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<mods:title id="3D5C536C3FD94065644C5A44F3E4DBB3">The return of the Duke - locality data for Megadytes ducalis Sharp, 1882, the world's largest diving beetle, with notes on related species (Coleoptera: Dytiscidae)</mods:title>
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<taxonomicName id="4786D8B3FFB2FFE3C1B06953FD494A75" ID-CoL="7VSTL" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[151,680,1876,1902]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="ducalis" subGenus="Bifurcitus">
<emphasis id="B2F27F22FFB2FFE3C1B06953FD494A75" bold="true" box="[151,680,1876,1902]" pageId="1" pageNumber="518">
<emphasis id="B2F27F22FFB2FFE3C1B06953FEF94A75" bold="true" box="[151,280,1876,1902]" italics="true" pageId="1" pageNumber="518">Megadytes</emphasis>
(
<emphasis id="B2F27F22FFB2FFE3C00F6953FE434A75" bold="true" box="[296,418,1876,1902]" italics="true" pageId="1" pageNumber="518">Bifurcitus</emphasis>
)
<emphasis id="B2F27F22FFB2FFE3C0936953FDEB4A75" bold="true" box="[436,522,1876,1902]" italics="true" pageId="1" pageNumber="518">ducalis</emphasis>
<bibRefCitation id="E417DEC1FFB2FFE3C3366953FD494A75" author="Sharp, D." box="[529,680,1876,1902]" pageId="1" pageNumber="518" pagination="179 - 1003" refId="ref7539" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
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<taxonomicName id="4786D8B3FFB2FFE3C1B0699DFDE94AAB" ID-CoL="7VSTL" authority="Sharp, 1882: 713" authorityName="Sharp" authorityPageNumber="713" authorityYear="1882" box="[151,520,1946,1968]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB2FFE3C1B0699DFEB54AAB" box="[151,340,1946,1968]" italics="true" pageId="1" pageNumber="518">Megadytes ducalis</emphasis>
<treatmentCitation id="01278521FFB2FFE3C07D699DFDE94AAB" author="Sharp, D." box="[346,520,1946,1968]" page="713" pageId="1" pageNumber="518" year="1882">
<bibRefCitation id="E417DEC1FFB2FFE3C07D699DFE394AAB" author="Sharp, D." box="[346,472,1946,1968]" pageId="1" pageNumber="518" pagination="179 - 1003" refId="ref7539" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
: 713
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(orig. descr.);
<treatmentCitation id="01278521FFB2FFE3C3BB699DFC744AAB" author="Zimmermann, A." box="[668,917,1946,1968]" page="256" pageId="1" pageNumber="518" year="1920">
<bibRefCitation id="E417DEC1FFB2FFE3C3BB699DFC824AAB" author="Zimmermann, A." box="[668,867,1946,1968]" pageId="1" pageNumber="518" refId="ref7847" refString="Zimmermann, A. (1920) Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Fol. 4. Pars 71. W. Junk, Berlin, 326 pp." type="book" year="1920">Zimmermann 1920</bibRefCitation>
: 256
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;
<treatmentCitation id="01278521FFB2FFE3C287699DFB634AAB" author="Blackwelder, R. E." box="[928,1154,1946,1968]" page="80" pageId="1" pageNumber="518" year="1944">
<bibRefCitation id="E417DEC1FFB2FFE3C287699DFBBD4AAB" author="Blackwelder, R. E." box="[928,1116,1946,1968]" pageId="1" pageNumber="518" pagination="1 - 188" refId="ref6239" refString="Blackwelder, R. E. (1944) Checklist of the coleopterous insects of Mexico, Central America, the West Indies and South America. Bulletin of the United States National Museum, 185 (1), XII + 1 - 188." type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
: 80
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.
<taxonomicName id="4786D8B3FFB2FFE3C1B069BEFDB34AD4" ID-CoL="7VSTL" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[151,594,1977,1999]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="518" phylum="Arthropoda" rank="species" species="ducalis" subGenus="Bifurcitus">
<emphasis id="B2F27F22FFB2FFE3C1B069BEFEE44AD4" box="[151,261,1977,1999]" italics="true" pageId="1" pageNumber="518">Megadytes</emphasis>
(
<emphasis id="B2F27F22FFB2FFE3C03469BEFE974AD4" box="[275,374,1977,1999]" italics="true" pageId="1" pageNumber="518">Bifurcitus</emphasis>
)
<emphasis id="B2F27F22FFB2FFE3C0A269BEFE2F4AD4" box="[389,462,1977,1999]" italics="true" pageId="1" pageNumber="518">ducalis</emphasis>
<bibRefCitation id="E417DEC1FFB2FFE3C0F369BEFDB34AD4" author="Sharp, D." box="[468,594,1977,1999]" pageId="1" pageNumber="518" pagination="179 - 1003" refId="ref7539" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="01278521FFB2FFE3C37B69BEFD174AD4" author="Brinck, P." box="[604,758,1977,1999]" page="8" pageId="1" pageNumber="518" year="1945">
<bibRefCitation id="E417DEC1FFB2FFE3C37B69BEFD3F4AD4" author="Brinck, P." box="[604,734,1977,1999]" pageId="1" pageNumber="518" pagination="1 - 20" refId="ref6407" refString="Brinck, P. (1945) Nomenklatorische und systematische Studien uber Dytisciden. III. Die Klassifikation der Cybisterinen. Kungliga Fysiografiska Sallskapets Handlingar, 56 (4), 1 - 20." type="journal article" year="1945">Brinck 1945</bibRefCitation>
: 8
</treatmentCitation>
; Trémouilles 1989b: 153;
