251 lines
33 KiB
XML
251 lines
33 KiB
XML
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<mods:title id="EBB042C8CD1736B1ABCC87696805806D">Trematochampsa taqueti as a nomen dubium and the crocodyliform diversity of the Upper Cretaceous In Beceten Formation of Niger</mods:title>
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<treatment id="9C3D87E6FF8BFFFEB747F9CE72093BF2" ID-DOI="http://doi.org/10.5281/zenodo.4725745" ID-GBIF-Taxon="181753737" ID-Zenodo-Dep="4725745" LSID="urn:lsid:plazi:treatment:9C3D87E6FF8BFFFEB747F9CE72093BF2" httpUri="http://treatment.plazi.org/id/9C3D87E6FF8BFFFEB747F9CE72093BF2" lastPageId="5" lastPageNumber="664" pageId="3" pageNumber="662">
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<subSubSection id="5C8E657BFF8BFFF8B747F9CE754E3B15" box="[870,1399,1605,1630]" pageId="3" pageNumber="662" type="nomenclature">
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<paragraph id="142B36F0FF8BFFF8B747F9CE754E3B15" blockId="3.[870,1399,1605,1630]" box="[870,1399,1605,1630]" pageId="3" pageNumber="662">
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<heading id="4F63819CFF8BFFF8B747F9CE754E3B15" box="[870,1399,1605,1630]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="662" reason="2">
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<taxonomicName id="D3944D73FF8BFFF8B747F9CE754E3B15" authority="TAQUETI BUFFETAUT, 1974" authorityName="TAQUETI BUFFETAUT" authorityYear="1974" box="[870,1399,1605,1630]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="3" pageNumber="662" phylum="Chordata" rank="genus">
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<emphasis id="26E0EAE2FF8BFFF8B747F9CE749F3B17" box="[870,1190,1605,1628]" italics="true" pageId="3" pageNumber="662">TREMATOCHAMPSA TAQUETI</emphasis>
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BUFFETAUT, 1974
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5C8E657BFF8BFFF8B008F9E075AD3BE2" pageId="3" pageNumber="662" type="description">
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<paragraph id="142B36F0FF8BFFF8B008F9E0748A3BCF" blockId="3.[1065,1203,1643,1668]" box="[1065,1203,1643,1668]" pageId="3" pageNumber="662">
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(
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<figureCitation id="8CAF2A75FF8BFFF8B013F9E074923BCF" box="[1074,1195,1643,1668]" captionStart="Figure 2" captionStartId="4.[145,224,1047,1069]" captionTargetBox="[147,1425,196,1007]" captionTargetId="figure-414@4.[145,1425,195,1008]" captionTargetPageId="4" captionText="Figure 2. Trematochampsa taqueti. Original cranial reconstruction modified from Buffetaut (1976) in (A) lateral and (B) dorsal views. Right lacrimal Ibc 231, holotype of T. taqueti in (C) lateral, (D) posterior and (E) medial view. F, G, H, outline drawings corresponding to each view. Montealtosuchus arrudacamposi in (F) left lateral and (K) close-up on the lacrimal region. J, L, outline drawings corresponding to each view. Abbreviations: antfe, antorbital fenestra; antfo, antorbital fossa; fo, foramen; j.s, suture for jugal; m.s, maxilla suture; mt, maxilla tooth; pf.s, suture for prefrontal. Scales are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5714644" httpUri="https://zenodo.org/record/5714644/files/figure.png" pageId="3" pageNumber="662">FIG. 2A–H</figureCitation>
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)
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</paragraph>
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<paragraph id="142B36F0FF8BFFF8B71AF91F75AD3BE2" blockId="3.[827,1428,1684,1706]" box="[827,1428,1684,1706]" pageId="3" pageNumber="662">
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<emphasis id="26E0EAE2FF8BFFF8B71AF91F739C3BE2" box="[827,933,1684,1705]" italics="true" pageId="3" pageNumber="662">
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<typeStatus id="CB2F8852FF8BFFF8B71AF91F73A63BE2" box="[827,927,1684,1705]" pageId="3" pageNumber="662" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Ibc 231, isolated right lacrimal (
