166 lines
22 KiB
XML
166 lines
22 KiB
XML
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<mods:title id="18E47C731EC3F08076232C478F903958">To the knowledge of the subtribe Pericalina Hope, 1838 (Coleoptera, Carabidae, Lebiinae), with description of new genus, new subgenus and species from Oriental region</mods:title>
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<mods:namePart id="5510240DB16BF173A7AD402E3E83C60D">Anichtchenko, Alexander</mods:namePart>
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<mods:date id="AE56ACC31F63844E22E785ADF5C01493">2024</mods:date>
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<treatment id="03EFA270FFBCFFB4FF345F614E47E09A" ID-DOI="http://doi.org/10.5281/zenodo.11234058" ID-GBIF-Taxon="230983558" ID-Zenodo-Dep="11234058" LSID="urn:lsid:plazi:treatment:03EFA270FFBCFFB4FF345F614E47E09A" httpUri="http://treatment.plazi.org/id/03EFA270FFBCFFB4FF345F614E47E09A" lastPageNumber="370" pageId="1" pageNumber="370">
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<subSubSection id="C35C40EDFFBCFFB4FF345F614E2DE4C3" box="[151,380,870,897]" pageId="1" pageNumber="370" type="nomenclature">
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<paragraph id="8BF91366FFBCFFB4FF345F614E2DE4C3" blockId="1.[151,380,870,897]" box="[151,380,870,897]" pageId="1" pageNumber="370">
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<heading id="D0B1A40AFFBCFFB4FF345F614E2DE4C3" bold="true" box="[151,380,870,897]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
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<taxonomicName id="4C4668E5FFBCFFB4FF345F614E2DE4C3" authority="Anichtchenko, 2024" authorityName="Anichtchenko" authorityYear="2024" box="[151,380,870,897]" class="Insecta" family="Carabidae" genus="Acoptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B932CF74FFBCFFB4FF345F614E2DE4C3" bold="true" box="[151,380,870,897]" pageId="1" pageNumber="370">
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<taxonomicNameLabel id="A201720FFFBCFFB4FF345F614E2DE4C3" box="[151,380,870,897]" pageId="1" pageNumber="370" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FF345F614E73E4C3" bold="true" box="[151,290,870,897]" italics="true" pageId="1" pageNumber="370">Acoptodera</emphasis>
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gen. n.
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</taxonomicNameLabel>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C40EDFFBCFFB4FF345FAB4DEBE481" box="[151,698,939,964]" pageId="1" pageNumber="370" type="type_taxon">
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<paragraph id="8BF91366FFBCFFB4FF345FAB4DEBE481" blockId="1.[151,698,939,964]" box="[151,698,939,964]" pageId="1" pageNumber="370">
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<typeStatus id="54FDADC4FFBCFFB4FF345FAB4F98E486" box="[151,201,940,964]" pageId="1" pageNumber="370">Type</typeStatus>
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species:
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<taxonomicName id="4C4668E5FFBCFFB4FE855FAC4DE7E481" authority="Chaudoir, 1883: 20" authorityName="Chaudoir" authorityPageNumber="20" authorityYear="1883" box="[294,694,939,963]" class="Insecta" family="Carabidae" genus="Coptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="piligera">
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<emphasis id="B932CF74FFBCFFB4FE855FAC4EB8E481" box="[294,489,939,963]" italics="true" pageId="1" pageNumber="370">Coptodera piligera</emphasis>
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<bibRefCitation id="EFD76E97FFBCFFB4FE535FAC4DE7E481" author="Chaudoir, B. M." box="[496,694,939,963]" pageId="1" pageNumber="370" pagination="17 - 39" refId="ref4100" refString="Chaudoir, B. M. (1883) Description de carabiques nouveaux. In: Coleopterorum novitates. Recueil specialement consacre a l'etude des Coleopteres. Tom. I. Livr. I. Oberthur, Rennes, pp. 17 - 39." type="book chapter" year="1883">Chaudoir, 1883: 20</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C40EDFFBCFFB4FF345FF64D24E12D" pageId="1" pageNumber="370" type="diagnosis">
