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<mods:title id="682CFE16BDFBA27793D6619109783E7B">Papuadocus blodiwai gen. nov., sp. nov. (Crustacea: Amphipoda: Maeridae), a new bathyal species associated with sunken wood in the Bismarck Sea (Papua New Guinea)</mods:title>
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<mods:namePart id="60BA4386E6BE569C8CFC55DE9B79FB11">Sorbe, Jean Claude</mods:namePart>
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<treatment id="1903B22DEE57FFECFF0AFEF241E10FFF" ID-DOI="http://doi.org/10.5281/zenodo.6118589" ID-GBIF-Taxon="119593147" ID-Zenodo-Dep="6118589" LSID="urn:lsid:plazi:treatment:1903B22DEE57FFECFF0AFEF241E10FFF" httpUri="http://treatment.plazi.org/id/1903B22DEE57FFECFF0AFEF241E10FFF" lastPageId="5" lastPageNumber="409" pageId="2" pageNumber="408">
<subSubSection id="D9B050B0EE57FFEBFF0AFEF244F30F40" box="[151,503,285,311]" pageId="2" pageNumber="408" type="nomenclature">
<paragraph id="9115033BEE57FFEBFF0AFEF244F30F40" blockId="2.[151,503,285,311]" box="[151,503,285,311]" pageId="2" pageNumber="408">
<heading id="CA5DB457EE57FFEBFF0AFEF244F30F40" bold="true" box="[151,503,285,311]" fontSize="11" level="1" pageId="2" pageNumber="408" reason="1">
<emphasis id="A3DEDF29EE57FFEBFF0AFEF244F30F40" bold="true" box="[151,503,285,311]" pageId="2" pageNumber="408">
Genus
<taxonomicName id="56AA78B8EE57FFEBFF73FEF244860F40" ID-CoL="7PRMK" box="[238,386,285,311]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A3DEDF29EE57FFEBFF73FEF244860F40" bold="true" box="[238,386,285,311]" italics="true" pageId="2" pageNumber="408">Papuadocus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8ED6252EE57FFEBFE14FEF244F30F40" box="[393,503,285,311]" pageId="2" pageNumber="408" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9B050B0EE57FFEBFF0AFE8B401B0CB7" pageId="2" pageNumber="408" type="diagnosis">
<paragraph id="9115033BEE57FFEBFF0AFE8B401B0CB7" blockId="2.[151,1437,356,2037]" pageId="2" pageNumber="408">
<emphasis id="A3DEDF29EE57FFEBFF0AFE8B44150F0A" bold="true" box="[151,273,356,381]" pageId="2" pageNumber="408">Diagnosis.</emphasis>
Head lacking rostrum. Eyes lacking. Antennae unequal, slender; accessory flagellum long, shorter than peduncle. Mandible robust, incisor dentate; left
<emphasis id="A3DEDF29EE57FFEBFD30FE6846510FD7" box="[685,853,391,416]" italics="true" pageId="2" pageNumber="408">lacinia mobilis</emphasis>
distally dentate and bigger than the right one; molar very well developed; palp article 3 falciform. Maxilla 1 with inner plate medially setose; left and right palps not symmetrical. Maxilla 2 inner plate with oblique setal row near mesial margin. Maxilliped inner plate long, with three or four apical teeth; palp robust, article 4 stout and inserted subdistally on article 3. Gnathopod 1 small, subchelate, with transverse palm. Gnathopod 2 larger and longer, subchelate, palmar margin multidentate, not sexually dimorphic. Pereopods 57 gradually increasing in length. Coxa 4 slightly concave posteriorly, clearly higher than coxa 5, posterodistal corner slightly produced. Urosome 12 with dorsolateral teeth. Uropods 12 rami with stout apical setae. Uropod 3 biramous and uniarticulate, rami subequal, distally dentate and setose.
<taxonomicName id="56AA78B8EE57FFEBFACFFD6B40980CEA" box="[1362,1436,644,669]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
deeply cleft, with two lateral setae near outer margin. Gills on coxae 26. Oostegites on pereonites 25.
</paragraph>
</subSubSection>
<subSubSection id="D9B050B0EE57FFEBFF5AFD2340720C92" box="[199,1398,716,741]" pageId="2" pageNumber="408" type="etymology">
<paragraph id="9115033BEE57FFEBFF5AFD2340720C92" blockId="2.[151,1437,356,2037]" box="[199,1398,716,741]" pageId="2" pageNumber="408">
<emphasis id="A3DEDF29EE57FFEBFF5AFD2344420C92" bold="true" box="[199,326,716,741]" pageId="2" pageNumber="408">Etymology</emphasis>
. Named after the geographical region where the specimens were collected,
<collectingCountry id="E9BD43ABEE57FFEBFB0BFD2240770C92" box="[1174,1395,716,741]" name="Papua New Guinea" pageId="2" pageNumber="408">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D9B050B0EE57FFECFF5AFD0041E10FFF" lastPageId="5" lastPageNumber="411" pageId="2" pageNumber="408" type="discussion">
<paragraph id="9115033BEE57FFEBFF5AFD00440C0BAF" blockId="2.[151,1437,356,2037]" pageId="2" pageNumber="408">
<emphasis id="A3DEDF29EE57FFEBFF5AFD00467C0D7F" bold="true" box="[199,888,751,776]" pageId="2" pageNumber="408">
Remarks on genus assignation and
<taxonomicName id="56AA78B8EE57FFEBFD1EFD0047F20D7F" box="[643,758,751,776]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
diagnosis.
</emphasis>
Following
<bibRefCitation id="F53B7ECAEE57FFEBFB9FFD0040380D7F" author="Barnard" box="[1026,1340,751,776]" pageId="2" pageNumber="408" refString="Barnard, J. L. &amp; Barnard, C. M. (1983) Freshwater Amphipoda of the World. I. Evolutionary patterns, II. Handbook and bibliography. Hayfield Associates, Mount Vernon, Virginia, 830 pp." type="book" year="1983">Barnard &amp; Barnard (1983)</bibRefCitation>
, Krapp-
<bibRefCitation id="F53B7ECAEE57FFEBFF0AFCFB444A0D5B" author="Schickel" box="[151,334,788,813]" pageId="2" pageNumber="408" refString="Krapp-Schickel, T. (2008) What has happened with the Maera - clade (Crustacea, Amphipoda) during the last decade? Bollettino del Museo Civico di Storia Naturale di Verona, 32, 3 - 32." type="journal article" year="2008">Schickel (2008)</bibRefCitation>
distinguished the
<emphasis id="A3DEDF29EE57FFEBFDB5FCFB467D0D5A" box="[552,889,788,813]" italics="true" pageId="2" pageNumber="408">Ceradocus-Maera-Elasmopus</emphasis>
group (the 'ceradocid' complex containing the three oldest genera of this group), leading to the creation of the family
<taxonomicName id="56AA78B8EE57FFEBFC2EFCD841190D27" box="[947,1053,823,848]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
on the basis of uropod 3 structure (subequal rami versus unequal rami in
<taxonomicName id="56AA78B8EE57FFEBFDFAFCB347DC0D02" box="[615,728,860,885]" class="Malacostraca" family="Melitidae" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Melitidae</taxonomicName>
). In Table 1, the BS specimens (
<taxonomicName id="56AA78B8EE57FFEBFBFCFCB341E80D02" box="[1121,1260,860,885]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A3DEDF29EE57FFEBFBFCFCB341E80D02" box="[1121,1260,860,885]" italics="true" pageId="2" pageNumber="408">Papuadocus</emphasis>
</taxonomicName>
<emphasis id="A3DEDF29EE57FFEBFB6AFCB3406A0D02" bold="true" box="[1271,1390,860,885]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="B8ED6252EE57FFEBFB6AFCB340620D02" box="[1271,1382,860,885]" pageId="2" pageNumber="408" rank="genus">gen. nov.</taxonomicNameLabel>
)
</emphasis>
are compared with these three first described genera, using genus diagnoses found in the literature (references in Table 1) as well as selected diagnostic characters given by
<bibRefCitation id="F53B7ECAEE57FFEBFD73FC4A46FD0DCB" author="Lowry" box="[750,1017,932,957]" pageId="2" pageNumber="408" refString="Lowry, J. K. &amp; Myers, A. A. (2013) A phylogeny and classification of the Senticaudata subord. nov. (Crustacea: Amphipoda). Zootaxa, 3610 (1), 1 - 80. http: // dx. doi. org / 10.11646 / zootaxa. 3610.1.1" type="journal article" year="2013">Lowry &amp; Myers (2013)</bibRefCitation>
for
<taxonomicName id="56AA78B8EE57FFEBFBB7FC4B41930DCA" box="[1066,1175,932,957]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
. The new genus shares most of these maerid diagnostic characters with the other genera (leading to its assignment to
<taxonomicName id="56AA78B8EE57FFEBFB36FC28401E0D97" box="[1195,1306,967,992]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
), except for maxillas 1 with asymmetrical right and left palps and gnathopods 2 without sexual dimorphism.