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<figureCitation id="8CAF2A75FF8BFFF8B138F91F75B53BE2" box="[1305,1420,1684,1706]" captionStart="Figure 2" captionStartId="4.[145,224,1047,1069]" captionTargetBox="[147,1425,196,1007]" captionTargetId="figure-414@4.[145,1425,195,1008]" captionTargetPageId="4" captionText="Figure 2. Trematochampsa taqueti. Original cranial reconstruction modified from Buffetaut (1976) in (A) lateral and (B) dorsal views. Right lacrimal Ibc 231, holotype of T. taqueti in (C) lateral, (D) posterior and (E) medial view. F, G, H, outline drawings corresponding to each view. Montealtosuchus arrudacamposi in (F) left lateral and (K) close-up on the lacrimal region. J, L, outline drawings corresponding to each view. Abbreviations: antfe, antorbital fenestra; antfo, antorbital fossa; fo, foramen; j.s, suture for jugal; m.s, maxilla suture; mt, maxilla tooth; pf.s, suture for prefrontal. Scales are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5714644" httpUri="https://zenodo.org/record/5714644/files/figure.png" pageId="3" pageNumber="662">Fig. 2C–H</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="5C8E657BFF8BFFFFB772F94972DF3829" lastPageId="4" lastPageNumber="663" pageId="3" pageNumber="662" type="discussion">
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<paragraph id="142B36F0FF8BFFFFB772F94972DF3829" blockId="3.[827,1443,1730,1906]" lastBlockId="4.[145,761,1234,1900]" lastPageId="4" lastPageNumber="663" pageId="3" pageNumber="662">
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<emphasis id="26E0EAE2FF8BFFF8B772F94973FF3B9C" box="[851,966,1730,1751]" italics="true" pageId="3" pageNumber="662">Reducing</emphasis>
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<taxonomicName id="D3944D73FF8BFFF8B7F0F94975353B93" authorityName="Buffetaut" authorityYear="1974" box="[977,1292,1730,1752]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="3" pageNumber="662" phylum="Chordata" rank="species" species="taqueti">Trematochampsa taqueti</taxonomicName>
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<emphasis id="26E0EAE2FF8BFFF8B136F94873A33BBD" italics="true" pageId="3" pageNumber="662">to a nomen dubium:</emphasis>
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The type of
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<taxonomicName id="D3944D73FF8BFFF8B003F96974B23BBC" authorityName="Buffetaut" authorityYear="1974" box="[1058,1163,1762,1783]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="3" pageNumber="662" phylum="Chordata" rank="species" species="taqueti">
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<emphasis id="26E0EAE2FF8BFFF8B003F96974B23BBC" box="[1058,1163,1762,1783]" italics="true" pageId="3" pageNumber="662">T. taqueti</emphasis>
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</taxonomicName>
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is an isolated right lacrimal, Ibc 231 (
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<figureCitation id="8CAF2A75FF8BFFF8B7EDF88B74023A5E" box="[972,1083,1792,1814]" captionStart="Figure 2" captionStartId="4.[145,224,1047,1069]" captionTargetBox="[147,1425,196,1007]" captionTargetId="figure-414@4.[145,1425,195,1008]" captionTargetPageId="4" captionText="Figure 2. Trematochampsa taqueti. Original cranial reconstruction modified from Buffetaut (1976) in (A) lateral and (B) dorsal views. Right lacrimal Ibc 231, holotype of T. taqueti in (C) lateral, (D) posterior and (E) medial view. F, G, H, outline drawings corresponding to each view. Montealtosuchus arrudacamposi in (F) left lateral and (K) close-up on the lacrimal region. J, L, outline drawings corresponding to each view. Abbreviations: antfe, antorbital fenestra; antfo, antorbital fossa; fo, foramen; j.s, suture for jugal; m.s, maxilla suture; mt, maxilla tooth; pf.s, suture for prefrontal. Scales are 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5714644" httpUri="https://zenodo.org/record/5714644/files/figure.png" pageId="3" pageNumber="662">Fig. 2C–H</figureCitation>
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). We propose the taxon
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<taxonomicName id="D3944D73FF8BFFF8B11DF88A759A3A5E" authorityName="Buffetaut" authorityYear="1974" box="[1340,1443,1792,1814]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="3" pageNumber="662" phylum="Chordata" rank="species" species="taqueti">