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<paragraph id="8BF91366FFBCFFB4FF345FF64D24E12D" blockId="1.[151,1437,1009,2008]" pageId="1" pageNumber="370">
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<emphasis id="B932CF74FFBCFFB4FF345FF64E43E349" bold="true" box="[151,274,1009,1035]" pageId="1" pageNumber="370">Diagnosis.</emphasis>
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The new genus is closely relative with five Oriental genera:
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<taxonomicName id="4C4668E5FFBCFFB4FC715FF64A7FE34E" authority="W. S. Macleay, 1825" authorityName="W. S. MacLeay" authorityYear="1825" box="[978,1326,1009,1036]" class="Insecta" family="Carabidae" genus="Pericalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FC715FF64B6CE349" box="[978,1085,1009,1035]" italics="true" pageId="1" pageNumber="370">Pericalus</emphasis>
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W.S. Macleay, 1825
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</taxonomicName>
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;
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<taxonomicName id="4C4668E5FFBCFFB4FA9F5FF54E1BE36D" authority="Chaudoir, 1869" authorityName="Chaudoir" authorityYear="1869" class="Insecta" family="Carabidae" genus="Lioptera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FA9F5FF54ACDE349" box="[1340,1436,1010,1035]" italics="true" pageId="1" pageNumber="370">Lioptera</emphasis>
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Chaudoir, 1869
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</taxonomicName>
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;
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<taxonomicName id="4C4668E5FFBCFFB4FEFB58124D20E372" authority="Dejean, 1825" authorityName="Dejean" authorityYear="1825" box="[344,625,1045,1072]" class="Insecta" family="Carabidae" genus="Coptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FEFB58124E81E36D" box="[344,464,1045,1071]" italics="true" pageId="1" pageNumber="370">Coptodera</emphasis>
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Dejean, 1825
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</taxonomicName>
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;
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<taxonomicName id="4C4668E5FFBCFFB4FDDC58124B56E372" authority="Louwerens, 1958" authorityName="Louwerens" authorityYear="1958" box="[639,1031,1045,1072]" class="Insecta" family="Carabidae" genus="Trichocoptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FDDC58124C6BE36D" box="[639,826,1045,1071]" italics="true" pageId="1" pageNumber="370">Trichocoptodera</emphasis>
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<bibRefCitation id="EFD76E97FFBCFFB4FCE058114B56E372" author="Louwerens, C. J." box="[835,1031,1045,1072]" pageId="1" pageNumber="370" pagination="251 - 261" refId="ref4188" refString="Louwerens, C. J. (1958) New Carabidae chiefly from Borneo and Celebes (Coleoptera). Treubia, 24 (2), 251 - 261." type="journal article" year="1958">Louwerens, 1958</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C4668E5FFBCFFB4FBE158124A1CE36D" authority="Andrewes, 1920" authorityName="Andrewes" authorityYear="1920" box="[1090,1357,1045,1071]" class="Insecta" family="Carabidae" genus="Gidda" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FBE158124BD9E36D" box="[1090,1160,1045,1071]" italics="true" pageId="1" pageNumber="370">Gidda</emphasis>
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Andrewes, 1920
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</taxonomicName>
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. These genera belong to the
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<taxonomicName id="4C4668E5FFBCFFB4FE23583E4EBAE311" authorityName="W. S. MacLeay" authorityYear="1825" box="[384,491,1081,1107]" class="Insecta" family="Carabidae" genus="Pericalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FE23583E4EBAE311" box="[384,491,1081,1107]" italics="true" pageId="1" pageNumber="370">Pericalus</emphasis>
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</taxonomicName>
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genus-group (
