37" bold="true" box="[1198,1262,895,920]" pageId="71" pageNumber="72">
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||
<taxonomicNameLabel id="6CA7D727D857FFACFB38467640D5FC37" box="[1198,1262,895,920]" pageId="71" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, det.
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<determinerName id="B0835A8AD857FFACFAB84676416CFC37" box="[1326,1367,895,920]" pageId="71" pageNumber="72">Yin</determinerName>
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||
&
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<determinerName id="B0835A8AD857FFACFAEA467641ACFC37" box="[1404,1431,895,920]" pageId="71" pageNumber="72">Li</determinerName>
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||
, 2014,
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<collectionCode id="23F12E8BD857FFACFF4846AD457BFC13" box="[222,320,932,956]" pageId="71" pageNumber="72">MHNG’</collectionCode>
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</materialsCitation>
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.
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<materialsCitation id="F588BC13D857FFACFEDB46AC4714FC12" ID-GBIF-Occurrence="3704617303" box="[333,815,932,957]" collectingDate="1987-06-19" collectionCode="MHNG" collectorName="Burckhardt- & Lobl" determinerName="Yin & Li" elevation="1200" location="Kota Kinabalu" municipality="Crocker Ra." pageId="71" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="SABAH" typeStatus="paratype">
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||
<typeStatus id="9A5B08ECD857FFACFEDB46AC4585FC12" box="[333,446,933,957]" pageId="71" pageNumber="72" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="53E67DC7D857FFACFE5C46AD45C3FC13" box="[458,504,932,956]" pageId="71" pageNumber="72" type="generic">1 ♂</specimenCount>
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, same data as the holotype
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||
</materialsCitation>
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||
;
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<materialsCitation id="F588BC13D857FFACFCAB46AD462FFC4F" ID-GBIF-Occurrence="3704617308" collectingDate="1987-06-17" collectionCode="MHNG" collectorName="Burckhardt- & Lobl" determinerName="Yin & Li" elevation="1270" location="km 60 rte Kota" municipality="Crocker Ra." pageId="71" pageNumber="72" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="SABAH" typeStatus="paratype">
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||
<specimenCount id="53E67DC7D857FFACFCAB46AD4751FC13" box="[829,874,932,956]" pageId="71" pageNumber="72" type="generic">1 ♂</specimenCount>
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||
,
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<specimenCount id="53E67DC7D857FFACFCEE46AD479FFC13" box="[888,932,932,956]" pageId="71" pageNumber="72" type="female">1 ♀</specimenCount>
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||
, same data except ‘
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||
<quantity id="82181BABD857FFACFB1246AD40DAFC13" box="[1156,1249,932,956]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="71" pageNumber="72" unit="m" value="1270.0">
|
||
<elevation id="CECD517DD857FFACFB1246AD40DAFC13" box="[1156,1249,932,956]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="71" pageNumber="72" unit="m" value="1270.0">1270 m</elevation>
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||
</quantity>
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||
,
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||
<location id="403FE095D857FFACFB7846AD41A7FC13" LSID="urn:lsid:plazi:treatment:CD490758D857FFACFF0147F846DBF85A:403FE095D857FFACFB7846AD41A7FC13" box="[1262,1436,932,957]" name="km 60 rte Kota" pageId="71" pageNumber="72">km 60 rte Kota</location>