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<emphasis id="26E0EAE2FF8BFFF8B11DF88A759A3A5E" box="[1340,1443,1792,1814]" italics="true" pageId="3" pageNumber="662">T. taqueti</emphasis>
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</taxonomicName>
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be reduced to a
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<emphasis id="26E0EAE2FF8BFFF8B7D1F89474A23A78" box="[1008,1179,1822,1844]" italics="true" pageId="3" pageNumber="662">nomen dubium</emphasis>
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based on the following description and discussion of this specimen that can find no morphologies to diagnose this taxon from other known crocodyliforms. Furthermore, because the type is a single isolated element, neither associated nor confidently combined with any other material in the In Becetin crocodyliform collection, nothing from this collection can be assigned to
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<taxonomicName id="D3944D73FF8CFFFFB5F4FAC672063829" authorityName="Buffetaut" authorityYear="1974" box="[469,575,1357,1378]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="taqueti">
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<emphasis id="26E0EAE2FF8CFFFFB5F4FAC672063829" box="[469,575,1357,1378]" italics="true" pageId="4" pageNumber="663">T. taqueti</emphasis>
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</taxonomicName>
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as a
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<typeStatus id="CB2F8852FF8CFFFFB658FAC772D83829" box="[633,737,1356,1378]" pageId="4" pageNumber="663" type="lectotype">lectotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<caption id="40EB6678FF8CFFFFB4B0FB9C750D39E8" ID-DOI="http://doi.org/10.5281/zenodo.5714644" ID-Zenodo-Dep="5714644" httpUri="https://zenodo.org/record/5714644/files/figure.png" pageId="4" pageNumber="663" startId="4.[145,224,1047,1069]" targetBox="[147,1425,196,1007]" targetPageId="4">
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<paragraph id="142B36F0FF8CFFFFB4B0FB9C750D39E8" blockId="4.[145,1425,1047,1187]" pageId="4" pageNumber="663">
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<emphasis id="26E0EAE2FF8CFFFFB4B0FB9C70C33967" bold="true" box="[145,250,1047,1069]" pageId="4" pageNumber="663">Figure 2.</emphasis>
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<taxonomicName id="D3944D73FF8CFFFFB524FB9372303965" authorityName="Buffetaut" authorityYear="1974" box="[261,521,1048,1070]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="taqueti">
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<emphasis id="26E0EAE2FF8CFFFFB524FB9372303965" box="[261,521,1048,1070]" italics="true" pageId="4" pageNumber="663">Trematochampsa taqueti</emphasis>
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</taxonomicName>
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. Original cranial reconstruction modified from
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<bibRefCitation id="70054B01FF8CFFFFB7DFFB9374953965" author="Buffetaut E" box="[1022,1196,1048,1070]" pageId="4" pageNumber="663" pagination="143 - 198" refId="ref13654" refString="Buffetaut E. 1976. Osteologie et affinities de Trematochampsa taqueti (Crocodylia, Mesosuchia) du Senonien inferieur d'In Beceten (Republique du Niger). Geobios 9: 143 - 198." type="journal article" year="1976">Buffetaut (1976)</bibRefCitation>
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in (A) lateral and (B) dorsal views. Right lacrimal Ibc 231, holotype of
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<taxonomicName id="D3944D73FF8CFFFFB6AFFBBD72C83900" authorityName="Buffetaut" authorityYear="1974" box="[654,753,1077,1099]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="taqueti">
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<emphasis id="26E0EAE2FF8CFFFFB6AFFBBD72C83900" box="[654,753,1077,1099]" italics="true" pageId="4" pageNumber="663">T. taqueti</emphasis>
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</taxonomicName>
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in (C) lateral, (D) posterior and (E) medial view. F, G, H, outline drawings corresponding to each view.
