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<mods:title id="A42E318EE6468F21474BDF8C7B0BEDF9">Redescription of the damsel-dragonfly Parafleckium senjituense on the basis of a more complete specimen (Odonata: Isophlebioptera: Campterophlebiidae)</mods:title>
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<mods:namePart id="8479A10F4C9EB2B31909B34966126A68">Li, Yongjun</mods:namePart>
<mods:affiliation id="71E1F3877A66142235571E02EC2BDCDF">College of Life Science, Capital Normal University, China &amp; Key Laboratory of Biodiversity Dynamics and Conservation of Guangdong Higher Education Institute, Sun Yat-Sen University, Guangzhou, China</mods:affiliation>
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<mods:namePart id="7966192B899BC3506EC16F4A4C6EDAC3">Nel, André</mods:namePart>
<mods:affiliation id="25B9BF1898FAA0CADF05BD86389DFBD3">CNRS UMR 7205, CP 50, Entomologie, Muséum National d'Histoire Naturelle, 45 rue Buffon, F- 75005 Paris, France Corresponding authors. E-mails: rendong @ mail. cnu. edu. cn or lsshpang @ mail. sysu. edu. cn</mods:affiliation>
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<mods:namePart id="F86AFD8C5F662BF681CD860E81BAAD8F">Ren, Dong</mods:namePart>
<mods:affiliation id="6514363D553A1C4B6DC26CE7620CB08F">College of Life Science, Capital Normal University, China</mods:affiliation>
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<mods:namePart id="85AD4E0A8E00270A2ED9A1FB7E0A757E">Pang, Hong</mods:namePart>
<mods:affiliation id="969CE67D31FA7EF412F8741A74C35850">Key Laboratory of Biodiversity Dynamics and Conservation of Guangdong Higher Education Institute, Sun Yat-Sen University, Guangzhou, China</mods:affiliation>
<mods:nameIdentifier id="892AD35C4869D1E76161C51E943E5C24" type="email">rendong@mail.cnu.edu.cn</mods:nameIdentifier>
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<subSubSection id="C37E653FFFA4FF8ED1AD6835FD91A231" box="[151,588,1861,1885]" pageId="0" pageNumber="53" type="nomenclature">
<paragraph id="8BDB36B4FFA4FF8ED1AD6835FD91A231" blockId="0.[151,588,1861,1919]" box="[151,588,1861,1885]" pageId="0" pageNumber="53">
<heading id="D09381D8FFA4FF8ED1AD6835FD91A231" bold="true" box="[151,588,1861,1885]" fontSize="10" level="2" pageId="0" pageNumber="53" reason="2">
<taxonomicName id="4C644D37FFA4FF8ED1AD6835FD91A231" authority="Li et al., 2012" authorityName="Li" authorityYear="2012" box="[151,588,1861,1885]" class="Insecta" family="Campterophlebiidae" genus="Parafleckium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="53" phylum="Arthropoda" rank="species" species="senjituense">
<emphasis id="B910EAA6FFA4FF8ED1AD6835FD91A231" bold="true" box="[151,588,1861,1885]" pageId="0" pageNumber="53">
<emphasis id="B910EAA6FFA4FF8ED1AD6835FE73A231" bold="true" box="[151,430,1861,1885]" italics="true" pageId="0" pageNumber="53">Parafleckium senjituense</emphasis>
<bibRefCitation id="EFF54B45FFA4FF8ED08E6835FD91A231" author="Li, Y. - J. &amp; Nel, A. &amp; Ren, D. &amp; Pang, H." box="[436,588,1861,1885]" pageId="0" pageNumber="53" pagination="340 - 343" refId="ref1865" refString="Li, Y. - J., Nel, A., Ren, D. &amp; Pang, H. (2012) A new damsel-dragonfly from the Lower Cretaceous of China enlightens the systematics of the Isophlebioidea (Odonata: Isophlebioptera: Campterophlebiidae). Cretaceous Research, 34, 340 - 343." type="journal article" year="2012">
Li
<emphasis id="B910EAA6FFA4FF8ED0EE6835FDDFA231" bold="true" box="[468,514,1861,1885]" italics="true" pageId="0" pageNumber="53">et al</emphasis>
., 2012
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</emphasis>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E653FFFA4FF8ED1AD6818FF3BA213" box="[151,230,1896,1919]" pageId="0" pageNumber="53" type="description">
<paragraph id="8BDB36B4FFA4FF8ED1AD6818FF3BA213" blockId="0.[151,588,1861,1919]" box="[151,230,1896,1919]" pageId="0" pageNumber="53">
(
<figureCitation id="135F2A31FFA4FF8ED1A56818FF00A213" box="[159,221,1896,1919]" captionStart="FIGURE 1" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[151,1436,195,1911]" captionTargetId="figure-74@2.[151,1436,195,1923]" captionTargetPageId="2" captionText="FIGURE 1. Parafleckium senjituense Li et al., 2012. A, Photo of specimen CNU-ODO-ND2011018, part only. B, Drawing of body. C, Drawing of forewing. D, Drawing of hindwing. E. Detail photo and drawing of hindwing base. F, Photo and drawing of the isolated forewing. Scale bars represent 10 mm in A to D and F, 5 mm in E." pageId="0" pageNumber="53">Fig. 1</figureCitation>
)
</paragraph>
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<subSubSection id="C37E653FFFA4FF8FD1AD68DDFDC2A41B" lastPageId="1" lastPageNumber="54" pageId="0" pageNumber="53" type="diagnosis">
<paragraph id="8BDB36B4FFA4FF8FD1AD68DDFB43A458" blockId="0.[151,1436,1965,2022]" lastBlockId="1.[151,1437,152,1275]" lastPageId="1" lastPageNumber="54" pageId="0" pageNumber="53">
<emphasis id="B910EAA6FFA4FF8ED1AD68DDFEA2A2A9" bold="true" box="[151,383,1965,1989]" inLineHeading="true" pageId="0" pageNumber="53" reason="1">Emended diagnosis.</emphasis>
The original diagnosis was established on forewing alone. Here we add some characters. In forewing: distal area between RA and RP1 with two rows of cells and slightly widened; many secondary intercalary veins originating from MP. In male hindwing: proximal part of area between MP and CuA more than three times as wide as proximal part of postdiscoidal area between MA and MP; cubital area very broad, with seven rows of cells between CuAa and posterior wing margin; CuAa with three main posterior branches; CuAb long and straight, not distally reaching AA, correlated with a very long and posteriorly opened subdiscoidal cell subdivided into several small cells by transverse crossveins; anal area triangular in shape, rather broad; male anal angle not very sharp.
</paragraph>
<paragraph id="8BDB36B4FFA5FF8FD1FD6E4EFDC2A41B" blockId="1.[151,1437,152,1275]" pageId="1" pageNumber="54">
<emphasis id="B910EAA6FFA5FF8FD1FD6E4EFE1DA43A" bold="true" box="[199,448,318,342]" inLineHeading="true" pageId="1" pageNumber="54" reason="1">Locality and horizon.</emphasis>
Baitugou, Nanyingpan Village, Sanbeigou Town, Duolun County,
<collectingRegion id="49A0F856FFA5FF8FD5A66E4EFA95A439" box="[1180,1352,318,341]" country="China" name="Nei Mongol" pageId="1" pageNumber="54">Inner Mongolia</collectingRegion>
,
<collectingCountry id="F3737624FFA5FF8FD46F6E4EFA45A439" box="[1365,1432,318,341]" name="China" pageId="1" pageNumber="54">China</collectingCountry>
. Yixian Formation, Early Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C37E653FFFA5FF8FD1FD6EF1FB5DA197" pageId="1" pageNumber="54" type="description">
<paragraph id="8BDB36B4FFA5FF8FD1FD6EF1FB5DA197" blockId="1.[151,1437,152,1275]" pageId="1" pageNumber="54">
<emphasis id="B910EAA6FFA5FF8FD1FD6EF1FE8FA4F5" bold="true" box="[199,338,385,409]" inLineHeading="true" pageId="1" pageNumber="54" reason="1">Description.</emphasis>
Specimen rather poorly preserved. On one slab, two hindwings and a forewing attached to the thorax (
<figureCitation id="135F2A31FFA5FF8FD1A56ED2FECDA4D5" box="[159,272,418,441]" captionStart="FIGURE 1" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[151,1436,195,1911]" captionTargetId="figure-74@2.[151,1436,195,1923]" captionTargetPageId="2" captionText="FIGURE 1. Parafleckium senjituense Li et al., 2012. A, Photo of specimen CNU-ODO-ND2011018, part only. B, Drawing of body. C, Drawing of forewing. D, Drawing of hindwing. E. Detail photo and drawing of hindwing base. F, Photo and drawing of the isolated forewing. Scale bars represent 10 mm in A to D and F, 5 mm in E." pageId="1" pageNumber="54">Fig. 1AE</figureCitation>
) and one isolated forewing some distance apart (
<figureCitation id="135F2A31FFA5FF8FD2246ED2FCB0A4D5" box="[798,877,418,441]" captionStart="FIGURE 1" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[151,1436,195,1911]" captionTargetId="figure-74@2.[151,1436,195,1923]" captionTargetPageId="2" captionText="FIGURE 1. Parafleckium senjituense Li et al., 2012. A, Photo of specimen CNU-ODO-ND2011018, part only. B, Drawing of body. C, Drawing of forewing. D, Drawing of hindwing. E. Detail photo and drawing of hindwing base. F, Photo and drawing of the isolated forewing. Scale bars represent 10 mm in A to D and F, 5 mm in E." pageId="1" pageNumber="54">Fig. 1F</figureCitation>
), they most probably belong to the same individual. Wing without trace of coloration, probably hyaline.
<emphasis id="B910EAA6FFA5FF8FD3F46EB3FCE7A4B7" bold="true" box="[718,826,451,475]" pageId="1" pageNumber="54">Forewing</emphasis>
(
<figureCitation id="135F2A31FFA5FF8FD2706EB4FC1BA4B7" box="[842,966,452,475]" captionStart="FIGURE 1" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[151,1436,195,1911]" captionTargetId="figure-74@2.[151,1436,195,1923]" captionTargetPageId="2" captionText="FIGURE 1. Parafleckium senjituense Li et al., 2012. A, Photo of specimen CNU-ODO-ND2011018, part only. B, Drawing of body. C, Drawing of forewing. D, Drawing of hindwing. E. Detail photo and drawing of hindwing base. F, Photo and drawing of the isolated forewing. Scale bars represent 10 mm in A to D and F, 5 mm in E." pageId="1" pageNumber="54">Figs. 1C, F</figureCitation>
) estimated wing length about
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long,
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wide at nodus level; distance between Ax1 and Ax2
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, between Ax1 and wing base 7.0 mm, between Ax2 and nodus 24.0 mm, between nodus and wing apex about 40.0 mm; Ax1
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basal of arculus; Ax1 and Ax2 slightly oblique; gaff elongated, basal area between MP and CuA (about
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wide) more than two times as broad as postdiscoidal area between MA and MP (
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wide); CuAa parallel with posterior wing margin for a short distance then vanishing into crossvein network, with one row of longitudinal cells below CuA; many secondary intercalary veins originating from MP; MP strongly undulating in mid part; MA slightly curved and zigzagged distally; postdiscoidal area basally with one row of cells, but 23 rows of cells at curvature of MP, distally with two rows of cells; RP3/4 strongly curved distally, distal area between it and MA strongly expanded; IR2 smoothly curved, more or less parallel with RP3/ 4; subnodus weakly oblique; RP2 not aligned but slightly basal to subnodus; preserved IR1 smoothly curved; distal area between RA and RP1 with two rows of cells and slightly widened.