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<bibRefCitation id="EFD76E97FFBCFFB4FD2F583E4C2DE311" author="Shpeley, D. & Ball, G. E." box="[652,892,1081,1107]" pageId="1" pageNumber="370" pagination="1 - 185" refId="ref4219" refString="Shpeley, D. & Ball, G. E. (2000) A taxonomic review of the subtribe Pericalina (Coleoptera: Carabi dae: Lebiini) in the Western Hemisphere, with descriptions of new species and notes about classification and zoogeography. Insecta Mundi, 14 (1 / 3), 1 - 185." type="journal article" year="2000">Shpeley & Ball 2000</bibRefCitation>
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), or subtribe
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<taxonomicName id="4C4668E5FFBCFFB4FBAE583E4BC6E311" authorityName="Chaudoir" authorityYear="1848" box="[1037,1175,1081,1107]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="subTribe" subTribe="Coptoderina">Coptoderina</taxonomicName>
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in sense of
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<bibRefCitation id="EFD76E97FFBCFFB4FAB8583E4E4CE33A" author="Basilewsky, P." pageId="1" pageNumber="370" pagination="525 - 559" refId="ref4061" refString="Basilewsky, P. (1984) Essai d'une classification supra-generique naturelle des Carabides Lebiens d'Afrique et de Madagascar (Coleoptera Carabidae Lebiinae). Revue de Zoologie Africaine, 98 (3), 525 - 559." type="journal article" year="1984">Basilewsky (1984: 542)</bibRefCitation>
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in having pectinate claws, the mentum without a median tooth, palpifer with one long seta, and the terminal labial palpomere fusiform. Of all the genera listed, in terms of presence of hairs on the head, pronotum and elytra, it is most similar to
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<emphasis id="B932CF74FFBCFFB4FDB558A24D88E3FD" box="[534,729,1189,1215]" italics="true" pageId="1" pageNumber="370">
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<taxonomicName id="4C4668E5FFBCFFB4FDB558A24D85E3FD" authorityName="Louwerens" authorityYear="1958" box="[534,724,1189,1215]" class="Insecta" family="Carabidae" genus="Trichocoptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">Trichocoptodera</taxonomicName>
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.
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</emphasis>
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These two genera can be easily distinguished by following features: in
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<taxonomicName id="4C4668E5FFBCFFB4FEB958CE4E84E3A1" authorityName="Louwerens" authorityYear="1958" box="[282,469,1225,1251]" class="Insecta" family="Carabidae" genus="Trichocoptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
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<emphasis id="B932CF74FFBCFFB4FEB958CE4E84E3A1" box="[282,469,1225,1251]" italics="true" pageId="1" pageNumber="370">Trichocoptodera</emphasis>
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</taxonomicName>
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antennae short, with only two terminal antennomeres surpassing base of elytra; labrum transverse, with widely rounded anterior angles and apex without incision (
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<figureCitation id="137D0FE3FFBCFFB4FC4E58EA4B11E245" box="[1005,1088,1261,1287]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[159,1428,1214,1711]" captionTargetId="figure-50@6.[151,1436,1207,1719]" captionTargetPageId="6" captionText="FIGURES 6–8. Labrum (a) and gonocoxite (b). (6) Trichocoptodera maculata Louwerens, 1958; (7) Acoptodera (s. str.) piligera (Chaudoir, 1883); (8) Acoptodera (Pericalocephala) paradoxa subgen. n., sp. n." figureDoi="http://doi.org/10.5281/zenodo.11234264" httpUri="https://zenodo.org/record/11234264/files/figure.png" pageId="1" pageNumber="370">Fig. 6a</figureCitation>