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<taxonomicName id="D3944D73FF8CFFFFB603FBD973453923" box="[546,892,1106,1128]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="arrudacamposi">
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<emphasis id="26E0EAE2FF8CFFFFB603FBD973453923" box="[546,892,1106,1128]" italics="true" pageId="4" pageNumber="663">Montealtosuchus arrudacamposi</emphasis>
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</taxonomicName>
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in (F) left lateral and (K) close-up on the lacrimal region. J, L, outline drawings corresponding to each view. Abbreviations: antfe, antorbital fenestra; antfo, antorbital fossa; fo, foramen; j.s, suture for jugal; m.s, maxilla suture; mt, maxilla tooth; pf.s, suture for prefrontal. Scales are 1 cm.
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</paragraph>
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</caption>
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<subSubSection id="5C8E657BFF8CFFFFB488FAE0727B3BDF" pageId="4" pageNumber="663" type="materials_examined">
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<paragraph id="142B36F0FF8CFFFFB488FAE0727B3BDF" blockId="4.[145,761,1234,1900]" pageId="4" pageNumber="663">
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The lacrimal Ibc 231 was described by Buffetaut in 1974 and 1976. The original diagnostic morphologies of
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<taxonomicName id="D3944D73FF8CFFFFB48AFA22712C38F5" authorityName="Buffetaut" authorityYear="1974" box="[171,277,1449,1470]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="taqueti">
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<emphasis id="26E0EAE2FF8CFFFFB48AFA22712C38F5" box="[171,277,1449,1470]" italics="true" pageId="4" pageNumber="663">T. taqueti</emphasis>
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</taxonomicName>
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that were present on the
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<typeStatus id="CB2F8852FF8CFFFFB617FA2372AE38F5" box="[566,663,1448,1470]" pageId="4" pageNumber="663" type="holotype">holotype</typeStatus>
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are only the presence of a large antorbital fenestra on the lacrimal and maxilla and, by inference, a relatively small to medium body size (
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<quantity id="D36C9B15FF8CFFFFB54FF98F718D3B51" box="[366,436,1540,1562]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="4" pageNumber="663" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1–3 m</quantity>
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) of the animal. The fenestra is relatively small, but the fossa is large. Both these morphologies are present in many small- to mediumsized Crocodyliformes and do not constitute unique diagnostic morphologies for
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<taxonomicName id="D3944D73FF8CFFFFB5F1F90B72023BDE" authorityName="Buffetaut" authorityYear="1974" box="[464,571,1664,1685]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="taqueti">
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<emphasis id="26E0EAE2FF8CFFFFB5F1F90B72023BDE" box="[464,571,1664,1685]" italics="true" pageId="4" pageNumber="663">T. taqueti</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="5C8E657BFF8CFFFEB488F91572CC3F67" lastPageId="5" lastPageNumber="664" pageId="4" pageNumber="663" type="discussion">
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<paragraph id="142B36F0FF8CFFFFB488F91575AB3A65" blockId="4.[145,761,1234,1900]" lastBlockId="4.[809,1426,1234,1899]" pageId="4" pageNumber="663">
|
||
