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<document id="96424D3A9B1B8B2EF077ECF8A72138C8" ID-DOI="10.11646/zootaxa.3850.1.1" ID-GBIF-Dataset="1bfc2e63-3da5-448a-8ebe-71f9d25c9c5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="286868" ID-ZooBank="BFD1F483-4255-429B-9E17-8D4A9E559C5F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460882059799" checkinUser="plazi" docAuthor="Yin, Zi-Wei &amp; Li, Li-Zhen" docDate="2014" docId="CD490758D82EFFD5FF0147AF4053F85D" docLanguage="en" docName="zt03850p083.pdf" docOrigin="Zootaxa 3850 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Horniella philippina Yin and Li, new species" docType="treatment" docVersion="5" lastPageNumber="63" masterDocId="31707F20D810FFEBFF964509443BFFAF" masterDocTitle="Revision of the Oriental genus Horniella Raffray (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="63" updateTime="1698358480273" updateUser="plazi">
<mods:mods id="722E138FEB03CA72956D30D686324DB7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="88C53DD2DF0C35F83A47E36CCBE73DD3">
<mods:title id="17D8000979D8F126243BE602B4CFE16F">Revision of the Oriental genus Horniella Raffray (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="6F4DBA11A2C4ACA408257CE22313133D" type="personal">
<mods:role id="56988DC827FA6D1A4E4DCDAAB3464480">
<mods:roleTerm id="6DCB5D3FCFAF57D5D875A8A6D1920134">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C915B8A332E72C39E418C7898ABCE08D">Yin, Zi-Wei</mods:namePart>
</mods:name>
<mods:name id="B28DD158EF21A9B17A813C33B5231BBC" type="personal">
<mods:role id="43A203F623D89825B985CA3EA91C9F0B">
<mods:roleTerm id="B4924F0D3FD5273D010BAE7483610BC2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5AE18FDD98DDEE9BAAEE772D778CB6B">Li, Li-Zhen</mods:namePart>
</mods:name>
<mods:typeOfResource id="2364FB36AD3A69FF02186B20CB3B075C">text</mods:typeOfResource>
<mods:relatedItem id="4B48E5A9E24F19DAEDF3D4B2CFE611AA" type="host">
<mods:titleInfo id="C58ED151696F20559A87FF6290688EE6">
<mods:title id="5BBF65019B262392605FB21B37C57A41">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="129864003EEFFD5DD28CF8BFE5CDF004">
<mods:date id="F24A7A01C9492E19396607C303818476">2014</mods:date>
<mods:detail id="95A4FC8FEEF9C9933FE88F452BD6CF8D" type="pubDate">
<mods:number id="3DCD8601A2B26A2872EF32E788C3C01B">2014-08-12</mods:number>
</mods:detail>
<mods:detail id="6599E27931AA0508030C73FAD7C03B50" type="volume">
<mods:number id="74ACA39D7AFB509BF2B9BBDEC41C87A9">3850</mods:number>
</mods:detail>
<mods:detail id="3FE1D3F32CC4CC417C035C90E8DF04C9" type="issue">
<mods:number id="79B8AF24895D5EFB02154DD0971BBBBF">1</mods:number>
</mods:detail>
<mods:extent id="12BEF87F7631F0B55430255F70D265B1" unit="page">
<mods:start id="CF2B858A7CEE583760EC493103E3A108">1</mods:start>
<mods:end id="0433D67CF73875FCB9E5A2A2D3A82F75">83</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="84BB22DDD40B5A79712029FEAB284186">journal article</mods:classification>
<mods:identifier id="427D14BF4566E48FEACD79F4A5060693" type="DOI">10.11646/zootaxa.3850.1.1</mods:identifier>
<mods:identifier id="0DBD893C2C025B24373D3ABF11AAF23D" type="GBIF-Dataset">1bfc2e63-3da5-448a-8ebe-71f9d25c9c5a</mods:identifier>
<mods:identifier id="2B2511304628233B6A269D1A472F6AD5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2C64995BFDE1D9F22D854D517220DE46" type="Zenodo-Dep">286868</mods:identifier>
<mods:identifier id="95A49567321BC8367FD43F06B1108D7A" type="ZooBank">BFD1F483-4255-429B-9E17-8D4A9E559C5F</mods:identifier>
</mods:mods>
<treatment id="CD490758D82EFFD5FF0147AF4053F85D" ID-DOI="http://doi.org/10.5281/zenodo.6142779" ID-GBIF-Taxon="193942098" ID-Zenodo-Dep="6142779" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D" httpUri="http://treatment.plazi.org/id/CD490758D82EFFD5FF0147AF4053F85D" lastPageNumber="63" pageId="62" pageNumber="63">
<subSubSection id="0DFAE5C5D82EFFD5FF0147AF46DCFD6F" box="[151,743,678,704]" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="455FB64ED82EFFD5FF0147AF46DCFD6F" blockId="62.[151,743,678,738]" box="[151,743,678,704]" pageId="62" pageNumber="63">
<heading id="1E170122D82EFFD5FF0147AF46DCFD6F" bold="true" box="[151,743,678,704]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="77946A5CD82EFFD5FF0147AF46DCFD6F" bold="true" box="[151,743,678,704]" pageId="62" pageNumber="63">
22.