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||
<location id="403FE095D857FFACFF0146CE45BEFC4F" LSID="urn:lsid:plazi:treatment:CD490758D857FFACFF0147F846DBF85A:403FE095D857FFACFF0146CE45BEFC4F" box="[151,389,967,992]" name="Kinabalu - Tambunan" pageId="71" pageNumber="72">Kinabalu–Tambunan</location>
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||
,
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||
<date id="315E908ED857FFACFE0646CE462BFC4F" box="[400,528,967,992]" pageId="71" pageNumber="72" value="1987-06-17">
|
||
<collectingDate id="211A6966D857FFACFE0646CE462BFC4F" box="[400,528,967,992]" pageId="71" pageNumber="72" value="1987-06-17">17.vi.1987</collectingDate>
|
||
</date>
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||
’
|
||
</materialsCitation>
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||
;
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||
<materialsCitation id="F588BC13D857FFACFDB746C146A6FBAB" ID-GBIF-Occurrence="3704617307" collectingDate="1988-08-17" collectionCode="MHNG" collectorName="A. Smetana" country="BORNEO" elevation="790" location="Sabah Mt. Kinab. N. P." pageId="71" pageNumber="72" specimenCode="B119" specimenCount="1" typeStatus="paratype">
|
||
<specimenCount id="53E67DC7D857FFACFDB746C14675FC70" box="[545,590,968,992]" pageId="71" pageNumber="72" type="generic">1 ♂</specimenCount>
|
||
, labeled ‘
|
||
<collectingCountry id="3DF7F6DED857FFACFD2146C14714FC4F" box="[695,815,968,992]" pageId="71" pageNumber="72">BORNEO</collectingCountry>
|
||
<location id="403FE095D857FFACFCA146CE4004FC70" LSID="urn:lsid:plazi:treatment:CD490758D857FFACFF0147F846DBF85A:403FE095D857FFACFCA146CE4004FC70" box="[823,1087,967,992]" country="BORNEO" name="Sabah Mt. Kinab. N. P." pageId="71" pageNumber="72">Sabah Mt. Kinab. N. P.</location>
|
||
,
|
||
<location id="403FE095D857FFACFBDD46C141A3FC4F" LSID="urn:lsid:plazi:treatment:CD490758D857FFACFF0147F846DBF85A:403FE095D857FFACFBDD46C141A3FC4F" box="[1099,1432,967,992]" country="BORNEO" name="Por. H. S. area Eastern Ridge" pageId="71" pageNumber="72">Por. H. S. area Eastern Ridge</location>
|
||
, Tr.
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||
<quantity id="82181BABD857FFACFF2946E54531FBAB" box="[191,266,1004,1028]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="71" pageNumber="72" unit="m" value="790.0">
|
||
<elevation id="CECD517DD857FFACFF2946E54531FBAB" box="[191,266,1004,1028]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="71" pageNumber="72" unit="m" value="790.0">790 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="315E908ED857FFACFE8E46E54599FBAA" box="[280,418,1004,1029]" pageId="71" pageNumber="72" value="1988-08-17">
|
||
<collectingDate id="211A6966D857FFACFE8E46E54599FBAA" box="[280,418,1004,1029]" pageId="71" pageNumber="72" value="1988-08-17">17.viii.1988</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="E815D398D857FFACFE2646E5460DFBAB" box="[432,566,1004,1028]" pageId="71" pageNumber="72">A. Smetana</collectorName>
|
||
[
|
||
<specimenCode id="15461E35D857FFACFDD046E546B7FBAB" box="[582,652,1004,1028]" pageId="71" pageNumber="72">B119</specimenCode>
|
||
]’.
|
||
</materialsCitation>
|
||
All
|
||
<typeStatus id="9A5B08ECD857FFACFD4546E44704FBAA" box="[723,831,1005,1029]" pageId="71" pageNumber="72" type="paratype">paratypes</typeStatus>
|
||
are housed in
|
||
<collectionCode id="23F12E8BD857FFACFC7D46E5407BFBAB" box="[1003,1088,1004,1028]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="71" pageNumber="72">MHNG</collectionCode>
|
||
, and each bears a yellow
|
||
<typeStatus id="9A5B08ECD857FFACFAFA46E441A7FBAA" box="[1388,1436,1005,1029]" pageId="71" pageNumber="72">type</typeStatus>
|
||
label similar to that of the
|
||