Anteriorly, Ibc 231 has a large antorbital fossa that expands well beyond a small antorbital fenestra that is defined by a small circular edge on the anterodorsal margin of the lacrimal. Although the entire fenestra is not preserved because it would have been bounded also by the maxilla, the acute circumference of the bor- der on the lacrimal suggests the fenestra was small. The fossa expands ventrally to the maxilla contact and dorsally to a broken edge. The ventral edge of the fossa is marked by a low but distinct near horizontal ridge that may have been the external limit to the maxillary contact. Below this ridge, the lacrimal is relatively smooth and lightly textured, possibly to contact a thin superficial overlap from the maxilla. A similar ventral antorbital fenestra morphology is present in
|
||
<taxonomicName id="D3944D73FF8CFFFFB708FA4C74773897" box="[809,1102,1479,1500]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="terrificus">
|
||
<emphasis id="26E0EAE2FF8CFFFFB708FA4C74773897" box="[809,1102,1479,1500]" italics="true" pageId="4" pageNumber="663">Uberabasuchus terrificus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3944D73FF8CFFFFB0A8FA4C739F38B0" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="arrudacamposi">
|
||
<emphasis id="26E0EAE2FF8CFFFFB0A8FA4C739F38B0" italics="true" pageId="4" pageNumber="663">Montealtosuchus arrudacamposi</emphasis>
|
||
</taxonomicName>
|
||
. This morphology suggests the antorbital fossa extended ventrally onto the maxilla. The remain- der of the external surface of the lacrimal is heavily ornamented anteroventrally with pitting and sulci extending onto the orbital margin. The posterior margin of the antorbital fenestra is sharply edged with the fossa placed deep from the lacrimal superficial surface, creating a sharp bony roof on the posterior margin of the fossa. This
|
||
<typeStatus id="CB2F8852FF8CFFFFB7F1F95074383BBB" box="[976,1025,1755,1776]" pageId="4" pageNumber="663">type</typeStatus>
|
||
of deeply inset antorbital fossa and ornamented orbital margin of the lacrimal is only otherwise present in
|
||
<taxonomicName id="D3944D73FF8CFFFFB7D3F892744E3A66" box="[1010,1143,1816,1838]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="terrificus">
|
||
<emphasis id="26E0EAE2FF8CFFFFB7D3F892744E3A66" box="[1010,1143,1816,1838]" italics="true" pageId="4" pageNumber="663">U. terrificus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3944D73FF8CFFFFB08EF89275B23A66" box="[1199,1419,1816,1838]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="663" phylum="Chordata" rank="species" species="arrudacamposi">
|
||
<emphasis id="26E0EAE2FF8CFFFFB08EF89275B23A66" box="[1199,1419,1816,1838]" italics="true" pageId="4" pageNumber="663">Mo. arrudacamposi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="142B36F0FF8CFFFEB760F8BC72CC3F67" blockId="4.[809,1426,1234,1899]" lastBlockId="5.[163,781,197,1722]" lastPageId="5" lastPageNumber="664" pageId="4" pageNumber="663">
|
||
The posteroventral corner of the lacrimal has a large, complex, ovate suture that contacted the jugal. The robust jugal contact posteroventrally and slender maxilla contact anteroventrally is shared with many Crocodyliformes, such as
|
||
<taxonomicName id="D3944D73FF8DFFFEB5EAFE89724A3C5C" box="[459,627,258,279]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="genus">
|
||
<emphasis id="26E0EAE2FF8DFFFEB5EAFE89724A3C5C" box="[459,627,258,279]" italics="true" pageId="5" pageNumber="664">Araripesuchus</emphasis>
|
||
</taxonomicName>
|
||
(all species), Sebecia and
|
||
<taxonomicName id="D3944D73FF8DFFFEB512FEAA71943C7C" authorityName="Clark" authorityYear="1988" box="[307,429,289,311]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neosuchia" pageId="5" pageNumber="664" phylum="Chordata" rank="order">Neosuchia</taxonomicName>
|
||
. However, many notosuchians, including
|
||
<taxonomicName id="D3944D73FF8DFFFEB533FECB718C3C1E" authorityName="Gomani" authorityYear="1997" box="[274,437,320,341]" class="Reptilia" family="Itasuchidae" genus="Malawisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="genus">
|
||
<emphasis id="26E0EAE2FF8DFFFEB533FECB718C3C1E" box="[274,437,320,341]" italics="true" pageId="5" pageNumber="664">Malawisuchus</emphasis>
|
||
</taxonomicName>
|
||
and derived notosuchians (following
|
||