<treatmentCitation id="01278521FFB2FFE3C52569BEFACB4AD4" author="Miller, K. B. &amp; Bergsten, J." box="[1026,1322,1977,1999]" page="108" pageId="1" pageNumber="518" year="2016">
<bibRefCitation id="E417DEC1FFB2FFE3C52569BEFB194AD4" author="Miller, K. B. &amp; Bergsten, J." box="[1026,1272,1977,1999]" pageId="1" pageNumber="518" refId="ref7159" refString="Miller, K. B. &amp; Bergsten, J. (2016) Diving beetles of the World. Systematics and biology of the Dytiscidae. Johns Hopkins University Press, Baltimore, 320 pp." type="book" year="2016">Miller &amp; Bergsten 2016</bibRefCitation>
: 108
</treatmentCitation>
;
<treatmentCitation id="01278521FFB2FFE3C41369BEFE874AF5" author="Nilsson, A. N. &amp; Hajek, J." page="80" pageId="1" pageNumber="518" year="2019">
<bibRefCitation id="E417DEC1FFB2FFE3C41369BEFEA14AF5" author="Nilsson, A. N. &amp; Hajek, J." pageId="1" pageNumber="518" refId="ref7262" refString="Nilsson, A. N. &amp; Hajek, J. (2019) A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Fersion 1. I. 2019. Distributed as a PDF file via Internet. Available from: http: // www. waterbeetles. eu (accessed 8 February 2019)" type="url" year="2019">Nilsson &amp; Hájek 2019</bibRefCitation>
: 80
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<caption id="D4F9F3B8FFB1FFE0C1B0688BFBA74BDB" httpUri="https://zenodo.org/record/2647352/files/figure.png" pageId="2" pageNumber="519" startId="2.[151,249,1676,1698]" targetBox="[171,1414,205,1629]" targetPageId="2">
<paragraph id="8039A330FFB1FFE0C1B0688BFBA74BDB" blockId="2.[151,1437,1676,1728]" pageId="2" pageNumber="519">
<emphasis id="B2F27F22FFB1FFE0C1B0688BFEF54BB9" bold="true" box="[151,276,1676,1698]" pageId="2" pageNumber="519">FIGURE 1.</emphasis>
Habitus in dorsal view of: A)
<taxonomicName id="4786D8B3FFB1FFE0C37D688AFD274BB9" authorityName="Sharp" authorityYear="1882" box="[602,710,1676,1698]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB1FFE0C37D688AFD274BB9" box="[602,710,1676,1698]" italics="true" pageId="2" pageNumber="519">M. ducalis</emphasis>
</taxonomicName>
, B)
<taxonomicName id="4786D8B3FFB1FFE0C3DF688AFC6F4BB9" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1829" box="[760,910,1676,1698]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="lherminieri">
<emphasis id="B2F27F22FFB1FFE0C3DF688AFC6F4BB9" box="[760,910,1676,1698]" italics="true" pageId="2" pageNumber="519">M. lherminieri</emphasis>
</taxonomicName>
, C)
<taxonomicName id="4786D8B3FFB1FFE0C298688AFBD34BB9" authorityName="Tremouilles &amp; Bachmann" authorityYear="1980" box="[959,1074,1677,1698]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B2F27F22FFB1FFE0C298688AFBD34BB9" box="[959,1074,1677,1698]" italics="true" pageId="2" pageNumber="519">M. magnus</emphasis>
</taxonomicName>
; habitus in ventral view of: D)
<taxonomicName id="4786D8B3FFB1FFE0C4A7688AFF014BDB" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB1FFE0C4A7688AFF014BDB" italics="true" pageId="2" pageNumber="519">M. ducalis</emphasis>
</taxonomicName>
, E)
<taxonomicName id="4786D8B3FFB1FFE0C02C68ACFE7F4BDB" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1829" box="[267,414,1706,1728]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="lherminieri">
<emphasis id="B2F27F22FFB1FFE0C02C68ACFE7F4BDB" box="[267,414,1706,1728]" italics="true" pageId="2" pageNumber="519">M. lherminieri</emphasis>
</taxonomicName>
(Ecuador, Esmeraldas), F)
<taxonomicName id="4786D8B3FFB1FFE0C39368ACFCC54BDB" authorityName="Tremouilles &amp; Bachmann" authorityYear="1980" box="[692,804,1707,1728]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B2F27F22FFB1FFE0C39368ACFCC54BDB" box="[692,804,1707,1728]" italics="true" pageId="2" pageNumber="519">M. magnus</emphasis>
</taxonomicName>
(Paraguay, Cerro Lambaré).
</paragraph>
</caption>
<subSubSection id="C89CF0BBFFB1FFE4C1B068E0FE014C93" lastPageId="6" lastPageNumber="523" pageId="2" pageNumber="519" type="materials_examined">
<paragraph id="8039A330FFB1FFE0C1B068E0FE694A1B" blockId="2.[151,1436,1767,2008]" box="[151,392,1767,1792]" pageId="2" pageNumber="519">
<materialsCitation id="30EEA96DFFB1FFE0C1B068E0FE654A1B" ID-GBIF-Occurrence="2236620147" box="[151,388,1767,1792]" country="Brazil" location="Brazil" pageId="2" pageNumber="519" specimenCount="1" typeStatus="holotype">
<emphasis id="B2F27F22FFB1FFE0C1B068E0FECC4A1B" bold="true" box="[151,301,1767,1792]" pageId="2" pageNumber="519">
<typeStatus id="5F3D1D92FFB1FFE0C1B068E0FF334A1B" box="[151,210,1767,1792]" pageId="2" pageNumber="519">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F891E3A0FFB1FFE0C01B68E0FE654A1B" box="[316,388,1767,1792]" name="Brazil" pageId="2" pageNumber="519">Brazil</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="8039A330FFB1FFE0C1E0690CFA7D4A3F" blockId="2.[151,1436,1767,2008]" box="[199,1436,1803,1828]" pageId="2" pageNumber="519">
<emphasis id="B2F27F22FFB1FFE0C1E0690CFD834A3F" bold="true" box="[199,610,1803,1828]" pageId="2" pageNumber="519">
Type material examined.