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); elytra and pronotum convex; lateral expansion of pronotum narrow in anterior half; anterior edge of pronotum almost straight, posterior margin protrudes backward; elytra elongate with subparallel sides; all known species (including those that will be described in the next article) have a characteristic color, ventral side yellow or light brown, the pronotum is light with a more or less wide dark longitudinal stripe in the middle, head and elytra dark brown with copper lustre and elytra with developed yellow pattern, the lateral edges of the elytra are always contrastingly light (
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<figureCitation id="137D0FE3FFBCFFB4FB2159A64B97E2F9" box="[1154,1222,1441,1467]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[275,1313,184,1911]" captionTargetId="figure-20@2.[269,1319,179,1917]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Trichocoptodera maculata Louwerens, 1958, Paratype." figureDoi="http://doi.org/10.5281/zenodo.11234252" httpUri="https://zenodo.org/record/11234252/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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); gonocoxite (
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<figureCitation id="137D0FE3FFBCFFB4FACC59A64FE7E29D" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[159,1428,1214,1711]" captionTargetId="figure-50@6.[151,1436,1207,1719]" captionTargetPageId="6" captionText="FIGURES 6–8. Labrum (a) and gonocoxite (b). (6) Trichocoptodera maculata Louwerens, 1958; (7) Acoptodera (s. str.) piligera (Chaudoir, 1883); (8) Acoptodera (Pericalocephala) paradoxa subgen. n., sp. n." figureDoi="http://doi.org/10.5281/zenodo.11234264" httpUri="https://zenodo.org/record/11234264/files/figure.png" pageId="1" pageNumber="370">Fig. 6b</figureCitation>
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); while in
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<taxonomicName id="4C4668E5FFBCFFB4FE8E59C24EE1E29D" authority="Anichtchenko, 2024" authorityName="Anichtchenko" authorityYear="2024" box="[301,432,1477,1503]" class="Insecta" family="Carabidae" genus="Acoptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B932CF74FFBCFFB4FE8E59C24EE1E29D" box="[301,432,1477,1503]" italics="true" pageId="1" pageNumber="370">Acoptodera</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B932CF74FFBCFFB4FE1559C24D54E29D" bold="true" box="[438,517,1477,1503]" pageId="1" pageNumber="370">
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||
<taxonomicNameLabel id="A201720FFFBCFFB4FE1559C24D54E29D" box="[438,517,1477,1503]" pageId="1" pageNumber="370" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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antennae long, with four terminal antennomeres surpassing base of elytra; labrum longer, with deeply incised apex (subgenus s. str.) or long, trapezoid (
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<taxonomicName id="4C4668E5FFBCFFB4FC1559EE4B3EE141" authority="Anichtchenko, 2024" authorityName="Anichtchenko" authorityYear="2024" box="[950,1135,1513,1539]" kingdom="Animalia" pageId="1" pageNumber="370" rank="subGenus" status="subgen. nov." subGenus="Pericalocephala">
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<emphasis id="B932CF74FFBCFFB4FC1559EE4B3EE141" box="[950,1135,1513,1539]" italics="true" pageId="1" pageNumber="370">Pericalocephala</emphasis>
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</taxonomicName>
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<emphasis id="B932CF74FFBCFFB4FBDB59EE4BA3E141" bold="true" box="[1144,1266,1513,1539]" pageId="1" pageNumber="370">
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||
<taxonomicNameLabel id="A201720FFFBCFFB4FBDB59EE4BA3E141" box="[1144,1266,1513,1539]" pageId="1" pageNumber="370" rank="subGenus">subgen. n.</taxonomicNameLabel>
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</emphasis>
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) (
|
||
<figureCitation id="137D0FE3FFBCFFB4FAAF59EE4ADFE141" box="[1292,1422,1513,1540]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[159,1428,1214,1711]" captionTargetId="figure-50@6.[151,1436,1207,1719]" captionTargetPageId="6" captionText="FIGURES 6–8. Labrum (a) and gonocoxite (b). (6) Trichocoptodera maculata Louwerens, 1958; (7) Acoptodera (s. str.) piligera (Chaudoir, 1883); (8) Acoptodera (Pericalocephala) paradoxa subgen. n., sp. n." figureDoi="http://doi.org/10.5281/zenodo.11234264" httpUri="https://zenodo.org/record/11234264/files/figure.png" pageId="1" pageNumber="370">Figs 7a, 8a</figureCitation>