<collectingCountry id="E9BD43ABEE57FFEBFB51FC0341EF0A73" box="[1228,1259,1004,1028]" name="American Samoa" pageId="2" pageNumber="408">As</collectingCountry>
shown in Table 1, the asymmetry of maxilla 1 palps is a distinguishing character shared with the genus
<emphasis id="A3DEDF29EE57FFEBFB35FBE040600A5F" box="[1192,1380,1039,1064]" italics="true" pageId="2" pageNumber="408">
<taxonomicName id="56AA78B8EE57FFEBFB35FBE0405B0A5F" box="[1192,1375,1039,1064]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">Bathyceradocus</taxonomicName>
,
</emphasis>
(see original genus diagnosis in
<bibRefCitation id="F53B7ECAEE57FFEBFE55FBDB47410A3B" author="Pirlot" box="[456,581,1076,1101]" pageId="2" pageNumber="408" refString="Pirlot, J. (1934) Les amphipodes de l'expedition du Siboga. Deuxieme partie. Les amphipodes gammarides. II. Les amphipodes de la mer profonde. 2. Siboga-Expeditie, 33 d, 167 - 235." type="journal article" year="1934">Pirlot 1934</bibRefCitation>
for
<taxonomicName id="56AA78B8EE57FFEBFDEFFBDA46100A3A" box="[626,788,1076,1101]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="species" species="stephenseni">
<emphasis id="A3DEDF29EE57FFEBFDEFFBDA46100A3A" box="[626,788,1076,1101]" italics="true" pageId="2" pageNumber="408">B. stephenseni</emphasis>
</taxonomicName>
; genus assigned to
<taxonomicName id="56AA78B8EE57FFEBFC70FBDB41530A3A" box="[1005,1111,1076,1101]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
by
<bibRefCitation id="F53B7ECAEE57FFEBFBE2FBDB40690A3B" author="Krapp-Schickel" box="[1151,1389,1076,1101]" pageId="2" pageNumber="408" refString="Krapp-Schickel, T. (2008) What has happened with the Maera - clade (Crustacea, Amphipoda) during the last decade? Bollettino del Museo Civico di Storia Naturale di Verona, 32, 3 - 32." type="journal article" year="2008">Krapp-Schickel 2008</bibRefCitation>
and
<bibRefCitation id="F53B7ECAEE57FFEBFF0AFBB6448B0A07" author="Lowry" box="[151,399,1112,1137]" pageId="2" pageNumber="408" refString="Lowry, J. K. &amp; Myers, A. A. (2013) A phylogeny and classification of the Senticaudata subord. nov. (Crustacea: Amphipoda). Zootaxa, 3610 (1), 1 - 80. http: // dx. doi. org / 10.11646 / zootaxa. 3610.1.1" type="journal article" year="2013">Lowry &amp; Myers 2013</bibRefCitation>
), a morphological feature confirmed in subsequent descriptions of new species (
<taxonomicName id="56AA78B8EE57FFEBFA8DFBB640980A07" box="[1296,1436,1111,1136]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="species" species="iberiensis">
<emphasis id="A3DEDF29EE57FFEBFA8DFBB640980A07" box="[1296,1436,1111,1136]" italics="true" pageId="2" pageNumber="408">B. iberiensis</emphasis>
</taxonomicName>
in
<bibRefCitation id="F53B7ECAEE57FFEBFF29FB93444C0AE3" author="Andres" box="[180,328,1148,1173]" pageId="2" pageNumber="408" refString="Andres, H. G. (1977) Gammaridea (Crustacea, Amphipoda) aus dem Iberischen Tiefseebecken. Auswertung des Materials der Fahrten 3 und 15 von F. S. Meteor. Meteor Forschungs-Ergebnisse, Reihe D, 25, 54 - 67." type="journal article" year="1977">Andres 1977</bibRefCitation>
;
<taxonomicName id="56AA78B8EE57FFEBFEC9FB9244C10AE3" box="[340,453,1149,1172]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="species" species="wuzzae">
<emphasis id="A3DEDF29EE57FFEBFEC9FB9244C10AE3" box="[340,453,1149,1172]" italics="true" pageId="2" pageNumber="408">B. wuzzae</emphasis>
</taxonomicName>
in
<bibRefCitation id="F53B7ECAEE57FFEBFE76FB9246550AE3" author="Larsen" box="[491,849,1148,1173]" pageId="2" pageNumber="408" refString="Larsen, K. &amp; Krapp-Schickel, T. (2007) Amphipoda (Crustacea: Peracarida) from chemically reduced habitats; the hydrothermal vent system of the north-east Pacific. Part II. Melitidae and Eusiridae. Journal of the Marine Biological Association of the United Kingdom, 87, 1207 - 1217. http: // dx. doi. org / 10.1017 / S 002531540705672 X" type="journal article" year="2007">Larsen &amp; Krapp-Schickel 2007</bibRefCitation>
). However, the BS specimens cannot be assigned to genus
<taxonomicName id="56AA78B8EE57FFEBFF7DFB7044900ACF" box="[224,404,1183,1208]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFF7DFB7044900ACF" box="[224,404,1183,1208]" italics="true" pageId="2" pageNumber="408">Bathyceradocus</emphasis>
</taxonomicName>
due to the absence of gill on coxae 7 (present in
<taxonomicName id="56AA78B8EE57FFEBFC5FFB7041720ACF" box="[962,1142,1183,1208]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFC5FFB7041720ACF" box="[962,1142,1183,1208]" italics="true" pageId="2" pageNumber="408">Bathyceradocus</emphasis>
</taxonomicName>
), a morphological feature shared with all other maerid genera (see
<bibRefCitation id="F53B7ECAEE57FFEBFD0CFB2A46A50AAB" author="Lowry" box="[657,929,1220,1245]" pageId="2" pageNumber="408" refString="Lowry, J. K. &amp; Myers, A. A. (2013) A phylogeny and classification of the Senticaudata subord. nov. (Crustacea: Amphipoda). Zootaxa, 3610 (1), 1 - 80. http: // dx. doi. org / 10.11646 / zootaxa. 3610.1.1" type="journal article" year="2013">Lowry &amp; Myers 2013</bibRefCitation>
).