<materialsCitation id="30EEA96DFFB1FFE0C0D2690CFD834A3F" ID-GBIF-Occurrence="2236620161" box="[501,610,1803,1828]" collectionCode="BMNH" country="Brazil" location="Saunders" municipality="Type" pageId="2" pageNumber="519" specimenCount="1" typeStatus="holotype">
<typeStatus id="5F3D1D92FFB1FFE0C0D2690CFD834A3F" box="[501,610,1803,1828]" pageId="2" pageNumber="519" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
,
<emphasis id="B2F27F22FFB1FFE0C357690CFCC24A3F" bold="true" box="[624,803,1803,1828]" pageId="2" pageNumber="519">male (BMNH):</emphasis>
&quot;
<collectingCountry id="F891E3A0FFB1FFE0C213690CFC9C4A3F" box="[820,893,1803,1828]" name="Brazil" pageId="2" pageNumber="519">Brazil</collectingCountry>
,
<location id="8559F5EBFFB1FFE0C2AF690CFC154A3F" LSID="urn:lsid:plazi:treatment:082F1226FFB2FFE4C1B06953FE464E18:8559F5EBFFB1FFE0C2AF690CFC154A3F" box="[904,1012,1803,1828]" country="Brazil" municipality="Type" name="Saunders" pageId="2" pageNumber="519">Saunders</location>
&quot;, &quot;
<collectingMunicipality id="605D394AFFB1FFE0C536690BFBB14A3F" box="[1041,1104,1804,1828]" pageId="2" pageNumber="519">Type</collectingMunicipality>
&quot;, &quot;
<taxonomicName id="4786D8B3FFB1FFE0C54A690CFA6E4A3F" authority="Sharp" authorityName="Sharp" authorityYear="1882" box="[1133,1423,1803,1828]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="519" phylum="Arthropoda" rank="species" species="ducalis">Megadytes ducalis Sharp</taxonomicName>
&quot;.
</paragraph>
<paragraph id="8039A330FFB1FFE0C1E06928FA7D4AC3" blockId="2.[151,1436,1767,2008]" pageId="2" pageNumber="519">
<emphasis id="B2F27F22FFB1FFE0C1E06928FE174A53" bold="true" box="[199,502,1839,1864]" pageId="2" pageNumber="519">Other material examined</emphasis>
.
<materialsCitation id="30EEA96DFFB1FFE0C3236928FD164A77" ID-GBIF-Occurrence="2236620151" country="Brazil" elevation="660" location="San Antonio da Barra" municipality="Prov. de Bahia" pageId="2" pageNumber="519" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<emphasis id="B2F27F22FFB1FFE0C3236928FDAC4A53" bold="true" box="[516,589,1839,1864]" pageId="2" pageNumber="519">
<collectingCountry id="F891E3A0FFB1FFE0C3236928FDAC4A53" box="[516,589,1839,1864]" name="Brazil" pageId="2" pageNumber="519">Brazil</collectingCountry>
</emphasis>
:
<specimenCount id="968068B9FFB1FFE0C37A6937FD4D4A53" box="[605,684,1839,1864]" pageId="2" pageNumber="519" type="male">1 male</specimenCount>
,
<specimenCount id="968068B9FFB1FFE0C39F6937FCFD4A53" box="[696,796,1839,1864]" pageId="2" pageNumber="519" type="female">1 female</specimenCount>
, &quot;
<location id="8559F5EBFFB1FFE0C2166937FBC84A5C" LSID="urn:lsid:plazi:treatment:082F1226FFB2FFE4C1B06953FE464E18:8559F5EBFFB1FFE0C2166937FBC84A5C" box="[817,1065,1839,1864]" country="Brazil" municipality="Prov. de Bahia" name="San Antonio da Barra" pageId="2" pageNumber="519">San Antônio da Barra</location>
[= Condeúba, City Center is at - 14.899° -41.968°,
<quantity id="477E0ED5FFB1FFE0C0546954FE244A70" box="[371,453,1875,1900]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="519" unit="m" value="660.0">
<elevation id="0BAB4403FFB1FFE0C0546954FE244A70" box="[371,453,1875,1900]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="519" unit="m" value="660.0">660 m</elevation>
</quantity>
],
<collectingMunicipality id="605D394AFFB1FFE0C0F86953FD724A77" box="[479,659,1875,1900]" pageId="2" pageNumber="519">Prov. de Bahia</collectingMunicipality>
, Brésil&quot;
</materialsCitation>
;
<materialsCitation id="30EEA96DFFB1FFE0C22D6953FE2B4A8B" ID-GBIF-Occurrence="2236620153" collectingDate="1888-11-12" collectorName="S. Antonia da Barra" country="Brazil" location="Bahia Gounelle" pageId="2" pageNumber="519" specimenCount="1" specimenCount-female="1">
<specimenCount id="968068B9FFB1FFE0C22D6953FC944A77" box="[778,885,1875,1900]" pageId="2" pageNumber="519" type="female">1 female</specimenCount>
, &quot;
<collectorName id="2D73C6E6FFB1FFE0C2B56953FB644A70" box="[914,1157,1875,1900]" pageId="2" pageNumber="519">S. Antonia da Barra</collectorName>
pr. de
<location id="8559F5EBFFB1FFE0C5C16954FA7D4A77" LSID="urn:lsid:plazi:treatment:082F1226FFB2FFE4C1B06953FE464E18:8559F5EBFFB1FFE0C5C16954FA7D4A77" box="[1254,1436,1875,1900]" country="Brazil" name="Bahia Gounelle" pageId="2" pageNumber="519">Bahia Gounelle</location>
<date id="F43885F0FFB1FFE0C1B0697FFEF64A8B" box="[151,279,1912,1936]" pageId="2" pageNumber="519" value="1888-11-12">
<collectingDate id="E47C7C18FFB1FFE0C1B0697FFEF64A8B" box="[151,279,1912,1936]" pageId="2" pageNumber="519" value="1888-11-12">11.12.1888</collectingDate>
</date>
&quot; [printed label]
</materialsCitation>
;
<materialsCitation id="30EEA96DFFB1FFE0C0F1697FFEA64AAF" ID-GBIF-Occurrence="2236620157" country="Brazil" location="Antonia da Barra" pageId="2" pageNumber="519" specimenCount="1" specimenCount-female="1">
<specimenCount id="968068B9FFB1FFE0C0F1697FFDDA4A8B" box="[470,571,1911,1936]" pageId="2" pageNumber="519" type="female">1 female</specimenCount>
, &quot;S.