|
||
); elytra and pronotum flattened; elytra wide with rounded sides; lateral expansion of pronotum wide; anterior edge of pronotum with prominent anterior angles, posterior margin straight; the main body color is black, elytra with red spots (
|
||
<figureCitation id="137D0FE3FFBCFFB4FF435A524E17E12D" box="[224,326,1621,1647]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,1943,1967]" captionStartId-1="4.[151,250,1943,1967]" captionTargetBox-0="[266,1321,188,1912]" captionTargetBox-1="[225,1342,181,1912]" captionTargetId-0="figure-15@3.[259,1328,181,1919]" captionTargetId-1="figure-20@4.[219,1369,181,1920]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Habitus of Acoptodera (s. str.) piligera (Chaudoir, 1883)." captionText-1="FIGURE 3. Habitus of Acoptodera (Pericalocephala) paradoxa subgen. n., sp. n. Holotype." figureDoi-0="http://doi.org/10.5281/zenodo.11234254" figureDoi-1="http://doi.org/10.5281/zenodo.11234258" httpUri-0="https://zenodo.org/record/11234254/files/figure.png" httpUri-1="https://zenodo.org/record/11234258/files/figure.png" pageId="1" pageNumber="370">Figs 2–3</figureCitation>
|
||
); gonocoxite (
|
||
<figureCitation id="137D0FE3FFBCFFB4FE455A524D39E12D" box="[486,616,1621,1647]" captionStart="FIGURES 6–8" captionStartId="6.[151,264,1742,1766]" captionTargetBox="[159,1428,1214,1711]" captionTargetId="figure-50@6.[151,1436,1207,1719]" captionTargetPageId="6" captionText="FIGURES 6–8. Labrum (a) and gonocoxite (b). (6) Trichocoptodera maculata Louwerens, 1958; (7) Acoptodera (s. str.) piligera (Chaudoir, 1883); (8) Acoptodera (Pericalocephala) paradoxa subgen. n., sp. n." figureDoi="http://doi.org/10.5281/zenodo.11234264" httpUri="https://zenodo.org/record/11234264/files/figure.png" pageId="1" pageNumber="370">Figs 7b, 8b</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C40EDFFBCFFB4FF645A7E4E00E0CD" pageId="1" pageNumber="370" type="description">
|
||
<paragraph id="8BF91366FFBCFFB4FF645A7E4E00E0CD" blockId="1.[151,1437,1009,2008]" pageId="1" pageNumber="370">
|
||
<emphasis id="B932CF74FFBCFFB4FF645A7E4E08E1D1" bold="true" box="[199,345,1657,1683]" pageId="1" pageNumber="370">Description.</emphasis>
|
||
Body black with yellow elytral pattern. Head, pronotum and elytra without microrreticulation. Pronotum and elytra flattened. Elytra wide and rounded. Lateral edge of elytra minutely or strongly serrate, with fringe of hairs. Head, pronotum and elytra covered with more or less long hairs. Antennae long, with four terminal antennomeres surpassing base of elytra. Labrum with 6 apical setae. Scape (first antennomere) with 4–10 additional setae. Anterior angles of pronotum slightly protruding forwards; lateral margins of pronotum wide, flattened and explanate. Elytra with 2 pores in third interval of elytra (basal and apical), discal pores missing; sutural stria long, not connected with first stria; scutellar pore situated near the base of first stria; umbilical series consist of 15 pores. Claws pectinate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C40EDFFBCFFB4FF645B9E4E47E09A" pageId="1" pageNumber="370" type="etymology">
|
||
<paragraph id="8BF91366FFBCFFB4FF645B9E4E47E09A" blockId="1.[151,1437,1009,2008]" pageId="1" pageNumber="370">
|
||
<emphasis id="B932CF74FFBCFFB4FF645B9E4E1FE0F1" bold="true" box="[199,334,1945,1971]" pageId="1" pageNumber="370">Etymology.</emphasis>
|
||
The prefix
|
||
<emphasis id="B932CF74FFBCFFB4FE495B9D4EA9E0F1" box="[490,504,1946,1971]" italics="true" pageId="1" pageNumber="370">a</emphasis>
|
||
- in Latin means “of, off, from, away” indicating a relationship with genus
|
||
<taxonomicName id="4C4668E5FFBCFFB4FF345BBA4E5EE095" authorityName="Dejean" authorityYear="1825" box="[151,271,1981,2007]" class="Insecta" family="Carabidae" genus="Coptodera" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B932CF74FFBCFFB4FF345BBA4E5EE095" box="[151,271,1981,2007]" italics="true" pageId="1" pageNumber="370">Coptodera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |