treatments-xml/data/03/91/7F/03917F7FFFF1FFDDA922FE1AFC5881D9.xml

<document id="8140B23D66B374F2EFECA6724F19DB05" ID-DOI="10.15298/rusentj.31.2.08" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179486" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722596157083" checkinUser="felipe" docAuthor="Gildenkov, M. Yu." docDate="2022" docId="03917F7FFFF1FFDDA922FE1AFC5881D9" docLanguage="en" docName="RussEntomolJ.31.2.140-143.pdf" docOrigin="Russian Entomological Journal 31 (2)" docSource="http://dx.doi.org/10.15298/rusentj.31.2.08" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Coprophilus maximus Gildenkov 2022, sp. n." docType="treatment" docVersion="2" lastPageNumber="143" masterDocId="FFA80707FFF0FFDEAA72FF95FF938065" masterDocTitle="New species of the genus Coprophilus Latreille, 1829 from North India (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="143" masterPageNumber="140" pageNumber="141" updateTime="1722637150028" updateUser="ExternalLinkService">
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<mods:title id="B3A4587E64BCE074E46FED9E1A74FBB7">New species of the genus Coprophilus Latreille, 1829 from North India (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:date id="DAD0118FC0E33E153A25E82D5F8EE81A">2022</mods:date>
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<taxonomicName id="4C38B5EAFFF1FFDFA922FE1AFA9881C2" ID-CoL="9GK74" authority="Gildenkov" authorityName="Gildenkov" authorityYear="2022" box="[848,1291,399,423]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="141" phylum="Arthropoda" rank="species" species="maximus" status="sp. nov.">
<emphasis id="B94C127BFFF1FFDFA922FE1AFC4681C2" box="[848,981,399,423]" italics="true" pageId="1" pageNumber="141">Coprophilus</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF1FFDFAE42FE05FB1C81C2" box="[1072,1167,400,423]" italics="true" pageId="1" pageNumber="141">maximus</emphasis>
Gildenkov
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, 
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<paragraph id="8B87CE69FFF1FFDFAE5DFE39FBE781A1" blockId="1.[812,1399,399,2053]" box="[1071,1140,428,452]" pageId="1" pageNumber="141">
<figureCitation id="1303D2ECFFF1FFDFAE5DFE39FBFD81A1" box="[1071,1134,428,452]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
.
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<paragraph id="8B87CE69FFF1FFDFA920FE44FAA78271" blockId="1.[812,1399,399,2053]" pageId="1" pageNumber="141">
<materialsCitation id="3B50C434FFF1FFDFA920FE44FAA78271" collectingDate="2003-06-27" collectingDateMax="2003-06-30" collectingDateMin="2003-06-27" country="India" county="Khati" elevation="2200" location="North India" municipality="Uttaranchal" pageId="1" pageNumber="141" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
MATERIAL. 
<typeStatus id="548370CBFFF1FFDFA9B9FE44FB8E8181" box="[971,1053,465,484]" pageId="1" pageNumber="141" type="holotype">Holotype</typeStatus>
, 
<specimenCount id="9D3E05E0FFF1FFDFAE57FE5AFBA28180" box="[1061,1073,463,485]" count="1" pageId="1" pageNumber="141" type="female">♀</specimenCount>
, 
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, 
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: with labels “N 
<collectingCountry id="F32F8EF9FFF1FFDFA93EFE7CFC1F8199" box="[844,908,489,508]" name="India" pageId="1" pageNumber="141">INDIA</collectingCountry>
: 
<collectingMunicipality id="6BE35413FFF1FFDFA9E7FE7CFC6A8199" box="[917,1017,489,508]" pageId="1" pageNumber="141">Uttaranchal</collectingMunicipality>
state, 
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<quantity id="4CC0638CFFF1FFDFAE41FE7CFBF98199" box="[1075,1130,489,508]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="141" unit="km" value="30.0">30 km</quantity>
N Bageshwar
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, 
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vill env., 
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, 
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, Kejval &amp; Tryzna” (cMSch).