<location id="8559F5EBFFB1FFE0C3576970FCD34A94" LSID="urn:lsid:plazi:treatment:082F1226FFB2FFE4C1B06953FE464E18:8559F5EBFFB1FFE0C3576970FCD34A94" box="[624,818,1911,1936]" country="Brazil" name="Antonia da Barra" pageId="2" pageNumber="519">Antonia da Barra</location>
pr. De Bahia CH Pujol 1890&quot; [printed label] (MNHN- CR, drawer 60)
</materialsCitation>
;
<materialsCitation id="30EEA96DFFB1FFE0C075699BFD874AAF" ID-GBIF-Occurrence="2236620174" box="[338,614,1947,1972]" country="Brazil" location="Bahia Bresil" pageId="2" pageNumber="519" specimenCount="1" specimenCount-female="1">
<specimenCount id="968068B9FFB1FFE0C075699BFE574AAF" box="[338,438,1947,1972]" pageId="2" pageNumber="519" type="female">1 female</specimenCount>
, &quot;
<location id="8559F5EBFFB1FFE0C0EE699CFDBC4AAF" LSID="urn:lsid:plazi:treatment:082F1226FFB2FFE4C1B06953FE464E18:8559F5EBFFB1FFE0C0EE699CFDBC4AAF" box="[457,605,1947,1972]" country="Brazil" name="Bahia Bresil" pageId="2" pageNumber="519">Bahia Brésil</location>
&quot;
</materialsCitation>
;
<materialsCitation id="30EEA96DFFB1FFE0C356699BFDE54AC3" ID-GBIF-Occurrence="2236620187" collectingDate="1888-11-12" collectionCode="MNHN-CG" country="Brazil" location="Antonia da Barra Pr. de Bahia Gounelle" pageId="2" pageNumber="519" specimenCount="1" specimenCount-male="1">
<specimenCount id="968068B9FFB1FFE0C356699BFD5F4AAF" box="[625,702,1947,1972]" pageId="2" pageNumber="519" type="male">1 male</specimenCount>
, &quot;S.
<location id="8559F5EBFFB1FFE0C3D5699CFB4C4AAF" LSID="urn:lsid:plazi:treatment:082F1226FFB2FFE4C1B06953FE464E18:8559F5EBFFB1FFE0C3D5699CFB4C4AAF" box="[754,1197,1947,1972]" country="Brazil" name="Antonia da Barra Pr. de Bahia Gounelle" pageId="2" pageNumber="519">Antonia da Barra Pr. de Bahia Gounelle</location>
<date id="F43885F0FFB1FFE0C593699BFAD54AAF" box="[1204,1332,1948,1972]" pageId="2" pageNumber="519" value="1888-11-12">
<collectingDate id="E47C7C18FFB1FFE0C593699BFAD54AAF" box="[1204,1332,1948,1972]" pageId="2" pageNumber="519" value="1888-11-12">11.12.1888</collectingDate>
</date>
&quot; [printed label] (
<collectionCode id="E6973BF5FFB1FFE0C1CD69C7FE904AC3" box="[234,369,1984,2008]" pageId="2" pageNumber="519">MNHN-CG</collectionCode>
, drawer 92)
</materialsCitation>
;
<materialsCitation id="30EEA96DFFB1FFE0C33369C7FA7D4AC3" ID-GBIF-Occurrence="2236620186" box="[532,1436,1983,2008]" country="Brazil" pageId="2" pageNumber="519" specimenCount="1" specimenCount-male="1">
<specimenCount id="968068B9FFB1FFE0C33369C7FD864AC3" box="[532,615,1983,2008]" pageId="2" pageNumber="519" type="male">1 male</specimenCount>
, &quot;Museum Paris Brésil Coll. E. Gounelle 1915&quot; [green printed label]
</materialsCitation>
</paragraph>
<paragraph id="8039A330FFB0FFE1C1B06E9FFDAC4C06" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="520">
[among indet. material in separate drawer];
<materialsCitation id="30EEA96DFFB0FFE1C3926E9FFDFB4DCF" ID-GBIF-Occurrence="2236620160" collectionCode="MNHN-CW" country="Brazil" location="Bahia" pageId="3" pageNumber="520" specimenCount="1" specimenCount-female="1">
<specimenCount id="968068B9FFB0FFE1C3926E9FFCC34DAB" box="[693,802,151,176]" pageId="3" pageNumber="520" type="female">1 female</specimenCount>
, &quot;
<collectingCountry id="F891E3A0FFB0FFE1C2196E90FC654DAB" box="[830,900,151,176]" name="Brazil" pageId="3" pageNumber="520">Brésil</collectingCountry>
<location id="8559F5EBFFB0FFE1C2B46E90FC344DAB" LSID="urn:lsid:plazi:treatment:082F1226FFB2FFE4C1B06953FE464E18:8559F5EBFFB0FFE1C2B46E90FC344DAB" box="[915,981,151,176]" country="Brazil" name="Bahia" pageId="3" pageNumber="520">Bahia</location>
intr&quot; [printed label], &quot;
<taxonomicName id="4786D8B3FFB0FFE1C5C96E90FA724DAB" authority="Sharp" authorityName="Sharp" authorityYear="1882" box="[1262,1427,151,176]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="ducalis">ducalis Sharp</taxonomicName>
&quot; [handwritten label] (
<collectionCode id="E6973BF5FFB0FFE1C0586EBBFDF34DCF" box="[383,530,188,212]" pageId="3" pageNumber="520">MNHN-CW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="30EEA96DFFB0FFE1C3006EBBFB4F4DCE" ID-GBIF-Occurrence="2236620168" box="[551,1198,188,213]" country="Brazil" location="Bresil" pageId="3" pageNumber="520" specimenCount="1" specimenCount-female="1">
<specimenCount id="968068B9FFB0FFE1C3006EBBFD6B4DCE" box="[551,650,188,213]" pageId="3" pageNumber="520" type="female">1 female</specimenCount>
&quot;
<taxonomicName id="4786D8B3FFB0FFE1C3BB6EBBFC934DCE" authorityName="Laporte" authorityYear="1835" box="[668,882,188,213]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="giganteus">Cybister giganteus</taxonomicName>
<collectingCountry id="F891E3A0FFB0FFE1C25C6EBBFC214DCE" box="[891,960,188,213]" name="Brazil" pageId="3" pageNumber="520">Brésil</collectingCountry>
&quot; [handwritten label]
</materialsCitation>
;
<materialsCitation id="30EEA96DFFB0FFE1C59C6EBBFDAB4C06" ID-GBIF-Occurrence="2236620164" collectorName="Coll. E. Gounelle" country="Brazil" location="Bresil" pageId="3" pageNumber="520" specimenCount="1" specimenCount-female="1">
<specimenCount id="968068B9FFB0FFE1C59C6EBBFAC04DCE" box="[1211,1313,188,213]" pageId="3" pageNumber="520" type="female">1 female</specimenCount>
, &quot;Museum Paris
<collectingCountry id="F891E3A0FFB0FFE1C1FB6ED8FEFE4DE3" box="[220,287,223,248]" name="Brazil" pageId="3" pageNumber="520">Brésil</collectingCountry>
<collectorName id="2D73C6E6FFB0FFE1C00B6ED8FE1A4DE3" box="[300,507,223,248]" pageId="3" pageNumber="520">Coll. E. Gounelle</collectorName>
1915&quot; [green printed label], &quot;
<taxonomicName id="4786D8B3FFB0FFE1C2426ED8FA754DE3" authority="Shp. San Antonio da Barra" box="[869,1428,223,248]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="ducalis">Megadytes ducalis Shp. San Antonio da Barra</taxonomicName>
&quot; [handwritten label] (MNHN-Magasin)
</materialsCitation>
.