<bibRefCitation id="F53B7ECAEE57FFEBFC26FB2A41DD0AAB" author="Lowry" box="[955,1241,1220,1245]" pageId="2" pageNumber="408" refString="Lowry, J. K. &amp; Myers, A. A. (2013) A phylogeny and classification of the Senticaudata subord. nov. (Crustacea: Amphipoda). Zootaxa, 3610 (1), 1 - 80. http: // dx. doi. org / 10.11646 / zootaxa. 3610.1.1" type="journal article" year="2013">Lowry &amp; Myers (2013)</bibRefCitation>
overlooked this distinguishing feature (gill on coxae 7) in their diagnosis of
<taxonomicName id="56AA78B8EE57FFEBFC1FFB0846EB0B77" box="[898,1007,1255,1280]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
, although mentioned for all known
<taxonomicName id="56AA78B8EE57FFEBFF0AFAE344480B52" box="[151,332,1292,1317]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFF0AFAE344480B52" box="[151,332,1292,1317]" italics="true" pageId="2" pageNumber="408">Bathyceradocus</emphasis>
</taxonomicName>
species (
<taxonomicName id="56AA78B8EE57FFEBFE2FFAE247570B52" box="[434,595,1292,1317]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="species" species="stephenseni">
<emphasis id="A3DEDF29EE57FFEBFE2FFAE247570B52" box="[434,595,1292,1317]" italics="true" pageId="2" pageNumber="408">B. stephenseni</emphasis>
</taxonomicName>
in
<bibRefCitation id="F53B7ECAEE57FFEBFDEBFAE347F20B53" author="Pirlot" box="[630,758,1292,1317]" pageId="2" pageNumber="408" refString="Pirlot, J. (1934) Les amphipodes de l'expedition du Siboga. Deuxieme partie. Les amphipodes gammarides. II. Les amphipodes de la mer profonde. 2. Siboga-Expeditie, 33 d, 167 - 235." type="journal article" year="1934">Pirlot 1934</bibRefCitation>
;
<taxonomicName id="56AA78B8EE57FFEBFC9CFAE246880B52" box="[769,908,1292,1317]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="species" species="iberiensis">
<emphasis id="A3DEDF29EE57FFEBFC9CFAE246880B52" box="[769,908,1292,1317]" italics="true" pageId="2" pageNumber="408">B. iberiensis</emphasis>
</taxonomicName>
in
<bibRefCitation id="F53B7ECAEE57FFEBFC33FAE341450B53" author="Andres" box="[942,1089,1292,1317]" pageId="2" pageNumber="408" refString="Andres, H. G. (1977) Gammaridea (Crustacea, Amphipoda) aus dem Iberischen Tiefseebecken. Auswertung des Materials der Fahrten 3 und 15 von F. S. Meteor. Meteor Forschungs-Ergebnisse, Reihe D, 25, 54 - 67." type="journal article" year="1977">Andres 1977</bibRefCitation>
;
<taxonomicName id="56AA78B8EE57FFEBFBD1FAE241B80B53" box="[1100,1212,1293,1316]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="species" species="wuzzae">
<emphasis id="A3DEDF29EE57FFEBFBD1FAE241B80B53" box="[1100,1212,1293,1316]" italics="true" pageId="2" pageNumber="408">B. wuzzae</emphasis>
</taxonomicName>
in Larsen &amp; Krapp- Schickel 2007). In accordance with the preceding genus comparisons, we conclude that the BS specimens belong to
<taxonomicName id="56AA78B8EE57FFEBFF28FABB441B0B1A" box="[181,287,1364,1389]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
and show a close morphological as well as genetic and ecological affinity to genus
<taxonomicName id="56AA78B8EE57FFEBFB7AFABB40980B1A" box="[1255,1436,1364,1389]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFB7AFABB40980B1A" box="[1255,1436,1364,1389]" italics="true" pageId="2" pageNumber="408">Bathyceradocus</emphasis>
</taxonomicName>
(see below). Accordingly, we propose to create the new genus
<taxonomicName id="56AA78B8EE57FFEBFCC9FA9846DB0BE7" box="[852,991,1399,1424]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFCC9FA9846DB0BE7" box="[852,991,1399,1424]" italics="true" pageId="2" pageNumber="408">Papuadocus</emphasis>
</taxonomicName>
for the BS specimens and to amend the Lowry &amp; Myerss diagnosis in order to maintain both genera
<taxonomicName id="56AA78B8EE57FFEBFCE3FA7341360BC2" box="[894,1074,1436,1461]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFCE3FA7341360BC2" box="[894,1074,1436,1461]" italics="true" pageId="2" pageNumber="408">Bathyceradocus</emphasis>
</taxonomicName>
and
<taxonomicName id="56AA78B8EE57FFEBFBE9FA7341FB0BC2" box="[1140,1279,1436,1461]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A3DEDF29EE57FFEBFBE9FA7341FB0BC2" box="[1140,1279,1436,1461]" italics="true" pageId="2" pageNumber="408">Papuadocus</emphasis>
</taxonomicName>
<emphasis id="A3DEDF29EE57FFEBFA96FA73407E0BC2" bold="true" box="[1291,1402,1436,1461]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="B8ED6252EE57FFEBFA96FA73407E0BC2" box="[1291,1402,1436,1461]" pageId="2" pageNumber="408" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
in
<taxonomicName id="56AA78B8EE57FFEBFF0AFA5044000BAF" box="[151,260,1471,1496]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
.
</paragraph>
<paragraph id="9115033BEE57FFEAFF5AFA0B43950CDD" blockId="2.[151,1437,356,2037]" lastBlockId="3.[119,1891,576,682]" lastPageId="3" lastPageNumber="409" pageId="2" pageNumber="408">
<emphasis id="A3DEDF29EE57FFEBFF5AFA0B410B0B8A" bold="true" box="[199,1039,1508,1533]" pageId="2" pageNumber="408">
Amended diagnosis of
<taxonomicName id="56AA78B8EE57FFEBFE60FA0B47740B8A" box="[509,624,1508,1533]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
(after
<bibRefCitation id="F53B7ECAEE57FFEBFD40FA0B41030B8A" author="Lowry" box="[733,1031,1508,1533]" pageId="2" pageNumber="408" refString="Lowry, J. K. &amp; Myers, A. A. (2013) A phylogeny and classification of the Senticaudata subord. nov. (Crustacea: Amphipoda). Zootaxa, 3610 (1), 1 - 80. http: // dx. doi. org / 10.11646 / zootaxa. 3610.1.1" type="journal article" year="2013">Lowry &amp; Myers 2013</bibRefCitation>
)
</emphasis>
. Body laterally compressed or subcylindrical. Eyes well developed or absent, if present then round, ovoid, reniform, lageniform or subrectangular. Antennae 12 calceoli absent. Antenna 1 subequal in length to, or longer than antenna 2; peduncular article 1 shorter than, subequal to, or longer than article 2; article 2 longer than article 3; article 3 shorter than article 1; peduncular articles 12 not geniculate; accessory flagellum long, short or minute. Antenna 2 peduncular article 1 not enlarged. Mandible molar triturative; palp symmetrical or absent. Maxilla 1 basal endite setose along medial margin or apically setose; palps symmetrical (most maerid genera) or
<emphasis id="A3DEDF29EE57FFEBFC2CF953415708A2" bold="true" box="[945,1107,1724,1749]" pageId="2" pageNumber="408">asymmetrical</emphasis>
(genera
<taxonomicName id="56AA78B8EE57FFEBFB2AF953406F08A2" box="[1207,1387,1724,1749]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFB2AF953406F08A2" box="[1207,1387,1724,1749]" italics="true" pageId="2" pageNumber="408">Bathyceradocus</emphasis>
</taxonomicName>
and
<taxonomicName id="56AA78B8EE57FFEBFF0AF9304426088F" box="[151,290,1759,1784]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A3DEDF29EE57FFEBFF0AF9304426088F" box="[151,290,1759,1784]" italics="true" pageId="2" pageNumber="408">Papuadocus</emphasis>
</taxonomicName>
<emphasis id="A3DEDF29EE57FFEBFEB1F930449D088F" bold="true" box="[300,409,1759,1784]" pageId="2" pageNumber="408">
<taxonomicNameLabel id="B8ED6252EE57FFEBFEB1F930449D088F" box="[300,409,1759,1784]" pageId="2" pageNumber="408" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
). Maxilla 2 basal endite with or without oblique setal row. Labium inner lobes present, vestigial or absent.