<taxonomicName id="82E0CDCDD82EFFD5FF5547AF4671FD6F" authority="Yin and Li" authorityName="Yin and Li" box="[195,586,678,704]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="philippina" status="sp. nov.">
<emphasis id="77946A5CD82EFFD5FF5547AF4580FD6F" bold="true" box="[195,443,678,704]" italics="true" pageId="62" pageNumber="63">Horniella philippina</emphasis>
Yin and Li
</taxonomicName>
,
<taxonomicNameLabel id="6CA7D727D82EFFD5FDC147AF46DCFD6F" box="[599,743,678,704]" pageId="62" pageNumber="63" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DFAE5C5D82EFFD5FF0147C0458CFD4D" box="[151,439,713,738]" pageId="62" pageNumber="63" type="description">
<paragraph id="455FB64ED82EFFD5FF0147C0458CFD4D" blockId="62.[151,743,678,738]" box="[151,439,713,738]" pageId="62" pageNumber="63">
<figureCitation id="DDDBAACBD82EFFD5FF0147C044D7FD4D" box="[151,236,713,738]" captionStart="FIGURE 37" captionStartId="60.[151,250,1990,2012]" captionTargetBox="[175,1410,763,1958]" captionTargetId="figure@60.[170,1416,756,1970]" captionTargetPageId="60" captionText="FIGURE 37. Male habitus of Horniella species. A. H. schwendingeri sp. n. B. H. philippina sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/286905/files/figure.png" pageId="62" pageNumber="63">Figs 37</figureCitation>
B, 39, 49F;
<figureCitation id="DDDBAACBD82EFFD5FEF947C3458CFD4D" box="[367,439,713,738]" captionStart="MAP 5" captionStartId="64.[151,209,1106,1127]" captionTargetBox="[151,1436,193,1085]" captionTargetId="figure@64.[151,1436,193,1085]" captionTargetPageId="64" captionText="MAP 5. Distribution of Horniella species in the Philippines." figureDoi="http://doi.org/10.5281/zenodo.6347828" httpUri="https://zenodo.org/record/6347828/files/figure.png" pageId="62" pageNumber="63">Map 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0DFAE5C5D82EFFD5FF01461847DCFC71" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="455FB64ED82EFFD5FF01461847DCFC71" blockId="62.[151,1437,785,2034]" pageId="62" pageNumber="63">
<emphasis id="77946A5CD82EFFD5FF0146184504FC85" bold="true" box="[151,319,785,810]" pageId="62" pageNumber="63">
<typeStatus id="9A5B08ECD82EFFD5FF01461844E9FC85" box="[151,210,785,810]" pageId="62" pageNumber="63">Type</typeStatus>
material
</emphasis>
(
<specimenCount id="53E67DC7D82EFFD5FEC4461B4546FC86" box="[338,381,786,810]" pageId="62" pageNumber="63" type="generic">1 ♂</specimenCount>
, 2 ♀♀).
<materialsCitation id="F588BC13D82EFFD5FE75461840DDFCE1" ID-GBIF-Occurrence="3700117324" collectingDate="1968-01" collectionCode="MHNG" collectorName="R. A. Mores" determinerName="Yin &amp; Li" location="Mt. Makiling" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="LUZON" typeStatus="holotype">
<typeStatus id="9A5B08ECD82EFFD5FE7546184675FC85" box="[483,590,785,810]" pageId="62" pageNumber="63" type="holotype">Holotype</typeStatus>
,
<specimenCount id="53E67DC7D82EFFD5FDCC461B464AFC86" box="[602,625,786,809]" pageId="62" pageNumber="63" type="male">♂</specimenCount>
, labeled ‘
<location id="403FE095D82EFFD5FD4A461B4741FC85" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D:403FE095D82EFFD5FD4A461B4741FC85" box="[732,890,785,810]" name="Mt. Makiling" pageId="62" pageNumber="63" stateProvince="LUZON">Mt. Makiling</location>
,
<location id="403FE095D82EFFD5FC10461B47D3FC85" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D:403FE095D82EFFD5FC10461B47D3FC85" box="[902,1000,786,810]" name="Lagunas" pageId="62" pageNumber="63" stateProvince="LUZON">Lagunas</location>
,
<collectingRegion id="872478ACD82EFFD5FC62461B406DFC85" box="[1012,1110,786,810]" pageId="62" pageNumber="63">LUZON</collectingRegion>
, rainfor. litter /
<collectorName id="E815D398D82EFFD5FA9E461B41A3FC86" box="[1288,1432,786,810]" pageId="62" pageNumber="63">R. A. Mores</collectorName>
,
<date id="315E908ED82EFFD5FF01463F453EFCE1" box="[151,261,821,846]" pageId="62" pageNumber="63" value="1968-01">
<collectingDate id="211A6966D82EFFD5FF01463F453EFCE1" box="[151,261,821,846]" pageId="62" pageNumber="63" value="1968-01">Jan. 1968</collectingDate>
</date>
, berlesate / Holotype [red], ♂,
<taxonomicName id="82E0CDCDD82EFFD5FDC9463C477DFCE2" box="[607,838,821,845]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="philippina" status="sp. nov.">
<emphasis id="77946A5CD82EFFD5FDC9463C477DFCE2" box="[607,838,821,845]" italics="true" pageId="62" pageNumber="63">Horniella philippina</emphasis>
</taxonomicName>
<emphasis id="77946A5CD82EFFD5FCDB463C47B7FCE1" bold="true" box="[845,908,821,846]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="6CA7D727D82EFFD5FCDB463C47B7FCE1" box="[845,908,821,846]" pageId="62" pageNumber="63" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, det.
<determinerName id="B0835A8AD82EFFD5FC5C463C47C8FCE1" box="[970,1011,821,846]" pageId="62" pageNumber="63">Yin</determinerName>
&amp;
<determinerName id="B0835A8AD82EFFD5FB80463C400AFCE1" box="[1046,1073,821,846]" pageId="62" pageNumber="63">Li</determinerName>
, 2014,
<collectionCode id="23F12E8BD82EFFD5FB15463F40DDFCE1" box="[1155,1254,822,846]" pageId="62" pageNumber="63">MHNG’</collectionCode>
</materialsCitation>
.
<materialsCitation id="F588BC13D82EFFD5FB79463F454CFC39" ID-GBIF-Occurrence="3700117311" collectingDate="1977-05-07" collectingMethod="berlese" collectionCode="MHNG" collectorName="L. E. Watrous'" country="Philippines" location="Mt., Makiling" pageId="62" pageNumber="63" specimenCount="1" specimenCount-female="1" stateProvince="Laguns Prov." typeStatus="paratype">
<typeStatus id="9A5B08ECD82EFFD5FB79463F415AFCE1" box="[1263,1377,822,846]" pageId="62" pageNumber="63" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="53E67DC7D82EFFD5FAFD463F41AEFCE2" box="[1387,1429,822,846]" pageId="62" pageNumber="63" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="3DF7F6DED82EFFD5FF634653459FFCDD" box="[245,420,858,882]" name="Philippines" pageId="62" pageNumber="63">PHILIPPINES</collectingCountry>
:
<location id="403FE095D82EFFD5FE2646534669FCDD" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D:403FE095D82EFFD5FE2646534669FCDD" box="[432,594,857,882]" country="Philippines" name="Mt., Makiling" pageId="62" pageNumber="63" stateProvince="Laguns Prov.">Mt., Makiling</location>
,
<collectingRegion id="872478ACD82EFFD5FDC9465346C2FCDE" box="[607,761,858,882]" pageId="62" pageNumber="63">Laguns Prov.</collectingRegion>
,
<locationDeviation id="54EDAF6CD82EFFD5FC9046534750FCDD" bearing="135.0" box="[774,875,857,882]" distance="4000.0" pageId="62" pageNumber="63">
<quantity id="82181BABD82EFFD5FC904653477AFCDD" box="[774,833,857,882]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="62" pageNumber="63" unit="km" value="4.0">4 km</quantity>
SE
</locationDeviation>
<location id="403FE095D82EFFD5FCE3465347CAFCDE" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D:403FE095D82EFFD5FCE3465347CAFCDE" box="[885,1009,858,881]" country="Philippines" name="Los Banos" pageId="62" pageNumber="63" stateProvince="Laguns Prov.">Los Banos</location>