<bibRefCitation id="70054B01FF8DFFFEB4D6FED5719A3C3F" author="Pol D & Nascimento PM & Carvalho AB & Riccomini C & Domingues RA & Zaher H" box="[247,419,350,372]" pageId="5" pageNumber="664" pagination="93105" refId="ref14548" refString="Pol D, Nascimento PM, Carvalho AB, Riccomini C, Pires- Domingues RA, Zaher H. 2014. A new notosuchian from the Late Cretaceous of Brazil and the phylogeny of advanced notosuchians. PLoS ONE 9: e 93105." type="journal article" year="2014">
|
||
Pol
|
||
<emphasis id="26E0EAE2FF8DFFFEB502FED471643C38" box="[291,349,350,372]" italics="true" pageId="5" pageNumber="664">et al.</emphasis>
|
||
, 2014
|
||
</bibRefCitation>
|
||
), lack a lacrimal–jugal contact implying no relationship to these taxa. The jugal and maxilla contact at the lacrimal of
|
||
<taxonomicName id="D3944D73FF8DFFFEB60BFE1772AF3CFA" authorityName="Buffetaut" authorityYear="1974" box="[554,662,412,433]" class="Reptilia" family="Trematochampsidae" genus="Trematochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="taqueti">
|
||
<emphasis id="26E0EAE2FF8DFFFEB60BFE1772AF3CFA" box="[554,662,412,433]" italics="true" pageId="5" pageNumber="664">T. taqueti</emphasis>
|
||
</taxonomicName>
|
||
compares closest to
|
||
<taxonomicName id="D3944D73FF8DFFFEB531FE3172253C9B" box="[272,540,442,464]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis id="26E0EAE2FF8DFFFEB531FE3172253C9B" box="[272,540,442,464]" italics="true" pageId="5" pageNumber="664">Araripesuchus wegeneri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D3944D73FF8DFFFEB609FE3072923C9B" box="[552,683,443,464]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="terrificus">
|
||
<emphasis id="26E0EAE2FF8DFFFEB609FE3072923C9B" box="[552,683,443,464]" italics="true" pageId="5" pageNumber="664">U. terrificus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3944D73FF8DFFFEB6C2FE3071763CA5" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="arrudacamposi">
|
||
<emphasis id="26E0EAE2FF8DFFFEB6C2FE3071763CA5" italics="true" pageId="5" pageNumber="664">Mo. arrudacamposi</emphasis>
|
||
</taxonomicName>
|
||
, with a robust butt lacrimal–jugal contact and a thin lacrimal–maxilla overlapping contact with the antorbital fossa extending onto the maxilla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5C8E657BFF8DFFFEB49AFDBE72523E91" pageId="5" pageNumber="664" type="description">
|
||
<paragraph id="142B36F0FF8DFFFEB49AFDBE72523E91" blockId="5.[163,781,197,1722]" pageId="5" pageNumber="664">Dorsolaterally, the lacrimal is deeply ornamented and forms a sharp posterior edge to the antorbital fossa. The fossa is smooth and the lacrimal portion of the fossa bounds the posteroventral corner of a small antorbital fenestra. The lateral surface of the lacrimal is relatively straight in posterior aspect. The robust prefrontal suture occupies much of the posterodorsal and posterodorsomedial surfaces of the lacrimal. Medially, a low, horizontal crest extends anteriorly from the base of the prefrontal contact and another smaller crest parallels this crest anterodorsally. A large lacrimal duct pierces the dorsal orbital margin and appears to have been bordered dorsally by the prefrontal. Two smaller foramina lie medial to the lacrimal duct.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5C8E657BFF8DFFFEB49AFC6872093BF2" pageId="5" pageNumber="664" type="discussion">
|
||
<paragraph id="142B36F0FF8DFFFEB49AFC6872093BF2" blockId="5.[163,781,197,1722]" pageId="5" pageNumber="664">
|
||
Many of these morphologies are present in
|
||
<taxonomicName id="D3944D73FF8DFFFEB690FC6870E1395D" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="terrificus">
|
||
<emphasis id="26E0EAE2FF8DFFFEB690FC6870E1395D" italics="true" pageId="5" pageNumber="664">U. terrificus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D3944D73FF8DFFFEB4C4FB8971FB395D" box="[229,450,1025,1047]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="arrudacamposi">
|
||
<emphasis id="26E0EAE2FF8DFFFEB4C4FB8971FB395D" box="[229,450,1025,1047]" italics="true" pageId="5" pageNumber="664">Mo. arrudacamposi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D3944D73FF8DFFFEB5EEFB8A7278395C" baseAuthorityName="Chiappe" baseAuthorityYear="1988" box="[463,577,1025,1047]" class="Reptilia" family="Trematochampsidae" genus="Amargasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="minor">
|
||
<emphasis id="26E0EAE2FF8DFFFEB5EEFB8A7278395C" box="[463,577,1025,1047]" italics="true" pageId="5" pageNumber="664">An. minor</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D3944D73FF8DFFFEB65AFB8A733C395C" box="[635,773,1025,1047]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis id="26E0EAE2FF8DFFFEB65AFB8A733C395C" box="[635,773,1025,1047]" italics="true" pageId="5" pageNumber="664">Ar. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
, but not in other
|
||
<taxonomicName id="D3944D73FF8DFFFEB57BFBAB726E397D" box="[346,599,1056,1078]" class="Reptilia" genus="Araripesuchus" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="26E0EAE2FF8DFFFEB57BFBAB71C5397E" box="[346,508,1056,1077]" italics="true" pageId="5" pageNumber="664">Araripesuchus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Specifically, the topology of bones surrounding the antorbital fenestra is shared with these taxa. Differences with
|
||
<taxonomicName id="D3944D73FF8DFFFEB67FFBD673333939" box="[606,778,1117,1138]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="genus">
|
||
<emphasis id="26E0EAE2FF8DFFFEB67FFBD673333939" box="[606,778,1117,1138]" italics="true" pageId="5" pageNumber="664">Uberabasuchus</emphasis>
|
||
</taxonomicName>
|
||
include the vertically oriented lacrimal edge to the antorbital fossa, as opposed to the curving, posteroventral margin in Ibc 231. Additionally, the orbital margin on the lacrimal of
|
||
<taxonomicName id="D3944D73FF8DFFFEB5BAFB53727039A6" box="[411,585,1240,1261]" class="Reptilia" family="Peirosauridae" genus="Uberabasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="genus">
|
||
<emphasis id="26E0EAE2FF8DFFFEB5BAFB53727039A6" box="[411,585,1240,1261]" italics="true" pageId="5" pageNumber="664">Uberabasuchus</emphasis>
|
||
</taxonomicName>
|
||
is linear with an angular dorsal margin, whereas the orbital margin in Ibc 231 is gently concave. Differences with
|
||
<taxonomicName id="D3944D73FF8DFFFEB65DFA9E73323861" box="[636,779,1301,1322]" class="Reptilia" family="Uruguaysuchidae" genus="Anatosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="genus">
|
||
<emphasis id="26E0EAE2FF8DFFFEB65DFA9E73323861" box="[636,779,1301,1322]" italics="true" pageId="5" pageNumber="664">Anatosuchus</emphasis>
|
||
</taxonomicName>
|
||
include the lack of ornamentation and relatively narrow proportions in this taxon’s lacrimal bar. There are few differences with
|
||
<taxonomicName id="D3944D73FF8DFFFEB5B3FAFA722538CC" box="[402,540,1393,1415]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="wegeneri">
|
||
<emphasis id="26E0EAE2FF8DFFFEB5B3FAFA722538CC" box="[402,540,1393,1415]" italics="true" pageId="5" pageNumber="664">Ar. wegeneri</emphasis>
|
||
</taxonomicName>
|
||
, but one notable difference is the lack of a sharp-edged and roofing posteroventral margin of the antorbital fenestra. However, the posterior sharp and roofed edge of the antorbital fossa, ornamentation and orbital margin is nearly identical to
|
||
<taxonomicName id="D3944D73FF8DFFFEB482F98071B83B54" box="[163,385,1546,1568]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="arrudacamposi">
|
||
<emphasis id="26E0EAE2FF8DFFFEB482F98071B83B54" box="[163,385,1546,1568]" italics="true" pageId="5" pageNumber="664">Mo. arrudacamposi</emphasis>
|
||
</taxonomicName>
|
||
. In fact, close examination of both finds no diagnostic features to separate the two. All morphologies described above for Ibc 231 are also present in the lacrimal of
|
||
<taxonomicName id="D3944D73FF8DFFFEB5BBF9EC724B3B30" box="[410,626,1638,1660]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="664" phylum="Chordata" rank="species" species="arrudacamposi">
|
||
<emphasis id="26E0EAE2FF8DFFFEB5BBF9EC724B3B30" box="[410,626,1638,1660]" italics="true" pageId="5" pageNumber="664">Mo. arrudacamposi</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, in light of these more recent crocodyliform discoveries, we find no autapomorphies for Ibc 231.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |