treatments-xml/data/67/35/62/673562BCADC9613492A9471A9EACC2FD.xml
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<mods:title id="6E4324DF65FDF15AC8AB35518DAC71F2">Australian ants of the genus Aphaenogaster (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="655CB11F052728807B40D29129E4E530">Shattuck, S. O.</mods:namePart>
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<treatment id="673562BCADC9613492A9471A9EACC2FD" ID-DOI="http://doi.org/10.5281/zenodo.6234136" ID-GBIF-Taxon="100122513" ID-Zenodo-Dep="6234136" LSID="urn:lsid:plazi:treatment:673562BCADC9613492A9471A9EACC2FD" httpUri="http://treatment.plazi.org/id/673562BCADC9613492A9471A9EACC2FD" lastPageNumber="28" pageNumber="27">
<subSubSection id="278FAF10D32B499B00675C32127B806D" type="nomenclature">
<paragraph id="AA037A451B6C2B923188D857DE196BFA" pageNumber="27">
<taxonomicName id="8EBBBB55B398A238B4418B6B3EEEDBF6" ID-CoL="Y85" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2351" class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Aphaenogaster</taxonomicName>
Mayr
</paragraph>
</subSubSection>
<subSubSection id="0CD4BCBEF3F26B4251A81B4279F3420D" type="diagnosis">
<paragraph id="337B10B34139CF9A44FE930CABC368B9" pageNumber="27">Diagnosis. Antennae 12 segmented (including the scape) with a 4 segmented club (Fig. 9). In side view the propodeum depressed below the level of the pronotum and anterior region of the mesonotum, these two regions being connected by the steeply sloping posterior section of the mesonotum (Fig. 2). Monomorphic.</paragraph>
<paragraph id="F2A428F41ACCB6F81906D8F00C09463C" pageNumber="27">
<taxonomicName id="F850E76B329E0B67D880FED6213FA9F7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2351" class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Aphaenogaster</taxonomicName>
is most likely to be confused with
<taxonomicName id="AF68BC6EAC0151C57EA9A36DA1C1494D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
or possibly
<taxonomicName id="ED2F27BDDE3D9C34D4D6D860FF30F0E6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24893" class="Insecta" family="Formicidae" genus="Pheidologeton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidologeton Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidologeton</taxonomicName>
. Workers of
<taxonomicName id="1A44B134380A3B802A2242C1D932DF71" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2351" class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Aphaenogaster</taxonomicName>
can be separated from those of
<taxonomicName id="E43BAE2F20CCD47592326AA6D47D8ED9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
by the 4 segmented rather than 3 segmented club and the larger body size (over 3.4mm long), and from
<taxonomicName id="C4C9689FF4B0BE3198046EFC933D70EA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24893" class="Insecta" family="Formicidae" genus="Pheidologeton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidologeton Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidologeton</taxonomicName>
by the 12 segmented antennae (11 segmented in
<taxonomicName id="EB827CDC7898CE6291E4928FCBC9646E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24893" class="Insecta" family="Formicidae" genus="Pheidologeton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidologeton Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidologeton</taxonomicName>
). Additionally, both
<taxonomicName id="572E831E62DEF87C035E5740BAABE7B9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
and
<taxonomicName id="2BB159B9D5E564E2BA571C7A6D064DB1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24893" class="Insecta" family="Formicidae" genus="Pheidologeton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidologeton Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidologeton</taxonomicName>
have polymorphic workers while
<taxonomicName id="9187A501E985E506943275CCA0427A43" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2351" class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Aphaenogaster</taxonomicName>
is monomorphic.
</paragraph>
</subSubSection>
<subSubSection id="ABD359E5A043E9771808AEC73415EB4E" type="discussion">
<paragraph id="72036843E56A5EE752F631046BE01302" pageNumber="27">
The Australian species of
<taxonomicName id="7A08D9EB70E91726A380C1627C5A48B5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2351" class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Aphaenogaster</taxonomicName>
show differences which are little more than &quot;variation on a theme.&quot; This is in contrast to the nearby Papua New Guinea fauna where morphological variation is considerable(Smith 1961). This difference suggests that the Australian fauna is composed of closely related species while that of PNG consists of several more distantly related lineages.