,
<date id="315E908ED82EFFD5FC6846534056FCDD" box="[1022,1133,858,882]" pageId="62" pageNumber="63" value="1977-05-07">
<collectingDate id="211A6966D82EFFD5FC6846534056FCDD" box="[1022,1133,858,882]" pageId="62" pageNumber="63" value="1977-05-07">07.v.1977</collectingDate>
</date>
/
<collectingMethod id="9CA1CE59D82EFFD5FB11465040E2FCDD" box="[1159,1241,857,882]" pageId="62" pageNumber="63">berlese</collectingMethod>
, rotten figs,
<collectorName id="E815D398D82EFFD5FAFA465344C6FC39" pageId="62" pageNumber="63">L.E. Watrous’</collectorName>
(
<collectionCode id="23F12E8BD82EFFD5FE8546774554FC39" box="[275,367,894,918]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="62" pageNumber="63">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F588BC13D82EFFD5FE1F467740A2FC15" ID-GBIF-Occurrence="3700117322" collectingDate="1995-11-26" collectionCode="MHNG" collectorName="J. Kodada" country="Philippines" elevation="500" location="Lagunas" municipality="Luzon" pageId="62" pageNumber="63" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="53E67DC7D82EFFD5FE1F46774581FC3A" box="[393,442,894,918]" pageId="62" pageNumber="63" type="female">1 ♀</specimenCount>
, labeled ‘
<collectingCountry id="3DF7F6DED82EFFD5FDA546744682FC39" box="[563,697,893,918]" name="Philippines" pageId="62" pageNumber="63">Philippines</collectingCountry>
,
<collectingMunicipality id="A53B2C34D82EFFD5FD5D4677472CFC3A" box="[715,791,894,917]" pageId="62" pageNumber="63">Luzon</collectingMunicipality>
:
<location id="403FE095D82EFFD5FCBE467747B1FC39" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D:403FE095D82EFFD5FCBE467747B1FC39" box="[808,906,894,918]" country="Philippines" municipality="Luzon" name="Lagunas" pageId="62" pageNumber="63">Lagunas</location>
,
<location id="403FE095D82EFFD5FC0B46774001FC39" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D:403FE095D82EFFD5FC0B46774001FC39" box="[925,1082,893,918]" country="Philippines" municipality="Luzon" name="Mt. Banahaw" pageId="62" pageNumber="63">Mt. Banahaw</location>
nr. school about
<quantity id="82181BABD82EFFD5FA8E46774163FC39" box="[1304,1368,893,918]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="62" pageNumber="63" unit="km" value="1.0">
<locationDeviation id="54EDAF6CD82EFFD5FA8E46774163FC39" box="[1304,1368,893,918]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="62" pageNumber="63" unit="km" value="1.0">1 km</locationDeviation>
</quantity>
from
<location id="403FE095D82EFFD5FF0146A84514FC15" LSID="urn:lsid:plazi:treatment:CD490758D82EFFD5FF0147AF4053F85D:403FE095D82EFFD5FF0146A84514FC15" box="[151,303,929,954]" country="Philippines" municipality="Luzon" name="Kinabuhayan" pageId="62" pageNumber="63">Kinabuhayan</location>
,
<quantity id="82181BABD82EFFD5FEAD46A845BEFC16" box="[315,389,929,954]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="62" pageNumber="63" unit="m" value="500.0">
<elevation id="CECD517DD82EFFD5FEAD46A845BEFC16" box="[315,389,929,954]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="62" pageNumber="63" unit="m" value="500.0">500 m</elevation>
</quantity>
, degraded rainforest,
<date id="315E908ED82EFFD5FDED46A846C8FC15" box="[635,755,929,954]" pageId="62" pageNumber="63" value="1995-11-26">
<collectingDate id="211A6966D82EFFD5FDED46A846C8FC15" box="[635,755,929,954]" pageId="62" pageNumber="63" value="1995-11-26">26.xi.1995</collectingDate>
</date>
[probably leg.
<collectorName id="E815D398D82EFFD5FC3746AB4021FC15" box="[929,1050,929,954]" pageId="62" pageNumber="63">J. Kodada</collectorName>
]’ (
<collectionCode id="23F12E8BD82EFFD5FBA346AB40AAFC15" box="[1077,1169,930,954]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="62" pageNumber="63">MHNG</collectionCode>
)
</materialsCitation>
. Each
<typeStatus id="9A5B08ECD82EFFD5FB7246AB417FFC15" box="[1252,1348,930,954]" pageId="62" pageNumber="63" type="paratype">paratype</typeStatus>
bears a yellow
<typeStatus id="9A5B08ECD82EFFD5FF7C46CF4521FC71" box="[234,282,966,990]" pageId="62" pageNumber="63">type</typeStatus>
label similar to that of the
<typeStatus id="9A5B08ECD82EFFD5FDDF46CC4690FC71" box="[585,683,965,990]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
except ‘
<typeStatus id="9A5B08ECD82EFFD5FC9046CF4756FC71" box="[774,877,966,990]" pageId="62" pageNumber="63" type="paratype">Paratype</typeStatus>
♂ (or ♀)’.
</paragraph>
</subSubSection>
<subSubSection id="0DFAE5C5D82EFFD5FF5146E0463FF9CA" pageId="62" pageNumber="63" type="description">
<paragraph id="455FB64ED82EFFD5FF5146E04620FA1E" blockId="62.[151,1437,785,2034]" pageId="62" pageNumber="63">
<emphasis id="77946A5CD82EFFD5FF5146E04562FBAD" bold="true" box="[199,345,1001,1026]" pageId="62" pageNumber="63">Description.</emphasis>
Male (
<figureCitation id="DDDBAACBD82EFFD5FE3F46E045C6FBAD" box="[425,509,1001,1026]" captionStart="FIGURE 37" captionStartId="60.[151,250,1990,2012]" captionTargetBox="[175,1410,763,1958]" captionTargetId="figure@60.[170,1416,756,1970]" captionTargetPageId="60" captionText="FIGURE 37. Male habitus of Horniella species. A. H. schwendingeri sp. n. B. H. philippina sp. n. Scales: 1.0 mm." httpUri="https://zenodo.org/record/286905/files/figure.png" pageId="62" pageNumber="63">Fig. 37</figureCitation>
B). Length
<quantity id="82181BABD82EFFD5FDEA46E346DFFBAE" box="[636,740,1002,1026]" metricMagnitude="-3" metricUnit="m" metricValue="3.09" pageId="62" pageNumber="63" unit="mm" value="3.09">3.09 mm</quantity>
. Head slightly longer than wide, HL
<quantity id="82181BABD82EFFD5FB1146E040D6FBAE" box="[1159,1261,1001,1026]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="62" pageNumber="63" unit="mm" value="0.62">0.62 mm</quantity>
, HW
<quantity id="82181BABD82EFFD5FAA646E041A3FBAE" box="[1328,1432,1001,1026]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="62" pageNumber="63" unit="mm" value="0.56">0.56 mm</quantity>
; anterolateral genal projections (
<figureCitation id="DDDBAACBD82EFFD5FD9D41044659FB89" box="[523,610,1037,1062]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
C) weakly indicated; median sulcus between antennal tubercles short; scapes (
<figureCitation id="DDDBAACBD82EFFD5FF6341384577FBE5" box="[245,332,1073,1098]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
B) lacking expansion at lateral margins; clubs (
<figureCitation id="DDDBAACBD82EFFD5FC15413847E2FBE5" box="[899,985,1073,1098]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
A) formed by apical three moderately enlarged antennomeres; venter lacking lateral spines (
<figureCitation id="DDDBAACBD82EFFD5FD60415C4772FBC1" box="[758,841,1109,1134]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
D). Maxillary palpomeres II slightly broadened from base toward apex. Each eye composed of about 45 facets. Pronotum slightly longer than wide, PL
<quantity id="82181BABD82EFFD5FB6B4170415EFB3E" box="[1277,1381,1145,1170]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="62" pageNumber="63" unit="mm" value="0.65">0.65 mm</quantity>
, PW
<quantity id="82181BABD82EFFD5FF014194453BFB1A" box="[151,256,1181,1206]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="62" pageNumber="63" unit="mm" value="0.62">0.62 mm</quantity>
. Elytra wider than long, EL
<quantity id="82181BABD82EFFD5FDDC41974689FB1A" box="[586,690,1182,1206]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="62" pageNumber="63" unit="mm" value="0.79">0.79 mm</quantity>
, EW
<quantity id="82181BABD82EFFD5FD6341974765FB1A" box="[757,862,1182,1206]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="62" pageNumber="63" unit="mm" value="1.17">1.17 mm</quantity>
; discal striae reaching apical 3/4 of elytral length. Protrochanters simple, profemora (
<figureCitation id="DDDBAACBD82EFFD5FDB041C84642FB75" box="[550,633,1217,1242]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
E) each with two small ventral spines near base, protibiae (
<figureCitation id="DDDBAACBD82EFFD5FABF41C84145FB75" box="[1321,1406,1217,1242]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
F), mesotrochanters, and mesofemora (
<figureCitation id="DDDBAACBD82EFFD5FDB041EC464DFB51" box="[550,630,1253,1278]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
G) simple, mesotibiae (
<figureCitation id="DDDBAACBD82EFFD5FCEA41EC47F6FB51" box="[892,973,1253,1278]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Fig. 39</figureCitation>
H) with indistinct apical spur; tarsomeres II normal, not extending to beneath tarsomeres III. Abdomen large, AL
<quantity id="82181BABD82EFFD5FC594003400CFA8E" box="[975,1079,1290,1314]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="62" pageNumber="63" unit="mm" value="1.03">1.03 mm</quantity>
, AW
<quantity id="82181BABD82EFFD5FBEC400340D9FA8E" box="[1146,1250,1290,1314]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="62" pageNumber="63" unit="mm" value="1.2">1.20 mm</quantity>
, tergite IV (first visible tergite) with median carina extending to near half of tergal length, lacking lateral discal carinae, tergite V lacking median carina. Sternite IX (
<figureCitation id="DDDBAACBD82EFFD5FDB34058464DFAC5" box="[549,630,1361,1386]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">
Fig.
<date id="315E908ED82EFFD5FDCF405B464DFAC5" box="[601,630,1362,1386]" pageId="62" pageNumber="63" value="1939-01">39</date>
</figureCitation>
I) elongate, well-sclerotized. AeL
<quantity id="82181BABD82EFFD5FC6340584060FAC6" box="[1013,1115,1361,1386]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="62" pageNumber="63" unit="mm" value="0.68">0.68 mm</quantity>
; aedeagus (
<figureCitation id="DDDBAACBD82EFFD5FB4A4058410BFAC5" box="[1244,1328,1361,1386]" captionStart="FIGURE 39" captionStartId="63.[151,250,1914,1936]" captionTargetBox="[210,1383,203,1877]" captionTargetId="figure@63.[202,1389,195,1884]" captionTargetPageId="63" captionText="FIGURE 39. Male diagnostic features of Horniella philippina sp. n. A. Antenna. B. Scape. C. Right half of head, in dorsal view. D. Head, in lateral view. E. Protrochanter and profemur. F. Protibia. G. Mesotrochanter and mesofemur. H. Mesotibia. I. Sternite IX. J. Aedeagus, in dorsal view. K. Same, in lateral view. L. Same, in ventral view. Scales: 0.3 mm, except B, I = 0.1 mm." httpUri="https://zenodo.org/record/286907/files/figure.png" pageId="62" pageNumber="63">Figs 39</figureCitation>
J–L) with elongate, nearly symmetric median lobe, apex rounded; endophallus composed of large, oval membranous part with many small denticles at apex.
</paragraph>
<paragraph id="455FB64ED82EFFD5FF5140B4463FF9CA" blockId="62.[151,1437,785,2034]" pageId="62" pageNumber="63">
Female. Similar to male in general appearance; each eye composed of about 40 facets; profemora each with two small ventral spines near base, metatibiae simple at apex. BL
<quantity id="82181BABD82EFFD5FC5640E84057FA56" box="[960,1132,1505,1530]" metricMagnitude="-3" metricUnit="m" metricValue="2.985" metricValueMax="3.05" metricValueMin="2.92" pageId="62" pageNumber="63" unit="mm" value="2.985" valueMax="3.05" valueMin="2.92">2.92–3.05 mm</quantity>
, HL
<quantity id="82181BABD82EFFD5FB2640E84167FA56" box="[1200,1372,1505,1530]" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.5" metricValueMin="6.2" pageId="62" pageNumber="63" unit="mm" value="0.635" valueMax="0.65" valueMin="0.62">0.62–0.65 mm</quantity>
, HW
<quantity id="82181BABD82EFFD5FF01430C4507F9B2" box="[151,316,1541,1566]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="5.8999999999999995" metricValueMin="5.6" pageId="62" pageNumber="63" unit="mm" value="0.575" valueMax="0.59" valueMin="0.56">0.56–0.59 mm</quantity>
, PL
<quantity id="82181BABD82EFFD5FEE7430C462DF9B2" box="[369,534,1541,1566]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="6.7" metricValueMin="6.2" pageId="62" pageNumber="63" unit="mm" value="0.645" valueMax="0.67" valueMin="0.62">0.62–0.67 mm</quantity>
, PW
<quantity id="82181BABD82EFFD5FDC2430C46C2F9B2" box="[596,761,1541,1566]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="62" pageNumber="63" unit="mm" value="0.6" valueMax="0.62" valueMin="0.58">0.58–0.62 mm</quantity>
, EL
<quantity id="82181BABD82EFFD5FCB9430F47EEF9B2" box="[815,981,1542,1566]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" metricValueMax="7.8" metricValueMin="7.4" pageId="62" pageNumber="63" unit="mm" value="0.76" valueMax="0.78" valueMin="0.74">0.74–0.78 mm</quantity>
, EW
<quantity id="82181BABD82EFFD5FB82430C4082F9B2" box="[1044,1209,1541,1566]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.22" metricValueMin="1.16" pageId="62" pageNumber="63" unit="mm" value="1.19" valueMax="1.22" valueMin="1.16">1.16–1.22 mm</quantity>
, AL
<quantity id="82181BABD82EFFD5FB64430C41A3F9B2" box="[1266,1432,1541,1566]" metricMagnitude="-4" metricUnit="m" metricValue="9.45" metricValueMax="9.5" metricValueMin="9.399999999999999" pageId="62" pageNumber="63" unit="mm" value="0.945" valueMax="0.95" valueMin="0.94">0.94–0.95 mm</quantity>
, AW
<quantity id="82181BABD82EFFD5FF5A43234548F9EE" box="[204,371,1578,1602]" metricMagnitude="-3" metricUnit="m" metricValue="1.2149999999999999" metricValueMax="1.23" metricValueMin="1.2" pageId="62" pageNumber="63" unit="mm" value="1.2149999999999999" valueMax="1.23" valueMin="1.2">1.20–1.23 mm</quantity>
. Genital complex (
<figureCitation id="DDDBAACBD82EFFD5FDDD432046A6F9ED" box="[587,669,1577,1602]" captionStart="FIGURE 49" captionStartId="78.[151,250,1557,1579]" captionTargetBox="[162,1432,197,1526]" captionTargetId="figure@78.[151,1435,193,1535]" captionTargetPageId="78" captionText="FIGURE 49. Female genital complex of Horniella species, in dorsal view. A. H. intricata. B. H. kaengkrachan. C. H. khaosabap. D. H. loebli. E. H. prolixo. F. H. philippina. G. H. pilosa. H. H. smetanai. I. H. cibodas. Scales: 0.2 mm." httpUri="https://zenodo.org/record/286916/files/figure.png" pageId="62" pageNumber="63">Fig. 49</figureCitation>
F) composed of transverse sclerite moderately sclerotized at middle and membranous at lateral parts.
</paragraph>
</subSubSection>
<subSubSection id="0DFAE5C5D82EFFD5FF5143784652F829" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="455FB64ED82EFFD5FF5143784652F829" blockId="62.[151,1437,785,2034]" pageId="62" pageNumber="63">
<emphasis id="77946A5CD82EFFD5FF51437845EAF925" bold="true" box="[199,465,1649,1674]" pageId="62" pageNumber="63">Differential diagnosis.</emphasis>
This species is placed as a member of the
<taxonomicName id="82E0CDCDD82EFFD5FC58437B407DF926" box="[974,1094,1649,1673]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="hirtella">
<emphasis id="77946A5CD82EFFD5FC58437B47D9F926" box="[974,994,1650,1673]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="77946A5CD82EFFD5FC624378407DF926" box="[1012,1094,1649,1673]" italics="true" pageId="62" pageNumber="63">hirtella</emphasis>
</taxonomicName>
group. Males share a similar general appearance with
<taxonomicName id="82E0CDCDD82EFFD5FE26439F4626F902" box="[432,541,1686,1709]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="awana">
<emphasis id="77946A5CD82EFFD5FE26439F45FFF902" box="[432,452,1686,1709]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="77946A5CD82EFFD5FE45439F4626F902" box="[467,541,1686,1709]" italics="true" pageId="62" pageNumber="63">awana</emphasis>
</taxonomicName>
and
<taxonomicName id="82E0CDCDD82EFFD5FDC3439F46F4F902" box="[597,719,1685,1709]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="cibodas">
<emphasis id="77946A5CD82EFFD5FDC3439F4652F902" box="[597,617,1686,1709]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="77946A5CD82EFFD5FDEE439C46F4F902" box="[632,719,1685,1709]" italics="true" pageId="62" pageNumber="63">cibodas</emphasis>
</taxonomicName>
(both described below), but can be readily separated from both by the presence of an elongate median lobe with a rounded apex of the aedeagus, and the endophallus having a large, oval, elongate membranous portion with many small denticles at the apex.
<taxonomicName id="82E0CDCDD82EFFD5FBD743D44139F95A" box="[1089,1282,1757,1781]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="awana">
<emphasis id="77946A5CD82EFFD5FBD743D44139F95A" box="[1089,1282,1757,1781]" italics="true" pageId="62" pageNumber="63">Horniella awana</emphasis>
</taxonomicName>
has a stouter form of the aedeagus, while
<taxonomicName id="82E0CDCDD82EFFD5FE7C420B465CF8B6" box="[490,615,1793,1817]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="cibodas">
<emphasis id="77946A5CD82EFFD5FE7C420B45C5F8B6" box="[490,510,1794,1817]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="77946A5CD82EFFD5FD864208465CF8B6" box="[528,615,1793,1817]" italics="true" pageId="62" pageNumber="63">cibodas</emphasis>
</taxonomicName>
has a small, sharp apical sclerite at the apex of the endophallus. The elongate median lobe and endophallus that lacks any sclerite of
<taxonomicName id="82E0CDCDD82EFFD5FCF0422F47C7F892" box="[870,1020,1829,1853]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="77946A5CD82EFFD5FCF0422F4741F892" box="[870,890,1830,1853]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="77946A5CD82EFFD5FC1F422C47C7F892" box="[905,1020,1829,1853]" italics="true" pageId="62" pageNumber="63">philippina</emphasis>
</taxonomicName>
is shared with
<taxonomicName id="82E0CDCDD82EFFD5FB31422F4122F892" box="[1191,1305,1829,1853]" class="Malacostraca" family="Melphidippidae" genus="Horniella" kingdom="Animalia" order="Amphipoda" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="prolixo">
<emphasis id="77946A5CD82EFFD5FB31422F4080F892" box="[1191,1211,1830,1853]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="77946A5CD82EFFD5FB5F422C4122F892" box="[1225,1305,1829,1853]" italics="true" pageId="62" pageNumber="63">prolixo</emphasis>
</taxonomicName>
. These two species can be easily separated by the simple metatibiae of the former species while the latter species has a strongly modified apical portion of the metatibiae.
</paragraph>
</subSubSection>
<subSubSection id="0DFAE5C5D82EFFD5FF514298468BF805" box="[199,688,1937,1962]" pageId="62" pageNumber="63" type="distribution">
<paragraph id="455FB64ED82EFFD5FF514298468BF805" blockId="62.[151,1437,785,2034]" box="[199,688,1937,1962]" pageId="62" pageNumber="63">
<emphasis id="77946A5CD82EFFD5FF514298455BF805" bold="true" box="[199,352,1937,1962]" pageId="62" pageNumber="63">Distribution.</emphasis>
<collectingCountry id="3DF7F6DED82EFFD5FEF1429845D1F805" box="[359,490,1937,1962]" name="Philippines" pageId="62" pageNumber="63">Philippines</collectingCountry>
: Laguna (
<figureCitation id="DDDBAACBD82EFFD5FDC0429B4699F805" box="[598,674,1937,1962]" captionStart="MAP 5" captionStartId="64.[151,209,1106,1127]" captionTargetBox="[151,1436,193,1085]" captionTargetId="figure@64.[151,1436,193,1085]" captionTargetPageId="64" captionText="MAP 5. Distribution of Horniella species in the Philippines." figureDoi="http://doi.org/10.5281/zenodo.6347828" httpUri="https://zenodo.org/record/6347828/files/figure.png" pageId="62" pageNumber="63">Map 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DFAE5C5D82EFFD5FF5142BC4162F861" box="[199,1369,1973,1998]" pageId="62" pageNumber="63" type="discussion">
<paragraph id="455FB64ED82EFFD5FF5142BC4162F861" blockId="62.[151,1437,785,2034]" box="[199,1369,1973,1998]" pageId="62" pageNumber="63">
<emphasis id="77946A5CD82EFFD5FF5142BC45B6F861" bold="true" box="[199,397,1973,1998]" pageId="62" pageNumber="63">Collection notes.</emphasis>
Individuals were collected from litter samples by use of Berlese funnels in rainforests.
</paragraph>
</subSubSection>
<subSubSection id="0DFAE5C5D82EFFD5FF5142D04053F85D" box="[199,1128,2009,2034]" pageId="62" pageNumber="63" type="etymology">
<paragraph id="455FB64ED82EFFD5FF5142D04053F85D" blockId="62.[151,1437,785,2034]" box="[199,1128,2009,2034]" pageId="62" pageNumber="63">
<emphasis id="77946A5CD82EFFD5FF5142D04575F85D" bold="true" box="[199,334,2009,2034]" pageId="62" pageNumber="63">Etymology.</emphasis>
The species is named after the
<collectingCountry id="3DF7F6DED82EFFD5FD3842D04717F85D" box="[686,812,2009,2034]" name="Philippines" pageId="62" pageNumber="63">Philippines</collectingCountry>
where the
<typeStatus id="9A5B08ECD82EFFD5FC3142D347ECF85D" box="[935,983,2010,2034]" pageId="62" pageNumber="63">type</typeStatus>
locality lies.
</paragraph>
</subSubSection>
</treatment>
</document>
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