<emphasis id="B910EAA6FFA5FF8FD2526C61FC05A645" bold="true" box="[872,984,785,809]" pageId="1" pageNumber="54">Hindwing</emphasis>
(
<figureCitation id="135F2A31FFA5FF8FD2D26C61FB80A644" box="[1000,1117,785,808]" captionStart="FIGURE 1" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[151,1436,195,1911]" captionTargetId="figure-74@2.[151,1436,195,1923]" captionTargetPageId="2" captionText="FIGURE 1. Parafleckium senjituense Li et al., 2012. A, Photo of specimen CNU-ODO-ND2011018, part only. B, Drawing of body. C, Drawing of forewing. D, Drawing of hindwing. E. Detail photo and drawing of hindwing base. F, Photo and drawing of the isolated forewing. Scale bars represent 10 mm in A to D and F, 5 mm in E." pageId="1" pageNumber="54">Figs. 1D,E</figureCitation>
) preserved wing length about 45.0 mm, widest area about
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; Ax1 not preserved; distance between Ax2 and wing base about
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; ScP very close to R; discoidal cell closed and simple; RP3/4 strongly curved; MA nearly straight; MP basally undulated; postdiscoidal area very narrow, with only one row of cells in preserved part; gaff very long and straight; proximal part of area between MP and CuA (about
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wide) more than three times as wide as proximal part of area between MA and MP (about
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wide); post-subdiscoidal area basally with four rows of cells; cubital area very broad, with seven rows of cell between CuAa and posterior wing margin; CuAa with three main posterior branches; CuAb long and straight, not distally reaching AA, correlated with a very long and posteriorly opened subdiscoidal cell; subdiscoidal cell divided into about nine cells; AA strongly bent towards posterior wing margin after CuP crossing; submedian space with at least one crossvein; anal triangle broad, with sixteen irregular cells; male anal angle not very sharp.
<emphasis id="B910EAA6FFA5FF8FD5A46B4CFB04A138" bold="true" box="[1182,1241,1084,1108]" pageId="1" pageNumber="54">Body</emphasis>
(
<figureCitation id="135F2A31FFA5FF8FD5D66B4DFA9FA138" box="[1260,1346,1085,1108]" captionStart="FIGURE 1" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[151,1436,195,1911]" captionTargetId="figure-74@2.[151,1436,195,1923]" captionTargetPageId="2" captionText="FIGURE 1. Parafleckium senjituense Li et al., 2012. A, Photo of specimen CNU-ODO-ND2011018, part only. B, Drawing of body. C, Drawing of forewing. D, Drawing of hindwing. E. Detail photo and drawing of hindwing base. F, Photo and drawing of the isolated forewing. Scale bars represent 10 mm in A to D and F, 5 mm in E." pageId="1" pageNumber="54">Fig. 1B</figureCitation>
) poorly preserved, lateral imprint; head, thorax, two segments of abdomen broad, with remnants of male secondary genital organs, and legs can only be discerned by outline; head rounded in shape,
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wide and
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long, with eyes well separated; length of thorax
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, of segment I
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and segment II about
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; foreleg: length of femur about
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, of tibia
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, of tarsus about
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; tarsi with two segments, equal in length; two claws; midleg: femur
<quantity id="4C9C9B51FFA5FF8FD4046BB2FA41A1B5" box="[1342,1436,1218,1241]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="1" pageNumber="54" unit="mm" value="14.5">14.5 mm</quantity>
long and tibia
<quantity id="4C9C9B51FFA5FF8FD00A6B94FE53A197" box="[304,398,1252,1275]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="1" pageNumber="54" unit="mm" value="10.1">10.1 mm</quantity>
long; hind leg with partly preserved femur, but coxa can be discerned.
</paragraph>
</subSubSection>
</treatment>
</document>