</paragraph>
</subSubSection>
<subSubSection id="0024FB56AE0DA30E6107C97C2297D52A" type="synonymic_list">
<paragraph id="B94363130211C2638C50224A415CDF48" pageNumber="27">List of Australian species</paragraph>
<paragraph id="3CF8ECEA0F79CEABF6261502C1C93E52" pageNumber="27">
<taxonomicName id="13FFFDAAD34FC5F78909871E345A7681" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238199" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster barbara Shattuck" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="barbara">barbara</taxonomicName>
<taxonomicNameLabel id="B858A4BAB922DA9EA2F486A8EF3A6523" rank="species">sp. n.</taxonomicNameLabel>
(Queensland)
</paragraph>
<paragraph id="13FB704B02EDACEFAC37ABA6A6E86120" pageNumber="27">
<taxonomicName id="4B1A095221BF93DA586703CEBE33D4FF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25388" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster barbigula Wheeler" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="barbigula">barbigula Wheeler</taxonomicName>
(New South Wales, Queensland, South Australia, Victoria)
</paragraph>
<paragraph id="DEF7EC28DC4699843E5C7D5CCA889753" pageNumber="27">
<taxonomicName id="7652D2DF602310D7B449062C099686A9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238200" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster kimberleyensis Shattuck" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="kimberleyensis">kimberleyensis</taxonomicName>
<taxonomicNameLabel id="D869A6569B1DE9021CE6D3F8C5432DDE" rank="species">sp. n.</taxonomicNameLabel>
(northern Northern Territory, northern Western Australia)
</paragraph>
<paragraph id="E7890AF3791923F312AE8ADCE2FA75F3" pageNumber="27">
<taxonomicName id="A839DE19E3F914B81E2F36F91B47AA4B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25453" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing, matching species" genus.innerRound="2" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster longiceps (Smith)" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="longiceps">longiceps (Smith)</taxonomicName>
(ACT, New South Wales, southern Queensland, south-east South Australia, Victoria)
</paragraph>
<paragraph id="A1F53951A23B4AE5E4714E6396C85466" pageNumber="27">
<taxonomicName id="267ED027717F4FDDFA72302FED7AB40C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:147880" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster longiceps flava Emery" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="flava">flava Emery</taxonomicName>
(new synonymy)
</paragraph>
<paragraph id="4CBABDB9F9CF9B1FCAAED5BEC54FD8EC" pageNumber="27">
<taxonomicName id="AC3216FB76CB627B39C7AB438D1CA2AD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:224485" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster ruginota (Forel)" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="ruginota">ruginota Forel</taxonomicName>
</paragraph>
<paragraph id="261AD6B0339D2B3FF738C46910E12C7A" pageNumber="27">
<taxonomicName id="48F1F81636CA415BA71EE245E45A68F9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238201" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster mediterrae Shattuck" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="mediterrae">mediterrae</taxonomicName>
<taxonomicNameLabel id="1BB8D549BACF79735217FF69E0EFF3BB" rank="species">sp. n.</taxonomicNameLabel>
(western South Australia, southern Western Australia)
</paragraph>
<paragraph id="5DB31E525097E2EDC835573433EE5D23" pageNumber="27">
<taxonomicName id="87CDFDF4C8C64EE8E4021D57BA2075B8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25482" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster poultoni Crawley" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="poultoni">poultoni Crawley</taxonomicName>
(south-western Western Australia)
</paragraph>
<paragraph id="1C81412440D1D133DA995A87A5B25DF5" pageNumber="27">
<taxonomicName id="13649640F0F4A03005D7A36BAB6C5F84" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25487" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster pythia Forel" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="pythia">pythia Forel</taxonomicName>
(Queensland, PNG)
</paragraph>
<paragraph id="6D3538EB97F5F8ACD5999EB93442D4F5" pageNumber="27">
<taxonomicName id="3D929FADA1BBEAAED5158F865037A4E5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238202" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster reichelae Shattuck" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="reichelae">reichelae</taxonomicName>
<taxonomicNameLabel id="B730003865BC3439F14258D067911595" rank="species">sp. n.</taxonomicNameLabel>
(northern Northern Territory)
</paragraph>
</subSubSection>
<subSubSection id="07B3BF701A408F006D58AF38E768FDFE" type="key">
<paragraph id="101522195972160F325D6DEC66839954" pageNumber="27">
Key to species of Australian
<taxonomicName id="46A1D342554BA86A66D143CFFF8881FA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:2351" class="Insecta" family="Formicidae" genus="Aphaenogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster Mayr" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Aphaenogaster</taxonomicName>
based on workers
</paragraph>
<paragraph id="4F84156217506D9A7B642DCBDD8898F0" pageNumber="27">1. Majority of hairs on venter of head located laterally and forming a distinct psammophore, only scattered hairs on central portion (Fig. 4) ................................................................................................................... 2</paragraph>
<paragraph id="18F0B325691EE13B19C2735E37D7DEB4" pageNumber="27">- Hairs on venter of head randomly distributed and not forming a distinct psammophore (Fig. 2)..............4</paragraph>
<paragraph id="87D404F642D60A6BA27C1A573A38828F" pageNumber="27">
2 Eye relatively large (EI greater than 21, Fig. 19); scape relatively long (SI greater than 106, Fig. 20)........ ......................................................................................................................................................
<taxonomicName id="0227A032DC4FC43F26F606C89668AD63" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238201" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster mediterrae Shattuck" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="mediterrae">mediterrae</taxonomicName>
</paragraph>
<paragraph id="63CA8516BE283C200C753C2FBBEACCE3" pageNumber="27">- Eye relatively small (EI less than 21, Fig. 19); scape relatively short (SI less than 106, Fig. 20)..............3</paragraph>
<paragraph id="2AFBE0606FA40B22C53BF4CCF33C6443" pageNumber="27">
3. Petiolar node (in dorsal view) wider than long; mandibular sculpture composed of irregularly sized striations(Fig. 6) (occurring in Western Australia).................................................................................
<taxonomicName id="E959B5A49E638F4F4CBC8432D94EA75C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25482" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster poultoni Crawley" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="poultoni">poultoni</taxonomicName>
</paragraph>
<paragraph id="46870751DCE969DE53DE3A0AF85B6540" pageNumber="27">
- Petiolar node (in dorsal view) approximately square; mandibular sculpture composed of regularly sized striations (Fig. 5) (occurring in South Australia and eastward)......................................................
<taxonomicName id="2155517254FB476E4A5ABA077B0A074B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25388" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster barbigula Wheeler" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="barbigula">barbigula</taxonomicName>
</paragraph>
<paragraph id="9C896C1A0ACF5723B2247E1236EEA99D" lastPageNumber="28" pageNumber="27">
4. Posterior margin of head nearly flat in full face view, extending laterally of the occipital collar before
<pageStartToken id="24322A8B8E1315C369F077A78B519212" pageNumber="28">passing</pageStartToken>
through a distinct posterolateral corner into the lateral margin of the head (Fig. 15).................... 5
</paragraph>
<paragraph id="B8994C0D35C351C2DBC4056AD1598AA3" pageNumber="28">- Posterior margin of head broadly arched in full face view, the arch beginning at the occipital collar and with at most a weak angle separating the posterior and lateral margins of the head (often posterior and lateral margins forming a continuous surface) (Fig. 9) ................................................................................... 6</paragraph>
<paragraph id="0BBC937B43D88D453CA8B1EA5A7EE449" pageNumber="28">
5. Scape relatively short (SI less than 125, Fig. 23) (occurring in e. Queensland and ne. New South Wales) .............................................................................................................................................................
<taxonomicName id="3E651A848DC823164A5E709C3A542746" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25487" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster pythia Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="pythia">pythia</taxonomicName>
</paragraph>
<paragraph id="49D1359C86DAD38D34394E0D58ADB5EC" pageNumber="28">
- Scape relatively long (SI greater than 135, Fig. 23) (occurring in Northern Territory) ..................
<taxonomicName id="62EA7DA9C9529E709BFDC83D8908E6D3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238202" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster reichelae Shattuck" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="reichelae">reichelae</taxonomicName>
</paragraph>
<paragraph id="E41BD45D7B680EBEDBDC443FA45A5B83" pageNumber="28">
6. Shorter erect hairs on mesosomal dorsum (especially those on mesonotum) with blunt tips; dorsal surfaces of propodeum and propodeal spines connected through a gentle concavity (so that the base of each spine is at approximately the same level as the dorsal surface of the propodeum) (Fig. 10)..................
<taxonomicName id="8AAE78442D1F5D7EFE8655F9E2279896" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:25453" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing, matching species" genus.innerRound="2" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster longiceps (Smith)" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="longiceps">longiceps</taxonomicName>
</paragraph>
<paragraph id="14CB7D8E0BE6106F1646CF2AEDF2244A" pageNumber="28">- Erect hairs on mesosomal dorsum tapering to sharp points; dorsal surfaces of propodeum and propodeal spines connected through a gentle concavity followed by a gentle convexity (so that the base of each spine is raised slightly above the dorsal surface of the propodeum) (Fig. 8)........................................................7</paragraph>
<paragraph id="8CFC40818912F9B9E03CCA997603B088" pageNumber="28">
7. Head relatively narrow (Fig. 21), scape relatively long (Fig. 22) (occurring in n. Northern Territory and n. Western Australia)..................................................................................................................
<taxonomicName id="D053A6AAD02BBD91E0A4287C30771560" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238200" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster kimberleyensis Shattuck" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="kimberleyensis">kimberleyensis</taxonomicName>
</paragraph>
<paragraph id="1C1A43BBAA0D42CB6535311393AF20F7" pageNumber="28">
- Head relatively broad (Fig. 21), scape relatively short (Fig. 22) (occurring in Queensland)...........
<taxonomicName id="B01EB720A9DBB3621A925274BF1081EC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:238199" class="Insecta" family="Formicidae" genus="Aphaenogaster" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Aphaenogaster barbara Shattuck" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="barbara">barbara</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>