<emphasis id="A3DEDF29EE57FFEBFEEFF8EB47710937" bold="true" pageId="2" pageNumber="408">
Coxal gills on pereopods 26 (most maerid genera, including
<taxonomicName id="56AA78B8EE57FFEBFBD0F8EC41DF096B" box="[1101,1243,1795,1820]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A3DEDF29EE57FFEBFBD0F8EC41DF096B" bold="true" box="[1101,1243,1795,1820]" italics="true" pageId="2" pageNumber="408">Papuadocus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8ED6252EE57FFEBFB7FF8EB4049096A" box="[1250,1357,1796,1821]" pageId="2" pageNumber="408" rank="genus">gen. nov.</taxonomicNameLabel>
) or on pereopods 27 (genus
<taxonomicName id="56AA78B8EE57FFEBFE2EF8C847680937" box="[435,620,1831,1856]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE57FFEBFE2EF8C847680937" bold="true" box="[435,620,1831,1856]" italics="true" pageId="2" pageNumber="408">Bathyceradocus</emphasis>
</taxonomicName>
)
</emphasis>
, not stalked; sternal gills absent; sternal blisters absent; oostegites fringing setae simple. Gnathopod 1 subchelate, not sexually dimorphic; smaller (or weaker) than or similar in size to gnathopod 2; propodus palm without robust setae along palmar margin. Gnathopod 2 subchelate;
<emphasis id="A3DEDF29EE57FFEBFAA1F880468009DA" bold="true" pageId="2" pageNumber="408">
sexually dimorphic (most maerid genera) or not (
<taxonomicName id="56AA78B8EE57FFEBFDE1F87C460D09DB" box="[636,777,1939,1964]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A3DEDF29EE57FFEBFDE1F87C460D09DB" bold="true" box="[636,777,1939,1964]" italics="true" pageId="2" pageNumber="408">Papuadocus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8ED6252EE57FFEBFC8CF87B467809DA" box="[785,892,1940,1965]" pageId="2" pageNumber="408" rank="genus">gen. nov.</taxonomicNameLabel>
)
</emphasis>
; carpus not produced along posterior margin of propodus, projecting between merus and propodus. Pereopods 34 not sexually dimorphic. Pereopod 4 with welldeveloped posteroventral lobe or with small posteroventral lobe or without posteroventral lobe. Pereopod 5 shorter
<emphasis id="A3DEDF29EE56FFEAFFEAFDA945E00C2B" bold="true" box="[119,228,582,604]" pageId="3" pageNumber="409">TABLE 1.</emphasis>
Morphological comparison of maerid genera belonging to the
<emphasis id="A3DEDF29EE56FFEAFCFDFDA941950C2C" box="[864,1169,582,604]" italics="true" pageId="3" pageNumber="409">
<taxonomicName id="56AA78B8EE56FFEAFCFDFDA946CA0C2C" box="[864,974,582,603]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus">Ceradocus</taxonomicName>
-
<taxonomicName id="56AA78B8EE56FFEAFC4BFDA8411D0C2B" box="[982,1049,583,604]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus">Maera</taxonomicName>
-
<taxonomicName id="56AA78B8EE56FFEAFBBFFDA941950C2C" box="[1058,1169,582,603]" class="Malacostraca" family="Melitidae" genus="Elasmopus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus">Elasmopus</taxonomicName>
</emphasis>
group (see
<bibRefCitation id="F53B7ECAEE56FFEAFA95FDA940E70C2B" author="Krapp-Schickel" box="[1288,1507,582,604]" pageId="3" pageNumber="409" refString="Krapp-Schickel, T. (2008) What has happened with the Maera - clade (Crustacea, Amphipoda) during the last decade? Bollettino del Museo Civico di Storia Naturale di Verona, 32, 3 - 32." type="journal article" year="2008">Krapp-Schickel 2008</bibRefCitation>
), according to published diagnoses (1:
<bibRefCitation id="F53B7ECAEE56FFEAFFEAFD8144110CF4" author="Karaman" box="[119,277,621,643]" pageId="3" pageNumber="409" refString="Karaman, G. S. (1982) Family Gammaridae (sensu lato). In: Ruffo, S. (Ed.), The Amphipoda of the Mediterranean. Part 1: Gammaridea (Acanthonotozomatidae to Gammaridae). Memoires de l'Institut Oceanographique, 13, pp. 1 - 364." type="journal article" year="1982">Karaman 1982</bibRefCitation>
; 2:
<bibRefCitation id="F53B7ECAEE56FFEAFEA3FD8247860CF4" author="Vader" box="[318,642,621,643]" pageId="3" pageNumber="409" refString="Vader, W. &amp; Krapp-Schickel, T. (2012) On some maerid and melitid material (Crustacea: Amphipoda) collected by the Hourglass Cruises (Florida). Part 2: Genera Dulichiella and Elasmopus, with a key to world Elasmopus. Journal of Natural History, 46, 1179 - 1218. http: // dx. doi. org / 10.1080 / 00222933.2011.652984" type="journal article" year="2012">Vader &amp; Krapp-Schickel 2012</bibRefCitation>
; 3:
<bibRefCitation id="F53B7ECAEE56FFEAFD36FD8246BC0CF4" author="Barnard" box="[683,952,621,643]" pageId="3" pageNumber="409" refString="Barnard, J. L. &amp; Barnard, C. M. (1983) Freshwater Amphipoda of the World. I. Evolutionary patterns, II. Handbook and bibliography. Hayfield Associates, Mount Vernon, Virginia, 830 pp." type="book" year="1983">Barnard &amp; Barnard 1983</bibRefCitation>
; 4:
<bibRefCitation id="F53B7ECAEE56FFEAFC7CFD8241C50CF4" author="Krapp-Schickel" box="[993,1217,621,643]" pageId="3" pageNumber="409" refString="Krapp-Schickel, T. (2008) What has happened with the Maera - clade (Crustacea, Amphipoda) during the last decade? Bollettino del Museo Civico di Storia Naturale di Verona, 32, 3 - 32." type="journal article" year="2008">Krapp-Schickel 2008</bibRefCitation>
; 5:
<bibRefCitation id="F53B7ECAEE56FFEAFB77FD82432A0CF4" author="Krapp-Schickel" box="[1258,1582,621,643]" pageId="3" pageNumber="409" refString="Krapp-Schickel, T. &amp; Vader, W. (2009) On some maerid genera (Crustacea, Amphipoda, Maeridae) collected by the Hourglass Cruises (Florida). Part I: Genera Anamaera, Ceradocus, Clessidra gen. nov., Jerbarnia, Maera, Meximaera, with a key to world Ceradocus. Journal of Natural History, 43, 2057 - 2086. http: // dx. doi. org / 10.1080 / 00222930903091007" type="journal article" year="2009">Krapp-Schickel &amp; Vader 2009</bibRefCitation>
; 6:
<bibRefCitation id="F53B7ECAEE56FFEAF9CAFD8243D40CF4" author="Pirlot" box="[1623,1744,621,643]" pageId="3" pageNumber="409" refString="Pirlot, J. (1934) Les amphipodes de l'expedition du Siboga. Deuxieme partie. Les amphipodes gammarides. II. Les amphipodes de la mer profonde. 2. Siboga-Expeditie, 33 d, 167 - 235." type="journal article" year="1934">Pirlot 1934</bibRefCitation>
; 7:
<bibRefCitation id="F53B7ECAEE56FFEAF964FD8144560CDD" author="Larsen" pageId="3" pageNumber="409" refString="Larsen, K. &amp; Krapp-Schickel, T. (2007) Amphipoda (Crustacea: Peracarida) from chemically reduced habitats; the hydrothermal vent system of the north-east Pacific. Part II. Melitidae and Eusiridae. Journal of the Marine Biological Association of the United Kingdom, 87, 1207 - 1217. http: // dx. doi. org / 10.1017 / S 002531540705672 X" type="journal article" year="2007">Larsen &amp; Krapp-Schickel 2007</bibRefCitation>
; 8:
<bibRefCitation id="F53B7ECAEE56FFEAFEEFFD7B44DB0CDD" author="Dahl" box="[370,479,660,682]" pageId="3" pageNumber="409" refString="Dahl, E. (1959) Amphipoda from depths exceeding 6000 meters. Galathea Report, 1, 211 - 240." type="journal article" year="1959">Dahl 1959</bibRefCitation>
). The underlined characters are those which differ from the diagnosis given by
<bibRefCitation id="F53B7ECAEE56FFEAFA9CFD7A43040CDD" author="Lowry" box="[1281,1536,660,682]" pageId="3" pageNumber="409" refString="Lowry, J. K. &amp; Myers, A. A. (2013) A phylogeny and classification of the Senticaudata subord. nov. (Crustacea: Amphipoda). Zootaxa, 3610 (1), 1 - 80. http: // dx. doi. org / 10.11646 / zootaxa. 3610.1.1" type="journal article" year="2013">Lowry and Myers (2013)</bibRefCitation>
for
<taxonomicName id="56AA78B8EE56FFEAF9B7FD7B43890CDD" box="[1578,1677,660,682]" class="Malacostraca" family="Maeridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="family">Maeridae</taxonomicName>
.
</paragraph>
<paragraph id="9115033BEE56FFEAFE30FD1243AF0864" pageId="3" pageNumber="409">
<table id="E3AAF19BEE560016FFEAFD1842580863" box="[119,1884,759,1556]" gridcols="6" gridrows="18" pageId="3" pageNumber="409">
<tr id="2F9A0179EE560016FFEAFD1842580D64" box="[119,1884,759,787]" gridrow="0" pageId="3" pageNumber="409" rowspan-0="1">
<th id="6C4B6805EE560016FE30FD1847750D64" box="[429,625,759,787]" gridcol="1" gridrow="0" pageId="3" pageNumber="409">
<taxonomicName id="56AA78B8EE56FFEAFE30FD1247240D64" box="[429,544,765,787]" class="Malacostraca" family="Melitidae" genus="Elasmopus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE56FFEAFE30FD1247240D64" bold="true" box="[429,544,765,787]" italics="true" pageId="3" pageNumber="409">Elasmopus</emphasis>
</taxonomicName>
<emphasis id="A3DEDF29EE56FFEAFDBDFD18473D0D71" bold="true" box="[544,569,759,774]" italics="true" pageId="3" pageNumber="409">1, 2</emphasis>
</th>
<th id="6C4B6805EE560016FD78FD1846CF0D64" box="[741,971,759,787]" gridcol="2" gridrow="0" pageId="3" pageNumber="409">
<taxonomicName id="56AA78B8EE56FFEAFD7BFD11462F0D64" box="[742,811,766,787]" class="Malacostraca" family="Melitidae" genus="Maera" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE56FFEAFD7BFD11462F0D64" bold="true" box="[742,811,766,787]" italics="true" pageId="3" pageNumber="409">Maera</emphasis>
</taxonomicName>
<emphasis id="A3DEDF29EE56FFEAFCB6FD1846400D71" bold="true" box="[811,836,759,774]" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFCB6FD18463C0D71" bold="true" box="[811,824,759,774]" italics="true" pageId="3" pageNumber="409">3,</emphasis>
4
</emphasis>
</th>
<th id="6C4B6805EE560016FB97FD18401C0D64" box="[1034,1304,759,787]" gridcol="3" gridrow="0" pageId="3" pageNumber="409">
<taxonomicName id="56AA78B8EE56FFEAFB91FD12417F0D64" box="[1036,1147,765,787]" class="Malacostraca" family="Melitidae" genus="Ceradocus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE56FFEAFB91FD12417F0D64" bold="true" box="[1036,1147,765,787]" italics="true" pageId="3" pageNumber="409">Ceradocus</emphasis>
</taxonomicName>
<emphasis id="A3DEDF29EE56FFEAFBE6FD1741970D71" bold="true" box="[1147,1171,760,774]" pageId="3" pageNumber="409">4, 5</emphasis>
</th>
<th id="6C4B6805EE560016FAD6FD1843190D64" box="[1355,1565,759,787]" gridcol="4" gridrow="0" pageId="3" pageNumber="409">
<taxonomicName id="56AA78B8EE56FFEAFAD0FD1240F00D64" box="[1357,1524,765,787]" class="Malacostraca" family="Maeridae" genus="Bathyceradocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus">
<emphasis id="A3DEDF29EE56FFEAFAD0FD1240F00D64" bold="true" box="[1357,1524,765,787]" italics="true" pageId="3" pageNumber="409">Bathyceradocus</emphasis>
</taxonomicName>
<emphasis id="A3DEDF29EE56FFEAFA69FD1743190D71" bold="true" box="[1524,1565,760,774]" pageId="3" pageNumber="409">6, 7, 8</emphasis>
</th>
<th id="6C4B6805EE560016F9D5FD1842580D64" box="[1608,1884,759,787]" gridcol="5" gridrow="0" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAF9D7FD1242350D64" bold="true" box="[1610,1841,765,787]" italics="true" pageId="3" pageNumber="409">
<taxonomicName id="56AA78B8EE56FFEAF9D7FD1243CE0D64" box="[1610,1738,765,787]" class="Malacostraca" family="Maeridae" genus="Papuadocus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="409" phylum="Arthropoda" rank="genus" status="gen. nov.">Papuadocus</taxonomicName>
<taxonomicNameLabel id="B8ED6252EE56FFEAF94DFD1242350D64" box="[1744,1841,765,787]" pageId="3" pageNumber="409" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="2F9A0179EE560016FFEAFCC442580D35" box="[119,1884,811,834]" gridrow="1" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFCC444780D35" box="[119,380,811,834]" gridcol="0" gridrow="1" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFCC345AD0D35" bold="true" box="[119,169,812,834]" pageId="3" pageNumber="409">Eyes</emphasis>
</th>
<td id="6C4B6805EE560016FE30FCC447750D35" box="[429,625,811,834]" gridcol="1" gridrow="1" pageId="3" pageNumber="409">present</td>
<td id="6C4B6805EE560016FD78FCC446CF0D35" box="[741,971,811,834]" gridcol="2" gridrow="1" pageId="3" pageNumber="409">present/absent</td>
<td id="6C4B6805EE560016FB97FCC4401C0D35" box="[1034,1304,811,834]" gridcol="3" gridrow="1" pageId="3" pageNumber="409">present</td>
<td id="6C4B6805EE560016FAD6FCC443190D35" box="[1355,1565,811,834]" gridcol="4" gridrow="1" pageId="3" pageNumber="409">absent</td>
<td id="6C4B6805EE560016F9D5FCC442580D35" box="[1608,1884,811,834]" gridcol="5" gridrow="1" pageId="3" pageNumber="409">absent</td>
</tr>
<tr id="2F9A0179EE560016FFEAFCB742580D18" box="[119,1884,856,879]" gridrow="2" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFCB744780D18" box="[119,380,856,879]" gridcol="0" gridrow="2" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFCB6441D0D18" bold="true" box="[119,281,857,879]" pageId="3" pageNumber="409">Cephalic notch</emphasis>
</th>
<td id="6C4B6805EE560016FE30FCB747750D18" box="[429,625,856,879]" gridcol="1" gridrow="2" pageId="3" pageNumber="409">present</td>
<td id="6C4B6805EE560016FD78FCB746CF0D18" box="[741,971,856,879]" gridcol="2" gridrow="2" pageId="3" pageNumber="409">present /absent</td>
<td id="6C4B6805EE560016FB97FCB7401C0D18" box="[1034,1304,856,879]" gridcol="3" gridrow="2" pageId="3" pageNumber="409">present /absent</td>
<td id="6C4B6805EE560016FAD6FCB743190D18" box="[1355,1565,856,879]" gridcol="4" gridrow="2" pageId="3" pageNumber="409">absent</td>
<td id="6C4B6805EE560016F9D5FCB742580D18" box="[1608,1884,856,879]" gridcol="5" gridrow="2" pageId="3" pageNumber="409">absent</td>
</tr>
<tr id="2F9A0179EE560016FFEAFC6942580DEB" box="[119,1884,902,924]" gridrow="3" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFC6944780DEB" box="[119,380,902,924]" gridcol="0" gridrow="3" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFC6945AB0DEB" bold="true" box="[119,175,902,924]" pageId="3" pageNumber="409">A12</emphasis>
</th>
<td id="6C4B6805EE560016FE30FC6947750DEB" box="[429,625,902,924]" gridcol="1" gridrow="3" pageId="3" pageNumber="409">A1&gt;A2</td>
<td id="6C4B6805EE560016FD78FC6946CF0DEB" box="[741,971,902,924]" gridcol="2" gridrow="3" pageId="3" pageNumber="409">A1&gt;A2</td>
<td id="6C4B6805EE560016FB97FC69401C0DEB" box="[1034,1304,902,924]" gridcol="3" gridrow="3" pageId="3" pageNumber="409">A1&gt;A2</td>
<td id="6C4B6805EE560016FAD6FC6943190DEB" box="[1355,1565,902,924]" gridcol="4" gridrow="3" pageId="3" pageNumber="409">A1=A2</td>
<td id="6C4B6805EE560016F9D5FC6942580DEB" box="[1608,1884,902,924]" gridcol="5" gridrow="3" pageId="3" pageNumber="409">A1&gt;A2</td>
</tr>
<tr id="2F9A0179EE560016FFEAFC5C42580DBE" box="[119,1884,947,969]" gridrow="4" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFC5C44780DBE" box="[119,380,947,969]" gridcol="0" gridrow="4" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFC5C444A0DBE" bold="true" box="[119,334,947,969]" pageId="3" pageNumber="409">Accessory flagellum</emphasis>
</th>
<td id="6C4B6805EE560016FE30FC5C47750DBE" box="[429,625,947,969]" gridcol="1" gridrow="4" pageId="3" pageNumber="409">&gt;1</td>
<td id="6C4B6805EE560016FD78FC5C46CF0DBE" box="[741,971,947,969]" gridcol="2" gridrow="4" pageId="3" pageNumber="409">&gt;2</td>
<td id="6C4B6805EE560016FB97FC5C401C0DBE" box="[1034,1304,947,969]" gridcol="3" gridrow="4" pageId="3" pageNumber="409">48</td>
<td id="6C4B6805EE560016FAD6FC5C43190DBE" box="[1355,1565,947,969]" gridcol="4" gridrow="4" pageId="3" pageNumber="409">610</td>
<td id="6C4B6805EE560016F9D5FC5C42580DBE" box="[1608,1884,947,969]" gridcol="5" gridrow="4" pageId="3" pageNumber="409">89</td>
</tr>
<tr id="2F9A0179EE560016FFEAFC0F42580D81" box="[119,1884,992,1014]" gridrow="5" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFC0F44780D81" box="[119,380,992,1014]" gridcol="0" gridrow="5" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFC0F44100D81" bold="true" box="[119,276,992,1014]" pageId="3" pageNumber="409">Md palp art. 1</emphasis>
</th>
<td id="6C4B6805EE560016FE30FC0F47750D81" box="[429,625,992,1014]" gridcol="1" gridrow="5" pageId="3" pageNumber="409">untoothed</td>
<td id="6C4B6805EE560016FD78FC0F46CF0D81" box="[741,971,992,1014]" gridcol="2" gridrow="5" pageId="3" pageNumber="409">toothed or not</td>
<td id="6C4B6805EE560016FB97FC0F401C0D81" box="[1034,1304,992,1014]" gridcol="3" gridrow="5" pageId="3" pageNumber="409">toothed</td>
<td id="6C4B6805EE560016FAD6FC0F43190D81" box="[1355,1565,992,1014]" gridcol="4" gridrow="5" pageId="3" pageNumber="409">untoothed</td>
<td id="6C4B6805EE560016F9D5FC0F42580D81" box="[1608,1884,992,1014]" gridcol="5" gridrow="5" pageId="3" pageNumber="409">untoothed</td>
</tr>
<tr id="2F9A0179EE560016FFEAFBE242580A54" box="[119,1884,1037,1059]" gridrow="6" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFBE244780A54" box="[119,380,1037,1059]" gridcol="0" gridrow="6" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFBE244100A54" bold="true" box="[119,276,1037,1059]" pageId="3" pageNumber="409">Md palp art. 3</emphasis>
</th>
<td id="6C4B6805EE560016FE30FBE247750A54" box="[429,625,1037,1059]" gridcol="1" gridrow="6" pageId="3" pageNumber="409">falcate</td>
<td id="6C4B6805EE560016FD78FBE246CF0A54" box="[741,971,1037,1059]" gridcol="2" gridrow="6" pageId="3" pageNumber="409">not falcate</td>
<td id="6C4B6805EE560016FB97FBE2401C0A54" box="[1034,1304,1037,1059]" gridcol="3" gridrow="6" pageId="3" pageNumber="409">not falcate</td>
<td id="6C4B6805EE560016FAD6FBE243190A54" box="[1355,1565,1037,1059]" gridcol="4" gridrow="6" pageId="3" pageNumber="409">not falcate</td>
<td id="6C4B6805EE560016F9D5FBE242580A54" box="[1608,1884,1037,1059]" gridcol="5" gridrow="6" pageId="3" pageNumber="409">falcate</td>
</tr>
<tr id="2F9A0179EE560016FFEAFBD542580A27" box="[119,1884,1082,1104]" gridrow="7" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFBD544780A27" box="[119,380,1082,1104]" gridcol="0" gridrow="7" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFBD5443D0A27" bold="true" box="[119,313,1082,1104]" pageId="3" pageNumber="409">Mx1-2 inner plate</emphasis>
</th>
<td id="6C4B6805EE560016FE30FBD547750A27" box="[429,625,1082,1104]" gridcol="1" gridrow="7" pageId="3" pageNumber="409">setose or not</td>
<td id="6C4B6805EE560016FD78FBD546CF0A27" box="[741,971,1082,1104]" gridcol="2" gridrow="7" pageId="3" pageNumber="409">medially poorly setose</td>
<td id="6C4B6805EE560016FB97FBD5401C0A27" box="[1034,1304,1082,1104]" gridcol="3" gridrow="7" pageId="3" pageNumber="409">medial setae</td>
<td id="6C4B6805EE560016FAD6FBD543190A27" box="[1355,1565,1082,1104]" gridcol="4" gridrow="7" pageId="3" pageNumber="409">setose</td>
<td id="6C4B6805EE560016F9D5FBD542580A27" box="[1608,1884,1082,1104]" gridcol="5" gridrow="7" pageId="3" pageNumber="409">medially setose</td>
</tr>
<tr id="2F9A0179EE560016FFEAFB8842580A0A" box="[119,1884,1127,1149]" gridrow="8" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFB8844780A0A" box="[119,380,1127,1149]" gridcol="0" gridrow="8" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFB8844200A0A" bold="true" box="[119,292,1127,1149]" pageId="3" pageNumber="409">Mx1 inner plate</emphasis>
</th>
<td id="6C4B6805EE560016FE30FB8847750A0A" box="[429,625,1127,1149]" gridcol="1" gridrow="8" pageId="3" pageNumber="409">conical</td>
<td id="6C4B6805EE560016FD78FB8846CF0A0A" box="[741,971,1127,1149]" gridcol="2" gridrow="8" pageId="3" pageNumber="409">ovate</td>