</materialsCitation>
</paragraph>
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DESCRIPTION (
<typeStatus id="548370CBFFF1FFDFA988FD8CFBC7824A" box="[1018,1108,537,559]" pageId="1" pageNumber="141" type="holotype">holotype</typeStatus>
). Length 
<quantity id="4CC0638CFFF1FFDFAEDEFD8CFB64824A" box="[1196,1271,537,559]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="141" unit="mm" value="8.8">8.8 mm</quantity>
. Body slightly convex, the integument integument slightly shining. Colouration brown, with a reddish tint; antennae black-brown (
<figureCitation id="1303D2ECFFF1FFDFA941FDFCFCF9821B" box="[819,874,617,639]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Head slightly transverse, widest across eyes. Ratio of its length (from base to anterior margin of clypeus) to maximum width (mm) about 1.07:1.39. Eyes moderately large, slightly convex (
<figureCitation id="1303D2ECFFF1FFDFA9A2FD2CFB9A82AB" box="[976,1033,697,719]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Forehead finely and sparsely punctate. (
<figureCitation id="1303D2ECFFF1FFDFA911FD41FC08828C" box="[867,923,724,746]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
); vertex with noticeably larger and denser punctation (
<figureCitation id="1303D2ECFFF1FFDFA900FD7AFC3F8361" box="[882,940,751,773]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Puncture diameter on the vertex is about 1.5 times more than diameter of the eye facet and diameter of points on the forehead, distances between punctures noticeably more than their diameter, interspaces smooth, slightly shining (
<figureCitation id="1303D2ECFFF1FFDFA9F0FCCCFC2F830B" box="[898,956,857,879]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Neck large, coarsely and densely punctate. Puncture diameter is about 3 times more than diameter of eye facet, distances between punctures noticeably smaller than their diameter, interspaces smooth, slightly shining. Antennae rather long (
<figureCitation id="1303D2ECFFF1FFDFA9BFFC51FB9583BC" box="[973,1030,964,986]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Antennal segment 1 elongate, conical, more than twice longer than its maximum width; segment 2 elongate, conical, about twice as long as its maximum broad, much shorter and narrower than segment 1; segment 3 conical, more than twice longer than its maximum broad; segments 4–6 cylindrical, slightly broaded to apex, only slightly longer than their maximum broad; segments 7–10 conical, much more massive than segments 4–6, slightly transverse; segment 11 pointed apically, about 1.5 times as long as broad. Last 5 segments form loose club (
<figureCitation id="1303D2ECFFF1FFDFAF7AFB21FAD284AC" box="[1288,1345,1204,1226]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B87CE69FFF1FFDFA920FB5AFB4F86BA" blockId="1.[812,1399,399,2053]" pageId="1" pageNumber="141">
Pronotum (
<figureCitation id="1303D2ECFFF1FFDFA9CCFB5AFC648481" box="[958,1015,1231,1253]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
) slightly convex, maximum broad after about 2/3 of length, measured from the base. Ratio of pronotum length to its maximum broad (mm) about 1.63:1.60. Front angles rounded, lateral margin serrated (
<figureCitation id="1303D2ECFFF1FFDFAE9AFA8AFAB38551" box="[1256,1312,1311,1333]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Pronotal disc with deep depressions: 2 paired oval depressions at base, on either side of midline; unpaired broad rhomboid depression at disc centre and 2 broad, deep, triangular depressions near lateral margins (
<figureCitation id="1303D2ECFFF1FFDFAE43FA1CFBFA85FB" box="[1073,1129,1417,1439]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Surface of pronotum with delicate, fine and sparsely punctation. Punctation irregular, the diameter of most punctures approximately 1.5 times more than diameter of eye facet, some punctures approximately equal to the diameter of eye facet; the distances between punctures are much longer than their diameter, interspaces smooth, slightly shining (
<figureCitation id="1303D2ECFFF1FFDFAE5FF9BCFBFA865B" box="[1069,1129,1577,1599]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Punctures at the base of pronotum and along the lateral margins are larger and denser; their diameter about 3 times more than diameter of eye facet, distance between punctures significantly smaller than their diameter. Punctation most dense at basal angles and at bottom of depressions near lateral margins, interspaces between these punctures much smaller than their diameter.
</paragraph>
<paragraph id="8B87CE69FFF1FFDFA920F971FB1487AA" blockId="1.[812,1399,399,2053]" pageId="1" pageNumber="141">
Elytra convex, ratio of length of elytra to their combined width (mm) about 2.20:2.46. Six rather shallow grooves are clearly visible on the surface of each elytron (
<figureCitation id="1303D2ECFFF1FFDFAE93F88CFA8B874B" box="[1249,1304,1817,1839]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Surface of elytra delicate, finely and sparsely punctate. Punctation irregular, diameter of most punctures approximately equal to diameter of eye facet, some punctures are 1.5 times more than diameter of eye facet, distances between punctures are much greater than their diameter, interspaces smooth, slightly shining (
<figureCitation id="1303D2ECFFF1FFDFA92BF82CFC0187AB" box="[857,914,1977,1999]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
). Elytra finely bordered.
</paragraph>
<paragraph id="8B87CE69FFF1FFDFA920F841FBC78861" blockId="1.[812,1399,399,2053]" pageId="1" pageNumber="141">
Abdomen with delicate, fine and sparsely punctation and delicately shagreened (
<figureCitation id="1303D2ECFFF1FFDFAE7EF87AFBD58861" box="[1036,1094,2031,2053]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="1" pageNumber="141">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF479EE1FFF2FFDCAA90F937FEE58763" ID-DOI="http://doi.org/10.5281/zenodo.13179488" ID-Zenodo-Dep="13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="2" pageNumber="142" startId="2.[226,263,1698,1722]" targetBox="[191,1938,608,1672]" targetPageId="2" targetType="figure">
<paragraph id="8B87CE69FFF2FFDCAA90F937FEEA86B7" blockId="2.[188,1944,1697,1798]" pageId="2" pageNumber="142">
Figs 1–4. 
<taxonomicName id="4C38B5EAFFF2FFDCAB45F937FE5D86DF" box="[311,462,1698,1722]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94C127BFFF2FFDCAB45F937FE3086DF" box="[311,419,1698,1722]" italics="true" pageId="2" pageNumber="142">Coprophilus</emphasis>
spp.
</taxonomicName>
, dorsal view: 1 — 
<taxonomicName id="4C38B5EAFFF2FFDCA81CF937FC8786DF" authorityName="Gildenkov" authorityYear="2022" box="[622,788,1698,1722]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="maximus" status="sp.n.">
<emphasis id="B94C127BFFF2FFDCA81CF937FD1186DF" box="[622,642,1698,1722]" italics="true" pageId="2" pageNumber="142">C.</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF2FFDCA8BAF937FC8786DF" box="[712,788,1698,1722]" italics="true" pageId="2" pageNumber="142">maximus</emphasis>
</taxonomicName>
<emphasis id="B94C127BFFF2FFDCA96BF937FCD786DF" bold="true" box="[793,836,1698,1722]" pageId="2" pageNumber="142">
<taxonomicNameLabel id="A27FAF00FFF2FFDCA96BF937FCD786DF" box="[793,836,1698,1722]" pageId="2" pageNumber="142" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, holotype, ♀; 2–3 — 
<taxonomicName id="4C38B5EAFFF2FFDCA989F937FB2986DF" authorityName="Y. Shibata" authorityYear="1993" box="[1019,1210,1698,1722]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B94C127BFFF2FFDCA989F937FB8386DF" box="[1019,1040,1698,1722]" italics="true" pageId="2" pageNumber="142">C.</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF2FFDCAE24F937FB2986DF" box="[1110,1210,1698,1722]" italics="true" pageId="2" pageNumber="142">formosanus</emphasis>
</taxonomicName>
(2 — paratype, ♀; 3 — holotype, ♂); 4 — 
<taxonomicName id="4C38B5EAFFF2FFDCAC5EF937F94486DF" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[1580,1751,1698,1722]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis id="B94C127BFFF2FFDCAC5EF937F9D286DF" box="[1580,1601,1698,1722]" italics="true" pageId="2" pageNumber="142">C.</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF2FFDCACF4F937F94486DF" box="[1670,1751,1698,1722]" italics="true" pageId="2" pageNumber="142">striatulus</emphasis>
</taxonomicName>
, ♀, Russia, Smolensk. Scale bar — 1.0 mm.
</paragraph>
<paragraph id="8B87CE69FFF2FFDCAA90F943FEE58763" blockId="2.[188,1944,1697,1798]" pageId="2" pageNumber="142">
Рис. 1–4. 
<taxonomicName id="4C38B5EAFFF2FFDCAB48F943FE42868B" box="[314,465,1750,1774]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94C127BFFF2FFDCAB48F943FE35868B" box="[314,422,1750,1774]" italics="true" pageId="2" pageNumber="142">Coprophilus</emphasis>
spp.
</taxonomicName>
, сверху: 1 — 
<taxonomicName id="4C38B5EAFFF2FFDCA83CF943FD65868B" authorityName="Gildenkov" authorityYear="2022" box="[590,758,1750,1774]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="maximus" status="sp.n.">
<emphasis id="B94C127BFFF2FFDCA83CF943FDF0868B" box="[590,611,1750,1774]" italics="true" pageId="2" pageNumber="142">C.</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF2FFDCA8D8F943FD65868B" box="[682,758,1750,1774]" italics="true" pageId="2" pageNumber="142">maximus</emphasis>
</taxonomicName>
<emphasis id="B94C127BFFF2FFDCA88EF943FCB4868B" bold="true" box="[764,807,1750,1774]" pageId="2" pageNumber="142">
<taxonomicNameLabel id="A27FAF00FFF2FFDCA88EF943FCB4868B" box="[764,807,1750,1774]" pageId="2" pageNumber="142" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, голотип, ♀; 2–3 — 
<taxonomicName id="4C38B5EAFFF2FFDCA992F943FB33868B" authorityName="Y. Shibata" authorityYear="1993" box="[992,1184,1750,1774]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B94C127BFFF2FFDCA992F943FC67868B" box="[992,1012,1750,1774]" italics="true" pageId="2" pageNumber="142">C.</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF2FFDCAE4EF943FB33868B" box="[1084,1184,1750,1774]" italics="true" pageId="2" pageNumber="142">formosanus</emphasis>
</taxonomicName>
(2 — паратип, ♀, 3 — голотип, ♂); 4 — 
<taxonomicName id="4C38B5EAFFF2FFDCAC6EF943F95A868B" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[1564,1737,1750,1774]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="142" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis id="B94C127BFFF2FFDCAC6EF943F9A2868B" box="[1564,1585,1750,1774]" italics="true" pageId="2" pageNumber="142">C.</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF2FFDCAC0AF943F95A868B" box="[1656,1737,1750,1774]" italics="true" pageId="2" pageNumber="142">striatulus</emphasis>
</taxonomicName>
, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм.
</paragraph>
</caption>
<paragraph id="8B87CE69FFF3FFDDAA90FEAEFE188134" blockId="3.[189,775,315,1137]" box="[226,395,315,337]" pageId="3" pageNumber="143">
<emphasis id="B94C127BFFF3FFDDAA90FEAEFE858135" bold="true" box="[226,278,315,336]" pageId="3" pageNumber="143">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B87CE69FFF3FFDDAA90FEC3FEF08445" blockId="3.[189,775,315,1137]" pageId="3" pageNumber="143">
COMPARATIVE NOTES. The new species is most similar in color and general habit to 
<taxonomicName id="4C38B5EAFFF3FFDDAB97FEE4FD2781E3" authorityName="Y. Shibata" authorityYear="1993" box="[485,692,369,390]" class="Insecta" family="Staphylinidae" genus="Coprophilus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B94C127BFFF3FFDDAB97FEE4FE6881E3" box="[485,507,369,390]" italics="true" pageId="3" pageNumber="143">C.</emphasis>
(s. str.) 
<emphasis id="B94C127BFFF3FFDDA834FEE4FD2781E3" box="[582,692,369,390]" italics="true" pageId="3" pageNumber="143">formosanus</emphasis>
</taxonomicName>
(
<figureCitation id="1303D2ECFFF3FFDDA8B2FEE4FF5981C5" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="3" pageNumber="143">Figs 1– 3</figureCitation>
). Well differs from it by significantly larger body sizes. Basic dimensions 
<taxonomicName id="4C38B5EAFFF3FFDDAB5FFE33FE0F81D9" authorityName="Gildenkov" authorityYear="2022" box="[301,412,422,444]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="maximus">
<emphasis id="B94C127BFFF3FFDDAB5FFE33FEAE81DE" box="[301,317,422,443]" italics="true" pageId="3" pageNumber="143">C</emphasis>
. 
<emphasis id="B94C127BFFF3FFDDAB3AFE32FE0F81D9" box="[328,412,423,444]" italics="true" pageId="3" pageNumber="143">maximus</emphasis>
</taxonomicName>
: body length 
<quantity id="4CC0638CFFF3FFDDA86EFE33FDF581DE" box="[540,614,422,443]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="3" pageNumber="143" unit="mm" value="8.8">8.8 mm</quantity>
; length of pronotum 
<quantity id="4CC0638CFFF3FFDDAA9AFE55FED381B3" box="[232,320,448,470]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="3" pageNumber="143" unit="mm" value="1.63">1.63 mm</quantity>
; maximum width of pronotum 
<quantity id="4CC0638CFFF3FFDDA818FE55FD5281B3" box="[618,705,448,470]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="143" unit="mm" value="1.6">1.60 mm</quantity>
, length of elytra 
<quantity id="4CC0638CFFF3FFDDAB61FE4EFEFF8194" box="[275,364,475,497]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="143" unit="mm" value="2.2">2.20 mm</quantity>
; maximum width of elytra 
<quantity id="4CC0638CFFF3FFDDA802FE4EFD5B8194" box="[624,712,475,497]" metricMagnitude="-3" metricUnit="m" metricValue="2.46" pageId="3" pageNumber="143" unit="mm" value="2.46">2.46 mm</quantity>
. Basic dimensions 
<taxonomicName id="4C38B5EAFFF3FFDDAB5DFE63FE28826E" authorityName="Y. Shibata" authorityYear="1993" box="[303,443,502,523]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B94C127BFFF3FFDDAB5DFE63FEAC826E" box="[303,319,502,523]" italics="true" pageId="3" pageNumber="143">C</emphasis>
. 
<emphasis id="B94C127BFFF3FFDDAB3FFE63FE28826E" box="[333,443,502,523]" italics="true" pageId="3" pageNumber="143">formosanus</emphasis>
</taxonomicName>
(
<specimenCount id="9D3E05E0FFF3FFDDABB8FE66FE44826E" box="[458,471,499,523]" count="1" pageId="3" pageNumber="143" type="female">♀</specimenCount>
, 
<typeStatus id="548370CBFFF3FFDDAB97FE63FDA9826E" box="[485,570,502,523]" pageId="3" pageNumber="143" type="paratype">Paratype</typeStatus>
, 
<figureCitation id="1303D2ECFFF3FFDDA837FE63FD13826E" box="[581,640,502,524]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="3" pageNumber="143">Fig. 2</figureCitation>
): body length 8.0 mm; length of pronotum 
<quantity id="4CC0638CFFF3FFDDAB93FD84FDAF8243" box="[481,572,529,550]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="143" unit="mm" value="1.23">1.23 mm</quantity>
; maximum width of pronotum 
<quantity id="4CC0638CFFF3FFDDAB6FFDBEFEE18224" box="[285,370,555,577]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="143" unit="mm" value="1.52">1.52 mm</quantity>
, length of elytra 
<quantity id="4CC0638CFFF3FFDDA87FFDBEFDF08224" box="[525,611,555,577]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" pageId="3" pageNumber="143" unit="mm" value="1.96">1.96 mm</quantity>
; maximum width of elytra 
<quantity id="4CC0638CFFF3FFDDAB6CFDD3FEEF823E" box="[286,380,582,604]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="3" pageNumber="143" unit="mm" value="2.15">2.15 mm</quantity>
. Basic dimensions 
<taxonomicName id="4C38B5EAFFF3FFDDA837FDD3FD45823E" authorityName="Y. Shibata" authorityYear="1993" box="[581,726,582,603]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B94C127BFFF3FFDDA837FDD3FDC6823E" box="[581,597,582,603]" italics="true" pageId="3" pageNumber="143">C</emphasis>
. 
<emphasis id="B94C127BFFF3FFDDA815FDD3FD45823E" box="[615,726,582,603]" italics="true" pageId="3" pageNumber="143">formosanus</emphasis>
</taxonomicName>
(
<specimenCount id="9D3E05E0FFF3FFDDA898FDD6FC93823E" box="[746,768,579,603]" count="1" pageId="3" pageNumber="143" type="male">♂</specimenCount>
, 
<typeStatus id="548370CBFFF3FFDDAACFFDF4FE8B8212" box="[189,280,609,631]" pageId="3" pageNumber="143" type="holotype">Holotype</typeStatus>
, 
<figureCitation id="1303D2ECFFF3FFDDAB51FDF4FECC8213" box="[291,351,609,631]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="3" pageNumber="143">Fig. 3</figureCitation>
): body length 7, 
<quantity id="4CC0638CFFF3FFDDA873FDF4FDAD8213" box="[513,574,609,630]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="143" unit="mm" value="7.0">7 mm</quantity>
; length of pronotum 
<quantity id="4CC0638CFFF3FFDDAACFFDEEFE8582F4" box="[189,278,635,657]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="3" pageNumber="143" unit="mm" value="1.24">1.24 mm</quantity>
; maximum width of pronotum 
<quantity id="4CC0638CFFF3FFDDA838FDEEFD3082F4" box="[586,675,635,657]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="3" pageNumber="143" unit="mm" value="1.54">1.54 mm</quantity>
, length of elytra 
<quantity id="4CC0638CFFF3FFDDAA88FD03FEC082CE" box="[250,339,662,684]" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="3" pageNumber="143" unit="mm" value="1.86">1.86 mm</quantity>
; maximum width of elytra 2.00 mm. From a rather large species of the genus and widely known in Europe 
<taxonomicName id="4C38B5EAFFF3FFDDAACFFD59FEA68285" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[189,309,715,737]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis id="B94C127BFFF3FFDDAACFFD59FEA68285" box="[189,309,715,737]" italics="true" pageId="3" pageNumber="143">C. striatulus</emphasis>
</taxonomicName>
, also well differs by significantly larger sizes: Basic dimensions 
<taxonomicName id="4C38B5EAFFF3FFDDAB1EFD73FE70829E" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[364,483,742,763]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis id="B94C127BFFF3FFDDAB1EFD73FEEF829E" box="[364,380,742,763]" italics="true" pageId="3" pageNumber="143">C</emphasis>
. 
<emphasis id="B94C127BFFF3FFDDABF8FD73FE70829E" box="[394,483,742,763]" italics="true" pageId="3" pageNumber="143">striatulus</emphasis>
</taxonomicName>
(
<specimenCount id="9D3E05E0FFF3FFDDAB81FD76FD93829E" box="[499,512,739,763]" count="1" pageId="3" pageNumber="143" type="female">♀</specimenCount>
, 
<collectingCountry id="F32F8EF9FFF3FFDDA87FFD73FDDC8299" box="[525,591,742,764]" name="Russia" pageId="3" pageNumber="143">Russia</collectingCountry>
, 
<collectingRegion id="49FC008BFFF3FFDDA828FD73FD288299" box="[602,699,742,764]" country="Russia" name="Smolensk" pageId="3" pageNumber="143">Smolensk</collectingRegion>
. 
<figureCitation id="1303D2ECFFF3FFDDA8B7FD73FC93829E" box="[709,768,742,764]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="3" pageNumber="143">Fig. 4</figureCitation>
) body length 7.0 mm; length of pronotum 
<quantity id="4CC0638CFFF3FFDDA837FC94FD0E8373" box="[581,669,769,790]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="3" pageNumber="143" unit="mm" value="1.13">1.13 mm</quantity>
; maximum width of pronotum 
<quantity id="4CC0638CFFF3FFDDAB07FC8EFE5F8354" box="[373,460,795,817]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="3" pageNumber="143" unit="mm" value="1.32">1.32 mm</quantity>
, length of elytra 
<quantity id="4CC0638CFFF3FFDDA81CFC8EFD568354" box="[622,709,795,817]" metricMagnitude="-3" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="143" unit="mm" value="1.72">1.72 mm</quantity>
; maximum width of elytra 
<quantity id="4CC0638CFFF3FFDDABFEFCA3FE76832E" box="[396,485,822,844]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="3" pageNumber="143" unit="mm" value="1.62">1.62 mm</quantity>
. The new species well differs from 
<taxonomicName id="4C38B5EAFFF3FFDDAA80FCC4FEEC8303" authorityName="Y. Shibata" authorityYear="1993" box="[242,383,849,870]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B94C127BFFF3FFDDAA80FCC4FEEC8303" box="[242,383,849,870]" italics="true" pageId="3" pageNumber="143">C. formosanus</emphasis>
</taxonomicName>
by more delicate and fine punctation of head, pronotum, and elytra, and by less deep striae on the elytra (
<figureCitation id="1303D2ECFFF3FFDDAAB1FC13FE8A83FE" box="[195,281,902,924]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="3" pageNumber="143">Figs 1–3</figureCitation>
). The new species well differs from 
<taxonomicName id="4C38B5EAFFF3FFDDA809FC13FC9483FE" authorityName="Y. Shibata" authorityYear="1993" box="[635,775,902,923]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B94C127BFFF3FFDDA809FC13FC9483FE" box="[635,775,902,923]" italics="true" pageId="3" pageNumber="143">C. formosanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C38B5EAFFF3FFDDAA97FC34FEC883D3" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[229,347,929,950]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis id="B94C127BFFF3FFDDAA97FC34FEC883D3" box="[229,347,929,950]" italics="true" pageId="3" pageNumber="143">C. striatulus</emphasis>
</taxonomicName>
by the wide, only slightly longer than maximum broad unpaired rhomboid impression in center of pronotal disc; in 
<taxonomicName id="4C38B5EAFFF3FFDDAB50FC43FE3F838E" authorityName="Y. Shibata" authorityYear="1993" box="[290,428,982,1004]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="formosanus">
C. 
<emphasis id="B94C127BFFF3FFDDAB4FFC43FE3F838E" box="[317,428,982,1003]" italics="true" pageId="3" pageNumber="143">formosanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C38B5EAFFF3FFDDABAAFC43FDDF838E" baseAuthorityName="Fabricius" baseAuthorityYear="1792" box="[472,588,982,1003]" class="Insecta" family="Staphylinidae" genus="Coprophilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="143" phylum="Arthropoda" rank="species" species="striatulus">
<emphasis id="B94C127BFFF3FFDDABAAFC43FDDF838E" box="[472,588,982,1003]" italics="true" pageId="3" pageNumber="143">C. striatulus</emphasis>
</taxonomicName>
impression is much narrower and more elongated, more than 2 times longer as broad (
<figureCitation id="1303D2ECFFF3FFDDAB72FB9EFEC48445" box="[256,343,1035,1057]" captionStart="Figs 1–4" captionStartId="2.[226,263,1698,1722]" captionTargetBox="[191,1938,608,1672]" captionTargetId="figure-108@2.[188,1942,584,1687]" captionTargetPageId="2" captionText="Figs 1–4. Coprophilus spp., dorsal view: 1 — C. (s. str.) maximus sp.n., holotype, ♀; 2–3 — C. (s. str.) formosanus (2 — paratype, ♀; 3 — holotype, ♂); 4 — C. (s. str.) striatulus, ♀, Russia, Smolensk. Scale bar — 1.0 mm. Рис. 1–4. Coprophilus spp., сверху: 1 — C. (s. str.) maximus sp.n., голотип, ♀; 2–3 — C. (s. str.) formosanus (2 — паратип, ♀, 3 — голотип, ♂); 4 — C. (s. str.) striatulus, ♀, РоссиЯ, Смоленск. МасШтаб — 1,0 мм." figureDoi="http://doi.org/10.5281/zenodo.13179488" httpUri="https://zenodo.org/record/13179488/files/figure.png" pageId="3" pageNumber="143">Figs 1–4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3229DE2FFF3FFDDAA90FBB3FD50845E" box="[226,707,1062,1084]" pageId="3" pageNumber="143" type="distribution">
<paragraph id="8B87CE69FFF3FFDDAA90FBB3FD50845E" blockId="3.[189,775,315,1137]" box="[226,707,1062,1084]" pageId="3" pageNumber="143">
DISTRIBUTION. North 
<collectingCountry id="F32F8EF9FFF3FFDDABAAFBB3FD9F8459" box="[472,524,1062,1084]" name="India" pageId="3" pageNumber="143">India</collectingCountry>
, Uttaranchal state.
</paragraph>
</subSubSection>
<subSubSection id="C3229DE2FFF3FFDDAA90FBD4FC5881D9" pageId="3" pageNumber="143" type="etymology">
<paragraph id="8B87CE69FFF3FFDDAA90FBD4FDD58414" blockId="3.[189,775,315,1137]" pageId="3" pageNumber="143">ETYMOLOGY. From Latin “maximum” (greatest); the name is given due to the very large size.</paragraph>
<paragraph id="8B87CE69FFF3FFDDAA90FB05FC9484A4" blockId="3.[189,775,1168,1217]" pageId="3" pageNumber="143">
<emphasis id="B94C127BFFF3FFDDAA90FB05FE2384C3" bold="true" box="[226,432,1168,1190]" pageId="3" pageNumber="143">Acknowledgements.</emphasis>
The author wishes to thank all colleagues for making material available for study: Michael
</paragraph>
<paragraph id="8B87CE69FFF3FFDDA95EFEAEFC5881D9" blockId="3.[812,1399,315,444]" pageId="3" pageNumber="143">
<materialsCitation id="3B50C434FFF3FFDDA95EFEAEFC5B81D9" collectionCode="TULE" country="Russia" county="Pedagogical State University" location="Daugavpils University Institute of Systematic Biology" municipality="Kirill Makarov" pageId="3" pageNumber="143" specimenCount="1" stateProvince="Moscow">
Schülke (Berlin), Yasutoshi Shibata (Machida, Tokyo),Alexey Shavrin (
<location id="8EE798B2FFF3FFDDA9F6FEC3FCC081E3" LSID="urn:lsid:plazi:treatment:03917F7FFFF1FFDDA922FE1AFC5881D9:8EE798B2FFF3FFDDA9F6FEC3FCC081E3" country="Russia" county="Pedagogical State University" municipality="Kirill Makarov" name="Daugavpils University Institute of Systematic Biology" pageId="3" pageNumber="143" stateProvince="Moscow">Daugavpils University Institute of Systematic Biology</location>
), Tadashi Ishikawa (
<collectionCode id="ED2956ACFFF3FFDDAE69FEE4FBCD81E3" box="[1051,1118,369,390]" country="Japan" httpUri="http://grbio.org/cool/fyp0-sb9p" name="Tokyo University of Agriculture and Technology" pageId="3" pageNumber="143">TULE</collectionCode>
). I also thank 
<collectingMunicipality id="6BE35413FFF3FFDDAE97FEE4FAE581E2" box="[1253,1398,369,391]" pageId="3" pageNumber="143">Kirill Makarov</collectingMunicipality>
for taking the photographs (
<collectingRegion id="49FC008BFFF3FFDDAE49FE1EFB0381C5" box="[1083,1168,395,416]" country="Russia" name="Moscow" pageId="3" pageNumber="143">Moscow</collectingRegion>
<collectingCounty id="62E6B6E5FFF3FFDDAEE4FE1EFCE381D9" pageId="3" pageNumber="143">Pedagogical State University</collectingCounty>
, 
<collectingCountry id="F32F8EF9FFF3FFDDA90EFE33FC5281D9" box="[892,961,422,444]" name="Russia" pageId="3" pageNumber="143">Russia</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>