<typeStatus id="9A5B08ECD857FFACFE284106461BFB87" box="[446,544,1039,1064]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
|
||
except ‘
|
||
<typeStatus id="9A5B08ECD857FFACFDEA411946D8FB87" box="[636,739,1040,1064]" pageId="71" pageNumber="72" type="paratype">Paratype</typeStatus>
|
||
♂ (or ♀)’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DFAE5C5D857FFACFF51413D4740F91F" pageId="71" pageNumber="72" type="description">
|
||
<paragraph id="455FB64ED857FFACFF51413D4577F98F" blockId="71.[151,1437,860,2037]" pageId="71" pageNumber="72">
|
||
<emphasis id="77946A5CD857FFACFF51413D4562FBE2" bold="true" box="[199,345,1076,1101]" pageId="71" pageNumber="72">Description.</emphasis>
|
||
Male (
|
||
<figureCitation id="DDDBAACBD857FFACFE3B413D4639FBE3" box="[429,514,1076,1101]" captionStart="FIGURE 43" captionStartId="69.[151,250,1552,1574]" captionTargetBox="[159,1426,201,1525]" captionTargetId="figure@69.[151,1435,193,1531]" captionTargetPageId="69" captionText="FIGURE 43. Male habitus of Horniella species. A. H. pilosa sp. n. B. H. smetanai sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/286911/files/figure.png" pageId="71" pageNumber="72">Fig. 43</figureCitation>
|
||
B). Length
|
||
<quantity id="82181BABD857FFACFD12413D4716FBE3" box="[644,813,1076,1101]" metricMagnitude="-3" metricUnit="m" metricValue="3.59" metricValueMax="3.64" metricValueMin="3.54" pageId="71" pageNumber="72" unit="mm" value="3.59" valueMax="3.64" valueMin="3.54">3.54–3.64 mm</quantity>
|
||
. Head distinctly longer than wide, HL
|
||
<quantity id="82181BABD857FFACFB66413D41A3FBE3" box="[1264,1432,1076,1100]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.1" metricValueMin="7.9" pageId="71" pageNumber="72" unit="mm" value="0.8" valueMax="0.81" valueMin="0.79">0.79–0.81 mm</quantity>
|
||
, HW
|
||
<quantity id="82181BABD857FFACFF58415E454EFBC0" box="[206,373,1111,1136]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="6.8" metricValueMin="6.2" pageId="71" pageNumber="72" unit="mm" value="0.65" valueMax="0.68" valueMin="0.62">0.62–0.68 mm</quantity>
|
||
; anterolateral genal projections (
|
||
<figureCitation id="DDDBAACBD857FFACFD7E415E4700FBDF" box="[744,827,1111,1136]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
C) weakly indicated; median sulcus between antennal tubercles short; scapes (
|
||
<figureCitation id="DDDBAACBD857FFACFE3C417545C6FB3A" box="[426,509,1148,1173]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
B) lacking expansion at lateral margins; clubs (
|
||
<figureCitation id="DDDBAACBD857FFACFB884175404AFB3A" box="[1054,1137,1148,1173]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
A) formed by apical three antennomeres, each distinctly elongate; venter lacking lateral spines (
|
||
<figureCitation id="DDDBAACBD857FFACFC274196403EFB17" box="[945,1029,1183,1208]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
D). Maxillary palpomeres II greatly elongate, slightly broadened at apical 1/3. Each eye composed of about 40–45 facets. Pronotum longer than wide, PL
|
||
<quantity id="82181BABD857FFACFF2841E14558FB50" box="[190,355,1256,1280]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.4" metricValueMin="8.1" pageId="71" pageNumber="72" unit="mm" value="0.825" valueMax="0.84" valueMin="0.81">0.81–0.84 mm</quantity>
|
||
, PW
|
||
<quantity id="82181BABD857FFACFE3641E1467DFB50" box="[416,582,1256,1280]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" metricValueMax="7.3" metricValueMin="7.1" pageId="71" pageNumber="72" unit="mm" value="0.72" valueMax="0.73" valueMin="0.71">0.71–0.73 mm</quantity>
|
||
. Elytra wider than long, EL
|
||
<quantity id="82181BABD857FFACFC1541E14013FB50" box="[899,1064,1256,1280]" metricMagnitude="-4" metricUnit="m" metricValue="8.95" metricValueMax="9.1" metricValueMin="8.8" pageId="71" pageNumber="72" unit="mm" value="0.895" valueMax="0.91" valueMin="0.88">0.88–0.91 mm</quantity>
|
||
, EW
|
||
<quantity id="82181BABD857FFACFBF041E14137FB50" box="[1126,1292,1256,1280]" metricMagnitude="-3" metricUnit="m" metricValue="1.3599999999999999" metricValueMax="1.38" metricValueMin="1.34" pageId="71" pageNumber="72" unit="mm" value="1.3599999999999999" valueMax="1.38" valueMin="1.34">1.34–1.38 mm</quantity>
|
||
; discal striae reaching apical 4/5 of elytral length. Protrochanters simple, profemora (
|
||
<figureCitation id="DDDBAACBD857FFACFC4840054009FA8A" box="[990,1074,1292,1317]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
E) each with two small ventral denticles at base and basal 1/3, protibiae (
|
||
<figureCitation id="DDDBAACBD857FFACFDF140264682FAE7" box="[615,697,1327,1352]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
F), mesotrochanters, mesofemora (
|
||
<figureCitation id="DDDBAACBD857FFACFBAA402640B5FAE7" box="[1084,1166,1327,1352]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
G), and mesotibiae (
|
||
<figureCitation id="DDDBAACBD857FFACFAF94026448FFAC2" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Fig. 45</figureCitation>
|
||
H) simple; tarsomeres II extending to midlength of tarsomeres III. Abdomen large, AL
|
||
<quantity id="82181BABD857FFACFB2E405D415BFAC3" box="[1208,1376,1364,1389]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" metricValueMax="1.08" metricValueMin="1.06" pageId="71" pageNumber="72" unit="mm" value="1.07" valueMax="1.08" valueMin="1.06">1.06–1.08 mm</quantity>
|
||
, AW
|
||
<quantity id="82181BABD857FFACFF0140714504FA3F" box="[151,319,1400,1424]" metricMagnitude="-3" metricUnit="m" metricValue="1.315" metricValueMax="1.32" metricValueMin="1.31" pageId="71" pageNumber="72" unit="mm" value="1.315" valueMax="1.32" valueMin="1.31">1.31–1.32 mm</quantity>
|
||
, tergite IV (first visible tergite) with median carina extending to near tergal apex, lacking lateral discal carinae, tergite V with thin median carina extending to half of tergal length. Sternite IX (
|
||
<figureCitation id="DDDBAACBD857FFACFB564095412AFA1A" box="[1216,1297,1436,1461]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">
|
||
Fig.
|
||
<date id="315E908ED857FFACFB624095412AFA1A" box="[1268,1297,1436,1461]" pageId="71" pageNumber="72" value="1945-01">45</date>
|
||
</figureCitation>
|
||
I) with wellsclerotized apical portion and less sclerotized basal portion. AeL
|
||
<quantity id="82181BABD857FFACFC1040B647D4FA77" box="[902,1007,1471,1496]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="71" pageNumber="72" unit="mm" value="0.61">0.61 mm</quantity>
|
||
; aedeagus (
|
||
<figureCitation id="DDDBAACBD857FFACFBE340B640F7FA77" box="[1141,1228,1471,1496]" captionStart="FIGURE 45" captionStartId="72.[151,250,1726,1748]" captionTargetBox="[160,1433,204,1695]" captionTargetId="figure@72.[151,1435,195,1704]" captionTargetPageId="72" captionText="FIGURE 45. Male diagnostic features of Horniella smetanai sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286913/files/figure.png" pageId="71" pageNumber="72">Figs 45</figureCitation>
|
||
J–L) with apically asymmetric median lobe, apex strongly extending; endophallus composed of single slender sclerite with broadened base.
|
||
</paragraph>
|
||
<paragraph id="455FB64ED857FFACFF5143254740F91F" blockId="71.[151,1437,860,2037]" pageId="71" pageNumber="72">
|
||
Female. Similar to male in general appearance; each eye composed of about 38–42 facets; Profemora simple. BL
|
||
<quantity id="82181BABD857FFACFF5543594552F9C7" box="[195,361,1616,1640]" metricMagnitude="-3" metricUnit="m" metricValue="3.7649999999999997" metricValueMax="3.8" metricValueMin="3.73" pageId="71" pageNumber="72" unit="mm" value="3.7649999999999997" valueMax="3.8" valueMin="3.73">3.73–3.80 mm</quantity>
|
||
, HL
|
||
<quantity id="82181BABD857FFACFE3243594671F9C7" box="[420,586,1616,1640]" metricMagnitude="-4" metricUnit="m" metricValue="8.55" metricValueMax="8.7" metricValueMin="8.4" pageId="71" pageNumber="72" unit="mm" value="0.855" valueMax="0.87" valueMin="0.84">0.84–0.87 mm</quantity>
|
||
, HW
|
||
<quantity id="82181BABD857FFACFD1B4346470FF9C7" box="[653,820,1615,1640]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="71" pageNumber="72" unit="mm" value="0.665" valueMax="0.68" valueMin="0.65">0.65–0.68 mm</quantity>
|
||
, PL
|
||
<quantity id="82181BABD857FFACFCFC4346402BF9C7" box="[874,1040,1615,1640]" metricMagnitude="-4" metricUnit="m" metricValue="8.55" metricValueMax="8.6" metricValueMin="8.5" pageId="71" pageNumber="72" unit="mm" value="0.855" valueMax="0.86" valueMin="0.85">0.85–0.86 mm</quantity>
|
||
, PW
|
||
<quantity id="82181BABD857FFACFBD9435940CEF9C7" box="[1103,1269,1616,1640]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.4" metricValueMin="7.1" pageId="71" pageNumber="72" unit="mm" value="0.725" valueMax="0.74" valueMin="0.71">0.71–0.74 mm</quantity>
|
||
, EL
|
||
<quantity id="82181BABD857FFACFABA435944FEF923" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.399999999999999" metricValueMin="9.1" pageId="71" pageNumber="72" unit="mm" value="0.925" valueMax="0.94" valueMin="0.91">0.91–0.94 mm</quantity>
|
||
, EW
|
||
<quantity id="82181BABD857FFACFE9D437D458FF923" box="[267,436,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" metricValueMax="1.44" metricValueMin="1.4" pageId="71" pageNumber="72" unit="mm" value="1.42" valueMax="1.44" valueMin="1.4">1.40–1.44 mm</quantity>
|
||
, AL
|
||
<quantity id="82181BABD857FFACFE63437D46A5F923" box="[501,670,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" metricValueMax="1.17" metricValueMin="1.09" pageId="71" pageNumber="72" unit="mm" value="1.13" valueMax="1.17" valueMin="1.09">1.09–1.17 mm</quantity>
|
||
, AW
|
||
<quantity id="82181BABD857FFACFD48437D47BEF923" box="[734,901,1652,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" metricValueMax="1.44" metricValueMin="1.38" pageId="71" pageNumber="72" unit="mm" value="1.41" valueMax="1.44" valueMin="1.38">1.38–1.44 mm</quantity>
|
||
. Genital complex (
|
||
<figureCitation id="DDDBAACBD857FFACFBF3437D4080F923" box="[1125,1211,1652,1677]" captionStart="FIGURE 49" captionStartId="78.[151,250,1557,1579]" captionTargetBox="[162,1432,197,1526]" captionTargetId="figure@78.[151,1435,193,1535]" captionTargetPageId="78" captionText="FIGURE 49. Female genital complex of Horniella species, in dorsal view. A. H. intricata. B. H. kaengkrachan. C. H. khaosabap. D. H. loebli. E. H. prolixo. F. H. philippina. G. H. pilosa. H. H. smetanai. I. H. cibodas. Scales: 0.2 mm." httpUri="https://zenodo.org/record/286916/files/figure.png" pageId="71" pageNumber="72">Fig. 49</figureCitation>
|
||
H) well-sclerotized, composed of apical sclerite and a pair of triangular basal sclerites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DFAE5C5D857FFACFF5143B54160F8EF" pageId="71" pageNumber="72" type="diagnosis">
|
||
<paragraph id="455FB64ED857FFACFF5143B54160F8EF" blockId="71.[151,1437,860,2037]" pageId="71" pageNumber="72">
|
||
<emphasis id="77946A5CD857FFACFF5143B545F4F97A" bold="true" box="[199,463,1724,1749]" pageId="71" pageNumber="72">Differential diagnosis.</emphasis>
|
||
This species belongs to the
|
||
<taxonomicName id="82E0CDCDD857FFACFC8743B4474BF97B" box="[785,880,1725,1748]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="77946A5CD857FFACFC8743B4474BF97B" box="[785,880,1725,1748]" italics="true" pageId="71" pageNumber="72">H. gigas</emphasis>
|
||
</taxonomicName>
|
||
group. Males can be separated from
|
||
<taxonomicName id="82E0CDCDD857FFACFA8E43B4414CF97B" box="[1304,1399,1725,1748]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="77946A5CD857FFACFA8E43B4414CF97B" box="[1304,1399,1725,1748]" italics="true" pageId="71" pageNumber="72">H. gigas</emphasis>