<td id="6C4B6805EE560016FB97FB88401C0A0A" box="[1034,1304,1127,1149]" gridcol="3" gridrow="8" pageId="3" pageNumber="409">triangular</td>
<td id="6C4B6805EE560016FAD6FB8843190A0A" box="[1355,1565,1127,1149]" gridcol="4" gridrow="8" pageId="3" pageNumber="409">triangular</td>
<td id="6C4B6805EE560016F9D5FB8842580A0A" box="[1608,1884,1127,1149]" gridcol="5" gridrow="8" pageId="3" pageNumber="409">triangular</td>
</tr>
<tr id="2F9A0179EE560016FFEAFB7B42580ADC" box="[119,1884,1172,1195]" gridrow="9" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFB7B44780ADC" box="[119,380,1172,1195]" gridcol="0" gridrow="9" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFB7A44780ADC" bold="true" box="[119,380,1173,1195]" pageId="3" pageNumber="409">Mx1 right and left palps</emphasis>
</th>
<td id="6C4B6805EE560016FE30FB7B47750ADC" box="[429,625,1172,1195]" gridcol="1" gridrow="9" pageId="3" pageNumber="409">symmetrical</td>
<td id="6C4B6805EE560016FD78FB7B46CF0ADC" box="[741,971,1172,1195]" gridcol="2" gridrow="9" pageId="3" pageNumber="409">symmetrical</td>
<td id="6C4B6805EE560016FB97FB7B401C0ADC" box="[1034,1304,1172,1195]" gridcol="3" gridrow="9" pageId="3" pageNumber="409">symmetrical</td>
<td id="6C4B6805EE560016FAD6FB7B43190ADC" box="[1355,1565,1172,1195]" gridcol="4" gridrow="9" pageId="3" pageNumber="409">asymmetrical</td>
<td id="6C4B6805EE560016F9D5FB7B42580ADC" box="[1608,1884,1172,1195]" gridcol="5" gridrow="9" pageId="3" pageNumber="409">asymmetrical</td>
</tr>
<tr id="2F9A0179EE560016FFEAFB2E42580AAF" box="[119,1884,1217,1240]" gridrow="10" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFB2E44780AAF" box="[119,380,1217,1240]" gridcol="0" gridrow="10" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFB2D442E0AAF" bold="true" box="[119,298,1218,1240]" pageId="3" pageNumber="409">Mx 2 inner plate</emphasis>
</th>
<td id="6C4B6805EE560016FE30FB2E47750AAF" box="[429,625,1217,1240]" gridcol="1" gridrow="10" pageId="3" pageNumber="409">without facial setae</td>
<td id="6C4B6805EE560016FD78FB2E46CF0AAF" box="[741,971,1217,1240]" gridcol="2" gridrow="10" pageId="3" pageNumber="409">without facial setae</td>
<td id="6C4B6805EE560016FB97FB2E401C0AAF" box="[1034,1304,1217,1240]" gridcol="3" gridrow="10" pageId="3" pageNumber="409">oblique row of facial setae</td>
<td id="6C4B6805EE560016FAD6FB2E43190AAF" box="[1355,1565,1217,1240]" gridcol="4" gridrow="10" pageId="3" pageNumber="409">without facial setae</td>
<td id="6C4B6805EE560016F9D5FB2E42580AAF" box="[1608,1884,1217,1240]" gridcol="5" gridrow="10" pageId="3" pageNumber="409">oblique row of facial setae</td>
</tr>
<tr id="2F9A0179EE560016FFEAFB0042580B72" box="[119,1884,1263,1285]" gridrow="11" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFB0044780B72" box="[119,380,1263,1285]" gridcol="0" gridrow="11" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFB00447D0B72" bold="true" box="[119,377,1263,1285]" pageId="3" pageNumber="409">Gn2 sexually dimorphic</emphasis>
</th>
<td id="6C4B6805EE560016FE30FB0047750B72" box="[429,625,1263,1285]" gridcol="1" gridrow="11" pageId="3" pageNumber="409">yes</td>
<td id="6C4B6805EE560016FD78FB0046CF0B72" box="[741,971,1263,1285]" gridcol="2" gridrow="11" pageId="3" pageNumber="409">yes</td>
<td id="6C4B6805EE560016FB97FB00401C0B72" box="[1034,1304,1263,1285]" gridcol="3" gridrow="11" pageId="3" pageNumber="409">more or less</td>
<td id="6C4B6805EE560016FAD6FB0043190B72" box="[1355,1565,1263,1285]" gridcol="4" gridrow="11" pageId="3" pageNumber="409">yes</td>
<td id="6C4B6805EE560016F9D5FB0042580B72" box="[1608,1884,1263,1285]" gridcol="5" gridrow="11" pageId="3" pageNumber="409">no</td>
</tr>
<tr id="2F9A0179EE560016FFEAFAF342580B45" box="[119,1884,1308,1330]" gridrow="12" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFAF344780B45" box="[119,380,1308,1330]" gridcol="0" gridrow="12" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFAF344420B45" bold="true" box="[119,326,1308,1330]" pageId="3" pageNumber="409">Gn2 male right/left</emphasis>
</th>
<td id="6C4B6805EE560016FE30FAF347750B45" box="[429,625,1308,1330]" gridcol="1" gridrow="12" pageId="3" pageNumber="409">symmetrical</td>
<td id="6C4B6805EE560016FD78FAF346CF0B45" box="[741,971,1308,1330]" gridcol="2" gridrow="12" pageId="3" pageNumber="409">symmetrical</td>
<td id="6C4B6805EE560016FB97FAF3401C0B45" box="[1034,1304,1308,1330]" gridcol="3" gridrow="12" pageId="3" pageNumber="409">symmetrical/asymmetrical</td>
<td id="6C4B6805EE560016FAD6FAF343190B45" box="[1355,1565,1308,1330]" gridcol="4" gridrow="12" pageId="3" pageNumber="409">symetrical</td>
<td id="6C4B6805EE560016F9D5FAF342580B45" box="[1608,1884,1308,1330]" gridcol="5" gridrow="12" pageId="3" pageNumber="409">symmetrical</td>
</tr>
<tr id="2F9A0179EE560016FFEAFAA642580B28" box="[119,1884,1353,1375]" gridrow="13" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFAA644780B28" box="[119,380,1353,1375]" gridcol="0" gridrow="13" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFAA644430B28" bold="true" box="[119,327,1353,1375]" pageId="3" pageNumber="409">Gn 2 dactylus setae</emphasis>
</th>
<td id="6C4B6805EE560016FE30FAA647750B28" box="[429,625,1353,1375]" gridcol="1" gridrow="13" pageId="3" pageNumber="409">01</td>
<td id="6C4B6805EE560016FD78FAA646CF0B28" box="[741,971,1353,1375]" gridcol="2" gridrow="13" pageId="3" pageNumber="409">denselysetose</td>
<td id="6C4B6805EE560016FB97FAA6401C0B28" box="[1034,1304,1353,1375]" gridcol="3" gridrow="13" pageId="3" pageNumber="409">0-1</td>
<td id="6C4B6805EE560016FAD6FAA643190B28" box="[1355,1565,1353,1375]" gridcol="4" gridrow="13" pageId="3" pageNumber="409">-</td>
<td id="6C4B6805EE560016F9D5FAA642580B28" box="[1608,1884,1353,1375]" gridcol="5" gridrow="13" pageId="3" pageNumber="409">1</td>
</tr>
<tr id="2F9A0179EE560016FFEAFA9942580BFB" box="[119,1884,1398,1420]" gridrow="14" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFA9944780BFB" box="[119,380,1398,1420]" gridcol="0" gridrow="14" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFA9944080BFB" bold="true" box="[119,268,1398,1420]" pageId="3" pageNumber="409">Gill on coxa 7</emphasis>
</th>
<td id="6C4B6805EE560016FE30FA9947750BFB" box="[429,625,1398,1420]" gridcol="1" gridrow="14" pageId="3" pageNumber="409">absent</td>
<td id="6C4B6805EE560016FD78FA9946CF0BFB" box="[741,971,1398,1420]" gridcol="2" gridrow="14" pageId="3" pageNumber="409">absent</td>
<td id="6C4B6805EE560016FB97FA99401C0BFB" box="[1034,1304,1398,1420]" gridcol="3" gridrow="14" pageId="3" pageNumber="409">absent</td>
<td id="6C4B6805EE560016FAD6FA9943190BFB" box="[1355,1565,1398,1420]" gridcol="4" gridrow="14" pageId="3" pageNumber="409">present</td>
<td id="6C4B6805EE560016F9D5FA9942580BFB" box="[1608,1884,1398,1420]" gridcol="5" gridrow="14" pageId="3" pageNumber="409">absent</td>
</tr>
<tr id="2F9A0179EE560016FFEAFA4C42580BCE" box="[119,1884,1443,1465]" gridrow="15" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFA4C44780BCE" box="[119,380,1443,1465]" gridcol="0" gridrow="15" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFA4C45C70BCE" bold="true" box="[119,195,1443,1465]" pageId="3" pageNumber="409">Coxa 1</emphasis>
</th>
<td id="6C4B6805EE560016FE30FA4C47750BCE" box="[429,625,1443,1465]" gridcol="1" gridrow="15" pageId="3" pageNumber="409">weakly produced</td>
<td id="6C4B6805EE560016FD78FA4C46CF0BCE" box="[741,971,1443,1465]" gridcol="2" gridrow="15" pageId="3" pageNumber="409">produced or not</td>
<td id="6C4B6805EE560016FB97FA4C401C0BCE" box="[1034,1304,1443,1465]" gridcol="3" gridrow="15" pageId="3" pageNumber="409">produced</td>
<td id="6C4B6805EE560016FAD6FA4C43190BCE" box="[1355,1565,1443,1465]" gridcol="4" gridrow="15" pageId="3" pageNumber="409">not produced</td>
<td id="6C4B6805EE560016F9D5FA4C42580BCE" box="[1608,1884,1443,1465]" gridcol="5" gridrow="15" pageId="3" pageNumber="409">not produced</td>
</tr>
<tr id="2F9A0179EE560016FFEAFA3F42580B91" box="[119,1884,1488,1510]" gridrow="16" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFA3F44780B91" box="[119,380,1488,1510]" gridcol="0" gridrow="16" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFA3F45C70B91" bold="true" box="[119,195,1488,1510]" pageId="3" pageNumber="409">Coxa 4</emphasis>
</th>
<td id="6C4B6805EE560016FE30FA3F47750B91" box="[429,625,1488,1510]" gridcol="1" gridrow="16" pageId="3" pageNumber="409">posterodistal lobe</td>
<td id="6C4B6805EE560016FD78FA3F46CF0B91" box="[741,971,1488,1510]" gridcol="2" gridrow="16" pageId="3" pageNumber="409">lobate or not</td>
<td id="6C4B6805EE560016FB97FA3F401C0B91" box="[1034,1304,1488,1510]" gridcol="3" gridrow="16" pageId="3" pageNumber="409">scarcely lobate</td>
<td id="6C4B6805EE560016FAD6FA3F43190B91" box="[1355,1565,1488,1510]" gridcol="4" gridrow="16" pageId="3" pageNumber="409">not lobate</td>
<td id="6C4B6805EE560016F9D5FA3F42580B91" box="[1608,1884,1488,1510]" gridcol="5" gridrow="16" pageId="3" pageNumber="409">not lobate</td>
</tr>
<tr id="2F9A0179EE560016FFEAFA1242580863" box="[119,1884,1533,1556]" gridrow="17" pageId="3" pageNumber="409">
<th id="6C4B6805EE560016FFEAFA1244780863" box="[119,380,1533,1556]" gridcol="0" gridrow="17" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAFFEAFA1145CB0863" bold="true" box="[119,207,1534,1556]" pageId="3" pageNumber="409">P7 basis</emphasis>
</th>
<td id="6C4B6805EE560016FE30FA1247750863" box="[429,625,1533,1556]" gridcol="1" gridrow="17" pageId="3" pageNumber="409">weakly expanded</td>
<td id="6C4B6805EE560016FD78FA1246CF0863" box="[741,971,1533,1556]" gridcol="2" gridrow="17" pageId="3" pageNumber="409">weakly expanded</td>
<td id="6C4B6805EE560016FB97FA12401C0863" box="[1034,1304,1533,1556]" gridcol="3" gridrow="17" pageId="3" pageNumber="409">weakly expanded</td>
<td id="6C4B6805EE560016FAD6FA1243190863" box="[1355,1565,1533,1556]" gridcol="4" gridrow="17" pageId="3" pageNumber="409">weakly expanded</td>
<td id="6C4B6805EE560016F9D5FA1242580863" box="[1608,1884,1533,1556]" gridcol="5" gridrow="17" pageId="3" pageNumber="409">expanded</td>
</tr>
</table>
</paragraph>
<paragraph id="9115033BEE56FFECF998F9C341E10FFF" blockId="3.[1541,1884,1580,1601]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="411" pageId="3" pageNumber="409">
<emphasis id="A3DEDF29EE56FFEAF998F9C342580836" box="[1541,1884,1580,1601]" italics="true" pageId="3" pageNumber="409">……continued on the next page</emphasis>
than or subequal in length to pereopod 6; coxa with posteroventral lobe or with large anteroventral lobe or with small anteroventral lobe or without lobes. Pereopod 7 subequal in length to, or longer than pereopod 5. Pleonites 13 without dorsal carina. Urosomites 13 free; with or without slender or robust dorsal setae. Urosomite 1 without large distoventral robust seta. Urosomite 2 with pair of dorsal concavities, each with 13 small setae or without dorsal setae. Uropod 1 with 1 or 2 basofacial robust setae or without basofacial robust setae. Uropod 3 sexually dimorphic or not; biramous, without plumose setae; endopod shorter than or subequal to exopod.
<taxonomicName id="56AA78B8EE50FFECFB63FEA3404C0F12" box="[1278,1352,332,357]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="5" pageNumber="418" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
deeply cleft to entire; dorsal or lateral robust setae present or absent; apical robust setae present or absent.
</paragraph>
</subSubSection>
</treatment>
</document>