</paragraph>
<paragraph id="8039A330FFB0FFE1C1E06F20FE934CB7" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="520">
<emphasis id="B2F27F22FFB0FFE1C1E06F20FE724C5B" bold="true" box="[199,403,295,320]" pageId="3" pageNumber="520">Descriptive notes</emphasis>
. Large beetle (TL&gt;
<quantity id="477E0ED5FFB0FFE1C35D6F2FFD3E4C5B" box="[634,735,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="3" pageNumber="520" unit="mm" value="42.9">42.9 mm</quantity>
) of very robust and &quot;thick&quot; appearance (
<figureCitation id="18BDBFB5FFB0FFE1C5866F20FB1A4C5B" box="[1185,1275,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,249,1959,1981]" captionTargetBox="[193,1409,484,1938]" captionTargetId="figure@3.[151,1436,481,1938]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in lateral view of: A) M. ducalis, B) M. lherminieri, C) M. magnus." httpUri="https://zenodo.org/record/2647354/files/figure.png" pageId="3" pageNumber="520">Fig. 2A</figureCitation>
). Descriptions of this species, based on the male
<typeStatus id="5F3D1D92FFB0FFE1C3096F4BFD724C7E" box="[558,659,332,357]" pageId="3" pageNumber="520" type="holotype">holotype</typeStatus>
, were provided by
<bibRefCitation id="E417DEC1FFB0FFE1C25F6F4BFBEE4C7F" author="Sharp, D." box="[888,1039,332,357]" pageId="3" pageNumber="520" pagination="179 - 1003" refId="ref7539" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series 2, 2, 179 - 1003, pls. 7 - 18." type="journal article" year="1882">Sharp (1882)</bibRefCitation>
and Trémouilles (1989). Here we provide photos of the habitus (
<figureCitation id="18BDBFB5FFB0FFE1C0D46F68FD974C9C" box="[499,630,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,249,1676,1698]" captionTargetBox="[171,1414,205,1629]" captionTargetId="figure@2.[166,1421,193,1654]" captionTargetPageId="2" captionText="FIGURE 1. Habitus in dorsal view of: A) M. ducalis, B) M. lherminieri, C) M. magnus; habitus in ventral view of: D) M. ducalis, E) M. lherminieri (Ecuador, Esmeraldas), F) M. magnus (Paraguay, Cerro Lambaré)." httpUri="https://zenodo.org/record/2647352/files/figure.png" pageId="3" pageNumber="520">Figs 1A, D</figureCitation>
,
<figureCitation id="18BDBFB5FFB0FFE1C3A46F77FD464C93" box="[643,679,368,392]" captionStart="FIGURE 2" captionStartId="3.[151,249,1959,1981]" captionTargetBox="[193,1409,484,1938]" captionTargetId="figure@3.[151,1436,481,1938]" captionTargetPageId="3" captionText="FIGURE 2. Habitus in lateral view of: A) M. ducalis, B) M. lherminieri, C) M. magnus." httpUri="https://zenodo.org/record/2647354/files/figure.png" pageId="3" pageNumber="520">2A</figureCitation>
,
<figureCitation id="18BDBFB5FFB0FFE1C3926F77FD224C93" box="[693,707,368,392]" captionStart="FIGURE 3" captionStartId="4.[151,249,1967,1989]" captionTargetBox="[316,1333,134,1856]" captionTargetId="figure@4.[189,1397,133,1946]" captionTargetPageId="4" captionText="FIGURE 3. Female of M. ducalis from collection Régimbart in MNHN [TL = 47.4 mm], the largest known diving beetle specimen in the world: &quot;San Antônio da Barra, Prov. de Bahia, Brésil&quot;." httpUri="https://zenodo.org/record/2647356/files/figure.png" pageId="3" pageNumber="520">3</figureCitation>
) and focus on morphological characters of the male in different views (
<figureCitation id="18BDBFB5FFB0FFE1C1C06F93FECC4CB7" box="[231,301,404,429]" captionStart="FIGURE 7" captionStartId="10.[151,249,1844,1866]" captionTargetBox="[166,1427,204,1822]" captionTargetId="figure@10.[151,1436,193,1822]" captionTargetPageId="10" captionText="FIGURE 7. Megadytes ducalis, male from collection Guignot: A) frontal view, B) ventral view of head and protarsus, C) left middle leg, D) dorso-lateral view of left side of head, pronotum and elytral base, E) ventro-lateral view of left hind leg. Scale bars = 5 mm." httpUri="https://zenodo.org/record/2647364/files/figure.png" pageId="3" pageNumber="520">Figs 7</figureCitation>
AE).
</paragraph>
<caption id="D4F9F3B8FFB0FFE1C1B069A0FBF74AA6" box="[151,1046,1959,1982]" httpUri="https://zenodo.org/record/2647354/files/figure.png" pageId="3" pageNumber="520" startId="3.[151,249,1959,1981]" targetBox="[193,1409,484,1938]" targetPageId="3">
<paragraph id="8039A330FFB0FFE1C1B069A0FBF74AA6" blockId="3.[151,1046,1959,1982]" box="[151,1046,1959,1982]" pageId="3" pageNumber="520">
<emphasis id="B2F27F22FFB0FFE1C1B069A0FEF34AA6" bold="true" box="[151,274,1959,1981]" pageId="3" pageNumber="520">FIGURE 2.</emphasis>
Habitus in lateral view of: A)
<taxonomicName id="4786D8B3FFB0FFE1C36F69AFFD524AA6" box="[584,691,1959,1981]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB0FFE1C36F69AFFD524AA6" box="[584,691,1959,1981]" italics="true" pageId="3" pageNumber="520">M. ducalis</emphasis>
</taxonomicName>
, B)
<taxonomicName id="4786D8B3FFB0FFE1C3F869AFFC924AA6" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1829" box="[735,883,1959,1981]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="lherminieri">
<emphasis id="B2F27F22FFB0FFE1C3F869AFFC924AA6" box="[735,883,1959,1981]" italics="true" pageId="3" pageNumber="520">M. lherminieri</emphasis>
</taxonomicName>
, C)
<taxonomicName id="4786D8B3FFB0FFE1C2B869AFFBF14AA6" authorityName="Tremouilles &amp; Bachmann" authorityYear="1980" box="[927,1040,1960,1981]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="520" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="B2F27F22FFB0FFE1C2B869AFFBF14AA6" box="[927,1040,1960,1981]" italics="true" pageId="3" pageNumber="520">M. magnus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="D4F9F3B8FFB7FFE6C1B069A8FC874AFF" httpUri="https://zenodo.org/record/2647356/files/figure.png" pageId="4" pageNumber="521" startId="4.[151,249,1967,1989]" targetBox="[316,1333,134,1856]" targetPageId="4">
<paragraph id="8039A330FFB7FFE6C1B069A8FC874AFF" blockId="4.[151,1436,1967,2020]" pageId="4" pageNumber="521">
<emphasis id="B2F27F22FFB7FFE6C1B069A8FEF44ADE" bold="true" box="[151,277,1967,1989]" pageId="4" pageNumber="521">FIGURE 3.</emphasis>
Female of
<taxonomicName id="4786D8B3FFB7FFE6C0B769B7FE1C4ADE" box="[400,509,1967,1989]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="521" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB7FFE6C0B769B7FE1C4ADE" box="[400,509,1967,1989]" italics="true" pageId="4" pageNumber="521">M. ducalis</emphasis>
</taxonomicName>
from collection Régimbart in MNHN [TL = 47.4 mm], the largest known diving beetle specimen in the world: &quot;San Antônio da Barra, Prov. de Bahia, Brésil&quot;.
</paragraph>
</caption>
<caption id="D4F9F3B8FFB6FFE7C1B06999FD244AE9" httpUri="https://zenodo.org/record/2647358/files/figure.png" pageId="5" pageNumber="522" startId="5.[151,249,1950,1972]" targetBox="[176,1412,193,1920]" targetPageId="5">
<paragraph id="8039A330FFB6FFE7C1B06999FD244AE9" blockId="5.[151,1436,1950,2034]" pageId="5" pageNumber="522">
<emphasis id="B2F27F22FFB6FFE7C1B06999FEF04AAF" bold="true" box="[151,273,1950,1972]" pageId="5" pageNumber="522">FIGURE 4.</emphasis>
<taxonomicName id="4786D8B3FFB6FFE7C0306999FE354AAF" box="[279,468,1950,1972]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="522" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB6FFE7C0306999FE354AAF" box="[279,468,1950,1972]" italics="true" pageId="5" pageNumber="522">Megadytes ducalis</emphasis>
</taxonomicName>
, male specimen from collection Guignot: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) anterior half median lobe in ventro-apical view;
<taxonomicName id="4786D8B3FFB6FFE7C2F569BAFB6F4AC8" box="[978,1166,1981,2003]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="522" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB6FFE7C2F569BAFB6F4AC8" box="[978,1166,1981,2003]" italics="true" pageId="5" pageNumber="522">Megadytes ducalis</emphasis>
</taxonomicName>
, holotype: D) median lobe in ventral view (Foto: R. Angus). Scale bars = 1.5 mm.
</paragraph>
</caption>
<paragraph id="8039A330FFB5FFE4C1E06E90FE014C93" blockId="6.[151,1437,151,771]" pageId="6" pageNumber="523">
The median lobe, with both parameres, of a male from Coll. Guignot is figured in ventral view (
<figureCitation id="18BDBFB5FFB5FFE4C42D6E90FA844DAB" box="[1290,1381,151,176]" captionStart="FIGURE 4" captionStartId="5.[151,249,1950,1972]" captionTargetBox="[176,1412,193,1920]" captionTargetId="figure@5.[176,1412,193,1920]" captionTargetPageId="5" captionText="FIGURE 4. Megadytes ducalis, male specimen from collection Guignot: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) anterior half median lobe in ventro-apical view; Megadytes ducalis, holotype: D) median lobe in ventral view (Foto: R. Angus). Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647358/files/figure.png" pageId="6" pageNumber="523">Fig. 4A</figureCitation>
), the anterior half of the median lobe in lateral view (
<figureCitation id="18BDBFB5FFB5FFE4C39B6EBBFCF24DCF" box="[700,787,188,213]" captionStart="FIGURE 4" captionStartId="5.[151,249,1950,1972]" captionTargetBox="[176,1412,193,1920]" captionTargetId="figure@5.[176,1412,193,1920]" captionTargetPageId="5" captionText="FIGURE 4. Megadytes ducalis, male specimen from collection Guignot: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) anterior half median lobe in ventro-apical view; Megadytes ducalis, holotype: D) median lobe in ventral view (Foto: R. Angus). Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647358/files/figure.png" pageId="6" pageNumber="523">Fig. 4B</figureCitation>
) and the tip of the median lobe in dorsal view (
<figureCitation id="18BDBFB5FFB5FFE4C4126EBBFA714DCF" box="[1333,1424,188,213]" captionStart="FIGURE 4" captionStartId="5.[151,249,1950,1972]" captionTargetBox="[176,1412,193,1920]" captionTargetId="figure@5.[176,1412,193,1920]" captionTargetPageId="5" captionText="FIGURE 4. Megadytes ducalis, male specimen from collection Guignot: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) anterior half median lobe in ventro-apical view; Megadytes ducalis, holotype: D) median lobe in ventral view (Foto: R. Angus). Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647358/files/figure.png" pageId="6" pageNumber="523">Fig. 4C</figureCitation>
). The median lobe of the
<typeStatus id="5F3D1D92FFB5FFE4C0806ED8FDE84DE3" box="[423,521,223,248]" pageId="6" pageNumber="523" type="holotype">holotype</typeStatus>
is presented in
<figureCitation id="18BDBFB5FFB5FFE4C39A6ED8FCF74DE3" box="[701,790,223,248]" captionStart="FIGURE 4" captionStartId="5.[151,249,1950,1972]" captionTargetBox="[176,1412,193,1920]" captionTargetId="figure@5.[176,1412,193,1920]" captionTargetPageId="5" captionText="FIGURE 4. Megadytes ducalis, male specimen from collection Guignot: A) median lobe with parameres in ventral view, B) tip of median lobe in lateral view, C) anterior half median lobe in ventro-apical view; Megadytes ducalis, holotype: D) median lobe in ventral view (Foto: R. Angus). Scale bars = 1.5 mm." httpUri="https://zenodo.org/record/2647358/files/figure.png" pageId="6" pageNumber="523">Fig. 4D</figureCitation>
. It appears much broader than in the other specimens we studied. This is apparently due to manipulation by previous researchers, perhaps J. Balfour-Browne, whereby cardboard wedges were inserted in between the main lobe and the ventral sclerite to expose the latter. This is actually seen in several other
<taxonomicName id="4786D8B3FFB5FFE4C0C56F4BFDBA4C7E" authorityName="Sharp" authorityYear="1882" box="[482,603,332,357]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis id="B2F27F22FFB5FFE4C0C56F4BFDBA4C7E" box="[482,603,332,357]" italics="true" pageId="6" pageNumber="523">Megadytes</emphasis>
</taxonomicName>
specimens in the BMNH collection, and can lead to significant changes to the shape of the median lobe.
</paragraph>
</subSubSection>
<subSubSection id="C89CF0BBFFB5FFE4C1E06F90FBC84FA8" pageId="6" pageNumber="523" type="description">
<paragraph id="8039A330FFB5FFE4C1E06F90FCC24CE3" blockId="6.[151,1437,151,771]" pageId="6" pageNumber="523">
<emphasis id="B2F27F22FFB5FFE4C1E06F90FEFF4CAB" bold="true" box="[199,286,407,432]" pageId="6" pageNumber="523">Female</emphasis>
. No traces of elytral striae or any other kind of sexual sculpture on pronotum or elytra. Without protarsal disks and yellow setae on ventral surface of mesotarsomere I; otherwise no external difference with male. Broadest part of femur in hind legs
<quantity id="477E0ED5FFB5FFE4C3016FD8FD9D4CEC" box="[550,636,479,504]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="523" unit="mm" value="5.3">5.3 mm</quantity>
(
<figureCitation id="18BDBFB5FFB5FFE4C3AD6FD8FD074CE3" box="[650,742,479,504]" captionStart="FIGURE 1" captionStartId="2.[151,249,1676,1698]" captionTargetBox="[171,1414,205,1629]" captionTargetId="figure@2.[166,1421,193,1654]" captionTargetPageId="2" captionText="FIGURE 1. Habitus in dorsal view of: A) M. ducalis, B) M. lherminieri, C) M. magnus; habitus in ventral view of: D) M. ducalis, E) M. lherminieri (Ecuador, Esmeraldas), F) M. magnus (Paraguay, Cerro Lambaré)." httpUri="https://zenodo.org/record/2647352/files/figure.png" pageId="6" pageNumber="523">Figs 1A</figureCitation>
,
<figureCitation id="18BDBFB5FFB5FFE4C3D46FE7FCF74CE3" box="[755,790,480,504]" captionStart="FIGURE 7" captionStartId="10.[151,249,1844,1866]" captionTargetBox="[166,1427,204,1822]" captionTargetId="figure@10.[151,1436,193,1822]" captionTargetPageId="10" captionText="FIGURE 7. Megadytes ducalis, male from collection Guignot: A) frontal view, B) ventral view of head and protarsus, C) left middle leg, D) dorso-lateral view of left side of head, pronotum and elytral base, E) ventro-lateral view of left hind leg. Scale bars = 5 mm." httpUri="https://zenodo.org/record/2647364/files/figure.png" pageId="6" pageNumber="523">7E</figureCitation>
).
</paragraph>
<paragraph id="8039A330FFB5FFE4C1E06C00FE1D4F7C" blockId="6.[151,1437,151,771]" pageId="6" pageNumber="523">
<emphasis id="B2F27F22FFB5FFE4C1E06C00FE974F3B" bold="true" box="[199,374,519,544]" pageId="6" pageNumber="523">Measurements</emphasis>
,
<emphasis id="B2F27F22FFB5FFE4C0A36C00FE0A4F3B" bold="true" box="[388,491,519,544]" pageId="6" pageNumber="523">
<typeStatus id="5F3D1D92FFB5FFE4C0A36C00FE0A4F3B" box="[388,491,519,544]" pageId="6" pageNumber="523" type="holotype">holotype</typeStatus>
</emphasis>
: TL =
<quantity id="477E0ED5FFB5FFE4C31B6C0FFD454F04" box="[572,676,520,544]" metricMagnitude="-2" metricUnit="m" metricValue="4.72" pageId="6" pageNumber="523" unit="mm" value="47.2">47.2 mm</quantity>
; TL-H =
<quantity id="477E0ED5FFB5FFE4C2286C0FFC964F04" box="[783,887,520,544]" metricMagnitude="-2" metricUnit="m" metricValue="4.41" pageId="6" pageNumber="523" unit="mm" value="44.1">44.1 mm</quantity>
; TW =
<quantity id="477E0ED5FFB5FFE4C2E96C00FBD44F04" box="[974,1077,519,544]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="6" pageNumber="523" unit="mm" value="26.4">26.4 mm</quantity>
.
<emphasis id="B2F27F22FFB5FFE4C5676C00FAA04F3B" bold="true" box="[1088,1345,519,544]" pageId="6" pageNumber="523">Additional specimens</emphasis>
,
<emphasis id="B2F27F22FFB5FFE4C4776C00FA754F3B" bold="true" box="[1360,1428,519,544]" pageId="6" pageNumber="523">males</emphasis>
: TL = 42.947.0 mm; TL-H =
<quantity id="477E0ED5FFB5FFE4C0CD6C2BFD704F58" box="[490,657,556,580]" metricMagnitude="-2" metricUnit="m" metricValue="4.255" metricValueMax="4.38" metricValueMin="4.13" pageId="6" pageNumber="523" unit="mm" value="42.55" valueMax="43.8" valueMin="41.3">41.343.8 mm</quantity>
; TW =
<quantity id="477E0ED5FFB5FFE4C3C06C2CFC6F4F58" box="[743,910,555,580]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.59" metricValueMin="2.51" pageId="6" pageNumber="523" unit="mm" value="25.5" valueMax="25.9" valueMin="25.1">25.125.9 mm</quantity>
.
<emphasis id="B2F27F22FFB5FFE4C2BE6C2CFC1A4F5F" bold="true" box="[921,1019,555,580]" pageId="6" pageNumber="523">Females</emphasis>
. TL =
<quantity id="477E0ED5FFB5FFE4C56C6C2CFB134F58" box="[1099,1266,555,580]" metricMagnitude="-2" metricUnit="m" metricValue="4.545" metricValueMax="4.74" metricValueMin="4.35" pageId="6" pageNumber="523" unit="mm" value="45.45" valueMax="47.4" valueMin="43.5">43.547.4 mm</quantity>
; TL-H =
<quantity id="477E0ED5FFB5FFE4C47A6C2BFF1F4F7C" metricMagnitude="-2" metricUnit="m" metricValue="4.24" metricValueMax="4.41" metricValueMin="4.07" pageId="6" pageNumber="523" unit="mm" value="42.4" valueMax="44.1" valueMin="40.7">40.7 44.1 mm</quantity>
; TW = 24.026,1 mm.
</paragraph>
<paragraph id="8039A330FFB5FFE4C1E06C70FBC84FA8" blockId="6.[151,1437,151,771]" pageId="6" pageNumber="523">
<emphasis id="B2F27F22FFB5FFE4C1E06C70FF1E4F8B" bold="true" box="[199,255,631,656]" pageId="6" pageNumber="523">Note</emphasis>
. The largest known specimen of
<taxonomicName id="4786D8B3FFB5FFE4C3AF6C7FFCE34F8B" box="[648,770,631,656]" class="Insecta" family="Dytiscidae" genus="Megadytes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="523" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="B2F27F22FFB5FFE4C3AF6C7FFCE34F8B" box="[648,770,631,656]" italics="true" pageId="6" pageNumber="523">M. ducalis</emphasis>
</taxonomicName>
(female, TL=
<quantity id="477E0ED5FFB5FFE4C2976C70FBF84F94" box="[944,1049,631,655]" metricMagnitude="-2" metricUnit="m" metricValue="4.74" pageId="6" pageNumber="523" unit="mm" value="47.4">47.4 mm</quantity>
) is deposited in the collection of
<collectingCountry id="F891E3A0FFB5FFE4C1B06C9CFF154FAF" box="[151,244,667,692]" name="Mauritius" pageId="6" pageNumber="523">Maurice</collectingCountry>
Régimbart in the Muséum National d'Histoire Naturelle in Paris (
<figureCitation id="18BDBFB5FFB5FFE4C2F16C9CFBFA4FA8" box="[982,1051,667,692]" captionStart="FIGURE 3" captionStartId="4.[151,249,1967,1989]" captionTargetBox="[316,1333,134,1856]" captionTargetId="figure@4.[189,1397,133,1946]" captionTargetPageId="4" captionText="FIGURE 3. Female of M. ducalis from collection Régimbart in MNHN [TL = 47.4 mm], the largest known diving beetle specimen in the world: &quot;San Antônio da Barra, Prov. de Bahia, Brésil&quot;." httpUri="https://zenodo.org/record/2647356/files/figure.png" pageId="6" pageNumber="523">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C89CF0BBFFB5FFE4C1E06CC5FE464E18" pageId="6" pageNumber="523" type="distribution">
<paragraph id="8039A330FFB5FFE4C1E06CC5FC584FC0" blockId="6.[151,1437,151,771]" box="[199,953,706,731]" pageId="6" pageNumber="523">
<emphasis id="B2F27F22FFB5FFE4C1E06CC5FEB84FC0" bold="true" box="[199,345,706,731]" pageId="6" pageNumber="523">Distribution</emphasis>
. Only known from Condeúba, Bahia,
<collectingCountry id="F891E3A0FFB5FFE4C2216CC5FCAB4FC0" box="[774,842,706,731]" name="Brazil" pageId="6" pageNumber="523">Brazil</collectingCountry>
(
<figureCitation id="18BDBFB5FFB5FFE4C27E6CC5FC4D4FC0" box="[857,940,706,731]" captionStart="FIGURE 11" captionStartId="16.[151,249,1904,1926]" captionTargetBox="[151,1436,193,1883]" captionTargetId="figure@16.[151,1436,193,1883]" captionTargetPageId="16" captionText="FIGURE 11. Distribution of the subgenus Bifurcitus in Central and South America: White line: distribution of M. lherminieri; black line: distribution of M. magnus; red spot: only known locality for M. ducalis." httpUri="https://zenodo.org/record/2647372/files/figure.png" pageId="6" pageNumber="523">Fig. 11</figureCitation>
).
</paragraph>
<paragraph id="8039A330FFB5FFE4C1E06CEDFE464E18" blockId="6.[151,1437,151,771]" box="[199,423,746,771]" pageId="6" pageNumber="523">
<emphasis id="B2F27F22FFB5FFE4C1E06CEDFEC24E18" bold="true" box="[199,291,746,771]" pageId="6" pageNumber="523">Habitat</emphasis>
. Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>