|
||
</taxonomicName>
|
||
by the much smaller body size, the simple protrochanters, and the more slender form of the aedeagus, while
|
||
<taxonomicName id="82E0CDCDD857FFACFAAA43E841A0F958" box="[1340,1435,1760,1784]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="77946A5CD857FFACFAAA43E841A0F958" box="[1340,1435,1760,1784]" italics="true" pageId="71" pageNumber="72">H. gigas</emphasis>
|
||
</taxonomicName>
|
||
is larger (over 4.0 mm), the protrochanters have a distinct ventral spine, and the aedeagus is much stouter in form. The lack of long hairs on the male mesofemora and metatibiae quickly separates
|
||
<taxonomicName id="82E0CDCDD857FFACFBB442204092F890" box="[1058,1193,1832,1856]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="smetanai">
|
||
<emphasis id="77946A5CD857FFACFBB442204092F890" box="[1058,1193,1832,1856]" italics="true" pageId="71" pageNumber="72">H. smetanai</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="82E0CDCDD857FFACFB784220416FF8EF" box="[1262,1364,1832,1856]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="pilosa">
|
||
<emphasis id="77946A5CD857FFACFB7842204139F8EF" box="[1262,1282,1833,1856]" italics="true" pageId="71" pageNumber="72">H</emphasis>
|
||
.
|
||
<emphasis id="77946A5CD857FFACFA864221416FF8EF" box="[1296,1364,1832,1856]" italics="true" pageId="71" pageNumber="72">pilosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DFAE5C5D857FFACFF51424546FAF8CA" box="[199,705,1868,1893]" pageId="71" pageNumber="72" type="distribution">
|
||
<paragraph id="455FB64ED857FFACFF51424546FAF8CA" blockId="71.[151,1437,860,2037]" box="[199,705,1868,1893]" pageId="71" pageNumber="72">
|
||
<emphasis id="77946A5CD857FFACFF514245455BF8CA" bold="true" box="[199,352,1868,1893]" pageId="71" pageNumber="72">Distribution.</emphasis>
|
||
East
|
||
<collectingCountry id="3DF7F6DED857FFACFE0842454632F8CA" box="[414,521,1868,1893]" name="Malaysia" pageId="71" pageNumber="72">Malaysia</collectingCountry>
|
||
: Sabah (
|
||
<figureCitation id="DDDBAACBD857FFACFDF142454688F8CA" box="[615,691,1868,1893]" captionStart="MAP 6" captionStartId="68.[151,209,1635,1656]" captionTargetBox="[151,1435,913,1613]" captionTargetId="figure@68.[151,1435,913,1614]" captionTargetPageId="68" captionText="MAP 6. Distribution of Horniella species in Malaysia." figureDoi="http://doi.org/10.5281/zenodo.6353210" httpUri="https://zenodo.org/record/6353210/files/figure.png" pageId="71" pageNumber="72">Map 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DFAE5C5D857FFACFF5142664536F803" pageId="71" pageNumber="72" type="discussion">
|
||
<paragraph id="455FB64ED857FFACFF5142664536F803" blockId="71.[151,1437,860,2037]" pageId="71" pageNumber="72">
|
||
<emphasis id="77946A5CD857FFACFF51426645B6F827" bold="true" box="[199,397,1903,1928]" pageId="71" pageNumber="72">Collection notes.</emphasis>
|
||
Individuals were collected from leaf litter samples by sifting and use of Winkler-Moczarski extractors.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0DFAE5C5D857FFACFF5142B146DBF85A" pageId="71" pageNumber="72" type="etymology">
|
||
<paragraph id="455FB64ED857FFACFF5142B146DBF85A" blockId="71.[151,1437,860,2037]" pageId="71" pageNumber="72">
|
||
<emphasis id="77946A5CD857FFACFF5142B14575F87E" bold="true" box="[199,334,1976,2001]" pageId="71" pageNumber="72">Etymology.</emphasis>
|
||
The new species is named after Aleš Smetana (Ottawa,
|
||
<collectingCountry id="3DF7F6DED857FFACFC6F42B1406AF87E" box="[1017,1105,1976,2001]" name="Canada" pageId="71" pageNumber="72">Canada</collectingCountry>
|
||
), in acknowledgment of his collection of some of the material used in this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |